248 lines
29 KiB
XML
248 lines
29 KiB
XML
<document id="B77F5E671CCB1280B4EC2086872BAFE9" ID-DOI="10.5281/zenodo.7911385" ID-ISSN="2305-2562" ID-Zenodo-Dep="7911385" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1683577494970" checkinUser="felipe" docAuthor="Plisko, Jadwiga Danuta" docDate="2004" docId="03D087EAFFB57C07FC9DFF78111FFD9E" docLanguage="en" docName="AfrInvertebr.45.1.287–313.pdf" docOrigin="African Invertebrates 45 (1998)" docStyle="DocumentStyle:C20B10C002986E51083406A4EACF003A.3:AfrInvertebr.2001-2007.journal_article" docStyleId="C20B10C002986E51083406A4EACF003A" docStyleName="AfrInvertebr.2001-2007.journal_article" docStyleVersion="3" docTitle="Udeina petrosi Plisko 2004, sp. n." docType="treatment" docVersion="5" lastPageNumber="307" masterDocId="FFE9FF92FFA77C13FD6AFFB6121FFF91" masterDocTitle="Review of the balantine genus Udeina Michaelsen, 1910 with descriptions of six new species in South Africa (Oligochaeta: Acanthodrilidae, Acanthodrilinae)" masterLastPageNumber="287" masterPageNumber="287" pageNumber="305" updateTime="1699498675842" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:mods id="3DF11A894C6FE993D45B6F68362EBCA4" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="143174FCD6E0DC4EED52F67FF6200114">
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<mods:title id="8BDB578FC97C5173FE377D9455356C02">Review of the balantine genus Udeina Michaelsen, 1910 with descriptions of six new species in South Africa (Oligochaeta: Acanthodrilidae, Acanthodrilinae)</mods:title>
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</mods:titleInfo>
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<mods:name id="F30F29D0E8608DB4E467541CB25D0077" type="personal">
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<mods:role id="8600D59E4C86EA4C4E3F59C8E68FAA20">
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<mods:roleTerm id="D02CBB53B6F9D1A55D462032CECC1158">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="ECC8DBCC695FE0FD9E2A378DFA3D9256">Plisko, Jadwiga Danuta</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="C4202FF74C71C13D4E693DA6FC1CA25D">text</mods:typeOfResource>
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<mods:relatedItem id="99F1BEB9ED3CA45941C4339B656ABCE3" type="host">
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<mods:titleInfo id="FEF9834E622DD7FA1BEEF413368C6A08">
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<mods:title id="7E3D9F7679448940FD0660E7AD550EE3">African Invertebrates</mods:title>
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</mods:titleInfo>
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<mods:part id="2CD9A608A904A8E831C7047BB02EFF69">
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<mods:date id="993B03CF1DE1DA21F2967040BB38B900">2004</mods:date>
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<mods:detail id="7533ADBDE65479C7BEC4971520BA4405" type="pubDate">
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<mods:number id="BC3644DDAFD616F86BA9BF86A23779CE">2004-12-31</mods:number>
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</mods:detail>
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||
<mods:detail id="05AED18EBF5B291991886F26E3DF53F1" type="volume">
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<mods:number id="294C10F3D01D6ABF9A43DD6F005A78E1">45</mods:number>
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</mods:detail>
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<mods:detail id="59A8109C4BCF042F586011F88F15F11F" type="issue">
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<mods:number id="5FA37DDBDF809B4DB6E5CEE5C37F4F97">1998</mods:number>
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</mods:detail>
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||
<mods:extent id="AF629F4CC8C6722BC501CDDDE7FA9EE2" unit="page">
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<mods:start id="A5C8686D5702F8B01A98A2F45B57148F">287</mods:start>
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<mods:end id="5BD0786A04D498E7551FCE49642DE2A1">287</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:classification id="232D1E703E0210F8365D2250A1657EE1">journal article</mods:classification>
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<mods:identifier id="EE07AF175E7860E4C0FAAA019BB45EC6" type="DOI">10.5281/zenodo.7911385</mods:identifier>
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<mods:identifier id="99CC8FD8B7232D778905A76B000375FB" type="ISSN">2305-2562</mods:identifier>
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<mods:identifier id="53575E8A6DC57D1D50BF185B56DCBCD7" type="Zenodo-Dep">7911385</mods:identifier>
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</mods:mods>
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<treatment id="03D087EAFFB57C07FC9DFF78111FFD9E" ID-DOI="http://doi.org/10.5281/zenodo.7910352" ID-Zenodo-Dep="7910352" LSID="urn:lsid:plazi:treatment:03D087EAFFB57C07FC9DFF78111FFD9E" httpUri="http://treatment.plazi.org/id/03D087EAFFB57C07FC9DFF78111FFD9E" lastPageId="20" lastPageNumber="307" pageId="18" pageNumber="305">
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<subSubSection id="C3636577FFB57C01FC9DFF7810F7FF76" box="[503,744,206,231]" pageId="18" pageNumber="305" type="nomenclature">
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<paragraph id="8BC636FCFFB57C01FC9DFF7810F7FF76" blockId="18.[144,1104,206,823]" box="[503,744,206,231]" pageId="18" pageNumber="305">
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<heading id="D08E8190FFB57C01FC9DFF7810F7FF76" box="[503,744,206,231]" centered="true" fontSize="11" level="2" pageId="18" pageNumber="305" reason="2">
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<taxonomicName id="4C794D7FFFB57C01FC9DFF7810B9FF76" ID-CoL="25fc9efd-5179-4a95-bbb2-4b6209d68402" authority="Plisko, 2004" authorityName="Plisko" authorityYear="2004" box="[503,678,206,231]" class="Clitellata" family="Acanthodrilidae" genus="Udeina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crassiclitellata" pageId="18" pageNumber="305" phylum="Annelida" rank="species" species="petrosi" status="sp. nov.">
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<emphasis id="B90DEAEEFFB57C01FC9DFF7810B9FF76" bold="true" box="[503,678,206,231]" pageId="18" pageNumber="305">Udeina petrosi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A23E5795FFB57C01FFC6FF7810F7FF76" box="[684,744,206,231]" pageId="18" pageNumber="305" rank="species">sp. n.</taxonomicNameLabel>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3636577FFB57C07FF5BFF40111FFD9E" lastPageId="20" lastPageNumber="307" pageId="18" pageNumber="305" type="description">
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<paragraph id="8BC636FCFFB57C01FF5BFF4010B1FE9E" blockId="18.[144,1104,206,823]" box="[561,686,246,271]" pageId="18" pageNumber="305">
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<figureCitation id="13422A79FFB57C01FF5BFF4010B1FE9E" box="[561,686,246,271]" captionStart="Figs 13–15" captionStartId="18.[144,183,1551,1571]" captionText="Figs 13–15. Udeina petrosi sp. n. Paratype. 13. External ventral area of segments 15–21, 90 X. Pp = prostatic pore; Sg = seminal groove. 14. Spermatheca, 380 X. A = ampulla; Dv = diverticulum; Spd = spermathecal duct. 15. Prostate of segment 19 and vestigial prostatic duct of segment 17. Pg = prostatic gland of segment 19; Pd19 = prostatic duct of segment 19; Pd17 = vestigial prostatic duct of segment 17." pageId="18" pageNumber="305">Figs 13–15</figureCitation>
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</paragraph>
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<paragraph id="8BC636FCFFB57C01FDFAFEA813F9FEC6" blockId="18.[144,1104,206,823]" pageId="18" pageNumber="305">
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Etymology: Named after Mr Petros Ngwenya of Ezemvelo KZN Wildlife, who assisted in collecting the
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<typeStatus id="54C2885EFFB57C01FC21FE881364FEC6" box="[331,379,318,343]" pageId="18" pageNumber="305">type</typeStatus>
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material.
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</paragraph>
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<paragraph id="8BC636FCFFB57C01FDFAFED0113AFE4E" blockId="18.[144,1104,206,823]" pageId="18" pageNumber="305">
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Type material:
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<materialsCitation id="3B113CA1FFB57C01FC58FED0107EFE2E" collectingDate="2001-03-14" collectionCode="NMSA, JDP" collectorName="A. J. Armstrong & P. Ngwenya" latitude="-29.309166" location="Highmoor State Forest" longLatPrecision="20" longitude="29.598333" pageId="18" pageNumber="305" specimenCount="1" stateProvince="KwaZulu-Natal" typeStatus="holotype">
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<typeStatus id="54C2885EFFB57C01FC58FED01382FEEE" box="[306,413,358,383]" pageId="18" pageNumber="305" type="holotype">Holotype</typeStatus>
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: clitellate
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<collectionCode id="ED68AE39FFB57C01FF7AFED0107CFEEE" box="[528,611,358,383]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="18" pageNumber="305" type="Museum">NMSA</collectionCode>
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/Olig.03371
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<emphasis id="B90DEAEEFFB57C01FF80FED01189FEEF" box="[746,918,358,382]" italics="true" pageId="18" pageNumber="305">
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<collectingRegion id="49BDF81EFFB57C01FF80FED01189FEEF" box="[746,918,358,382]" country="South Africa" name="KwaZulu-Natal" pageId="18" pageNumber="305">KwaZuluNatal</collectingRegion>
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</emphasis>
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:
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<location id="8EA66027FFB57C01FEC8FED012C9FE0E" LSID="urn:lsid:plazi:treatment:03D087EAFFB57C07FC9DFF78111FFD9E:8EA66027FFB57C01FEC8FED012C9FE0E" latitude="-29.309166" longLatPrecision="20" longitude="29.598333" name="Highmoor State Forest" pageId="18" pageNumber="305" stateProvince="KwaZulu-Natal">Highmoor State Forest</location>
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(
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<geoCoordinate id="EE4D503BFFB57C01FD89FE30137FFE0E" box="[227,352,390,415]" degrees="29" direction="south" minutes="18" orientation="latitude" pageId="18" pageNumber="305" precision="15" seconds="33" value="-29.309166">29º18'33"S</geoCoordinate>
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:
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<geoCoordinate id="EE4D503BFFB57C01FC0EFE3013F5FE0E" box="[356,490,390,415]" degrees="29" direction="east" minutes="35" orientation="longitude" pageId="18" pageNumber="305" precision="15" seconds="54" value="29.598333">29º35'54"E</geoCoordinate>
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), grassland at the edge of marsh, in moist soil,
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<date id="FFC7103CFFB57C01FE8FFE3012D4FE2E" pageId="18" pageNumber="305" value="2001-03-14">
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<collectingDate id="EF83E9D4FFB57C01FE8FFE3012D4FE2E" pageId="18" pageNumber="305" value="2001-03-14">14 March 2001</collectingDate>
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</date>
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,
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<collectionCode id="ED68AE39FFB57C01FDBDFE10131AFE2E" box="[215,261,422,447]" pageId="18" pageNumber="305">JDP</collectionCode>
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,
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<collectorName id="268C532AFFB57C01FC7BFE1013D6FE2E" box="[273,457,422,447]" pageId="18" pageNumber="305">A. J. Armstrong</collectorName>
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,
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<collectorName id="268C532AFFB57C01FCBCFE10107EFE2E" box="[470,609,422,447]" pageId="18" pageNumber="305">P. Ngwenya</collectorName>
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</materialsCitation>
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.
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<materialsCitation id="3B113CA1FFB57C01FF07FE101653FE2E" box="[621,1100,422,447]" collectionCode="NMSA" pageId="18" pageNumber="305" specimenCount="1" stateProvince="KwaZulu-Natal" typeStatus="paratype">
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<typeStatus id="54C2885EFFB57C01FF07FE1010C1FE2E" box="[621,734,422,447]" pageId="18" pageNumber="305" type="paratype">Paratypes</typeStatus>
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:
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<collectionCode id="ED68AE39FFB57C01FF80FE101121FE2E" box="[746,830,422,447]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="18" pageNumber="305" type="Museum">NMSA</collectionCode>
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/Olig.03616, 7 clitellate
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</materialsCitation>
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;
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<materialsCitation id="3B113CA1FFB57C01FDFAFE70113EFE4E" box="[144,801,454,479]" collectingDate="2001-03-14" collectionCode="NMSA" collectorName="A. J. Armstrong & P. Ngwenya" latitude="-29.309166" location="Highmoor State Forest" longLatPrecision="20" longitude="29.598333" pageId="18" pageNumber="305" specimenCount="7" specimenCount-juvenile="7" stateProvince="KwaZulu-Natal" typeStatus="paratype">
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<collectionCode id="ED68AE39FFB57C01FDFAFE7012FBFE4E" box="[144,228,454,479]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="18" pageNumber="305" type="Museum">NMSA</collectionCode>
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/Olig.03617,
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<specimenCount id="9D7FFD75FFB57C01FC1FFE7013F7FE4E" box="[373,488,454,479]" pageId="18" pageNumber="305" type="juvenile">7 juvenile</specimenCount>
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; all collected with holotype
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</materialsCitation>
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.
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</paragraph>
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<paragraph id="8BC636FCFFB57C01FDFAFE581304FD96" blockId="18.[144,1104,206,823]" box="[144,283,494,519]" pageId="18" pageNumber="305">Description:</paragraph>
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<paragraph id="8BC636FCFFB57C01FDFAFDA01348FDBE" blockId="18.[144,1104,206,823]" box="[144,343,534,559]" pageId="18" pageNumber="305">External features:</paragraph>
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<paragraph id="8BC636FCFFB57C00FDFAFD88113BFDB6" blockId="18.[144,1104,206,823]" lastBlockId="19.[181,1143,206,1682]" lastPageId="19" lastPageNumber="306" pageId="18" pageNumber="305">
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Body cylindrical, in life dorsally violet, ventrally yellowishgrey; alcoholpreserved dorsally violet, with stronger tint on preclitellar segments; ventrally yellowishgray. Clitellate
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<quantity id="4C819B19FFB57C01FC68FDC81392FD06" box="[258,397,638,663]" metricMagnitude="-1" metricUnit="m" metricValue="1.09" metricValueMax="1.28" metricValueMin="0.9" pageId="18" pageNumber="305" unit="mm" value="109.0" valueMax="128.0" valueMin="90.0">90–128 mm</quantity>
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long,
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<quantity id="4C819B19FFB57C01FCB1FDC81000FD06" box="[475,543,638,663]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="18" pageNumber="305" unit="mm" value="4.0">4 mm</quantity>
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wide.
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<emphasis id="B90DEAEEFFB57C01FF18FDC8112DFD07" box="[626,818,638,662]" italics="true" pageId="18" pageNumber="305">Segment number</emphasis>
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:
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<typeStatus id="54C2885EFFB57C01FE2EFDC811B8FD06" box="[836,935,638,663]" pageId="18" pageNumber="305" type="holotype">holotype</typeStatus>
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160; clitellate
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<typeStatus id="54C2885EFFB57C01FDFAFD2812E3FD26" box="[144,252,670,695]" pageId="18" pageNumber="305" type="paratype">paratypes</typeStatus>
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158–166, aclitellate 117–161.
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<emphasis id="B90DEAEEFFB57C01FF38FD2810C4FD27" box="[594,731,670,694]" italics="true" pageId="18" pageNumber="305">Prostomium</emphasis>
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: tanylobous with evident sutures.
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<emphasis id="B90DEAEEFFB57C01FDFAFD0812D2FD47" box="[144,205,702,726]" italics="true" pageId="18" pageNumber="305">Setae</emphasis>
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: widely paired; on postclitellar segments
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<emphasis id="B90DEAEEFFB57C01FFD8FD0810D1FD47" box="[690,718,702,726]" italics="true" pageId="18" pageNumber="305">aa</emphasis>
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<
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<emphasis id="B90DEAEEFFB57C01FF9AFD081114FD47" box="[752,779,702,726]" italics="true" pageId="18" pageNumber="305">bc</emphasis>
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;
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<emphasis id="B90DEAEEFFB57C01FE76FD081127FD47" box="[796,824,702,726]" italics="true" pageId="18" pageNumber="305">ab</emphasis>
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<
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<emphasis id="B90DEAEEFFB57C01FE30FD08116AFD47" box="[858,885,702,726]" italics="true" pageId="18" pageNumber="305">cd</emphasis>
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.
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<emphasis id="B90DEAEEFFB57C01FEEFFD081604FD47" box="[901,1051,702,726]" italics="true" pageId="18" pageNumber="305">Dorsal pores</emphasis>
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: not observed.
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<emphasis id="B90DEAEEFFB57C01FC62FD6813D3FD67" box="[264,460,734,758]" italics="true" pageId="18" pageNumber="305">Nephridial pores</emphasis>
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: on
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<typeStatus id="54C2885EFFB57C01FF6FFD681077FD66" box="[517,616,734,759]" pageId="18" pageNumber="305" type="holotype">holotype</typeStatus>
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not observed; on
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<specimenCount id="9D7FFD75FFB57C01FE29FD6811FAFD66" box="[835,997,734,759]" pageId="18" pageNumber="305" type="generic" typeStatus="paratypes">two paratypes</specimenCount>
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noted in
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<emphasis id="B90DEAEEFFB57C01FDFAFD481281FC87" box="[144,158,766,790]" italics="true" pageId="18" pageNumber="305">b</emphasis>
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setal lines in intersegmental furrow 21/22, and in selected posterior furrows.
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<emphasis id="B90DEAEEFFB57C01FDFAFCA81370FCA7" box="[144,367,798,822]" italics="true" pageId="18" pageNumber="305">Spermathecal pores</emphasis>
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: in intersegmental furrow 7/8, in front of
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<emphasis id="B90DEAEEFFB57C01FE2CFCA8114BFCA7" box="[838,852,798,822]" italics="true" pageId="18" pageNumber="305">b</emphasis>
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.
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<emphasis id="B90DEAEEFFB57C01FE08FCA811E5FCA7" box="[866,1018,798,822]" italics="true" pageId="18" pageNumber="305">Female pores</emphasis>
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: paired in 14, in front of
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<emphasis id="B90DEAEEFFB47C00FCEBFF781390FF77" box="[385,399,206,230]" italics="true" pageId="19" pageNumber="306">b</emphasis>
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setae.
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<taxonomicName id="4C794D7FFFB47C00FC8BFF781055FF77" box="[481,586,206,230]" class="Ascidiacea" family="Didemnidae" genus="Clitellum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Aplousobranchia" pageId="19" pageNumber="306" phylum="Chordata" rank="genus">
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<emphasis id="B90DEAEEFFB47C00FC8BFF781055FF77" box="[481,586,206,230]" italics="true" pageId="19" pageNumber="306">Clitellum</emphasis>
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</taxonomicName>
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: saddleshaped, on 13–1/n18; anterior border at intersegmental furrow 12/13, clitellar tissue on segment 13 slightly thinner than on 14–17; posterior border on 1/n18; lateral edges extend to rows of papillae in zigzag lines, bordering papillae located on 13–17.
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<emphasis id="B90DEAEEFFB47C00FFC0FE981147FED7" box="[682,856,302,326]" italics="true" pageId="19" pageNumber="306">Prostatic pores</emphasis>
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: one pair in small round papillae encircling
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<emphasis id="B90DEAEEFFB47C00FCEFFEF8138CFEF7" box="[389,403,334,358]" italics="true" pageId="19" pageNumber="306">b</emphasis>
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setae, in segment 19.
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<emphasis id="B90DEAEEFFB47C00FF15FEF8112BFEF7" box="[639,820,334,358]" italics="true" pageId="19" pageNumber="306">Seminal grooves</emphasis>
|
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(
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<figureCitation id="13422A79FFB47C00FE2BFEF81192FEF6" box="[833,909,334,359]" captionStart="Figs 13–15" captionStartId="18.[144,183,1551,1571]" captionText="Figs 13–15. Udeina petrosi sp. n. Paratype. 13. External ventral area of segments 15–21, 90 X. Pp = prostatic pore; Sg = seminal groove. 14. Spermatheca, 380 X. A = ampulla; Dv = diverticulum; Spd = spermathecal duct. 15. Prostate of segment 19 and vestigial prostatic duct of segment 17. Pg = prostatic gland of segment 19; Pd19 = prostatic duct of segment 19; Pd17 = vestigial prostatic duct of segment 17." pageId="19" pageNumber="306">Fig.13</figureCitation>
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): paired, longitudinal slightly curved, between posterior part of 17 and prostatic pore of 19; in 18 noted small depression being male pore.
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<emphasis id="B90DEAEEFFB47C00FF51FE3810A6FE37" box="[571,697,398,422]" italics="true" pageId="19" pageNumber="306">Male pores</emphasis>
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: paired, small openings in 18, lateral to
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<emphasis id="B90DEAEEFFB47C00FDDFFE1812DCFE57" box="[181,195,430,454]" italics="true" pageId="19" pageNumber="306">b</emphasis>
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setae; indicated by vas deferens entering body wall behind
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<emphasis id="B90DEAEEFFB47C00FE1BFE181192FE57" box="[881,909,430,454]" italics="true" pageId="19" pageNumber="306">ab</emphasis>
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setae, posteriorly to 17.
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<emphasis id="B90DEAEEFFB47C00FDB5FE781320FE77" box="[223,319,462,486]" italics="true" pageId="19" pageNumber="306">Papillae</emphasis>
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: paired, below
|
||
<emphasis id="B90DEAEEFFB47C00FC87FE7813E4FE77" box="[493,507,462,486]" italics="true" pageId="19" pageNumber="306">b</emphasis>
|
||
setae, parallel with ventral clitellar borders on segments 13–17; small swellings, oval, flat. It is not known if papillae of 17 support ectal prostatic ducts of the evidently reduced prostates of segment 17.
|
||
</paragraph>
|
||
<caption id="DF066674FFB57C01FDFAF9B913BCF912" pageId="18" pageNumber="305" startId="18.[144,183,1551,1571]" targetType="figure">
|
||
<paragraph id="8BC636FCFFB57C01FDFAF9B913BCF912" blockId="18.[144,1105,1551,1667]" pageId="18" pageNumber="305">
|
||
Figs 13–15.
|
||
<taxonomicName id="4C794D7FFFB57C01FD97F9B9139DF9B2" authority="Plisko, 2004" authorityName="Plisko" authorityYear="2004" box="[253,386,1551,1571]" class="Clitellata" family="Acanthodrilidae" genus="Udeina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crassiclitellata" pageId="18" pageNumber="305" phylum="Annelida" rank="species" species="petrosi" status="sp. nov.">
|
||
<emphasis id="B90DEAEEFFB57C01FD97F9B9139DF9B2" box="[253,386,1551,1571]" italics="true" pageId="18" pageNumber="305">Udeina petrosi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23E5795FFB57C01FCECF9B913AAF9B2" box="[390,437,1551,1571]" pageId="18" pageNumber="305" rank="species">sp. n.</taxonomicNameLabel>
|
||
Paratype. 13. External ventral area of segments 15–21, 90 X. Pp = prostatic pore; Sg = seminal groove. 14. Spermatheca, 380 X. A = ampulla; Dv = diverticulum; Spd = spermathecal duct. 15. Prostate of segment 19 and vestigial prostatic duct of segment 17. Pg = prostatic gland of segment 19; Pd19 = prostatic duct of segment 19; Pd17 = vestigial prostatic duct of segment 17.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BC636FCFFB47C00FDDFFD821392FDDC" blockId="19.[181,1143,206,1682]" box="[181,397,564,589]" pageId="19" pageNumber="306">Internal characters:</paragraph>
|
||
<paragraph id="8BC636FCFFB47C00FDDFFDEF164EF903" blockId="19.[181,1143,206,1682]" pageId="19" pageNumber="306">
|
||
<emphasis id="B90DEAEEFFB47C00FDDFFDEF1376FDE0" box="[181,361,601,625]" italics="true" pageId="19" pageNumber="306">Salivary glands</emphasis>
|
||
: not extending beyond septum 4/5.
|
||
<emphasis id="B90DEAEEFFB47C00FE7EFDEF1173FDE0" box="[788,876,601,625]" italics="true" pageId="19" pageNumber="306">Gizzard</emphasis>
|
||
: well developed, in 5, muscular, cylindrical with anterior soft fold.
|
||
<emphasis id="B90DEAEEFFB47C00FFC2FDCF10F9FD00" box="[680,742,633,657]" italics="true" pageId="19" pageNumber="306">Septa</emphasis>
|
||
: 5/6–6/7 thin, 7/8–12/13 increasing in thickness, muscular, 13/14–15/16 thinner; some variations in thickness in dissected individuals were observed.
|
||
<emphasis id="B90DEAEEFFB47C00FC8DFD0F1058FD40" box="[487,583,697,721]" italics="true" pageId="19" pageNumber="306">Intestine</emphasis>
|
||
: commences in 16.
|
||
<emphasis id="B90DEAEEFFB47C00FE4FFD0F11DCFD40" box="[805,963,697,721]" italics="true" pageId="19" pageNumber="306">Lateral hearts</emphasis>
|
||
: four pairs, thin vessels, last pair in 12.
|
||
<emphasis id="B90DEAEEFFB47C00FCC6FD6F1003FD60" box="[428,540,729,753]" italics="true" pageId="19" pageNumber="306">Nephridia</emphasis>
|
||
: large elongated coiled loops without terminal vesicles, posterior smaller.
|
||
<emphasis id="B90DEAEEFFB47C00FC10FD4F13CDFC80" box="[378,466,761,785]" italics="true" pageId="19" pageNumber="306">Ovaries</emphasis>
|
||
: large funnels attached to intersegmental furrow 13/14.
|
||
<emphasis id="B90DEAEEFFB47C00F959FD4F1669FC80" box="[1075,1142,761,785]" italics="true" pageId="19" pageNumber="306">Testes</emphasis>
|
||
and
|
||
<emphasis id="B90DEAEEFFB47C00FD82FCAF13CBFCA0" box="[232,468,793,817]" italics="true" pageId="19" pageNumber="306">spermiductal funnels</emphasis>
|
||
: spermiductal funnels large, free, medially in segment 10, iridescent.
|
||
<taxonomicName id="4C794D7FFFB47C00FC5BFC8F13C2FCC0" box="[305,477,825,849]" class="Aves" genus="Vasa" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="19" pageNumber="306" phylum="Chordata" rank="species" species="deferentia">
|
||
<emphasis id="B90DEAEEFFB47C00FC5BFC8F13C2FCC0" box="[305,477,825,849]" italics="true" pageId="19" pageNumber="306">Vasa deferentia</emphasis>
|
||
</taxonomicName>
|
||
: one pair; anteriorly covered by thick tissues of body wall, run backwards parallel to axis of the body to 15, where become evident as single, soft tube, extending to posterior part of 17, and entering into body wall. In majority of dissected
|
||
<typeStatus id="54C2885EFFB47C00FC74FC2F1396FC23" box="[286,393,921,946]" pageId="19" pageNumber="306" type="paratype">paratypes</typeStatus>
|
||
, after passing vestigial parts of muscular ducts of the reduced prostatic gland of segment 17, run backwards and enter into body wall near intersegmental furrow 17/18. It is, however, not known if posterior part of vas deferens enters into wellmarked externally small depression in intersegmental furrow 17/18, or extends into noticeable depression in 18 that might be male pore.
|
||
<emphasis id="B90DEAEEFFB47C00FFC8FBAF1142FBA0" box="[674,861,1049,1073]" italics="true" pageId="19" pageNumber="306">Seminal vesicles</emphasis>
|
||
: two pairs; in 9 and 11, differing in size; in 9 very small, laterally attached to 9/10; in 11 larger, connected at septum 10/11, composite sacs extending dorsally. In dissected
|
||
<typeStatus id="54C2885EFFB47C00FE33FBEF11DCFBE3" box="[857,963,1113,1138]" pageId="19" pageNumber="306" type="paratype">paratypes</typeStatus>
|
||
some variations in size and shape were observed, which might be due to different states of individual maturity.
|
||
<emphasis id="B90DEAEEFFB47C00FC75FB2F13A1FB20" box="[287,446,1177,1201]" italics="true" pageId="19" pageNumber="306">Spermathecae</emphasis>
|
||
(
|
||
<figureCitation id="13422A79FFB47C00FCA1FB2F1002FB23" box="[459,541,1177,1202]" captionStart="Figs 13–15" captionStartId="18.[144,183,1551,1571]" captionText="Figs 13–15. Udeina petrosi sp. n. Paratype. 13. External ventral area of segments 15–21, 90 X. Pp = prostatic pore; Sg = seminal groove. 14. Spermatheca, 380 X. A = ampulla; Dv = diverticulum; Spd = spermathecal duct. 15. Prostate of segment 19 and vestigial prostatic duct of segment 17. Pg = prostatic gland of segment 19; Pd19 = prostatic duct of segment 19; Pd17 = vestigial prostatic duct of segment 17." pageId="19" pageNumber="306">Fig. 14</figureCitation>
|
||
): one pair in segment 8.Ampulla elongated, connected with duct above diverticulum. Duct moderately thick with ectal parts entering body wall in intersegmental furrow 7/8; short, not longer than
|
||
<subScript id="17FD34B9FFB47C00FE2CFB531151FB62" attach="right" box="[838,846,1253,1267]" fontSize="6" pageId="19" pageNumber="306">4</subScript>
|
||
<superScript id="7C0C9BB4FFB47C00FE2CFB6E1151FB77" attach="right" box="[838,846,1240,1254]" fontSize="6" pageId="19" pageNumber="306">1</superScript>
|
||
of the length of ampulla. Diverticulum attached at basal region of spermathecal duct, much smaller than ampulla, variable shapes and folds indicating two partially separated divisions.
|
||
<emphasis id="B90DEAEEFFB47C00FEA5FAAF1627FAA0" box="[975,1080,1305,1329]" italics="true" pageId="19" pageNumber="306">Prostates</emphasis>
|
||
(
|
||
<figureCitation id="13422A79FFB47C00F922FAAF12CBFAC3" captionStart="Figs 13–15" captionStartId="18.[144,183,1551,1571]" captionText="Figs 13–15. Udeina petrosi sp. n. Paratype. 13. External ventral area of segments 15–21, 90 X. Pp = prostatic pore; Sg = seminal groove. 14. Spermatheca, 380 X. A = ampulla; Dv = diverticulum; Spd = spermathecal duct. 15. Prostate of segment 19 and vestigial prostatic duct of segment 17. Pg = prostatic gland of segment 19; Pd19 = prostatic duct of segment 19; Pd17 = vestigial prostatic duct of segment 17." pageId="19" pageNumber="306">Fig. 15</figureCitation>
|
||
): one welldeveloped pair in 19, and one pair of small, vestigial prostatic ducts in 17; ectal openings in 17 not confirmed. Prostatic duct in segment 19 muscular, coiled or looped several times in the same segment, extending backwards into 20, evidently entering prostatic pore in segment 19. Prostatic gland much thicker than duct, multifolded, looped, passing irregularly backwards and forwards through segments 18–23; septa are pierced, or only conically extended. The rudimentary ducts of segment 17 differ in size and states of development, being however similar to ectal parts of ducts in segment 19, in six dissected individuals. These vestiges can occur paired, or single, either at left or right side of the body, but occur only in segment 17.
|
||
<emphasis id="B90DEAEEFFB47C00FED5F98F1655F9C0" box="[959,1098,1593,1617]" italics="true" pageId="19" pageNumber="306">Penial setae</emphasis>
|
||
:
|
||
<emphasis id="B90DEAEEFFB47C00F930F98F1669F9C0" box="[1114,1142,1593,1617]" italics="true" pageId="19" pageNumber="306">ab</emphasis>
|
||
setae of segment 19 transferred into penial setae. Differ in size, being probably in different states of development.
|
||
<emphasis id="B90DEAEEFFB47C00FCD2F9CF1117F900" box="[440,776,1657,1681]" italics="true" pageId="19" pageNumber="306">Penial setal retractor muscles</emphasis>
|
||
: commence at septum 18/19.
|
||
</paragraph>
|
||
<paragraph id="8BC636FCFFB37C07FDFAFF781391FED6" blockId="20.[144,1104,206,527]" pageId="20" pageNumber="307">
|
||
Biological notes and distribution: This species was collected in moist grassland soil, at edge of muddy depression, in the foothills of Drakensberg escarpment.Associated with the microchaetid
|
||
<taxonomicName id="4C794D7FFFB37C07FC3EFEB81118FEB6" authority="Plisko 2000" authorityName="Plisko" authorityYear="2000" box="[340,775,270,295]" class="Clitellata" family="Microchaetidae" genus="Proandricus" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Crassiclitellata" pageId="20" pageNumber="307" phylum="Annelida" rank="species" species="bergvillensis">
|
||
<emphasis id="B90DEAEEFFB37C07FC3EFEB8106AFEB7" box="[340,629,270,294]" italics="true" pageId="20" pageNumber="307">Proandricus bergvillensis</emphasis>
|
||
Plisko 2000
|
||
</taxonomicName>
|
||
, and other acanthodrilines of the genus
|
||
<taxonomicName id="4C794D7FFFB37C07FC6BFE981398FED7" authorityName="Pickford" authorityYear="1937" box="[257,391,302,326]" class="Clitellata" family="Acanthodrilidae" genus="Parachilota" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crassiclitellata" pageId="20" pageNumber="307" phylum="Annelida" rank="genus">
|
||
<emphasis id="B90DEAEEFFB37C07FC6BFE981398FED7" box="[257,391,302,326]" italics="true" pageId="20" pageNumber="307">Parachilota</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BC636FCFFB37C07FDFAFEE0111FFD9E" blockId="20.[144,1104,206,527]" pageId="20" pageNumber="307">
|
||
Discussion: A distinctive species, having in addition to the welldeveloped prostates in segment 19, the additional pair of vestigial parts of the prostatic ducts in 17. No developed prostatic glands in 17. These vestigial ducts may suggest reduction of the anterior pair of prostates, which are present in the species of the sister genus
|
||
<taxonomicName id="4C794D7FFFB37C07FEE6FE00160CFE5F" authorityName="Pickford" authorityYear="1937" box="[908,1043,438,462]" class="Clitellata" family="Acanthodrilidae" genus="Parachilota" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crassiclitellata" pageId="20" pageNumber="307" phylum="Annelida" rank="genus">
|
||
<emphasis id="B90DEAEEFFB37C07FEE6FE00160CFE5F" box="[908,1043,438,462]" italics="true" pageId="20" pageNumber="307">Parachilota</emphasis>
|
||
</taxonomicName>
|
||
. The spermathecal diverticulum is small, attached to the basal region of the spermathecal duct, with partially separated divisions, variable in size.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |