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<document id="7A9F35115ADE6363476DC5E00CA7B64D" ID-DOI="10.1093/zoolinnean/zlad115" ID-ISSN="0024-4082" ID-Zenodo-Dep="11241094" ID-ZooBank="E6D9AF7E-F0AD-4656-B2F2-7FBFAA0312B3R" IM.bibliography_approvedBy="julia" IM.illustrations_approvedBy="julia" IM.materialsCitations_approvedBy="julia" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="julia" IM.taxonomicNames_approvedBy="julia" IM.treatmentCitations_approvedBy="julia" IM.treatments_approvedBy="julia" checkinTime="1716363659947" checkinUser="plazi" docAuthor="Cherman, Mariana A., Basílio, Daniel S., Clarkson, Bruno, Agostinis, André O., Smith, Andrew B. T., Vaz-De-Mello, Fernando Z. &amp; Almeida, Lúcia M." docDate="2024" docId="03D18442FFDEFF9C317DFEEC5BB8FC27" docLanguage="en" docName="zlad115.pdf" docOrigin="Zoological Journal of the Linnean Society 201 (1)" docSource="http://dx.doi.org/10.1093/zoolinnean/zlad115" docStyle="DocumentStyle:4F230B9370E98E256D973D6DFB57F36C.5:ZoolJLinnSoc.2023-.journal_article" docStyleId="4F230B9370E98E256D973D6DFB57F36C" docStyleName="ZoolJLinnSoc.2023-.journal_article" docStyleVersion="5" docTitle="Liogenys niger Cherman &amp; Basílio &amp; Clarkson &amp; Agostinis &amp; Smith &amp; Vaz-De-Mello &amp; Almeida 2024" docType="treatment" docVersion="1" lastPageNumber="119" masterDocId="FFE8FC3AFFCCFF893202FFE1597BFF90" masterDocTitle="New and revised taxa of Neotropical Diplotaxini (Coleoptera: Melolonthidae): do they change the existing relationships? Revisiting systematics with morphological and molecular data" masterLastPageNumber="135" masterPageNumber="98" pageNumber="116" updateTime="1716408306816" updateUser="julia">
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<mods:title id="584D44DE67816E49EB19FB5B07272083">New and revised taxa of Neotropical Diplotaxini (Coleoptera: Melolonthidae): do they change the existing relationships? Revisiting systematics with morphological and molecular data</mods:title>
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<mods:namePart id="4D7179CB2193BCEBDB034956ABF3456C">Cherman, Mariana A.</mods:namePart>
<mods:affiliation id="9CC6DC8C50C67BFC8642E0FDF01AE429">Departamento de Zoologia, Universidade Federal do Paraná, Caixa Postal 19020, 81531 - 980, Curitiba, Paraná, Brazil &amp; Departamento de Biologia e Zoologia, Instituto de Biociências, Universidade Federal de Mato Grosso, Av. Fernando Correia, no. 2367, Boa Esperança, 78060 - 900, Cuiabá, Mato Grosso, Brazil</mods:affiliation>
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<mods:namePart id="7B724178A362565F4D9553002E6B805E">Basílio, Daniel S.</mods:namePart>
<mods:affiliation id="316F8E32CAC573925370B1769A3D5595">Departamento de Zoologia, Universidade Federal do Paraná, Caixa Postal 19020, 81531 - 980, Curitiba, Paraná, Brazil</mods:affiliation>
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<mods:namePart id="5650EB95907AB08C4C5249DBC1BDE62B">Clarkson, Bruno</mods:namePart>
<mods:affiliation id="459A58ADE08811912D20C7FAD09883B3">Departamento de Zoologia, Universidade Federal do Paraná, Caixa Postal 19020, 81531 - 980, Curitiba, Paraná, Brazil &amp; Instituto Oswaldo Cruz, Fundação Oswaldo Cruz, Av. Brasil, 4365, Pavilhão Mourisco, Manguinhos, 21040 - 360, Rio de Janeiro, Rio de Janeiro, Brazil &amp; Laboratório de Entomologia e Fitopatologia, Universidade Estadual do Norte Fluminense Darcy Ribeiro, 28013 - 602 Campos dos Goytacazes, Rio de Janeiro, Brazil</mods:affiliation>
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<mods:namePart id="09B0CD795B3DBE5B48167C80BBDFD3D2">Agostinis, André O.</mods:namePart>
<mods:affiliation id="E4EF22C3FCACDB6E089CF376D9FAC374">Departamento de Zoologia, Universidade Federal do Paraná, Caixa Postal 19020, 81531 - 980, Curitiba, Paraná, Brazil</mods:affiliation>
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<mods:namePart id="DF089E183CBD94343937FC097409A2F0">Vaz-De-Mello, Fernando Z.</mods:namePart>
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<mods:title id="975B92010C8EA03766F43C66FCDEF700">Zoological Journal of the Linnean Society</mods:title>
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<mods:date id="538DEBF92B3A30540B0A4DFACC8BAB73">2024</mods:date>
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<taxonomicName id="4C784ED7FFDEFF9B317DFEEC5D92FEB7" ID-CoL="BV644" authority="(Blanchard, 1851)" authorityName="Cherman &amp; Basílio &amp; Clarkson &amp; Agostinis &amp; Smith &amp; Vaz-De-Mello &amp; Almeida" authorityYear="2024" baseAuthorityName="Blanchard" baseAuthorityYear="1851" box="[895,1257,269,295]" class="Insecta" family="Scarabaeidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="116" phylum="Arthropoda" rank="species" species="niger" status="comb. nov.">
<emphasis id="B90CE946FFDEFF9B317DFEEC5D64FEB7" bold="true" box="[895,1055,269,295]" italics="true" pageId="18" pageNumber="116">Liogenys niger</emphasis>
(
<bibRefCitation id="EFE948A5FFDEFF9B3629FEEC5DA6FEB7" author="Blanchard ME" box="[1067,1245,269,295]" pageId="18" pageNumber="116" refId="ref26799" refString="Blanchard ME. Museum d'Histoire Naturelle de Paris. Catalogue de la Collection Entomologique. Classe des Insectes. Ordre des Coleopteres. Tome I. Part 2. Paris: Gide et Baudry, 1851." type="book" year="1851">Blanchard, 1851</bibRefCitation>
)
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comb. nov. (
<figureCitation id="134329D1FFDEFF9B363BFED35DD8FEDC" box="[1081,1187,306,332]" captionStart="Figure 7" captionStartId="19.[129,194,1836,1860]" captionTargetBox="[181,1423,149,1808]" captionTargetId="figure-7@19.[181,1423,144,1808]" captionTargetPageId="19" captionText="Figure 7. Liogenys niger (Blanchard) comb. nov.. Male neotype habitus (A), labels (E), and complete series at MLUH (D); female lectotype (B) and labels (F); non-uniform body colour variation (C); male in lateral (G) and posterior (I) views; detail of ventrites II and II with medial tubercle (H); parameres in dorsal (J) and lateral (K) views." figureDoi="http://doi.org/10.5281/zenodo.11241108" httpUri="https://zenodo.org/record/11241108/files/figure.png" pageId="18" pageNumber="116">Fig. 7A–K</figureCitation>
)
</emphasis>
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<treatmentCitation id="0AD91345FFDEFF9B3128FE8C5DBCFE15" author="Blanchard ME" box="[810,1223,365,389]" page="172" pageId="18" pageNumber="116" year="1851">
<taxonomicName id="4C784ED7FFDEFF9B3128FE8C5DBCFE15" ID-CoL="3MD22" authority="Blanchard 1851: 172" authorityName="Blanchard" authorityPageNumber="172" authorityYear="1851" box="[810,1223,365,389]" class="Insecta" family="Scarabaeidae" genus="Homalochilus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="116" phylum="Arthropoda" rank="species" species="niger">
<emphasis id="B90CE946FFDEFF9B3128FE8C5A9CFE15" box="[810,999,365,389]" italics="true" pageId="18" pageNumber="116">Homalochilus niger</emphasis>
<bibRefCitation id="EFE948A5FFDEFF9B31EDFE8C5DBCFE15" author="Blanchard ME" box="[1007,1223,365,389]" pageId="18" pageNumber="116" refId="ref26799" refString="Blanchard ME. Museum d'Histoire Naturelle de Paris. Catalogue de la Collection Entomologique. Classe des Insectes. Ordre des Coleopteres. Tome I. Part 2. Paris: Gide et Baudry, 1851." type="book" year="1851">Blanchard 1851: 172</bibRefCitation>
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(original description); Harold 1869: 1141 (catalogue);
<bibRefCitation id="EFE948A5FFDEFF9B3676FE6C5D9CFE34" author="Borre P" box="[1140,1255,396,420]" pageId="18" pageNumber="116" refId="ref26836" refString="Borre P. Comptes-rendus des seances de la Societe entomologique de Belgique. Assemblee mensuelle du 4 mars 1871. Annales de la Societe Entomologique de Belgique 1871; 14: XVIII - XV." type="journal volume" year="1871">Borre 1871</bibRefCitation>
: XXIII (catalogue); Kolbe 1907: 65 (catalogue);
<treatmentCitation id="0AD91345FFDEFF9B364FFE4D5D8CFE54" author="Bruch C" box="[1101,1271,428,452]" page="201" pageId="18" pageNumber="116" year="1911">
<bibRefCitation id="EFE948A5FFDEFF9B364FFE4D5D8CFE54" author="Bruch C" box="[1101,1271,428,452]" pageId="18" pageNumber="116" pagination="181 - 225" refId="ref26889" refString="Bruch C. Catalogo sistematico de los coleopteros de la Republica Argentina. Revista del Museo de la Plata 1911; 17: 181 - 225." type="journal article" year="1911">Bruch 1911: 201</bibRefCitation>
</treatmentCitation>
(catalogue); Dalla Torre 1913: 319;
<treatmentCitation id="0AD91345FFDEFF9B31DCFE2A5DB2FE73" author="Blackwelder RE" box="[990,1225,459,483]" page="228" pageId="18" pageNumber="116" year="1944">
<bibRefCitation id="EFE948A5FFDEFF9B31DCFE2A5DB2FE73" author="Blackwelder RE" box="[990,1225,459,483]" pageId="18" pageNumber="116" pagination="189 - 341" refId="ref26760" refString="Blackwelder RE. Checklist of the coleopterous insects of Mexico, Central America, the West Indies and South America, Part 2. Bulletin of the United States National Museum 1944; 185: 189 - 341." type="journal article" year="1944">Blackwelder 1944: 228</bibRefCitation>
</treatmentCitation>
(catalogue);
<treatmentCitation id="0AD91345FFDEFF9B3753FE2A5AE0FD92" author="Gutierrez R" page="213" pageId="18" pageNumber="116" year="1952">
<bibRefCitation id="EFE948A5FFDEFF9B3753FE2A5AE0FD92" author="Gutierrez R" pageId="18" pageNumber="116" pagination="207 - 27" refId="ref27805" refString="Gutierrez R. Notas sobre Scarabaeidae Neotropicos. Revista Chilena de Entomologia 1952; 2: 207 - 27." type="journal article" year="1952">Gutiérrez 1952: 213</bibRefCitation>
</treatmentCitation>
, 214 (key, redescription);
<treatmentCitation id="0AD91345FFDEFF9B36C3FE0B5C16FD92" author="Smith KGV" box="[1217,1389,490,514]" page="62" pageId="18" pageNumber="116" year="1987">
<bibRefCitation id="EFE948A5FFDEFF9B36C3FE0B5C16FD92" author="Smith KGV" box="[1217,1389,490,514]" pageId="18" pageNumber="116" pagination="1 - 143" refId="ref29036" refString="Smith KGV. Darwin's insects: Charles Darwin's entomological notes. In Duncan M. Porter (ed.), Bulletin of the British Museum of Natural History (Historical Series) 1987; 14: 1 - 143." type="journal article" year="1987">Smith 1987: 62</bibRefCitation>
</treatmentCitation>
(catalogue);
<treatmentCitation id="0AD91345FFDEFF9B317EFDEB5D65FDB2" author="Evans AV" box="[892,1054,522,546]" page="11" pageId="18" pageNumber="116" year="2003">
<bibRefCitation id="EFE948A5FFDEFF9B317EFDEB5D65FDB2" author="Evans AV" box="[892,1054,522,546]" pageId="18" pageNumber="116" pagination="1 - 458" refId="ref27431" refString="Evans AV. A checklist of the New World chafers (Coleoptera: Scarabaeidae: Melolonthinae). Zootaxa 2003; 211: 1 - 458." type="journal article" year="2003">Evans 2003: 11</bibRefCitation>
</treatmentCitation>
, 206 (systematics, catalog);
<treatmentCitation id="0AD91345FFDEFF9B374BFDEB5AAEFDD1" author="Evans AV &amp; Smith ABT" page="170" pageId="18" pageNumber="116" year="2005">
<bibRefCitation id="EFE948A5FFDEFF9B374BFDEB5AAEFDD1" author="Evans AV &amp; Smith ABT" pageId="18" pageNumber="116" pagination="1 - 344" refId="ref27458" refString="Evans AV, Smith ABT. An electronic checklist of the New World chafers (Coleoptera: Scarabaeidae: Melolonthinae), v. 1. Papers in Entomology 2005; 1: 1 - 344." type="journal article" year="2005">Evans and Smith 2005: 170</bibRefCitation>
</treatmentCitation>
(catalogue);
<treatmentCitation id="0AD91345FFDEFF9B365EFDC85C08FDD1" author="Evans AV &amp; Smith ABT" box="[1116,1395,553,577]" page="174" pageId="18" pageNumber="116" year="2009">
<bibRefCitation id="EFE948A5FFDEFF9B365EFDC85C08FDD1" author="Evans AV &amp; Smith ABT" box="[1116,1395,553,577]" pageId="18" pageNumber="116" pagination="1 - 344" refId="ref27495" refString="Evans AV, Smith ABT. An electronic checklist of the New World chafers (Coleoptera: Scarabaeidae: Melolonthinae), v. 3, Papers in Entomology 2009; 1: 1 - 344." type="journal article" year="2009">Evans and Smith 2009: 174</bibRefCitation>
</treatmentCitation>
(catalogue);
<treatmentCitation id="0AD91345FFDEFF9B317BFDA95D55FDF0" author="Krajcik M" box="[889,1070,584,608]" page="128" pageId="18" pageNumber="116" year="2012">
<bibRefCitation id="EFE948A5FFDEFF9B317BFDA95D55FDF0" author="Krajcik M" box="[889,1070,584,608]" pageId="18" pageNumber="116" refId="ref28103" refString="Krajcik M. Checklist of the World Scarabaeoidea. Plzen: Animma. x, 2012." type="book" year="2012">Krajčík 2012: 128</bibRefCitation>
</treatmentCitation>
(catalogue);
<treatmentCitation id="0AD91345FFDEFF9B36B6FDA95CC8FDF0" author="Cherman MA &amp; Moron MA &amp; Almeida LM" box="[1204,1459,584,608]" page="765" pageId="18" pageNumber="116" year="2016">
<bibRefCitation id="EFE948A5FFDEFF9B36B6FDA95CC8FDF0" author="Cherman MA &amp; Moron MA &amp; Almeida LM" box="[1204,1459,584,608]" pageId="18" pageNumber="116" pagination="744 - 70" refId="ref27020" refString="Cherman MA, Moron MA, Almeida LM. Phylogenetic relationships within Diplotaxini Kirby (Coleoptera: Melolonthidae: Melolonthinae) with emphasis on Liogenys Guerin-Meneville. Systematic Entomology 2016; 41: 744 - 70." type="journal article" year="2016">
Cherman
<emphasis id="B90CE946FFDEFF9B3718FDA85C33FDF0" box="[1306,1352,584,608]" italics="true" pageId="18" pageNumber="116">et al.</emphasis>
2016: 765
</bibRefCitation>
</treatmentCitation>
(systematics);
<treatmentCitation id="0AD91345FFDEFF9B31BDFD895DD5FD10" author="Cherman MA &amp; Mise KM &amp; Moron MA" box="[959,1198,616,640]" page="18" pageId="18" pageNumber="116" year="2017">
<bibRefCitation id="EFE948A5FFDEFF9B31BDFD895DD5FD10" author="Cherman MA &amp; Mise KM &amp; Moron MA" box="[959,1198,616,640]" pageId="18" pageNumber="116" pagination="1 - 120" refId="ref27057" refString="Cherman MA, Mise KM, Moron MA et al. A taxonomic revision of Liogenys occurring in Brazil with an interactive key and remarks on New World Diplotaxini (Coleoptera, Melolonthidae). ZooKeys 2017; 699: 1 - 120." type="journal article" year="2017">
Cherman
<emphasis id="B90CE946FFDEFF9B3627FD895D2FFD10" box="[1061,1108,616,640]" italics="true" pageId="18" pageNumber="116">et al.</emphasis>
2017:18
</bibRefCitation>
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, 34 (systematics).
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</paragraph>
<paragraph id="8BC73554FFDEFF9B3147FD695C2BFC0A" blockId="18.[810,1459,365,922]" pageId="18" pageNumber="116">
<treatmentCitationGroup id="AB68127AFFDEFF9B3147FD695C2BFC0A" pageId="18" pageNumber="116">
<treatmentCitation id="0AD91345FFDEFF9B3147FD695DB9FD0F" author="Burmeister H" box="[837,1218,647,671]" page="16" pageId="18" pageNumber="116" year="1855">
<taxonomicName id="4C784ED7FFDEFF9B3147FD695DB9FD0F" ID-CoL="3V5Y2" authority="Burmeister 1855: 16" authorityName="Burmeister" authorityPageNumber="16" authorityYear="1855" box="[837,1218,647,671]" class="Insecta" family="Scarabaeidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="116" phylum="Arthropoda" rank="species" species="morio">
<emphasis id="B90CE946FFDEFF9B3147FD695AA6FD0F" box="[837,989,648,671]" italics="true" pageId="18" pageNumber="116">Liogenys morio</emphasis>
<bibRefCitation id="EFE948A5FFDEFF9B31E5FD665DB9FD0F" author="Burmeister H" box="[999,1218,647,671]" pageId="18" pageNumber="116" refId="ref26916" refString="Burmeister H. Handbuch der entomologie (Coleoptera Lamellicornia Phyllophaga Chaenochela). Vol. 4, Part 2. Berlin: Enslin, T. C. F., 1855." type="book" year="1855">Burmeister 1855: 16</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(original description); Harold 1869: 1141 (catalogue); Bates 1887: 155 (redescription); Kolbe 1907: 65 (catalogue);
<treatmentCitation id="0AD91345FFDEFF9B364FFD245D8CFD4E" author="Bruch C" box="[1101,1271,709,734]" page="200" pageId="18" pageNumber="116" year="1911">
<bibRefCitation id="EFE948A5FFDEFF9B364FFD245D8CFD4E" author="Bruch C" box="[1101,1271,709,734]" pageId="18" pageNumber="116" pagination="181 - 225" refId="ref26889" refString="Bruch C. Catalogo sistematico de los coleopteros de la Republica Argentina. Revista del Museo de la Plata 1911; 17: 181 - 225." type="journal article" year="1911">Bruch 1911: 200</bibRefCitation>
</treatmentCitation>
(catalogue); Dalla Torre 1913: 318 (catalogue);
<treatmentCitation id="0AD91345FFDEFF9B3676FD045C17FD6D" author="Blackwelder RE" box="[1140,1388,741,765]" page="227" pageId="18" pageNumber="116" year="1944">
<bibRefCitation id="EFE948A5FFDEFF9B3676FD045C17FD6D" author="Blackwelder RE" box="[1140,1388,741,765]" pageId="18" pageNumber="116" pagination="189 - 341" refId="ref26760" refString="Blackwelder RE. Checklist of the coleopterous insects of Mexico, Central America, the West Indies and South America, Part 2. Bulletin of the United States National Museum 1944; 185: 189 - 341." type="journal article" year="1944">Blackwelder 1944: 227</bibRefCitation>
</treatmentCitation>
(catalogue);
<treatmentCitation id="0AD91345FFDEFF9B3186FCE55D12FC8C" author="Gutierrez R" box="[900,1129,772,796]" page="211" pageId="18" pageNumber="116" year="1952">
<bibRefCitation id="EFE948A5FFDEFF9B3186FCE55D12FC8C" author="Gutierrez R" box="[900,1129,772,796]" pageId="18" pageNumber="116" pagination="207 - 27" refId="ref27805" refString="Gutierrez R. Notas sobre Scarabaeidae Neotropicos. Revista Chilena de Entomologia 1952; 2: 207 - 27." type="journal article" year="1952">Gutiérrez 1952: 211</bibRefCitation>
</treatmentCitation>
(
<emphasis id="B90CE946FFDEFF9B3686FCE55C71FC8C" box="[1156,1290,772,796]" italics="true" pageId="18" pageNumber="116">Homalochilus</emphasis>
redescription);
<treatmentCitation id="0AD91345FFDEFF9B3128FCC55A98FCAC" author="Evans AV" box="[810,995,804,828]" page="211" pageId="18" pageNumber="116" year="2003">
<bibRefCitation id="EFE948A5FFDEFF9B3128FCC55A98FCAC" author="Evans AV" box="[810,995,804,828]" pageId="18" pageNumber="116" pagination="1 - 458" refId="ref27431" refString="Evans AV. A checklist of the New World chafers (Coleoptera: Scarabaeidae: Melolonthinae). Zootaxa 2003; 211: 1 - 458." type="journal article" year="2003">Evans 2003: 211</bibRefCitation>
</treatmentCitation>
(catalogue);
<treatmentCitation id="0AD91345FFDEFF9B3679FCC55CC8FCAB" author="Evans AV &amp; Smith ABT" box="[1147,1459,803,827]" page="175" pageId="18" pageNumber="116" year="2005">
<bibRefCitation id="EFE948A5FFDEFF9B3679FCC55CC8FCAB" author="Evans AV &amp; Smith ABT" box="[1147,1459,803,827]" pageId="18" pageNumber="116" pagination="1 - 344" refId="ref27458" refString="Evans AV, Smith ABT. An electronic checklist of the New World chafers (Coleoptera: Scarabaeidae: Melolonthinae), v. 1. Papers in Entomology 2005; 1: 1 - 344." type="journal article" year="2005">Evans and Smith 2005: 175</bibRefCitation>
</treatmentCitation>
(catalogue);
<treatmentCitation id="0AD91345FFDEFF9B31B3FCA25DA2FCCB" author="Evans AV &amp; Smith ABT" box="[945,1241,835,859]" page="179" pageId="18" pageNumber="116" year="2009">
<bibRefCitation id="EFE948A5FFDEFF9B31B3FCA25DA2FCCB" author="Evans AV &amp; Smith ABT" box="[945,1241,835,859]" pageId="18" pageNumber="116" pagination="1 - 344" refId="ref27495" refString="Evans AV, Smith ABT. An electronic checklist of the New World chafers (Coleoptera: Scarabaeidae: Melolonthinae), v. 3, Papers in Entomology 2009; 1: 1 - 344." type="journal article" year="2009">Evans and Smith 2009: 179</bibRefCitation>
</treatmentCitation>
(catalogue);
<treatmentCitation id="0AD91345FFDEFF9B376BFCA25AE9FCEA" author="Krajcik M" page="145" pageId="18" pageNumber="116" year="2012">
<bibRefCitation id="EFE948A5FFDEFF9B376BFCA25AE9FCEA" author="Krajcik M" pageId="18" pageNumber="116" refId="ref28103" refString="Krajcik M. Checklist of the World Scarabaeoidea. Plzen: Animma. x, 2012." type="book" year="2012">Krajčík 2012: 145</bibRefCitation>
</treatmentCitation>
(catalogue);
<treatmentCitation id="0AD91345FFDEFF9B3618FC835C66FCEA" author="Cherman MA &amp; Moron MA &amp; Almeida LM" box="[1050,1309,866,890]" page="765" pageId="18" pageNumber="116" year="2016">
<bibRefCitation id="EFE948A5FFDEFF9B3618FC835C66FCEA" author="Cherman MA &amp; Moron MA &amp; Almeida LM" box="[1050,1309,866,890]" pageId="18" pageNumber="116" pagination="744 - 70" refId="ref27020" refString="Cherman MA, Moron MA, Almeida LM. Phylogenetic relationships within Diplotaxini Kirby (Coleoptera: Melolonthidae: Melolonthinae) with emphasis on Liogenys Guerin-Meneville. Systematic Entomology 2016; 41: 744 - 70." type="journal article" year="2016">
Cherman
<emphasis id="B90CE946FFDEFF9B3683FC825DCAFCEA" box="[1153,1201,866,890]" italics="true" pageId="18" pageNumber="116">et al.</emphasis>
2016: 765
</bibRefCitation>
</treatmentCitation>
(systematics);
<treatmentCitation id="0AD91345FFDEFF9B3128FC605D6AFC0A" author="Cherman MA &amp; Mise KM &amp; Moron MA" box="[810,1041,897,922]" page="4" pageId="18" pageNumber="116" year="2017">
<bibRefCitation id="EFE948A5FFDEFF9B3128FC605D6AFC0A" author="Cherman MA &amp; Mise KM &amp; Moron MA" box="[810,1041,897,922]" pageId="18" pageNumber="116" pagination="1 - 120" refId="ref27057" refString="Cherman MA, Mise KM, Moron MA et al. A taxonomic revision of Liogenys occurring in Brazil with an interactive key and remarks on New World Diplotaxini (Coleoptera, Melolonthidae). ZooKeys 2017; 699: 1 - 120." type="journal article" year="2017">
Cherman
<emphasis id="B90CE946FFDEFF9B3192FC635AC4FC09" box="[912,959,897,921]" italics="true" pageId="18" pageNumber="116">et al.</emphasis>
2017: 4
</bibRefCitation>
</treatmentCitation>
(taxonomic history).
<emphasis id="B90CE946FFDEFF9B36F2FC635C2BFC0A" bold="true" box="[1264,1360,898,922]" pageId="18" pageNumber="116">syn. nov.</emphasis>
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C36266DFFFDEFF9D31FCFC5B5B55FF43" lastPageId="20" lastPageNumber="118" pageId="18" pageNumber="116" type="materials_examined">
<paragraph id="8BC73554FFDEFF9B31FCFC5B5DA4FC43" blockId="18.[1022,1247,953,979]" box="[1022,1247,953,979]" pageId="18" pageNumber="116">
<emphasis id="B90CE946FFDEFF9B31FCFC5B5DA4FC43" box="[1022,1247,953,979]" italics="true" pageId="18" pageNumber="116">Type material examined</emphasis>
</paragraph>
<paragraph id="8BC73554FFDEFF9B3128FC015CEBFA82" blockId="18.[810,1460,992,1454]" pageId="18" pageNumber="116">
<taxonomicName id="4C784ED7FFDEFF9B3128FC015A9DFC67" ID-CoL="3MD22" authorityName="Blanchard" authorityYear="1851" box="[810,998,992,1016]" class="Insecta" family="Scarabaeidae" genus="Homalochilus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="116" phylum="Arthropoda" rank="species" species="niger">
<emphasis id="B90CE946FFDEFF9B3128FC015A9DFC67" box="[810,998,992,1016]" italics="true" pageId="18" pageNumber="116">Homalochilus niger</emphasis>
</taxonomicName>
<specimenCount id="9D7EFEDDFFDEFF9B31E9FC025D7EFC67" box="[1003,1029,995,1015]" count="1" pageId="18" pageNumber="116" type="female">♀</specimenCount>
<typeStatus id="54C38BF6FFDEFF9B3609FC015D17FC68" box="[1035,1132,992,1016]" pageId="18" pageNumber="116" type="lectotype">lectotype</typeStatus>
, present designation (MNHN): [white handwritten] ‘6045/34’, [light green typeset] ‘MUSÉUM PARIS/[handwritten] Patagonie/d’Orbigny’, [green handwritten] ‘O.
<taxonomicName id="4C784ED7FFDEFF9B31BAFBDF5D55FBC6" ID-CoL="3MD22" authorityName="Blanchard" authorityYear="1851" box="[952,1070,1086,1110]" class="Insecta" family="Scarabaeidae" genus="Homalochilus" isUncertain="true" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="116" phylum="Arthropoda" rank="species" species="niger">[sic] niger</taxonomicName>
/Cat Mus/Patagonie/M. d’Orbigny’, [red typeset] ‘SYNTYPE’, [white, outlined in red, typeset] ‘
<typeStatus id="54C38BF6FFDEFF9B312CFB9D5AB3FB04" box="[814,968,1148,1172]" pageId="18" pageNumber="116" type="lectotype">LECTOTYPE</typeStatus>
/
<taxonomicName id="4C784ED7FFDEFF9B31D2FB9D5DDEFB04" ID-CoL="3MD22" authorityName="Blanchard" authorityYear="1851" box="[976,1189,1148,1172]" class="Insecta" family="Scarabaeidae" genus="Homalochilus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="116" phylum="Arthropoda" rank="species" species="niger">
<emphasis id="B90CE946FFDEFF9B31D2FB9D5DDEFB04" box="[976,1189,1148,1172]" italics="true" pageId="18" pageNumber="116">Homalochilus niger</emphasis>
</taxonomicName>
/
<bibRefCitation id="EFE948A5FFDEFF9B36AEFB9D5C0BFB04" author="Blanchard ME" box="[1196,1392,1148,1172]" pageId="18" pageNumber="116" refId="ref26799" refString="Blanchard ME. Museum d'Histoire Naturelle de Paris. Catalogue de la Collection Entomologique. Classe des Insectes. Ordre des Coleopteres. Tome I. Part 2. Paris: Gide et Baudry, 1851." type="book" year="1851">Blanchard 1851</bibRefCitation>
/[des.] M. A.
<bibRefCitation id="EFE948A5FFDEFF9B316EFB7D5D7CFB23" author="Cherman MA &amp; Almeida LM" box="[876,1031,1179,1204]" pageId="18" pageNumber="116" pagination="171 - 8" refId="ref26985" refString="Cherman MA, Almeida LM. New Brazilian species of Liogenys Guerin-Meneville (Coleoptera: Melolonthidae: Melolonthinae) and redescription of two related species. Dugesiana 2015; 22: 171 - 8." type="journal article" year="2015">Cherman 2015</bibRefCitation>
’. Female
<typeStatus id="54C38BF6FFDEFF9B3664FB7D5D8AFB24" box="[1126,1265,1180,1204]" pageId="18" pageNumber="116" type="paralectotype">paralectotype</typeStatus>
(MNHN): [white handwritten] ‘6045/34’, [light green typeset] ‘MUSÉUM PARIS/PATAGONIE/(PATAGONES)/D’ORBIGNY 1834’, [white, typeset in red] ‘COTYPE’, [red typeset] ‘SYNTYPE’.
</paragraph>
<paragraph id="8BC73554FFDEFF9B3147FAFB5AB4FA3E" blockId="18.[810,1460,992,1454]" pageId="18" pageNumber="116">
<taxonomicName id="4C784ED7FFDEFF9B3147FAFB5AA5FAA1" ID-CoL="3V5Y2" box="[837,990,1306,1329]" class="Insecta" family="Scarabaeidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="116" phylum="Arthropoda" rank="species" species="morio">
<emphasis id="B90CE946FFDEFF9B3147FAFB5AA5FAA1" box="[837,990,1306,1329]" italics="true" pageId="18" pageNumber="116">Liogenys morio</emphasis>
</taxonomicName>
<specimenCount id="9D7EFEDDFFDEFF9B31EAFAFD5D79FAA0" box="[1000,1026,1308,1328]" count="1" pageId="18" pageNumber="116" type="male">♂</specimenCount>
<typeStatus id="54C38BF6FFDEFF9B360FFAF85D18FAA1" box="[1037,1123,1305,1329]" pageId="18" pageNumber="116" type="neotype">neotype</typeStatus>
, present designation (MLUH): [green handwritten] ‘morio [underlined] Burm/Bras. Dp [Dupont]’, [white handwritten by Darwin] ‘B. Blanca’, [red handwritten] ‘
<taxonomicName id="4C784ED7FFDEFF9B3188FA965D6AFA1F" ID-CoL="5F3D" box="[906,1041,1399,1423]" class="Insecta" family="Scarabaeidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="116" phylum="Arthropoda" rank="genus">LIOGENYS</taxonomicName>
/MORIO/Burm, 1855/
<typeStatus id="54C38BF6FFDEFF9B3706FA965CFFFA1F" box="[1284,1412,1399,1423]" pageId="18" pageNumber="116" type="neotype">NEOTYPE</typeStatus>
/des. Cherman M. A.’.
</paragraph>
<paragraph id="8BC73554FFDEFF9B31EAFA245D8EFA4E" blockId="18.[810,1460,1476,1977]" box="[1000,1269,1476,1502]" pageId="18" pageNumber="116">
<emphasis id="B90CE946FFDEFF9B31EAFA245D8EFA4E" box="[1000,1269,1476,1502]" italics="true" pageId="18" pageNumber="116">Non-type material examined</emphasis>
</paragraph>
<paragraph id="8BC73554FFDEFF9B3128FA0A5CCFF9B2" blockId="18.[810,1460,1476,1977]" pageId="18" pageNumber="116">
<materialsCitation id="3B103F09FFDEFF9B3128FA0A5CCBF9B2" collectingDate="1983-02" collectionCode="ZMHB, DZUP" collectorName="J. N. Pedrosa Macedo" country="Brazil" location="Itapena" municipality="Without" pageId="18" pageNumber="116" specimenCount="7" specimenCount-female="3" specimenCount-male="4" stateProvince="Sao Paulo">
<collectingCountry id="F36F75C4FFDEFF9B3128FA0A5AFDF993" box="[810,902,1515,1539]" name="Brazil" pageId="18" pageNumber="116">BRAZIL</collectingCountry>
.
<collectingMunicipality id="6BA3AF2EFFDEFF9B3193FA0B5A9DF992" box="[913,998,1514,1538]" pageId="18" pageNumber="116">Without</collectingMunicipality>
locality and date,
<specimenCount id="9D7EFEDDFFDEFF9B36A5FA0B5DB5F991" box="[1191,1230,1514,1538]" count="3" pageId="18" pageNumber="116" type="female">3♀</specimenCount>
(
<collectionCode id="ED69AD91FFDEFF9B36DDFA0B5C49F992" box="[1247,1330,1514,1538]" pageId="18" pageNumber="116">ZMHB</collectionCode>
);
<collectingRegion id="49BCFBB6FFDEFF9B3744FA0B5CCBF992" box="[1350,1456,1514,1538]" country="Brazil" name="Sao Paulo" pageId="18" pageNumber="116">São Paulo</collectingRegion>
:
<location id="8EA7638FFFDEFF9B3128F9EB5A0CF9B1" LSID="urn:lsid:plazi:treatment:03D18442FFDEFF9C317DFEEC5BB8FC27:8EA7638FFFDEFF9B3128F9EB5A0CF9B1" box="[810,887,1546,1569]" country="Brazil" municipality="Without" name="Itapena" pageId="18" pageNumber="116" stateProvince="Sao Paulo">Itapena</location>
,
<date id="FFC61394FFDEFF9B3188F9EB5AAEF9B2" box="[906,981,1546,1570]" pageId="18" pageNumber="116" value="1983-02">
<collectingDate id="EF82EA7CFFDEFF9B3188F9EB5AAEF9B2" box="[906,981,1546,1570]" pageId="18" pageNumber="116" value="1983-02">II.1983</collectingDate>
</date>
,
<collectorName id="268D5082FFDEFF9B31EBF9EB5DB4F9B2" box="[1001,1231,1546,1570]" pageId="18" pageNumber="116">J.N. Pedrosa Macedo</collectorName>
coll.,
<specimenCount id="9D7EFEDDFFDEFF9B371CF9EB5C3EF9B1" box="[1310,1349,1546,1570]" count="4" pageId="18" pageNumber="116" type="male">4♂</specimenCount>
(
<collectionCode id="ED69AD91FFDEFF9B375EF9EB5CD3F9B2" box="[1372,1448,1546,1570]" country="Brazil" httpUri="http://grbio.org/cool/5xp9-edpx" name="Universidade Federal do Parana, Colecao de Entomologia Pe. Jesus Santiago Moure" pageId="18" pageNumber="116" type="University or college">DZUP</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BC73554FFDEFF9B3128F9C85CC8F94E" blockId="18.[810,1460,1476,1977]" pageId="18" pageNumber="116">
<materialsCitation id="3B103F09FFDEFF9B3128F9C85CCBF9D1" box="[810,1456,1577,1601]" collectingDate="1955-10" collectorName="Bosq" country="Argentina" location="Pindapoy" municipality="Misiones" pageId="18" pageNumber="116" specimenCount="1">
<collectingCountry id="F36F75C4FFDEFF9B3128F9C85AB8F9D1" box="[810,963,1577,1601]" name="Argentina" pageId="18" pageNumber="116">ARGENTINA</collectingCountry>
.
<collectingMunicipality id="6BA3AF2EFFDEFF9B31DFF9C85D46F9D1" box="[989,1085,1577,1601]" pageId="18" pageNumber="116">Misiones</collectingMunicipality>
:
<location id="8EA7638FFFDEFF9B3655F9C85DCCF9D1" LSID="urn:lsid:plazi:treatment:03D18442FFDEFF9C317DFEEC5BB8FC27:8EA7638FFFDEFF9B3655F9C85DCCF9D1" box="[1111,1207,1577,1601]" country="Argentina" municipality="Misiones" name="Pindapoy" pageId="18" pageNumber="116">Pindapoy</location>
,
<date id="FFC61394FFDEFF9B36D0F9C85C65F9D1" box="[1234,1310,1577,1601]" pageId="18" pageNumber="116" value="1955-10">
<collectingDate id="EF82EA7CFFDEFF9B36D0F9C85C65F9D1" box="[1234,1310,1577,1601]" pageId="18" pageNumber="116" value="1955-10">X.1955</collectingDate>
</date>
,
<collectorName id="268D5082FFDEFF9B373BF9C85C17F9D0" box="[1337,1388,1577,1600]" pageId="18" pageNumber="116">Bosq</collectorName>
coll.
</materialsCitation>
,
<materialsCitation id="3B103F09FFDEFF9B3128F9A95D22F90F" collectingDate="1945-11" collectingDateMax="1959-09" collectingDateMin="1945-11" collectionCode="MZUC, CMNC, II" collectorName="ExperimentalStation, A" country="Argentina" location="Pindapoy" pageId="18" pageNumber="116" specimenCount="6" specimenCount-female="5" specimenCount-male="1">
<specimenCount id="9D7EFEDDFFDEFF9B3128F9A95A2AF9CF" box="[810,849,1608,1632]" count="1" pageId="18" pageNumber="116" type="female">1♀</specimenCount>
(
<collectionCode id="ED69AD91FFDEFF9B3168F9A95AC4F9F0" box="[874,959,1608,1632]" country="Chile" httpUri="http://grbio.org/cool/70mh-t5r1" name="Museo de Zoologia, Universidad de Concepcion" pageId="18" pageNumber="116">MZUC</collectionCode>
);
<location id="8EA7638FFFDEFF9B31D9F9A95D40F9F0" LSID="urn:lsid:plazi:treatment:03D18442FFDEFF9C317DFEEC5BB8FC27:8EA7638FFFDEFF9B31D9F9A95D40F9F0" box="[987,1083,1608,1632]" country="Argentina" name="Pindapoy" pageId="18" pageNumber="116">Pindapoy</location>
,
<date id="FFC61394FFDEFF9B364DF9A95DD9F9F0" box="[1103,1186,1608,1632]" pageId="18" pageNumber="116" value="1945-11">
<collectingDate id="EF82EA7CFFDEFF9B364DF9A95DD9F9F0" box="[1103,1186,1608,1632]" pageId="18" pageNumber="116" value="1945-11">XI.1945</collectingDate>
</date>
,
<specimenCount id="9D7EFEDDFFDEFF9B36B5F9A95DA5F9CF" box="[1207,1246,1608,1632]" count="2" pageId="18" pageNumber="116" type="female">2♀</specimenCount>
(
<collectionCode id="ED69AD91FFDEFF9B36F5F9A95C35F9F0" box="[1271,1358,1608,1632]" pageId="18" pageNumber="116">CMNC</collectionCode>
); Loreto,
<collectorName id="268D5082FFDEFF9B3128F9895D63F910" box="[810,1048,1640,1664]" pageId="18" pageNumber="116">ExperimentalStation,A</collectorName>
. Ogloblin,
<specimenCount id="9D7EFEDDFFDEFF9B3683F9895DDCF9EF" box="[1153,1191,1640,1664]" count="2" pageId="18" pageNumber="116" type="female">2♀</specimenCount>
(
<collectionCode id="ED69AD91FFDEFF9B36B0F9895C72F910" box="[1202,1289,1640,1664]" pageId="18" pageNumber="116">CMNC</collectionCode>
); Loreto, without collector,
<date id="FFC61394FFDEFF9B3196F9665A83F90F" box="[916,1016,1671,1695]" pageId="18" pageNumber="116" value="1959-09">
<collectingDate id="EF82EA7CFFDEFF9B3196F9665A83F90F" box="[916,1016,1671,1695]" pageId="18" pageNumber="116" value="1959-09">IX. 1959</collectingDate>
</date>
,
<specimenCount id="9D7EFEDDFFDEFF9B3608F9665D4AF90E" box="[1034,1073,1671,1695]" count="1" pageId="18" pageNumber="116" type="male">1♂</specimenCount>
;
<date id="FFC61394FFDEFF9B3645F9665D22F90F" box="[1095,1113,1671,1695]" pageId="18" pageNumber="116" value="1960-02">
<collectionCode id="ED69AD91FFDEFF9B3645F9665D22F90F" box="[1095,1113,1671,1695]" pageId="18" pageNumber="116">II</collectionCode>
</date>
</materialsCitation>
<materialsCitation id="3B103F09FFDEFF9B365BF9665CCBF94E" collectingDate="1938-01" collectionCode="NHMB, MLPA" collectorName="de Cordoba &amp; C. Bruch" country="Argentina" county="Cordoba" location="Sierras de Cordoba" municipality="Alta Gracia" pageId="18" pageNumber="116" specimenCount="3" specimenCount-female="1" specimenCount-male="2" stateProvince="Entre Rios">
.1960,
<specimenCount id="9D7EFEDDFFDEFF9B36A6F9665DB0F90E" box="[1188,1227,1671,1695]" count="1" pageId="18" pageNumber="116" type="male">1♂</specimenCount>
(
<collectionCode id="ED69AD91FFDEFF9B36E0F9665C43F90F" box="[1250,1336,1671,1695]" country="0" httpUri="http://grbio.org/cool/1d5a-4efs" name="Natural History Museum Bucharest" pageId="18" pageNumber="116">NHMB</collectionCode>
);
<collectingCounty id="62A64DD8FFDEFF9B3750F9665CCBF90F" box="[1362,1456,1671,1695]" pageId="18" pageNumber="116">Córdoba</collectingCounty>
:
<collectingMunicipality id="6BA3AF2EFFDEFF9B3128F9475AE4F92E" box="[810,927,1702,1726]" pageId="18" pageNumber="116">Alta Gracia</collectingMunicipality>
,
<location id="8EA7638FFFDEFF9B31A8F9475D14F92E" LSID="urn:lsid:plazi:treatment:03D18442FFDEFF9C317DFEEC5BB8FC27:8EA7638FFFDEFF9B31A8F9475D14F92E" box="[938,1135,1702,1726]" country="Argentina" county="Cordoba" municipality="Alta Gracia" name="Sierras de Cordoba" pageId="18" pageNumber="116" stateProvince="Entre Rios">
Sierras
<collectorName id="268D5082FFDEFF9B31F6F9475D14F92E" box="[1012,1135,1702,1726]" pageId="18" pageNumber="116">de Córdoba</collectorName>
</location>
,
<date id="FFC61394FFDEFF9B3678F9475DC7F92E" box="[1146,1212,1702,1726]" pageId="18" pageNumber="116" value="1938-01">
<collectingDate id="EF82EA7CFFDEFF9B3678F9475DC7F92E" box="[1146,1212,1702,1726]" pageId="18" pageNumber="116" value="1938-01">I.1938</collectingDate>
</date>
,
<collectorName id="268D5082FFDEFF9B36CAF9475C5EF92E" box="[1224,1317,1702,1726]" pageId="18" pageNumber="116">C. Bruch</collectorName>
,
<specimenCount id="9D7EFEDDFFDEFF9B3733F9475C2CF92D" box="[1329,1367,1702,1726]" count="1" pageId="18" pageNumber="116" type="male">1♂</specimenCount>
and
<specimenCount id="9D7EFEDDFFDEFF9B378EF9475CC8F92D" box="[1420,1459,1702,1726]" count="1" pageId="18" pageNumber="116" type="female">1♀</specimenCount>
(
<collectionCode id="ED69AD91FFDEFF9B3133F9275A04F94E" box="[817,895,1734,1758]" pageId="18" pageNumber="116">MLPA</collectionCode>
);
<collectingRegion id="49BCFBB6FFDEFF9B3191F9275D7DF94E" box="[915,1030,1734,1758]" country="Argentina" name="Entre Rios" pageId="18" pageNumber="116">Entre Ríos</collectingRegion>
:
<location id="8EA7638FFFDEFF9B3610F9275C61F94E" LSID="urn:lsid:plazi:treatment:03D18442FFDEFF9C317DFEEC5BB8FC27:8EA7638FFFDEFF9B3610F9275C61F94E" box="[1042,1306,1734,1758]" country="Argentina" county="Cordoba" municipality="Alta Gracia" name="Departamento Concordia" pageId="18" pageNumber="116" stateProvince="Entre Rios">Departamento Concordia</location>
,
<location id="8EA7638FFFDEFF9B3724F9275CCBF94E" LSID="urn:lsid:plazi:treatment:03D18442FFDEFF9C317DFEEC5BB8FC27:8EA7638FFFDEFF9B3724F9275CCBF94E" box="[1318,1456,1734,1758]" country="Argentina" county="Cordoba" municipality="Alta Gracia" name="Salto Grande" pageId="18" pageNumber="116" stateProvince="Entre Rios">Salto Grande</location>
</materialsCitation>
,
</paragraph>
<paragraph id="8BC73554FFDEFF9D3128F9045B55FF43" blockId="18.[810,1460,1476,1977]" lastBlockId="20.[113,763,155,211]" lastPageId="20" lastPageNumber="118" pageId="18" pageNumber="116">
<materialsCitation id="3B103F09FFDEFF9B3128F9045D55F96D" box="[810,1070,1765,1789]" collectingDate="1979-08" collectionCode="CMNC" pageId="18" pageNumber="116" specimenCount="1" specimenCount-female="1">
<date id="FFC61394FFDEFF9B3128F9045AF5F96D" box="[810,910,1765,1789]" pageId="18" pageNumber="116" value="1979-08">
<collectingDate id="EF82EA7CFFDEFF9B3128F9045AF5F96D" box="[810,910,1765,1789]" pageId="18" pageNumber="116" value="1979-08">VIII.1979</collectingDate>
</date>
,
<specimenCount id="9D7EFEDDFFDEFF9B3198F9045ABBF96C" box="[922,960,1765,1789]" count="1" pageId="18" pageNumber="116" type="female">1♀</specimenCount>
(
<collectionCode id="ED69AD91FFDEFF9B31D2F9045D5CF96D" box="[976,1063,1765,1789]" pageId="18" pageNumber="116">CMNC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B103F09FFDEFF9B363BF9045AF3F88C" collectorName="Departamento Capital &amp; San Jeronimo" location="San Luis" pageId="18" pageNumber="116" specimenCount="1">
<location id="8EA7638FFFDEFF9B363BF9045DEFF96D" LSID="urn:lsid:plazi:treatment:03D18442FFDEFF9C317DFEEC5BB8FC27:8EA7638FFFDEFF9B363BF9045DEFF96D" box="[1081,1172,1765,1789]" name="San Luis" pageId="18" pageNumber="116">San Luis</location>
:
<collectorName id="268D5082FFDEFF9B369DF9045CF8F96D" box="[1183,1411,1765,1789]" pageId="18" pageNumber="116">Departamento Capital</collectorName>
,
<collectorName id="268D5082FFDEFF9B378CF9045AF3F88C" pageId="18" pageNumber="116">San Jerónimo</collectorName>
</materialsCitation>
,
<materialsCitation id="3B103F09FFDEFF9B3196F8E55C62F88C" box="[916,1305,1796,1820]" collectingDate="1979-01" collectionCode="CMNC" collectorName="Guillimer" pageId="18" pageNumber="116" specimenCount="1" specimenCount-female="1">
<date id="FFC61394FFDEFF9B3196F8E55AADF88C" box="[916,982,1796,1820]" pageId="18" pageNumber="116" value="1979-01">
<collectingDate id="EF82EA7CFFDEFF9B3196F8E55AADF88C" box="[916,982,1796,1820]" pageId="18" pageNumber="116" value="1979-01">I.1979</collectingDate>
</date>
,
<collectorName id="268D5082FFDEFF9B31E0F8E55D38F88C" box="[994,1091,1796,1820]" pageId="18" pageNumber="116">Guillimer</collectorName>
coll.,
<specimenCount id="9D7EFEDDFFDEFF9B3681F8E55DD1F88B" box="[1155,1194,1796,1820]" count="1" pageId="18" pageNumber="116" type="female">1♀</specimenCount>
(
<collectionCode id="ED69AD91FFDEFF9B36B8F8E55C6AF88C" box="[1210,1297,1796,1820]" pageId="18" pageNumber="116">CMNC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B103F09FFDEFF9B3727F8E45C70F8AC" collectionCode="BMNH" collectorName="C. Darwin" location="Bahia Blanca" municipality="Buenos Aires" pageId="18" pageNumber="116" specimenCount="9" specimenCount-female="1" specimenCount-male="8">
<collectingMunicipality id="6BA3AF2EFFDEFF9B3727F8E45CCBF88C" box="[1317,1456,1796,1820]" pageId="18" pageNumber="116">Buenos Aires</collectingMunicipality>
:
<location id="8EA7638FFFDEFF9B3128F8C55AD7F8AC" LSID="urn:lsid:plazi:treatment:03D18442FFDEFF9C317DFEEC5BB8FC27:8EA7638FFFDEFF9B3128F8C55AD7F8AC" box="[810,940,1828,1852]" municipality="Buenos Aires" name="Bahia Blanca" pageId="18" pageNumber="116">Bahia Blanca</location>
,
<collectorName id="268D5082FFDEFF9B31B1F8C55D67F8AC" box="[947,1052,1828,1852]" pageId="18" pageNumber="116">C. Darwin</collectorName>
coll.,
<specimenCount id="9D7EFEDDFFDEFF9B3650F8C55D02F8AB" box="[1106,1145,1828,1852]" count="8" pageId="18" pageNumber="116" type="male">8♂</specimenCount>
<specimenCount id="9D7EFEDDFFDEFF9B367BF8C55DDBF8AB" box="[1145,1184,1828,1852]" count="1" pageId="18" pageNumber="116" type="female">1♀</specimenCount>
(
<collectionCode id="ED69AD91FFDEFF9B36AEF8C55C78F8AC" box="[1196,1283,1828,1852]" pageId="18" pageNumber="116">BMNH</collectionCode>
)
</materialsCitation>
; without locality,
<materialsCitation id="3B103F09FFDEFF9B3128F8A25DE1F8CB" box="[810,1178,1859,1883]" collectingDate="1938-01" collectionCode="MLPA" collectorName="C. Bruch" pageId="18" pageNumber="116" specimenCount="1" specimenCount-male="1">
<date id="FFC61394FFDEFF9B3128F8A25A17F8CB" box="[810,876,1859,1883]" pageId="18" pageNumber="116" value="1938-01">
<collectingDate id="EF82EA7CFFDEFF9B3128F8A25A17F8CB" box="[810,876,1859,1883]" pageId="18" pageNumber="116" value="1938-01">I.1938</collectingDate>
</date>
,
<collectorName id="268D5082FFDEFF9B3175F8A25AA8F8CB" box="[887,979,1859,1883]" pageId="18" pageNumber="116">C. Bruch</collectorName>
coll.,
<specimenCount id="9D7EFEDDFFDEFF9B3612F8A25D4DF8CA" box="[1040,1078,1859,1883]" count="1" pageId="18" pageNumber="116" type="male">1♂</specimenCount>
(
<collectionCode id="ED69AD91FFDEFF9B3647F8A25DE8F8CB" box="[1093,1171,1859,1883]" pageId="18" pageNumber="116">MLPA</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B103F09FFDEFF9B36A6F8A25AC7F8EA" collectionCode="ZMHB" collectorName="Frobert" pageId="18" pageNumber="116" specimenCount="1" specimenCount-female="1">
without date,
<collectorName id="268D5082FFDEFF9B3733F8A25C06F8CB" box="[1329,1405,1859,1883]" pageId="18" pageNumber="116">Frobert</collectorName>
coll.,
<specimenCount id="9D7EFEDDFFDEFF9B3128F8835A2AF8E9" box="[810,849,1890,1914]" count="1" pageId="18" pageNumber="116" type="female">1♀</specimenCount>
(
<collectionCode id="ED69AD91FFDEFF9B3163F8835ACFF8EA" box="[865,948,1890,1914]" pageId="18" pageNumber="116">ZMHB</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B103F09FFDEFF9B31CBF8835D21F8EA" box="[969,1114,1890,1914]" collectionCode="II" location="Felipe Sola" pageId="18" pageNumber="116" specimenCount="1">
<location id="8EA7638FFFDEFF9B31CBF8835D47F8EA" LSID="urn:lsid:plazi:treatment:03D18442FFDEFF9C317DFEEC5BB8FC27:8EA7638FFFDEFF9B31CBF8835D47F8EA" box="[969,1084,1890,1914]" name="Felipe Sola" pageId="18" pageNumber="116">Felipe Solá</location>
,
<date id="FFC61394FFDEFF9B364AF8835D21F8EA" box="[1096,1114,1890,1914]" pageId="18" pageNumber="116" value="1942-02">
<collectionCode id="ED69AD91FFDEFF9B364AF8835D21F8EA" box="[1096,1114,1890,1914]" pageId="18" pageNumber="116">II</collectionCode>
</date>
</materialsCitation>
<materialsCitation id="3B103F09FFDEFF9B3658F8835AEAF80A" collectionCode="CMNC" location="Martin Coll." pageId="18" pageNumber="116" specimenCount="7" specimenCount-female="6" specimenCount-male="1">
.1942,
<location id="8EA7638FFFDEFF9B36A2F8835C59F8EA" LSID="urn:lsid:plazi:treatment:03D18442FFDEFF9C317DFEEC5BB8FC27:8EA7638FFFDEFF9B36A2F8835C59F8EA" box="[1184,1314,1890,1914]" name="Martin Coll." pageId="18" pageNumber="116">Martin Coll.</location>
,
<specimenCount id="9D7EFEDDFFDEFF9B372CF8835C2FF8E9" box="[1326,1364,1890,1914]" count="1" pageId="18" pageNumber="116" type="male">1♂</specimenCount>
and
<specimenCount id="9D7EFEDDFFDEFF9B378EF8835CC8F8E9" box="[1420,1459,1890,1914]" count="6" pageId="18" pageNumber="116" type="female">6♀</specimenCount>
(
<collectionCode id="ED69AD91FFDEFF9B3130F8635AF2F80A" box="[818,905,1922,1946]" pageId="18" pageNumber="116">CMNC</collectionCode>
)
</materialsCitation>
; Felipe Solá,
<materialsCitation id="3B103F09FFDEFF9B3616F8635C71F80A" box="[1044,1290,1922,1946]" collectingDate="1944-12" collectionCode="MZUC" pageId="18" pageNumber="116" specimenCount="1" specimenCount-female="1">
<date id="FFC61394FFDEFF9B3616F8635D0BF80A" box="[1044,1136,1922,1946]" pageId="18" pageNumber="116" value="1944-12">
<collectingDate id="EF82EA7CFFDEFF9B3616F8635D0BF80A" box="[1044,1136,1922,1946]" pageId="18" pageNumber="116" value="1944-12">XII.1944</collectingDate>
</date>
,
<specimenCount id="9D7EFEDDFFDEFF9B367BF8635DDBF809" box="[1145,1184,1922,1946]" count="1" pageId="18" pageNumber="116" type="female">1♀</specimenCount>
(
<collectionCode id="ED69AD91FFDEFF9B36ACF8635C78F80A" box="[1198,1283,1922,1946]" country="Chile" httpUri="http://grbio.org/cool/70mh-t5r1" name="Museo de Zoologia, Universidad de Concepcion" pageId="18" pageNumber="116">MZUC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B103F09FFDEFF9B3716F8635C32F80A" box="[1300,1353,1922,1946]" location="Puan" pageId="18" pageNumber="116" specimenCount="1">
<location id="8EA7638FFFDEFF9B3716F8635C32F80A" LSID="urn:lsid:plazi:treatment:03D18442FFDEFF9C317DFEEC5BB8FC27:8EA7638FFFDEFF9B3716F8635C32F80A" box="[1300,1353,1922,1946]" name="Puan" pageId="18" pageNumber="116">Puán</location>
</materialsCitation>
,
<materialsCitation id="3B103F09FFDEFF9B3751F8605DD1F829" collectingDate="1961-12" collectionCode="MZSP" collectorName="Martinezcoll." pageId="18" pageNumber="116" specimenCount="4" specimenCount-female="3" specimenCount-male="1">
<date id="FFC61394FFDEFF9B3751F8605CD4F809" box="[1363,1455,1921,1945]" pageId="18" pageNumber="116" value="1961-12">
<collectingDate id="EF82EA7CFFDEFF9B3751F8605CD4F809" box="[1363,1455,1921,1945]" pageId="18" pageNumber="116" value="1961-12">XII.1961</collectingDate>
</date>
, A.
<collectorName id="268D5082FFDEFF9B3146F8405AB4F829" box="[836,975,1953,1977]" pageId="18" pageNumber="116">Martinezcoll.</collectorName>
,
<specimenCount id="9D7EFEDDFFDEFF9B31D6F8405A80F828" box="[980,1019,1953,1977]" count="1" pageId="18" pageNumber="116" type="male">1♂</specimenCount>
and
<specimenCount id="9D7EFEDDFFDEFF9B3624F8405D37F828" box="[1062,1100,1953,1977]" count="3" pageId="18" pageNumber="116" type="female">3♀</specimenCount>
(
<collectionCode id="ED69AD91FFDEFF9B3655F8405DD9F829" box="[1111,1186,1953,1977]" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="18" pageNumber="116" type="Museum">MZSP</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B103F09FFDEFF9D36B2F8405891FF24" collectionCode="MLUH" collectorName="Darwin" lastPageId="20" lastPageNumber="118" location="Patagonia" pageId="18" pageNumber="116" specimenCount="3" specimenCount-female="3">
<location id="8EA7638FFFDEFF9B36B2F8405C6AF829" LSID="urn:lsid:plazi:treatment:03D18442FFDEFF9C317DFEEC5BB8FC27:8EA7638FFFDEFF9B36B2F8405C6AF829" box="[1200,1297,1953,1977]" name="Patagonia" pageId="18" pageNumber="116">Patagonia</location>
[probablyBahia Blanca],
<collectorName id="268D5082FFD8FF9D32C9FF7A5863FF23" box="[203,280,155,179]" pageId="20" pageNumber="118">Darwin</collectorName>
coll.,
<specimenCount id="9D7EFEDDFFD8FF9D335AFF7D5805FF23" box="[344,382,156,180]" count="3" pageId="20" pageNumber="118" type="female">3♀</specimenCount>
(
<collectionCode id="ED69AD91FFD8FF9D338CFF7D5898FF24" box="[398,483,156,180]" country="Germany" httpUri="http://grbio.org/cool/pvva-bewi" name="Martin Luther Universitaet" pageId="20" pageNumber="118">MLUH</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B103F09FFD8FF9D33F4FF7D5B51FF43" collectionCode="MLPA" collectorName="Tremoleras" country="Uruguay" location="Montevideo" pageId="20" pageNumber="118" specimenCount="1" specimenCount-male="1" stateProvince="Montevideo">
<collectingCountry id="F36F75C4FFD8FF9D33F4FF7D5B0BFF24" box="[502,624,156,180]" name="Uruguay" pageId="20" pageNumber="118">URUGUAY</collectingCountry>
.
<collectingRegion id="49BCFBB6FFD8FF9D3079FF7A5B8CFF23" box="[635,759,155,179]" country="Uruguay" name="Montevideo" pageId="20" pageNumber="118">Montevideo</collectingRegion>
, without date,
<collectorName id="268D5082FFD8FF9D32FBFF5A5811FF43" box="[249,362,187,211]" pageId="20" pageNumber="118">Tremoleras</collectorName>
coll.,
<specimenCount id="9D7EFEDDFFD8FF9D33A1FF5A58B1FF42" box="[419,458,187,211]" count="1" pageId="20" pageNumber="118" type="male">1♂</specimenCount>
(
<collectionCode id="ED69AD91FFD8FF9D33D4FF5A5B58FF43" box="[470,547,187,211]" pageId="20" pageNumber="118">MLPA</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="DF0765DCFFDFFF9A3283F8CD5BCAF8EC" ID-DOI="http://doi.org/10.5281/zenodo.11241108" ID-Zenodo-Dep="11241108" httpUri="https://zenodo.org/record/11241108/files/figure.png" pageId="19" pageNumber="117" startId="19.[129,194,1836,1860]" targetBox="[181,1423,149,1808]" targetPageId="19" targetType="figure">
<paragraph id="8BC73554FFDFFF9A3283F8CD5BCAF8EC" blockId="19.[129,1464,1836,1916]" pageId="19" pageNumber="117">
<emphasis id="B90CE946FFDFFF9A3283F8CD59A2F8D4" bold="true" box="[129,217,1836,1860]" pageId="19" pageNumber="117">Figure 7.</emphasis>
<taxonomicName id="4C784ED7FFDFFF9A32DCF8CC58AEF8D4" authority="(Blanchard)" authorityName="Cherman &amp; Basílio &amp; Clarkson &amp; Agostinis &amp; Smith &amp; Vaz-De-Mello &amp; Almeida" authorityYear="2024" baseAuthorityName="Blanchard" baseAuthorityYear="1851" box="[222,469,1836,1860]" class="Insecta" family="Scarabaeidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="117" phylum="Arthropoda" rank="species" species="niger" status="comb. nov.">
<emphasis id="B90CE946FFDFFF9A32DCF8CC5827F8D4" box="[222,348,1837,1860]" italics="true" pageId="19" pageNumber="117">Liogenys niger</emphasis>
(Blanchard)
</taxonomicName>
<taxonomicNameLabel id="A23F543DFFDFFF9A33D8F8CD5B47F8D4" box="[474,572,1836,1860]" pageId="19" pageNumber="117" rank="species">comb. nov</taxonomicNameLabel>
.. Male neotype habitus (A), labels (E), and complete series at MLUH (D); female lectotype (B) and labels (F); non-uniform body colour variation (C); male in lateral (G) and posterior (I) views; detail of ventrites II and II with medial tubercle (H); parameres in dorsal (J) and lateral (K) views.
</paragraph>
</caption>
<subSubSection id="C36266DFFFD8FF9D3385FF1D5838FA51" pageId="20" pageNumber="118" type="diagnosis">
<paragraph id="8BC73554FFD8FF9D3385FF1D589FFE85" blockId="20.[113,763,252,1473]" box="[391,484,252,277]" pageId="20" pageNumber="118">
<emphasis id="B90CE946FFD8FF9D3385FF1D589FFE85" box="[391,484,252,277]" italics="true" pageId="20" pageNumber="118">Diagnosis</emphasis>
</paragraph>
<paragraph id="8BC73554FFD8FF9D3273FEC05838FA51" blockId="20.[113,763,252,1473]" pageId="20" pageNumber="118">
<emphasis id="B90CE946FFD8FF9D3273FEC059C7FEA9" box="[113,188,289,313]" italics="true" pageId="20" pageNumber="118">Length:</emphasis>
<quantity id="4C8098B1FFD8FF9D32C5FEC0582CFEA9" box="[199,343,289,313]" metricMagnitude="-3" metricUnit="m" metricValue="9.6" metricValueMax="10.5" metricValueMin="8.7" pageId="20" pageNumber="118" unit="mm" value="9.6" valueMax="10.5" valueMin="8.7">8.7–10.5 mm</quantity>
; width:
<quantity id="4C8098B1FFD8FF9D33B1FEC05B4FFEA9" box="[435,564,289,313]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" metricValueMax="6.2" metricValueMin="4.8" pageId="20" pageNumber="118" unit="mm" value="5.5" valueMax="6.2" valueMin="4.8">4.8–6.2 mm</quantity>
. Body short, oval; body and elytra in general black (
<figureCitation id="134329D1FFD8FF9D33DAFEA05B39FEC8" box="[472,578,321,345]" captionStart="Figure 7" captionStartId="19.[129,194,1836,1860]" captionTargetBox="[181,1423,149,1808]" captionTargetId="figure-7@19.[181,1423,144,1808]" captionTargetPageId="19" captionText="Figure 7. Liogenys niger (Blanchard) comb. nov.. Male neotype habitus (A), labels (E), and complete series at MLUH (D); female lectotype (B) and labels (F); non-uniform body colour variation (C); male in lateral (G) and posterior (I) views; detail of ventrites II and II with medial tubercle (H); parameres in dorsal (J) and lateral (K) views." figureDoi="http://doi.org/10.5281/zenodo.11241108" httpUri="https://zenodo.org/record/11241108/files/figure.png" pageId="20" pageNumber="118">Fig. 7A, B</figureCitation>
), elytra glabrous, eventually reddish-brown on disc and black on scutellum and surrounding (
<figureCitation id="134329D1FFD8FF9D32FDFE9E5831FE08" box="[255,330,383,408]" captionStart="Figure 7" captionStartId="19.[129,194,1836,1860]" captionTargetBox="[181,1423,149,1808]" captionTargetId="figure-7@19.[181,1423,144,1808]" captionTargetPageId="19" captionText="Figure 7. Liogenys niger (Blanchard) comb. nov.. Male neotype habitus (A), labels (E), and complete series at MLUH (D); female lectotype (B) and labels (F); non-uniform body colour variation (C); male in lateral (G) and posterior (I) views; detail of ventrites II and II with medial tubercle (H); parameres in dorsal (J) and lateral (K) views." figureDoi="http://doi.org/10.5281/zenodo.11241108" httpUri="https://zenodo.org/record/11241108/files/figure.png" pageId="20" pageNumber="118">Fig. 7C</figureCitation>
); shiny or semiopaque, coarsely punctate; pronotum with slight iridescent green reflections; distance between eyes more than five times wider than one eye; clypeus and frons almost coplanar; clypeus very wide, subemarginate; clypeal emargination small, shallow, rounded, and narrow; clypeal lateral margin convex; ocular canthus exceeding the outer margin of the eye; galea of maxilla with four strong teeth; distal maxillary palpomere oval, fovea very shallow, hardly reaching the midline of the palpomere; labium transversely carinate, as wide as it is long; two labial palpomeres equal in length; antenna with 10 antennomeres; club unicolorous with the funicle and as long as; pronotal anterior margin concave and slightly produced medially; pronotal disc strongly and coarsely punctate, posterior corners rounded; distance between mesocoxae and metacoxae slightly longer than metacoxa; scutellum wide; elytra convex dorsoventrally (
<figureCitation id="134329D1FFD8FF9D32B6FCB45985FCFD" box="[180,254,853,877]" captionStart="Figure 7" captionStartId="19.[129,194,1836,1860]" captionTargetBox="[181,1423,149,1808]" captionTargetId="figure-7@19.[181,1423,144,1808]" captionTargetPageId="19" captionText="Figure 7. Liogenys niger (Blanchard) comb. nov.. Male neotype habitus (A), labels (E), and complete series at MLUH (D); female lectotype (B) and labels (F); non-uniform body colour variation (C); male in lateral (G) and posterior (I) views; detail of ventrites II and II with medial tubercle (H); parameres in dorsal (J) and lateral (K) views." figureDoi="http://doi.org/10.5281/zenodo.11241108" httpUri="https://zenodo.org/record/11241108/files/figure.png" pageId="20" pageNumber="118">Fig. 7G</figureCitation>
), nearly 2.5 times longer than the pronotum; three elytral ridges noticeable (I,
<collectionCode id="ED69AD91FFD8FF9D3390FC9458DEFC1C" box="[402,421,885,908]" pageId="20" pageNumber="118">II</collectionCode>
, and IV); subapical callus above the level of the distal inner corner of the sutural ridge; mesotibia cylindrical and with transverse carina complete in both sexes; metatibial spurs slightly uneven, the longest slightly shorter than the diameter of the tibial apex; ventrites
<collectionCode id="ED69AD91FFD8FF9D300DFC135B5AFB99" box="[527,545,1010,1033]" pageId="20" pageNumber="118">II</collectionCode>
and
<collectionCode id="ED69AD91FFD8FF9D3051FC135B16FB99" box="[595,621,1010,1033]" pageId="20" pageNumber="118">III</collectionCode>
with tubercle at the midline (
<figureCitation id="134329D1FFD8FF9D3313FBF05825FBB9" box="[273,350,1041,1065]" captionStart="Figure 7" captionStartId="19.[129,194,1836,1860]" captionTargetBox="[181,1423,149,1808]" captionTargetId="figure-7@19.[181,1423,144,1808]" captionTargetPageId="19" captionText="Figure 7. Liogenys niger (Blanchard) comb. nov.. Male neotype habitus (A), labels (E), and complete series at MLUH (D); female lectotype (B) and labels (F); non-uniform body colour variation (C); male in lateral (G) and posterior (I) views; detail of ventrites II and II with medial tubercle (H); parameres in dorsal (J) and lateral (K) views." figureDoi="http://doi.org/10.5281/zenodo.11241108" httpUri="https://zenodo.org/record/11241108/files/figure.png" pageId="20" pageNumber="118">Fig. 7H</figureCitation>
); propygidium glabrous, almost entirely covered by the elytra; pygidium convex, especially in females; wide; subtrapezoidal; pygidial disc glabrous, coarse umbilicate punctures (
<figureCitation id="134329D1FFD8FF9D32E5FB8E5852FB17" box="[231,297,1135,1159]" captionStart="Figure 7" captionStartId="19.[129,194,1836,1860]" captionTargetBox="[181,1423,149,1808]" captionTargetId="figure-7@19.[181,1423,144,1808]" captionTargetPageId="19" captionText="Figure 7. Liogenys niger (Blanchard) comb. nov.. Male neotype habitus (A), labels (E), and complete series at MLUH (D); female lectotype (B) and labels (F); non-uniform body colour variation (C); male in lateral (G) and posterior (I) views; detail of ventrites II and II with medial tubercle (H); parameres in dorsal (J) and lateral (K) views." figureDoi="http://doi.org/10.5281/zenodo.11241108" httpUri="https://zenodo.org/record/11241108/files/figure.png" pageId="20" pageNumber="118">Fig. 7I</figureCitation>
); in males inner margin of metatibia strongly carinate and produced from medial portion towards apex; inner surface with dense, thick bristles, bristles almost as long as the metatibial width; pro- and mesotarsomeres I to IV enlarged; protarsomere
<collectionCode id="ED69AD91FFD8FF9D3302FB0C5869FA94" box="[256,274,1261,1284]" pageId="20" pageNumber="118">II</collectionCode>
as wide as it is long; claw bifid on all legs, symmetrical; basal region of parameres long, more or less compressed proximally, abruptly bulged subbasally, swollen, and narrowed at the midline; parameral split after the midline; longitudinal sulcus deep; inner margins of parameres straight; expanded apically, apical margin rounded; parameres in lateral view slightly concave, coplanar (
<figureCitation id="134329D1FFD8FF9D32DAFA495849FA50" box="[216,306,1448,1472]" captionStart="Figure 7" captionStartId="19.[129,194,1836,1860]" captionTargetBox="[181,1423,149,1808]" captionTargetId="figure-7@19.[181,1423,144,1808]" captionTargetPageId="19" captionText="Figure 7. Liogenys niger (Blanchard) comb. nov.. Male neotype habitus (A), labels (E), and complete series at MLUH (D); female lectotype (B) and labels (F); non-uniform body colour variation (C); male in lateral (G) and posterior (I) views; detail of ventrites II and II with medial tubercle (H); parameres in dorsal (J) and lateral (K) views." figureDoi="http://doi.org/10.5281/zenodo.11241108" httpUri="https://zenodo.org/record/11241108/files/figure.png" pageId="20" pageNumber="118">Fig. 7J, K</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C36266DFFFD8FF9D3379FA0D5B08F9D9" pageId="20" pageNumber="118" type="description">
<paragraph id="8BC73554FFD8FF9D3379FA0D588AF996" blockId="20.[113,763,1516,1610]" box="[379,497,1516,1542]" pageId="20" pageNumber="118">
<emphasis id="B90CE946FFD8FF9D3379FA0D588AF996" box="[379,497,1516,1542]" italics="true" pageId="20" pageNumber="118">Type locality</emphasis>
</paragraph>
<paragraph id="8BC73554FFD8FF9D3273F9F35B08F9D9" blockId="20.[113,763,1516,1610]" pageId="20" pageNumber="118">
<emphasis id="B90CE946FFD8FF9D3273F9F3584DF9BA" box="[113,310,1554,1578]" italics="true" pageId="20" pageNumber="118">
<taxonomicName id="4C784ED7FFD8FF9D3273F9F35855F9BA" authorityName="Blanchard" authorityYear="1851" box="[113,302,1554,1578]" class="Insecta" family="Scarabaeidae" genus="Homalochilus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="118" phylum="Arthropoda" rank="species" species="niger">Homalochilus niger</taxonomicName>
:
</emphasis>
<collectingCountry id="F36F75C4FFD8FF9D333EF9F358ACF9BA" box="[316,471,1554,1578]" name="Argentina" pageId="20" pageNumber="118">ARGENTINA</collectingCountry>
.
<collectingRegion id="49BCFBB6FFD8FF9D33E3F9F25B10F9BA" box="[481,619,1554,1578]" country="Argentina" name="Ciudad Autonoma de Buenos Aires" pageId="20" pageNumber="118">Buenos Aires</collectingRegion>
, Bahía de San Blas.
<taxonomicName id="4C784ED7FFD8FF9D32A4F9D35842F9D9" box="[166,313,1586,1609]" class="Insecta" family="Scarabaeidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="118" phylum="Arthropoda" rank="species" species="morio">
<emphasis id="B90CE946FFD8FF9D32A4F9D35842F9D9" box="[166,313,1586,1609]" italics="true" pageId="20" pageNumber="118">Liogenys morio</emphasis>
</taxonomicName>
:
<collectingCountry id="F36F75C4FFD8FF9D3347F9D3589BF9DA" box="[325,480,1586,1610]" name="Argentina" pageId="20" pageNumber="118">ARGENTINA</collectingCountry>
. Bahia Blanca.
</paragraph>
</subSubSection>
<subSubSection id="C36266DFFFD8FF9D333DF9885B80F956" pageId="20" pageNumber="118" type="distribution">
<paragraph id="8BC73554FFD8FF9D333DF9885B56F913" blockId="20.[113,763,1641,1735]" box="[319,557,1641,1667]" pageId="20" pageNumber="118">
<emphasis id="B90CE946FFD8FF9D333DF9885B56F913" box="[319,557,1641,1667]" italics="true" pageId="20" pageNumber="118">Geographical distribution</emphasis>
</paragraph>
<paragraph id="8BC73554FFD8FF9D3273F96E5B80F956" blockId="20.[113,763,1641,1735]" pageId="20" pageNumber="118">
<emphasis id="B90CE946FFD8FF9D3273F96E582DF937" bold="true" box="[113,342,1679,1703]" pageId="20" pageNumber="118">
<collectingCountry id="F36F75C4FFD8FF9D3273F96E59B4F937" box="[113,207,1679,1703]" name="Brazil" pageId="20" pageNumber="118">BRAZIL</collectingCountry>
(
<collectingRegion id="49BCFBB6FFD8FF9D32DFF96E5834F937" box="[221,335,1679,1703]" country="Brazil" name="Sao Paulo" pageId="20" pageNumber="118">São Paulo</collectingRegion>
)
</emphasis>
; ARGENTINA
<emphasis id="B90CE946FFD8FF9D3000F96E5845F957" bold="true" pageId="20" pageNumber="118">(Misiones, Entre Ríos, Córdoba, San Luis,</emphasis>
Buenos Aires);
<emphasis id="B90CE946FFD8FF9D33D9F94E5B8FF956" bold="true" box="[475,756,1710,1735]" pageId="20" pageNumber="118">
<collectingCountry id="F36F75C4FFD8FF9D33D9F94E5B22F957" box="[475,601,1711,1735]" name="Uruguay" pageId="20" pageNumber="118">URUGUAY</collectingCountry>
(
<collectingRegion id="49BCFBB6FFD8FF9D3061F94F5B96F956" box="[611,749,1710,1734]" country="Uruguay" name="Montevideo" pageId="20" pageNumber="118">Montevideo</collectingRegion>
)
</emphasis>
.
</paragraph>
</subSubSection>
<subSubSection id="C36266DFFFD8FF9C3294F9065BB8FC27" lastPageId="21" lastPageNumber="119" pageId="20" pageNumber="118" type="discussion">
<paragraph id="8BC73554FFD8FF9D3294F9065BAEF890" blockId="20.[150,725,1766,1792]" box="[150,725,1766,1792]" pageId="20" pageNumber="118">
<emphasis id="B90CE946FFD8FF9D3294F9065BAEF890" box="[150,725,1766,1792]" italics="true" pageId="20" pageNumber="118">
<taxonomicName id="4C784ED7FFD8FF9D3294F9065853F890" authorityName="Cherman &amp; Basílio &amp; Clarkson &amp; Agostinis &amp; Smith &amp; Vaz-De-Mello &amp; Almeida" authorityYear="2024" baseAuthorityName="Blanchard" baseAuthorityYear="1851" box="[150,296,1767,1792]" class="Insecta" family="Scarabaeidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="118" phylum="Arthropoda" rank="species" species="niger">Liogenys niger</taxonomicName>
taxonomic history and remarks on systematics
</emphasis>
</paragraph>
<paragraph id="8BC73554FFD8FF9D3273F8EC5B51F851" blockId="20.[113,762,1805,1985]" pageId="20" pageNumber="118">
<bibRefCitation id="EFE948A5FFD8FF9D3273F8EC584BF8B5" author="Blanchard ME" box="[113,304,1805,1829]" pageId="20" pageNumber="118" refId="ref26799" refString="Blanchard ME. Museum d'Histoire Naturelle de Paris. Catalogue de la Collection Entomologique. Classe des Insectes. Ordre des Coleopteres. Tome I. Part 2. Paris: Gide et Baudry, 1851." type="book" year="1851">Blanchard (1851)</bibRefCitation>
described
<taxonomicName id="4C784ED7FFD8FF9D33ADF8EC5B09F8B5" authorityName="Blanchard" authorityYear="1851" box="[431,626,1805,1829]" class="Insecta" family="Scarabaeidae" genus="Homalochilus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="118" phylum="Arthropoda" rank="species" species="niger">
<emphasis id="B90CE946FFD8FF9D33ADF8EC5B09F8B5" box="[431,626,1805,1829]" italics="true" pageId="20" pageNumber="118">Homalochilus niger</emphasis>
</taxonomicName>
based on a series of
<specimenCount id="9D7EFEDDFFD8FF9D32C8F8CC5838F8D4" box="[202,323,1836,1860]" count="2" pageId="20" pageNumber="118" type="female">two females</specimenCount>
collected by A.D. d’Orbigny from
<collectingRegion id="49BCFBB6FFD8FF9D30A0F8CD5BA7F8D4" box="[674,732,1836,1860]" country="Brazil" name="Bahia" pageId="20" pageNumber="118">Bahía</collectingRegion>
de San Blas,
<collectingRegion id="49BCFBB6FFD8FF9D32D5F8AD5819F8F3" box="[215,354,1867,1891]" country="Argentina" name="Ciudad Autonoma de Buenos Aires" pageId="20" pageNumber="118">Buenos Aires</collectingRegion>
. The features mentioned by the author were the body coarsely punctate, entirely black; clypeus poorly emarginated; scutellum almost smooth; elytra short and wide, convex; protibiae tridentate and tarsi black.
</paragraph>
<paragraph id="8BC73554FFD8FF9D3147FF715DDBFDB0" blockId="20.[810,1460,144,1985]" pageId="20" pageNumber="118">
<bibRefCitation id="EFE948A5FFD8FF9D3147FF715D70FF38" author="Burmeister H" box="[837,1035,144,168]" pageId="20" pageNumber="118" refId="ref26916" refString="Burmeister H. Handbuch der entomologie (Coleoptera Lamellicornia Phyllophaga Chaenochela). Vol. 4, Part 2. Berlin: Enslin, T. C. F., 1855." type="book" year="1855">Burmeister (1855)</bibRefCitation>
described a male specimen of
<taxonomicName id="4C784ED7FFD8FF9D375DFF705A1FFF57" class="Insecta" family="Scarabaeidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="118" phylum="Arthropoda" rank="species" species="morio">
<emphasis id="B90CE946FFD8FF9D375DFF705A1FFF57" italics="true" pageId="20" pageNumber="118">Liogenys morio</emphasis>
</taxonomicName>
as oval, shiny, punctate; black with copper reflections on pronotum; and clypeus emarginated, subindented. He stated that the species resembles
<taxonomicName id="4C784ED7FFD8FF9D368BFF0E5C2DFE96" authority="Burmeister" authorityName="Burmeister" authorityYear="1855" box="[1161,1366,238,262]" class="Insecta" family="Scarabaeidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="118" phylum="Arthropoda" rank="species" species="obesa">
<emphasis id="B90CE946FFD8FF9D368BFF0E5DA7FE96" box="[1161,1244,238,262]" italics="true" pageId="20" pageNumber="118">L. obesa</emphasis>
Burmeister
</taxonomicName>
(current synonym of
<taxonomicName id="4C784ED7FFD8FF9D31ABFEEF5DF8FEB5" authority="Blanchard" authorityName="Blanchard" authorityYear="1850" box="[937,1155,269,293]" class="Insecta" family="Scarabaeidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="118" phylum="Arthropoda" rank="species" species="concolor">
<emphasis id="B90CE946FFD8FF9D31ABFEEF5D69FEB5" box="[937,1042,269,293]" italics="true" pageId="20" pageNumber="118">L. concolor</emphasis>
Blanchard
</taxonomicName>
) (
<bibRefCitation id="EFE948A5FFD8FF9D3698FEEC5C13FEB5" author="Cherman MA &amp; Mise KM &amp; Moron MA" box="[1178,1384,269,293]" pageId="20" pageNumber="118" pagination="1 - 120" refId="ref27057" refString="Cherman MA, Mise KM, Moron MA et al. A taxonomic revision of Liogenys occurring in Brazil with an interactive key and remarks on New World Diplotaxini (Coleoptera, Melolonthidae). ZooKeys 2017; 699: 1 - 120." type="journal article" year="2017">
Cherman
<emphasis id="B90CE946FFD8FF9D3702FEEF5C54FEB5" box="[1280,1327,269,293]" italics="true" pageId="20" pageNumber="118">et al.</emphasis>
2017
</bibRefCitation>
) in the short, convex, oval body. This short body is due to the distance between mesocoxae and metacoxae slightly longer than the metacoxa, an uncommon feature in most
<taxonomicName id="4C784ED7FFD8FF9D36D4FE8D5C51FE13" box="[1238,1322,364,387]" class="Insecta" family="Scarabaeidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="118" phylum="Arthropoda" rank="genus">
<emphasis id="B90CE946FFD8FF9D36D4FE8D5C51FE13" box="[1238,1322,364,387]" italics="true" pageId="20" pageNumber="118">Liogenys</emphasis>
</taxonomicName>
(
<bibRefCitation id="EFE948A5FFD8FF9D3738FE8A5A07FE32" author="Cherman MA &amp; Moron MA &amp; Almeida LM" pageId="20" pageNumber="118" pagination="744 - 70" refId="ref27020" refString="Cherman MA, Moron MA, Almeida LM. Phylogenetic relationships within Diplotaxini Kirby (Coleoptera: Melolonthidae: Melolonthinae) with emphasis on Liogenys Guerin-Meneville. Systematic Entomology 2016; 41: 744 - 70." type="journal article" year="2016">
Cherman
<emphasis id="B90CE946FFD8FF9D37A0FE8D5A38FE32" italics="true" pageId="20" pageNumber="118">et al.</emphasis>
2016
</bibRefCitation>
,
<bibRefCitation id="EFE948A5FFD8FF9D318BFE6A5AC6FE33" author="Cherman MA &amp; Mise KM &amp; Moron MA" box="[905,957,395,419]" pageId="20" pageNumber="118" pagination="1 - 120" refId="ref27057" refString="Cherman MA, Mise KM, Moron MA et al. A taxonomic revision of Liogenys occurring in Brazil with an interactive key and remarks on New World Diplotaxini (Coleoptera, Melolonthidae). ZooKeys 2017; 699: 1 - 120." type="journal article" year="2017">2017</bibRefCitation>
) which have this distance two times longer than the metacoxa. Other morphological features that these species share are the elytra convex, less than three times the length of the pronotum; distal maxillary palpomere oval; and antenna almost entirely black (
<bibRefCitation id="EFE948A5FFD8FF9D31C0FDE95DF4FDB0" author="Cherman MA &amp; Mise KM &amp; Moron MA" box="[962,1167,520,544]" pageId="20" pageNumber="118" pagination="1 - 120" refId="ref27057" refString="Cherman MA, Mise KM, Moron MA et al. A taxonomic revision of Liogenys occurring in Brazil with an interactive key and remarks on New World Diplotaxini (Coleoptera, Melolonthidae). ZooKeys 2017; 699: 1 - 120." type="journal article" year="2017">
Cherman
<emphasis id="B90CE946FFD8FF9D362AFDE85D2CFDB0" box="[1064,1111,520,544]" italics="true" pageId="20" pageNumber="118">et al.</emphasis>
2017
</bibRefCitation>
).
</paragraph>
<paragraph id="8BC73554FFD8FF9D3147FDC65DC3FDCE" blockId="20.[810,1460,144,1985]" pageId="20" pageNumber="118">
<bibRefCitation id="EFE948A5FFD8FF9D3147FDC65AA9FDAF" author="Bruch C" box="[837,978,551,575]" pageId="20" pageNumber="118" pagination="181 - 225" refId="ref26889" refString="Bruch C. Catalogo sistematico de los coleopteros de la Republica Argentina. Revista del Museo de la Plata 1911; 17: 181 - 225." type="journal article" year="1911">Bruch (1911)</bibRefCitation>
expanded the distribution of
<taxonomicName id="4C784ED7FFD8FF9D36FAFDC95C31FDAF" box="[1272,1354,552,575]" class="Insecta" family="Scarabaeidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="118" phylum="Arthropoda" rank="species" species="morio">
<emphasis id="B90CE946FFD8FF9D36FAFDC95C31FDAF" box="[1272,1354,552,575]" italics="true" pageId="20" pageNumber="118">L. morio</emphasis>
</taxonomicName>
to
<collectingRegion id="49BCFBB6FFD8FF9D3764FDC95D6FFDCE" isEnumeration="true" pageId="20" pageNumber="118">Buenos Aires, Cordoba, Chaco</collectingRegion>
, and ‘Patagonia’.
</paragraph>
<paragraph id="8BC73554FFD8FF9D3147FD875AE5FD2C" blockId="20.[810,1460,144,1985]" pageId="20" pageNumber="118">
<bibRefCitation id="EFE948A5FFD8FF9D3147FD875ABDFDEE" author="Frey G" box="[837,966,614,638]" pageId="20" pageNumber="118" pagination="38 - 64" refId="ref27637" refString="Frey G. Bestimmungstabelle und revision der gattung Liogenys Guerrez (Coleoptera - Melolonthinae - Macrodactylini). Entomologische Arbeiten Aus Dem Museum G. Frey 1969; 20: 38 - 64." type="journal article" year="1969">Frey (1969)</bibRefCitation>
stated that the
<typeStatus id="54C38BF6FFD8FF9D366FFD875DE0FDEE" box="[1133,1179,614,638]" pageId="20" pageNumber="118">type</typeStatus>
of
<taxonomicName id="4C784ED7FFD8FF9D36C3FD865C6CFDEE" box="[1217,1303,615,638]" class="Insecta" family="Scarabaeidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="118" phylum="Arthropoda" rank="species" species="morio">
<emphasis id="B90CE946FFD8FF9D36C3FD865C6CFDEE" box="[1217,1303,615,638]" italics="true" pageId="20" pageNumber="118">L. morio</emphasis>
</taxonomicName>
had been lost, and therefore he was unable to treat the species in his revision of
<taxonomicName id="4C784ED7FFD8FF9D3146FD445AE3FD2C" box="[836,920,677,700]" class="Insecta" family="Scarabaeidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="118" phylum="Arthropoda" rank="genus">
<emphasis id="B90CE946FFD8FF9D3146FD445AE3FD2C" box="[836,920,677,700]" italics="true" pageId="20" pageNumber="118">Liogenys</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BC73554FFD8FF9D3147FD255C54F8B4" blockId="20.[810,1460,144,1985]" pageId="20" pageNumber="118">
At
<collectionCode id="ED69AD91FFD8FF9D3167FD255AC8FD4B" box="[869,947,708,731]" country="Germany" httpUri="http://grbio.org/cool/pvva-bewi" name="Martin Luther Universitaet" pageId="20" pageNumber="118">MLUH</collectionCode>
we found a series of
<specimenCount id="9D7EFEDDFFD8FF9D3690FD255D87FD4C" box="[1170,1276,708,732]" count="2" pageId="20" pageNumber="118" type="male">two males</specimenCount>
and
<specimenCount id="9D7EFEDDFFD8FF9D3733FD255CD4FD4C" box="[1329,1455,708,732]" count="2" pageId="20" pageNumber="118" type="female">two females</specimenCount>
, accompanied by the typical green square label of Burmeister’s types at the bottom of the drawer (
<figureCitation id="134329D1FFD8FF9D36A9FCE35D81FC8A" box="[1195,1274,770,794]" captionStart="Figure 7" captionStartId="19.[129,194,1836,1860]" captionTargetBox="[181,1423,149,1808]" captionTargetId="figure-7@19.[181,1423,144,1808]" captionTargetPageId="19" captionText="Figure 7. Liogenys niger (Blanchard) comb. nov.. Male neotype habitus (A), labels (E), and complete series at MLUH (D); female lectotype (B) and labels (F); non-uniform body colour variation (C); male in lateral (G) and posterior (I) views; detail of ventrites II and II with medial tubercle (H); parameres in dorsal (J) and lateral (K) views." figureDoi="http://doi.org/10.5281/zenodo.11241108" httpUri="https://zenodo.org/record/11241108/files/figure.png" pageId="20" pageNumber="118">Fig. 7D</figureCitation>
). On the label it is handwritten:
<taxonomicName id="4C784ED7FFD8FF9D31E3FCC35D08FCAA" authority="Burm" authorityName="Burm" box="[993,1139,802,826]" class="Insecta" family="Scarabaeidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="118" phylum="Arthropoda" rank="species" species="morio">‘morio Burm</taxonomicName>
/Bras [
<collectingCountry id="F36F75C4FFD8FF9D36C3FCC35C7EFCAA" box="[1217,1285,802,826]" name="Brazil" pageId="20" pageNumber="118">Brazil</collectingCountry>
] Dp [probably Dupont]’, and this locality does not match with the one stated at the original description: ‘Mittel Amerika’. Moreover, two out of the
<specimenCount id="9D7EFEDDFFD8FF9D3173FC615D6AFC08" box="[881,1041,896,920]" count="4" pageId="20" pageNumber="118" type="generic">four specimens</specimenCount>
bear a handwritten, white label: ‘
<collectingRegion id="49BCFBB6FFD8FF9D3775FC615CC8FC08" box="[1399,1459,896,920]" country="Brazil" name="Bahia" pageId="20" pageNumber="118">Bahia</collectingRegion>
Blanca’ and ‘Patagonia, Darwin’, respectively, which does not match either with the description, or the drawer’s label. In the original ‘Handbuch’ housed at the
<collectionCode id="ED69AD91FFD8FF9D3695FC3F5D9CFC65" box="[1175,1255,990,1013]" country="Germany" httpUri="http://grbio.org/cool/pvva-bewi" name="Martin Luther Universitaet" pageId="20" pageNumber="118">MLUH</collectionCode>
, ‘Mittel Amerika’ is struck through and ‘Patagonien’ has been handwritten above. Using handwriting comparisons, this information might have been written by Harold (
<bibRefCitation id="EFE948A5FFD8FF9D363FFBDD5C56FBC4" author="Horn W &amp; Kahle I" box="[1085,1325,1084,1108]" pageId="20" pageNumber="118" pagination="1935 - 37" refId="ref27901" refString="Horn W, Kahle I. Uber entomologische Sammlungen, Entomologen &amp; Entomo-Museologie. Entomologische Beihefte, Band 2 - 4, 1935 - 37. Berlin: Deutsches Entomologisches Institut, 1937." type="journal article" year="1937">Horn and Kahle 1937</bibRefCitation>
). Moreover, this locality record was used consistently since Harold (1869: 1141), so we can infer that the series we found has been there, at least, since that date. Bates (1887) mentioned that no specimens were found in Central America matching with
<taxonomicName id="4C784ED7FFD8FF9D3755FB5B5CD6FB41" box="[1367,1453,1210,1233]" class="Insecta" family="Scarabaeidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="118" phylum="Arthropoda" rank="species" species="morio">
<emphasis id="B90CE946FFD8FF9D3755FB5B5CD6FB41" box="[1367,1453,1210,1233]" italics="true" pageId="20" pageNumber="118">L. morio</emphasis>
</taxonomicName>
. There is another species,
<taxonomicName id="4C784ED7FFD8FF9D3644FB395D68FA80" authority="(Burmeister) (Cherman et al. 2016)" authorityName="Cherman et al." authorityYear="2016" baseAuthorityName="Burmeister" baseAuthorityYear="1855" class="Insecta" family="Scarabaeidae" genus="Diplotaxis" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="118" phylum="Arthropoda" rank="species" species="micropyga">
<emphasis id="B90CE946FFD8FF9D3644FB395C64FB60" box="[1094,1311,1240,1264]" italics="true" pageId="20" pageNumber="118">Diplotaxis micropyga</emphasis>
(Burmeister) (
<bibRefCitation id="EFE948A5FFD8FF9D3136FB195D7CFA80" author="Cherman MA &amp; Moron MA &amp; Almeida LM" box="[820,1031,1272,1296]" pageId="20" pageNumber="118" pagination="744 - 70" refId="ref27020" refString="Cherman MA, Moron MA, Almeida LM. Phylogenetic relationships within Diplotaxini Kirby (Coleoptera: Melolonthidae: Melolonthinae) with emphasis on Liogenys Guerin-Meneville. Systematic Entomology 2016; 41: 744 - 70." type="journal article" year="2016">
Cherman
<emphasis id="B90CE946FFD8FF9D319FFB195AB5FA80" box="[925,974,1272,1296]" italics="true" pageId="20" pageNumber="118">et al.</emphasis>
2016
</bibRefCitation>
)
</taxonomicName>
, which was described even on the same page as
<taxonomicName id="4C784ED7FFD8FF9D3180FAF95AA0FABF" box="[898,987,1304,1327]" class="Insecta" family="Scarabaeidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="118" phylum="Arthropoda" rank="species" species="morio">
<emphasis id="B90CE946FFD8FF9D3180FAF95AA0FABF" box="[898,987,1304,1327]" italics="true" pageId="20" pageNumber="118">L. morio</emphasis>
</taxonomicName>
and its type locality is ‘Brasil’.
<taxonomicName id="4C784ED7FFD8FF9D3730FAF65CE2FABF" authorityName="Kirby" authorityYear="1837" box="[1330,1433,1303,1327]" class="Insecta" family="Scarabaeidae" genus="Diplotaxis" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="118" phylum="Arthropoda" rank="genus">
<emphasis id="B90CE946FFD8FF9D3730FAF65CE2FABF" box="[1330,1433,1303,1327]" italics="true" pageId="20" pageNumber="118">Diplotaxis</emphasis>
</taxonomicName>
is mostly Nearctic, although one species was recently recorded in
<collectingCountry id="F36F75C4FFD8FF9D3128FAB75AEFFAFE" box="[810,916,1366,1390]" name="Colombia" pageId="20" pageNumber="118">Colombia</collectingCountry>
, becoming its southernmost occurrence (
<bibRefCitation id="EFE948A5FFD8FF9D3750FAB75D4BFA1D" author="Mendoza IA &amp; Garcia-Atencia SP" pageId="20" pageNumber="118" pagination="435 - 9" refId="ref28381" refString="Mendoza IA, Garcia-Atencia SP. First record of the genus Diplotaxis Kirby, 1837 (Coleoptera, scarabaeidae, diplotaxini) in South America. Check List 2019; 15: 435 - 9." type="journal article" year="2019">Mendoza and García-Atencia 2019</bibRefCitation>
). Germar wrote the square labels to Burmeister, so he might have accidentally switched the type labels of
<taxonomicName id="4C784ED7FFD8FF9D318CFA555D69FA5B" authorityName="Cherman et al." authorityYear="2016" baseAuthorityName="Burmeister" baseAuthorityYear="1855" box="[910,1042,1460,1483]" class="Insecta" family="Scarabaeidae" genus="Diplotaxis" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="118" phylum="Arthropoda" rank="species" species="micropyga">
<emphasis id="B90CE946FFD8FF9D318CFA555D69FA5B" box="[910,1042,1460,1483]" italics="true" pageId="20" pageNumber="118">L. micropyga</emphasis>
</taxonomicName>
and
<taxonomicName id="4C784ED7FFD8FF9D364EFA555DDEFA5B" box="[1100,1189,1460,1483]" class="Insecta" family="Scarabaeidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="118" phylum="Arthropoda" rank="species" species="morio">
<emphasis id="B90CE946FFD8FF9D364EFA555DDEFA5B" box="[1100,1189,1460,1483]" italics="true" pageId="20" pageNumber="118">L. morio</emphasis>
</taxonomicName>
. In the description of
<taxonomicName id="4C784ED7FFD8FF9D379DFA555A98FA7B" authority=", Burmeister" authorityName="Burmeister" class="Insecta" family="Scarabaeidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="118" phylum="Arthropoda" rank="species" species="morio">
<emphasis id="B90CE946FFD8FF9D379DFA555A1FFA7B" italics="true" pageId="20" pageNumber="118">L. morio</emphasis>
, Burmeister
</taxonomicName>
did not mention any information about who was either the collector or the owner of the material. However, judging from what is written in the green label of the drawer, Burmeister received those specimens from Henry Dupont (1798–1873), who was a prolific dealer of beetles during the first half of the nineteenth century, and probably the seller of Darwin’s specimens deposited at the
<collectionCode id="ED69AD91FFD8FF9D36A7F96E5D8EF936" box="[1189,1269,1679,1702]" country="Germany" httpUri="http://grbio.org/cool/pvva-bewi" name="Martin Luther Universitaet" pageId="20" pageNumber="118">MLUH</collectionCode>
. In order to clarify the taxonomic status of the name by fixing it to a specimen, according to Article 75.3 of the International Code of Zoological Nomenclature (1999), the specimen found at the
<collectionCode id="ED69AD91FFD8FF9D373AF90C5CFDF894" box="[1336,1414,1773,1796]" country="Germany" httpUri="http://grbio.org/cool/pvva-bewi" name="Martin Luther Universitaet" pageId="20" pageNumber="118">MLUH</collectionCode>
was selected as the
<typeStatus id="54C38BF6FFD8FF9D31C1F8EC5D63F8B5" box="[963,1048,1805,1829]" pageId="20" pageNumber="118" type="neotype">neotype</typeStatus>
of
<taxonomicName id="4C784ED7FFD8FF9D363AF8EC5DF7F8B4" box="[1080,1164,1805,1828]" class="Insecta" family="Scarabaeidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="118" phylum="Arthropoda" rank="species" species="morio">
<emphasis id="B90CE946FFD8FF9D363AF8EC5DF7F8B4" box="[1080,1164,1805,1828]" italics="true" pageId="20" pageNumber="118">L. morio</emphasis>
</taxonomicName>
(
<figureCitation id="134329D1FFD8FF9D369EF8ED5C65F8B4" box="[1180,1310,1804,1828]" captionStart="Figure 7" captionStartId="19.[129,194,1836,1860]" captionTargetBox="[181,1423,149,1808]" captionTargetId="figure-7@19.[181,1423,144,1808]" captionTargetPageId="19" captionText="Figure 7. Liogenys niger (Blanchard) comb. nov.. Male neotype habitus (A), labels (E), and complete series at MLUH (D); female lectotype (B) and labels (F); non-uniform body colour variation (C); male in lateral (G) and posterior (I) views; detail of ventrites II and II with medial tubercle (H); parameres in dorsal (J) and lateral (K) views." figureDoi="http://doi.org/10.5281/zenodo.11241108" httpUri="https://zenodo.org/record/11241108/files/figure.png" pageId="20" pageNumber="118">Fig. 7A, D, E</figureCitation>
).
</paragraph>
<paragraph id="8BC73554FFD8FF9C3147F8CA5A71FF57" blockId="20.[810,1460,144,1985]" lastBlockId="21.[129,779,144,951]" lastPageId="21" lastPageNumber="119" pageId="20" pageNumber="118">
After the study of
<taxonomicName id="4C784ED7FFD8FF9D3601F8CA5C57F8D4" authority="(MNHN)" baseAuthorityName="MNHN" box="[1027,1324,1835,1860]" class="Insecta" family="Scarabaeidae" genus="Homalochilus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="118" phylum="Arthropoda" rank="species" species="niger">
<emphasis id="B90CE946FFD8FF9D3601F8CA5DC7F8D3" box="[1027,1212,1835,1859]" italics="true" pageId="20" pageNumber="118">Homalochilus niger</emphasis>
(MNHN)
</taxonomicName>
and
<taxonomicName id="4C784ED7FFD8FF9D3763F8CD5AB0F8F3" authority="(MLUH)" baseAuthorityName="MLUH" class="Insecta" family="Scarabaeidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="118" phylum="Arthropoda" rank="species" species="morio">
<emphasis id="B90CE946FFD8FF9D3763F8CD5A19F8F2" italics="true" pageId="20" pageNumber="118">Liogenys morio</emphasis>
(MLUH)
</taxonomicName>
<typeStatus id="54C38BF6FFD8FF9D31D0F8AA5A84F8F3" box="[978,1023,1867,1891]" pageId="20" pageNumber="118">type</typeStatus>
series, we concluded that they are conspecific, with
<taxonomicName id="4C784ED7FFD8FF9D3191F88A5D58F812" box="[915,1059,1899,1922]" class="Insecta" family="Scarabaeidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="118" phylum="Arthropoda" rank="species" species="morio">
<emphasis id="B90CE946FFD8FF9D3191F88A5D58F812" box="[915,1059,1899,1922]" italics="true" pageId="20" pageNumber="118">Liogenys morio</emphasis>
</taxonomicName>
being the junior subjective synonym of
<taxonomicName id="4C784ED7FFD8FF9D3128F8685A9AF831" box="[810,993,1929,1953]" class="Insecta" family="Scarabaeidae" genus="Homalochilus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="118" phylum="Arthropoda" rank="species" species="niger">
<emphasis id="B90CE946FFD8FF9D3128F8685A9AF831" box="[810,993,1929,1953]" italics="true" pageId="20" pageNumber="118">Homalochilus niger</emphasis>
</taxonomicName>
(
<taxonomicName id="4C784ED7FFD8FF9D31F3F86B5D42F831" authorityName="Cherman &amp; Basílio &amp; Clarkson &amp; Agostinis &amp; Smith &amp; Vaz-De-Mello &amp; Almeida" authorityYear="2024" baseAuthorityName="Cherman &amp; Basílio &amp; Clarkson &amp; Agostinis &amp; Smith &amp; Vaz-De-Mello &amp; Almeida" baseAuthorityYear="1851" box="[1009,1081,1930,1953]" class="Insecta" family="Scarabaeidae" genus="Homalochilus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="118" phylum="Arthropoda" rank="species" species="niger" status="comb. nov.">
<emphasis id="B90CE946FFD8FF9D31F3F86B5D42F831" box="[1009,1081,1930,1953]" italics="true" pageId="20" pageNumber="118">L. niger</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23F543DFFD8FF9D363CF8685DCBF832" box="[1086,1200,1929,1954]" pageId="20" pageNumber="118" rank="species">comb. nov.</taxonomicNameLabel>
). Although the specimens from ‘Brazil’ and Misiones (
<collectingCountry id="F36F75C4FFD8FF9D3635F8485DDAF851" box="[1079,1185,1961,1985]" name="Argentina" pageId="20" pageNumber="118">Argentina</collectingCountry>
) do not have the ventrites
<collectionCode id="ED69AD91FFD8FF9D37A3F8485CC8F850" box="[1441,1459,1961,1984]" pageId="20" pageNumber="118">II</collectionCode>
and
<collectionCode id="ED69AD91FFD9FF9C32A8FF7159BFFF37" box="[170,196,144,167]" pageId="21" pageNumber="119">III</collectionCode>
with tubercles at the midline and the parameres show some variations in shape, we considered those as intraspecific variations.
</paragraph>
<paragraph id="8BC73554FFD9FF9C329EFF2E5B45FD0D" blockId="21.[129,779,144,951]" pageId="21" pageNumber="119">
In the phylogenetic hypothesis of
<bibRefCitation id="EFE948A5FFD9FF9C33F9FF2E5B99FF77" author="Cherman MA &amp; Moron MA &amp; Almeida LM" box="[507,738,207,231]" pageId="21" pageNumber="119" pagination="744 - 70" refId="ref27020" refString="Cherman MA, Moron MA, Almeida LM. Phylogenetic relationships within Diplotaxini Kirby (Coleoptera: Melolonthidae: Melolonthinae) with emphasis on Liogenys Guerin-Meneville. Systematic Entomology 2016; 41: 744 - 70." type="journal article" year="2016">
Cherman
<emphasis id="B90CE946FFD9FF9C3061FF2E5BE8FF77" box="[611,659,207,231]" italics="true" pageId="21" pageNumber="119">et al.</emphasis>
(2016)
</bibRefCitation>
,
<taxonomicName id="4C784ED7FFD9FF9C30F3FF2E59D4FE96" authorityName="Cherman &amp; Basílio &amp; Clarkson &amp; Agostinis &amp; Smith &amp; Vaz-De-Mello &amp; Almeida" authorityYear="2024" baseAuthorityName="Cherman &amp; Basílio &amp; Clarkson &amp; Agostinis &amp; Smith &amp; Vaz-De-Mello &amp; Almeida" baseAuthorityYear="1851" class="Insecta" family="Scarabaeidae" genus="Homalochilus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="119" phylum="Arthropoda" rank="species" species="niger">
<emphasis id="B90CE946FFD9FF9C30F3FF2E59D4FE96" italics="true" pageId="21" pageNumber="119">H. niger</emphasis>
</taxonomicName>
(hereinafter
<taxonomicName id="4C784ED7FFD9FF9C3329FF0E580BFE96" authorityName="Cherman &amp; Basílio &amp; Clarkson &amp; Agostinis &amp; Smith &amp; Vaz-De-Mello &amp; Almeida" authorityYear="2024" baseAuthorityName="Cherman &amp; Basílio &amp; Clarkson &amp; Agostinis &amp; Smith &amp; Vaz-De-Mello &amp; Almeida" baseAuthorityYear="1851" box="[299,368,239,262]" class="Insecta" family="Scarabaeidae" genus="Homalochilus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="119" phylum="Arthropoda" rank="species" species="niger">
<emphasis id="B90CE946FFD9FF9C3329FF0E580BFE96" box="[299,368,239,262]" italics="true" pageId="21" pageNumber="119">L. niger</emphasis>
</taxonomicName>
) and
<taxonomicName id="4C784ED7FFD9FF9C33A4FF0E5B71FE96" authorityName="Blanchard" authorityYear="1850" box="[422,522,238,262]" class="Insecta" family="Scarabaeidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="119" phylum="Arthropoda" rank="species" species="concolor">
<emphasis id="B90CE946FFD9FF9C33A4FF0E5B71FE96" box="[422,522,238,262]" italics="true" pageId="21" pageNumber="119">L.concolor</emphasis>
</taxonomicName>
are not closely related species, due to miscoded characters already discussed in this text. The present hypotheses from equal weighting and implied weighting analysis show support (
<collectionCode id="ED69AD91FFD9FF9C3366FEAD58EFFEF4" box="[356,404,332,356]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32937" lsid="urn:lsid:biocol.org:col:32937" name="Archbold Biological Station" pageId="21" pageNumber="119" type="Other type of research institution/biorepository">ABS</collectionCode>
= 4/SR = 56;
<collectionCode id="ED69AD91FFD9FF9C3027FEAD5B29FEF4" box="[549,594,332,356]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13945" lsid="urn:lsid:biocol.org:col:13945" name="Royal Botanic Society" pageId="21" pageNumber="119" type="Herbarium">RBS</collectionCode>
= 35/SR = 78, respectively) in
<taxonomicName id="4C784ED7FFD9FF9C3309FE8D5829FE13" authorityName="Cherman &amp; Basílio &amp; Clarkson &amp; Agostinis &amp; Smith &amp; Vaz-De-Mello &amp; Almeida" authorityYear="2024" baseAuthorityName="Cherman &amp; Basílio &amp; Clarkson &amp; Agostinis &amp; Smith &amp; Vaz-De-Mello &amp; Almeida" baseAuthorityYear="1851" box="[267,338,364,387]" class="Insecta" family="Scarabaeidae" genus="Homalochilus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="119" phylum="Arthropoda" rank="species" species="niger">
<emphasis id="B90CE946FFD9FF9C3309FE8D5829FE13" box="[267,338,364,387]" italics="true" pageId="21" pageNumber="119">L. niger</emphasis>
</taxonomicName>
being nested within
<taxonomicName id="4C784ED7FFD9FF9C3020FE8D5B08FE13" box="[546,627,364,387]" class="Insecta" family="Scarabaeidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="119" phylum="Arthropoda" rank="genus">
<emphasis id="B90CE946FFD9FF9C3020FE8D5B08FE13" box="[546,627,364,387]" italics="true" pageId="21" pageNumber="119">Liogenys</emphasis>
</taxonomicName>
. This inclusion does not significantly change the diagnosis of the genus (
<bibRefCitation id="EFE948A5FFD9FF9C30A9FE6A599CFE52" author="Cherman MA &amp; Mise KM &amp; Moron MA" pageId="21" pageNumber="119" pagination="1 - 120" refId="ref27057" refString="Cherman MA, Mise KM, Moron MA et al. A taxonomic revision of Liogenys occurring in Brazil with an interactive key and remarks on New World Diplotaxini (Coleoptera, Melolonthidae). ZooKeys 2017; 699: 1 - 120." type="journal article" year="2017">
Cherman
<emphasis id="B90CE946FFD9FF9C3283FE4A59D2FE52" box="[129,169,426,450]" italics="true" pageId="21" pageNumber="119">et al</emphasis>
. 2017
</bibRefCitation>
), though
<taxonomicName id="4C784ED7FFD9FF9C3349FE4A58EFFE52" authorityName="Cherman &amp; Basílio &amp; Clarkson &amp; Agostinis &amp; Smith &amp; Vaz-De-Mello &amp; Almeida" authorityYear="2024" baseAuthorityName="Cherman &amp; Basílio &amp; Clarkson &amp; Agostinis &amp; Smith &amp; Vaz-De-Mello &amp; Almeida" baseAuthorityYear="1851" box="[331,404,427,450]" class="Insecta" family="Scarabaeidae" genus="Homalochilus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="119" phylum="Arthropoda" rank="species" species="niger">
<emphasis id="B90CE946FFD9FF9C3349FE4A58EFFE52" box="[331,404,427,450]" italics="true" pageId="21" pageNumber="119">L. niger</emphasis>
</taxonomicName>
includes some exceptions among the genus
<taxonomicName id="4C784ED7FFD9FF9C32BCFE2B5874FE71" box="[190,271,458,481]" class="Insecta" family="Scarabaeidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="119" phylum="Arthropoda" rank="genus">
<emphasis id="B90CE946FFD9FF9C32BCFE2B5874FE71" box="[190,271,458,481]" italics="true" pageId="21" pageNumber="119">Liogenys</emphasis>
</taxonomicName>
: clypeus in lateral view straight, not sinuous (12:1); labrum bulging on superior margin (38:0); and setae on mesotibial carina longer than the apical (84:0). All characters supporting
<taxonomicName id="4C784ED7FFD9FF9C3283FDC959A9FDAF" box="[129,210,552,575]" class="Insecta" family="Scarabaeidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="119" phylum="Arthropoda" rank="genus">
<emphasis id="B90CE946FFD9FF9C3283FDC959A9FDAF" box="[129,210,552,575]" italics="true" pageId="21" pageNumber="119">Liogenys</emphasis>
</taxonomicName>
in
<bibRefCitation id="EFE948A5FFD9FF9C32F0FDC658A8FDAF" author="Cherman MA &amp; Moron MA &amp; Almeida LM" box="[242,467,551,575]" pageId="21" pageNumber="119" pagination="744 - 70" refId="ref27020" refString="Cherman MA, Moron MA, Almeida LM. Phylogenetic relationships within Diplotaxini Kirby (Coleoptera: Melolonthidae: Melolonthinae) with emphasis on Liogenys Guerin-Meneville. Systematic Entomology 2016; 41: 744 - 70." type="journal article" year="2016">
Cherman
<emphasis id="B90CE946FFD9FF9C3354FDC958FEFDAF" box="[342,389,551,575]" italics="true" pageId="21" pageNumber="119">et al.</emphasis>
(2016)
</bibRefCitation>
are also in the present hypotheses, including its synapomorphy (
<tableCitation id="C6FA00EFFFD9FF9C33D5FDA75B5BFDCF" box="[471,544,582,607]" captionStart="Table 1" captionStartId="6.[113,168,143,167]" captionTargetPageId="6" captionText="Table 1. List of common synapomorphies for the main clades of Neotropical Diplotaxini analysis, comparing equal and implied weights to the characters." pageId="21" pageNumber="119">Table 1</tableCitation>
). Character 156:1 (
<figureCitation id="134329D1FFD9FF9C30E7FDA759D0FDEE" captionStart="Figure 12" captionStartId="26.[113,178,1401,1425]" captionTargetBox="[116,1458,147,1372]" captionTargetId="figure-360@26.[114,1458,144,1374]" captionTargetPageId="26" captionText="Figure 12. Hindwing in Careocallus, detail of radial sector.Careocallus tehuelche male (A) and female (B)." figureDoi="http://doi.org/10.5281/zenodo.11241118" httpUri="https://zenodo.org/record/11241118/files/figure.png" pageId="21" pageNumber="119">Fig. 12C</figureCitation>
), coded for the first time, also supports the clade, and thus, is a new contribution to the diagnosis of
<taxonomicName id="4C784ED7FFD9FF9C33EAFD675B42FD0D" box="[488,569,646,669]" class="Insecta" family="Scarabaeidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="119" phylum="Arthropoda" rank="genus">
<emphasis id="B90CE946FFD9FF9C33EAFD675B42FD0D" box="[488,569,646,669]" italics="true" pageId="21" pageNumber="119">Liogenys</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BC73554FFD9FF9C329EFD455BB8FC27" blockId="21.[129,779,144,951]" pageId="21" pageNumber="119">
The implied weighting analysis (
<figureCitation id="134329D1FFD9FF9C33F3FD455B57FD2D" box="[497,556,676,701]" captionStart="Figure 2" captionStartId="5.[129,194,1455,1479]" captionTargetBox="[131,1470,147,1426]" captionTargetId="figure-304@5.[129,1473,144,1427]" captionTargetPageId="5" captionText="Figure 2. Implied weighting analysis, one tree obtained with concavity constant value k = 8.750. The numbers displayed in each node correspond to the value of relative Bremer and symmetric resampling supports (above and below, respectively)." figureDoi="http://doi.org/10.5281/zenodo.11241098" httpUri="https://zenodo.org/record/11241098/files/figure.png" pageId="21" pageNumber="119">Fig. 2</figureCitation>
) solved the relationships within
<taxonomicName id="4C784ED7FFD9FF9C330AFD245827FD4C" box="[264,348,709,732]" class="Insecta" family="Scarabaeidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="119" phylum="Arthropoda" rank="genus">
<emphasis id="B90CE946FFD9FF9C330AFD245827FD4C" box="[264,348,709,732]" italics="true" pageId="21" pageNumber="119">Liogenys</emphasis>
</taxonomicName>
better than the equal weighting analysis, where the clade containing
<taxonomicName id="4C784ED7FFD9FF9C339FFD05589CFD6B" authorityName="Cherman &amp; Basílio &amp; Clarkson &amp; Agostinis &amp; Smith &amp; Vaz-De-Mello &amp; Almeida" authorityYear="2024" baseAuthorityName="Cherman &amp; Basílio &amp; Clarkson &amp; Agostinis &amp; Smith &amp; Vaz-De-Mello &amp; Almeida" baseAuthorityYear="1851" box="[413,487,740,763]" class="Insecta" family="Scarabaeidae" genus="Homalochilus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="119" phylum="Arthropoda" rank="species" species="niger">
<emphasis id="B90CE946FFD9FF9C339FFD05589CFD6B" box="[413,487,740,763]" italics="true" pageId="21" pageNumber="119">L. niger</emphasis>
</taxonomicName>
collapsed with 10 other lineages (
<collectionCode id="ED69AD91FFD9FF9C32C1FCE2598FFC8B" box="[195,244,771,795]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32937" lsid="urn:lsid:biocol.org:col:32937" name="Archbold Biological Station" pageId="21" pageNumber="119" type="Other type of research institution/biorepository">ABS</collectionCode>
= 1/21). In the former analysis,
<taxonomicName id="4C784ED7FFD9FF9C3040FCE25BF7FC8A" authorityName="Cherman &amp; Basílio &amp; Clarkson &amp; Agostinis &amp; Smith &amp; Vaz-De-Mello &amp; Almeida" authorityYear="2024" baseAuthorityName="Cherman &amp; Basílio &amp; Clarkson &amp; Agostinis &amp; Smith &amp; Vaz-De-Mello &amp; Almeida" baseAuthorityYear="1851" box="[578,652,771,794]" class="Insecta" family="Scarabaeidae" genus="Homalochilus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="119" phylum="Arthropoda" rank="species" species="niger">
<emphasis id="B90CE946FFD9FF9C3040FCE25BF7FC8A" box="[578,652,771,794]" italics="true" pageId="21" pageNumber="119">L. niger</emphasis>
</taxonomicName>
is the sister-clade of
<taxonomicName id="4C784ED7FFD9FF9C32E2FCC2582BFCAA" authorityName="Blanchard" authorityYear="1850" box="[224,336,802,826]" class="Insecta" family="Scarabaeidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="119" phylum="Arthropoda" rank="species" species="concolor">
<emphasis id="B90CE946FFD9FF9C32E2FCC2582BFCAA" box="[224,336,802,826]" italics="true" pageId="21" pageNumber="119">L. concolor</emphasis>
</taxonomicName>
(
<collectionCode id="ED69AD91FFD9FF9C3360FCC358E8FCAA" box="[354,403,802,826]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13945" lsid="urn:lsid:biocol.org:col:13945" name="Royal Botanic Society" pageId="21" pageNumber="119" type="Herbarium">RBS</collectionCode>
= 28/SR = 26), supported by four transformations (
<tableCitation id="C6FA00EFFFD9FF9C3336FCA058FBFCC9" box="[308,384,833,857]" captionStart="Table 1" captionStartId="6.[113,168,143,167]" captionTargetPageId="6" captionText="Table 1. List of common synapomorphies for the main clades of Neotropical Diplotaxini analysis, comparing equal and implied weights to the characters." pageId="21" pageNumber="119">Table 1</tableCitation>
). The body convex, slightly depressed posteriorly (59:0); the pronotum with posterior angles slightly projected (48:0); and the apex of the scutellum rounded (56:0), are similarities already mentioned by
<bibRefCitation id="EFE948A5FFD9FF9C33FEFC7E5BC6FC27" author="Burmeister H" box="[508,701,927,951]" pageId="21" pageNumber="119" refId="ref26916" refString="Burmeister H. Handbuch der entomologie (Coleoptera Lamellicornia Phyllophaga Chaenochela). Vol. 4, Part 2. Berlin: Enslin, T. C. F., 1855." type="book" year="1855">Burmeister (1855)</bibRefCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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