124 lines
17 KiB
XML
124 lines
17 KiB
XML
<document id="50EAACA4342C27519865422C7FF2FC9A" ID="10.11646/zootaxa.4254.2.6SLASH10796" ID-CLB-Dataset="33235" ID-DOI="10.11646/zootaxa.4254.2.6" ID-GBIF-Dataset="bc040c13-0b57-44fc-9a7f-c236756231b3" ID-ISSN="1175-5326" ID-Zenodo-Dep="545901" ID-ZooBank="C8BA1773-6527-4533-9F61-01E421D47266" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1492070477963" checkinUser="plazi" docAuthor="Bilton, David T." docDate="2017" docId="03D87B73FFC9FFF240E191DAA78FFCA2" docLanguage="en" docName="zootaxa.4254.2.6SLASH10796.pdf" docOrigin="Zootaxa 4254 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Leielmis Deleve 1964" docType="treatment" docVersion="7" lastPageNumber="257" masterDocId="FFE1030BFFC8FFF04076920BA507FF99" masterDocTitle="A revision of the South African riffle beetle genus Leielmis Delève, 1964 (Coleoptera: Elmidae)" masterLastPageNumber="268" masterPageNumber="255" pageNumber="256" updateTime="1698438892865" updateUser="ExternalLinkService">
|
|
<mods:mods id="0A31F2EAA2405879A4FAF8736F682BE2" xmlns:mods="http://www.loc.gov/mods/v3">
|
|
<mods:titleInfo id="F5320C803872B4F5E8795C7B6B29D1EB">
|
|
<mods:title id="7D6E54E90FCD671ECA6293D8E86B0208">A revision of the South African riffle beetle genus Leielmis Delève, 1964 (Coleoptera: Elmidae)</mods:title>
|
|
</mods:titleInfo>
|
|
<mods:name id="DE73FA15D2D304442D9D68B3D9474936" type="personal">
|
|
<mods:role id="459CE391CC978120B7426C922F65D17C">
|
|
<mods:roleTerm id="728645F8E37EAE6753206795524F286E">Author</mods:roleTerm>
|
|
</mods:role>
|
|
<mods:namePart id="9462C9D17623FEE52562E6022602E5E0">Bilton, David T.</mods:namePart>
|
|
</mods:name>
|
|
<mods:typeOfResource id="F6AA0C7A9F3669AAF0D3495D70AD54E7">text</mods:typeOfResource>
|
|
<mods:relatedItem id="2BCEC8DE7B38342D999D7D5EA8AC8EC2" type="host">
|
|
<mods:titleInfo id="C8E72505A97A517FD494548F76392C4C">
|
|
<mods:title id="719178C667BFCCDB43619B577E5CCFFE">Zootaxa</mods:title>
|
|
</mods:titleInfo>
|
|
<mods:part id="E7656129565C674A361323BD5F164FB4">
|
|
<mods:date id="0CC21B3A28CB290D0AFD673A1ECE32B7">2017</mods:date>
|
|
<mods:detail id="4ED7F06444F4290BADDDDF44B37C8DBC" type="volume">
|
|
<mods:number id="B723F58F99BB358B07AFE088B01B03DF">4254</mods:number>
|
|
</mods:detail>
|
|
<mods:detail id="F8DC7D99C37B5AD248B297F39960D8C2" type="issue">
|
|
<mods:number id="2AEB66EDEB8C7D8CD679C3F5DCB78D5D">2</mods:number>
|
|
</mods:detail>
|
|
<mods:extent id="8371DB3DAE07600D5015E7AFE708762C" unit="page">
|
|
<mods:start id="56112FB8B74572CB8AB200EA62D016BB">255</mods:start>
|
|
<mods:end id="CA5C5335B8F0C1ABA1263D0C3779D3C8">268</mods:end>
|
|
</mods:extent>
|
|
</mods:part>
|
|
</mods:relatedItem>
|
|
<mods:classification id="3C249F29CC8D28F53164C4717E13BA72">journal article</mods:classification>
|
|
<mods:identifier id="FBA0EF6D57D7BE10AABE76D041CFB03B" type="CLB-Dataset">33235</mods:identifier>
|
|
<mods:identifier id="2F979C1D0D620161206C5E9A8CD95CC3" type="DOI">10.11646/zootaxa.4254.2.6</mods:identifier>
|
|
<mods:identifier id="EFF8501FB24BFFDB6BD1B476F47D838A" type="GBIF-Dataset">bc040c13-0b57-44fc-9a7f-c236756231b3</mods:identifier>
|
|
<mods:identifier id="4BA4BE16E7F5042A8A5B875191B05127" type="ISSN">1175-5326</mods:identifier>
|
|
<mods:identifier id="D04EE4CF571BD8860B10809A156554B1" type="Zenodo-Dep">545901</mods:identifier>
|
|
<mods:identifier id="57A4EB723FADE3921019C7AA724B7B56" type="ZooBank">C8BA1773-6527-4533-9F61-01E421D47266</mods:identifier>
|
|
</mods:mods>
|
|
<treatment id="03D87B73FFC9FFF240E191DAA78FFCA2" ID-DOI="http://doi.org/10.5281/zenodo.5999279" ID-GBIF-Taxon="128527411" ID-Zenodo-Dep="5999279" LSID="urn:lsid:plazi:treatment:03D87B73FFC9FFF240E191DAA78FFCA2" httpUri="http://treatment.plazi.org/id/03D87B73FFC9FFF240E191DAA78FFCA2" lastPageId="2" lastPageNumber="257" pageId="1" pageNumber="256">
|
|
<subSubSection id="C36B99EEFFC9FFF140E191DAA4A6FC72" box="[151,417,977,1003]" pageId="1" pageNumber="256" type="nomenclature">
|
|
<paragraph id="8BCECA65FFC9FFF140E191DAA4A6FC72" blockId="1.[151,417,977,1003]" box="[151,417,977,1003]" pageId="1" pageNumber="256">
|
|
<heading id="D0867D09FFC9FFF140E191DAA4A6FC72" bold="true" box="[151,417,977,1003]" fontSize="11" level="1" pageId="1" pageNumber="256" reason="1">
|
|
<taxonomicName id="4C71B1E6FFC9FFF140E191DAA4A6FC72" ID-CoL="5BV3" authority="Deleve, 1964" authorityName="Deleve" authorityYear="1964" box="[151,417,977,1003]" class="Insecta" family="Elmidae" genus="Leielmis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="256" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="B9051677FFC9FFF140E191DAA4A6FC72" bold="true" box="[151,417,977,1003]" pageId="1" pageNumber="256">
|
|
<emphasis id="B9051677FFC9FFF140E191DAA5FAFC72" bold="true" box="[151,253,977,1003]" italics="true" pageId="1" pageNumber="256">Leielmis</emphasis>
|
|
<bibRefCitation id="EFE0B794FFC9FFF1417291DAA4A6FC72" author="Deleve" box="[260,417,977,1003]" pageId="1" pageNumber="256" refString="Deleve, J. (1964) Contribution a l'etude des Dryopoidea. VIII. Cinq nouveaux genres d'Elminthidae (Coleoptera) de l'Afrique australe. Bulletin et Annales de la Societe Royale d'Enromologie de Belgique, 100, 155 - 176." type="journal article" year="1964">Delève, 1964</bibRefCitation>
|
|
</emphasis>
|
|
</taxonomicName>
|
|
</heading>
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="C36B99EEFFC9FFF140E19612A7A6FBB7" box="[151,673,1048,1070]" pageId="1" pageNumber="256" type="materials_examined">
|
|
<paragraph id="8BCECA65FFC9FFF140E19612A7A6FBB7" blockId="1.[151,673,1048,1070]" box="[151,673,1048,1070]" pageId="1" pageNumber="256">
|
|
Type species:
|
|
<taxonomicName id="4C71B1E6FFC9FFF141509613A7A6FBB7" authority="Grouvelle, 1890" authorityName="Grouvelle" authorityYear="1890" box="[294,673,1048,1070]" class="Insecta" family="Elmidae" genus="Helmis" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="256" phylum="Arthropoda" rank="species" species="georyssoides">
|
|
<emphasis id="B9051677FFC9FFF141509613A4F0FBB4" box="[294,503,1048,1069]" italics="true" pageId="1" pageNumber="256">Helmis georyssoides</emphasis>
|
|
<bibRefCitation id="EFE0B794FFC9FFF1418A9613A7A6FBB7" author="Grouvelle" box="[508,673,1048,1070]" pageId="1" pageNumber="256" refString="Grouvelle, A. (1890) [Diagnoses de trois Helmis nouveau]. Annales de la Societe Entomologique de France. Series 6. Fol. 10. Bulletin des Seances et Bulletin bibliographique de la Societe Entomologique de France, 1890 (XIV), CCXII." type="journal article" year="1890">Grouvelle, 1890</bibRefCitation>
|
|
</taxonomicName>
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="C36B99EEFFC9FFF240E19651A4EDFDFA" lastPageId="2" lastPageNumber="257" pageId="1" pageNumber="256" type="description">
|
|
<paragraph id="8BCECA65FFC9FFF140E19651A1C5FBEA" blockId="1.[151,1437,1114,2003]" box="[151,1218,1114,1139]" pageId="1" pageNumber="256">
|
|
<emphasis id="B9051677FFC9FFF140E19651A42EFBEA" bold="true" box="[151,297,1114,1139]" pageId="1" pageNumber="256">Description.</emphasis>
|
|
Body form (
|
|
<figureCitation id="134AD6E0FFC9FFF141CF9651A4FEFBEA" box="[441,505,1114,1139]" captionStart="FIGURE 1" captionStartId="3.[151,250,1980,2002]" captionTargetBox="[166,1414,644,1940]" captionTargetId="figure@3.[151,1436,625,1958]" captionTargetPageId="3" captionText="FIGURE 1. Leielmis species, male habitus. A) L. georyssoides; B) L. gibbosus sp. nov., holotype; C) L. hirsutus sp. nov., holotype. Scale bar = 1 mm." httpUri="https://zenodo.org/record/545902/files/figure.png" pageId="1" pageNumber="256">Fig. 1</figureCitation>
|
|
) short and convex, strongly narrowed both in front and behind.
|
|
</paragraph>
|
|
<paragraph id="8BCECA65FFC9FFF140B19675A021FA2E" blockId="1.[151,1437,1114,2003]" pageId="1" pageNumber="256">Head: Relatively narrow, partly retractable, orientated ventrally in life. Labrum short, broader than long and strongly arcuate at anterior margin. Clypeus distinctly broader and longer than labrum, convex, weakly arcuate at lateral margins, concave anteriorly. Frontoclypeal suture weakly arcuate. Antennae 11-segmented, all segments elongate, somewhat flattened dorsoventrally. Mandibles acuminate at apex, prostheca spinose. Maxillary palpi short, spinose, 4-segmented, dorsoventrally flattened, especially expanded segment 3. Palpifer well-developed, galea 2-segmented, long and narrow, flattened. Lacinia larger than galea, flattened, elongate, setose at apex, trapezoidal, with mesal and apical fringes of long setae. Labial palps short, 3-segmented, flattened dorsoventrally, second segment expanded; first segment much shorter than second; third shorter and narrower than second. Mentum transverse, broadly emarginate anteriorly. Ligula trapezoidal, broadly expanded anterolaterally.</paragraph>
|
|
<paragraph id="8BCECA65FFC9FFF140B197C8A13DF9DE" blockId="1.[151,1437,1114,2003]" pageId="1" pageNumber="256">Pronotum: Transverse, broadest at base, evenly narrowed to anterior margin; anterior margin approximately half width of posterior margin. Anterior margin arched, arcuate, hood-like, partly shielding head. Lateral margins strongly bordered. Sulci, carinae and gibbosities entirely absent; surface punctate and granulate, granules also present along lateral margins. Granules lost to varying degrees on older specimens.</paragraph>
|
|
<paragraph id="8BCECA65FFC9FFF140B19459A4F7F962" blockId="1.[151,1437,1114,2003]" pageId="1" pageNumber="256">Elytra: Slightly elongate, strongly bordered, widest approximately one third to one half behind shoulder, narrowing to apex, abruptly so in posterior third. Elytral apices cojointly acuminate. Each elytron with 9 puncture rows, moderately to weakly striate impressed in part; rows 7 and 8 confluent in anterior and posterior thirds. Intervals flat or slightly convex, punctate, with granules apicolaterally. Scutellum trapezoidal, front margin broadly arcuate, sides weakly rounded.</paragraph>
|
|
<paragraph id="8BCECA65FFC9FFF240B1950CA5EBFF61" blockId="1.[151,1437,1114,2003]" lastBlockId="2.[151,1437,151,827]" lastPageId="2" lastPageNumber="257" pageId="1" pageNumber="256">
|
|
<materialsCitation id="3B19C038FFC9FFF240B1950CA5EBFF61" ID-GBIF-Occurrence="1455889602" collectingDate="3 x" county="Venter" lastPageId="2" lastPageNumber="257" location="Prosternal" municipality="Prosternum" pageId="1" pageNumber="256" specimenCount="1" stateProvince="Centre">
|
|
<collectingCounty id="62AFB2E9FFC9FFF140B1950CA412F887" box="[199,277,1799,1822]" pageId="1" pageNumber="256">Venter</collectingCounty>
|
|
:
|
|
<collectingMunicipality id="6BAA501FFFC9FFF14152950CA4A2F887" box="[292,421,1799,1822]" pageId="1" pageNumber="256">Prosternum</collectingMunicipality>
|
|
broadly emarginate anteriorly, to receive ventral margin of retracted head.
|
|
<location id="8EAE9CBEFFC9FFF1455C950DA09BF886" LSID="urn:lsid:plazi:treatment:03D87B73FFC9FFF240E191DAA78FFCA2:8EAE9CBEFFC9FFF1455C950DA09BF886" box="[1322,1436,1798,1823]" county="Venter" municipality="Prosternum" name="Prosternal" pageId="1" pageNumber="256" stateProvince="Centre">Prosternal</location>
|
|
process slightly longer than broad, almost parallel-sided over most of length; lateral margins strongly raised.
|
|
<collectingRegion id="49B50487FFC9FFF140E19544A5E5F8FE" box="[151,226,1871,1895]" country="France" name="Centre" pageId="1" pageNumber="256">Centre</collectingRegion>
|
|
of mesoventrite broadly emarginate to receive prosternal process.
|
|
<location id="8EAE9CBEFFC9FFF143A59545A135F8FE" LSID="urn:lsid:plazi:treatment:03D87B73FFC9FFF240E191DAA78FFCA2:8EAE9CBEFFC9FFF143A59545A135F8FE" box="[979,1074,1870,1895]" county="Venter" municipality="Prosternum" name="Junction" pageId="1" pageNumber="256" stateProvince="Centre">Junction</location>
|
|
of meso and metaventrites with lateral foveae.
|
|
<location id="8EAE9CBEFFC9FFF141489579A4CAF812" LSID="urn:lsid:plazi:treatment:03D87B73FFC9FFF240E191DAA78FFCA2:8EAE9CBEFFC9FFF141489579A4CAF812" box="[318,461,1906,1931]" county="Venter" municipality="Prosternum" name="Metaventrite" pageId="1" pageNumber="256" stateProvince="Centre">Metaventrite</location>
|
|
punctate, broadly concave in middle, more strongly so in males than females; median longitudinal suture evident.
|
|
<location id="8EAE9CBEFFC9FFF141A4959DA747F836" LSID="urn:lsid:plazi:treatment:03D87B73FFC9FFF240E191DAA78FFCA2:8EAE9CBEFFC9FFF141A4959DA747F836" box="[466,576,1942,1967]" county="Venter" municipality="Prosternum" name="Abdomen" pageId="1" pageNumber="256" stateProvince="Centre">Abdomen</location>
|
|
with 5 ventrites.
|
|
<location id="8EAE9CBEFFC9FFF14370959DA63DF836" LSID="urn:lsid:plazi:treatment:03D87B73FFC9FFF240E191DAA78FFCA2:8EAE9CBEFFC9FFF14370959DA63DF836" box="[774,826,1942,1967]" county="Venter" municipality="Prosternum" name="First" pageId="1" pageNumber="256" stateProvince="Centre">First</location>
|
|
ventrite with semicircular carinae encircling posterior margins of metacoxae, fringed with irregular row of coarse, rugose punctures.
|
|
<location id="8EAE9CBEFFC9FFF1447095B1A12DF84A" LSID="urn:lsid:plazi:treatment:03D87B73FFC9FFF240E191DAA78FFCA2:8EAE9CBEFFC9FFF1447095B1A12DF84A" box="[1030,1066,1978,2003]" county="Venter" municipality="Prosternum" name="All" pageId="1" pageNumber="256" stateProvince="Centre">All</location>
|
|
ventrites punctate and granulate, at least laterally.
|
|
<location id="8EAE9CBEFFCAFFF24128929CA4B2FF29" LSID="urn:lsid:plazi:treatment:03D87B73FFC9FFF240E191DAA78FFCA2:8EAE9CBEFFCAFFF24128929CA4B2FF29" box="[350,437,151,176]" county="Venter" municipality="Prosternum" name="Sternite" pageId="2" pageNumber="257" stateProvince="Centre">Sternite</location>
|
|
8 with long, narrow apodeme, approximately
|
|
<collectingDate id="EF8B154DFFCAFFF243A29293A6F7FF29" box="[980,1008,152,176]" pageId="2" pageNumber="257">
|
|
<date id="FFCFECA5FFCAFFF243A29293A6F7FF29" box="[980,1008,152,176]" pageId="2" pageNumber="257">3x</date>
|
|
</collectingDate>
|
|
length of plate in females; apodeme triangular, similar length to plate, in males. Tergite 8 with broadly rounded apex in both sexes, with apical setal fringes.
|
|
</materialsCitation>
|
|
</paragraph>
|
|
<paragraph id="8BCECA65FFCAFFF240B1930EA5EDFED9" blockId="2.[151,1437,151,827]" pageId="2" pageNumber="257">Legs: Rather short, tibiae with golden setae on inner surfaces. Meso and metatibiae modified in males. Claws simple.</paragraph>
|
|
<paragraph id="8BCECA65FFCAFFF240B19347A4AAFE4A" blockId="2.[151,1437,151,827]" pageId="2" pageNumber="257">Hindwing: Approximately 1.5x body length; apical field large, occupying half of total wing length; medial spur narrow, weakly pigmented, not reaching hind margin; medial field occupying two-thirds of basal half of wing, with three distinct veins (MP3, MP4, Cu); anal field small, occupying one third of basal half; hind wing margin with long trichiae throughout.</paragraph>
|
|
<paragraph id="8BCECA65FFCAFFF240B193D5A715FD82" blockId="2.[151,1437,151,827]" pageId="2" pageNumber="257">Aedeagus: Long and slender, corona and fibula absent. Phallobase asymmetrical, shorter than penis and parameres; penis apex acuminate.</paragraph>
|
|
<paragraph id="8BCECA65FFCAFFF240B1902DA4EDFDFA" blockId="2.[151,1437,151,827]" pageId="2" pageNumber="257">Ovipositor: Symmetrical, elongate; proximal gonocoxite as long as or longer than distal; distal gonocoxite elongate, with terminal stylus.</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="C36B99EEFFCAFFF240B19065A78FFCA2" pageId="2" pageNumber="257" type="diagnosis">
|
|
<paragraph id="8BCECA65FFCAFFF240B19065A76CFD6A" blockId="2.[151,1437,151,827]" pageId="2" pageNumber="257">
|
|
<emphasis id="B9051677FFCAFFF240B19065A4C0FD1E" bold="true" box="[199,455,622,647]" pageId="2" pageNumber="257">Differential diagnosis</emphasis>
|
|
.
|
|
<taxonomicName id="4C71B1E6FFCAFFF241A19064A731FD1E" box="[471,566,623,647]" class="Insecta" family="Elmidae" genus="Leielmis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="257" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="B9051677FFCAFFF241A19064A731FD1E" box="[471,566,623,647]" italics="true" pageId="2" pageNumber="257">Leielmis</emphasis>
|
|
</taxonomicName>
|
|
can be recognised by the combination of short and strongly convex habitus, strongly narrowed anteriorly and posteriorly; absence of sulci, carinae or gibbosities on the pronotum, which is expanded into a narrow anterior hood; sub-parallel prosternal process; elytra with 9 puncture rows, lacking costae or gibbosities and the simple tarsal claws.
|
|
</paragraph>
|
|
<paragraph id="8BCECA65FFCAFFF240B190F5A78FFCA2" blockId="2.[151,1437,151,827]" pageId="2" pageNumber="257">
|
|
<materialsCitation id="3B19C038FFCAFFF240B190F5A78FFCA2" ID-GBIF-Occurrence="1455889623" country="South Africa" location="Cape" pageId="2" pageNumber="257" specimenCount="1" stateProvince="Cape">
|
|
The genus currently includes one described species, plus two new species described below, all of which are endemic to the
|
|
<collectingRegion id="49B50487FFCAFFF241359128A47AFCA2" box="[323,381,803,827]" country="South Africa" name="Western Cape" pageId="2" pageNumber="257">Cape</collectingRegion>
|
|
region of
|
|
<collectingCountry id="F3668AF5FFCAFFF241869129A784FCA2" box="[496,643,802,827]" name="South Africa" pageId="2" pageNumber="257">South Africa</collectingCountry>
|
|
.
|
|
</materialsCitation>
|
|
</paragraph>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |