277 lines
37 KiB
XML
277 lines
37 KiB
XML
<document id="C2A5FB957495CED8812C822976B4431F" ID-DOI="10.2478/if-2019-0003" ID-ISSN="2533-4069" ID-Zenodo-Dep="4586641" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1614972125604" checkinUser="felipe" docAuthor="Štorch, Petr & Manda, Štěpán" docDate="2019" docId="039187E8022DB54A64D7FB9F5A32DFF2" docLanguage="en" docName="FossilImprint.75.1.44-58.pdf" docOrigin="Fossil Imprint 75 (1)" docSource="http://dx.doi.org/10.2478/if-2019-0003" docStyle="DocumentStyle:0EC54BF35E227CA4627F894F6BA30634.4:FossilImprint.2016-.journal_article" docStyleId="0EC54BF35E227CA4627F894F6BA30634" docStyleName="FossilImprint.2016-.journal_article" docStyleVersion="4" docTitle="Monograptus ambiguus JAEGER 1991" docType="treatment" docVersion="3" lastPageNumber="53" masterDocId="FFA8FF90022AB543676DFFD05852DB63" masterDocTitle="Little Known Homerian (Lower Silurian) Graptolites From Kosov Quarry Near Beroun, The Czech Republic" masterLastPageNumber="58" masterPageNumber="44" pageNumber="51" updateTime="1698881558317" updateUser="plazi" zenodo-license-document="CC-BY-NC-ND-3.0">
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<mods:title id="8C27FE66AE8D13F65091ED183B5BA7BF">Little Known Homerian (Lower Silurian) Graptolites From Kosov Quarry Near Beroun, The Czech Republic</mods:title>
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<mods:namePart id="922BB67EAB4F8F0F37DB107B58B43E6E">Štorch, Petr</mods:namePart>
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<mods:affiliation id="5DED8040B393B836AD3E44B27DBA3E61">Institute of Geology of the Czech Academy of Sciences, v. v. i., Rozvojová 269, 165 00 Praha 6, the Czech Republic; e-mail: storch @ gli. cas. cz.</mods:affiliation>
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<mods:namePart id="98C67FE6186EE33FAECC6E00E944F1C0">Manda, Štěpán</mods:namePart>
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<mods:affiliation id="CEF18284992C08EBEE852B9E1158C9EA">Czech Geological Survey, Klárov 3, 118 21 Praha 1, the Czech Republic; e-mail: stepan. manda @ geology. cz.</mods:affiliation>
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<mods:title id="B60D8BB31B0FEFB0F796DD985010745F">Fossil Imprint</mods:title>
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<mods:date id="19579BD7BCE3E9A2BFDA3425917E9154">2019</mods:date>
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<treatment id="039187E8022DB54A64D7FB9F5A32DFF2" LSID="urn:lsid:plazi:treatment:039187E8022DB54A64D7FB9F5A32DFF2" httpUri="http://treatment.plazi.org/id/039187E8022DB54A64D7FB9F5A32DFF2" lastPageId="9" lastPageNumber="53" pageId="7" pageNumber="51">
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<subSubSection id="C3226575022DB54464D7FB9F5D08DF05" box="[954,1370,1102,1127]" pageId="7" pageNumber="51" type="nomenclature">
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<paragraph id="8B8736FE022DB54464D7FB9F5D08DF05" blockId="7.[954,1370,1102,1127]" box="[954,1370,1102,1127]" pageId="7" pageNumber="51">
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<heading id="D0CF8192022DB54464D7FB9F5D08DF05" box="[954,1370,1102,1127]" centered="true" fontSize="8" level="2" pageId="7" pageNumber="51" reason="2">
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<taxonomicName id="4C384D7D022DB54464D7FB9F5D08DF05" authority="JAEGER, 1991" authorityName="JAEGER" authorityYear="1991" box="[954,1370,1102,1127]" class="Graptolithina" family="Monograptidae" genus="Monograptus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Graptoloidea" pageId="7" pageNumber="51" phylum="Hemichordata" rank="species" species="ambiguus">
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<emphasis id="B94CEAEC022DB54464D7FB9F5D08DF05" bold="true" box="[954,1370,1102,1127]" pageId="7" pageNumber="51">
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<emphasis id="B94CEAEC022DB54464D7FB9F5C93DF05" bold="true" box="[954,1217,1102,1126]" italics="true" pageId="7" pageNumber="51">Monograptus ambiguus</emphasis>
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<bibRefCitation id="EFA94B0F022DB54463A5FB9F5D45DF06" author="Jaeger, H." box="[1224,1303,1103,1127]" pageId="7" pageNumber="51" pagination="303 - 354" refId="ref9898" refString="Jaeger, H. (1991): Neue Standard-Graptolithenzonenfolge nach der " Grossen Krise " an der Wenlock / Ludlow-Gren- ze (Silur). - Neues Jahrbuch fur Geologie und Palaontologie, Abhandlungen, 182: 303 - 354. https: // doi. org / 10.1127 / njgpa / 182 / 1991 / 303" type="journal article" year="1991">JAEGER</bibRefCitation>
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, 1991
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</emphasis>
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||
</taxonomicName>
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||
</heading>
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</paragraph>
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||
</subSubSection>
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<subSubSection id="C3226575022DB54A64D7FBBF5A32DFF2" lastPageId="9" lastPageNumber="53" pageId="7" pageNumber="51" type="description">
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<paragraph id="8B8736FE022DB54464D7FBBF5D09DFE4" blockId="7.[954,1371,1134,1159]" box="[954,1371,1134,1159]" pageId="7" pageNumber="51">
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<figureCitation id="13032A7B022DB54464D7FBBF5C07DFE5" box="[954,1109,1134,1159]" captionStart="Text-fig" captionStartId="6.[125,206,1901,1922]" captionTargetBox="[127,1424,151,1877]" captionTargetId="figure-3@6.[125,1440,151,1950]" captionTargetPageId="6" captionText="Text-fig. 5. Monograptid graptolites from the upper lundgreni (a–h, l–m) and lower ludensis-gerhardi (i–k) biozones of the Kosov quarry.?a, c–d, f – Monograptus subflexilis PŘIBYL, 1941: a – PŠ 3874, slender juvenile specimen from sample K2/11; c – PŠ 3871/1, sample K2/13; d – PŠ 3871/2, sample K2/13; f – PŠ 3864, sample K2/12. b, e, g–h, l–m – Monograptus ambiguus JAEGER, 1991: b – PŠ 3899, sample K2/7; e – PŠ 3896, least ventrally curved specimen, sample K2/7; g – PŠ 3873, specimen showing small apertural spines in proximal thecae, sample K2/6; h – PŠ 3851, most ventrally curved specimen from sample K2/6; l – PŠ 3883, sample K2/6; m – PŠ 3866, sample K2/7. i–k – Pristiograptus auctus RICKARDS, 1965: i – PŠ 3848/2, sample K1/32; j – PŠ 3848/1, sample K1/32; k – PŠ 3839, sample K1/32. All specimens × 6, scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4586651" httpUri="https://zenodo.org/record/4586651/files/figure.png" pageId="7" pageNumber="51">Text-figs 5b, e</figureCitation>
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, g–h, l–m, 6c, e, 7b, d–e
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</paragraph>
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<paragraph id="8B8736FE022DB5446426FB4D5C10DFB2" blockId="7.[843,1482,1181,1322]" pageId="7" pageNumber="51">
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1965
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<taxonomicName id="4C384D7D022DB54464FBFB4E5C9EDFD1" authority="PRIBYL" authorityName="PRIBYL" authorityYear="1941" box="[918,1228,1181,1203]" class="Graptolithina" family="Monograptidae" genus="Monograptus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Graptoloidea" pageId="7" pageNumber="51" phylum="Hemichordata" rank="species" species="subflexilis">
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<emphasis id="B94CEAEC022DB54464FBFB4E5CD0DFD1" box="[918,1154,1181,1203]" italics="true" pageId="7" pageNumber="51">Monograptus subflexilis</emphasis>
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PŘIBYL
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</taxonomicName>
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; Jaworowski, text-fig. 3, fig. 6, pl. 1, fig. 6.
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</paragraph>
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<paragraph id="8B8736FE022DB5446426FB095C7EDE68" blockId="7.[843,1482,1181,1322]" pageId="7" pageNumber="51">
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||
1991
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||
<taxonomicName id="4C384D7D022DB54464FBFB095C2FDF8E" authority="Štorch & Manda, 2019" authorityName="Štorch & Manda" authorityYear="2019" box="[918,1149,1240,1262]" class="Graptolithina" family="Monograptidae" genus="Monograptus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Graptoloidea" pageId="7" pageNumber="51" phylum="Hemichordata" rank="species" species="ambiguus" status="sp. nov.">
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<emphasis id="B94CEAEC022DB54464FBFB095C2FDF8E" box="[918,1149,1240,1262]" italics="true" pageId="7" pageNumber="51">Monograptus ambiguus</emphasis>
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</taxonomicName>
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||
<taxonomicNameLabel id="A27F5797022DB54463EBFB095CEFDF8D" box="[1158,1213,1241,1262]" pageId="7" pageNumber="51" rank="species">sp. n.</taxonomicNameLabel>
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; Jaeger, pp. 343, 345, 347, figs 1, 29/8, 10.
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</paragraph>
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||
<paragraph id="8B8736FE022DB5446426FAC45D98DE4A" blockId="7.[843,1482,1181,1322]" box="[843,1482,1300,1322]" pageId="7" pageNumber="51">
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2019
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||
<taxonomicName id="4C384D7D022DB54464FBFAC45C92DE44" authority="JAEGER" authorityName="JAEGER" authorityYear="1991" box="[918,1216,1300,1321]" class="Graptolithina" family="Monograptidae" genus="Monograptus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Graptoloidea" pageId="7" pageNumber="51" phylum="Hemichordata" rank="species" species="ambiguus">
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<emphasis id="B94CEAEC022DB54464FBFAC45C28DE4A" box="[918,1146,1300,1321]" italics="true" pageId="7" pageNumber="51">Monograptus ambiguus</emphasis>
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JAEGER
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</taxonomicName>
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; Manda et al., figs 7a, 9d, e.
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</paragraph>
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<paragraph id="8B8736FE022DB544641CFA925D98DEF9" blockId="7.[843,1482,1345,1434]" pageId="7" pageNumber="51">
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H o l o t y p e. Specimen no MBg705.1 from the uppermost
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<emphasis id="B94CEAEC022DB54464ABFAB15C7FDE1A" box="[966,1069,1377,1401]" italics="true" pageId="7" pageNumber="51">lundgreni</emphasis>
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Biozone of the Gräfenwarth section,
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<collectingRegion id="49FCF81C022DB5446426FA525BE5DEF9" box="[843,951,1410,1434]" country="Germany" name="Thueringen" pageId="7" pageNumber="51">Thuringia</collectingRegion>
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,
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<collectingCountry id="F32F766E022DB54464ACFA525C7ADEF9" box="[961,1064,1410,1434]" name="Germany" pageId="7" pageNumber="51">Germany</collectingCountry>
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; figured by
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<bibRefCitation id="EFA94B0F022DB54463C9FA525D63DEF9" author="Jaeger, H." box="[1188,1329,1410,1434]" pageId="7" pageNumber="51" pagination="303 - 354" refId="ref9898" refString="Jaeger, H. (1991): Neue Standard-Graptolithenzonenfolge nach der " Grossen Krise " an der Wenlock / Ludlow-Gren- ze (Silur). - Neues Jahrbuch fur Geologie und Palaontologie, Abhandlungen, 182: 303 - 354. https: // doi. org / 10.1127 / njgpa / 182 / 1991 / 303" type="journal article" year="1991">Jaeger (1991</bibRefCitation>
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||
: fig. 29/8, 10).
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</paragraph>
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||
<paragraph id="8B8736FE022DB544641CFA625D92DD69" blockId="7.[843,1482,1457,1546]" pageId="7" pageNumber="51">
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M a t e r i a l. 59 mostly complete flattened rhabdosomes from the uppermost
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<emphasis id="B94CEAEC022DB5446357FA015CF3DE8A" box="[1082,1185,1489,1513]" italics="true" pageId="7" pageNumber="51">lundgreni</emphasis>
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Biozone of the Kosov 2 section (samples K2/8–K2/5; see
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||
<bibRefCitation id="EFA94B0F022DB54463DCFA225D24DD69" author="Manda, S. & Storch, P. & Fryda, J. & Slavik, L. & Tasaryova, Z." box="[1201,1398,1522,1546]" pageId="7" pageNumber="51" pagination="14 - 34" refId="ref10736" refString="Manda, S., Storch, P., Fryda, J., Slavik, L., Tasaryova, Z. (2019): The mid-Homerian (Silurian) biotic crisis in offshore settings of the Prague Synform, Czech Republic: integration of the graptolite fossil record with conodonts, shelly fauna and carbon isotope data. - Palaeogeogra- phy, Palaeoclimatology, Palaeoecology, 528: 14 - 34. https: // doi. org / 10.1016 / j. palaeo. 2019.04.026" type="journal article" year="2019">Manda et al. 2019</bibRefCitation>
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||
: fig. 2)
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||
</paragraph>
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<paragraph id="8B8736FE022DB544641CF9F25CC4DC1F" blockId="7.[843,1482,1569,1916]" pageId="7" pageNumber="51">
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D i a g n o s i s. After
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<bibRefCitation id="EFA94B0F022DB54463E0F9F25D6EDD5A" author="Jaeger, H." box="[1165,1340,1569,1594]" pageId="7" pageNumber="51" pagination="303 - 354" refId="ref9898" refString="Jaeger, H. (1991): Neue Standard-Graptolithenzonenfolge nach der " Grossen Krise " an der Wenlock / Ludlow-Gren- ze (Silur). - Neues Jahrbuch fur Geologie und Palaontologie, Abhandlungen, 182: 303 - 354. https: // doi. org / 10.1127 / njgpa / 182 / 1991 / 303" type="journal article" year="1991">Jaeger (1991)</bibRefCitation>
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, emended. Rhabdosome ventrally curved with slight dorsal curvature at the proximal end comprising th1–th2. Sicula straight or slightly dorsally curved. Thecae hooked, overlapping for about one-quarter their length, with steeply inclined interthecal septa, curved prothecae and ventral prothecal walls parallel to the rhabdosome axis. Dorso-ventral width increases gradually from about
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<quantity id="4CC09B1B022DB5446384F8D35D6EDC79" box="[1257,1340,1795,1819]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="7" pageNumber="51" unit="mm" value="0.7">0.7 mm</quantity>
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at th1 to the maximum
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<quantity id="4CC09B1B022DB54464D0F8F35C5FDC58" box="[957,1037,1827,1851]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="7" pageNumber="51" unit="mm" value="1.3">1.3 mm</quantity>
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most distally. Metathecal hooks comprise half the rhabdosome’s width in proximal thecae and onethird the width in distal thecae.
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</paragraph>
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<paragraph id="8B8736FE022DB54B641CF84B5D4BD350" blockId="7.[843,1483,1947,2100]" lastBlockId="8.[820,1460,1748,2100]" lastPageId="8" lastPageNumber="52" pageId="7" pageNumber="51">
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||
D e s c r i p t i o n. Rhabdosome is very gently dorsally curved over th1–2. Thereafter, it is moderately ventrally curved, until at least th14–33. Ventral curvature varies among specimens, but becomes almost negligible distally, which is well seen in more than
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<quantity id="4CC09B1B022DB5446323F7CC5CC9D350" box="[1102,1179,2076,2100]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" pageId="7" pageNumber="51" unit="mm" value="30.0">30 mm</quantity>
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||
long, mature rhabdosomes. The sicula is
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<quantity id="4CC09B1B0222B54B666EF90459C7DD8F" box="[259,405,1748,1772]" metricMagnitude="-3" metricUnit="m" metricValue="1.675" metricValueMax="1.85" metricValueMin="1.5" pageId="8" pageNumber="52" unit="mm" value="1.675" valueMax="1.85" valueMin="1.5">1.5–1.85 mm</quantity>
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||
long and
|
||
<quantity id="4CC09B1B0222B54B6566F9045AF8DD8F" box="[523,682,1748,1772]" metricMagnitude="-4" metricUnit="m" metricValue="2.75" metricValueMax="3.3" metricValueMin="2.2" pageId="8" pageNumber="52" unit="mm" value="0.275" valueMax="0.33" valueMin="0.22">0.22–0.33 mm</quantity>
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||
wide across its concave aperture, which is furnished with a tiny virgella. Sicular apex attains the middle or the dorsal wall of th2. Protheca of th1 begins with a prominent bulge
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<quantity id="4CC09B1B0222B54B65E6F8E658C1DC0C" metricMagnitude="-4" metricUnit="m" metricValue="1.85" metricValueMax="2.0" metricValueMin="1.7" pageId="8" pageNumber="52" unit="mm" value="0.185" valueMax="0.2" valueMin="0.17">0.17–0.2 mm</quantity>
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from the sicular aperture (e.g. see
|
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<figureCitation id="13032A7B0222B54B657CF8875A90DC0C" box="[529,706,1879,1903]" captionStart="Text-fig" captionStartId="6.[125,206,1901,1922]" captionTargetBox="[127,1424,151,1877]" captionTargetId="figure-3@6.[125,1440,151,1950]" captionTargetPageId="6" captionText="Text-fig. 5. Monograptid graptolites from the upper lundgreni (a–h, l–m) and lower ludensis-gerhardi (i–k) biozones of the Kosov quarry.?a, c–d, f – Monograptus subflexilis PŘIBYL, 1941: a – PŠ 3874, slender juvenile specimen from sample K2/11; c – PŠ 3871/1, sample K2/13; d – PŠ 3871/2, sample K2/13; f – PŠ 3864, sample K2/12. b, e, g–h, l–m – Monograptus ambiguus JAEGER, 1991: b – PŠ 3899, sample K2/7; e – PŠ 3896, least ventrally curved specimen, sample K2/7; g – PŠ 3873, specimen showing small apertural spines in proximal thecae, sample K2/6; h – PŠ 3851, most ventrally curved specimen from sample K2/6; l – PŠ 3883, sample K2/6; m – PŠ 3866, sample K2/7. i–k – Pristiograptus auctus RICKARDS, 1965: i – PŠ 3848/2, sample K1/32; j – PŠ 3848/1, sample K1/32; k – PŠ 3839, sample K1/32. All specimens × 6, scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4586651" httpUri="https://zenodo.org/record/4586651/files/figure.png" pageId="8" pageNumber="52">Text-fig. 5b, e, l</figureCitation>
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). It is strongly curved in a similar manner to all other prothecae, which have ventral walls either parallel to the rhabdosome axis or very gently (≤2°) inclined. Thecae overlap for about one-quarter their length; interthecal septa are steeply inclined at an angle of 50–85°. Metathecae are hooked, with proximally facing apertures, in proximal thecae probably furnished by short, paired lateral spines. Metathecal hook comprises from one-half the rhabdosome’s dorso-ventral width in proximal thecae to one-third in distal thecae. Dorso-ventral width of the rhabdosome increases gradually from
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<quantity id="4CC09B1B0222B54B6403F8875C55DC0C" box="[878,1031,1879,1903]" metricMagnitude="-4" metricUnit="m" metricValue="7.4" metricValueMax="8.4" metricValueMin="6.4" pageId="8" pageNumber="52" unit="mm" value="0.74" valueMax="0.84" valueMin="0.64">0.64–0.84 mm</quantity>
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at th1 (M = 0.72, n = 10) to 0.72–0.84 at th3 (M =
|
||
<quantity id="4CC09B1B0222B54B64F4F8A85BA8DCEC" box="[921,1018,1912,1936]" metricMagnitude="-4" metricUnit="m" metricValue="7.6" pageId="8" pageNumber="52" unit="mm" value="0.76">0.76 mm</quantity>
|
||
, n = 10),
|
||
<quantity id="4CC09B1B0222B54B6333F8A85CAADCEC" box="[1118,1272,1912,1936]" metricMagnitude="-4" metricUnit="m" metricValue="8.5" metricValueMax="9.3" metricValueMin="7.7" pageId="8" pageNumber="52" unit="mm" value="0.85" valueMax="0.93" valueMin="0.77">0.77–0.93 mm</quantity>
|
||
at th5 (M = 0.85, n = 10), and
|
||
<quantity id="4CC09B1B0222B54B64AFF8495C03DCD3" box="[962,1105,1945,1969]" metricMagnitude="-3" metricUnit="m" metricValue="1.06" metricValueMax="1.2" metricValueMin="0.92" pageId="8" pageNumber="52" unit="mm" value="1.06" valueMax="1.2" valueMin="0.92">0.92–1.2 mm</quantity>
|
||
at th10 (M = 0.99, n = 9) to the maximum
|
||
<quantity id="4CC09B1B0222B54B64C5F86A5C55DCB2" box="[936,1031,1978,2002]" metricMagnitude="-3" metricUnit="m" metricValue="1.27" pageId="8" pageNumber="52" unit="mm" value="1.27">1.27 mm</quantity>
|
||
attained by th40. 2TRD increases from
|
||
<quantity id="4CC09B1B0222B54B6459F80A5B9CDC91" box="[820,974,2010,2034]" metricMagnitude="-3" metricUnit="m" metricValue="1.505" metricValueMax="1.73" metricValueMin="1.28" pageId="8" pageNumber="52" unit="mm" value="1.505" valueMax="1.73" valueMin="1.28">1.28–1.73 mm</quantity>
|
||
at th2 (M =
|
||
<quantity id="4CC09B1B0222B54B633DF80A5CF1DC91" box="[1104,1187,2010,2034]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" pageId="8" pageNumber="52" unit="mm" value="1.6">1.6 mm</quantity>
|
||
, n = 10) to
|
||
<quantity id="4CC09B1B0222B54B6274F80A5DE0DC91" box="[1305,1458,2010,2034]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" metricValueMax="1.95" metricValueMin="1.45" pageId="8" pageNumber="52" unit="mm" value="1.7" valueMax="1.95" valueMin="1.45">1.45–1.95 mm</quantity>
|
||
at th10 (M = 1.71, n = 10). Distal thecae number
|
||
<quantity id="4CC09B1B0222B54B623AF82B5DC2D370" box="[1367,1424,2043,2067]" metricMagnitude="-1" metricUnit="m" metricValue="2.54" pageId="8" pageNumber="52" unit="in" value="10.0">10 in</quantity>
|
||
<quantity id="4CC09B1B0222B54B62F5F82B5B32D350" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="8" pageNumber="52" unit="mm" value="10.0">10 mm</quantity>
|
||
, with 2TRD ranging from
|
||
<quantity id="4CC09B1B0222B54B6310F7CC5D47D350" box="[1149,1301,2076,2100]" metricMagnitude="-3" metricUnit="m" metricValue="1.95" metricValueMax="2.1" metricValueMin="1.8" pageId="8" pageNumber="52" unit="mm" value="1.95" valueMax="2.1" valueMin="1.8">1.8 to 2.1 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
<caption id="DF4766760222B54B6710F9F05A37DDEE" ID-DOI="http://doi.org/10.5281/zenodo.4586653" ID-Zenodo-Dep="4586653" httpUri="https://zenodo.org/record/4586653/files/figure.png" pageId="8" pageNumber="52" startId="8.[125,206,1568,1589]" targetBox="[125,1440,151,1546]" targetPageId="8">
|
||
<paragraph id="8B8736FE0222B54B6710F9F05A37DDEE" blockId="8.[125,1440,1568,1677]" pageId="8" pageNumber="52">
|
||
<emphasis id="B94CEAEC0222B54B6710F9F05A27DD13" bold="true" pageId="8" pageNumber="52">
|
||
Text-fig. 6. Monograptid graptolites from the upper
|
||
<emphasis id="B94CEAEC0222B54B65E2F9F05AA2DD56" bold="true" box="[655,752,1568,1589]" italics="true" pageId="8" pageNumber="52">lundgreni</emphasis>
|
||
(b–e) and lower
|
||
<emphasis id="B94CEAEC0222B54B64F6F9F05C18DD56" bold="true" box="[923,1098,1568,1589]" italics="true" pageId="8" pageNumber="52">ludensis-gerhardi</emphasis>
|
||
(a) biozones of the Kosov quarry. a –
|
||
<taxonomicName id="4C384D7D0222B54B67C9F9ED5A72DD31" authority="RICKARDS, 1965" authorityName="RICKARDS" authorityYear="1965" box="[164,544,1597,1618]" class="Graptolithina" family="Rhabdinoporidae" genus="Pristiograptus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Graptoloidea" pageId="8" pageNumber="52" phylum="Hemichordata" rank="species" species="auctus">
|
||
<emphasis id="B94CEAEC0222B54B67C9F9ED592BDD31" bold="true" box="[164,377,1597,1618]" italics="true" pageId="8" pageNumber="52">Pristiograptus auctus</emphasis>
|
||
<bibRefCitation id="EFA94B0F0222B54B66ECF9ED59B2DD32" author="Rickards, R. B." box="[385,480,1597,1618]" pageId="8" pageNumber="52" pagination="247 - 271" refId="ref11182" refString="Rickards, R. B. (1965): New Silurian graptolites from the Howgill Fells (Northern England). - Palaeontology, 8 (2): 247 - 271." type="journal article" year="1965">RICKARDS</bibRefCitation>
|
||
, 1965
|
||
</taxonomicName>
|
||
: PŠ 3848/1, sample K1/32. b, d –
|
||
<taxonomicName id="4C384D7D0222B54B64EDF9ED5CACDD31" authority="PRIBYL, 1941" authorityName="PRIBYL" authorityYear="1941" box="[896,1278,1597,1618]" class="Graptolithina" family="Monograptidae" genus="Monograptus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Graptoloidea" pageId="8" pageNumber="52" phylum="Hemichordata" rank="species" species="subflexilis">
|
||
<emphasis id="B94CEAEC0222B54B64EDF9ED5C20DD31" bold="true" box="[896,1138,1597,1618]" italics="true" pageId="8" pageNumber="52">Monograptus subflexilis</emphasis>
|
||
