334 lines
57 KiB
XML
334 lines
57 KiB
XML
<document id="1AB54D2A423056C1C3170CF3731AA4ED" ID="10.11646/zootaxa.4178.3.2" ID-DOI="10.11646/zootaxa.4178.3.2" ID-GBIF-Dataset="e5f947fa-f818-4487-93d6-eadb93d8f537" ID-ISSN="1175-5326" ID-Zenodo-Dep="263560" ID-ZooBank="A3990425-8A2A-4419-B2F0-986154E7173C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1477464814593" checkinUser="plazi" docAuthor="Korn, Michael & Hundsdoerfer, Anna K." docDate="2016" docId="039187C9C6471F1BD3C4FA9AAF82FBE5" docLanguage="en" docName="zootaxa.4178.3.2.pdf" docOrigin="Zootaxa 4178 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Triops multifidus Korn, sp. nov." docType="treatment" docVersion="9" lastPageNumber="343" masterDocId="FFA8FFB1C64B1F14D353FFC4AC13FFDA" masterDocTitle="Molecular phylogeny, morphology and taxonomy of Moroccan Triops granarius (Lucas, 1864) (Crustacea: Notostraca), with the description of two new species" masterLastPageNumber="346" masterPageNumber="328" pageNumber="340" updateTime="1698390773422" updateUser="plazi">
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<mods:title id="44DC0CB631FC90BD3B99ECA1D1D85119">Molecular phylogeny, morphology and taxonomy of Moroccan Triops granarius (Lucas, 1864) (Crustacea: Notostraca), with the description of two new species</mods:title>
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<mods:namePart id="8151EEEB619FAA79BC4849E6DBC537D2">Korn, Michael</mods:namePart>
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<mods:namePart id="E45413A22DF735F5DE029614ED156B68">Hundsdoerfer, Anna K.</mods:namePart>
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<mods:typeOfResource id="B441522219F23AFEEE9841BED40C22EF">text</mods:typeOfResource>
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<mods:title id="4B60CEC2A15488D1354BE822E5909772">Zootaxa</mods:title>
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<mods:date id="844E463D0E3CB550C91423C557ECF224">2016</mods:date>
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<mods:number id="4881CAC3081C9214E201F0C9DC24723B">4178</mods:number>
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<mods:identifier id="B69E3763D40359E7DB76F4F15EB65539" type="DOI">10.11646/zootaxa.4178.3.2</mods:identifier>
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<treatment id="039187C9C6471F1BD3C4FA9AAF82FBE5" ID-DOI="http://doi.org/10.5281/zenodo.5625930" ID-GBIF-Taxon="124558128" ID-Zenodo-Dep="5625930" LSID="urn:lsid:plazi:treatment:039187C9C6471F1BD3C4FA9AAF82FBE5" httpUri="http://treatment.plazi.org/id/039187C9C6471F1BD3C4FA9AAF82FBE5" lastPageId="15" lastPageNumber="343" pageId="12" pageNumber="340">
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<subSubSection id="C3226554C6471F18D3C4FA9AAD14FA43" pageId="12" pageNumber="340" type="nomenclature">
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<paragraph id="8B8736DFC6471F18D3C4FA9AAE05FAA2" blockId="12.[151,534,1374,1434]" box="[151,534,1374,1400]" pageId="12" pageNumber="340">
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<heading id="D0CF81B3C6471F18D3C4FA9AAE05FAA2" bold="true" box="[151,534,1374,1400]" fontSize="11" level="1" pageId="12" pageNumber="340" reason="1">
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<emphasis id="B94CEACDC6471F18D3C4FA9AAE05FAA2" bold="true" box="[151,534,1374,1400]" pageId="12" pageNumber="340">
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<taxonomicName id="4C384D5CC6471F18D3C4FA9AADA2FAA2" ID-CoL="58WLT" authority="Korn" authorityName="Korn" box="[151,433,1374,1400]" class="Branchiopoda" family="Triopsidae" genus="Triops" kingdom="Animalia" order="Notostraca" pageId="12" pageNumber="340" phylum="Arthropoda" rank="species" species="multifidus" status="sp. nov.">
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<emphasis id="B94CEACDC6471F18D3C4FA9AAD74FAA2" bold="true" box="[151,359,1374,1400]" italics="true" pageId="12" pageNumber="340">Triops multifidus</emphasis>
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Korn
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</taxonomicName>
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<taxonomicNameLabel id="A27F57B6C6471F18D2EBFA9AAE05FAA2" box="[440,534,1374,1400]" pageId="12" pageNumber="340" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8B8736DFC6471F18D3C4FA45AD14FA43" blockId="12.[151,534,1374,1434]" box="[151,263,1409,1434]" pageId="12" pageNumber="340">
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(
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<figureCitation id="13032A5AC6471F18D3CCFA45ACF0FA43" box="[159,227,1409,1434]" captionStart="FIGURE 8" captionStartId="14.[151,250,1778,1800]" captionTargetBox="[263,1305,193,1723]" captionTargetId="figure@14.[263,1305,193,1725]" captionTargetPageId="14" captionText="FIGURE 8. Triops multifidus sp. nov. A, endopodite and 5 th endite of 2 nd trunk limb of the holotype (male), anterior view. B, endopodite of 2 nd trunk limb of a female of 2.3 mm telson width (from the type locality), anterior view. C, endopodite of 6 th trunk limb of a male of 2.3 mm telson width (from the type locality), anterior view. D, endopodite and 5 th endite of 2 nd trunk limb of a male of 2.2 mm telson width (from population 10), anterior view. E, second antenna of a second instar larva (from the type locality) as seen in a shed exuvium, ventral view. F, general aspect of adult male (showing a specimen of population 13). G, variation in the shape of the nuchal organ: examples (from left to right) from specimen of 2.2 (population 10) 1.8 (population 12, female) 2.4 (holotype) and 2.2 mm telson width (population 8). H, telson of the holotype, dorsal view. I, telson of a male of 2.3 mm telson width (from the type locality), lateral view." httpUri="https://zenodo.org/record/263566/files/figure.png" pageId="12" pageNumber="340">Fig. 8</figureCitation>
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,
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<figureCitation id="13032A5AC6471F18D3BCFA45ACEDFA43" box="[239,254,1409,1433]" captionStart="FIGURE 9" captionStartId="15.[151,250,896,918]" captionTargetBox="[153,1433,195,871]" captionTargetId="figure@15.[151,1436,193,875]" captionTargetPageId="15" captionText="FIGURE 9. Triops multifidus sp. nov. A, carapace of the holotype (male), right lateral view. B, second maxilla of the holotype, anterior view. C, detail of the 1 st trunk limb of the holotype, anterior view (note that the epipodite is missing in this sample, possibly due to injury)." httpUri="https://zenodo.org/record/263567/files/figure.png" pageId="12" pageNumber="340">9</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C3226554C6471F18D3C4FA07A835FA06" box="[151,1062,1475,1500]" pageId="12" pageNumber="340" type="description">
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<paragraph id="8B8736DFC6471F18D3C4FA07A835FA06" blockId="12.[151,1437,1475,2009]" box="[151,1062,1475,1500]" pageId="12" pageNumber="340">
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This species refers to the phylogenetic lineage ‘
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<taxonomicName id="4C384D5CC6471F18D1FEFA01AF79FA06" box="[685,874,1477,1500]" class="Branchiopoda" family="Triopsidae" genus="Triops" kingdom="Animalia" order="Notostraca" pageId="12" pageNumber="340" phylum="Arthropoda" rank="species" species="granarius">
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<emphasis id="B94CEACDC6471F18D1FEFA01AF79FA06" box="[685,874,1477,1500]" italics="true" pageId="12" pageNumber="340">Triops granarius</emphasis>
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</taxonomicName>
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8’ (see
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<tableCitation id="C6BA0364C6471F18D090FA07A806FA06" box="[963,1045,1475,1500]" captionStart="TABLE 1" captionStartId="2.[151,239,152,175]" captionTargetBox="[159,1415,298,1685]" captionTargetPageId="2" captionText="TABLE 1. Ingroup samples from western Morocco included in our molecular analyses. Voucher numbers refer to tissue vouchers in the ‘ MTD-TW’ tissue collection of the Museum of Zoology, Senckenberg Dresden (Germany). Accession numbers in italics refer to sequences previously published in GenBank. Locations of sampled populations are shown in Fig. 1 B. Previously studied sampling sites are indicated referring to Thiéry (1987)." httpUri="http://table.plazi.org/id/DF476657C6491F16D3C4FF5CA81AFED5" pageId="12" pageNumber="340" tableUuid="DF476657C6491F16D3C4FF5CA81AFED5">Table 1</tableCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="C3226554C6471F18D394FA23A8BFF999" pageId="12" pageNumber="340" type="etymology">
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<paragraph id="8B8736DFC6471F18D394FA23A8BFF999" blockId="12.[151,1437,1475,2009]" pageId="12" pageNumber="340">
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<emphasis id="B94CEACDC6471F18D394FA23AD5DF9DA" bold="true" box="[199,334,1511,1536]" pageId="12" pageNumber="340">Etymology.</emphasis>
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The name refers to the strongly increased number of swimming setae on the exopod of the 2nd antenna as seen in early instars. With a count of up to 15, the species shows a doubling or tripling of the usual set of 5–7 setae that has hitherto been reported for
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<taxonomicName id="4C384D5CC6471F18D1D9F9EFAF14F998" box="[650,775,1579,1602]" class="Branchiopoda" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notostraca" pageId="12" pageNumber="340" phylum="Arthropoda" rank="order">Notostraca</taxonomicName>
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. This name is formed as an adjective.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3226554C6471F18D394F988AF5CF8F6" pageId="12" pageNumber="340" type="materials_examined">
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<paragraph id="8B8736DFC6471F18D394F988AE8EF95F" blockId="12.[151,1437,1475,2009]" pageId="12" pageNumber="340">
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<materialsCitation id="3B503C82C6471F18D394F988AE8EF95F" ID-GBIF-Occurrence="1324200584" country="Morocco" county="Tiznit" location="Temporary" municipality="Agadir" pageId="12" pageNumber="340" specimenCount="1" typeStatus="holotype">
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<emphasis id="B94CEACDC6471F18D394F988AD4CF9BF" bold="true" box="[199,351,1612,1637]" pageId="12" pageNumber="340">
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<typeStatus id="5483887DC6471F18D394F988AD11F9BF" box="[199,258,1612,1637]" pageId="12" pageNumber="340">Type</typeStatus>
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locality
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</emphasis>
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.
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<location id="8EE76004C6471F18D23DF989ADFAF9BF" LSID="urn:lsid:plazi:treatment:039187C9C6471F1BD3C4FA9AAF82FBE5:8EE76004C6471F18D23DF989ADFAF9BF" box="[366,489,1613,1637]" country="Morocco" county="Tiznit" municipality="Agadir" name="Temporary" pageId="12" pageNumber="340">Temporary</location>
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pond located at the crossroads of road ‘N1’ and the road leading to
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<location id="8EE76004C6471F18D7A5F988A929F9BF" LSID="urn:lsid:plazi:treatment:039187C9C6471F1BD3C4FA9AAF82FBE5:8EE76004C6471F18D7A5F988A929F9BF" box="[1270,1338,1612,1637]" country="Morocco" county="Tiznit" municipality="Agadir" name="Imjad" pageId="12" pageNumber="340">Imjad</location>
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, approx. halfway between
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<collectingMunicipality id="6BE3ACA5C6471F18D20FF9A9ADB9F95C" box="[348,426,1645,1670]" pageId="12" pageNumber="340">Agadir</collectingMunicipality>
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and
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<collectingCounty id="62E64E53C6471F18D2B3F9A9AE36F95C" box="[480,549,1645,1670]" pageId="12" pageNumber="340">Tiznit</collectingCounty>
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,
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<collectingCountry id="F32F764FC6471F18D163F9AAAE8AF95F" box="[560,665,1646,1669]" name="Morocco" pageId="12" pageNumber="340">Morocco</collectingCountry>
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.
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</materialsCitation>
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</paragraph>
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<paragraph id="8B8736DFC6471F18D394F94AADFBF912" blockId="12.[151,1437,1475,2009]" pageId="12" pageNumber="340">
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<emphasis id="B94CEACDC6471F18D394F94AADFCF97D" bold="true" box="[199,495,1678,1703]" pageId="12" pageNumber="340">Type material. Holotype.</emphasis>
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Adult male, Museum of Zoology Dresden (MTD-Cru S 185, population no. 17, leg. Michael Korn
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<date id="FF86101FC6471F18D269F96BADCFF912" box="[314,476,1711,1736]" pageId="12" pageNumber="340" value="2005-04-15">15 April 2005</date>
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).
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</paragraph>
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<paragraph id="8B8736DFC6471F18D394F915AF5CF8F6" blockId="12.[151,1437,1475,2009]" pageId="12" pageNumber="340">
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<emphasis id="B94CEACDC6471F18D394F915ADEEF930" bold="true" box="[199,509,1745,1770]" pageId="12" pageNumber="340">Other material examined.</emphasis>
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Population no. 8, 5ƋƋ, 1♀; population no. 9, 1♀; population no. 10, 6ƋƋ, 2♀♀; population no. 11, 2ƋƋ, 1♀; population no. 12, 5ƋƋ, 4♀♀; population no. 13, 5ƋƋ; population no. 14, 2ƋƋ, 1♀; population no. 16, 2ƋƋ, 2♀♀; population no. 17, 3ƋƋ, 3♀♀.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3226554C6471F18D394F8F1AD3DF8B5" pageId="12" pageNumber="340" type="diagnosis">
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<paragraph id="8B8736DFC6471F18D394F8F1AD3DF8B5" blockId="12.[151,1437,1475,2009]" pageId="12" pageNumber="340">
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<emphasis id="B94CEACDC6471F18D394F8F1AD52F894" bold="true" box="[199,321,1845,1870]" pageId="12" pageNumber="340">Diagnosis.</emphasis>
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<emphasis id="B94CEACDC6471F18D21AF8F2AD9FF897" box="[329,396,1846,1869]" italics="true" pageId="12" pageNumber="340">Larva</emphasis>
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(
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<figureCitation id="13032A5AC6471F18D2CFF8F1ADCCF894" box="[412,479,1845,1870]" captionStart="FIGURE 8" captionStartId="14.[151,250,1778,1800]" captionTargetBox="[263,1305,193,1723]" captionTargetId="figure@14.[263,1305,193,1725]" captionTargetPageId="14" captionText="FIGURE 8. Triops multifidus sp. nov. A, endopodite and 5 th endite of 2 nd trunk limb of the holotype (male), anterior view. B, endopodite of 2 nd trunk limb of a female of 2.3 mm telson width (from the type locality), anterior view. C, endopodite of 6 th trunk limb of a male of 2.3 mm telson width (from the type locality), anterior view. D, endopodite and 5 th endite of 2 nd trunk limb of a male of 2.2 mm telson width (from population 10), anterior view. E, second antenna of a second instar larva (from the type locality) as seen in a shed exuvium, ventral view. F, general aspect of adult male (showing a specimen of population 13). G, variation in the shape of the nuchal organ: examples (from left to right) from specimen of 2.2 (population 10) 1.8 (population 12, female) 2.4 (holotype) and 2.2 mm telson width (population 8). H, telson of the holotype, dorsal view. I, telson of a male of 2.3 mm telson width (from the type locality), lateral view." httpUri="https://zenodo.org/record/263566/files/figure.png" pageId="12" pageNumber="340">Fig. 8</figureCitation>
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E). Exopodite of the second antenna with 11–15 setae. Distal masticatory spine short (see
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<figureCitation id="13032A5AC6471F18D398F892AD1FF8B5" box="[203,268,1878,1903]" captionStart="FIGURE 8" captionStartId="14.[151,250,1778,1800]" captionTargetBox="[263,1305,193,1723]" captionTargetId="figure@14.[263,1305,193,1725]" captionTargetPageId="14" captionText="FIGURE 8. Triops multifidus sp. nov. A, endopodite and 5 th endite of 2 nd trunk limb of the holotype (male), anterior view. B, endopodite of 2 nd trunk limb of a female of 2.3 mm telson width (from the type locality), anterior view. C, endopodite of 6 th trunk limb of a male of 2.3 mm telson width (from the type locality), anterior view. D, endopodite and 5 th endite of 2 nd trunk limb of a male of 2.2 mm telson width (from population 10), anterior view. E, second antenna of a second instar larva (from the type locality) as seen in a shed exuvium, ventral view. F, general aspect of adult male (showing a specimen of population 13). G, variation in the shape of the nuchal organ: examples (from left to right) from specimen of 2.2 (population 10) 1.8 (population 12, female) 2.4 (holotype) and 2.2 mm telson width (population 8). H, telson of the holotype, dorsal view. I, telson of a male of 2.3 mm telson width (from the type locality), lateral view." httpUri="https://zenodo.org/record/263566/files/figure.png" pageId="12" pageNumber="340">Fig. 8</figureCitation>
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E).
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</paragraph>
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</subSubSection>
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||
<subSubSection id="C3226554C6471F19D394F8B3AD80F9D0" lastPageId="13" lastPageNumber="341" pageId="12" pageNumber="340" type="description">
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<paragraph id="8B8736DFC6471F18D394F8B3A98FF803" blockId="12.[151,1437,1475,2009]" pageId="12" pageNumber="340">
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||
<emphasis id="B94CEACDC6471F18D394F8B3ADC5F84A" bold="true" box="[199,470,1911,1936]" pageId="12" pageNumber="340">
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Description.
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<typeStatus id="5483887DC6471F18D231F8B3ADC2F84A" box="[354,465,1911,1936]" pageId="12" pageNumber="340" type="holotype">Holotype</typeStatus>
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.
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</emphasis>
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<emphasis id="B94CEACDC6471F18D2B3F8B3AE4FF84A" box="[480,604,1911,1936]" italics="true" pageId="12" pageNumber="340">Adult male</emphasis>
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. Body rings approx. 35.3 (i.e. 35 + 1 incomplete ring); total number of trunk limbs 44. 1st trunk limb (
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<figureCitation id="13032A5AC6471F18D2B5F858AE3AF86E" box="[486,553,1948,1973]" captionStart="FIGURE 9" captionStartId="15.[151,250,896,918]" captionTargetBox="[153,1433,195,871]" captionTargetId="figure@15.[151,1436,193,875]" captionTargetPageId="15" captionText="FIGURE 9. Triops multifidus sp. nov. A, carapace of the holotype (male), right lateral view. B, second maxilla of the holotype, anterior view. C, detail of the 1 st trunk limb of the holotype, anterior view (note that the epipodite is missing in this sample, possibly due to injury)." httpUri="https://zenodo.org/record/263567/files/figure.png" pageId="12" pageNumber="340">Fig. 9</figureCitation>
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C): distal portion of the limb corm (see Fig. S.
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<quantity id="4CC09B3AC6471F18D77BF858A846F86F" box="[1064,1109,1948,1973]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="12" pageNumber="340" unit="in" value="2.0">2 in</quantity>
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<bibRefCitation id="EFA94B2EC6471F18D708F859A900F86E" author="Korn" box="[1115,1299,1948,1973]" pageId="12" pageNumber="340" refString="Korn, M., Rabet, N., Ghate, H. V., Marrone, F. & Hundsdoerfer, A. K. (2013) Molecular phylogeny of the Notostraca. Molecular phylogenetics and evolution, 69 (3), 1159 - 1171. http: // dx. doi. org / 10.1016 / j. ympev. 2013.08.006" type="journal article" year="2013">
|
||
Korn
|
||
<emphasis id="B94CEACDC6471F18D7C8F859A8C1F86F" box="[1179,1234,1948,1973]" italics="true" pageId="12" pageNumber="340">et al.</emphasis>
|
||
2013
|
||
</bibRefCitation>
|
||
) broadened, its maximum diameter
|
||
<quantity id="4CC09B3AC6471F18D2CBF804ADEDF802" box="[408,510,1984,2008]" metricMagnitude="-3" metricUnit="m" metricValue="1.38" pageId="12" pageNumber="340" unit="mm" value="1.38">1.38 mm</quantity>
|
||
, length of exopodite
|
||
<quantity id="4CC09B3AC6471F18D1BAF804AF5FF802" box="[745,844,1984,2009]" metricMagnitude="-3" metricUnit="m" metricValue="3.59" pageId="12" pageNumber="340" unit="mm" value="3.59">3.59 mm</quantity>
|
||
from anterior to posterior corner. 2nd trunk limb (
|
||
<figureCitation id="13032A5AC6471F18D63DF804A98FF803" box="[1390,1436,1984,2009]" captionStart="FIGURE 8" captionStartId="14.[151,250,1778,1800]" captionTargetBox="[263,1305,193,1723]" captionTargetId="figure@14.[263,1305,193,1725]" captionTargetPageId="14" captionText="FIGURE 8. Triops multifidus sp. nov. A, endopodite and 5 th endite of 2 nd trunk limb of the holotype (male), anterior view. B, endopodite of 2 nd trunk limb of a female of 2.3 mm telson width (from the type locality), anterior view. C, endopodite of 6 th trunk limb of a male of 2.3 mm telson width (from the type locality), anterior view. D, endopodite and 5 th endite of 2 nd trunk limb of a male of 2.2 mm telson width (from population 10), anterior view. E, second antenna of a second instar larva (from the type locality) as seen in a shed exuvium, ventral view. F, general aspect of adult male (showing a specimen of population 13). G, variation in the shape of the nuchal organ: examples (from left to right) from specimen of 2.2 (population 10) 1.8 (population 12, female) 2.4 (holotype) and 2.2 mm telson width (population 8). H, telson of the holotype, dorsal view. I, telson of a male of 2.3 mm telson width (from the type locality), lateral view." httpUri="https://zenodo.org/record/263566/files/figure.png" pageId="12" pageNumber="340">Fig.</figureCitation>
|
||
</paragraph>
|
||
<paragraph id="8B8736DFC6461F19D3C4FF5CAEE7FE1E" blockId="13.[151,1437,151,1983]" pageId="13" pageNumber="341">
|
||
8A): length of endopodite (excluding distal claw)
|
||
<quantity id="4CC09B3AC6461F19D190FF5CAF3BFF75" box="[707,808,152,176]" metricMagnitude="-3" metricUnit="m" metricValue="3.27" pageId="13" pageNumber="341" unit="mm" value="3.27">3.27 mm</quantity>
|
||
, length of distal claw
|
||
<quantity id="4CC09B3AC6461F19D74FFF5CA892FF75" box="[1052,1153,152,176]" metricMagnitude="-4" metricUnit="m" metricValue="1.7" pageId="13" pageNumber="341" unit="mm" value="0.17">0.17 mm</quantity>
|
||
, length of largest digging spine
|
||
<quantity id="4CC09B3AC6461F19D38FFF78AD56FF0E" box="[220,325,188,212]" metricMagnitude="-5" metricUnit="m" metricValue="9.0" pageId="13" pageNumber="341" unit="mm" value="0.09">0.09 mm</quantity>
|
||
, length of 5th endite
|
||
<quantity id="4CC09B3AC6461F19D164FF78AEB1FF0E" box="[567,674,188,213]" metricMagnitude="-3" metricUnit="m" metricValue="3.76" pageId="13" pageNumber="341" unit="mm" value="3.76">3.76 mm</quantity>
|
||
. 10th trunk limb: 6 anterior meshwork spines on 3rd endite, 8 submarginal spines on 4th endite. Carapace (
|
||
<figureCitation id="13032A5AC6461F19D132FF24AEB6FF22" box="[609,677,224,249]" captionStart="FIGURE 9" captionStartId="15.[151,250,896,918]" captionTargetBox="[153,1433,195,871]" captionTargetId="figure@15.[151,1436,193,875]" captionTargetPageId="15" captionText="FIGURE 9. Triops multifidus sp. nov. A, carapace of the holotype (male), right lateral view. B, second maxilla of the holotype, anterior view. C, detail of the 1 st trunk limb of the holotype, anterior view (note that the epipodite is missing in this sample, possibly due to injury)." httpUri="https://zenodo.org/record/263567/files/figure.png" pageId="13" pageNumber="341">Fig. 9</figureCitation>
|
||
A): length of carapace
|
||
<quantity id="4CC09B3AC6461F19D0FBFF24A808FF22" box="[936,1051,224,249]" metricMagnitude="-2" metricUnit="m" metricValue="1.6019999999999999" pageId="13" pageNumber="341" unit="mm" value="16.02">16.02 mm</quantity>
|
||
(measured along its median line), no dorsal carina spines, sulcus with 38 spines, length of nuchal organ
|
||
<quantity id="4CC09B3AC6461F19D0E9FEC1A832FEC6" box="[954,1057,261,285]" metricMagnitude="-4" metricUnit="m" metricValue="8.9" pageId="13" pageNumber="341" unit="mm" value="0.89">0.89 mm</quantity>
|
||
, mystax present. 2nd maxilla well developed (
|
||
<figureCitation id="13032A5AC6461F19D248FEE1AD72FEE4" box="[283,353,293,318]" captionStart="FIGURE 9" captionStartId="15.[151,250,896,918]" captionTargetBox="[153,1433,195,871]" captionTargetId="figure@15.[151,1436,193,875]" captionTargetPageId="15" captionText="FIGURE 9. Triops multifidus sp. nov. A, carapace of the holotype (male), right lateral view. B, second maxilla of the holotype, anterior view. C, detail of the 1 st trunk limb of the holotype, anterior view (note that the epipodite is missing in this sample, possibly due to injury)." httpUri="https://zenodo.org/record/263567/files/figure.png" pageId="13" pageNumber="341">Fig. 9</figureCitation>
|
||
B). Telson (
|
||
<figureCitation id="13032A5AC6461F19D2BBFEE1AE3FFEE4" box="[488,556,293,318]" captionStart="FIGURE 8" captionStartId="14.[151,250,1778,1800]" captionTargetBox="[263,1305,193,1723]" captionTargetId="figure@14.[263,1305,193,1725]" captionTargetPageId="14" captionText="FIGURE 8. Triops multifidus sp. nov. A, endopodite and 5 th endite of 2 nd trunk limb of the holotype (male), anterior view. B, endopodite of 2 nd trunk limb of a female of 2.3 mm telson width (from the type locality), anterior view. C, endopodite of 6 th trunk limb of a male of 2.3 mm telson width (from the type locality), anterior view. D, endopodite and 5 th endite of 2 nd trunk limb of a male of 2.2 mm telson width (from population 10), anterior view. E, second antenna of a second instar larva (from the type locality) as seen in a shed exuvium, ventral view. F, general aspect of adult male (showing a specimen of population 13). G, variation in the shape of the nuchal organ: examples (from left to right) from specimen of 2.2 (population 10) 1.8 (population 12, female) 2.4 (holotype) and 2.2 mm telson width (population 8). H, telson of the holotype, dorsal view. I, telson of a male of 2.3 mm telson width (from the type locality), lateral view." httpUri="https://zenodo.org/record/263566/files/figure.png" pageId="13" pageNumber="341">Fig. 8</figureCitation>
|
||
H): telson width
|
||
<quantity id="4CC09B3AC6461F19D1BCFEE2AF4BFEE7" box="[751,856,294,318]" metricMagnitude="-3" metricUnit="m" metricValue="2.38" pageId="13" pageNumber="341" unit="mm" value="2.38">2.38 mm</quantity>
|
||
, length of largest furcal spines on each side of the telson 0.14 and
|
||
<quantity id="4CC09B3AC6461F19D21DFE82ADA5FE85" box="[334,438,326,351]" metricMagnitude="-4" metricUnit="m" metricValue="1.6" pageId="13" pageNumber="341" unit="mm" value="0.16">0.16 mm</quantity>
|
||
, 7 posterior marginal spines, i.e. 5 additional secondary posterior marginals developed, posterior marginal spines small, 4 median spines (1 of these very small), one additional spine approx. half-way between median and setal spines. Apodous abdominal segments approx. 10.4 (10.1 + 1 incomplete segment); numerous supernumerary spines, broad, scale-shaped.
|
||
</paragraph>
|
||
<paragraph id="8B8736DFC6461F19D394FE0BAEE2FCDD" blockId="13.[151,1437,151,1983]" pageId="13" pageNumber="341">
|
||
<emphasis id="B94CEACDC6461F19D394FE0BAE07FE32" bold="true" box="[199,532,463,488]" pageId="13" pageNumber="341">Adult (and sub-adult) male</emphasis>
|
||
(
|
||
<figureCitation id="13032A5AC6461F19D176FE0BAE79FE3D" box="[549,618,463,488]" captionStart="FIGURE 8" captionStartId="14.[151,250,1778,1800]" captionTargetBox="[263,1305,193,1723]" captionTargetId="figure@14.[263,1305,193,1725]" captionTargetPageId="14" captionText="FIGURE 8. Triops multifidus sp. nov. A, endopodite and 5 th endite of 2 nd trunk limb of the holotype (male), anterior view. B, endopodite of 2 nd trunk limb of a female of 2.3 mm telson width (from the type locality), anterior view. C, endopodite of 6 th trunk limb of a male of 2.3 mm telson width (from the type locality), anterior view. D, endopodite and 5 th endite of 2 nd trunk limb of a male of 2.2 mm telson width (from population 10), anterior view. E, second antenna of a second instar larva (from the type locality) as seen in a shed exuvium, ventral view. F, general aspect of adult male (showing a specimen of population 13). G, variation in the shape of the nuchal organ: examples (from left to right) from specimen of 2.2 (population 10) 1.8 (population 12, female) 2.4 (holotype) and 2.2 mm telson width (population 8). H, telson of the holotype, dorsal view. I, telson of a male of 2.3 mm telson width (from the type locality), lateral view." httpUri="https://zenodo.org/record/263566/files/figure.png" pageId="13" pageNumber="341">Fig. 8</figureCitation>
|
||
A, C, D, F–I, 9).
|
||
<emphasis id="B94CEACDC6461F19D06BFE0BAFC5FE32" box="[824,982,459,488]" italics="true" pageId="13" pageNumber="341">2nd trunk limb</emphasis>
|
||
(
|
||
<figureCitation id="13032A5AC6461F19D0BBFE0BA83FFE3D" box="[1000,1068,463,488]" captionStart="FIGURE 8" captionStartId="14.[151,250,1778,1800]" captionTargetBox="[263,1305,193,1723]" captionTargetId="figure@14.[263,1305,193,1725]" captionTargetPageId="14" captionText="FIGURE 8. Triops multifidus sp. nov. A, endopodite and 5 th endite of 2 nd trunk limb of the holotype (male), anterior view. B, endopodite of 2 nd trunk limb of a female of 2.3 mm telson width (from the type locality), anterior view. C, endopodite of 6 th trunk limb of a male of 2.3 mm telson width (from the type locality), anterior view. D, endopodite and 5 th endite of 2 nd trunk limb of a male of 2.2 mm telson width (from population 10), anterior view. E, second antenna of a second instar larva (from the type locality) as seen in a shed exuvium, ventral view. F, general aspect of adult male (showing a specimen of population 13). G, variation in the shape of the nuchal organ: examples (from left to right) from specimen of 2.2 (population 10) 1.8 (population 12, female) 2.4 (holotype) and 2.2 mm telson width (population 8). H, telson of the holotype, dorsal view. I, telson of a male of 2.3 mm telson width (from the type locality), lateral view." httpUri="https://zenodo.org/record/263566/files/figure.png" pageId="13" pageNumber="341">Fig. 8</figureCitation>
|
||
A, D). ‘Proportional endopodite length’ 80.2–96.5% (mean=89.5%; SD=3.4; n=31); row of digging spines on endopodite of rather homogeneous appearance (see
|
||
<figureCitation id="13032A5AC6461F19D20AFDD3AD8EFDF5" box="[345,413,535,560]" captionStart="FIGURE 8" captionStartId="14.[151,250,1778,1800]" captionTargetBox="[263,1305,193,1723]" captionTargetId="figure@14.[263,1305,193,1725]" captionTargetPageId="14" captionText="FIGURE 8. Triops multifidus sp. nov. A, endopodite and 5 th endite of 2 nd trunk limb of the holotype (male), anterior view. B, endopodite of 2 nd trunk limb of a female of 2.3 mm telson width (from the type locality), anterior view. C, endopodite of 6 th trunk limb of a male of 2.3 mm telson width (from the type locality), anterior view. D, endopodite and 5 th endite of 2 nd trunk limb of a male of 2.2 mm telson width (from population 10), anterior view. E, second antenna of a second instar larva (from the type locality) as seen in a shed exuvium, ventral view. F, general aspect of adult male (showing a specimen of population 13). G, variation in the shape of the nuchal organ: examples (from left to right) from specimen of 2.2 (population 10) 1.8 (population 12, female) 2.4 (holotype) and 2.2 mm telson width (population 8). H, telson of the holotype, dorsal view. I, telson of a male of 2.3 mm telson width (from the type locality), lateral view." httpUri="https://zenodo.org/record/263566/files/figure.png" pageId="13" pageNumber="341">Fig. 8</figureCitation>
|
||
A, D); ‘proportional length of digging spines’ on endopodite 3.52–4.76% (standardised values; mean=4.36%; SD=0.26; n=31); absolute size of largest digging spines on endopodite 54–115 µm (range of telson width in studied specimens 1.2–4.0 mm); distal claw of endopodite 2–6% the length of the endopodite (excluding the distal claw), stronger curved than is usual in the majority of specimens with its tip bent proximally giving a hook-shaped appearance (in 71% of the
|
||
<specimenCount id="9D3EFD56C6461F19D184FD63AF61FD1A" box="[727,882,679,704]" pageId="13" pageNumber="341" type="generic">31 specimens</specimenCount>
|
||
studied; this character expression was found in specimens of all 8 populations studied morphologically), distal claw thereby often reaching a position of approx. a right angle to the row of digging spines (see
|
||
<figureCitation id="13032A5AC6461F19D1D8FD2BAEDFFCDD" box="[651,716,751,776]" captionStart="FIGURE 8" captionStartId="14.[151,250,1778,1800]" captionTargetBox="[263,1305,193,1723]" captionTargetId="figure@14.[263,1305,193,1725]" captionTargetPageId="14" captionText="FIGURE 8. Triops multifidus sp. nov. A, endopodite and 5 th endite of 2 nd trunk limb of the holotype (male), anterior view. B, endopodite of 2 nd trunk limb of a female of 2.3 mm telson width (from the type locality), anterior view. C, endopodite of 6 th trunk limb of a male of 2.3 mm telson width (from the type locality), anterior view. D, endopodite and 5 th endite of 2 nd trunk limb of a male of 2.2 mm telson width (from population 10), anterior view. E, second antenna of a second instar larva (from the type locality) as seen in a shed exuvium, ventral view. F, general aspect of adult male (showing a specimen of population 13). G, variation in the shape of the nuchal organ: examples (from left to right) from specimen of 2.2 (population 10) 1.8 (population 12, female) 2.4 (holotype) and 2.2 mm telson width (population 8). H, telson of the holotype, dorsal view. I, telson of a male of 2.3 mm telson width (from the type locality), lateral view." httpUri="https://zenodo.org/record/263566/files/figure.png" pageId="13" pageNumber="341">Fig. 8</figureCitation>
|
||
D).
|
||
</paragraph>
|
||
<paragraph id="8B8736DFC6461F19D394FCD2AF8DFC89" blockId="13.[151,1437,151,1983]" pageId="13" pageNumber="341">
|
||
<emphasis id="B94CEACDC6461F19D394FCD2AD50FCF4" box="[199,323,786,814]" italics="true" pageId="13" pageNumber="341">1st maxilla.</emphasis>
|
||
The row of spines not confined to the centrally directed face but extending to a part of the ventral margin; spines on the ventral margin typically bent postero-ventrally.
|
||
</paragraph>
|
||
<paragraph id="8B8736DFC6461F19D39DFCA5AE7EFCA3" blockId="13.[151,1437,151,1983]" box="[206,621,861,889]" pageId="13" pageNumber="341">
|
||
<emphasis id="B94CEACDC6461F19D39DFCA5AD55FCA3" box="[206,326,861,889]" italics="true" pageId="13" pageNumber="341">2nd maxilla</emphasis>
|
||
(
|
||
<figureCitation id="13032A5AC6461F19D206FCA4AD8BFCA3" box="[341,408,864,889]" captionStart="FIGURE 9" captionStartId="15.[151,250,896,918]" captionTargetBox="[153,1433,195,871]" captionTargetId="figure@15.[151,1436,193,875]" captionTargetPageId="15" captionText="FIGURE 9. Triops multifidus sp. nov. A, carapace of the holotype (male), right lateral view. B, second maxilla of the holotype, anterior view. C, detail of the 1 st trunk limb of the holotype, anterior view (note that the epipodite is missing in this sample, possibly due to injury)." httpUri="https://zenodo.org/record/263567/files/figure.png" pageId="13" pageNumber="341">Fig. 9</figureCitation>
|
||
B) well developed.
|
||
</paragraph>
|
||
<paragraph id="8B8736DFC6461F19D394FC41AD54FBD2" blockId="13.[151,1437,151,1983]" pageId="13" pageNumber="341">
|
||
<emphasis id="B94CEACDC6461F19D394FC41AD26FC46" box="[199,309,901,924]" italics="true" pageId="13" pageNumber="341">Carapace</emphasis>
|
||
. Shape of nuchal organ as shown in
|
||
<figureCitation id="13032A5AC6461F19D19FFC40AF1FFC47" box="[716,780,900,925]" captionStart="FIGURE 8" captionStartId="14.[151,250,1778,1800]" captionTargetBox="[263,1305,193,1723]" captionTargetId="figure@14.[263,1305,193,1725]" captionTargetPageId="14" captionText="FIGURE 8. Triops multifidus sp. nov. A, endopodite and 5 th endite of 2 nd trunk limb of the holotype (male), anterior view. B, endopodite of 2 nd trunk limb of a female of 2.3 mm telson width (from the type locality), anterior view. C, endopodite of 6 th trunk limb of a male of 2.3 mm telson width (from the type locality), anterior view. D, endopodite and 5 th endite of 2 nd trunk limb of a male of 2.2 mm telson width (from population 10), anterior view. E, second antenna of a second instar larva (from the type locality) as seen in a shed exuvium, ventral view. F, general aspect of adult male (showing a specimen of population 13). G, variation in the shape of the nuchal organ: examples (from left to right) from specimen of 2.2 (population 10) 1.8 (population 12, female) 2.4 (holotype) and 2.2 mm telson width (population 8). H, telson of the holotype, dorsal view. I, telson of a male of 2.3 mm telson width (from the type locality), lateral view." httpUri="https://zenodo.org/record/263566/files/figure.png" pageId="13" pageNumber="341">Fig. 8</figureCitation>
|
||
G; dorsal carina usually smooth (
|
||
<figureCitation id="13032A5AC6461F19D72EFC40A8ACFC47" box="[1149,1215,900,925]" captionStart="FIGURE 9" captionStartId="15.[151,250,896,918]" captionTargetBox="[153,1433,195,871]" captionTargetId="figure@15.[151,1436,193,875]" captionTargetPageId="15" captionText="FIGURE 9. Triops multifidus sp. nov. A, carapace of the holotype (male), right lateral view. B, second maxilla of the holotype, anterior view. C, detail of the 1 st trunk limb of the holotype, anterior view (note that the epipodite is missing in this sample, possibly due to injury)." httpUri="https://zenodo.org/record/263567/files/figure.png" pageId="13" pageNumber="341">Fig. 9</figureCitation>
|
||
A), but a few dorsal carina spines may be present (1 to 2 recorded); terminal carina spine small, sometimes completely reduced; sulcus spines highly variable in size, from very prominent strongly chitinised structures to tiny, almost reduced bulges; mystax present.
|
||
</paragraph>
|
||
<paragraph id="8B8736DFC6461F19D39DFBD0A901FBF7" blockId="13.[151,1437,151,1983]" box="[206,1298,1044,1069]" pageId="13" pageNumber="341">
|
||
<emphasis id="B94CEACDC6461F19D39DFBD0AD07FBF7" box="[206,276,1044,1069]" italics="true" pageId="13" pageNumber="341">Telson</emphasis>
|
||
(
|
||
<figureCitation id="13032A5AC6461F19D276FBD0AD74FBF7" box="[293,359,1044,1069]" captionStart="FIGURE 8" captionStartId="14.[151,250,1778,1800]" captionTargetBox="[263,1305,193,1723]" captionTargetId="figure@14.[263,1305,193,1725]" captionTargetPageId="14" captionText="FIGURE 8. Triops multifidus sp. nov. A, endopodite and 5 th endite of 2 nd trunk limb of the holotype (male), anterior view. B, endopodite of 2 nd trunk limb of a female of 2.3 mm telson width (from the type locality), anterior view. C, endopodite of 6 th trunk limb of a male of 2.3 mm telson width (from the type locality), anterior view. D, endopodite and 5 th endite of 2 nd trunk limb of a male of 2.2 mm telson width (from population 10), anterior view. E, second antenna of a second instar larva (from the type locality) as seen in a shed exuvium, ventral view. F, general aspect of adult male (showing a specimen of population 13). G, variation in the shape of the nuchal organ: examples (from left to right) from specimen of 2.2 (population 10) 1.8 (population 12, female) 2.4 (holotype) and 2.2 mm telson width (population 8). H, telson of the holotype, dorsal view. I, telson of a male of 2.3 mm telson width (from the type locality), lateral view." httpUri="https://zenodo.org/record/263566/files/figure.png" pageId="13" pageNumber="341">Fig. 8</figureCitation>
|
||
H, I). ‘Proportional length of furcal spines’ 2.0–6.3% (mean=3.7%; SD=0.9; n=31).
|
||
</paragraph>
|
||
<paragraph id="8B8736DFC6461F19D39DFBFCAF01FB8B" blockId="13.[151,1437,151,1983]" box="[206,786,1080,1105]" pageId="13" pageNumber="341">
|
||
<emphasis id="B94CEACDC6461F19D39DFBFCAD54FB8B" box="[206,327,1080,1105]" italics="true" pageId="13" pageNumber="341">Body rings</emphasis>
|
||
. 33.0–37.0 (mean=34.9; SD=0.9; n=31).
|
||
</paragraph>
|
||
<paragraph id="8B8736DFC6461F19D394FB98AD80FB43" blockId="13.[151,1437,151,1983]" pageId="13" pageNumber="341">
|
||
<emphasis id="B94CEACDC6461F19D394FB98AE37FBAF" box="[199,548,1116,1141]" italics="true" pageId="13" pageNumber="341">Apodous abdominal segments</emphasis>
|
||
9.9–12.5 (mean=10.8; SD=0.6; n=31), with medium to high numbers of supernumerary spines.
|
||
</paragraph>
|
||
<paragraph id="8B8736DFC6461F19D394FB63AF4FFAD2" blockId="13.[151,1437,151,1983]" pageId="13" pageNumber="341">
|
||
<emphasis id="B94CEACDC6461F19D394FB63AE35FB1A" bold="true" box="[199,550,1191,1216]" pageId="13" pageNumber="341">Adult (and sub-adult) female</emphasis>
|
||
:
|
||
<emphasis id="B94CEACDC6461F19D164FB6CAEC0FB1A" box="[567,723,1188,1216]" italics="true" pageId="13" pageNumber="341">2nd trunk limb</emphasis>
|
||
(
|
||
<figureCitation id="13032A5AC6461F19D1B7FB63AF3BFB1A" box="[740,808,1191,1216]" captionStart="FIGURE 8" captionStartId="14.[151,250,1778,1800]" captionTargetBox="[263,1305,193,1723]" captionTargetId="figure@14.[263,1305,193,1725]" captionTargetPageId="14" captionText="FIGURE 8. Triops multifidus sp. nov. A, endopodite and 5 th endite of 2 nd trunk limb of the holotype (male), anterior view. B, endopodite of 2 nd trunk limb of a female of 2.3 mm telson width (from the type locality), anterior view. C, endopodite of 6 th trunk limb of a male of 2.3 mm telson width (from the type locality), anterior view. D, endopodite and 5 th endite of 2 nd trunk limb of a male of 2.2 mm telson width (from population 10), anterior view. E, second antenna of a second instar larva (from the type locality) as seen in a shed exuvium, ventral view. F, general aspect of adult male (showing a specimen of population 13). G, variation in the shape of the nuchal organ: examples (from left to right) from specimen of 2.2 (population 10) 1.8 (population 12, female) 2.4 (holotype) and 2.2 mm telson width (population 8). H, telson of the holotype, dorsal view. I, telson of a male of 2.3 mm telson width (from the type locality), lateral view." httpUri="https://zenodo.org/record/263566/files/figure.png" pageId="13" pageNumber="341">Fig. 8</figureCitation>
|
||
B). Digging spines on endopodite show a more regular pattern of size distribution than in preceding species, with large sturdy spines being separated by only few smaller spines, density of spines usually higher towards the distal claw.
|
||
</paragraph>
|
||
<paragraph id="8B8736DFC6461F19D394FAD3ADD6FAF5" blockId="13.[151,1437,151,1983]" box="[199,453,1298,1327]" pageId="13" pageNumber="341">
|
||
<emphasis id="B94CEACDC6461F19D394FAD3AD29FAF5" box="[199,314,1298,1327]" italics="true" pageId="13" pageNumber="341">1st maxilla</emphasis>
|
||
as in males.
|
||
</paragraph>
|
||
<paragraph id="8B8736DFC6461F19D394FAFAADEBFA8C" blockId="13.[151,1437,151,1983]" box="[199,504,1337,1366]" pageId="13" pageNumber="341">
|
||
<emphasis id="B94CEACDC6461F19D394FAFAAD2CFA8C" box="[199,319,1337,1366]" italics="true" pageId="13" pageNumber="341">2nd maxilla</emphasis>
|
||
well developed.
|
||
</paragraph>
|
||
<paragraph id="8B8736DFC6461F19D394FAA6AD19FA44" blockId="13.[151,1437,151,1983]" pageId="13" pageNumber="341">
|
||
<emphasis id="B94CEACDC6461F19D394FAA6AD26FAA3" box="[199,309,1378,1401]" italics="true" pageId="13" pageNumber="341">Carapace</emphasis>
|
||
. Nuchal organ as shown in
|
||
<figureCitation id="13032A5AC6461F19D127FAA5AEA4FAA3" box="[628,695,1377,1402]" captionStart="FIGURE 8" captionStartId="14.[151,250,1778,1800]" captionTargetBox="[263,1305,193,1723]" captionTargetId="figure@14.[263,1305,193,1725]" captionTargetPageId="14" captionText="FIGURE 8. Triops multifidus sp. nov. A, endopodite and 5 th endite of 2 nd trunk limb of the holotype (male), anterior view. B, endopodite of 2 nd trunk limb of a female of 2.3 mm telson width (from the type locality), anterior view. C, endopodite of 6 th trunk limb of a male of 2.3 mm telson width (from the type locality), anterior view. D, endopodite and 5 th endite of 2 nd trunk limb of a male of 2.2 mm telson width (from population 10), anterior view. E, second antenna of a second instar larva (from the type locality) as seen in a shed exuvium, ventral view. F, general aspect of adult male (showing a specimen of population 13). G, variation in the shape of the nuchal organ: examples (from left to right) from specimen of 2.2 (population 10) 1.8 (population 12, female) 2.4 (holotype) and 2.2 mm telson width (population 8). H, telson of the holotype, dorsal view. I, telson of a male of 2.3 mm telson width (from the type locality), lateral view." httpUri="https://zenodo.org/record/263566/files/figure.png" pageId="13" pageNumber="341">Fig. 8</figureCitation>
|
||
G; dorsal carina smooth or with few dorsal carina spines (up to 3 recorded).
|
||
</paragraph>
|
||
<paragraph id="8B8736DFC6461F19D394FA6DAF18FA18" blockId="13.[151,1437,151,1983]" box="[199,779,1449,1474]" pageId="13" pageNumber="341">
|
||
<emphasis id="B94CEACDC6461F19D394FA6DAD52FA1B" box="[199,321,1449,1474]" italics="true" pageId="13" pageNumber="341">Body rings</emphasis>
|
||
. 33.9–35.6 (mean=34.6; SD=0.5; n=15).
|
||
</paragraph>
|
||
<paragraph id="8B8736DFC6461F19D394FA09AD80F9D0" blockId="13.[151,1437,151,1983]" pageId="13" pageNumber="341">
|
||
<emphasis id="B94CEACDC6461F19D394FA09AE38FA3F" box="[199,555,1485,1510]" italics="true" pageId="13" pageNumber="341">Apodous abdominal segments</emphasis>
|
||
7.2–9.1 (mean=8.3; SD=0.6; n=15) with medium to high numbers of supernumerary spines.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3226554C6461F1BD394F9D1AF82FBE5" lastPageId="15" lastPageNumber="343" pageId="13" pageNumber="341" type="diagnosis">
|
||
<paragraph id="8B8736DFC6461F19D394F9D1AF45F865" blockId="13.[151,1437,151,1983]" pageId="13" pageNumber="341">
|
||
<emphasis id="B94CEACDC6461F19D394F9D1ADC3F9F4" bold="true" box="[199,464,1557,1582]" pageId="13" pageNumber="341">Differential diagnosis.</emphasis>
|
||
The species is morphologically well differentiated from remaining species of
|
||
<taxonomicName id="4C384D5CC6461F19D607F9D2A989F9F7" box="[1364,1434,1558,1581]" class="Branchiopoda" family="Triopsidae" genus="Triops" kingdom="Animalia" order="Notostraca" pageId="13" pageNumber="341" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B94CEACDC6461F19D607F9D2A989F9F7" box="[1364,1434,1558,1581]" italics="true" pageId="13" pageNumber="341">Triops</emphasis>
|
||
</taxonomicName>
|
||
(except
|
||
<taxonomicName id="4C384D5CC6461F19D3BCF9FEADB2F98B" box="[239,417,1594,1617]" class="Branchiopoda" family="Triopsidae" genus="Triops" kingdom="Animalia" order="Notostraca" pageId="13" pageNumber="341" phylum="Arthropoda" rank="species" species="maximus" status="sp. nov.">
|
||
<emphasis id="B94CEACDC6461F19D3BCF9FEADB2F98B" box="[239,417,1594,1617]" italics="true" pageId="13" pageNumber="341">Triops maximus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27F57B6C6461F19D2F4F9FDAE13F988" box="[423,512,1593,1618]" pageId="13" pageNumber="341" rank="species">
|
||
<emphasis id="B94CEACDC6461F19D2F4F9FDAE13F988" bold="true" box="[423,512,1593,1618]" pageId="13" pageNumber="341">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
) by a strongly increased number of setae (11–15 instead of the hitherto reported 5– 7) on the exopodite of the second antenna in early larval stages. In adult females the largest digging spines in the 2nd trunk limb (usually located in the proximal half of the row of digging spines) are larger and/or broader than in females of other species (except
|
||
<taxonomicName id="4C384D5CC6461F19D146F96DAED8F91A" box="[533,715,1705,1728]" class="Branchiopoda" family="Triopsidae" genus="Triops" kingdom="Animalia" order="Notostraca" pageId="13" pageNumber="341" phylum="Arthropoda" rank="species" species="maximus" status="sp. nov.">
|
||
<emphasis id="B94CEACDC6461F19D146F96DAED8F91A" box="[533,715,1705,1728]" italics="true" pageId="13" pageNumber="341">Triops maximus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27F57B6C6461F19D187F963AF22F91A" box="[724,817,1703,1728]" pageId="13" pageNumber="341" rank="species">
|
||
<emphasis id="B94CEACDC6461F19D187F963AF22F91A" bold="true" box="[724,817,1703,1728]" pageId="13" pageNumber="341">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
) and digging spines show a lower density within the proximal half of the row of spines (
|
||
<figureCitation id="13032A5AC6461F19D177F90AAE74F93D" box="[548,615,1742,1767]" captionStart="FIGURE 8" captionStartId="14.[151,250,1778,1800]" captionTargetBox="[263,1305,193,1723]" captionTargetId="figure@14.[263,1305,193,1725]" captionTargetPageId="14" captionText="FIGURE 8. Triops multifidus sp. nov. A, endopodite and 5 th endite of 2 nd trunk limb of the holotype (male), anterior view. B, endopodite of 2 nd trunk limb of a female of 2.3 mm telson width (from the type locality), anterior view. C, endopodite of 6 th trunk limb of a male of 2.3 mm telson width (from the type locality), anterior view. D, endopodite and 5 th endite of 2 nd trunk limb of a male of 2.2 mm telson width (from population 10), anterior view. E, second antenna of a second instar larva (from the type locality) as seen in a shed exuvium, ventral view. F, general aspect of adult male (showing a specimen of population 13). G, variation in the shape of the nuchal organ: examples (from left to right) from specimen of 2.2 (population 10) 1.8 (population 12, female) 2.4 (holotype) and 2.2 mm telson width (population 8). H, telson of the holotype, dorsal view. I, telson of a male of 2.3 mm telson width (from the type locality), lateral view." httpUri="https://zenodo.org/record/263566/files/figure.png" pageId="13" pageNumber="341">Fig. 8</figureCitation>
|
||
B). In adult males, the 2nd trunk limbs show comparatively high values of ‘proportional endopodite length’ and, in the majority of specimens, a hook-shaped distal claw in the endopodite (though it should be noted that a similar morphology was also observed in '
|
||
<taxonomicName id="4C384D5CC6461F19D0B0F8DCA87DF8F5" box="[995,1134,1816,1839]" class="Branchiopoda" family="Triopsidae" genus="Triops" kingdom="Animalia" order="Notostraca" pageId="13" pageNumber="341" phylum="Arthropoda" rank="species" species="granarius">
|
||
<emphasis id="B94CEACDC6461F19D0B0F8DCA87DF8F5" box="[995,1134,1816,1839]" italics="true" pageId="13" pageNumber="341">T. granarius</emphasis>
|
||
</taxonomicName>
|
||
1 and 5’, M.K. pers. obs.). The largest digging spines on the margin of the endopodite are relatively smaller than in
|
||
<taxonomicName id="4C384D5CC6461F19D7D5F8F8A929F889" box="[1158,1338,1852,1875]" class="Branchiopoda" family="Triopsidae" genus="Triops" kingdom="Animalia" order="Notostraca" pageId="13" pageNumber="341" phylum="Arthropoda" rank="species" species="maximus" status="sp. nov.">
|
||
<emphasis id="B94CEACDC6461F19D7D5F8F8A929F889" box="[1158,1338,1852,1875]" italics="true" pageId="13" pageNumber="341">Triops maximus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27F57B6C6461F19D612F8FEA98FF889" box="[1345,1436,1850,1875]" pageId="13" pageNumber="341" rank="species">
|
||
<emphasis id="B94CEACDC6461F19D612F8FEA98FF889" bold="true" box="[1345,1436,1850,1875]" pageId="13" pageNumber="341">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
(see Fig. 4C, D;
|
||
<figureCitation id="13032A5AC6461F19D201F89AAD80F8AD" box="[338,403,1886,1911]" captionStart="FIGURE 8" captionStartId="14.[151,250,1778,1800]" captionTargetBox="[263,1305,193,1723]" captionTargetId="figure@14.[263,1305,193,1725]" captionTargetPageId="14" captionText="FIGURE 8. Triops multifidus sp. nov. A, endopodite and 5 th endite of 2 nd trunk limb of the holotype (male), anterior view. B, endopodite of 2 nd trunk limb of a female of 2.3 mm telson width (from the type locality), anterior view. C, endopodite of 6 th trunk limb of a male of 2.3 mm telson width (from the type locality), anterior view. D, endopodite and 5 th endite of 2 nd trunk limb of a male of 2.2 mm telson width (from population 10), anterior view. E, second antenna of a second instar larva (from the type locality) as seen in a shed exuvium, ventral view. F, general aspect of adult male (showing a specimen of population 13). G, variation in the shape of the nuchal organ: examples (from left to right) from specimen of 2.2 (population 10) 1.8 (population 12, female) 2.4 (holotype) and 2.2 mm telson width (population 8). H, telson of the holotype, dorsal view. I, telson of a male of 2.3 mm telson width (from the type locality), lateral view." httpUri="https://zenodo.org/record/263566/files/figure.png" pageId="13" pageNumber="341">Fig. 8</figureCitation>
|
||
A, D). The species can be distinguished from
|
||
<taxonomicName id="4C384D5CC6461F19D0CAF8A4A845F8AD" box="[921,1110,1888,1911]" class="Branchiopoda" family="Triopsidae" genus="Triops" kingdom="Animalia" order="Notostraca" pageId="13" pageNumber="341" phylum="Arthropoda" rank="species" sensu="stricto" species="granarius">
|
||
<emphasis id="B94CEACDC6461F19D0CAF8A4A845F8AD" box="[921,1110,1888,1911]" italics="true" pageId="13" pageNumber="341">Triops granarius</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27F57B6C6461F19D70DF8A4A891F8AD" box="[1118,1154,1888,1911]" pageId="13" pageNumber="341" sensu="stricto">s.s.</taxonomicNameLabel>
|
||
by the slender elongated shape of the endopodites (with pointed tips) in mid-thoracic appendages (see
|
||
<figureCitation id="13032A5AC6461F19D747F846A84BF841" box="[1044,1112,1922,1947]" captionStart="FIGURE 8" captionStartId="14.[151,250,1778,1800]" captionTargetBox="[263,1305,193,1723]" captionTargetId="figure@14.[263,1305,193,1725]" captionTargetPageId="14" captionText="FIGURE 8. Triops multifidus sp. nov. A, endopodite and 5 th endite of 2 nd trunk limb of the holotype (male), anterior view. B, endopodite of 2 nd trunk limb of a female of 2.3 mm telson width (from the type locality), anterior view. C, endopodite of 6 th trunk limb of a male of 2.3 mm telson width (from the type locality), anterior view. D, endopodite and 5 th endite of 2 nd trunk limb of a male of 2.2 mm telson width (from population 10), anterior view. E, second antenna of a second instar larva (from the type locality) as seen in a shed exuvium, ventral view. F, general aspect of adult male (showing a specimen of population 13). G, variation in the shape of the nuchal organ: examples (from left to right) from specimen of 2.2 (population 10) 1.8 (population 12, female) 2.4 (holotype) and 2.2 mm telson width (population 8). H, telson of the holotype, dorsal view. I, telson of a male of 2.3 mm telson width (from the type locality), lateral view." httpUri="https://zenodo.org/record/263566/files/figure.png" pageId="13" pageNumber="341">Fig. 8</figureCitation>
|
||
C), these are indicated to be more rounded in
|
||
<taxonomicName id="4C384D5CC6461F19D204F86CAE00F865" box="[343,531,1960,1983]" class="Branchiopoda" family="Triopsidae" genus="Triops" kingdom="Animalia" order="Notostraca" pageId="13" pageNumber="341" phylum="Arthropoda" rank="species" sensu="stricto" species="granarius">
|
||
<emphasis id="B94CEACDC6461F19D204F86CAE00F865" box="[343,531,1960,1983]" italics="true" pageId="13" pageNumber="341">Triops granarius</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27F57B6C6461F19D148F86CAE2CF865" box="[539,575,1960,1983]" pageId="13" pageNumber="341" sensu="stricto">s.s.</taxonomicNameLabel>
|
||
(see discussion section).
|
||
</paragraph>
|
||
<caption id="DF476657C6451F1AD3C4F936AEF1F83E" httpUri="https://zenodo.org/record/263566/files/figure.png" pageId="14" pageNumber="342" targetBox="[263,1305,193,1723]" targetPageId="14">
|
||
<paragraph id="8B8736DFC6451F1AD3C4F936AEF1F83E" blockId="14.[151,1436,1775,2020]" pageId="14" pageNumber="342">
|
||
<emphasis id="B94CEACDC6451F1AD3C4F936AD00F8DD" bold="true" box="[151,275,1778,1800]" pageId="14" pageNumber="342">FIGURE 8.</emphasis>
|
||
<taxonomicName id="4C384D5CC6451F1AD248F937ADDBF8D2" box="[283,456,1778,1800]" class="Branchiopoda" family="Triopsidae" genus="Triops" kingdom="Animalia" order="Notostraca" pageId="14" pageNumber="342" phylum="Arthropoda" rank="species" species="multifidus" status="sp. nov.">
|
||
<emphasis id="B94CEACDC6451F1AD248F937ADDBF8D2" box="[283,456,1778,1800]" italics="true" pageId="14" pageNumber="342">Triops multifidus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27F57B6C6451F1AD29CF937AE33F8D2" box="[463,544,1779,1800]" pageId="14" pageNumber="342" rank="species">
|
||
<emphasis id="B94CEACDC6451F1AD29CF937AE33F8D2" bold="true" box="[463,544,1779,1800]" pageId="14" pageNumber="342">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
A, endopodite and 5th endite of 2nd trunk limb of the holotype (male), anterior view. B, endopodite of 2nd trunk limb of a female of 2.3 mm telson width (from the type locality), anterior view. C, endopodite of 6th trunk limb of a male of 2.3 mm telson width (from the type locality), anterior view. D, endopodite and 5th endite of 2nd trunk limb of a male of 2.2 mm telson width (from population 10), anterior view. E, second antenna of a second instar larva (from the type locality) as seen in a shed exuvium, ventral view. F, general aspect of adult male (showing a specimen of population 13). G, variation in the shape of the nuchal organ: examples (from left to right) from specimen of 2.2 (population 10) 1.8 (population 12, female) 2.4 (holotype) and 2.2 mm telson width (population 8). H, telson of the holotype, dorsal view. I, telson of a male of 2.3 mm telson width (from the type locality), lateral view.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF476657C6441F1BD3C4FC44AD91FC0C" httpUri="https://zenodo.org/record/263567/files/figure.png" pageId="15" pageNumber="343" targetBox="[153,1433,195,871]" targetPageId="15">
|
||
<paragraph id="8B8736DFC6441F1BD3C4FC44AD91FC0C" blockId="15.[151,1436,896,982]" pageId="15" pageNumber="343">
|
||
<emphasis id="B94CEACDC6441F1BD3C4FC44AD01FC4F" bold="true" box="[151,274,896,918]" pageId="15" pageNumber="343">FIGURE 9.</emphasis>
|
||
<taxonomicName id="4C384D5CC6441F1BD24BFC45ADD0FC4C" box="[280,451,896,918]" class="Branchiopoda" family="Triopsidae" genus="Triops" kingdom="Animalia" order="Notostraca" pageId="15" pageNumber="343" phylum="Arthropoda" rank="species" species="multifidus" status="sp. nov.">
|
||
<emphasis id="B94CEACDC6441F1BD24BFC45ADD0FC4C" box="[280,451,896,918]" italics="true" pageId="15" pageNumber="343">Triops multifidus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27F57B6C6441F1BD299FC44AE09FC4F" box="[458,538,896,917]" pageId="15" pageNumber="343" rank="species">
|
||
<emphasis id="B94CEACDC6441F1BD299FC44AE09FC4F" bold="true" box="[458,538,896,917]" pageId="15" pageNumber="343">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
A, carapace of the holotype (male), right lateral view. B, second maxilla of the holotype, anterior view. C, detail of the 1st trunk limb of the holotype, anterior view (note that the epipodite is missing in this sample, possibly due to injury).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B8736DFC6441F1BD394FBC6AF82FBE5" blockId="15.[151,1436,1026,1087]" pageId="15" pageNumber="343">
|
||
<emphasis id="B94CEACDC6441F1BD394FBC6AD09FBC1" bold="true" box="[199,282,1026,1051]" pageId="15" pageNumber="343">Range.</emphasis>
|
||
Lowlands west of the High Atlas Mountains between Casablanca and Anti-Atlas Mountains, except for the area of the Chaouia plain inhabited by the preceding species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |