treatments-xml/data/03/91/87/039187B1FF976A69FF51FF3F8FDB4739.xml

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<mods:title id="8DC238FBB48AE6A67B228A35B251DF6B">Nemacheilus pezidion, a new species of loach from southern Laos (Teleostei: Nemacheilidae)</mods:title>
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<mods:namePart id="0287C27B13494318A5B94CDBDC88080E">Kottelat, Maurice</mods:namePart>
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<mods:affiliation id="5D94991502625DCF3F6F839D38017ED1">Rue des Rauraques 6, 2800 Delémont, Switzerland (permanent address); and Lee Kong Chian Natural History Museum, National University of Singapore, 2 Conservatory Drive, Singapore 117377. mkottelat @ dplanet. ch; https: // orcid. org / 0000 - 0003 - 3638 - 2824</mods:affiliation>
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<mods:date id="03141D126B01F7FD1AC409789EDC15B0">2022</mods:date>
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<mods:number id="707A13027C8871B212B800348078010A">2022-04-26</mods:number>
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<treatment id="039187B1FF976A69FF51FF3F8FDB4739" ID-DOI="http://doi.org/10.5281/zenodo.6502471" ID-GBIF-Taxon="194854837" ID-Zenodo-Dep="6502471" LSID="urn:lsid:plazi:treatment:039187B1FF976A69FF51FF3F8FDB4739" httpUri="http://treatment.plazi.org/id/039187B1FF976A69FF51FF3F8FDB4739" lastPageId="10" lastPageNumber="102" pageId="1" pageNumber="93">
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<taxonomicName id="4C384D24FF976A62FF51FF3F8CD144B2" authority="Kottelat, 2022" authorityName="Kottelat" authorityYear="2022" box="[151,418,151,178]" class="Actinopterygii" family="Nemacheilidae" genus="Nemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="1" pageNumber="93" phylum="Chordata" rank="species" species="pezidion" status="sp. nov.">
<emphasis id="B94CEAB5FF976A62FF51FF3F8CD144B2" bold="true" box="[151,418,151,178]" italics="true" pageId="1" pageNumber="93">Nemacheilus pezidion</emphasis>
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, 
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(
<figureCitation id="13032A22FF976A62FF66FF138D9344D5" box="[160,224,187,213]" captionStart="FIGURE 1" captionStartId="2.[151,250,1943,1967]" captionTargetBox="[275,1313,181,1920]" captionTargetId="figure-16@2.[275,1313,181,1920]" captionTargetPageId="2" captionText="FIGURE 1. Nemacheilus pezidion, Laos: Xe Nam Noi: Xe Pian; a, MHNG 2787.088, holotype, 57.8 mm SL, male; b, CMK 15687, paratype, 53.8 mm SL, female; c, CMK 21330, paratype, 54.2 mm SL; female." figureDoi="http://doi.org/10.5281/zenodo.6488210" httpUri="https://zenodo.org/record/6488210/files/figure.png" pageId="1" pageNumber="93">Fig. 1</figureCitation>
)
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<subSubSection id="C322652CFF976A62FF51FEAB8C094661" pageId="1" pageNumber="93" type="materials_examined">
<paragraph id="8B8736A7FF976A62FF51FEAB8E464541" blockId="1.[151,1437,259,1978]" pageId="1" pageNumber="93">
<materialsCitation id="3B503CFAFF976A62FF51FEAB8E464541" ID-GBIF-Occurrence="3757640305" collectingDate="1999-05-22" collectionCode="MHNG" collectorName="M. Kottelat" country="Laos" county="Xe Namnoy" latitude="14.706111" location="Ban Mai" longLatPrecision="21" longitude="106.49611" municipality="Xe Pian" pageId="1" pageNumber="93" specimenCode="MHNG 2787.088" specimenCount="1" stateProvince="Attapu" typeStatus="holotype">
<emphasis id="B94CEAB5FF976A62FF51FEAB8C78451D" bold="true" box="[151,267,259,285]" pageId="1" pageNumber="93">
<typeStatus id="54838805FF976A62FF51FEAB8C75451D" box="[151,262,259,285]" pageId="1" pageNumber="93" type="holotype">Holotype</typeStatus>
.
</emphasis>
<specimenCode id="DB9E9EDCFF976A62FED1FEAB8C97451D" box="[279,484,259,285]" collectionCode="MHNG" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="1" pageNumber="93" type="Museum">MHNG 2787.088</specimenCode>
, 
<quantity id="4CC09B42FF976A62FE32FEAB8F2E451E" box="[500,605,259,286]" metricMagnitude="-2" metricUnit="m" metricValue="5.779999999999999" pageId="1" pageNumber="93" unit="mm" value="57.8">57.8 mm</quantity>
SL; 
<collectingCountry id="F32F7637FF976A62FD5BFEAC8FAB451E" box="[669,728,260,286]" name="Laos" pageId="1" pageNumber="93">Laos</collectingCountry>
: 
<collectingRegion id="49FCF845FF976A62FD20FEAB8EFA451E" box="[742,905,259,286]" country="Laos" name="Attapu" pageId="1" pageNumber="93">Attapeu Prov.</collectingRegion>
: 
<collectingCounty id="62E64E2BFF976A62FC5EFEAC8957451E" box="[920,1060,260,286]" pageId="1" pageNumber="93">Xe Namnoy</collectingCounty>
drainage: 
<collectingMunicipality id="6BE3ACDDFF976A62FB63FEAC8870451D" box="[1189,1283,259,286]" pageId="1" pageNumber="93">Xe Pian</collectingMunicipality>
at 
<location id="8EE7607CFF976A62FAE9FEAC88EB451D" LSID="urn:lsid:plazi:treatment:039187B1FF976A69FF51FF3F8FDB4739:8EE7607CFF976A62FAE9FEAC88EB451D" box="[1327,1432,259,286]" country="Laos" county="Xe Namnoy" latitude="14.706111" longLatPrecision="21" longitude="106.49611" municipality="Xe Pian" name="Ban Mai" pageId="1" pageNumber="93" stateProvince="Attapu">Ban Mai</location>
, 
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<geoCoordinate id="EE0C5060FF976A62FEE1FE8F8CCD4541" box="[295,446,295,321]" degrees="106" direction="east" minutes="29" orientation="longitude" pageId="1" pageNumber="93" precision="15" seconds="46" value="106.49611">106°29’46”E</geoCoordinate>
; 
<collectorName id="26CD5371FF976A62FE0CFE808F384541" box="[458,587,295,322]" pageId="1" pageNumber="93">M. Kottelat</collectorName>
<emphasis id="B94CEAB5FF976A62FD94FE808FF04541" box="[594,643,295,321]" italics="true" pageId="1" pageNumber="93">et al</emphasis>
., 
<date id="FF861067FF976A62FD5EFE8F8E424541" box="[664,817,295,322]" pageId="1" pageNumber="93" value="1999-05-22">
<collectingDate id="EFC2E98FFF976A62FD5EFE8F8E424541" box="[664,817,295,322]" pageId="1" pageNumber="93" value="1999-05-22">22 May 1999</collectingDate>
</date>
.
</materialsCitation>
</paragraph>
<paragraph id="8B8736A7FF976A62FF01FEE38C094661" blockId="1.[151,1437,259,1978]" pageId="1" pageNumber="93">
<materialsCitation id="3B503CFAFF976A62FF01FEE389774565" ID-GBIF-Occurrence="3757640312" box="[199,1028,331,358]" collectorName="Xe Namnoy &amp; Xe Pian" country="Laos" location="All" pageId="1" pageNumber="93" specimenCount="1" stateProvince="Attapu" typeStatus="paratype">
<emphasis id="B94CEAB5FF976A62FF01FEE38C354565" bold="true" box="[199,326,331,357]" pageId="1" pageNumber="93">
<typeStatus id="54838805FF976A62FF01FEE38C324565" box="[199,321,331,357]" pageId="1" pageNumber="93" type="paratype">Paratypes</typeStatus>
.
</emphasis>
<location id="8EE7607CFF976A62FE8AFEE38C034565" LSID="urn:lsid:plazi:treatment:039187B1FF976A69FF51FF3F8FDB4739:8EE7607CFF976A62FE8AFEE38C034565" box="[332,368,331,357]" country="Laos" name="All" pageId="1" pageNumber="93" stateProvince="Attapu">All</location>
from 
<collectingCountry id="F32F7637FF976A62FE70FEE48C824566" box="[438,497,332,358]" name="Laos" pageId="1" pageNumber="93">Laos</collectingCountry>
: 
<collectingRegion id="49FCF845FF976A62FE3DFEE38FE94566" box="[507,666,331,358]" country="Laos" name="Attapu" pageId="1" pageNumber="93">Attapeu Prov.</collectingRegion>
: 
<collectorName id="26CD5371FF976A62FD63FEE48E5E4566" box="[677,813,332,358]" pageId="1" pageNumber="93">Xe Namnoy</collectorName>
drainage: 
<collectorName id="26CD5371FF976A62FC63FEE489774565" box="[933,1028,331,358]" pageId="1" pageNumber="93">Xe Pian</collectorName>
</materialsCitation>
: 
<materialsCitation id="3B503CFAFF976A62FBC9FEE38CE14589" ID-GBIF-Occurrence="3757640322" accessionNumber="CMK 15687" collectingDate="1999-05-22" collectionCode="CMK" collectorName="M. Kottelat" country="Laos" county="Xe Namnoy" latitude="14.706111" location="Ban Mai" longLatPrecision="21" longitude="106.49611" municipality="Xe Pian" pageId="1" pageNumber="93" specimenCode="2" specimenCount="1" stateProvince="Attapu" typeStatus="paratype">
<collectionCode id="ED29AE62FF976A62FBC9FEE3893C4565" box="[1039,1103,331,357]" pageId="1" pageNumber="93">CMK</collectionCode>
15687, 
<specimenCode id="DB9E9EDCFF976A62FB6AFEE389CF4565" box="[1196,1212,331,357]" collectionCode="CMK" pageId="1" pageNumber="93">2</specimenCode>
, 
<quantity id="4CC09B42FF976A62FB0EFEE3881F4566" box="[1224,1388,331,358]" metricMagnitude="-2" metricUnit="m" metricValue="5.109999999999999" metricValueMax="5.38" metricValueMin="4.84" pageId="1" pageNumber="93" unit="mm" value="51.099999999999994" valueMax="53.8" valueMin="48.4">48.4–53.8 mm</quantity>
SL; same data as holotype
</materialsCitation>
.— 
<materialsCitation id="3B503CFAFF976A62FE74FEC78E214589" ID-GBIF-Occurrence="3757640313" accessionNumber="CMK 21246" box="[434,850,367,394]" collectionCode="CMK" country="Laos" pageId="1" pageNumber="93" specimenCode="15" specimenCount="1" stateProvince="Attapu" typeStatus="paratype">
<collectionCode id="ED29AE62FF976A62FE74FEC78C804589" box="[434,499,367,393]" pageId="1" pageNumber="93">CMK</collectionCode>
21246, 
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, 
<quantity id="4CC09B42FF976A62FDB8FEC78E56458A" box="[638,805,367,394]" metricMagnitude="-2" metricUnit="m" metricValue="5.18" metricValueMax="5.6899999999999995" metricValueMin="4.67" pageId="1" pageNumber="93" unit="mm" value="51.8" valueMax="56.9" valueMin="46.7">46.7–56.9 mm</quantity>
SL
</materialsCitation>
; 
<materialsCitation id="3B503CFAFF976A62FC98FEC7890C45AD" ID-GBIF-Occurrence="3757640303" accessionNumber="ZRC 62559" collectingDate="2009-05-20" collectionCode="ZRC" collectorName="M. Kottelat" country="Laos" latitude="14.725277" location="Ban Mai" longLatPrecision="21" longitude="106.494995" municipality="Xe Pian" pageId="1" pageNumber="93" specimenCode="6" specimenCount="1" stateProvince="Attapu" typeStatus="paratype">
<collectionCode id="ED29AE62FF976A62FC98FEC78EE64589" box="[862,917,367,393]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="1" pageNumber="93">ZRC</collectionCode>
62559, 
<specimenCode id="DB9E9EDCFF976A62FC32FEC789774589" box="[1012,1028,367,393]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="1" pageNumber="93">6</specimenCode>
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<quantity id="4CC09B42FF976A62FBD4FEC789CB458A" box="[1042,1208,367,394]" metricMagnitude="-2" metricUnit="m" metricValue="5.15" metricValueMax="5.39" metricValueMin="4.91" pageId="1" pageNumber="93" unit="mm" value="51.5" valueMax="53.9" valueMin="49.1">49.1–53.9 mm</quantity>
SL; 
<collectingMunicipality id="6BE3ACDDFF976A62FB37FED888224589" box="[1265,1361,367,394]" pageId="1" pageNumber="93">Xe Pian</collectingMunicipality>
, about 
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upstream of 
<location id="8EE7607CFF976A62FEA1FE3C8CBD45AD" LSID="urn:lsid:plazi:treatment:039187B1FF976A69FF51FF3F8FDB4739:8EE7607CFF976A62FEA1FE3C8CBD45AD" box="[359,462,403,430]" country="Laos" latitude="14.725277" longLatPrecision="21" longitude="106.494995" municipality="Xe Pian" name="Ban Mai" pageId="1" pageNumber="93" stateProvince="Attapu">Ban Mai</location>
, 
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; 
<collectorName id="26CD5371FF976A62FCD5FE3C8EE545AD" box="[787,918,403,430]" pageId="1" pageNumber="93">M. Kottelat</collectorName>
<emphasis id="B94CEAB5FF976A62FC58FE3C8EA345AD" box="[926,976,403,429]" italics="true" pageId="1" pageNumber="93">et al</emphasis>
., 
<date id="FF861067FF976A62FC20FE3B890C45AD" box="[998,1151,403,430]" pageId="1" pageNumber="93" value="2009-05-20">
<collectingDate id="EFC2E98FFF976A62FC20FE3B890C45AD" box="[998,1151,403,430]" pageId="1" pageNumber="93" value="2009-05-20">20 May 2009</collectingDate>
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</materialsCitation>
.— 
<materialsCitation id="3B503CFAFF976A62FB66FE3B8DA345F5" ID-GBIF-Occurrence="3757640314" accessionNumber="CMK 21280" collectingDate="2009-05-20" collectionCode="CMK" collectorName="M. Kottelat" country="Laos" latitude="14.6925" location="Ban Mai" longLatPrecision="21" longitude="106.47806" municipality="Xe Pian" pageId="1" pageNumber="93" specimenCode="3" specimenCount="1" stateProvince="Attapu" typeStatus="paratype">
<collectionCode id="ED29AE62FF976A62FB66FE3B899145AD" box="[1184,1250,403,429]" pageId="1" pageNumber="93">CMK</collectionCode>
21280, 
<specimenCode id="DB9E9EDCFF976A62FAF9FE3B883C45AD" box="[1343,1359,403,429]" collectionCode="CMK" pageId="1" pageNumber="93">3</specimenCode>
, 
<quantity id="4CC09B42FF976A62FA9AFE3B8D8E45D2" metricMagnitude="-2" metricUnit="m" metricValue="4.54" metricValueMax="4.71" metricValueMin="4.37" pageId="1" pageNumber="93" unit="mm" value="45.4" valueMax="47.1" valueMin="43.7">43.7– 47.1 mm</quantity>
SL; 
<collectingMunicipality id="6BE3ACDDFF976A62FEF2FE108CE245D1" box="[308,401,439,466]" pageId="1" pageNumber="93">Xe Pian</collectingMunicipality>
, about 
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downstream of 
<location id="8EE7607CFF976A62FD17FE108E4545D1" LSID="urn:lsid:plazi:treatment:039187B1FF976A69FF51FF3F8FDB4739:8EE7607CFF976A62FD17FE108E4545D1" box="[721,822,439,466]" country="Laos" latitude="14.6925" longLatPrecision="21" longitude="106.47806" municipality="Xe Pian" name="Ban Mai" pageId="1" pageNumber="93" stateProvince="Attapu">Ban Mai</location>
, 
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<geoCoordinate id="EE0C5060FF976A62FC14FE1F891945D1" box="[978,1130,439,465]" degrees="106" direction="east" minutes="28" orientation="longitude" pageId="1" pageNumber="93" precision="15" seconds="41" value="106.47806">106°28’41”E</geoCoordinate>
; 
<collectorName id="26CD5371FF976A62FBB3FE10898445D1" box="[1141,1271,439,466]" pageId="1" pageNumber="93">M. Kottelat</collectorName>
<emphasis id="B94CEAB5FF976A62FB39FE10884345D1" box="[1279,1328,439,465]" italics="true" pageId="1" pageNumber="93">et al</emphasis>
., 
<date id="FF861067FF976A62FA83FE1F8DA345F5" pageId="1" pageNumber="93" value="2009-05-20">
<collectingDate id="EFC2E98FFF976A62FA83FE1F8DA345F5" pageId="1" pageNumber="93" value="2009-05-20">20 May 2009</collectingDate>
</date>
</materialsCitation>
.— 
<materialsCitation id="3B503CFAFF976A62FF37FE7388494619" ID-GBIF-Occurrence="3757640319" accessionNumber="CMK 21330" collectingDate="2009-05-21" collectionCode="CMK" collectorName="M. Kottelat" country="Laos" county="Houai Pin" elevation="300" latitude="14.679444" location="Ban Mai" longLatPrecision="21" longitude="106.48416" municipality="Xe Pian" pageId="1" pageNumber="93" specimenCode="14" specimenCount="1" stateProvince="Attapu" typeStatus="paratype">
<collectionCode id="ED29AE62FF976A62FF37FE738C4145F5" box="[241,306,475,501]" pageId="1" pageNumber="93">CMK</collectionCode>
21330, 
<specimenCode id="DB9E9EDCFF976A62FE4CFE738CDB45F5" box="[394,424,475,501]" collectionCode="CMK" pageId="1" pageNumber="93">14</specimenCode>
, 
<quantity id="4CC09B42FF976A62FE75FE738F2645F6" box="[435,597,475,502]" metricMagnitude="-2" metricUnit="m" metricValue="4.745" metricValueMax="5.4799999999999995" metricValueMin="4.01" pageId="1" pageNumber="93" unit="mm" value="47.45" valueMax="54.8" valueMin="40.1">40.1–54.8 mm</quantity>
SL; 
<collectingCounty id="62E64E2BFF976A62FD4EFE738F8945F5" box="[648,762,475,501]" pageId="1" pageNumber="93">Houai Pin</collectingCounty>
, a tributary of 
<collectingMunicipality id="6BE3ACDDFF976A62FC5CFE748E8645F5" box="[922,1013,475,502]" pageId="1" pageNumber="93">Xe Pian</collectingMunicipality>
, entering it about 
<quantity id="4CC09B42FF976A62FB7FFE73898245F5" box="[1209,1265,475,502]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="1" pageNumber="93" unit="km" value="5.0">5 km</quantity>
downstream of 
<location id="8EE7607CFF976A62FF51FDA88D8F4619" LSID="urn:lsid:plazi:treatment:039187B1FF976A69FF51FF3F8FDB4739:8EE7607CFF976A62FF51FDA88D8F4619" box="[151,252,511,538]" country="Laos" county="Houai Pin" latitude="14.679444" longLatPrecision="21" longitude="106.48416" municipality="Xe Pian" name="Ban Mai" pageId="1" pageNumber="93" stateProvince="Attapu">Ban Mai</location>
; about 
<quantity id="4CC09B42FF976A62FE8CFE578CE3461A" box="[330,400,511,538]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="1" pageNumber="93" unit="m" value="300.0">
<elevation id="0015D194FF976A62FE8CFE578CE3461A" box="[330,400,511,538]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="1" pageNumber="93" unit="m" value="300.0">300 m</elevation>
</quantity>
upstream of confluence, 
<geoCoordinate id="EE0C5060FF976A62FD6FFE578E404619" box="[681,819,511,537]" degrees="14" direction="north" minutes="40" orientation="latitude" pageId="1" pageNumber="93" precision="15" seconds="46" value="14.679444">14°40’46”N</geoCoordinate>
<geoCoordinate id="EE0C5060FF976A62FCFEFE578EA34619" box="[824,976,511,537]" degrees="106" direction="east" minutes="29" orientation="longitude" pageId="1" pageNumber="93" precision="15" seconds="03" value="106.48416">106°29’03”E</geoCoordinate>
; 
<collectorName id="26CD5371FF976A62FC1CFDA889284619" box="[986,1115,511,538]" pageId="1" pageNumber="93">M. Kottelat</collectorName>
<emphasis id="B94CEAB5FF976A62FBA7FDA889E14619" box="[1121,1170,511,537]" italics="true" pageId="1" pageNumber="93">et al</emphasis>
., 
<date id="FF861067FF976A62FB60FE5788494619" box="[1190,1338,511,538]" pageId="1" pageNumber="93" value="2009-05-21">
<collectingDate id="EFC2E98FFF976A62FB60FE5788494619" box="[1190,1338,511,538]" pageId="1" pageNumber="93" value="2009-05-21">21 May 2009</collectingDate>
</date>
</materialsCitation>
.— 
<materialsCitation id="3B503CFAFF976A62FA9DFE578C054661" ID-GBIF-Occurrence="3757640317" accessionNumber="CMK 21315" collectingDate="2009-05-21" collectionCode="CMK" collectorName="M. Kottelat" country="Laos" latitude="14.657222" location="Ban Mai" longLatPrecision="21" longitude="106.47417" municipality="Xe Pian" pageId="1" pageNumber="93" specimenCode="1" specimenCount="1" stateProvince="Attapu" typeStatus="paratype">
<collectionCode id="ED29AE62FF976A62FA9DFE5788EF4619" box="[1371,1436,511,537]" pageId="1" pageNumber="93">CMK</collectionCode>
21315, 
<specimenCode id="DB9E9EDCFF976A62FF2CFD8B8D8B463D" box="[234,248,547,573]" collectionCode="CMK" pageId="1" pageNumber="93">1</specimenCode>
, 
<quantity id="4CC09B42FF976A62FEC0FD8B8C1A463E" box="[262,361,547,574]" metricMagnitude="-2" metricUnit="m" metricValue="3.92" pageId="1" pageNumber="93" unit="mm" value="39.2">39.2 mm</quantity>
SL; 
<collectingMunicipality id="6BE3ACDDFF976A62FE5BFD8C8C8A463D" box="[413,505,547,574]" pageId="1" pageNumber="93">Xe Pian</collectingMunicipality>
, rapids about 
<quantity id="4CC09B42FF976A62FD54FD8B8FB9463D" box="[658,714,547,573]" metricMagnitude="3" metricUnit="m" metricValue="6.0" pageId="1" pageNumber="93" unit="km" value="6.0">6 km</quantity>
downstream of 
<location id="8EE7607CFF976A62FCBAFD8C8E92463D" LSID="urn:lsid:plazi:treatment:039187B1FF976A69FF51FF3F8FDB4739:8EE7607CFF976A62FCBAFD8C8E92463D" box="[892,993,547,574]" country="Laos" latitude="14.657222" longLatPrecision="21" longitude="106.47417" municipality="Xe Pian" name="Ban Mai" pageId="1" pageNumber="93" stateProvince="Attapu">Ban Mai</location>
; 
<geoCoordinate id="EE0C5060FF976A62FC2DFD8B8907463D" box="[1003,1140,547,573]" degrees="14" direction="north" minutes="39" orientation="latitude" pageId="1" pageNumber="93" precision="15" seconds="26" value="14.657222">14°39’26”N</geoCoordinate>
<geoCoordinate id="EE0C5060FF976A62FBBCFD8B8861463D" box="[1146,1298,547,573]" degrees="106" direction="east" minutes="28" orientation="longitude" pageId="1" pageNumber="93" precision="15" seconds="27" value="106.47417">106°28’27”E</geoCoordinate>
; 
<collectorName id="26CD5371FF976A62FADAFD8C88EF463D" box="[1308,1436,547,574]" pageId="1" pageNumber="93">M. Kottelat</collectorName>
<emphasis id="B94CEAB5FF976A62FF51FDE08DBB4661" box="[151,200,583,609]" italics="true" pageId="1" pageNumber="93">et al</emphasis>
., 
<date id="FF861067FF976A62FF1BFDEF8C054661" box="[221,374,583,610]" pageId="1" pageNumber="93" value="2009-05-21">
<collectingDate id="EFC2E98FFF976A62FF1BFDEF8C054661" box="[221,374,583,610]" pageId="1" pageNumber="93" value="2009-05-21">21 May 2009</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C322652CFF976A62FF01FDC388FA47A6" pageId="1" pageNumber="93" type="diagnosis">
<paragraph id="8B8736A7FF976A62FF01FDC388FA47A6" blockId="1.[151,1437,259,1978]" pageId="1" pageNumber="93">
<emphasis id="B94CEAB5FF976A62FF01FDC38C314685" bold="true" box="[199,322,619,645]" pageId="1" pageNumber="93">Diagnosis.</emphasis>
<taxonomicName id="4C384D24FF976A62FE8CFDC38F304685" authorityName="Kottelat" authorityYear="2022" box="[330,579,619,645]" class="Actinopterygii" family="Nemacheilidae" genus="Nemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="1" pageNumber="93" phylum="Chordata" rank="species" species="pezidion">
<emphasis id="B94CEAB5FF976A62FE8CFDC38F304685" box="[330,579,619,645]" italics="true" pageId="1" pageNumber="93">Nemacheilus pezidion</emphasis>
</taxonomicName>
is distinguished from most species of 
<taxonomicName id="4C384D24FF976A62FBC6FDC389DB4685" box="[1024,1192,619,645]" class="Actinopterygii" family="Nemacheilidae" kingdom="Animalia" order="Cypriniformes" pageId="1" pageNumber="93" phylum="Chordata" rank="family">Nemacheilidae</taxonomicName>
in Southeast Asia by having an obvious dark brown to black midlateral stripe. A somewhat similar pattern is present only in 
<taxonomicName id="4C384D24FF976A62FACBFD3888E646A9" box="[1293,1429,655,681]" class="Actinopterygii" family="Nemacheilidae" genus="Nemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="1" pageNumber="93" phylum="Chordata" rank="species" species="binotatus">
<emphasis id="B94CEAB5FF976A62FACBFD3888E646A9" box="[1293,1429,655,681]" italics="true" pageId="1" pageNumber="93">N. binotatus</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C384D24FF976A62FF51FD1C8C3B46CD" authorityName="Kottelat" authorityYear="1990" box="[151,328,691,717]" class="Actinopterygii" family="Nemacheilidae" genus="Nemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="1" pageNumber="93" phylum="Chordata" rank="species" species="longistriatus">
<emphasis id="B94CEAB5FF976A62FF51FD1C8C3B46CD" box="[151,328,691,717]" italics="true" pageId="1" pageNumber="93">N. longistriatus</emphasis>
</taxonomicName>
, 
<emphasis id="B94CEAB5FF976A62FE91FD1C8F6A46CD" box="[343,537,692,718]" italics="true" pageId="1" pageNumber="93">‘N.’ argyrogaster</emphasis>
, 
<taxonomicName id="4C384D24FF976A62FDEFFD1C8FAC46CD" box="[553,735,691,717]" class="Actinopterygii" family="Nemacheilidae" genus="Nemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="1" pageNumber="93" phylum="Chordata" rank="species" species="nandingensis">
<emphasis id="B94CEAB5FF976A62FDEFFD1C8FAC46CD" box="[553,735,691,717]" italics="true" pageId="1" pageNumber="93">N. nandingensis</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C384D24FF976A62FD29FD1B897746CD" authorityName="Bohlen &amp; Slechtova" authorityYear="2013" box="[751,1028,691,717]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="1" pageNumber="93" phylum="Chordata" rank="species" species="rubrimaculata">
<emphasis id="B94CEAB5FF976A62FD29FD1B897746CD" box="[751,1028,691,717]" italics="true" pageId="1" pageNumber="93">Schistura rubrimaculata</emphasis>
</taxonomicName>
and 
<emphasis id="B94CEAB5FF976A62FBFBFD1B89BA46CD" box="[1085,1225,691,717]" italics="true" pageId="1" pageNumber="93">
<taxonomicName id="4C384D24FF976A62FBFBFD1B89B646CD" authorityName="Bohlen &amp; Slechtova" authorityYear="2013" box="[1085,1221,691,717]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="1" pageNumber="93" phylum="Chordata" rank="species" species="pawensis">S. pawensis</taxonomicName>
.
</emphasis>
It is distinguished from them in having a forked caudal fin with the upper lobe distinctly longer than the lower one (vs. subequal), the pectoral-fin rays with short filamentous extensions in both sexes (vs. without extensions), the presence of an elongate spot on lower ¼ of first branched dorsal-fin ray and adjacent membranes (vs. absence), and the presence of 5 supraorbital pores (vs. 6). In addition, males of 
<taxonomicName id="4C384D24FF976A62FC46FCEC8976475D" authorityName="Kottelat" authorityYear="2022" box="[896,1029,835,861]" class="Actinopterygii" family="Nemacheilidae" genus="Nemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="1" pageNumber="93" phylum="Chordata" rank="species" species="pezidion">
<emphasis id="B94CEAB5FF976A62FC46FCEC8976475D" box="[896,1029,835,861]" italics="true" pageId="1" pageNumber="93">N. pezidion</emphasis>
</taxonomicName>
have a modified pectoral fin with conspicuous tubercles on anterior rays and wide unculiferous pads (vs. minute tubercles, narrow unculiferous pads), and numerous small tubercles on the middle and posterior part of the flank of large males (vs. without tubercles).
</paragraph>
</subSubSection>
<subSubSection id="C322652CFF976A66FF01FC078E9A475E" lastPageId="5" lastPageNumber="97" pageId="1" pageNumber="93" type="description">
<paragraph id="8B8736A7FF976A62FF01FC07896840C6" blockId="1.[151,1437,259,1978]" pageId="1" pageNumber="93">
<emphasis id="B94CEAB5FF976A62FF01FC078C2A47C9" bold="true" box="[199,345,943,969]" pageId="1" pageNumber="93">Description.</emphasis>
See 
<figureCitation id="13032A22FF976A62FE51FC078C8947C9" box="[407,506,943,969]" captionStart="FIGURE 1" captionStartId="2.[151,250,1943,1967]" captionTargetBox="[275,1313,181,1920]" captionTargetId="figure-16@2.[275,1313,181,1920]" captionTargetPageId="2" captionText="FIGURE 1. Nemacheilus pezidion, Laos: Xe Nam Noi: Xe Pian; a, MHNG 2787.088, holotype, 57.8 mm SL, male; b, CMK 15687, paratype, 53.8 mm SL, female; c, CMK 21330, paratype, 54.2 mm SL; female." figureDoi="http://doi.org/10.5281/zenodo.6488210" httpUri="https://zenodo.org/record/6488210/files/figure.png" pageId="1" pageNumber="93">Figure 1</figureCitation>
for general appearance and 
<tableCitation id="C6BA031CFF976A62FC8DFC078ED147C9" box="[843,930,943,969]" captionStart="TABLE 1" captionStartId="3.[151,239,151,175]" captionTargetPageId="3" captionText="TABLE 1. Morphometric data of type material of Nemacheilus pezidion (n=11). Range and mean include holotype data." httpUri="http://table.plazi.org/id/DF47662FFF956A60FF51FF3F88EE44AF" pageId="1" pageNumber="93" tableUuid="DF47662FFF956A60FF51FF3F88EE44AF">Table 1</tableCitation>
for morphometric data of 
<typeStatus id="54838805FF976A62FB19FC07883247C9" box="[1247,1345,943,969]" pageId="1" pageNumber="93" type="holotype">holotype</typeStatus>
and 
<specimenCount id="9D3EFD2EFF976A62FA46FC078C7547EE" pageId="1" pageNumber="93" type="generic" typeStatus="paratypes">10 paratypes</specimenCount>
. An elongate nemacheilid with body depth gradually increasing up to slightly in front of dorsal-fin origin. Behind dorsal-fin origin, body depth decreasing slowly to caudal-fin base. Dorsal profile continuous between head and body. Head very slightly depressed; body slightly compressed anteriorly to compressed posteriorly. Interorbital area convex. In lateral view, eye usually below dorsal profile of head (flushed in a few specimens). Cheeks not swollen. Snout rounded. Depth of caudal peduncle 1.7–2.0 times in its length, tapering posteriorly. Low dorsal keel on posterior fourth of post-dorsal area. No ventral keel on caudal peduncle. Depth of caudal peduncle 1.5–1.9 times in body depth at dorsal-fin origin. Largest recorded size 
<quantity id="4CC09B42FF976A62FCC8FB038E0140C6" box="[782,882,1195,1222]" metricMagnitude="-2" metricUnit="m" metricValue="5.779999999999999" pageId="1" pageNumber="93" unit="mm" value="57.8">57.8 mm</quantity>
SL (
<typeStatus id="54838805FF976A62FC61FB03896340C6" box="[935,1040,1195,1222]" pageId="1" pageNumber="93" type="holotype">holotype</typeStatus>
).
</paragraph>
<paragraph id="8B8736A7FF976A62FF01FB678FC5422E" blockId="1.[151,1437,259,1978]" pageId="1" pageNumber="93">
Dorsal fin with 4 unbranched and 9½ branched rays; distal margin straight to slightly concave. First branched ray longest. Pectoral fin with 1 unbranched and 10 (1), 11 (7*) or 12 (3) branched rays (including small last ray, usually unbranched), falcate, reaching about ¾ of distance to pelvic-fin base; all rays with short filamentous extensions (
<figureCitation id="13032A22FF976A62FF66FA938D9B4155" box="[160,232,1339,1365]" captionStart="FIGURE 2" captionStartId="4.[151,250,744,768]" captionTargetBox="[151,1403,181,719]" captionTargetId="figure-16@4.[151,1435,181,719]" captionTargetPageId="4" captionText="FIGURE 2. Nemacheilus pezidion, pectoral fin in dorsal view of: a, male, MHNG 2787.088, holotype, 57.8 mm SL (note small tubercles on branched rays 1–6); and b, female, CMK 21246, 52.9 mm SL." figureDoi="http://doi.org/10.5281/zenodo.6488212" httpUri="https://zenodo.org/record/6488212/files/figure.png" pageId="1" pageNumber="93">Fig. 2</figureCitation>
). No axillary pectoral lobe. Pelvic fin with 1 unbranched and 7 branched rays (including small last ray, usually unbranched); reaching ³⁄ to ¾ of distance to anus; triangular; posterior margin straight; origin below base of branched dorsal-fin rays 1–2; axillary lobe present, free, small (equal to base of fin). Anus situated about 1–1½ eye diameter in front of anal fin, behind posterior extremity of pelvic fin. Anal fin with 3 unbranched and 5½ branched rays; distal margin almost straight. Caudal fin with 9+8 branched rays; about 6–7 dorsal and 6–7 ventral procurrent rays (cannot be counted with accuracy, possibly more), forked, lobes pointed, upper lobe 1.1–1.3 times length of lower lobe, 1.9–2.2 times length of median rays.
</paragraph>
<paragraph id="8B8736A7FF976A62FF01F99F8EDB42BE" blockId="1.[151,1437,259,1978]" pageId="1" pageNumber="93">Body entirely covered by scales. Scales embedded. Lateral line complete, with about 93–102 pores (due to shrinkage, lateral line sunk in gutter and impossible to count with accuracy). Scales on flank in large males with tubercles (see details under Sexual dimorphism below). Cephalic lateral line system with 5 supraorbital, 4 + 10–11 infraorbital, 8–10 preoperculo-mandibular and 3 supratemporal pores.</paragraph>
<paragraph id="8B8736A7FF976A62FF01F96F8C5B43B9" blockId="1.[151,1437,259,1978]" pageId="1" pageNumber="93">
Anterior naris pierced in front side of a pointed flap-like tube. Posterior naris adjacent to anterior one. Mouth U-shaped, gape about 2 times wider than long (
<figureCitation id="13032A22FF976A62FD7BF9438E774305" box="[701,772,1771,1797]" captionStart="FIGURE 3" captionStartId="4.[151,250,1636,1660]" captionTargetId="figure-53@4.[297,1291,828,1612]" captionTargetPageId="4" captionText="FIGURE 3. Nemacheilus pezidion, CMK 21246, 52.9 mm SL; mouth." figureDoi="http://doi.org/10.5281/zenodo.6502469" httpUri="https://zenodo.org/record/6502469/files/figure.png" pageId="1" pageNumber="93">Fig. 3</figureCitation>
). Lips thin but fleshy. Upper lip without median notch or with a small, shallow median notch, numerous wrinkles; edge crenulated. Processus dentiformis present. Lower lip without median interruption or small notch in outer part of lip; median part with 4–5 sulci, lateral parts smooth; posterior margin of lip thin, smooth. Tip of lower jaw not exposed. No median concavity in lower jaw. Inner rostral barbel reaching vertical of middle of eye. Outer rostral barbel and maxillary barbel reaching middle of postorbital area of head.
</paragraph>
<caption id="DF47662FFF946A61FF51F83F897043D4" ID-DOI="http://doi.org/10.5281/zenodo.6488210" ID-Zenodo-Dep="6488210" httpUri="https://zenodo.org/record/6488210/files/figure.png" pageId="2" pageNumber="94" startId="2.[151,250,1943,1967]" targetBox="[275,1313,181,1920]" targetPageId="2">
<paragraph id="8B8736A7FF946A61FF51F83F897043D4" blockId="2.[151,1436,1943,2004]" pageId="2" pageNumber="94">
<emphasis id="B94CEAB5FF946A61FF51F83F8C6743AF" bold="true" box="[151,276,1943,1967]" pageId="2" pageNumber="94">FIGURE 1.</emphasis>
<taxonomicName id="4C384D24FF946A61FEDDF8308C8843B0" authorityName="Kottelat" authorityYear="2022" box="[283,507,1944,1968]" class="Actinopterygii" family="Nemacheilidae" genus="Nemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="94" phylum="Chordata" rank="species" species="pezidion">
<emphasis id="B94CEAB5FF946A61FEDDF8308C8843B0" box="[283,507,1944,1968]" italics="true" pageId="2" pageNumber="94">Nemacheilus pezidion</emphasis>
</taxonomicName>
, Laos: Xe Nam Noi: Xe Pian; 
<emphasis id="B94CEAB5FF946A61FCFFF8308E3F43B0" bold="true" box="[825,844,1944,1968]" pageId="2" pageNumber="94">a,</emphasis>
MHNG 2787.088, holotype, 57.8 mm SL, male; 
<emphasis id="B94CEAB5FF946A61FA81F830882843B0" bold="true" box="[1351,1371,1944,1968]" pageId="2" pageNumber="94">b,</emphasis>
CMK 15687, paratype, 53.8 mm SL, female; 
<emphasis id="B94CEAB5FF946A61FDE2F8148F4543D4" bold="true" box="[548,566,1980,2004]" pageId="2" pageNumber="94">c,</emphasis>
CMK 21330, paratype, 54.2 mm SL; female.
</paragraph>
</caption>
<caption id="DF47662FFF956A60FF51FF3F88EE44AF" ID-Table-UUID="DF47662FFF956A60FF51FF3F88EE44AF" box="[151,1437,151,176]" httpUri="http://table.plazi.org/id/DF47662FFF956A60FF51FF3F88EE44AF" pageId="3" pageNumber="95" startId="3.[151,239,151,175]" targetBox="[166,1357,192,1584]" targetIsTable="true" targetPageId="3">
<paragraph id="8B8736A7FF956A60FF51FF3F88EE44AF" blockId="3.[151,1437,151,176]" box="[151,1437,151,176]" pageId="3" pageNumber="95">
<emphasis id="B94CEAB5FF956A60FF51FF3F8C7944AF" bold="true" box="[151,266,151,175]" pageId="3" pageNumber="95">TABLE 1.</emphasis>
Morphometric data of type material of 
<taxonomicName id="4C384D24FF956A60FD77FF3F8EE844AF" authorityName="Kottelat" authorityYear="2022" box="[689,923,151,175]" class="Actinopterygii" family="Nemacheilidae" genus="Nemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="3" pageNumber="95" phylum="Chordata" rank="species" species="pezidion">
<emphasis id="B94CEAB5FF956A60FD77FF3F8EE844AF" box="[689,923,151,175]" italics="true" pageId="3" pageNumber="95">Nemacheilus pezidion</emphasis>
</taxonomicName>
(n=11). Range and mean include holotype data.
</paragraph>
</caption>
<paragraph id="8B8736A7FF956A60FD5CFF68884E4230" pageId="3" pageNumber="95">
<table id="F938C407FF95959CFF60FF68883E4230" box="[166,1357,192,1584]" gridcols="4" gridrows="35" pageId="3" pageNumber="95">
<tr id="350834E5FF95959CFF60FF68883E44D8" box="[166,1357,192,216]" gridrow="0" pageId="3" pageNumber="95" rowspan-0="1">
<th id="76D95D99FF95959CFD5CFF688F8044D8" box="[666,755,192,216]" gridcol="1" gridrow="0" pageId="3" pageNumber="95">holotype</th>
<th id="76D95D99FF95959CFC0DFF68893944D8" box="[971,1098,192,216]" gridcol="2" gridrow="0" pageId="3" pageNumber="95">range</th>
<th id="76D95D99FF95959CFAD2FF68883E44D8" box="[1300,1357,192,216]" gridcol="3" gridrow="0" pageId="3" pageNumber="95">mean</th>
</tr>
<tr id="350834E5FF95959CFF60FF40883E4500" box="[166,1357,232,256]" gridrow="1" pageId="3" pageNumber="95" rowspan-3="1">
<th id="76D95D99FF95959CFF60FF408F774500" box="[166,516,232,256]" gridcol="0" gridrow="1" pageId="3" pageNumber="95">Standard length (mm)</th>
<td id="76D95D99FF95959CFD5CFF408F804500" box="[666,755,232,256]" gridcol="1" gridrow="1" pageId="3" pageNumber="95">57.8</td>
<td id="76D95D99FF95959CFC0DFF4089394500" box="[971,1098,232,256]" gridcol="2" gridrow="1" pageId="3" pageNumber="95">46.6–57.8</td>
</tr>
<tr id="350834E5FF95959CFF60FEB8883E4528" box="[166,1357,272,296]" gridrow="2" pageId="3" pageNumber="95" rowspan-3="1">
<th id="76D95D99FF95959CFF60FEB88F774528" box="[166,516,272,296]" gridcol="0" gridrow="2" pageId="3" pageNumber="95">Total length (mm)</th>
<td id="76D95D99FF95959CFD5CFEB88F804528" box="[666,755,272,296]" gridcol="1" gridrow="2" pageId="3" pageNumber="95">76.7</td>
<td id="76D95D99FF95959CFC0DFEB889394528" box="[971,1098,272,296]" gridcol="2" gridrow="2" pageId="3" pageNumber="95">59.6–76.7</td>
</tr>
<tr id="350834E5FF95959CFF60FE90883E4550" box="[166,1357,312,336]" gridrow="3" pageId="3" pageNumber="95">
<th id="76D95D99FF95959CFF60FE90883E4550" box="[166,1357,312,336]" colspan="4" colspanRight="3" gridcol="0" gridrow="3" pageId="3" pageNumber="95">In percent of standard length</th>
</tr>
<tr id="350834E5FF95959CFF60FEC9883E4579" box="[166,1357,353,377]" gridrow="4" pageId="3" pageNumber="95">
<th id="76D95D99FF95959CFF60FEC98F774579" box="[166,516,353,377]" gridcol="0" gridrow="4" pageId="3" pageNumber="95">Total length</th>
<td id="76D95D99FF95959CFD5CFEC98F804579" box="[666,755,353,377]" gridcol="1" gridrow="4" pageId="3" pageNumber="95">132.8</td>
<td id="76D95D99FF95959CFC0DFEC989394579" box="[971,1098,353,377]" gridcol="2" gridrow="4" pageId="3" pageNumber="95">126.7–133.4</td>
<td id="76D95D99FF95959CFAD2FEC9883E4579" box="[1300,1357,353,377]" gridcol="3" gridrow="4" pageId="3" pageNumber="95">130.1</td>
</tr>
<tr id="350834E5FF95959CFF60FE21883E45A1" box="[166,1357,393,417]" gridrow="5" pageId="3" pageNumber="95">
<th id="76D95D99FF95959CFF60FE218F7745A1" box="[166,516,393,417]" gridcol="0" gridrow="5" pageId="3" pageNumber="95">Head length (dorsal)</th>
<td id="76D95D99FF95959CFD5CFE218F8045A1" box="[666,755,393,417]" gridcol="1" gridrow="5" pageId="3" pageNumber="95">18.7</td>
<td id="76D95D99FF95959CFC0DFE21893945A1" box="[971,1098,393,417]" gridcol="2" gridrow="5" pageId="3" pageNumber="95">18.0–20.9</td>
<td id="76D95D99FF95959CFAD2FE21883E45A1" box="[1300,1357,393,417]" gridcol="3" gridrow="5" pageId="3" pageNumber="95">19.4</td>
</tr>
<tr id="350834E5FF95959CFF60FE19883E45C9" box="[166,1357,433,457]" gridrow="6" pageId="3" pageNumber="95">
<th id="76D95D99FF95959CFF60FE198F7745C9" box="[166,516,433,457]" gridcol="0" gridrow="6" pageId="3" pageNumber="95">Head length (lateral)</th>
<td id="76D95D99FF95959CFD5CFE198F8045C9" box="[666,755,433,457]" gridcol="1" gridrow="6" pageId="3" pageNumber="95">21.8</td>
<td id="76D95D99FF95959CFC0DFE19893945C9" box="[971,1098,433,457]" gridcol="2" gridrow="6" pageId="3" pageNumber="95">21.2–22.9</td>
<td id="76D95D99FF95959CFAD2FE19883E45C9" box="[1300,1357,433,457]" gridcol="3" gridrow="6" pageId="3" pageNumber="95">22.0</td>
</tr>
<tr id="350834E5FF95959CFF60FE71883E45F2" box="[166,1357,473,498]" gridrow="7" pageId="3" pageNumber="95">
<th id="76D95D99FF95959CFF60FE718F7745F2" box="[166,516,473,498]" gridcol="0" gridrow="7" pageId="3" pageNumber="95">Predorsal length</th>
<td id="76D95D99FF95959CFD5CFE718F8045F2" box="[666,755,473,498]" gridcol="1" gridrow="7" pageId="3" pageNumber="95">47.9</td>
<td id="76D95D99FF95959CFC0DFE71893945F2" box="[971,1098,473,498]" gridcol="2" gridrow="7" pageId="3" pageNumber="95">45.2–48.2</td>
<td id="76D95D99FF95959CFAD2FE71883E45F2" box="[1300,1357,473,498]" gridcol="3" gridrow="7" pageId="3" pageNumber="95">46.9</td>
</tr>
<tr id="350834E5FF95959CFF60FDAA883E461A" box="[166,1357,514,538]" gridrow="8" pageId="3" pageNumber="95">
<th id="76D95D99FF95959CFF60FDAA8F77461A" box="[166,516,514,538]" gridcol="0" gridrow="8" pageId="3" pageNumber="95">Prepelvic length</th>
<td id="76D95D99FF95959CFD5CFDAA8F80461A" box="[666,755,514,538]" gridcol="1" gridrow="8" pageId="3" pageNumber="95">48.6</td>
<td id="76D95D99FF95959CFC0DFDAA8939461A" box="[971,1098,514,538]" gridcol="2" gridrow="8" pageId="3" pageNumber="95">47.6–49.4</td>
<td id="76D95D99FF95959CFAD2FDAA883E461A" box="[1300,1357,514,538]" gridcol="3" gridrow="8" pageId="3" pageNumber="95">48.7</td>
</tr>
<tr id="350834E5FF95959CFF60FD82883E4642" box="[166,1357,554,578]" gridrow="9" pageId="3" pageNumber="95">
<th id="76D95D99FF95959CFF60FD828F774642" box="[166,516,554,578]" gridcol="0" gridrow="9" pageId="3" pageNumber="95">Pre-anus length</th>
<td id="76D95D99FF95959CFD5CFD828F804642" box="[666,755,554,578]" gridcol="1" gridrow="9" pageId="3" pageNumber="95">69.4</td>
<td id="76D95D99FF95959CFC0DFD8289394642" box="[971,1098,554,578]" gridcol="2" gridrow="9" pageId="3" pageNumber="95">64.2–69.4</td>
<td id="76D95D99FF95959CFAD2FD82883E4642" box="[1300,1357,554,578]" gridcol="3" gridrow="9" pageId="3" pageNumber="95">67.7</td>
</tr>
<tr id="350834E5FF95959CFF60FDFA883E466A" box="[166,1357,594,618]" gridrow="10" pageId="3" pageNumber="95">
<th id="76D95D99FF95959CFF60FDFA8F77466A" box="[166,516,594,618]" gridcol="0" gridrow="10" pageId="3" pageNumber="95">Pre-anal length</th>
<td id="76D95D99FF95959CFD5CFDFA8F80466A" box="[666,755,594,618]" gridcol="1" gridrow="10" pageId="3" pageNumber="95">76.9</td>
<td id="76D95D99FF95959CFC0DFDFA8939466A" box="[971,1098,594,618]" gridcol="2" gridrow="10" pageId="3" pageNumber="95">74.6–77.6</td>
<td id="76D95D99FF95959CFAD2FDFA883E466A" box="[1300,1357,594,618]" gridcol="3" gridrow="10" pageId="3" pageNumber="95">76.3</td>
</tr>
<tr id="350834E5FF95959CFF60FDD2883E4693" box="[166,1357,634,659]" gridrow="11" pageId="3" pageNumber="95">
<th id="76D95D99FF95959CFF60FDD28F774693" box="[166,516,634,659]" gridcol="0" gridrow="11" pageId="3" pageNumber="95">Head depth</th>
<td id="76D95D99FF95959CFD5CFDD28F804693" box="[666,755,634,659]" gridcol="1" gridrow="11" pageId="3" pageNumber="95">11.4</td>
<td id="76D95D99FF95959CFC0DFDD289394693" box="[971,1098,634,659]" gridcol="2" gridrow="11" pageId="3" pageNumber="95">10.7–12.2</td>
<td id="76D95D99FF95959CFAD2FDD2883E4693" box="[1300,1357,634,659]" gridcol="3" gridrow="11" pageId="3" pageNumber="95">11.5</td>
</tr>
<tr id="350834E5FF95959CFF60FD0B883E46BB" box="[166,1357,675,699]" gridrow="12" pageId="3" pageNumber="95">
<th id="76D95D99FF95959CFF60FD0B8F7746BB" box="[166,516,675,699]" gridcol="0" gridrow="12" pageId="3" pageNumber="95">Body depth at dorsal-fin origin</th>
<td id="76D95D99FF95959CFD5CFD0B8F8046BB" box="[666,755,675,699]" gridcol="1" gridrow="12" pageId="3" pageNumber="95">15.4</td>
<td id="76D95D99FF95959CFC0DFD0B893946BB" box="[971,1098,675,699]" gridcol="2" gridrow="12" pageId="3" pageNumber="95">15.0–19.0</td>
<td id="76D95D99FF95959CFAD2FD0B883E46BB" box="[1300,1357,675,699]" gridcol="3" gridrow="12" pageId="3" pageNumber="95">16.9</td>
</tr>
<tr id="350834E5FF95959CFF60FD63883E46E3" box="[166,1357,715,739]" gridrow="13" pageId="3" pageNumber="95">
<th id="76D95D99FF95959CFF60FD638F7746E3" box="[166,516,715,739]" gridcol="0" gridrow="13" pageId="3" pageNumber="95">Depth of caudal peduncle</th>
<td id="76D95D99FF95959CFD5CFD638F8046E3" box="[666,755,715,739]" gridcol="1" gridrow="13" pageId="3" pageNumber="95">10.0</td>
<td id="76D95D99FF95959CFC0DFD63893946E3" box="[971,1098,715,739]" gridcol="2" gridrow="13" pageId="3" pageNumber="95">9.3–10.5</td>
<td id="76D95D99FF95959CFAD2FD63883E46E3" box="[1300,1357,715,739]" gridcol="3" gridrow="13" pageId="3" pageNumber="95">9.9</td>
</tr>
<tr id="350834E5FF95959CFF60FD5B883E470B" box="[166,1357,755,779]" gridrow="14" pageId="3" pageNumber="95">
<th id="76D95D99FF95959CFF60FD5B8F77470B" box="[166,516,755,779]" gridcol="0" gridrow="14" pageId="3" pageNumber="95">Length of caudal peduncle</th>
<td id="76D95D99FF95959CFD5CFD5B8F80470B" box="[666,755,755,779]" gridcol="1" gridrow="14" pageId="3" pageNumber="95">17.5</td>
<td id="76D95D99FF95959CFC0DFD5B8939470B" box="[971,1098,755,779]" gridcol="2" gridrow="14" pageId="3" pageNumber="95">17.0–20.3</td>
<td id="76D95D99FF95959CFAD2FD5B883E470B" box="[1300,1357,755,779]" gridcol="3" gridrow="14" pageId="3" pageNumber="95">18.4</td>
</tr>
<tr id="350834E5FF95959CFF60FCB3883E4734" box="[166,1357,795,820]" gridrow="15" pageId="3" pageNumber="95">
<th id="76D95D99FF95959CFF60FCB38F774734" box="[166,516,795,820]" gridcol="0" gridrow="15" pageId="3" pageNumber="95">Head width</th>
<td id="76D95D99FF95959CFD5CFCB38F804734" box="[666,755,795,820]" gridcol="1" gridrow="15" pageId="3" pageNumber="95">13.0</td>
<td id="76D95D99FF95959CFC0DFCB389394734" box="[971,1098,795,820]" gridcol="2" gridrow="15" pageId="3" pageNumber="95">11.9–13.7</td>
<td id="76D95D99FF95959CFAD2FCB3883E4734" box="[1300,1357,795,820]" gridcol="3" gridrow="15" pageId="3" pageNumber="95">12.6</td>
</tr>
<tr id="350834E5FF95959CFF60FCEB883E475C" box="[166,1357,835,860]" gridrow="16" pageId="3" pageNumber="95">
<th id="76D95D99FF95959CFF60FCEB8F77475C" box="[166,516,835,860]" gridcol="0" gridrow="16" pageId="3" pageNumber="95">Body width at dorsal-fin origin</th>
<td id="76D95D99FF95959CFD5CFCEB8F80475C" box="[666,755,835,860]" gridcol="1" gridrow="16" pageId="3" pageNumber="95">11.0</td>
<td id="76D95D99FF95959CFC0DFCEB8939475C" box="[971,1098,835,860]" gridcol="2" gridrow="16" pageId="3" pageNumber="95">9.8–13.5</td>
<td id="76D95D99FF95959CFAD2FCEB883E475C" box="[1300,1357,835,860]" gridcol="3" gridrow="16" pageId="3" pageNumber="95">12.1</td>
</tr>
<tr id="350834E5FF95959CFF60FCC4883E4784" box="[166,1357,876,900]" gridrow="17" pageId="3" pageNumber="95">
<th id="76D95D99FF95959CFF60FCC48F774784" box="[166,516,876,900]" gridcol="0" gridrow="17" pageId="3" pageNumber="95">Snout length</th>
<td id="76D95D99FF95959CFD5CFCC48F804784" box="[666,755,876,900]" gridcol="1" gridrow="17" pageId="3" pageNumber="95">8.6</td>
<td id="76D95D99FF95959CFC0DFCC489394784" box="[971,1098,876,900]" gridcol="2" gridrow="17" pageId="3" pageNumber="95">8.1–8.6</td>
<td id="76D95D99FF95959CFAD2FCC4883E4784" box="[1300,1357,876,900]" gridcol="3" gridrow="17" pageId="3" pageNumber="95">8.3</td>
</tr>
<tr id="350834E5FF95959CFF60FC3C883E47AD" box="[166,1357,916,941]" gridrow="18" pageId="3" pageNumber="95">
<th id="76D95D99FF95959CFF60FC3C8F7747AD" box="[166,516,916,941]" gridcol="0" gridrow="18" pageId="3" pageNumber="95">Eye diameter</th>
<td id="76D95D99FF95959CFD5CFC3C8F8047AD" box="[666,755,916,941]" gridcol="1" gridrow="18" pageId="3" pageNumber="95">5.1</td>
<td id="76D95D99FF95959CFC0DFC3C893947AD" box="[971,1098,916,941]" gridcol="2" gridrow="18" pageId="3" pageNumber="95">4.3–5.7</td>
<td id="76D95D99FF95959CFAD2FC3C883E47AD" box="[1300,1357,916,941]" gridcol="3" gridrow="18" pageId="3" pageNumber="95">5.0</td>
</tr>
<tr id="350834E5FF95959CFF60FC14883E47D4" box="[166,1357,956,980]" gridrow="19" pageId="3" pageNumber="95">
<th id="76D95D99FF95959CFF60FC148F7747D4" box="[166,516,956,980]" gridcol="0" gridrow="19" pageId="3" pageNumber="95">Interorbital width</th>
<td id="76D95D99FF95959CFD5CFC148F8047D4" box="[666,755,956,980]" gridcol="1" gridrow="19" pageId="3" pageNumber="95">5.6</td>
<td id="76D95D99FF95959CFC0DFC14893947D4" box="[971,1098,956,980]" gridcol="2" gridrow="19" pageId="3" pageNumber="95">5.6–6.9</td>
<td id="76D95D99FF95959CFAD2FC14883E47D4" box="[1300,1357,956,980]" gridcol="3" gridrow="19" pageId="3" pageNumber="95">6.3</td>
</tr>
<tr id="350834E5FF95959CFF60FC4C883E47FD" box="[166,1357,996,1021]" gridrow="20" pageId="3" pageNumber="95">
<th id="76D95D99FF95959CFF60FC4C8F7747FD" box="[166,516,996,1021]" gridcol="0" gridrow="20" pageId="3" pageNumber="95">Length of dorsal fin</th>
<td id="76D95D99FF95959CFD5CFC4C8F8047FD" box="[666,755,996,1021]" gridcol="1" gridrow="20" pageId="3" pageNumber="95">20.4</td>
<td id="76D95D99FF95959CFC0DFC4C893947FD" box="[971,1098,996,1021]" gridcol="2" gridrow="20" pageId="3" pageNumber="95">19.5–22.4</td>
<td id="76D95D99FF95959CFAD2FC4C883E47FD" box="[1300,1357,996,1021]" gridcol="3" gridrow="20" pageId="3" pageNumber="95">20.5</td>
</tr>
<tr id="350834E5FF95959CFF60FBA5883E4025" box="[166,1357,1037,1061]" gridrow="21" pageId="3" pageNumber="95">
<th id="76D95D99FF95959CFF60FBA58F774025" box="[166,516,1037,1061]" gridcol="0" gridrow="21" pageId="3" pageNumber="95">Length of upper caudal-fin lobe</th>
<td id="76D95D99FF95959CFD5CFBA58F804025" box="[666,755,1037,1061]" gridcol="1" gridrow="21" pageId="3" pageNumber="95">32.6</td>
<td id="76D95D99FF95959CFC0DFBA589394025" box="[971,1098,1037,1061]" gridcol="2" gridrow="21" pageId="3" pageNumber="95">27.1–32.6</td>
<td id="76D95D99FF95959CFAD2FBA5883E4025" box="[1300,1357,1037,1061]" gridcol="3" gridrow="21" pageId="3" pageNumber="95">30.4</td>
</tr>
<tr id="350834E5FF95959CFF60FB9D883E404E" box="[166,1357,1077,1102]" gridrow="22" pageId="3" pageNumber="95">
<th id="76D95D99FF95959CFF60FB9D8F77404E" box="[166,516,1077,1102]" gridcol="0" gridrow="22" pageId="3" pageNumber="95">Length of median caudal-fin rays</th>
<td id="76D95D99FF95959CFD5CFB9D8F80404E" box="[666,755,1077,1102]" gridcol="1" gridrow="22" pageId="3" pageNumber="95">15.3</td>
<td id="76D95D99FF95959CFC0DFB9D8939404E" box="[971,1098,1077,1102]" gridcol="2" gridrow="22" pageId="3" pageNumber="95">13.6–16.6</td>
<td id="76D95D99FF95959CFAD2FB9D883E404E" box="[1300,1357,1077,1102]" gridcol="3" gridrow="22" pageId="3" pageNumber="95">15.1</td>
</tr>
<tr id="350834E5FF95959CFF60FBF5883E4075" box="[166,1357,1117,1141]" gridrow="23" pageId="3" pageNumber="95">
<th id="76D95D99FF95959CFF60FBF58F774075" box="[166,516,1117,1141]" gridcol="0" gridrow="23" pageId="3" pageNumber="95">Length of lower caudal-fin lobe</th>
<td id="76D95D99FF95959CFD5CFBF58F804075" box="[666,755,1117,1141]" gridcol="1" gridrow="23" pageId="3" pageNumber="95">26.5</td>
<td id="76D95D99FF95959CFC0DFBF589394075" box="[971,1098,1117,1141]" gridcol="2" gridrow="23" pageId="3" pageNumber="95">22.3–29.6</td>
<td id="76D95D99FF95959CFAD2FBF5883E4075" box="[1300,1357,1117,1141]" gridcol="3" gridrow="23" pageId="3" pageNumber="95">25.6</td>
</tr>
<tr id="350834E5FF95959CFF60FB2D883E409E" box="[166,1357,1157,1182]" gridrow="24" pageId="3" pageNumber="95">
<th id="76D95D99FF95959CFF60FB2D8F77409E" box="[166,516,1157,1182]" gridcol="0" gridrow="24" pageId="3" pageNumber="95">Length of anal fin</th>
<td id="76D95D99FF95959CFD5CFB2D8F80409E" box="[666,755,1157,1182]" gridcol="1" gridrow="24" pageId="3" pageNumber="95">16.1</td>
<td id="76D95D99FF95959CFC0DFB2D8939409E" box="[971,1098,1157,1182]" gridcol="2" gridrow="24" pageId="3" pageNumber="95">15.5–17.8</td>
<td id="76D95D99FF95959CFAD2FB2D883E409E" box="[1300,1357,1157,1182]" gridcol="3" gridrow="24" pageId="3" pageNumber="95">16.5</td>
</tr>
<tr id="350834E5FF95959CFF60FB06883E40C6" box="[166,1357,1198,1222]" gridrow="25" pageId="3" pageNumber="95">
<th id="76D95D99FF95959CFF60FB068F7740C6" box="[166,516,1198,1222]" gridcol="0" gridrow="25" pageId="3" pageNumber="95">Length of pelvic fin</th>
<td id="76D95D99FF95959CFD5CFB068F8040C6" box="[666,755,1198,1222]" gridcol="1" gridrow="25" pageId="3" pageNumber="95">16.3</td>
<td id="76D95D99FF95959CFC0DFB06893940C6" box="[971,1098,1198,1222]" gridcol="2" gridrow="25" pageId="3" pageNumber="95">13.9–16.8</td>
<td id="76D95D99FF95959CFAD2FB06883E40C6" box="[1300,1357,1198,1222]" gridcol="3" gridrow="25" pageId="3" pageNumber="95">15.6</td>
</tr>
<tr id="350834E5FF95959CFF60FB7E883E40EE" box="[166,1357,1238,1262]" gridrow="26" pageId="3" pageNumber="95">
<th id="76D95D99FF95959CFF60FB7E8F7740EE" box="[166,516,1238,1262]" gridcol="0" gridrow="26" pageId="3" pageNumber="95">Length of pectoral fin</th>
<td id="76D95D99FF95959CFD5CFB7E8F8040EE" box="[666,755,1238,1262]" gridcol="1" gridrow="26" pageId="3" pageNumber="95">22.8</td>
<td id="76D95D99FF95959CFC0DFB7E893940EE" box="[971,1098,1238,1262]" gridcol="2" gridrow="26" pageId="3" pageNumber="95">18.0–23.9</td>
<td id="76D95D99FF95959CFAD2FB7E883E40EE" box="[1300,1357,1238,1262]" gridcol="3" gridrow="26" pageId="3" pageNumber="95">20.1</td>
</tr>
<tr id="350834E5FF95959CFF60FB56883E4116" box="[166,1357,1278,1302]" gridrow="27" pageId="3" pageNumber="95">
<th id="76D95D99FF95959CFF60FB56883E4116" box="[166,1357,1278,1302]" colspan="4" colspanRight="3" gridcol="0" gridrow="27" pageId="3" pageNumber="95">In percent of dorsal head length</th>
</tr>
<tr id="350834E5FF95959CFF60FA8E883E413F" box="[166,1357,1318,1343]" gridrow="28" pageId="3" pageNumber="95">
<th id="76D95D99FF95959CFF60FA8E8F77413F" box="[166,516,1318,1343]" gridcol="0" gridrow="28" pageId="3" pageNumber="95">Snout length</th>
<td id="76D95D99FF95959CFD5CFA8E8F80413F" box="[666,755,1318,1343]" gridcol="1" gridrow="28" pageId="3" pageNumber="95">46</td>
<td id="76D95D99FF95959CFC0DFA8E8939413F" box="[971,1098,1318,1343]" gridcol="2" gridrow="28" pageId="3" pageNumber="95">40–46</td>
<td id="76D95D99FF95959CFAD2FA8E883E413F" box="[1300,1357,1318,1343]" gridcol="3" gridrow="28" pageId="3" pageNumber="95">43</td>
</tr>
<tr id="350834E5FF95959CFF60FAE7883E4167" box="[166,1357,1359,1383]" gridrow="29" pageId="3" pageNumber="95">
<th id="76D95D99FF95959CFF60FAE78F774167" box="[166,516,1359,1383]" gridcol="0" gridrow="29" pageId="3" pageNumber="95">Eye diameter</th>
<td id="76D95D99FF95959CFD5CFAE78F804167" box="[666,755,1359,1383]" gridcol="1" gridrow="29" pageId="3" pageNumber="95">27</td>
<td id="76D95D99FF95959CFC0DFAE789394167" box="[971,1098,1359,1383]" gridcol="2" gridrow="29" pageId="3" pageNumber="95">22–29</td>
<td id="76D95D99FF95959CFAD2FAE7883E4167" box="[1300,1357,1359,1383]" gridcol="3" gridrow="29" pageId="3" pageNumber="95">26</td>
</tr>
<tr id="350834E5FF95959CFF60FADF883E418F" box="[166,1357,1399,1423]" gridrow="30" pageId="3" pageNumber="95">
<th id="76D95D99FF95959CFF60FADF8F77418F" box="[166,516,1399,1423]" gridcol="0" gridrow="30" pageId="3" pageNumber="95">Interorbital width</th>
<td id="76D95D99FF95959CFD5CFADF8F80418F" box="[666,755,1399,1423]" gridcol="1" gridrow="30" pageId="3" pageNumber="95">30</td>
<td id="76D95D99FF95959CFC0DFADF8939418F" box="[971,1098,1399,1423]" gridcol="2" gridrow="30" pageId="3" pageNumber="95">28–36</td>
<td id="76D95D99FF95959CFAD2FADF883E418F" box="[1300,1357,1399,1423]" gridcol="3" gridrow="30" pageId="3" pageNumber="95">33</td>
</tr>
<tr id="350834E5FF95959CFF60FA37883E41B7" box="[166,1357,1439,1463]" gridrow="31" pageId="3" pageNumber="95">
<th id="76D95D99FF95959CFF60FA37883E41B7" box="[166,1357,1439,1463]" colspan="4" colspanRight="3" gridcol="0" gridrow="31" pageId="3" pageNumber="95">In percent of lateral head length</th>
</tr>
<tr id="350834E5FF95959CFF60FA6F883E41E0" box="[166,1357,1479,1504]" gridrow="32" pageId="3" pageNumber="95">
<th id="76D95D99FF95959CFF60FA6F8F7741E0" box="[166,516,1479,1504]" gridcol="0" gridrow="32" pageId="3" pageNumber="95">Snout length</th>
<td id="76D95D99FF95959CFD5CFA6F8F8041E0" box="[666,755,1479,1504]" gridcol="1" gridrow="32" pageId="3" pageNumber="95">40</td>
<td id="76D95D99FF95959CFC0DFA6F893941E0" box="[971,1098,1479,1504]" gridcol="2" gridrow="32" pageId="3" pageNumber="95">36–40</td>
<td id="76D95D99FF95959CFAD2FA6F883E41E0" box="[1300,1357,1479,1504]" gridcol="3" gridrow="32" pageId="3" pageNumber="95">38</td>
</tr>
<tr id="350834E5FF95959CFF60FA58883E4208" box="[166,1357,1520,1544]" gridrow="33" pageId="3" pageNumber="95">
<th id="76D95D99FF95959CFF60FA588F774208" box="[166,516,1520,1544]" gridcol="0" gridrow="33" pageId="3" pageNumber="95">Eye diameter</th>
<td id="76D95D99FF95959CFD5CFA588F804208" box="[666,755,1520,1544]" gridcol="1" gridrow="33" pageId="3" pageNumber="95">23</td>
<td id="76D95D99FF95959CFC0DFA5889394208" box="[971,1098,1520,1544]" gridcol="2" gridrow="33" pageId="3" pageNumber="95">20–27</td>
<td id="76D95D99FF95959CFAD2FA58883E4208" box="[1300,1357,1520,1544]" gridcol="3" gridrow="33" pageId="3" pageNumber="95">23</td>
</tr>
<tr id="350834E5FF95959CFF60F9B0883E4230" box="[166,1357,1560,1584]" gridrow="34" pageId="3" pageNumber="95">
<th id="76D95D99FF95959CFF60F9B08F774230" box="[166,516,1560,1584]" gridcol="0" gridrow="34" pageId="3" pageNumber="95">Interorbital width</th>
<td id="76D95D99FF95959CFD5CF9B08F804230" box="[666,755,1560,1584]" gridcol="1" gridrow="34" pageId="3" pageNumber="95">26</td>
<td id="76D95D99FF95959CFC0DF9B089394230" box="[971,1098,1560,1584]" gridcol="2" gridrow="34" pageId="3" pageNumber="95">24–32</td>
<td id="76D95D99FF95959CFAD2F9B0883E4230" box="[1300,1357,1560,1584]" gridcol="3" gridrow="34" pageId="3" pageNumber="95">29</td>
</tr>
</table>
</paragraph>
<paragraph id="8B8736A7FF956A60FF01F9F7882C4279" blockId="3.[151,1437,1631,2018]" box="[199,1375,1631,1658]" pageId="3" pageNumber="95">Intestine with a loop shortly behind stomach. Air bladder without posterior chamber in abdominal cavity.</paragraph>
<paragraph id="8B8736A7FF956A60FF01F92B89ED43E1" blockId="3.[151,1437,1631,2018]" pageId="3" pageNumber="95">
<emphasis id="B94CEAB5FF956A60FF01F92B8CC9429D" bold="true" box="[199,442,1667,1693]" pageId="3" pageNumber="95">Sexual dimorphism.</emphasis>
Males distinguishable by presence (vs. absence) of a suborbital flap (modified lateral ethmoid; 
<figureCitation id="13032A22FF956A60FEC4F90F8C2042C2" box="[258,339,1703,1730]" captionStart="FIGURE 4" captionStartId="4.[151,250,1972,1996]" captionTargetBox="[151,1435,1684,1948]" captionTargetId="figure-67@4.[151,1435,1684,1948]" captionTargetPageId="4" captionText="FIGURE 4. Nemacheilus pezidion, MHNG 2787.088, holotype, 57.8 mm SL, male: a, nostril and suborbital flap; and b, flank with small tubercles on scales." figureDoi="http://doi.org/10.5281/zenodo.6488214" httpUri="https://zenodo.org/record/6488214/files/figure.png" pageId="3" pageNumber="95">Fig. 4a</figureCitation>
), with a few tiny tubercles at extremity and on inner surface. In males, dorsal surface of branched pectoral-fin rays 1–6 with densely-set tubercles on ray itself and on fleshy pad extending along its posterior margin (only behind rays 1–3); on last rays, tubercles restricted to a single row along proximal part of ray (
<figureCitation id="13032A22FF956A60FADCF947881D430A" box="[1306,1390,1775,1802]" captionStart="FIGURE 2" captionStartId="4.[151,250,744,768]" captionTargetBox="[151,1403,181,719]" captionTargetId="figure-16@4.[151,1435,181,719]" captionTargetPageId="4" captionText="FIGURE 2. Nemacheilus pezidion, pectoral fin in dorsal view of: a, male, MHNG 2787.088, holotype, 57.8 mm SL (note small tubercles on branched rays 1–6); and b, female, CMK 21246, 52.9 mm SL." figureDoi="http://doi.org/10.5281/zenodo.6488212" httpUri="https://zenodo.org/record/6488212/files/figure.png" pageId="3" pageNumber="95">Fig. 2a</figureCitation>
). In females, fin somewhat shorter, no tubercles (
<figureCitation id="13032A22FF956A60FD41F8BB8FA9432E" box="[647,730,1811,1838]" captionStart="FIGURE 2" captionStartId="4.[151,250,744,768]" captionTargetBox="[151,1403,181,719]" captionTargetId="figure-16@4.[151,1435,181,719]" captionTargetPageId="4" captionText="FIGURE 2. Nemacheilus pezidion, pectoral fin in dorsal view of: a, male, MHNG 2787.088, holotype, 57.8 mm SL (note small tubercles on branched rays 1–6); and b, female, CMK 21246, 52.9 mm SL." figureDoi="http://doi.org/10.5281/zenodo.6488212" httpUri="https://zenodo.org/record/6488212/files/figure.png" pageId="3" pageNumber="95">Fig. 2b</figureCitation>
). In males, a line of black pigments along anterior and posterior edge of branched ray 1 and behind branched rays 2 and 3; in large females, black pigments on branched ray 1 and on membranes between it and adjacent rays. Tubercles on first branched ray already visible in smallest available male (CMK 21315, 
<quantity id="4CC09B42FF956A60FEFBF8288CD1439A" box="[317,418,1920,1946]" metricMagnitude="-2" metricUnit="m" metricValue="3.92" pageId="3" pageNumber="95" unit="mm" value="39.2">39.2 mm</quantity>
SL). In males, a patch of scales with minute tubercles on flank, longitudinally from about tip of pectoral fin to above anal-fin base, on lateral line scales, and on about 3 rows of scales above and about ⅔ of flank below lateral line (
<figureCitation id="13032A22FF956A60FE6FF86F8C8D43E2" box="[425,510,1991,2018]" captionStart="FIGURE 4" captionStartId="4.[151,250,1972,1996]" captionTargetBox="[151,1435,1684,1948]" captionTargetId="figure-67@4.[151,1435,1684,1948]" captionTargetPageId="4" captionText="FIGURE 4. Nemacheilus pezidion, MHNG 2787.088, holotype, 57.8 mm SL, male: a, nostril and suborbital flap; and b, flank with small tubercles on scales." figureDoi="http://doi.org/10.5281/zenodo.6488214" httpUri="https://zenodo.org/record/6488214/files/figure.png" pageId="3" pageNumber="95">Fig. 4b</figureCitation>
); tubercles missing in females. Ripe females deeper bodied.
</paragraph>
<caption id="DF47662FFF926A67FF51FD408EE34724" ID-DOI="http://doi.org/10.5281/zenodo.6488212" ID-Zenodo-Dep="6488212" httpUri="https://zenodo.org/record/6488212/files/figure.png" pageId="4" pageNumber="96" startId="4.[151,250,744,768]" targetBox="[151,1403,181,719]" targetPageId="4">
<paragraph id="8B8736A7FF926A67FF51FD408EE34724" blockId="4.[151,1436,744,804]" pageId="4" pageNumber="96">
<emphasis id="B94CEAB5FF926A67FF51FD408C614700" bold="true" box="[151,274,744,768]" pageId="4" pageNumber="96">FIGURE 2.</emphasis>
<taxonomicName id="4C384D24FF926A67FEDEFD408C844700" authorityName="Kottelat" authorityYear="2022" box="[280,503,744,768]" class="Actinopterygii" family="Nemacheilidae" genus="Nemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="96" phylum="Chordata" rank="species" species="pezidion">
<emphasis id="B94CEAB5FF926A67FEDEFD408C844700" box="[280,503,744,768]" italics="true" pageId="4" pageNumber="96">Nemacheilus pezidion</emphasis>
</taxonomicName>
, pectoral fin in dorsal view of: 
<emphasis id="B94CEAB5FF926A67FCF7FD408E374700" bold="true" box="[817,836,744,768]" pageId="4" pageNumber="96">a,</emphasis>
male, MHNG 2787.088, holotype, 57.8 mm SL (note small tubercles on branched rays 1–6); and 
<emphasis id="B94CEAB5FF926A67FDD5FCA48F544724" bold="true" box="[531,551,780,804]" pageId="4" pageNumber="96">b,</emphasis>
female, CMK 21246, 52.9 mm SL.
</paragraph>
</caption>
<caption id="DF47662FFF926A67FF51F9CC8E16427C" ID-DOI="http://doi.org/10.5281/zenodo.6502469" ID-Zenodo-Dep="6502469" box="[151,869,1636,1660]" httpUri="https://zenodo.org/record/6502469/files/figure.png" pageId="4" pageNumber="96" startId="4.[151,250,1636,1660]" targetBox="[297,1291,828,1612]" targetPageId="4">
<paragraph id="8B8736A7FF926A67FF51F9CC8E16427C" blockId="4.[151,869,1636,1660]" box="[151,869,1636,1660]" pageId="4" pageNumber="96">
<emphasis id="B94CEAB5FF926A67FF51F9CC8C60427C" bold="true" box="[151,275,1636,1660]" pageId="4" pageNumber="96">FIGURE 3.</emphasis>
<taxonomicName id="4C384D24FF926A67FEDFF9CC8F4C427C" authority=", CMK" authorityName="CMK" box="[281,575,1636,1660]" class="Actinopterygii" family="Nemacheilidae" genus="Nemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="96" phylum="Chordata" rank="species" species="pezidion">
<emphasis id="B94CEAB5FF926A67FEDFF9CC8C8A427C" box="[281,505,1636,1660]" italics="true" pageId="4" pageNumber="96">Nemacheilus pezidion</emphasis>
, CMK
</taxonomicName>
21246, 52.9 mm SL; mouth.
</paragraph>
</caption>
<caption id="DF47662FFF926A67FF51F81C8CBE43F0" ID-DOI="http://doi.org/10.5281/zenodo.6488214" ID-Zenodo-Dep="6488214" httpUri="https://zenodo.org/record/6488214/files/figure.png" pageId="4" pageNumber="96" startId="4.[151,250,1972,1996]" targetBox="[151,1435,1684,1948]" targetPageId="4">
<paragraph id="8B8736A7FF926A67FF51F81C8CBE43F0" blockId="4.[151,1436,1972,2032]" pageId="4" pageNumber="96">
<emphasis id="B94CEAB5FF926A67FF51F81C8C6043CC" bold="true" box="[151,275,1972,1996]" pageId="4" pageNumber="96">FIGURE 4.</emphasis>
<taxonomicName id="4C384D24FF926A67FEDCF81C8F2043CD" authority=", MHNG" authorityName="MHNG" box="[282,595,1972,1997]" class="Actinopterygii" family="Nemacheilidae" genus="Nemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="96" phylum="Chordata" rank="species" species="pezidion">
<emphasis id="B94CEAB5FF926A67FEDCF81C8C8A43CC" box="[282,505,1972,1996]" italics="true" pageId="4" pageNumber="96">Nemacheilus pezidion</emphasis>
, MHNG
</taxonomicName>
2787.088, holotype, 57.8 mm SL, male: 
<emphasis id="B94CEAB5FF926A67FC33F81D897B43CD" bold="true" box="[1013,1032,1973,1997]" pageId="4" pageNumber="96">a,</emphasis>
nostril and suborbital flap; and 
<emphasis id="B94CEAB5FF926A67FA8BF81D881243CD" bold="true" box="[1357,1377,1973,1997]" pageId="4" pageNumber="96">b,</emphasis>
flank with small tubercles on scales.
</paragraph>
</caption>
<paragraph id="8B8736A7FF936A66FF01FF3E883A4589" blockId="5.[151,1437,150,1006]" pageId="5" pageNumber="97">
<emphasis id="B94CEAB5FF936A66FF01FF3E8C3C44B0" bold="true" box="[199,335,150,176]" pageId="5" pageNumber="97">Coloration.</emphasis>
Preserved: Upper half of head and body background colour pale olive brown, lower half yellowish; unless otherwise stated, markings dark brown to black. Two transverse lines in interorbital area and on posterior part of head. One spot between tip of snout and nostril. Two spots between nostrils. A midlateral stripe from base of outer rostral barbel to eye and to upper part of opercle. Body with a row of 11–15 narrow saddles along back (4–5 predorsal, 2–3 subdorsal, 5–7 postdorsal); saddles of regular width and shape, sometimes paired or with paler central area. A stripe from upper extremity of gill opening to end of caudal peduncle; anteriorly as a very regular band of pigments of same intensity, width about equal to eye diameter; posteriorly becoming narrower, less regular.
</paragraph>
<paragraph id="8B8736A7FF936A66FF01FE3B8EBA45F5" blockId="5.[151,1437,150,1006]" pageId="5" pageNumber="97">Pattern at caudal-fin base: A conspicuous black spot at base of fin, slightly lower than middle, extending on lower 2 rays of upper lobe and upper 6 rays of lower lobe. Spot aligned with midlateral stripe, separated from it by narrow unpigmented patch, black, darker than rest of patterning on body.</paragraph>
<paragraph id="8B8736A7FF936A66FF01FE578D9846CE" blockId="5.[151,1437,150,1006]" pageId="5" pageNumber="97">Dorsal fin hyaline, with: an elongate spot on lower ¼ of first branched ray and adjacent membranes; upper half of last unbranched ray black; 2 or 3 irregular rows of spots, usually at branching points of rays; membranes hyaline. Caudal fin hyaline with dusky membranes near base, or with a few spots on median rays, or 1 or 2 vague vertical rows of spots corresponding to pigments on rays approximately at branching points. Anal and pelvic fins hyaline. Pectoral fin hyaline, dark band on first and second branched rays (see more details under sexual dimorphism, above).</paragraph>
<paragraph id="8B8736A7FF936A66FF01FD7F8F1346F1" blockId="5.[151,1437,150,1006]" box="[199,608,727,753]" pageId="5" pageNumber="97">Colour pattern of juvenile unknown.</paragraph>
<paragraph id="8B8736A7FF936A66FF01FD53880F4715" blockId="5.[151,1437,150,1006]" box="[199,1404,763,789]" pageId="5" pageNumber="97">In life: yellowish, patterning dark brown; spot at caudal-fin base black, contrasting with pale area around it.</paragraph>
<paragraph id="8B8736A7FF936A66FF01FCB78E9A475E" blockId="5.[151,1437,150,1006]" pageId="5" pageNumber="97">
<emphasis id="B94CEAB5FF936A66FF01FCB78CE04738" bold="true" box="[199,403,798,825]" pageId="5" pageNumber="97">Notes on biology.</emphasis>
A dissected female (CMK 21330, 
<quantity id="4CC09B42FF936A66FCDCFCB68E0C4739" box="[794,895,798,825]" metricMagnitude="-2" metricUnit="m" metricValue="5.36" pageId="5" pageNumber="97" unit="mm" value="53.6">53.6 mm</quantity>
SL) had unripe ovaries with eggs about 
<quantity id="4CC09B42FF936A66FA8EFCB78DB0475D" metricMagnitude="-4" metricUnit="m" metricValue="6.499999999999999" metricValueMax="7.0" metricValueMin="6.0" pageId="5" pageNumber="97" unit="mm" value="0.6499999999999999" valueMax="0.7" valueMin="0.6">0.6–0.7 mm</quantity>
diameter. Its stomach contained unidentifiable material (possibly mud).
</paragraph>
</subSubSection>
<subSubSection id="C322652CFF936A66FF01FCCE8F5D47EE" pageId="5" pageNumber="97" type="distribution">
<paragraph id="8B8736A7FF936A66FF01FCCE8F5D47EE" blockId="5.[151,1437,150,1006]" pageId="5" pageNumber="97">
<emphasis id="B94CEAB5FF936A66FF01FCCE8C124780" bold="true" box="[199,353,870,896]" pageId="5" pageNumber="97">Distribution.</emphasis>
<taxonomicName id="4C384D24FF936A66FEA1FCCF8F2E4781" authority="Kottelat, 2022" box="[359,605,871,897]" class="Actinopterygii" family="Nemacheilidae" genus="Nemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="97" phylum="Chordata" rank="species" species="pezidion">
<emphasis id="B94CEAB5FF936A66FEA1FCCF8F2E4781" box="[359,605,871,897]" italics="true" pageId="5" pageNumber="97">Nemacheilus pezidion</emphasis>
</taxonomicName>
has been observed only in the Xe Pian, a tributary of Xe Kong, flowing on the southern slope of the Bolaven plateau (
<figureCitation id="13032A22FF936A66FDB1FC238FCF47A5" box="[631,700,906,933]" captionStart="FIGURE 5" captionStartId="5.[151,250,1894,1918]" captionTargetBox="[170,1414,1029,1867]" captionTargetId="figure-417@5.[170,1417,1029,1870]" captionTargetPageId="5" captionText="FIGURE 5. Southern Laos, showing localities of examined material of Nemacheilus pezidion (circles) and ‘N’. argyrogaster (triangles). Each symbol may represent several localities. White circle is type locality of N. pezidion. Dams, reservoirs and other alterations of original hydrographic network are not shown." figureDoi="http://doi.org/10.5281/zenodo.6488216" httpUri="https://zenodo.org/record/6488216/files/figure.png" pageId="5" pageNumber="97">Fig. 5</figureCitation>
). All localities are in the floodplain; at the time of collecting (end of dry season–beginning of wet season), the water of the Xe Pian was turbid at the sampling sites, with the bottom varying from gravel to mud (
<figureCitation id="13032A22FF936A66FE1AFC7B8F5347EE" box="[476,544,979,1006]" captionStart="FIGURE 6" captionStartId="6.[151,250,1395,1419]" captionTargetBox="[159,1429,669,1372]" captionTargetId="figure-255@6.[159,1429,669,1372]" captionTargetPageId="6" captionText="FIGURE 6. Laos: Xe Kong watershed: Houai Pin, tributary of Xe Pian, entering it about 5 km downstream of Ban Mai, 21 May 2009, locality of Nemacheilus pezidion, CMK 21330." figureDoi="http://doi.org/10.5281/zenodo.6488218" httpUri="https://zenodo.org/record/6488218/files/figure.png" pageId="5" pageNumber="97">Fig. 6</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="DF47662FFF936A66FF51F8CE8F8743C6" ID-DOI="http://doi.org/10.5281/zenodo.6488216" ID-Zenodo-Dep="6488216" httpUri="https://zenodo.org/record/6488216/files/figure.png" pageId="5" pageNumber="97" startId="5.[151,250,1894,1918]" targetBox="[170,1414,1029,1867]" targetPageId="5">
<paragraph id="8B8736A7FF936A66FF51F8CE8F8743C6" blockId="5.[151,1436,1894,1990]" pageId="5" pageNumber="97">
<emphasis id="B94CEAB5FF936A66FF51F8CE8C67437E" bold="true" box="[151,276,1894,1918]" pageId="5" pageNumber="97">FIGURE 5.</emphasis>
Southern Laos, showing localities of examined material of 
<taxonomicName id="4C384D24FF936A66FCBAF8CE892F437E" box="[892,1116,1894,1918]" class="Actinopterygii" family="Nemacheilidae" genus="Nemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="97" phylum="Chordata" rank="species" species="pezidion">
<emphasis id="B94CEAB5FF936A66FCBAF8CE892F437E" box="[892,1116,1894,1918]" italics="true" pageId="5" pageNumber="97">Nemacheilus pezidion</emphasis>
</taxonomicName>
(circles) and 
<emphasis id="B94CEAB5FF936A66FB2CF8CE88EF437E" box="[1258,1436,1894,1918]" italics="true" pageId="5" pageNumber="97">‘N’. argyrogaster</emphasis>
(triangles). Each symbol may represent several localities. White circle is type locality of 
<taxonomicName id="4C384D24FF936A66FBC9F82389F143A2" box="[1039,1154,1930,1954]" class="Actinopterygii" family="Nemacheilidae" genus="Nemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="97" phylum="Chordata" rank="species" species="pezidion">
<emphasis id="B94CEAB5FF936A66FBC9F82389F143A2" box="[1039,1154,1930,1954]" italics="true" pageId="5" pageNumber="97">N. pezidion</emphasis>
</taxonomicName>
. Dams, reservoirs and other alterations of original hydrographic network are not shown.
</paragraph>
</caption>
<subSubSection id="C322652CFF906A65FF01FF3F8CFA44D5" pageId="6" pageNumber="98" type="etymology">
<paragraph id="8B8736A7FF906A65FF01FF3F8CFA44D5" blockId="6.[151,1437,150,645]" pageId="6" pageNumber="98">
<emphasis id="B94CEAB5FF906A65FF01FF3F8C3D44B1" bold="true" box="[199,334,151,177]" pageId="6" pageNumber="98">Etymology.</emphasis>
From the ancient Greek 
<emphasis id="B94CEAB5FF906A65FDA2FF3E8FCC44B1" box="[612,703,150,177]" italics="true" pageId="6" pageNumber="98">πεζίΔΙΟΝ</emphasis>
(pezidion), a ribbon, reference to the black stripe along the flank. A noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="C322652CFF906A69FF01FF778FDB4739" lastPageId="10" lastPageNumber="102" pageId="6" pageNumber="98" type="discussion">
<paragraph id="8B8736A7FF906A65FF01FF7789B54685" blockId="6.[151,1437,150,645]" pageId="6" pageNumber="98">
<emphasis id="B94CEAB5FF906A65FF01FF778C4844F9" bold="true" box="[199,315,223,249]" pageId="6" pageNumber="98">Remarks.</emphasis>
As mentioned in the diagnosis, 
<taxonomicName id="4C384D24FF906A65FD51FF778E6744F9" authority="Kottelat, 2022" box="[663,788,223,249]" class="Actinopterygii" family="Nemacheilidae" genus="Nemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="98" phylum="Chordata" rank="species" species="pezidion">
<emphasis id="B94CEAB5FF906A65FD51FF778E6744F9" box="[663,788,223,249]" italics="true" pageId="6" pageNumber="98">N. pezidion</emphasis>
</taxonomicName>
is distinguished from most other species of 
<taxonomicName id="4C384D24FF906A65FB32FF7788EF44F9" box="[1268,1436,223,249]" class="Actinopterygii" family="Nemacheilidae" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="98" phylum="Chordata" rank="family">Nemacheilidae</taxonomicName>
in Southeast Asia in having an obvious dark brown to black midlateral stripe. A somewhat similar colour pattern is observed in 
<taxonomicName id="4C384D24FF906A65FE85FE8F8CBC4541" box="[323,463,295,321]" class="Actinopterygii" family="Nemacheilidae" genus="Nemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="98" phylum="Chordata" rank="species" species="binotatus">
<emphasis id="B94CEAB5FF906A65FE85FE8F8CBC4541" box="[323,463,295,321]" italics="true" pageId="6" pageNumber="98">N. binotatus</emphasis>
</taxonomicName>
(
<figureCitation id="13032A22FF906A65FE24FE8F8F5B4541" box="[482,552,295,321]" captionStart="FIGURE 7" captionStartId="6.[151,250,1863,1887]" captionTargetBox="[151,1436,1479,1840]" captionTargetId="figure-289@6.[151,1436,1479,1840]" captionTargetPageId="6" captionText="FIGURE 7. Nemacheilus binotatus, CMK 16044, 34.6 mm SL; Thailand: Chao Phraya drainage: Nan River." figureDoi="http://doi.org/10.5281/zenodo.6488220" httpUri="https://zenodo.org/record/6488220/files/figure.png" pageId="6" pageNumber="98">Fig. 7</figureCitation>
) known from the Chao Phraya and Mae Khlong drainages in 
<collectingCountry id="F32F7637FF906A65FB3AFE8F88104541" box="[1276,1379,295,321]" name="Thailand" pageId="6" pageNumber="98">Thailand</collectingCountry>
. The two species are distinguished by a forked caudal fin with the upper lobe distinctly longer than the lower one in 
<taxonomicName id="4C384D24FF906A65FA45FEE38D854589" authority="Kottelat, 2022" class="Actinopterygii" family="Nemacheilidae" genus="Nemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="98" phylum="Chordata" rank="species" species="pezidion">
<emphasis id="B94CEAB5FF906A65FA45FEE38D854589" italics="true" pageId="6" pageNumber="98">N. pezidion</emphasis>
</taxonomicName>
(vs. subequal in 
<taxonomicName id="4C384D24FF906A65FE7EFEC78F324589" box="[440,577,367,393]" class="Actinopterygii" family="Nemacheilidae" genus="Nemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="98" phylum="Chordata" rank="species" species="binotatus">
<emphasis id="B94CEAB5FF906A65FE7EFEC78F324589" box="[440,577,367,393]" italics="true" pageId="6" pageNumber="98">N. binotatus</emphasis>
</taxonomicName>
), the pectoral-fin rays with filamentous extensions (vs. without extensions), the mid-dorsal series of saddles (vs. a black stripe from top of head to caudal-fin base), the presence of an elongate spot on lower ¼ of first branched dorsal-fin ray and adjacent membranes (vs. absence) and the absence of small vertically elongated spots above and in contact with the midlateral stripe (vs. presence of one, rarely two or three, spots at about level of the tip of the pectoral fin). Further, male 
<taxonomicName id="4C384D24FF906A65FC4CFE57897E4619" authority="Kottelat, 2022" box="[906,1037,511,537]" class="Actinopterygii" family="Nemacheilidae" genus="Nemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="98" phylum="Chordata" rank="species" species="pezidion">
<emphasis id="B94CEAB5FF906A65FC4CFE57897E4619" box="[906,1037,511,537]" italics="true" pageId="6" pageNumber="98">N. pezidion</emphasis>
</taxonomicName>
have a modified pectoral fin with conspicuous tubercles on anterior rays and wide unculiferous pads (vs. minute tubercles, narrow unculiferous pads), and numerous small tubercles on the middle and posterior part of the flank of large males (vs. without tubercles). Data on 
<taxonomicName id="4C384D24FF906A65FF3EFDC38CF74685" box="[248,388,619,645]" class="Actinopterygii" family="Nemacheilidae" genus="Nemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="98" phylum="Chordata" rank="species" species="binotatus">
<emphasis id="B94CEAB5FF906A65FF3EFDC38CF74685" box="[248,388,619,645]" italics="true" pageId="6" pageNumber="98">N. binotatus</emphasis>
</taxonomicName>
are from 
<bibRefCitation id="EFA94B56FF906A65FE33FDC38FD64685" author="Kottelat, M." box="[501,677,619,645]" pageId="6" pageNumber="98" refId="ref6499" refString="Kottelat, M. (1990) Indochinese nemacheilines. A revision of nemacheiline loaches (Pisces: Cypriniformes) of Thailand, Burma, Laos, Cambodia and southern Viet Nam. Pfeil, Munchen, 262 pp." type="book" year="1990">Kottelat (1990)</bibRefCitation>
and re-examination of specimens cited therein.
</paragraph>
<caption id="DF47662FFF906A65FF51FADB8FC441AF" ID-DOI="http://doi.org/10.5281/zenodo.6488218" ID-Zenodo-Dep="6488218" httpUri="https://zenodo.org/record/6488218/files/figure.png" pageId="6" pageNumber="98" startId="6.[151,250,1395,1419]" targetBox="[159,1429,669,1372]" targetPageId="6">
<paragraph id="8B8736A7FF906A65FF51FADB8FC441AF" blockId="6.[151,1436,1395,1455]" pageId="6" pageNumber="98">
<emphasis id="B94CEAB5FF906A65FF51FADB8C61418B" bold="true" box="[151,274,1395,1419]" pageId="6" pageNumber="98">FIGURE 6.</emphasis>
Laos: Xe Kong watershed: Houai Pin, tributary of Xe Pian, entering it about 5 km downstream of Ban Mai, 21 May 2009, locality of 
<taxonomicName id="4C384D24FF906A65FE82FA3F8F1941AF" authority=", CMK" authorityName="CMK" box="[324,618,1431,1455]" class="Actinopterygii" family="Nemacheilidae" genus="Nemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="98" phylum="Chordata" rank="species" species="pezidion">
<emphasis id="B94CEAB5FF906A65FE82FA3F8F5041AF" box="[324,547,1431,1455]" italics="true" pageId="6" pageNumber="98">Nemacheilus pezidion</emphasis>
, CMK
</taxonomicName>
21330.
</paragraph>
</caption>
<caption id="DF47662FFF906A65FF51F8EF899B4360" ID-DOI="http://doi.org/10.5281/zenodo.6488220" ID-Zenodo-Dep="6488220" box="[151,1256,1863,1888]" httpUri="https://zenodo.org/record/6488220/files/figure.png" pageId="6" pageNumber="98" startId="6.[151,250,1863,1887]" targetBox="[151,1436,1479,1840]" targetPageId="6">
<paragraph id="8B8736A7FF906A65FF51F8EF899B4360" blockId="6.[151,1256,1863,1888]" box="[151,1256,1863,1888]" pageId="6" pageNumber="98">
<emphasis id="B94CEAB5FF906A65FF51F8EF8C604360" bold="true" box="[151,275,1863,1888]" pageId="6" pageNumber="98">FIGURE 7.</emphasis>
<taxonomicName id="4C384D24FF906A65FEDFF8E08F3B4360" authority=", CMK" authorityName="CMK" box="[281,584,1864,1888]" class="Actinopterygii" family="Nemacheilidae" genus="Nemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="98" phylum="Chordata" rank="species" species="binotatus">
<emphasis id="B94CEAB5FF906A65FEDFF8E08F724360" box="[281,513,1864,1888]" italics="true" pageId="6" pageNumber="98">Nemacheilus binotatus</emphasis>
, CMK
</taxonomicName>
16044, 34.6 mm SL; Thailand: Chao Phraya drainage: Nan River.
</paragraph>
</caption>
<paragraph id="8B8736A7FF906A64FF01F8268D884661" blockId="6.[151,1437,1934,2032]" lastBlockId="7.[151,1437,151,609]" lastPageId="7" lastPageNumber="99" pageId="6" pageNumber="98">
<taxonomicName id="4C384D24FF906A65FF01F8268CB243A8" authority="Kottelat, 2022" box="[199,449,1934,1960]" class="Actinopterygii" family="Nemacheilidae" genus="Nemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="98" phylum="Chordata" rank="species" species="pezidion">
<emphasis id="B94CEAB5FF906A65FF01F8268CB243A8" box="[199,449,1934,1960]" italics="true" pageId="6" pageNumber="98">Nemacheilus pezidion</emphasis>
</taxonomicName>
had at first been misidentified as 
<taxonomicName id="4C384D24FF906A65FC96F827897243A8" authorityName="Kottelat" authorityYear="1990" box="[848,1025,1934,1960]" class="Actinopterygii" family="Nemacheilidae" genus="Nemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="98" phylum="Chordata" rank="species" species="longistriatus">
<emphasis id="B94CEAB5FF906A65FC96F827897243A8" box="[848,1025,1934,1960]" italics="true" pageId="6" pageNumber="98">N. longistriatus</emphasis>
</taxonomicName>
(
<figureCitation id="13032A22FF906A65FBD5F826892A43A8" box="[1043,1113,1934,1960]" captionStart="FIGURE 8" captionStartId="7.[151,250,1788,1812]" captionTargetBox="[151,1436,633,1765]" captionTargetId="figure-249@7.[151,1436,633,1765]" captionTargetPageId="7" captionText="FIGURE 8. Nemacheilus longistriatus, Mekong drainage: a, CAS 62547, holotype, 53.2 mm SL; Thailand: Mekong at Chiang Khan; b, CMK 23085, 37.5 mm SL; Laos: Xe Bangfai." figureDoi="http://doi.org/10.5281/zenodo.6488224" httpUri="https://zenodo.org/record/6488224/files/figure.png" pageId="6" pageNumber="98">Fig. 8</figureCitation>
), a species known from the Mekong drainage upstream of the Khone falls, because of the presence of the midlateral stripe (
<bibRefCitation id="EFA94B56FF906A65FB2EF81A88E243CC" author="Kottelat, M." box="[1256,1425,1970,1996]" pageId="6" pageNumber="98" refId="ref6664" refString="Kottelat, M. (2009) Fishes of the Xe Kong drainage in Laos. Aquatic resources management to improve rural livelihoods of the Xe Kong basin (ARL Xe Kong Basin Project), WWF Laos, Vientiane, 29 pp." type="book" year="2009">Kottelat, 2009</bibRefCitation>
). The two species are distinguished by a forked caudal fin with the upper lobe distinctly longer than the lower one in 
<taxonomicName id="4C384D24FF916A64FF7EFF3F8C4C44B1" authority="Kottelat, 2022" box="[184,319,151,177]" class="Actinopterygii" family="Nemacheilidae" genus="Nemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="99" phylum="Chordata" rank="species" species="pezidion">
<emphasis id="B94CEAB5FF916A64FF7EFF3F8C4C44B1" box="[184,319,151,177]" italics="true" pageId="7" pageNumber="99">N. pezidion</emphasis>
</taxonomicName>
(vs. subequal in 
<taxonomicName id="4C384D24FF916A64FDD5FF3F8FD144B1" box="[531,674,151,177]" class="Actinopterygii" family="Nemacheilidae" genus="Nemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="99" phylum="Chordata" rank="species" species="binotatus">
<emphasis id="B94CEAB5FF916A64FDD5FF3F8FD144B1" box="[531,674,151,177]" italics="true" pageId="7" pageNumber="99">N. binotatus</emphasis>
</taxonomicName>
), the pectoral-fin rays with filamentous extensions (vs. without extensions), the presence of an elongate spot on lower ¼ of first branched dorsal-fin ray and adjacent membranes (vs. absence). Further, male 
<taxonomicName id="4C384D24FF916A64FE1CFF778F2F44F9" authority="Kottelat, 2022" box="[474,604,223,249]" class="Actinopterygii" family="Nemacheilidae" genus="Nemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="99" phylum="Chordata" rank="species" species="pezidion">
<emphasis id="B94CEAB5FF916A64FE1CFF778F2F44F9" box="[474,604,223,249]" italics="true" pageId="7" pageNumber="99">N. pezidion</emphasis>
</taxonomicName>
have a modified pectoral fin with conspicuous tubercles on anterior rays and wide unculiferous pads (vs. pectoral fins not modified, tubercles minute and few, on a single row along posterior margin of branched rays 1–3), and numerous small tubercles on the middle and posterior part of the flank of large males (vs. without tubercles), and the suborbital flap well developed (vs. rudimentary). The smallest observed specimen of 
<taxonomicName id="4C384D24FF916A64FEE3FEC78CD74589" authority="Kottelat, 2022" box="[293,420,367,393]" class="Actinopterygii" family="Nemacheilidae" genus="Nemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="99" phylum="Chordata" rank="species" species="pezidion">
<emphasis id="B94CEAB5FF916A64FEE3FEC78CD74589" box="[293,420,367,393]" italics="true" pageId="7" pageNumber="99">N. pezidion</emphasis>
</taxonomicName>
is 
<quantity id="4CC09B42FF916A64FE05FEC78F554589" box="[451,550,367,393]" metricMagnitude="-2" metricUnit="m" metricValue="3.92" pageId="7" pageNumber="99" unit="mm" value="39.2">39.2 mm</quantity>
SL and all specimens have the colour pattern described above of a quite regular stripe. In specimens of 
<taxonomicName id="4C384D24FF916A64FE58FE3B8F3E45AD" authorityName="Kottelat" authorityYear="1990" box="[414,589,403,429]" class="Actinopterygii" family="Nemacheilidae" genus="Nemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="99" phylum="Chordata" rank="species" species="longistriatus">
<emphasis id="B94CEAB5FF916A64FE58FE3B8F3E45AD" box="[414,589,403,429]" italics="true" pageId="7" pageNumber="99">N. longistriatus</emphasis>
</taxonomicName>
the stripe is less regular and small specimens have small bars on the flank, connected with the saddles on the back; the bars then become fainter, and only remain as spots on the course of the lateral line, becoming connected by a midlateral stripe; vestiges of the spots persist, more or less distinct, until adult, when the stripe is irregular (
<figureCitation id="13032A22FF916A64FE1AFE578F5C4619" box="[476,559,511,537]" captionStart="FIGURE 8" captionStartId="7.[151,250,1788,1812]" captionTargetBox="[151,1436,633,1765]" captionTargetId="figure-249@7.[151,1436,633,1765]" captionTargetPageId="7" captionText="FIGURE 8. Nemacheilus longistriatus, Mekong drainage: a, CAS 62547, holotype, 53.2 mm SL; Thailand: Mekong at Chiang Khan; b, CMK 23085, 37.5 mm SL; Laos: Xe Bangfai." figureDoi="http://doi.org/10.5281/zenodo.6488224" httpUri="https://zenodo.org/record/6488224/files/figure.png" pageId="7" pageNumber="99">Fig. 8a</figureCitation>
). 
<taxonomicName id="4C384D24FF916A64FD80FE578E4D4619" authority="Kottelat, 2022" box="[582,830,511,537]" class="Actinopterygii" family="Nemacheilidae" genus="Nemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="99" phylum="Chordata" rank="species" species="pezidion">
<emphasis id="B94CEAB5FF916A64FD80FE578E4D4619" box="[582,830,511,537]" italics="true" pageId="7" pageNumber="99">Nemacheilus pezidion</emphasis>
</taxonomicName>
has up to now been recorded only from the Mekong drainage downstream of Khone falls. Data on 
<taxonomicName id="4C384D24FF916A64FD6CFD8B8E28463D" authorityName="Kottelat" authorityYear="1990" box="[682,859,547,573]" class="Actinopterygii" family="Nemacheilidae" genus="Nemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="99" phylum="Chordata" rank="species" species="longistriatus">
<emphasis id="B94CEAB5FF916A64FD6CFD8B8E28463D" box="[682,859,547,573]" italics="true" pageId="7" pageNumber="99">N. longistriatus</emphasis>
</taxonomicName>
are from 
<bibRefCitation id="EFA94B56FF916A64FC08FD8B890F463D" author="Kottelat, M." box="[974,1148,547,573]" pageId="7" pageNumber="99" refId="ref6499" refString="Kottelat, M. (1990) Indochinese nemacheilines. A revision of nemacheiline loaches (Pisces: Cypriniformes) of Thailand, Burma, Laos, Cambodia and southern Viet Nam. Pfeil, Munchen, 262 pp." type="book" year="1990">Kottelat (1990)</bibRefCitation>
and additional examined material.
</paragraph>
<caption id="DF47662FFF916A64FF51F9548FBB4338" ID-DOI="http://doi.org/10.5281/zenodo.6488224" ID-Zenodo-Dep="6488224" httpUri="https://zenodo.org/record/6488224/files/figure.png" pageId="7" pageNumber="99" startId="7.[151,250,1788,1812]" targetBox="[151,1436,633,1765]" targetPageId="7">
<paragraph id="8B8736A7FF916A64FF51F9548FBB4338" blockId="7.[151,1436,1788,1848]" pageId="7" pageNumber="99">
<emphasis id="B94CEAB5FF916A64FF51F9548C604314" bold="true" box="[151,275,1788,1812]" pageId="7" pageNumber="99">FIGURE 8.</emphasis>
<taxonomicName id="4C384D24FF916A64FEDFF9548FF14314" authority=", Mekong" authorityName="Mekong" box="[281,642,1788,1812]" class="Actinopterygii" family="Nemacheilidae" genus="Nemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="99" phylum="Chordata" rank="species" species="longistriatus">
<emphasis id="B94CEAB5FF916A64FEDFF9548F514314" box="[281,546,1788,1812]" italics="true" pageId="7" pageNumber="99">Nemacheilus longistriatus</emphasis>
, Mekong
</taxonomicName>
drainage: 
<emphasis id="B94CEAB5FF916A64FD2BF9548E734314" bold="true" box="[749,768,1788,1812]" pageId="7" pageNumber="99">a,</emphasis>
CAS 62547, holotype, 53.2 mm SL; Thailand: Mekong at Chiang Khan; 
<emphasis id="B94CEAB5FF916A64FF1DF8888D9C4338" bold="true" box="[219,239,1824,1848]" pageId="7" pageNumber="99">b,</emphasis>
CMK 23085, 37.5 mm SL; Laos: Xe Bangfai.
</paragraph>
</caption>
<paragraph id="8B8736A7FF916A6BFF01F8C88CA845F6" blockId="7.[151,1436,1888,2024]" lastBlockId="8.[151,1437,151,502]" lastPageId="8" lastPageNumber="100" pageId="7" pageNumber="99">
<emphasis id="B94CEAB5FF916A64FF01F8C88F704379" box="[199,515,1888,1914]" italics="true" pageId="7" pageNumber="99">‘Nemacheilus’ argyrogaster</emphasis>
(
<figureCitation id="13032A22FF916A64FDD1F8C88F13437A" box="[535,608,1888,1914]" captionStart="FIGURE 9" captionStartId="8.[151,250,1246,1270]" captionTargetBox="[151,1436,525,1223]" captionTargetId="figure-190@8.[151,1436,525,1223]" captionTargetPageId="8" captionText="FIGURE 9. ‘Nemacheilus’ argyrogaster, MHNG 2785.032, holotype, 63.0 mm SL; Laos:Mekong drainage:Xe Kong watershed." figureDoi="http://doi.org/10.5281/zenodo.6488226" httpUri="https://zenodo.org/record/6488226/files/figure.png" pageId="7" pageNumber="99">Fig. 9</figureCitation>
) is a second species from the Xe Kong watershed that had also been misidentified as 
<taxonomicName id="4C384D24FF916A64FE8DF82C8C8B439E" box="[331,504,1924,1950]" class="Actinopterygii" family="Nemacheilidae" genus="Nemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="99" phylum="Chordata" rank="species" species="longistriatus">
<emphasis id="B94CEAB5FF916A64FE8DF82C8C8B439E" box="[331,504,1924,1950]" italics="true" pageId="7" pageNumber="99">N. longistriatus</emphasis>
</taxonomicName>
(e.g., 
<bibRefCitation id="EFA94B56FF916A64FDFFF82C8E6A439E" author="Baird, I. G. &amp; Inthaphaisy, V. &amp; Kisouvannalath, P. &amp; Phylavanh, B. &amp; Mounsouphom, B." box="[569,793,1924,1950]" pageId="7" pageNumber="99" refId="ref5819" refString="Baird, I. G., Inthaphaisy, V., Kisouvannalath, P., Phylavanh, B. &amp; Mounsouphom, B. (1999) [The fishes of southern Lao]. Lao Community Fisheries and Dolphin Protection Project, Pakse, 161 pp. [in Lao]" type="book" year="1999">
Baird 
<emphasis id="B94CEAB5FF916A64FDBAF82C8FD8439E" box="[636,683,1924,1950]" italics="true" pageId="7" pageNumber="99">et al</emphasis>
. 1999: 70
</bibRefCitation>
; 
<bibRefCitation id="EFA94B56FF916A64FCE5F82C8977439E" author="Rainboth, W. J. &amp; Vidthayanon, C. &amp; Mai, D. Y." box="[803,1028,1924,1950]" pageId="7" pageNumber="99" pagination="1 - 173" refId="ref7568" refString="Rainboth, W. J., Vidthayanon, C. &amp; Mai, D. Y. (2012) Fishes of the Greater Mekong ecosystem with species list and photographic atlas. Miscellaneous Publications, Museum of Zoology, University of Michigan, 201, i - vi + 1 - 173, 121 pls." type="journal article" year="2012">
Rainboth 
<emphasis id="B94CEAB5FF916A64FC48F82C8ECE439E" box="[910,957,1924,1950]" italics="true" pageId="7" pageNumber="99">et al</emphasis>
. 2012
</bibRefCitation>
: pl. 33 fig. 685). When first collected, it was identified as a 
<taxonomicName id="4C384D24FF916A64FE47F8008F6143C2" authorityName=", Kottelat" authorityYear="2012" box="[385,530,1960,1986]" class="Actinopterygii" family="Nemacheilidae" genus="Nemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="99" phylum="Chordata" rank="genus">
<emphasis id="B94CEAB5FF916A64FE47F8008F6143C2" box="[385,530,1960,1986]" italics="true" pageId="7" pageNumber="99">Nemacheilus</emphasis>
</taxonomicName>
, in fact not even distinguished from 
<taxonomicName id="4C384D24FF916A64FC6CF800895A43C2" authority="Kottelat, 2022" box="[938,1065,1960,1986]" class="Actinopterygii" family="Nemacheilidae" genus="Nemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="99" phylum="Chordata" rank="species" species="pezidion">
<emphasis id="B94CEAB5FF916A64FC6CF800895A43C2" box="[938,1065,1960,1986]" italics="true" pageId="7" pageNumber="99">N. pezidion</emphasis>
</taxonomicName>
. Later examination showed them to be distinct, with the molecular study by 
<bibRefCitation id="EFA94B56FF916A64FD40F8658EF843E6" author="Slechtova, V. &amp; Musilova, Z. &amp; Tan, H. H. &amp; Kottelat, M. &amp; Bohlen, J." box="[646,907,1996,2024]" pageId="7" pageNumber="99" pagination="1 - 13" refId="ref7718" refString="Slechtova, V., Musilova, Z., Tan, H. H., Kottelat, M. &amp; Bohlen, J. (2021) One northward, one southward: contrasting biogeographical history in two benthic freshwater fish genera across Southeast Asia (Teleostei: Cobitoidea: Nemacheilus, Pangio). Molecular Phylogenetics and Evolution, 161 (107139), 1 - 13. https: // doi. org / 10.1016 / j. ympev. 2021.107139" type="journal article" year="2021">
Šlechtová 
<emphasis id="B94CEAB5FF916A64FD38F8648E4B43E6" box="[766,824,1996,2022]" italics="true" pageId="7" pageNumber="99">et al.</emphasis>
(2021)
</bibRefCitation>
showing that, if included in 
<taxonomicName id="4C384D24FF916A64FB1FF864881943E6" authorityName=", Kottelat" authorityYear="2012" box="[1241,1386,1996,2022]" class="Actinopterygii" family="Nemacheilidae" genus="Nemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="99" phylum="Chordata" rank="genus">
<emphasis id="B94CEAB5FF916A64FB1FF864881943E6" box="[1241,1386,1996,2022]" italics="true" pageId="7" pageNumber="99">Nemacheilus</emphasis>
</taxonomicName>
, the latter genus ceases to be monophyletic. It therefore falls within 
<taxonomicName id="4C384D24FF9E6A6BFCA9FF3F8EAA44B1" box="[879,985,151,177]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="8" pageNumber="100" phylum="Chordata" rank="genus" sensu="lato">
<emphasis id="B94CEAB5FF9E6A6BFCA9FF3F8EAA44B1" box="[879,985,151,177]" italics="true" pageId="8" pageNumber="100">Schistura</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27F57CEFF9E6A6BFC24FF3F892044B1" box="[994,1107,151,177]" pageId="8" pageNumber="100" sensu="lato">sensu lato</taxonomicNameLabel>
and a more accurate generic placement will only be possible after relationships within 
<taxonomicName id="4C384D24FF9E6A6BFCE8FF138EEB44D5" box="[814,920,187,213]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="8" pageNumber="100" phylum="Chordata" rank="genus" sensu="lato">
<emphasis id="B94CEAB5FF9E6A6BFCE8FF138EEB44D5" box="[814,920,187,213]" italics="true" pageId="8" pageNumber="100">Schistura</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27F57CEFF9E6A6BFC66FF138EB244D5" box="[928,961,187,213]" pageId="8" pageNumber="100" sensu="lato">s.l.</taxonomicNameLabel>
are better understood; 
<taxonomicName id="4C384D24FF9E6A6BFB0CFF13884744D5" box="[1226,1332,187,213]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="8" pageNumber="100" phylum="Chordata" rank="genus">
<emphasis id="B94CEAB5FF9E6A6BFB0CFF13884744D5" box="[1226,1332,187,213]" italics="true" pageId="8" pageNumber="100">Schistura</emphasis>
</taxonomicName>
has long been known to be a polyphyletic catchall for Southeast and South Asian nemacheilids (e.g., 
<bibRefCitation id="EFA94B56FF9E6A6BFB5EFF77884844F9" author="Kottelat, M." box="[1176,1339,223,249]" pageId="8" pageNumber="100" refId="ref6499" refString="Kottelat, M. (1990) Indochinese nemacheilines. A revision of nemacheiline loaches (Pisces: Cypriniformes) of Thailand, Burma, Laos, Cambodia and southern Viet Nam. Pfeil, Munchen, 262 pp." type="book" year="1990">Kottelat, 1990</bibRefCitation>
; 
<bibRefCitation id="EFA94B56FF9E6A6BFA81FF778C6A451D" author="Sember, A. &amp; Bohlen, J. &amp; Slechtova, V. &amp; Altmanova, M. &amp; Symonova, R. &amp; Rab, P." pageId="8" pageNumber="100" pagination="1 - 22" refId="ref7626" refString="Sember, A., Bohlen, J., Slechtova, V., Altmanova, M., Symonova, R. &amp; Rab, P. (2015) Karyotype differentiation in 19 species of river loach fishes (Nemacheilidae, Teleostei): extensive variability associated with rDNA and heterochromatin distribution and its phylogenetic and ecological interpretation. BMC Evolutionary Biology, 15 (251), 1 - 22. https: // doi. org / 10.1186 / s 12862 - 015 - 0532 - 9" type="journal article" year="2015">
Sember 
<emphasis id="B94CEAB5FF9E6A6BFF51FEAB8DBA451D" box="[151,201,259,285]" italics="true" pageId="8" pageNumber="100">et al</emphasis>
., 2015
</bibRefCitation>
; 
<bibRefCitation id="EFA94B56FF9E6A6BFEE3FEAB8F74451D" author="Freyhof, J. &amp; Geiger, M. F. &amp; Golzarianpour, K. &amp; Patimar, R." box="[293,519,259,285]" pageId="8" pageNumber="100" pagination="65 - 80" refId="ref6379" refString="Freyhof, J., Geiger, M. F., Golzarianpour, K. &amp; Patimar, R. 2016. Sasanidus, a new generic name for Noemacheilus kermanshahensis Banarescu &amp; Nalbant, with discussion of Ilamnemacheilus and Schistura (Teleostei; Nemacheilidae). Zootaxa, 4107 (1), 65 - 80. https: // doi. org / 10.11646 / zootaxa. 4107.1.3" type="journal article" year="2016">
Freyhof 
<emphasis id="B94CEAB5FF9E6A6BFE43FEAB8CC5451D" box="[389,438,259,285]" italics="true" pageId="8" pageNumber="100">et al</emphasis>
., 2016
</bibRefCitation>
; 
<bibRefCitation id="EFA94B56FF9E6A6BFDD4FEAB8FAB451D" author="Chen, W. &amp; Yang, J. - P. &amp; Li, Y. - F. &amp; Li, X. - H." box="[530,728,259,285]" pageId="8" pageNumber="100" pagination="1 - 11" refId="ref6143" refString="Chen, W., Yang, J. - P., Li, Y. - F. &amp; Li, X. - H. (2019) Exploring taxonomic diversity and biogeography of the family Nemacheilinae (Cypriniformes). Ecology and Evolution, 9 (10343), 1 - 11. https: // doi. org / 10.1002 / ece 3.5553" type="journal article" year="2019">
Chen 
<emphasis id="B94CEAB5FF9E6A6BFD93FEAB8FF5451D" box="[597,646,259,285]" italics="true" pageId="8" pageNumber="100">et al</emphasis>
., 2019
</bibRefCitation>
). The two species are distinguished by a forked caudal fin with the upper lobe distinctly longer than the lower one in 
<taxonomicName id="4C384D24FF9E6A6BFD35FE8F8E074541" authority="Kottelat, 2022" box="[755,884,295,321]" class="Actinopterygii" family="Nemacheilidae" genus="Nemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="8" pageNumber="100" phylum="Chordata" rank="species" species="pezidion">
<emphasis id="B94CEAB5FF9E6A6BFD35FE8F8E074541" box="[755,884,295,321]" italics="true" pageId="8" pageNumber="100">N. pezidion</emphasis>
</taxonomicName>
(vs. subequal in 
<emphasis id="B94CEAB5FF9E6A6BFBF2FE8F89844540" box="[1076,1271,295,321]" italics="true" pageId="8" pageNumber="100">‘N’. argyrogaster</emphasis>
), the pectoralfin rays with filamentous extensions (vs. without extensions), the presence of an elongate spot on lower ¼ of first branched dorsal-fin ray and adjacent membranes (vs. absence), and the flank colour about the same above and below the midlateral stripe (vs. the flank above the midlateral stripe being yellowish brown, pale yellow to silvery below the stripe). Further, large male 
<taxonomicName id="4C384D24FF9E6A6BFE31FE1F8F0B45D1" authority="Kottelat, 2022" box="[503,632,439,465]" class="Actinopterygii" family="Nemacheilidae" genus="Nemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="8" pageNumber="100" phylum="Chordata" rank="species" species="pezidion">
<emphasis id="B94CEAB5FF9E6A6BFE31FE1F8F0B45D1" box="[503,632,439,465]" italics="true" pageId="8" pageNumber="100">N. pezidion</emphasis>
</taxonomicName>
have numerous small tubercles on the middle and posterior part of the flank (vs. without tubercles).
</paragraph>
<caption id="DF47662FFF9E6A6BFF51FB7688EE40F6" ID-DOI="http://doi.org/10.5281/zenodo.6488226" ID-Zenodo-Dep="6488226" box="[151,1437,1246,1271]" httpUri="https://zenodo.org/record/6488226/files/figure.png" pageId="8" pageNumber="100" startId="8.[151,250,1246,1270]" targetBox="[151,1436,525,1223]" targetPageId="8">
<paragraph id="8B8736A7FF9E6A6BFF51FB7688EE40F6" blockId="8.[151,1437,1246,1271]" box="[151,1437,1246,1271]" pageId="8" pageNumber="100">
<emphasis id="B94CEAB5FF9E6A6BFF51FB768C6240F6" bold="true" box="[151,273,1246,1270]" pageId="8" pageNumber="100">FIGURE 9.</emphasis>
<emphasis id="B94CEAB5FF9E6A6BFED3FB768F5F40F6" box="[277,556,1246,1270]" italics="true" pageId="8" pageNumber="100">‘Nemacheilus’ argyrogaster</emphasis>
, MHNG 2785.032, holotype, 63.0 mm SL; Laos: Mekong drainage: Xe Kong watershed.
</paragraph>
</caption>
<paragraph id="8B8736A7FF9E6A6BFF01FA8C8FDA4256" blockId="8.[151,1437,1316,2042]" pageId="8" pageNumber="100">
Based on the present data, 
<taxonomicName id="4C384D24FF9E6A6BFDC6FA8D8FF0413E" authority="Kottelat, 2022" box="[512,643,1316,1342]" class="Actinopterygii" family="Nemacheilidae" genus="Nemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="8" pageNumber="100" phylum="Chordata" rank="species" species="pezidion">
<emphasis id="B94CEAB5FF9E6A6BFDC6FA8D8FF0413E" box="[512,643,1316,1342]" italics="true" pageId="8" pageNumber="100">N. pezidion</emphasis>
</taxonomicName>
and 
<emphasis id="B94CEAB5FF9E6A6BFD7BFA8D8EF0413E" box="[701,899,1317,1343]" italics="true" pageId="8" pageNumber="100">‘N.’ argyrogaster</emphasis>
have allopatric distributions, with 
<taxonomicName id="4C384D24FF9E6A6BFADFFA8D88EF413E" authority="Kottelat, 2022" box="[1305,1436,1316,1342]" class="Actinopterygii" family="Nemacheilidae" genus="Nemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="8" pageNumber="100" phylum="Chordata" rank="species" species="pezidion">
<emphasis id="B94CEAB5FF9E6A6BFADFFA8D88EF413E" box="[1305,1436,1316,1342]" italics="true" pageId="8" pageNumber="100">N. pezidion</emphasis>
</taxonomicName>
observed only in the Xe Pian (a tributary of Xe Kong), and 
<emphasis id="B94CEAB5FF9E6A6BFCF7FAE08E874161" box="[817,1012,1352,1378]" italics="true" pageId="8" pageNumber="100">‘N.’ argyrogaster</emphasis>
observed only in the Xe Kong proper and in tributaries upriver of 
<collectingRegion id="49FCF845FF9E6A6BFE1CFAC38F464185" box="[474,565,1387,1413]" country="Laos" name="Attapu" pageId="8" pageNumber="100">Attapeu</collectingRegion>
(
<figureCitation id="13032A22FF9E6A6BFD82FAC28FF94185" box="[580,650,1386,1413]" captionStart="FIGURE 5" captionStartId="5.[151,250,1894,1918]" captionTargetBox="[170,1414,1029,1867]" captionTargetId="figure-417@5.[170,1417,1029,1870]" captionTargetPageId="5" captionText="FIGURE 5. Southern Laos, showing localities of examined material of Nemacheilus pezidion (circles) and ‘N’. argyrogaster (triangles). Each symbol may represent several localities. White circle is type locality of N. pezidion. Dams, reservoirs and other alterations of original hydrographic network are not shown." figureDoi="http://doi.org/10.5281/zenodo.6488216" httpUri="https://zenodo.org/record/6488216/files/figure.png" pageId="8" pageNumber="100">Fig. 5</figureCitation>
); this may, however, be only a sampling artefact, related to different habitats targeted by different components of the surveys. At the time (
<date id="FF861067FF9E6A6BFC0EFA2589E341A7" box="[968,1168,1421,1448]" pageId="8" pageNumber="100" value="1999-05-20" valueMax="1999-05-22" valueMin="1999-05-20">20–22 May 1999</date>
and 2009) and sites of sampling, 
<taxonomicName id="4C384D24FF9E6A6BFED6FA198CE641CA" authority="Kottelat, 2022" box="[272,405,1456,1482]" class="Actinopterygii" family="Nemacheilidae" genus="Nemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="8" pageNumber="100" phylum="Chordata" rank="species" species="pezidion">
<emphasis id="B94CEAB5FF9E6A6BFED6FA198CE641CA" box="[272,405,1456,1482]" italics="true" pageId="8" pageNumber="100">N. pezidion</emphasis>
</taxonomicName>
was in flowing turbid water over mostly gravel to mud bottom, in low gradient streams in the flood plain. Specimens of 
<emphasis id="B94CEAB5FF9E6A6BFDC9FA7C8FA441ED" box="[527,727,1492,1518]" italics="true" pageId="8" pageNumber="100">‘N’. argyrogaster</emphasis>
were collected during the same period (
<date id="FF861067FF9E6A6BFB60FA7B881F41ED" box="[1190,1388,1491,1518]" pageId="8" pageNumber="100" value="1999-05-20" valueMax="1999-05-28" valueMin="1999-05-20">20–28 May 1999</date>
and 2009), in streams with faster gradient, in rapids, with substrate comprised of stones to boulders. [CMK 15687, 
<typeStatus id="54838805FF9E6A6BFF51F9B28DA74234" box="[151,212,1562,1588]" pageId="8" pageNumber="100">types</typeStatus>
of 
<taxonomicName id="4C384D24FF9E6A6BFF3AF9B28C0C4233" authority="Kottelat, 2022" box="[252,383,1561,1587]" class="Actinopterygii" family="Nemacheilidae" genus="Nemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="8" pageNumber="100" phylum="Chordata" rank="species" species="pezidion">
<emphasis id="B94CEAB5FF9E6A6BFF3AF9B28C0C4233" box="[252,383,1561,1587]" italics="true" pageId="8" pageNumber="100">N. pezidion</emphasis>
</taxonomicName>
, were erroneously listed as 
<emphasis id="B94CEAB5FF9E6A6BFD02F9B18EFE4233" box="[708,909,1561,1587]" italics="true" pageId="8" pageNumber="100">‘N’. argyrogaster</emphasis>
in 
<bibRefCitation id="EFA94B56FF9E6A6BFC72F9B189E44233" author="Kottelat, M." box="[948,1175,1561,1587]" pageId="8" pageNumber="100" pagination="277 - 288" refId="ref7231" refString="Kottelat, M. (2021) ' Nemacheilus' argyrogaster, a new species of loach from southern Laos (Teleostei: Nemacheilidae). Zootaxa, 4933 (2), 277 - 288. https: // doi. org / 10.11646 / zootaxa. 4933.2.6" type="journal article" year="2021">Kottelat, 2021: 277</bibRefCitation>
; this removes the only record of 
<emphasis id="B94CEAB5FF9E6A6BFEC1F9948CB84256" box="[263,459,1596,1622]" italics="true" pageId="8" pageNumber="100">‘N.’ argyrogaster</emphasis>
from the Xe Pian].
</paragraph>
<paragraph id="8B8736A7FF9E6A6AFF01F9F7882944F7" blockId="8.[151,1437,1316,2042]" lastBlockId="9.[151,1437,151,423]" lastPageId="9" lastPageNumber="101" pageId="8" pageNumber="100">
<taxonomicName id="4C384D24FF9E6A6BFF01F9F78FFE4279" authority="Zhu &amp; Wang" authorityName="Zhu &amp; Wang" box="[199,653,1631,1657]" class="Actinopterygii" family="Nemacheilidae" genus="Nemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="8" pageNumber="100" phylum="Chordata" rank="species" species="nandingensis">
<emphasis id="B94CEAB5FF9E6A6BFF01F9F78C804279" box="[199,499,1631,1657]" italics="true" pageId="8" pageNumber="100">Nemacheilus nandingensis</emphasis>
Zhu &amp; Wang
</taxonomicName>
, described from the Salween drainage in 
<collectingRegion id="49FCF845FF9E6A6BFB9FF9F789C14279" box="[1113,1202,1631,1657]" country="China" name="Yunnan" pageId="8" pageNumber="100">Yunnan</collectingRegion>
, also has a yellowish to whitish body with a conspicuous black midlateral stripe and a middorsal series of 16–19 squarish saddles (
<bibRefCitation id="EFA94B56FF9E6A6BFAABF92A8C3342C0" author="Zhu, S. - Q. &amp; Wang, S. - H." pageId="8" pageNumber="100" pagination="208 - 220" refId="ref7848" refString="Zhu, S. - Q. &amp; Wang, S. - H. (1985) [The noemacheiline fishes from Yunnan Province, China]. Acta Zootaxonomica Sinica, 10 (2), 208 - 220. [in Chinese, English summary]" type="journal article" year="1985">Zhu &amp; Wang, 1985</bibRefCitation>
). 
<taxonomicName id="4C384D24FF9E6A6BFE92F90D8FF242BF" box="[340,641,1701,1727]" class="Actinopterygii" family="Nemacheilidae" genus="Nemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="8" pageNumber="100" phylum="Chordata" rank="species" species="nandingensis">
<emphasis id="B94CEAB5FF9E6A6BFE92F90D8FF242BF" box="[340,641,1701,1727]" italics="true" pageId="8" pageNumber="100">Nemacheilus nandingensis</emphasis>
</taxonomicName>
was treated as a synonym of 
<taxonomicName id="4C384D24FF9E6A6BFC14F90D880B42C0" authority="(Zhu)" baseAuthorityName="Zhu" baseAuthorityYear="1982" box="[978,1400,1701,1728]" class="Actinopterygii" family="Nemacheilidae" genus="Pteronemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="8" pageNumber="100" phylum="Chordata" rank="species" species="meridionalis">
<emphasis id="B94CEAB5FF9E6A6BFC14F90D884242BF" box="[978,1329,1701,1727]" italics="true" pageId="8" pageNumber="100">Pteronemacheilus meridionalis</emphasis>
(Zhu)
</taxonomicName>
by Yang (in 
<bibRefCitation id="EFA94B56FF9E6A6BFF39F9608C8842E2" author="Chu, X. - L. &amp; Chen, Y. - R." box="[255,507,1736,1762]" pageId="8" pageNumber="100" refId="ref6337" refString="Chu, X. - L. &amp; Chen, Y. - R. (1990) [The fishes of Yunnan, China. Part II]. Science Press, Beijing, 313 pp. [in Chinese]" type="book" year="1990">Chu &amp; Chen 1990: 49</bibRefCitation>
), as a valid species of 
<taxonomicName id="4C384D24FF9E6A6BFD3EF9608E1142E2" box="[760,866,1736,1762]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="8" pageNumber="100" phylum="Chordata" rank="genus">
<emphasis id="B94CEAB5FF9E6A6BFD3EF9608E1142E2" box="[760,866,1736,1762]" italics="true" pageId="8" pageNumber="100">Schistura</emphasis>
</taxonomicName>
by 
<bibRefCitation id="EFA94B56FF9E6A6BFC4BF960894942E3" author="Zhu, S. - Q." box="[909,1082,1736,1763]" pageId="8" pageNumber="100" refId="ref7803" refString="Zhu, S. - Q. (1989) [The loaches of the subfamily Nemacheilinae in China (Cypriniformes: Cobitidae)]. Jiangsu Science and Technology Publishing House, Nanjing, 150 pp. [in Chinese, English summary]" type="book" year="1989">Zhu (1989: 54)</bibRefCitation>
, 
<bibRefCitation id="EFA94B56FF9E6A6BFB80F960885942E2" author="Kottelat, M." box="[1094,1322,1736,1762]" pageId="8" pageNumber="100" pagination="1 - 199" refId="ref6708" refString="Kottelat, M. (2012) Conspectus cobitidum: an inventory of the loaches of the world (Teleostei: Cypriniformes: Cobitoidea). Raffles Bulletin of Zoology, Supplement, 26, 1 - 199." type="journal article" year="2012">Kottelat (2012: 113)</bibRefCitation>
and 
<bibRefCitation id="EFA94B56FF9E6A6BFAA7F9608C534306" author="Chen, X. - Y." pageId="8" pageNumber="100" pagination="281 - 343" refId="ref6211" refString="Chen, X. - Y. (2013) [Checklist of fishes of Yunnan]. Zoological Research, 34 (4), 281 - 343. [in Chinese, English summary]" type="journal article" year="2013">Chen (2013: 296)</bibRefCitation>
, and as a valid species of 
<taxonomicName id="4C384D24FF9E6A6BFD90F9438F944305" authorityName=", Kottelat" authorityYear="2012" box="[598,743,1771,1797]" class="Actinopterygii" family="Nemacheilidae" genus="Nemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="8" pageNumber="100" phylum="Chordata" rank="genus">
<emphasis id="B94CEAB5FF9E6A6BFD90F9438F944305" box="[598,743,1771,1797]" italics="true" pageId="8" pageNumber="100">Nemacheilus</emphasis>
</taxonomicName>
by 
<bibRefCitation id="EFA94B56FF9E6A6BFCD1F94389774305" author="Kottelat, M." box="[791,1028,1770,1797]" pageId="8" pageNumber="100" pagination="277 - 288" refId="ref7231" refString="Kottelat, M. (2021) ' Nemacheilus' argyrogaster, a new species of loach from southern Laos (Teleostei: Nemacheilidae). Zootaxa, 4933 (2), 277 - 288. https: // doi. org / 10.11646 / zootaxa. 4933.2.6" type="journal article" year="2021">Kottelat (2021: 285)</bibRefCitation>
. No topotypical specimen could be examined. A species apparently similar to 
<taxonomicName id="4C384D24FF9E6A6BFDB5F8A68E5B4328" box="[627,808,1806,1832]" class="Actinopterygii" family="Nemacheilidae" genus="Nemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="8" pageNumber="100" phylum="Chordata" rank="species" species="nandingensis">
<emphasis id="B94CEAB5FF9E6A6BFDB5F8A68E5B4328" box="[627,808,1806,1832]" italics="true" pageId="8" pageNumber="100">N. nandingensis</emphasis>
</taxonomicName>
, or the same species, is present in the Mekong drainage in Xishuangbanna (
<collectingRegion id="49FCF845FF9E6A6BFEBFF8998CA1434B" box="[377,466,1841,1867]" country="China" name="Yunnan" pageId="8" pageNumber="100">Yunnan</collectingRegion>
) (
<figureCitation id="13032A22FF9E6A6BFE2BF8988F37434B" box="[493,580,1840,1867]" captionStart="FIGURE 10" captionStartId="9.[151,250,1024,1048]" captionTargetBox="[151,1436,446,1000]" captionTargetId="figure-174@9.[151,1436,446,1000]" captionTargetPageId="9" captionText="FIGURE 10 Nemacheilus aff. nandingensis, CMK 23949, 56.6 mm SL; China: Xishuangbanna: Mekong drainage: Nam Xing." figureDoi="http://doi.org/10.5281/zenodo.6488234" httpUri="https://zenodo.org/record/6488234/files/figure.png" pageId="8" pageNumber="100">Fig. 10</figureCitation>
); the single available specimen, a male, has a small triangular suborbital flap and a pectoral fin with no tubercles (the first branched ray is thickened and its branches are not secondarily branched). The two species are distinguished by a forked caudal fin with the upper lobe distinctly longer than the lower one in 
<taxonomicName id="4C384D24FF9E6A6BFEE8F8328CDC43B3" authority="Kottelat, 2022" box="[302,431,1945,1971]" class="Actinopterygii" family="Nemacheilidae" genus="Nemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="8" pageNumber="100" phylum="Chordata" rank="species" species="pezidion">
<emphasis id="B94CEAB5FF9E6A6BFEE8F8328CDC43B3" box="[302,431,1945,1971]" italics="true" pageId="8" pageNumber="100">N. pezidion</emphasis>
</taxonomicName>
(vs. subequal in 
<taxonomicName id="4C384D24FF9E6A6BFDB5F8328E5A43B3" box="[627,809,1945,1971]" class="Actinopterygii" family="Nemacheilidae" genus="Nemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="8" pageNumber="100" phylum="Chordata" rank="species" species="nandingensis">
<emphasis id="B94CEAB5FF9E6A6BFDB5F8328E5A43B3" box="[627,809,1945,1971]" italics="true" pageId="8" pageNumber="100">N. nandingensis</emphasis>
</taxonomicName>
), the pectoral-fin rays with filamentous extensions (vs. without extensions), the presence of an elongate spot on the lower ¼ of the first branched dorsal-fin ray and adjacent membranes (vs. absence), and a thinner midlateral stripe, wider anteriorly, where clearly less than eye diameter (vs. width uniform from head to caudal-fin base, about equal to eye diameter). Further, male 
<taxonomicName id="4C384D24FF9F6A6AFBA0FF3F899644B1" authority="Kottelat, 2022" box="[1126,1253,151,177]" class="Actinopterygii" family="Nemacheilidae" genus="Nemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="9" pageNumber="101" phylum="Chordata" rank="species" species="pezidion">
<emphasis id="B94CEAB5FF9F6A6AFBA0FF3F899644B1" box="[1126,1253,151,177]" italics="true" pageId="9" pageNumber="101">N. pezidion</emphasis>
</taxonomicName>
have a modified pectoral fin with conspicuous tubercles on the anterior rays and wide unculiferous pads (vs. without), in addition to numerous small tubercles on the middle and posterior part of the flank of large males (vs. without tubercles).
</paragraph>
<paragraph id="8B8736A7FF9F6A6AFF01FF5788EE45A6" blockId="9.[151,1437,151,423]" pageId="9" pageNumber="101">
The mouth of 
<taxonomicName id="4C384D24FF9F6A6AFEA7FEA88F674519" box="[353,532,255,281]" class="Actinopterygii" family="Nemacheilidae" genus="Nemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="9" pageNumber="101" phylum="Chordata" rank="species" species="nandingensis">
<emphasis id="B94CEAB5FF9F6A6AFEA7FEA88F674519" box="[353,532,255,281]" italics="true" pageId="9" pageNumber="101">N. nandingensis</emphasis>
</taxonomicName>
has the characters of 
<taxonomicName id="4C384D24FF9F6A6AFD38FF578EFC4519" authorityName=", Kottelat" authorityYear="2012" box="[766,911,255,281]" class="Actinopterygii" family="Nemacheilidae" genus="Nemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="9" pageNumber="101" phylum="Chordata" rank="genus">
<emphasis id="B94CEAB5FF9F6A6AFD38FF578EFC4519" box="[766,911,255,281]" italics="true" pageId="9" pageNumber="101">Nemacheilus</emphasis>
</taxonomicName>
(see 
<bibRefCitation id="EFA94B56FF9F6A6AFC00FF578915451A" author="Kottelat, M." box="[966,1126,255,282]" pageId="9" pageNumber="101" refId="ref6499" refString="Kottelat, M. (1990) Indochinese nemacheilines. A revision of nemacheiline loaches (Pisces: Cypriniformes) of Thailand, Burma, Laos, Cambodia and southern Viet Nam. Pfeil, Munchen, 262 pp." type="book" year="1990">Kottelat, 1990</bibRefCitation>
) and definitively neither the conspicuous median interruption in the lower lip nor the deep sulci in its median parts typical of 
<taxonomicName id="4C384D24FF9F6A6AFB14FE8A884F453C" box="[1234,1340,290,316]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="9" pageNumber="101" phylum="Chordata" rank="genus" sensu="lato">
<emphasis id="B94CEAB5FF9F6A6AFB14FE8A884F453C" box="[1234,1340,290,316]" italics="true" pageId="9" pageNumber="101">Schistura</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27F57CEFF9F6A6AFA85FE8A8817453C" box="[1347,1380,290,316]" pageId="9" pageNumber="101" sensu="lato">s.l.</taxonomicNameLabel>
This is possibly the species identified as 
<taxonomicName id="4C384D24FF9F6A6AFDEBFEEE8FAE455F" box="[557,733,325,351]" class="Actinopterygii" family="Nemacheilidae" genus="Nemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="9" pageNumber="101" phylum="Chordata" rank="species" species="longistriatus">
<emphasis id="B94CEAB5FF9F6A6AFDEBFEEE8FAE455F" box="[557,733,325,351]" italics="true" pageId="9" pageNumber="101">N. longistriatus</emphasis>
</taxonomicName>
by 
<bibRefCitation id="EFA94B56FF9F6A6AFCCFFEED8E8E455F" author="Lin, F. &amp; Zhang, Y. &amp; Fu, G. - Q. &amp; Chen, X. - Y." box="[777,1021,325,352]" pageId="9" pageNumber="101" pagination="871 - 881" refId="ref7353" refString="Lin, F., Zhang, Y., Fu, G. - Q. &amp; Chen, X. - Y. (2021) [Three new records of fish species (Nemacheilidae) in the lower Lancang River, Yunnan Province]. Chinese Journal of Zoology, 56 (6), 871 - 881. [in Chinese, English summary]" type="journal article" year="2021">
Lin 
<emphasis id="B94CEAB5FF9F6A6AFCFEFEEE8E19455F" box="[824,874,325,351]" italics="true" pageId="9" pageNumber="101">et al</emphasis>
. (2021: 873)
</bibRefCitation>
from the Mekong River mainstream at Menghan, 
<collectingRegion id="49FCF845FF9F6A6AFEECFEC18CF04583" box="[298,387,361,387]" country="China" name="Yunnan" pageId="9" pageNumber="101">Yunnan</collectingRegion>
, 
<collectingCountry id="F32F7637FF9F6A6AFE48FEC08CA64582" box="[398,469,360,386]" name="China" pageId="9" pageNumber="101">China</collectingCountry>
. Their pictures definitively do not depict 
<taxonomicName id="4C384D24FF9F6A6AFC6EFEC189244582" box="[936,1111,360,386]" class="Actinopterygii" family="Nemacheilidae" genus="Nemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="9" pageNumber="101" phylum="Chordata" rank="species" species="longistriatus">
<emphasis id="B94CEAB5FF9F6A6AFC6EFEC189244582" box="[936,1111,360,386]" italics="true" pageId="9" pageNumber="101">N. longistriatus</emphasis>
</taxonomicName>
(compare their figure 2 with 
<figureCitation id="13032A22FF9F6A6AFF51FE248D9445A6" box="[151,231,396,422]" captionStart="FIGURE 10" captionStartId="9.[151,250,1024,1048]" captionTargetBox="[151,1436,446,1000]" captionTargetId="figure-174@9.[151,1436,446,1000]" captionTargetPageId="9" captionText="FIGURE 10 Nemacheilus aff. nandingensis, CMK 23949, 56.6 mm SL; China: Xishuangbanna: Mekong drainage: Nam Xing." figureDoi="http://doi.org/10.5281/zenodo.6488234" httpUri="https://zenodo.org/record/6488234/files/figure.png" pageId="9" pageNumber="101">Fig. 10</figureCitation>
of the present article); the suboptimal fixation of their specimen does not allow a more detailed discussion.
</paragraph>
<caption id="DF47662FFF9F6A6AFF51FBA88DA2403C" ID-DOI="http://doi.org/10.5281/zenodo.6488234" ID-Zenodo-Dep="6488234" httpUri="https://zenodo.org/record/6488234/files/figure.png" pageId="9" pageNumber="101" startId="9.[151,250,1024,1048]" targetBox="[151,1436,446,1000]" targetPageId="9">
<paragraph id="8B8736A7FF9F6A6AFF51FBA88DA2403C" blockId="9.[151,1436,1024,1084]" pageId="9" pageNumber="101">
<emphasis id="B94CEAB5FF9F6A6AFF51FBA88C6E4018" bold="true" box="[151,285,1024,1048]" pageId="9" pageNumber="101">FIGURE 10</emphasis>
<taxonomicName id="4C384D24FF9F6A6AFEEEFBA88FC04018" authority=", CMK" authorityName="CMK" box="[296,691,1024,1048]" class="Actinopterygii" family="Nemacheilidae" genus="Nemacheilus" isUncertain="true" kingdom="Animalia" order="Cypriniformes" pageId="9" pageNumber="101" phylum="Chordata" rank="species" species="nandingensis">
<emphasis id="B94CEAB5FF9F6A6AFEEEFBA88CD84018" box="[296,427,1024,1048]" italics="true" pageId="9" pageNumber="101">Nemacheilus</emphasis>
aff. 
<emphasis id="B94CEAB5FF9F6A6AFE24FBA88F1B4018" box="[482,616,1024,1048]" italics="true" pageId="9" pageNumber="101">nandingensis</emphasis>
, CMK
</taxonomicName>
23949, 56.6 mm SL; China: Xishuangbanna: Mekong drainage: Nam Xing.
</paragraph>
</caption>
<paragraph id="8B8736A7FF9F6A6AFF01FBC28C1A4139" blockId="9.[151,1437,1130,1624]" pageId="9" pageNumber="101">
<taxonomicName id="4C384D24FF9F6A6AFF01FBC28FB44087" authority="Bohlen &amp; Slechtova" authorityName="Bohlen &amp; Slechtova" authorityYear="2013" box="[199,711,1130,1159]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="9" pageNumber="101" phylum="Chordata" rank="species" species="rubrimaculata">
<emphasis id="B94CEAB5FF9F6A6AFF01FBC28CA84084" box="[199,475,1130,1156]" italics="true" pageId="9" pageNumber="101">Schistura rubrimaculata</emphasis>
Bohlen &amp; Šlechtová
</taxonomicName>
(from the 
<collectingRegion id="49FCF845FF9F6A6AFC86FBC28EE14084" box="[832,914,1130,1156]" country="Myanmar" name="Rakhine" pageId="9" pageNumber="101">Arakan</collectingRegion>
range in 
<collectingRegion id="49FCF845FF9F6A6AFC38FBC2892D4084" box="[1022,1118,1130,1156]" country="Myanmar" name="Rakhine" pageId="9" pageNumber="101">Rakhine</collectingRegion>
, 
<collectingCountry id="F32F7637FF9F6A6AFBACFBC389AA4085" box="[1130,1241,1131,1157]" name="Myanmar" pageId="9" pageNumber="101">Myanmar</collectingCountry>
) and 
<taxonomicName id="4C384D24FF9F6A6AFADFFBC38C0E40AB" authority="Bohlen &amp; Slechtova" authorityName="Bohlen &amp; Slechtova" authorityYear="2013" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="9" pageNumber="101" phylum="Chordata" rank="species" species="pawensis">
<emphasis id="B94CEAB5FF9F6A6AFADFFBC388EE4084" box="[1305,1437,1131,1157]" italics="true" pageId="9" pageNumber="101">S. pawensis</emphasis>
Bohlen &amp; Šlechtová
</taxonomicName>
(from 
<collectingRegion id="49FCF845FF9F6A6AFE09FB268F3140A8" box="[463,578,1166,1192]" country="Myanmar" name="Ayeyawady" pageId="9" pageNumber="101">Irrawaddy</collectingRegion>
drainage in 
<collectingCountry id="F32F7637FF9F6A6AFD14FB278E3240A9" box="[722,833,1167,1193]" name="Myanmar" pageId="9" pageNumber="101">Myanmar</collectingCountry>
) too, have a colour pattern that includes a midlateral stripe and a pale belly, but the back is dark brown, with males lacking modified pectoral-fin rays. Besides, 
<taxonomicName id="4C384D24FF9F6A6AFA41FB1B8C4A40F0" authorityName="Bohlen &amp; Slechtova" authorityYear="2013" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="9" pageNumber="101" phylum="Chordata" rank="species" species="rubrimaculata">
<emphasis id="B94CEAB5FF9F6A6AFA41FB1B8C4A40F0" italics="true" pageId="9" pageNumber="101">S. rubrimaculata</emphasis>
</taxonomicName>
has up to 6 saddles on the back and a maximum known size of 
<quantity id="4CC09B42FF9F6A6AFC3EFB7F893540F1" box="[1016,1094,1239,1265]" metricMagnitude="-2" metricUnit="m" metricValue="2.8" pageId="9" pageNumber="101" unit="mm" value="28.0">28 mm</quantity>
SL, while 
<taxonomicName id="4C384D24FF9F6A6AFB78FB7F883240F0" authorityName="Bohlen &amp; Slechtova" authorityYear="2013" box="[1214,1345,1239,1265]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="9" pageNumber="101" phylum="Chordata" rank="species" species="pawensis">
<emphasis id="B94CEAB5FF9F6A6AFB78FB7F883240F0" box="[1214,1345,1239,1265]" italics="true" pageId="9" pageNumber="101">S. pawensis</emphasis>
</taxonomicName>
has 6–7 weakly marked saddles on the back, no suborbital flap, and 7–8 + 7 branched caudal-fin rays (data from 
<bibRefCitation id="EFA94B56FF9F6A6AFAE8FB528C2E4139" author="Bohlen, J. &amp; Slechtova, V." pageId="9" pageNumber="101" pagination="21 - 30" refId="ref5963" refString="Bohlen, J. &amp; Slechtova, V. (2013 a) Two new species of Schistura from Myanmar (Cypriniformes: Nemacheilidae). Ichthyological Exploration of Freshwaters, 24 (1), 21 - 30." type="journal article" year="2013">Bohlen &amp; Šlechtová, 2013a</bibRefCitation>
).
</paragraph>
<paragraph id="8B8736A7FF9F6A6AFF01FAEA8F0C4258" blockId="9.[151,1437,1130,1624]" pageId="9" pageNumber="101">
<taxonomicName id="4C384D24FF9F6A6AFF01FAEA8CCC415C" authority="Kottelat, 2022" box="[199,447,1346,1372]" class="Actinopterygii" family="Nemacheilidae" genus="Nemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="9" pageNumber="101" phylum="Chordata" rank="species" species="pezidion">
<emphasis id="B94CEAB5FF9F6A6AFF01FAEA8CCC415C" box="[199,447,1346,1372]" italics="true" pageId="9" pageNumber="101">Nemacheilus pezidion</emphasis>
</taxonomicName>
was tentatively identified as 
<taxonomicName id="4C384D24FF9F6A6AFCCAFAEB8EC5415C" authorityName="Kottelat" authorityYear="1990" box="[780,950,1346,1372]" class="Actinopterygii" family="Nemacheilidae" genus="Nemacheilus" isUncertain="true" kingdom="Animalia" order="Cypriniformes" pageId="9" pageNumber="101" phylum="Chordata" rank="species" species="pallidus">
<emphasis id="B94CEAB5FF9F6A6AFCCAFAEB8E54415C" box="[780,807,1347,1372]" italics="true" pageId="9" pageNumber="101">N.</emphasis>
aff. 
<emphasis id="B94CEAB5FF9F6A6AFC9AFAEA8EC5415C" box="[860,950,1346,1372]" italics="true" pageId="9" pageNumber="101">pallidus</emphasis>
</taxonomicName>
by 
<bibRefCitation id="EFA94B56FF9F6A6AFC24FAEA89BF415D" author="Kottelat, M." box="[994,1228,1346,1373]" pageId="9" pageNumber="101" pagination="277 - 288" refId="ref7231" refString="Kottelat, M. (2021) ' Nemacheilus' argyrogaster, a new species of loach from southern Laos (Teleostei: Nemacheilidae). Zootaxa, 4933 (2), 277 - 288. https: // doi. org / 10.11646 / zootaxa. 4933.2.6" type="journal article" year="2021">Kottelat (2021: 285)</bibRefCitation>
. In addition to the presence of the bold black midlateral stripe (vs. a midlateral row of 17–19 blackish spots and a middorsal row of 14–17 narrow saddles), 
<taxonomicName id="4C384D24FF9F6A6AFE6DFA238F5E41A4" authority="Kottelat, 2022" box="[427,557,1418,1444]" class="Actinopterygii" family="Nemacheilidae" genus="Nemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="9" pageNumber="101" phylum="Chordata" rank="species" species="pezidion">
<emphasis id="B94CEAB5FF9F6A6AFE6DFA238F5E41A4" box="[427,557,1418,1444]" italics="true" pageId="9" pageNumber="101">N. pezidion</emphasis>
</taxonomicName>
is distinguished from 
<taxonomicName id="4C384D24FF9F6A6AFCF7FA238EDD41A4" authorityName="Kottelat" authorityYear="1990" box="[817,942,1418,1444]" class="Actinopterygii" family="Nemacheilidae" genus="Nemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="9" pageNumber="101" phylum="Chordata" rank="species" species="pallidus">
<emphasis id="B94CEAB5FF9F6A6AFCF7FA238EDD41A4" box="[817,942,1418,1444]" italics="true" pageId="9" pageNumber="101">N. pallidus</emphasis>
</taxonomicName>
(s.s., from Chao Phraya drainage; 
<figureCitation id="13032A22FF9F6A6AFA84FA2288E041A5" box="[1346,1427,1418,1445]" captionStart="FIGURE 11" captionStartId="9.[151,250,2007,2031]" captionTargetBox="[151,1436,1648,1983]" captionTargetId="figure-424@9.[151,1436,1648,1983]" captionTargetPageId="9" captionText="FIGURE 11. Nemacheilus pallidus, ZRC 38468, holotype, 54.0 mm SL; Thailand: Mae Nam Yom." figureDoi="http://doi.org/10.5281/zenodo.6488237" httpUri="https://zenodo.org/record/6488237/files/figure.png" pageId="9" pageNumber="101">Fig. 11</figureCitation>
) by a more slender caudal peduncle (length 17.0–20.3 % SL vs 14.1–16.3; depth 9.3–10.5 % SL vs. 11.6–12.4, 1.7–2.0 (vs. 1.2–1.4) times in its length, and 1.5–1.9 (vs 1.4–1.6) times in depth at dorsal-fin origin). The fish widespread in the Mekong drainage (from at least the Xe Banghiang watershed northwards to at least the Nam Ngum watershed) identified as 
<taxonomicName id="4C384D24FF9F6A6AFE39F9B38F0F4234" authorityName="Kottelat" authorityYear="1990" box="[511,636,1562,1588]" class="Actinopterygii" family="Nemacheilidae" genus="Nemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="9" pageNumber="101" phylum="Chordata" rank="species" species="pallidus">
<emphasis id="B94CEAB5FF9F6A6AFE39F9B38F0F4234" box="[511,636,1562,1588]" italics="true" pageId="9" pageNumber="101">N. pallidus</emphasis>
</taxonomicName>
by 
<bibRefCitation id="EFA94B56FF9F6A6AFD6CF9B28E0D4234" author="Kottelat, M." box="[682,894,1562,1589]" pageId="9" pageNumber="101" refId="ref6499" refString="Kottelat, M. (1990) Indochinese nemacheilines. A revision of nemacheiline loaches (Pisces: Cypriniformes) of Thailand, Burma, Laos, Cambodia and southern Viet Nam. Pfeil, Munchen, 262 pp." type="book" year="1990">Kottelat (1990: 63</bibRefCitation>
, fig. 36b, 1998: 69, 2001: 100), is an unnamed species whose description is in preparation.
</paragraph>
<caption id="DF47662FFF9F6A6AFF51F87F89FA43EF" ID-DOI="http://doi.org/10.5281/zenodo.6488237" ID-Zenodo-Dep="6488237" box="[151,1161,2007,2031]" httpUri="https://zenodo.org/record/6488237/files/figure.png" pageId="9" pageNumber="101" startId="9.[151,250,2007,2031]" targetBox="[151,1436,1648,1983]" targetPageId="9">
<paragraph id="8B8736A7FF9F6A6AFF51F87F89FA43EF" blockId="9.[151,1161,2007,2031]" box="[151,1161,2007,2031]" pageId="9" pageNumber="101">
<emphasis id="B94CEAB5FF9F6A6AFF51F87F8C6D43EF" bold="true" box="[151,286,2007,2031]" pageId="9" pageNumber="101">FIGURE 11.</emphasis>
<taxonomicName id="4C384D24FF9F6A6AFEE3F87F8F4E43EF" authority=", ZRC" authorityName="ZRC" box="[293,573,2007,2031]" class="Actinopterygii" family="Nemacheilidae" genus="Nemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="9" pageNumber="101" phylum="Chordata" rank="species" species="pallidus">
<emphasis id="B94CEAB5FF9F6A6AFEE3F87F8F7343EF" box="[293,512,2007,2031]" italics="true" pageId="9" pageNumber="101">Nemacheilus pallidus</emphasis>
, ZRC
</taxonomicName>
38468, holotype, 54.0 mm SL; Thailand: Mae Nam Yom.
</paragraph>
</caption>
<paragraph id="8B8736A7FF9C6A69FF01FF3E88EE44B1" blockId="10.[151,1437,150,213]" box="[199,1437,150,177]" pageId="10" pageNumber="102">
<emphasis id="B94CEAB5FF9C6A69FF01FF3E8CBF44B0" bold="true" box="[199,460,150,176]" pageId="10" pageNumber="102">Comparison material.</emphasis>
List not exhaustive. 
<taxonomicName id="4C384D24FF9C6A69FD69FF3F894944B1" authority=": CMK 1768" authorityName="CMK" authorityYear="1768" box="[687,1082,151,177]" class="Actinopterygii" family="Nemacheilidae" genus="Nemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="10" pageNumber="102" phylum="Chordata" rank="species" species="binotatus">
<emphasis id="B94CEAB5FF9C6A69FD69FF3F8EDE44B1" box="[687,941,151,177]" italics="true" pageId="10" pageNumber="102">Nemacheilus binotatus</emphasis>
: CMK 1768
</taxonomicName>
, 6, 
<quantity id="4CC09B42FF9C6A69FB98FF3F898C44B1" box="[1118,1279,151,177]" metricMagnitude="-2" metricUnit="m" metricValue="3.63" metricValueMax="3.9799999999999995" metricValueMin="3.28" pageId="10" pageNumber="102" unit="mm" value="36.3" valueMax="39.8" valueMin="32.8">32.8–39.8 mm</quantity>
SL; 
<collectingCountry id="F32F7637FF9C6A69FAF7FF3F88EA44B1" box="[1329,1433,151,177]" name="Thailand" pageId="10" pageNumber="102">Thailand</collectingCountry>
:
</paragraph>
<paragraph id="8B8736A7FF9C6A69FF51FF138E3144D5" blockId="10.[151,1437,150,213]" box="[151,834,187,213]" pageId="10" pageNumber="102">
<materialsCitation id="3B503CFAFF9C6A69FF51FF138E4D44D5" ID-GBIF-Occurrence="3757640315" box="[151,830,187,213]" location="Mae Nam Ping in Chiang Dao" pageId="10" pageNumber="102" specimenCount="1" stateProvince="Chiang Mai">
<collectingRegion id="49FCF845FF9C6A69FF51FF138CF844D5" box="[151,395,187,213]" country="Thailand" name="Chiang Mai" pageId="10" pageNumber="102">Chiang Mai Province</collectingRegion>
: 
<location id="8EE7607CFF9C6A69FE50FF138F9944D5" LSID="urn:lsid:plazi:treatment:039187B1FF976A69FF51FF3F8FDB4739:8EE7607CFF9C6A69FE50FF138F9944D5" box="[406,746,187,213]" name="Mae Nam Ping in Chiang Dao" pageId="10" pageNumber="102" stateProvince="Chiang Mai">Mae Nam Ping in Chiang Dao</location>
gorges
</materialsCitation>
.
</paragraph>
<paragraph id="8B8736A7FF9C6A69FF51FEAB8CB545F5" blockId="10.[151,1437,258,825]" pageId="10" pageNumber="102">
<taxonomicName id="4C384D24FF9C6A69FF51FEAB8CE4451D" authority=": CMK" authorityName="CMK" box="[151,407,259,285]" class="Actinopterygii" family="Nemacheilidae" genus="Nemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="10" pageNumber="102" phylum="Chordata" rank="species" species="longistriatus">
<emphasis id="B94CEAB5FF9C6A69FF51FEAB8C34451D" box="[151,327,259,285]" italics="true" pageId="10" pageNumber="102">N. longistriatus</emphasis>
: CMK
</taxonomicName>
5402, 
<specimenCount id="9D3EFD2EFF9C6A69FE20FEAB8F19451D" box="[486,618,259,285]" pageId="10" pageNumber="102" type="generic" typeStatus="paratypes">4 paratypes</specimenCount>
, 36.0– 
<quantity id="4CC09B42FF9C6A69FD70FEAA8E68451D" box="[694,795,258,285]" metricMagnitude="-2" metricUnit="m" metricValue="4.65" pageId="10" pageNumber="102" unit="mm" value="46.5">46.5 mm</quantity>
SL; 
<materialsCitation id="3B503CFAFF9C6A69FC95FEAB896F4541" ID-GBIF-Occurrence="3757640320" accessionNumber="CMK 13704" collectionCode="CMK" country="Thailand" location="Chiang Khan" municipality="Mekong" pageId="10" pageNumber="102" specimenCode="22" specimenCount="1" stateProvince="Loei">
<collectingCountry id="F32F7637FF9C6A69FC95FEAB8EC8451D" box="[851,955,259,285]" name="Thailand" pageId="10" pageNumber="102">Thailand</collectingCountry>
: 
<collectingRegion id="49FCF845FF9C6A69FC00FEAB8919451D" box="[966,1130,259,285]" country="Thailand" name="Loei" pageId="10" pageNumber="102">Loei Province</collectingRegion>
: 
<collectingMunicipality id="6BE3ACDDFF9C6A69FBB3FEAB89A1451D" box="[1141,1234,259,285]" pageId="10" pageNumber="102">Mekong</collectingMunicipality>
mainstream from 
<location id="8EE7607CFF9C6A69FF00FE8F8C2E4541" LSID="urn:lsid:plazi:treatment:039187B1FF976A69FF51FF3F8FDB4739:8EE7607CFF9C6A69FF00FE8F8C2E4541" box="[198,349,295,321]" country="Thailand" municipality="Mekong" name="Chiang Khan" pageId="10" pageNumber="102" stateProvince="Loei">Chiang Khan</location>
to 
<quantity id="4CC09B42FF9C6A69FE45FE8F8CBF4541" box="[387,460,295,321]" metricMagnitude="4" metricUnit="m" metricValue="7.0" pageId="10" pageNumber="102" unit="km" value="70.0">70 km</quantity>
downstream.— 
<collectionCode id="ED29AE62FF9C6A69FD46FE8F8FB24541" box="[640,705,295,321]" pageId="10" pageNumber="102">CMK</collectionCode>
13704, 
<specimenCode id="DB9E9EDCFF9C6A69FCD9FE8F8E4D4541" box="[799,830,295,321]" collectionCode="CMK" pageId="10" pageNumber="102">22</specimenCode>
, 
<quantity id="4CC09B42FF9C6A69FC8DFE8F8E9C4541" box="[843,1007,295,321]" metricMagnitude="-2" metricUnit="m" metricValue="3.47" metricValueMax="3.88" metricValueMin="3.06" pageId="10" pageNumber="102" unit="mm" value="34.7" valueMax="38.8" valueMin="30.6">30.6–38.8 mm</quantity>
SL
</materialsCitation>
; 
<materialsCitation id="3B503CFAFF9C6A69FBEFFE8F89C74565" ID-GBIF-Occurrence="3757640304" accessionNumber="CMK 13777" collectionCode="CMK" collectorName="Xepon." country="Laos" location="Xe Bang Hiang" pageId="10" pageNumber="102" specimenCode="7" specimenCount="1" stateProvince="Savannakhet">
<collectingCountry id="F32F7637FF9C6A69FBEFFE8F89174541" box="[1065,1124,295,321]" name="Laos" pageId="10" pageNumber="102">Laos</collectingCountry>
: 
<collectingRegion id="49FCF845FF9C6A69FBB6FE8F88034541" box="[1136,1392,295,321]" country="Laos" name="Savannakhet" pageId="10" pageNumber="102">Savannakhet Province</collectingRegion>
: 
<location id="8EE7607CFF9C6A69FABAFE8F8C204565" LSID="urn:lsid:plazi:treatment:039187B1FF976A69FF51FF3F8FDB4739:8EE7607CFF9C6A69FABAFE8F8C204565" country="Laos" name="Xe Bang Hiang" pageId="10" pageNumber="102" stateProvince="Savannakhet">Xe Bang Hiang</location>
, about 
<quantity id="4CC09B42FF9C6A69FE6BFEE38C994565" box="[429,490,331,357]" metricMagnitude="3" metricUnit="m" metricValue="6.0" pageId="10" pageNumber="102" unit="km" value="6.0">6 km</quantity>
downstream of 
<collectorName id="26CD5371FF9C6A69FD6AFEE38F8F4565" box="[684,764,331,357]" pageId="10" pageNumber="102">Xepon.</collectorName>
— 
<collectionCode id="ED29AE62FF9C6A69FCDFFEE38E294565" box="[793,858,331,357]" pageId="10" pageNumber="102">CMK</collectionCode>
13777, 
<specimenCode id="DB9E9EDCFF9C6A69FC7BFEE38EBE4565" box="[957,973,331,357]" collectionCode="CMK" pageId="10" pageNumber="102">7</specimenCode>
, 
<quantity id="4CC09B42FF9C6A69FC1BFEE389F64565" box="[989,1157,331,357]" metricMagnitude="-2" metricUnit="m" metricValue="3.4849999999999994" metricValueMax="3.7299999999999995" metricValueMin="3.2399999999999998" pageId="10" pageNumber="102" unit="mm" value="34.849999999999994" valueMax="37.3" valueMin="32.4">32.4–37.3 mm</quantity>
SL
</materialsCitation>
; 
<materialsCitation id="3B503CFAFF9C6A69FB02FEE389614589" ID-GBIF-Occurrence="3757640301" accessionNumber="CMK 15950" collectionCode="CMK" collectorName="Ban Tat Hai Xe." country="Laos" location="Xe Bang Hiang" pageId="10" pageNumber="102" specimenCode="1" specimenCount="1" stateProvince="Savannakhet">
<collectingCountry id="F32F7637FF9C6A69FB02FEE3898C4565" box="[1220,1279,331,357]" name="Laos" pageId="10" pageNumber="102">Laos</collectingCountry>
: 
<collectingRegion id="49FCF845FF9C6A69FACBFEE38C5D4589" country="Laos" name="Savannakhet" pageId="10" pageNumber="102">Savannakhet Province</collectingRegion>
: 
<location id="8EE7607CFF9C6A69FEF1FEC78C904589" LSID="urn:lsid:plazi:treatment:039187B1FF976A69FF51FF3F8FDB4739:8EE7607CFF9C6A69FEF1FEC78C904589" box="[311,483,367,393]" country="Laos" name="Xe Bang Hiang" pageId="10" pageNumber="102" stateProvince="Savannakhet">Xe Bang Hiang</location>
at 
<collectorName id="26CD5371FF9C6A69FDC2FEC78FC04589" box="[516,691,367,393]" pageId="10" pageNumber="102">Ban Tat Hai Xe.</collectorName>
— 
<collectionCode id="ED29AE62FF9C6A69FD16FEC78E624589" box="[720,785,367,393]" pageId="10" pageNumber="102">CMK</collectionCode>
15950, 
<specimenCode id="DB9E9EDCFF9C6A69FCADFEC78E0A4589" box="[875,889,367,393]" collectionCode="CMK" pageId="10" pageNumber="102">1</specimenCode>
, 
<quantity id="4CC09B42FF9C6A69FC40FEC78E9A4589" box="[902,1001,367,393]" metricMagnitude="-2" metricUnit="m" metricValue="6.72" pageId="10" pageNumber="102" unit="mm" value="67.2">67.2 mm</quantity>
SL
</materialsCitation>
; 
<materialsCitation id="3B503CFAFF9C6A69FBDBFEC78EB245AD" ID-GBIF-Occurrence="3757640309" accessionNumber="CMK 19312" collectionCode="CMK" collectorName="Nam Ngiep." country="Laos" location="Nam Xao" pageId="10" pageNumber="102" specimenCode="7" specimenCount="1" stateProvince="Bolikhamxai">
<collectingCountry id="F32F7637FF9C6A69FBDBFEC7892B4589" box="[1053,1112,367,393]" name="Laos" pageId="10" pageNumber="102">Laos</collectingCountry>
: 
<collectingRegion id="49FCF845FF9C6A69FBA7FEC7882E4589" box="[1121,1373,367,393]" country="Laos" name="Bolikhamxai" pageId="10" pageNumber="102">Bolikhamsai Province</collectingRegion>
: 
<location id="8EE7607CFF9C6A69FAA0FEC78D8445AD" LSID="urn:lsid:plazi:treatment:039187B1FF976A69FF51FF3F8FDB4739:8EE7607CFF9C6A69FAA0FEC78D8445AD" country="Laos" name="Nam Xao" pageId="10" pageNumber="102" stateProvince="Bolikhamxai">Nam Xao</location>
, a tributary of 
<collectorName id="26CD5371FF9C6A69FE5AFE3B8F5045AD" box="[412,547,403,429]" pageId="10" pageNumber="102">Nam Ngiep.</collectorName>
— 
<collectionCode id="ED29AE62FF9C6A69FD86FE3B8FF245AD" box="[576,641,403,429]" pageId="10" pageNumber="102">CMK</collectionCode>
19312, 
<specimenCode id="DB9E9EDCFF9C6A69FD1AFE3B8F9F45AD" box="[732,748,403,429]" collectionCode="CMK" pageId="10" pageNumber="102">7</specimenCode>
, 
<quantity id="4CC09B42FF9C6A69FD3EFE3A8EE545AD" box="[760,918,402,429]" metricMagnitude="-2" metricUnit="m" metricValue="3.315" metricValueMax="3.65" metricValueMin="2.98" pageId="10" pageNumber="102" unit="mm" value="33.15" valueMax="36.5" valueMin="29.8">29.8-36.5 mm</quantity>
SL
</materialsCitation>
; 
<materialsCitation id="3B503CFAFF9C6A69FC0BFE3B8E3C45D1" ID-GBIF-Occurrence="3757640311" accessionNumber="CMK 23085" collectionCode="CMK" collectorName="Keng Mou Man." country="Laos" location="Xe Bangfai" pageId="10" pageNumber="102" specimenCode="56" specimenCount="1" stateProvince="Khammouan">
<collectingCountry id="F32F7637FF9C6A69FC0BFE3B897B45AD" box="[973,1032,403,429]" name="Laos" pageId="10" pageNumber="102">Laos</collectingCountry>
: 
<collectingRegion id="49FCF845FF9C6A69FBD4FE3B886245AD" box="[1042,1297,403,429]" country="Laos" name="Khammouan" pageId="10" pageNumber="102">Khammouan Province</collectingRegion>
: 
<location id="8EE7607CFF9C6A69FADAFE3B88EE45AD" LSID="urn:lsid:plazi:treatment:039187B1FF976A69FF51FF3F8FDB4739:8EE7607CFF9C6A69FADAFE3B88EE45AD" box="[1308,1437,403,429]" country="Laos" name="Xe Bangfai" pageId="10" pageNumber="102" stateProvince="Khammouan">Xe Bangfai</location>
at 
<collectorName id="26CD5371FF9C6A69FF24FE1F8CE845D1" box="[226,411,439,465]" pageId="10" pageNumber="102">Keng Mou Man.</collectorName>
— 
<collectionCode id="ED29AE62FF9C6A69FE7EFE1F8C8A45D1" box="[440,505,439,465]" pageId="10" pageNumber="102">CMK</collectionCode>
23085, 
<specimenCode id="DB9E9EDCFF9C6A69FD90FE1E8F0745D1" box="[598,628,438,465]" collectionCode="CMK" pageId="10" pageNumber="102">56</specimenCode>
, 
<quantity id="4CC09B42FF9C6A69FD46FE1F8E5745D1" box="[640,804,439,465]" metricMagnitude="-2" metricUnit="m" metricValue="3.815" metricValueMax="4.66" metricValueMin="2.9699999999999998" pageId="10" pageNumber="102" unit="mm" value="38.15" valueMax="46.6" valueMin="29.7">29.7–46.6 mm</quantity>
SL
</materialsCitation>
; 
<materialsCitation id="3B503CFAFF9C6A69FC9DFE1F8CB045F5" ID-GBIF-Occurrence="3757640307" country="Laos" location="Savannakhet Province" pageId="10" pageNumber="102" specimenCount="1" stateProvince="Savannakhet">
<collectingCountry id="F32F7637FF9C6A69FC9DFE1F8EE545D1" box="[859,918,439,465]" name="Laos" pageId="10" pageNumber="102">Laos</collectingCountry>
: 
<collectingRegion id="49FCF845FF9C6A69FC67FE1F89D345D1" box="[929,1184,439,465]" country="Laos" name="Savannakhet" pageId="10" pageNumber="102">Savannakhet Province</collectingRegion>
: 
<collectorName id="26CD5371FF9C6A69FB6DFE1F885E45D1" box="[1195,1325,439,465]" pageId="10" pageNumber="102">Xe Bangfai</collectorName>
drainage: 
<collectorName id="26CD5371FF9C6A69FF00FE738C6D45F5" box="[198,286,475,501]" pageId="10" pageNumber="102">Xe Noy</collectorName>
at 
<collectorName id="26CD5371FF9C6A69FE86FE738CB045F5" box="[320,451,475,501]" pageId="10" pageNumber="102">Keng Boua</collectorName>
</materialsCitation>
.
</paragraph>
<paragraph id="8B8736A7FF9C6A69FF51FE5789AD4619" blockId="10.[151,1437,258,825]" box="[151,1246,510,537]" pageId="10" pageNumber="102">
<taxonomicName id="4C384D24FF9C6A69FF51FE578CE94619" authority=": CMK" authorityName="CMK" box="[151,410,511,537]" class="Actinopterygii" family="Nemacheilidae" genus="Nemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="10" pageNumber="102" phylum="Chordata" rank="species" species="nandingensis">
<emphasis id="B94CEAB5FF9C6A69FF51FE578C3F4619" box="[151,332,511,537]" italics="true" pageId="10" pageNumber="102">N. nandingensis</emphasis>
: 
<materialsCitation id="3B503CFAFF9C6A69FE9CFE578CE94619" ID-GBIF-Occurrence="3757640308" accessionNumber="CMK 23949" box="[346,410,511,537]" country="China" location="Nam Xing River" municipality="Xishuangbanna" pageId="10" pageNumber="102" specimenCount="1" stateProvince="Yunnan">CMK</materialsCitation>
</taxonomicName>
23949, 1, 
<quantity id="4CC09B42FF9C6A69FDD7FE568F064619" box="[529,629,510,537]" metricMagnitude="-2" metricUnit="m" metricValue="5.66" pageId="10" pageNumber="102" unit="mm" value="56.6">56.6 mm</quantity>
SL; 
<collectingCountry id="F32F7637FF9C6A69FD6DFE578F814619" box="[683,754,511,537]" name="China" pageId="10" pageNumber="102">China</collectingCountry>
: 
<collectingRegion id="49FCF845FF9C6A69FD3AFE578E254619" box="[764,854,511,537]" country="China" name="Yunnan" pageId="10" pageNumber="102">Yunnan</collectingRegion>
: 
<collectingMunicipality id="6BE3ACDDFF9C6A69FCA6FE5789604619" box="[864,1043,511,537]" pageId="10" pageNumber="102">Xishuangbanna</collectingMunicipality>
: 
<location id="8EE7607CFF9C6A69FBDBFE5789A94619" LSID="urn:lsid:plazi:treatment:039187B1FF976A69FF51FF3F8FDB4739:8EE7607CFF9C6A69FBDBFE5789A94619" box="[1053,1242,511,537]" country="China" municipality="Xishuangbanna" name="Nam Xing River" pageId="10" pageNumber="102" stateProvince="Yunnan">Nam Xing River</location>
.
</paragraph>
<paragraph id="8B8736A7FF9C6A69FF51FD8B8804463D" blockId="10.[151,1437,258,825]" box="[151,1399,546,573]" pageId="10" pageNumber="102">
<taxonomicName id="4C384D24FF9C6A69FF51FD8B8C2F463D" authority=": CMK" authorityName="CMK" box="[151,348,547,573]" class="Actinopterygii" family="Nemacheilidae" genus="Nemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="10" pageNumber="102" phylum="Chordata" rank="species" species="zonatus">
<emphasis id="B94CEAB5FF9C6A69FF51FD8B8C7D463C" box="[151,270,547,572]" italics="true" pageId="10" pageNumber="102">N. zonatus</emphasis>
: 
<materialsCitation id="3B503CFAFF9C6A69FEDAFD8B8C2F463D" ID-GBIF-Occurrence="3757640302" accessionNumber="CMK 19373" box="[284,348,547,573]" country="Laos" location="Ban Tha Ponsaoe" municipality="Xe Bangfai" pageId="10" pageNumber="102" specimenCount="1" stateProvince="Khammouan">CMK</materialsCitation>
</taxonomicName>
19373, 13, 
<quantity id="4CC09B42FF9C6A69FE27FD8A8FF7463D" box="[481,644,546,573]" metricMagnitude="-2" metricUnit="m" metricValue="2.7649999999999997" metricValueMax="3.02" metricValueMin="2.51" pageId="10" pageNumber="102" unit="mm" value="27.65" valueMax="30.2" valueMin="25.1">25.1–30.2 mm</quantity>
SL; 
<collectingCountry id="F32F7637FF9C6A69FD7CFD8B8F86463D" box="[698,757,547,573]" name="Laos" pageId="10" pageNumber="102">Laos</collectingCountry>
: 
<collectingRegion id="49FCF845FF9C6A69FD39FD8B8E8C463D" box="[767,1023,547,573]" country="Laos" name="Khammouan" pageId="10" pageNumber="102">Khammouan Province</collectingRegion>
: 
<collectingMunicipality id="6BE3ACDDFF9C6A69FBCFFD8B89F9463D" box="[1033,1162,547,573]" pageId="10" pageNumber="102">Xe Bangfai</collectingMunicipality>
at 
<location id="8EE7607CFF9C6A69FB6AFD8B8800463D" LSID="urn:lsid:plazi:treatment:039187B1FF976A69FF51FF3F8FDB4739:8EE7607CFF9C6A69FB6AFD8B8800463D" box="[1196,1395,547,573]" country="Laos" municipality="Xe Bangfai" name="Ban Tha Ponsaoe" pageId="10" pageNumber="102" stateProvince="Khammouan">Ban Tha Ponsaoe</location>
.
</paragraph>
<paragraph id="8B8736A7FF9C6A69FF51FDEF8F3E4685" blockId="10.[151,1437,258,825]" pageId="10" pageNumber="102">
<taxonomicName id="4C384D24FF9C6A69FF51FDEF8C174661" authority=": CMK" authorityName="CMK" box="[151,356,583,609]" class="Actinopterygii" family="Nemacheilidae" genus="Nemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="10" pageNumber="102" phylum="Chordata" rank="species" species="pallidus">
<emphasis id="B94CEAB5FF9C6A69FF51FDEF8C674661" box="[151,276,583,609]" italics="true" pageId="10" pageNumber="102">N. pallidus</emphasis>
: CMK
</taxonomicName>
4835, 
<specimenCount id="9D3EFD2EFF9C6A69FE72FDEF8F4A4661" box="[436,569,583,609]" pageId="10" pageNumber="102" type="generic" typeStatus="paratypes">7 paratypes</specimenCount>
, 34.9–50.0 mm SL; 
<collectingCountry id="F32F7637FF9C6A69FCE5FDEF8EF84661" box="[803,907,583,609]" name="Thailand" pageId="10" pageNumber="102">Thailand</collectingCountry>
: 
<collectingRegion id="49FCF845FF9C6A69FC51FDEF89014661" box="[919,1138,583,609]" country="Thailand" name="Lampang" pageId="10" pageNumber="102">Lampang Province</collectingRegion>
: Mae Nam Yom drainage: Huai Mae Phlung near Ban Phong.
</paragraph>
<paragraph id="8B8736A7FF9C6A69FF51FD27880146CD" blockId="10.[151,1437,258,825]" pageId="10" pageNumber="102">
<taxonomicName id="4C384D24FF9C6A69FF51FD278C2946A9" authority=": CMK" authorityName="CMK" box="[151,346,655,681]" class="Actinopterygii" family="Nemacheilidae" genus="Nemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="10" pageNumber="102" phylum="Chordata" rank="species" species="masyae">
<emphasis id="B94CEAB5FF9C6A69FF51FD278C7F46A8" box="[151,268,655,680]" italics="true" pageId="10" pageNumber="102">N. masyae</emphasis>
: CMK
</taxonomicName>
5180, 7, 
<quantity id="4CC09B42FF9C6A69FE04FD268F1646A9" box="[450,613,654,681]" metricMagnitude="-2" metricUnit="m" metricValue="5.695" metricValueMax="6.58" metricValueMin="4.81" pageId="10" pageNumber="102" unit="mm" value="56.95" valueMax="65.8" valueMin="48.1">48.1–65.8 mm</quantity>
SL; 
<materialsCitation id="3B503CFAFF9C6A69FD5CFD278C8E46CD" ID-GBIF-Occurrence="3757640310" collectionCode="CMK" country="Thailand" location="Tapi River" pageId="10" pageNumber="102" specimenCode="CMK 5165, 1" specimenCount="1" stateProvince="Surat Thani">
<collectingCountry id="F32F7637FF9C6A69FD5CFD278E7146A9" box="[666,770,655,681]" name="Thailand" pageId="10" pageNumber="102">Thailand</collectingCountry>
: 
<collectingRegion id="49FCF845FF9C6A69FCCAFD278E8E46A9" box="[780,1021,655,681]" country="Thailand" name="Surat Thani" pageId="10" pageNumber="102">Surat Thani Province</collectingRegion>
: 
<location id="8EE7607CFF9C6A69FBC1FD27890E46A9" LSID="urn:lsid:plazi:treatment:039187B1FF976A69FF51FF3F8FDB4739:8EE7607CFF9C6A69FBC1FD27890E46A9" box="[1031,1149,655,681]" country="Thailand" name="Tapi River" pageId="10" pageNumber="102" stateProvince="Surat Thani">Tapi River</location>
drainage: Khlong Sok.— 
<specimenCode id="DB9E9EDCFF9C6A69FF00FD1B8C3B46CD" box="[198,328,690,717]" collectionCode="CMK" pageId="10" pageNumber="102">CMK 5165</specimenCode>
, 
<specimenCode id="DB9E9EDCFF9C6A69FE95FD1B8C1246CD" box="[339,353,691,717]" collectionCode="CMK" pageId="10" pageNumber="102">1</specimenCode>
, 
<quantity id="4CC09B42FF9C6A69FEA9FD1A8CA046CD" box="[367,467,690,717]" metricMagnitude="-2" metricUnit="m" metricValue="6.15" pageId="10" pageNumber="102" unit="mm" value="61.5">61.5 mm</quantity>
SL
</materialsCitation>
; 
<materialsCitation id="3B503CFAFF9C6A69FDCEFD1B881D46CD" ID-GBIF-Occurrence="3757640306" box="[520,1390,691,717]" country="Thailand" location="Surat Thani Province" pageId="10" pageNumber="102" specimenCount="1" stateProvince="Surat Thani">
<collectingCountry id="F32F7637FF9C6A69FDCEFD1B8F0346CD" box="[520,624,691,717]" name="Thailand" pageId="10" pageNumber="102">Thailand</collectingCountry>
: 
<collectingRegion id="49FCF845FF9C6A69FDBDFD1B8E1F46CD" box="[635,876,691,717]" country="Thailand" name="Surat Thani" pageId="10" pageNumber="102">Surat Thani Province</collectingRegion>
: Khlong Phum Duang, upstream of Phunphin
</materialsCitation>
.
</paragraph>
<paragraph id="8B8736A7FF9C6A69FF51FD7F8FDB4739" blockId="10.[151,1437,258,825]" pageId="10" pageNumber="102">
<emphasis id="B94CEAB5FF9C6A69FF51FD7F8C2A46F0" box="[151,345,727,753]" italics="true" pageId="10" pageNumber="102">‘N.’ argyrogaster</emphasis>
: 
<materialsCitation id="3B503CFAFF9C6A69FEAFFD7F8E3246F1" ID-GBIF-Occurrence="3757640316" box="[361,833,726,753]" collectionCode="MHNG" pageId="10" pageNumber="102" specimenCode="MHNG 2785.032" specimenCount="1" typeStatus="holotype">
<specimenCode id="DB9E9EDCFF9C6A69FEAFFD7F8F4046F1" box="[361,563,726,753]" collectionCode="MHNG" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="10" pageNumber="102" type="Museum">MHNG 2785.032</specimenCode>
, 
<typeStatus id="54838805FF9C6A69FDF9FD7F8FD746F1" box="[575,676,727,753]" pageId="10" pageNumber="102" type="holotype">holotype</typeStatus>
, 63.0 mm SL
</materialsCitation>
; 
<materialsCitation id="3B503CFAFF9C6A69FC88FD7F885A46F1" ID-GBIF-Occurrence="3757640318" accessionNumber="CMK 21663" box="[846,1321,726,753]" collectionCode="CMK, ZRC" pageId="10" pageNumber="102" specimenCode="26" specimenCount="1" typeStatus="holotype">
<collectionCode id="ED29AE62FF9C6A69FC88FD7F8EFD46F1" box="[846,910,727,753]" pageId="10" pageNumber="102">CMK</collectionCode>
21663, 
<specimenCode id="DB9E9EDCFF9C6A69FC2DFD7F897A46F1" box="[1003,1033,727,753]" collectionCode="CMK" pageId="10" pageNumber="102">26</specimenCode>
, 
<quantity id="4CC09B42FF9C6A69FBD0FD7E89C846F1" box="[1046,1211,726,753]" metricMagnitude="-2" metricUnit="m" metricValue="6.11" metricValueMax="7.69" metricValueMin="4.529999999999999" pageId="10" pageNumber="102" unit="mm" value="61.1" valueMax="76.9" valueMin="45.3">45.3–76.9 mm</quantity>
SL; 
<collectionCode id="ED29AE62FF9C6A69FB35FD7F885A46F1" box="[1267,1321,727,753]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="10" pageNumber="102">ZRC</collectionCode>
</materialsCitation>
<materialsCitation id="3B503CFAFF9C6A69FAF4FD7F8FD74739" ID-GBIF-Occurrence="3757640321" collectionCode="ZRC" country="Laos" county="Ban Kaleum" location="Mekong" municipality="Houai Peung" pageId="10" pageNumber="102" specimenCode="6" specimenCount="1" typeStatus="holotype">
61628, 
<specimenCode id="DB9E9EDCFF9C6A69FA41FD7F88E446F1" box="[1415,1431,727,753]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="10" pageNumber="102">6</specimenCode>
, 
<quantity id="4CC09B42FF9C6A69FF00FD528C194715" box="[198,362,762,789]" metricMagnitude="-2" metricUnit="m" metricValue="5.37" metricValueMax="6.18" metricValueMin="4.56" pageId="10" pageNumber="102" unit="mm" value="53.7" valueMax="61.8" valueMin="45.6">45.6–61.8 mm</quantity>
SL; 
<collectingCountry id="F32F7637FF9C6A69FE67FD538CAF4715" box="[417,476,763,789]" name="Laos" pageId="10" pageNumber="102">Laos</collectingCountry>
: 
<location id="8EE7607CFF9C6A69FE21FD538F374715" LSID="urn:lsid:plazi:treatment:039187B1FF976A69FF51FF3F8FDB4739:8EE7607CFF9C6A69FE21FD538F374715" box="[487,580,763,789]" country="Laos" county="Ban Kaleum" municipality="Houai Peung" name="Mekong" pageId="10" pageNumber="102">Mekong</location>
drainage: 
<location id="8EE7607CFF9C6A69FD7AFD538ED24715" LSID="urn:lsid:plazi:treatment:039187B1FF976A69FF51FF3F8FDB4739:8EE7607CFF9C6A69FD7AFD538ED24715" box="[700,929,763,789]" country="Laos" county="Ban Kaleum" municipality="Houai Peung" name="Xe Kong watershed" pageId="10" pageNumber="102">Xe Kong watershed</location>
: 
<location id="8EE7607CFF9C6A69FC6AFD53892A4715" LSID="urn:lsid:plazi:treatment:039187B1FF976A69FF51FF3F8FDB4739:8EE7607CFF9C6A69FC6AFD53892A4715" box="[940,1113,763,789]" country="Laos" county="Ban Kaleum" municipality="Houai Peung" name="Xe Kong Prov." pageId="10" pageNumber="102">Xe Kong Prov.</location>
: confluence of 
<location id="8EE7607CFF9C6A69FAC3FD53881F4715" LSID="urn:lsid:plazi:treatment:039187B1FF976A69FF51FF3F8FDB4739:8EE7607CFF9C6A69FAC3FD53881F4715" box="[1285,1388,763,789]" country="Laos" county="Ban Kaleum" municipality="Houai Peung" name="Xe Kong" pageId="10" pageNumber="102">Xe Kong</location>
and 
<collectingMunicipality id="6BE3ACDDFF9C6A69FF00FCB78C294739" box="[198,346,799,825]" pageId="10" pageNumber="102">Houai Peung</collectingMunicipality>
, downstream of 
<collectingCounty id="62E64E2BFF9C6A69FDD2FCB78FD74739" box="[532,676,799,825]" pageId="10" pageNumber="102">Ban Kaleum</collectingCounty>
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>