<bibRefCitation id="EFA94B0F0222B54B6317F9ED5CEFDD32" author="Pribyl, A." box="[1146,1213,1597,1618]" pageId="8" pageNumber="52" pagination="1 - 10" refId="ref11096" refString="Pribyl, A. (1941): O nekolika novych druzich graptolitu z ceskeho siluru [On several new graptolite species from the Silurian of Bohemia]. - Rozpravy 2. Tridy Ceske akademie, 51 (7): 1 - 10. (in Czech)" type="journal article" year="1941">PŘIBYL</bibRefCitation>
|
||
, 1941
|
||
</taxonomicName>
|
||
: b – PŠ 3871/2, sample K2/13; d – PŠ 3864, sample K2/12. c, e –
|
||
</emphasis>
|
||
<taxonomicName id="4C384D7D0222B54B6510F98B5BA9DD13" authority="JAEGER, 1991" authorityName="JAEGER" authorityYear="1991" box="[637,1019,1627,1648]" class="Graptolithina" family="Monograptidae" genus="Monograptus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Graptoloidea" pageId="8" pageNumber="52" phylum="Hemichordata" rank="species" species="ambiguus">
|
||
<emphasis id="B94CEAEC0222B54B6510F98B5BE9DD0D" bold="true" box="[637,955,1627,1648]" pageId="8" pageNumber="52">
|
||
<emphasis id="B94CEAEC0222B54B6510F98B5B39DD13" bold="true" box="[637,875,1627,1648]" italics="true" pageId="8" pageNumber="52">Monograptus ambiguus</emphasis>
|
||
<bibRefCitation id="EFA94B0F0222B54B641EF98B5BE9DD0D" author="Jaeger, H." box="[883,955,1627,1648]" pageId="8" pageNumber="52" pagination="303 - 354" refId="ref9898" refString="Jaeger, H. (1991): Neue Standard-Graptolithenzonenfolge nach der " Grossen Krise " an der Wenlock / Ludlow-Gren- ze (Silur). - Neues Jahrbuch fur Geologie und Palaontologie, Abhandlungen, 182: 303 - 354. https: // doi. org / 10.1127 / njgpa / 182 / 1991 / 303" type="journal article" year="1991">JAEGER</bibRefCitation>
|
||
</emphasis>
|
||
, 1991
|
||
</taxonomicName>
|
||
: c – PŠ 3866, sample K2/7; e – PŠ 3896, sample K2/7. All specimens × 6, scale bar 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B8736FE0223B54A67CBFF485A32DFF2" blockId="9.[128,768,151,1169]" pageId="9" pageNumber="53">
|
||
R e m a r k s. Rich material of
|
||
<taxonomicName id="4C384D7D0223B54A6690FF485B52DBCC" authorityName="JAEGER" authorityYear="1991" box="[509,768,151,175]" class="Graptolithina" family="Monograptidae" genus="Monograptus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Graptoloidea" pageId="9" pageNumber="53" phylum="Hemichordata" rank="species" species="ambiguus">
|
||
<emphasis id="B94CEAEC0223B54A6690FF485B52DBCC" box="[509,768,151,175]" italics="true" pageId="9" pageNumber="53">Monograptus ambiguus</emphasis>
|
||
</taxonomicName>
|
||
found in the uppermost
|
||
<emphasis id="B94CEAEC0223B54A66E4FF6859A2DBB3" box="[393,496,184,208]" italics="true" pageId="9" pageNumber="53">lundgreni</emphasis>
|
||
Biozone of the Kosov 2 section (
|
||
<bibRefCitation id="EFA94B0F0223B54A67B5FF0859C5DB93" author="Manda, S. & Storch, P. & Fryda, J. & Slavik, L. & Tasaryova, Z." box="[216,407,216,240]" pageId="9" pageNumber="53" pagination="14 - 34" refId="ref10736" refString="Manda, S., Storch, P., Fryda, J., Slavik, L., Tasaryova, Z. (2019): The mid-Homerian (Silurian) biotic crisis in offshore settings of the Prague Synform, Czech Republic: integration of the graptolite fossil record with conodonts, shelly fauna and carbon isotope data. - Palaeogeogra- phy, Palaeoclimatology, Palaeoecology, 528: 14 - 34. https: // doi. org / 10.1016 / j. palaeo. 2019.04.026" type="journal article" year="2019">Manda et al. 2019</bibRefCitation>
|
||
) matches the limited
|
||
<typeStatus id="5483885C0223B54A6518FF085AF1DB93" box="[629,675,216,240]" pageId="9" pageNumber="53">type</typeStatus>
|
||
material of the species described by
|
||
<bibRefCitation id="EFA94B0F0223B54A66F1FF285A7EDA73" author="Jaeger, H." box="[412,556,248,272]" pageId="9" pageNumber="53" pagination="303 - 354" refId="ref9898" refString="Jaeger, H. (1991): Neue Standard-Graptolithenzonenfolge nach der " Grossen Krise " an der Wenlock / Ludlow-Gren- ze (Silur). - Neues Jahrbuch fur Geologie und Palaontologie, Abhandlungen, 182: 303 - 354. https: // doi. org / 10.1127 / njgpa / 182 / 1991 / 303" type="journal article" year="1991">Jaeger (1991)</bibRefCitation>
|
||
from the uppermost
|
||
<emphasis id="B94CEAEC0223B54A67ECFEC85915DA4C" box="[129,327,280,303]" italics="true" pageId="9" pageNumber="53">
|
||
<taxonomicName id="4C384D7D0223B54A67ECFEC85958DA4C" authorityName="Prybil" authorityYear="1967" box="[129,266,280,303]" class="Graptolithina" genus="Testograptus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="9" pageNumber="53" phylum="Hemichordata" rank="genus">Testograptus</taxonomicName>
|
||
testis
|
||
</emphasis>
|
||
Subzone (uppermost
|
||
<emphasis id="B94CEAEC0223B54A655DFEC85AC5DA53" box="[560,663,280,304]" italics="true" pageId="9" pageNumber="53">lundgreni</emphasis>
|
||
Biozone) of the Gräfenwarth section, Thuringia in rhabdosome form, thecal form and measured parameters. The present material from the Kosov quarry, however, revealed remarkable variability in the ventral curvature of the rhabdosome, and has shown that specimens from the uppermost
|
||
<emphasis id="B94CEAEC0223B54A65F4FE685B52DAB3" box="[665,768,440,464]" italics="true" pageId="9" pageNumber="53">lundgreni</emphasis>
|
||
Biozone of Bartoszyce and Gołdap boreholes in NE
|
||
<collectingCountry id="F32F766E0223B54A67ECFE28589CD973" box="[129,206,504,528]" name="Poland" pageId="9" pageNumber="53">Poland</collectingCountry>
|
||
, refered to
|
||
<taxonomicName id="4C384D7D0223B54A6624FE295A8FD973" authority="PRIBYL, 1941" authorityName="PRIBYL" authorityYear="1941" box="[329,733,504,528]" class="Graptolithina" family="Monograptidae" genus="Monograptus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Graptoloidea" pageId="9" pageNumber="53" phylum="Hemichordata" rank="species" species="subflexilis">
|
||
<emphasis id="B94CEAEC0223B54A6624FE295A19D973" box="[329,587,504,528]" italics="true" pageId="9" pageNumber="53">Monograptus subflexilis</emphasis>
|
||
<bibRefCitation id="EFA94B0F0223B54A653FFE285A8FD973" author="Pribyl, A." box="[594,733,504,528]" pageId="9" pageNumber="53" pagination="1 - 10" refId="ref11096" refString="Pribyl, A. (1941): O nekolika novych druzich graptolitu z ceskeho siluru [On several new graptolite species from the Silurian of Bohemia]. - Rozpravy 2. Tridy Ceske akademie, 51 (7): 1 - 10. (in Czech)" type="journal article" year="1941">PŘIBYL, 1941</bibRefCitation>
|
||
</taxonomicName>
|
||
by
|
||
<bibRefCitation id="EFA94B0F0223B54A67ECFDC85903D953" author="Jaworowski, K." box="[129,337,536,560]" pageId="9" pageNumber="53" pagination="511 - 526" refId="ref9992" refString="Jaworowski, K. (1965): Strop warstw posleckich w obnaze- niu litewskim a granica wenlok - ludlow [Top of the Paslek Beds in the Lithuanian depression and the Wen- lockian - Ludlowian boundary]. - Kwartalnik Geologiczny, 9: 511 - 526. (in Polish)" type="journal article" year="1965">Jaworowski (1965)</bibRefCitation>
|
||
, also belong to
|
||
<taxonomicName id="4C384D7D0223B54A6692FDC95ADED953" authorityName="JAEGER" authorityYear="1991" box="[511,652,536,560]" class="Graptolithina" family="Monograptidae" genus="Monograptus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Graptoloidea" pageId="9" pageNumber="53" phylum="Hemichordata" rank="species" species="ambiguus">
|
||
<emphasis id="B94CEAEC0223B54A6692FDC95ADED953" box="[511,652,536,560]" italics="true" pageId="9" pageNumber="53">M. ambiguus</emphasis>
|
||
</taxonomicName>
|
||
. Thus, the species, so far recorded in
|
||
<collectingCountry id="F32F766E0223B54A66DEFDE85A45D933" box="[435,535,568,592]" name="Germany" pageId="9" pageNumber="53">Germany</collectingCountry>
|
||
, the
|
||
<collectingCountry id="F32F766E0223B54A653CFDE85AADD933" box="[593,767,568,592]" name="Czech Republic" pageId="9" pageNumber="53">Czech Republic</collectingCountry>
|
||
and
|
||
<collectingCountry id="F32F766E0223B54A67DEFD885952D913" box="[179,256,600,624]" name="Poland" pageId="9" pageNumber="53">Poland</collectingCountry>
|
||
, qualifies as a useful biostratigraphic marker of the uppermost part of the
|
||
<emphasis id="B94CEAEC0223B54A66D0FDA85A76D9F3" box="[445,548,632,656]" italics="true" pageId="9" pageNumber="53">lundgreni</emphasis>
|
||
Biozone in Europe.
|
||
<taxonomicName id="4C384D7D0223B54A67ECFD4959D0D9D3" authorityName="JAEGER" authorityYear="1991" box="[129,386,664,688]" class="Graptolithina" family="Monograptidae" genus="Monograptus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Graptoloidea" pageId="9" pageNumber="53" phylum="Hemichordata" rank="species" species="ambiguus">
|
||
<emphasis id="B94CEAEC0223B54A67ECFD4959D0D9D3" box="[129,386,664,688]" italics="true" pageId="9" pageNumber="53">Monograptus ambiguus</emphasis>
|
||
</taxonomicName>
|
||
can be readily distinguished from
|
||
<taxonomicName id="4C384D7D0223B54A67ECFD695945D9B3" authorityName="PRIBYL" authorityYear="1941" box="[129,279,696,720]" class="Graptolithina" family="Monograptidae" genus="Monograptus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Graptoloidea" pageId="9" pageNumber="53" phylum="Hemichordata" rank="species" species="subflexilis">
|
||
<emphasis id="B94CEAEC0223B54A67ECFD695945D9B3" box="[129,279,696,720]" italics="true" pageId="9" pageNumber="53">M. subflexilis</emphasis>
|
||
</taxonomicName>
|
||
by its very short, gently dorsally curved proximal portion of the rhabdosome, more prominent subsequent ventral curvature, much narrower and more slowly increasing dorso-ventral width, relatively longer ventral prothecal walls, almost parallel to the stipe, and much smaller apertural hooks. Maximum width of the rhabdosome does not exceed
|
||
<quantity id="4CC09B1B0223B54A6643FCA8592CD8F3" box="[302,382,888,912]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="9" pageNumber="53" unit="mm" value="1.3">1.3 mm</quantity>
|
||
in
|
||
<taxonomicName id="4C384D7D0223B54A66F2FCA95A7BD8F3" authorityName="JAEGER" authorityYear="1991" box="[415,553,888,912]" class="Graptolithina" family="Monograptidae" genus="Monograptus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Graptoloidea" pageId="9" pageNumber="53" phylum="Hemichordata" rank="species" species="ambiguus">
|
||
<emphasis id="B94CEAEC0223B54A66F2FCA95A7BD8F3" box="[415,553,888,912]" italics="true" pageId="9" pageNumber="53">M. ambiguus</emphasis>
|
||
</taxonomicName>
|
||
, in contrast to about
|
||
<quantity id="4CC09B1B0223B54A67ECFC4858EED8D3" box="[129,188,920,944]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="9" pageNumber="53" unit="mm" value="3.0">3 mm</quantity>
|
||
in
|
||
<taxonomicName id="4C384D7D0223B54A67B0FC495939D8D3" authorityName="PRIBYL" authorityYear="1941" box="[221,363,920,944]" class="Graptolithina" family="Monograptidae" genus="Monograptus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Graptoloidea" pageId="9" pageNumber="53" phylum="Hemichordata" rank="species" species="subflexilis">
|
||
<emphasis id="B94CEAEC0223B54A67B0FC495939D8D3" box="[221,363,920,944]" italics="true" pageId="9" pageNumber="53">M. subflexilis</emphasis>
|
||
</taxonomicName>
|
||
. The simply hooked thecal form of
|
||
<taxonomicName id="4C384D7D0223B54A658FFC4958BBD8B3" authorityName="JAEGER" authorityYear="1991" class="Graptolithina" family="Monograptidae" genus="Monograptus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Graptoloidea" pageId="9" pageNumber="53" phylum="Hemichordata" rank="species" species="ambiguus">
|
||
<emphasis id="B94CEAEC0223B54A658FFC4958BBD8B3" italics="true" pageId="9" pageNumber="53">M. ambiguus</emphasis>
|
||
</taxonomicName>
|
||
is closely similar to that of
|
||
<taxonomicName id="4C384D7D0223B54A6572FC695AFDD8B3" authorityName="PRIBYL" authorityYear="1941" box="[543,687,952,976]" class="Graptolithina" family="Monograptidae" genus="Monograptus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Graptoloidea" pageId="9" pageNumber="53" phylum="Hemichordata" rank="species" species="subflexilis">
|
||
<emphasis id="B94CEAEC0223B54A6572FC695AFDD8B3" box="[543,687,952,976]" italics="true" pageId="9" pageNumber="53">M. subflexilis</emphasis>
|
||
</taxonomicName>
|
||
, which indicates that, despite superficial similarity to Homerian species with S-shaped or ventrally curved rhabdosomes and coiled, streptograptid-like thecae (
|
||
<emphasis id="B94CEAEC0223B54A669BFBC95AADDF53" box="[502,767,1048,1072]" italics="true" pageId="9" pageNumber="53">
|
||
<taxonomicName id="4C384D7D0223B54A669BFBC95AD4DF53" authorityName="GEINITZ" authorityYear="1852" box="[502,646,1049,1072]" class="Graptolithina" family="Monograptidae" genus="Monograptus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Graptoloidea" pageId="9" pageNumber="53" phylum="Hemichordata" rank="genus">Monograptus</taxonomicName>
|
||
retroflexus
|
||
</emphasis>
|
||
<bibRefCitation id="EFA94B0F0223B54A67ECFBE9597ADF32" author="Tullberg, S. A." box="[129,296,1081,1105]" pageId="9" pageNumber="53" pagination="1 - 43" refId="ref11545" refString="Tullberg, S. A. (1883): Skanes Graptoliter, 2 [Graptolites of Scania, 2]. - Sveriges geologiska Undersokning, Serie C, 55: 1 - 43. (in Swedish)" type="journal article" year="1883">TULLBERG, 1883</bibRefCitation>
|
||
and
|
||
<taxonomicName id="4C384D7D0223B54A6637FBE95AA0DF32" authority="JAEGER, 1991" authorityName="JAEGER" authorityYear="1991" box="[346,754,1081,1105]" class="Graptolithina" family="Monograptidae" genus="Monograptus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Graptoloidea" pageId="9" pageNumber="53" phylum="Hemichordata" rank="species" species="serexiguus">
|
||
<emphasis id="B94CEAEC0223B54A6637FBE95A32DF33" box="[346,608,1081,1104]" italics="true" pageId="9" pageNumber="53">Monograptus serexiguus</emphasis>
|
||
<bibRefCitation id="EFA94B0F0223B54A650BFBE95AA0DF32" author="Jaeger, H." box="[614,754,1081,1105]" pageId="9" pageNumber="53" pagination="303 - 354" refId="ref9898" refString="Jaeger, H. (1991): Neue Standard-Graptolithenzonenfolge nach der " Grossen Krise " an der Wenlock / Ludlow-Gren- ze (Silur). - Neues Jahrbuch fur Geologie und Palaontologie, Abhandlungen, 182: 303 - 354. https: // doi. org / 10.1127 / njgpa / 182 / 1991 / 303" type="journal article" year="1991">JAEGER, 1991</bibRefCitation>
|
||
</taxonomicName>
|
||
),
|
||
<taxonomicName id="4C384D7D0223B54A67EDFB89595EDF12" authorityName="JAEGER" authorityYear="1991" box="[128,268,1113,1137]" class="Graptolithina" family="Monograptidae" genus="Monograptus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Graptoloidea" pageId="9" pageNumber="53" phylum="Hemichordata" rank="species" species="ambiguus">
|
||
<emphasis id="B94CEAEC0223B54A67EDFB89595EDF12" box="[128,268,1113,1137]" italics="true" pageId="9" pageNumber="53">M. ambiguus</emphasis>
|
||
</taxonomicName>
|
||
is a true, although more derived offshoot from the
|
||
<emphasis id="B94CEAEC0223B54A67C5FBA95931DFF2" box="[168,355,1145,1169]" italics="true" pageId="9" pageNumber="53">priodon-flemingii</emphasis>
|
||
monograptid stem line.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |