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<document id="8BD5CD4AF3CA93C2CAD8349496A6049F" ID-DOI="10.11646/zootaxa.5325.4.9" ID-ISSN="1175-5326" ID-Zenodo-Dep="8243787" ID-ZooBank="1E98E639-F78B-41D9-BDE9-39F075A4644F" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1691943785067" checkinUser="plazi" docAuthor="Guo, Liang, Huang, Si-Yao, Wang, Min &amp; Ma, Fang-Zhou" docDate="2023" docId="038BC051FD36842BFF513878A7B2FCA4" docLanguage="en" docName="zootaxa.5325.4.9.pdf" docOrigin="Zootaxa 5325 (4)" docSource="http://dx.doi.org/10.11646/zootaxa.5325.4.9" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Chonala laurae subsp. wuyishana Huang, Guo &amp; Wang 2023, ssp. n." docType="treatment" docVersion="2" lastPageNumber="596" masterDocId="FFB2B829FD378428FFC63C41A30CFFF9" masterDocTitle="Description of a new subspecies of Chonala laurae Bozano, 1999 from Southeastern China (Lepidoptera, Nymphalidae, Satyrinae)" masterLastPageNumber="600" masterPageNumber="593" pageNumber="594" updateTime="1691944186330" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="604CD1036BA5D53012722BFB903DD52C" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="E72E32922361E7F0F765FFF3DC603214">
<mods:title id="DEAB2F14A7000C84107A3F8D607785B7">Description of a new subspecies of Chonala laurae Bozano, 1999 from Southeastern China (Lepidoptera, Nymphalidae, Satyrinae)</mods:title>
</mods:titleInfo>
<mods:name id="B647A536F1A06473A6B9BD763E8C175B" type="personal">
<mods:role id="23F4941DBA08EFC18DA894B68F84471E">
<mods:roleTerm id="1998B898EEDD73520B1EB5A7EEB190B9">Author</mods:roleTerm>
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<mods:namePart id="35812A7E02826879D6AECA252D3DF757">Guo, Liang</mods:namePart>
<mods:nameIdentifier id="DD92686EDDFE1C2D62225063EF7EA739" type="ORCID">0009-0004-2436-6151</mods:nameIdentifier>
<mods:affiliation id="B202146A315BB2A9AD81A3D3893FCBF9">Department of Entomology, College of Plant Protection, South China Agricultural University, Guangzhou 510642, Guangdong, China &amp; guoljxty @ 163. com; https: // orcid. org / 0009 - 0004 - 2436 - 6151</mods:affiliation>
</mods:name>
<mods:name id="EA06BFF85F5BDA80E28F4C1B2981D49A" type="personal">
<mods:role id="79D08ABE7184985ED4DA52084942A0BC">
<mods:roleTerm id="969B02C358D1BAFE4CAD8AE05C555CA1">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="4ACFA9DF3E88350FB483C2A7F65723D9">Huang, Si-Yao</mods:namePart>
<mods:nameIdentifier id="515EBFE5D012D11859DB6E25F13EDCC9" type="ORCID">0000-0002-9859-9212</mods:nameIdentifier>
<mods:affiliation id="5CE975A33FCB718DF37C15B7A6262154">Arthropoda Department, Leibniz Institute for the Analysis of Biodiversity Change, Zoological Research Museum Alexander Koenig, Adenauer Allee 127, Bonn 53113, Germany &amp; S. Huang @ leibniz-lib. de; huangsiyao 2007 @ aliyun. com; https: // orcid. org / 0000 - 0002 - 9859 - 9212</mods:affiliation>
<mods:nameIdentifier id="446278C62C1F55A58886FA53F4407365" type="email">uang@leibniz-lib.de</mods:nameIdentifier>
</mods:name>
<mods:name id="4C18DD32BD82A9A324F78E35A51B6E25" type="personal">
<mods:role id="E77AC46812FCFFAB01653B1DB8A31F13">
<mods:roleTerm id="DD03C7F884652C4DE521982846863449">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="AAFF183658956854930FA192EDEF37A6">Wang, Min</mods:namePart>
<mods:nameIdentifier id="693774904D0987A002C7BCFB3FB0A0B2" type="ORCID">0000-0001-5834-4058</mods:nameIdentifier>
<mods:affiliation id="28AD9F53B3DA8AFE00EC54FA2A5D2164">Department of Entomology, College of Plant Protection, South China Agricultural University, Guangzhou 510642, Guangdong, China &amp; Institute of International Rivers and Eco-Security, Yunnan University, Kunming 650500, Yunnan, China &amp; minwang @ scau. edu. cn; https: // orcid. org / 0000 - 0001 - 5834 - 4058</mods:affiliation>
<mods:nameIdentifier id="FFEBB2B7FBF8D14F92C49F1732E73424" type="email">minwang@scau.edu.cn</mods:nameIdentifier>
</mods:name>
<mods:name id="879AC9742F0DB276226FAF742FBA9BB5" type="personal">
<mods:role id="885FC1494F785CD20A5355C64F7737E7">
<mods:roleTerm id="00357A348842644F8FA95F5B6FBB4C2B">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="72C6D938D0069038BD24C2572B9F1771">Ma, Fang-Zhou</mods:namePart>
<mods:nameIdentifier id="AD6701F7F71B1B6AD833E66794CF66F6" type="ORCID">0000-0001-8219-7967</mods:nameIdentifier>
<mods:affiliation id="C4D1AE8B188BC8945C51B7853D759FDD">State Environmental Protection Scientific Observation and Research Station for Ecology and Environment of Wuyi Mountains, Ministry of Ecology and Environment, Nanjing, 210042, China &amp; Nanjing Institute of Environmental Sciences, Ministry of Ecology and Environment; National Key Laboratory of Biosafety, Nanjing, 210042, China &amp; mfz @ nies. org; https: // orcid. org / 0000 - 0001 - 8219 - 7967</mods:affiliation>
<mods:nameIdentifier id="17B949470CAAE29381646F9D145595B8" type="email">mfz@nies.org</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="8EC37065CB3B7F1B53210C2697C0665F">text</mods:typeOfResource>
<mods:relatedItem id="279DD7BCA296204C48E634FC0F597DC0" type="host">
<mods:titleInfo id="161D110AC82F1FD41EA1F219CFC4D682">
<mods:title id="6BBF9F660E6B24B96892EDEE06B586A2">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="42DEDEEC7AB442FFFD122A7E3103A963">
<mods:date id="E073B5B80E224AB2D8ACFF8027ADC240">2023</mods:date>
<mods:detail id="1C56B7B952463A139229F02AA3F5C6BA" type="pubDate">
<mods:number id="A52E62FC22D6BCF1CD6A675D36021726">2023-08-10</mods:number>
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<mods:number id="4D1A31494EB93070ECD99A8A58710E5C">5325</mods:number>
</mods:detail>
<mods:detail id="9150BD7A3374E7691DB2F1F42680DC82" type="issue">
<mods:number id="B7BDE3F96B3324E5FA06D0EB4CACE58B">4</mods:number>
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<mods:start id="9EB380AB8C387C5CF28F768BC7C6C694">593</mods:start>
<mods:end id="DED69BCE52E88F88CDBED91E872A5212">600</mods:end>
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<mods:location id="4579D2C125D37B03DB76BBA108124582">
<mods:url id="449C63FC51A5ADEF1E9ACC3CEAA9D2E2">http://dx.doi.org/10.11646/zootaxa.5325.4.9</mods:url>
</mods:location>
<mods:classification id="F84F1268E8E5800CBA557C64DD7685E4">journal article</mods:classification>
<mods:identifier id="59E87E8C912779BAC963C58A2A73B5FE" type="DOI">10.11646/zootaxa.5325.4.9</mods:identifier>
<mods:identifier id="4732981F8C1508BB77DEF99DAD30A88A" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="6F0134DAEB6564EFCEA24961C790D865" type="Zenodo-Dep">8243787</mods:identifier>
<mods:identifier id="29E3C5C9F5FBCE99A8B31AF8361862A6" type="ZooBank">1E98E639-F78B-41D9-BDE9-39F075A4644F</mods:identifier>
</mods:mods>
<treatment id="038BC051FD36842BFF513878A7B2FCA4" ID-DOI="http://doi.org/10.5281/zenodo.8243799" ID-Zenodo-Dep="8243799" LSID="urn:lsid:plazi:treatment:038BC051FD36842BFF513878A7B2FCA4" httpUri="http://treatment.plazi.org/id/038BC051FD36842BFF513878A7B2FCA4" lastPageId="3" lastPageNumber="596" pageId="1" pageNumber="594">
<subSubSection id="C33822CCFD368429FF513878A040FBAA" box="[151,844,1081,1108]" pageId="1" pageNumber="594" type="nomenclature">
<paragraph id="8B9D7147FD368429FF513878A040FBAA" blockId="1.[151,844,1081,1143]" box="[151,844,1081,1108]" pageId="1" pageNumber="594">
<heading id="D0D5C62BFD368429FF513878A040FBAA" bold="true" box="[151,844,1081,1108]" fontSize="11" level="1" pageId="1" pageNumber="594" reason="1">
<emphasis id="B956AD55FD368429FF513878A040FBAA" bold="true" box="[151,844,1081,1108]" pageId="1" pageNumber="594">
<taxonomicName id="4C220AC4FD368429FF513878A1FDFBAA" authority="Huang, Guo &amp; Wang" authorityName="Huang, Guo &amp; Wang" authorityYear="2023" box="[151,753,1081,1108]" class="Insecta" family="Nymphalidae" genus="Chonala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="594" phylum="Arthropoda" rank="subSpecies" species="laurae" status="ssp. n." subSpecies="wuyishana">
<emphasis id="B956AD55FD368429FF513878A2D2FBAD" bold="true" box="[151,478,1081,1108]" italics="true" pageId="1" pageNumber="594">Chonala laurae wuyishana</emphasis>
Huang, Guo &amp; Wang
</taxonomicName>
,
<taxonomicNameLabel id="A265102EFD368429FD383878A040FBAA" box="[766,844,1081,1107]" pageId="1" pageNumber="594" rank="subSpecies">ssp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C33822CCFD368429FF51381CA27AFB8E" box="[151,374,1117,1143]" pageId="1" pageNumber="594" type="description">
<paragraph id="8B9D7147FD368429FF51381CA27AFB8E" blockId="1.[151,844,1081,1143]" box="[151,374,1117,1143]" pageId="1" pageNumber="594">
<figureCitation id="13196DC2FD368429FF51381CA3F6FB8E" box="[151,250,1117,1143]" captionStart="FIGURES 1–8" captionStartId="2.[151,264,1943,1967]" captionTargetBox="[203,1384,181,1919]" captionTargetId="figure-21@2.[203,1384,181,1919]" captionTargetPageId="2" captionText="FIGURES 1–8. Male of Chonala sspp. 1–2, C. laurae wuyishana ssp. nov., holotype, Huanggangshan, Mt. Wuyi, Fujian (SCAU); 3–4, ditto, paratype, ditto (SCAU); 5–6, C. laurae laurae, holotype, Qinling, Shaanxi (ZFMK); 7–8, ditto, Shennongjia, Hubei, SATY0700 (CSYL). Scale=10 mm." figureDoi="http://doi.org/10.5281/zenodo.8243789" httpUri="https://zenodo.org/record/8243789/files/figure.png" pageId="1" pageNumber="594">Figs 1–4</figureCitation>
,
<figureCitation id="13196DC2FD368429FEC0381CA242FB8E" box="[262,334,1117,1143]" captionStart="FIGURES 9–16" captionStartId="4.[151,264,1943,1967]" captionTargetBox="[203,1384,181,1919]" captionTargetId="figure-21@4.[203,1384,181,1919]" captionTargetPageId="4" captionText="FIGURES 9–16. Adults of Chonala spp. 9–10, C. houae, holotype, Mt. Jinfo, Chongqing, SATY1156 (CQMNH); 11–12, ditto, paratype, ditto, SATY0466 (CQMNH); 13–14, C. laurae wuyishana ssp. nov., paratype, Mt. Wuyi, Fujian (SCAU); 15–16, C. laurae laurae, Shennongjia, Hubei (CSYL); 9–12 male, 13–16 female. Scale=10 mm." figureDoi="http://doi.org/10.5281/zenodo.8243793" httpUri="https://zenodo.org/record/8243793/files/figure.png" pageId="1" pageNumber="594">13, 14</figureCitation>
,
<figureCitation id="13196DC2FD368429FE9C381CA27AFB8E" box="[346,374,1117,1143]" captionStart="FIGURES 17–20" captionStartId="5.[151,264,1943,1967]" captionTargetBox="[197,1391,194,1907]" captionTargetId="figure-20@5.[184,1403,181,1919]" captionTargetPageId="5" captionText="FIGURES 17–20. Male genitalia of Chonala sspp. Depository of the genitalia: 17 in SCAU, 18 in CSYL, 19 &amp; 20 in CQMNH. a=genitalia capsule with left valva and aedeagus removed; b=aedeagus in dorsal view; c=aedeagus in lateral view; d=left valva in dorsal view. Scale=1 mm." figureDoi="http://doi.org/10.5281/zenodo.8243795" httpUri="https://zenodo.org/record/8243795/files/figure.png" pageId="1" pageNumber="594">17</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C33822CCFD368429FF5138E1A187FA97" pageId="1" pageNumber="594" type="materials_examined">
<paragraph id="8B9D7147FD368429FF5138E1A187FA97" blockId="1.[151,1437,1184,2039]" pageId="1" pageNumber="594">
<emphasis id="B956AD55FD368429FF5138E1A248FB43" bold="true" box="[151,324,1184,1210]" pageId="1" pageNumber="594">Type material.</emphasis>
<materialsCitation id="3B4A7B1AFD368429FE8C38E1A1B5FB27" collectingDate="2022-07-11" collectionCode="SCAU" collectorName="Teng-da Liang &amp; Min Wang" country="China" elevation="1850" location="Mt. Wuyi" pageId="1" pageNumber="594" specimenCount="1" stateProvince="Fujian" typeStatus="holotype">
<emphasis id="B956AD55FD368429FE8C38E1A2CCFB43" bold="true" box="[330,448,1184,1210]" pageId="1" pageNumber="594">
<typeStatus id="5499CFE5FD368429FE8C38E1A2B7FB43" box="[330,443,1184,1210]" pageId="1" pageNumber="594">Holotype</typeStatus>
:
</emphasis>
male, altitude
<quantity id="4CDADCA2FD368429FDA238E1A00CFB42" box="[612,768,1184,1211]" metricMagnitude="3" metricUnit="m" metricValue="1.85" metricValueMax="1.9" metricValueMin="1.8" pageId="1" pageNumber="594" unit="m" value="1850.0" valueMax="1900.0" valueMin="1800.0">
<elevation id="000F9674FD368429FDA238E1A00CFB42" box="[612,768,1184,1211]" metricMagnitude="3" metricUnit="m" metricValue="1.85" metricValueMax="1.9" metricValueMin="1.8" pageId="1" pageNumber="594" unit="m" value="1850.0" valueMax="1900.0" valueMin="1800.0">1800–1900 m</elevation>
</quantity>
,
<date id="FF9C5787FD368429FCCD38E1A0ADFB43" box="[779,929,1184,1211]" pageId="1" pageNumber="594" value="2022-07-11">
<collectingDate id="EFD8AE6FFD368429FCCD38E1A0ADFB43" box="[779,929,1184,1211]" pageId="1" pageNumber="594" value="2022-07-11">11. VII. 2022</collectingDate>
</date>
, Huanggangshan,
<location id="8EFD279CFD368429FBAB38E0A7D6FB43" LSID="urn:lsid:plazi:treatment:038BC051FD36842BFF513878A7B2FCA4:8EFD279CFD368429FBAB38E0A7D6FB43" box="[1133,1242,1184,1211]" country="China" name="Mt. Wuyi" pageId="1" pageNumber="594" stateProvince="Fujian">Mt. Wuyi</location>
,
<collectingRegion id="49E6BFA5FD368429FB2238E1A694FB43" box="[1252,1432,1184,1210]" country="China" name="Fujian" pageId="1" pageNumber="594">Fujian Province</collectingRegion>
,
<collectingCountry id="F33531D7FD368429FF513885A3D1FB27" box="[151,221,1220,1246]" name="China" pageId="1" pageNumber="594">China</collectingCountry>
, leg.
<collectorName id="26D71491FD368429FED13885A2B6FB27" box="[279,442,1220,1246]" pageId="1" pageNumber="594">Teng-da Liang</collectorName>
&amp;
<collectorName id="26D71491FD368429FE1B3885A15EFB26" box="[477,594,1220,1247]" pageId="1" pageNumber="594">Min Wang</collectorName>
(
<collectionCode id="ED33E982FD368429FDA63885A1BDFB27" box="[608,689,1220,1246]" pageId="1" pageNumber="594">SCAU</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B4A7B1AFD368429FD053885A7F3FB27" box="[707,1279,1220,1247]" collectingDate="2022-07-11" collectionCode="SCAU" collectorName="Teng-da Liang &amp; Min Wang" country="China" elevation="1850" location="Mt. Wuyi" pageId="1" pageNumber="594" specimenCount="1" specimenCount-male="1" stateProvince="Fujian" typeStatus="paratype">
<emphasis id="B956AD55FD368429FD053885A048FB27" bold="true" box="[707,836,1220,1246]" pageId="1" pageNumber="594">
<typeStatus id="5499CFE5FD368429FD053885A033FB27" box="[707,831,1220,1246]" pageId="1" pageNumber="594">Paratypes</typeStatus>
:
</emphasis>
<specimenCount id="9D24BACEFD368429FC8D3885A094FB27" box="[843,920,1220,1246]" pageId="1" pageNumber="594" type="male">1 male</specimenCount>
, same data as holotype (
<collectionCode id="ED33E982FD368429FB613885A7F4FB27" box="[1191,1272,1220,1246]" pageId="1" pageNumber="594">SCAU</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B4A7B1AFD368429FACC3885A76AFAFB" collectingDate="2022-07-08" collectionCode="SCAU" collectorName="Min Wang &amp; Jun-ru He" country="China" elevation="1650" location="Mt. Wuyi" pageId="1" pageNumber="594" specimenCount="1" specimenCount-male="1" stateProvince="Fujian" typeStatus="paratype">
<specimenCount id="9D24BACEFD368429FACC3885A654FB27" box="[1290,1368,1220,1246]" pageId="1" pageNumber="594" type="male">1 male</specimenCount>
, same locality, but altitude
<quantity id="4CDADCA2FD368429FEB938A9A111FAFA" box="[383,541,1256,1283]" metricMagnitude="3" metricUnit="m" metricValue="1.65" metricValueMax="1.7" metricValueMin="1.6" pageId="1" pageNumber="594" unit="m" value="1650.0" valueMax="1700.0" valueMin="1600.0">
<elevation id="000F9674FD368429FEB938A9A111FAFA" box="[383,541,1256,1283]" metricMagnitude="3" metricUnit="m" metricValue="1.65" metricValueMax="1.7" metricValueMin="1.6" pageId="1" pageNumber="594" unit="m" value="1650.0" valueMax="1700.0" valueMin="1600.0">1600–1700 m</elevation>
</quantity>
,
<date id="FF9C5787FD368429FDEF38A9A1BAFAFB" box="[553,694,1256,1283]" pageId="1" pageNumber="594" value="2022-07-08">
<collectingDate id="EFD8AE6FFD368429FDEF38A9A1BAFAFB" box="[553,694,1256,1283]" pageId="1" pageNumber="594" value="2022-07-08">8. VII. 2022</collectingDate>
</date>
, leg.
<collectorName id="26D71491FD368429FD3438A9A065FAFA" box="[754,873,1256,1283]" pageId="1" pageNumber="594">Min Wang</collectorName>
&amp;
<collectorName id="26D71491FD368429FC4B38A8A0F2FAFA" box="[909,1022,1257,1283]" pageId="1" pageNumber="594">Jun-ru He</collectorName>
(
<collectionCode id="ED33E982FD368429FBCB38A9A752FAFB" box="[1037,1118,1256,1282]" pageId="1" pageNumber="594">SCAU</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B4A7B1AFD368429FBB438A9A196FADF" collectingDate="2022-07-08" collectionCode="SCAU" collectorName="Min Wang &amp; Li-ping Zhou" country="China" elevation="1650" location="Mt. Wuyi" pageId="1" pageNumber="594" specimenCount="2" specimenCount-male="2" stateProvince="Fujian" typeStatus="paratype">
<specimenCount id="9D24BACEFD368429FBB438A9A7C7FAFB" box="[1138,1227,1256,1282]" pageId="1" pageNumber="594" type="male">2 males</specimenCount>
, same locality and date, leg.
<collectorName id="26D71491FD368429FEC2394DA275FADE" box="[260,377,1292,1319]" pageId="1" pageNumber="594">Min Wang</collectorName>
&amp;
<collectorName id="26D71491FD368429FE5A394DA13EFADF" box="[412,562,1292,1318]" pageId="1" pageNumber="594">Li-ping Zhou</collectorName>
(
<collectionCode id="ED33E982FD368429FD87394DA19EFADF" box="[577,658,1292,1318]" pageId="1" pageNumber="594">SCAU</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B4A7B1AFD368429FD63394DA1A3FAB3" collectingDate="2022-07-13" collectionCode="SCAU" collectorName="Min Wang &amp; Yu-long Zhang" country="China" elevation="1900" location="Mt. Wuyi" pageId="1" pageNumber="594" specimenCount="3" specimenCount-female="2" specimenCount-male="1" stateProvince="Fujian" typeStatus="paratype">
<specimenCount id="9D24BACEFD368429FD63394DA1FFFADF" box="[677,755,1292,1318]" pageId="1" pageNumber="594" type="male">1 male</specimenCount>
,
<specimenCount id="9D24BACEFD368429FD38394DA061FADF" box="[766,877,1292,1318]" pageId="1" pageNumber="594" type="female">2 females</specimenCount>
, same locality, but altitude
<quantity id="4CDADCA2FD368429FB5B394DA636FADE" box="[1181,1338,1292,1319]" metricMagnitude="3" metricUnit="m" metricValue="1.9" metricValueMax="2.0" metricValueMin="1.8" pageId="1" pageNumber="594" unit="m" value="1900.0" valueMax="2000.0" valueMin="1800.0">
<elevation id="000F9674FD368429FB5B394DA636FADE" box="[1181,1338,1292,1319]" metricMagnitude="3" metricUnit="m" metricValue="1.9" metricValueMax="2.0" metricValueMin="1.8" pageId="1" pageNumber="594" unit="m" value="1900.0" valueMax="2000.0" valueMin="1800.0">1800–2000 m</elevation>
</quantity>
,
<date id="FF9C5787FD368429FA83394DA3DEFAB3" pageId="1" pageNumber="594" value="2022-07-13">
<collectingDate id="EFD8AE6FFD368429FA83394DA3DEFAB3" pageId="1" pageNumber="594" value="2022-07-13">13. VII. 2022</collectingDate>
</date>
, leg.
<collectorName id="26D71491FD368429FECD3971A28CFAB2" box="[267,384,1328,1355]" pageId="1" pageNumber="594">Min Wang</collectorName>
&amp;
<collectorName id="26D71491FD368429FE663971A145FAB3" box="[416,585,1328,1354]" pageId="1" pageNumber="594">Yu-long Zhang</collectorName>
(
<collectionCode id="ED33E982FD368429FD903971A1ABFAB3" box="[598,679,1328,1354]" pageId="1" pageNumber="594">SCAU</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B4A7B1AFD368429FD7F3971A18BFA97" collectingDate="2022-07-29" collectionCode="SCAU" collectorName="Min Wang &amp; Liang Guo" country="China" elevation="1800" location="Mt. Wuyi" pageId="1" pageNumber="594" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Fujian" typeStatus="paratype">
<specimenCount id="9D24BACEFD368429FD7F3971A00AFAB3" box="[697,774,1328,1354]" pageId="1" pageNumber="594" type="male">1 male</specimenCount>
,
<specimenCount id="9D24BACEFD368429FCD63971A07FFAB3" box="[784,883,1328,1354]" pageId="1" pageNumber="594" type="female">1 female</specimenCount>
, same locality, but altitude
<quantity id="4CDADCA2FD368429FB593971A637FAB2" box="[1183,1339,1328,1355]" metricMagnitude="3" metricUnit="m" metricValue="1.8" metricValueMax="1.9" metricValueMin="1.7" pageId="1" pageNumber="594" unit="m" value="1800.0" valueMax="1900.0" valueMin="1700.0">
<elevation id="000F9674FD368429FB593971A637FAB2" box="[1183,1339,1328,1355]" metricMagnitude="3" metricUnit="m" metricValue="1.8" metricValueMax="1.9" metricValueMin="1.7" pageId="1" pageNumber="594" unit="m" value="1800.0" valueMax="1900.0" valueMin="1700.0">1700–1900 m</elevation>
</quantity>
,
<date id="FF9C5787FD368429FA803971A3DEFA97" pageId="1" pageNumber="594" value="2022-07-29">
<collectingDate id="EFD8AE6FFD368429FA803971A3DEFA97" pageId="1" pageNumber="594" value="2022-07-29">29. VII. 2022</collectingDate>
</date>
, leg.
<collectorName id="26D71491FD368429FECB3915A288FA96" box="[269,388,1364,1391]" pageId="1" pageNumber="594">Min Wang</collectorName>
&amp;
<collectorName id="26D71491FD368429FE613915A113FA97" box="[423,543,1364,1390]" pageId="1" pageNumber="594">Liang Guo</collectorName>
(
<collectionCode id="ED33E982FD368429FDE83915A173FA97" box="[558,639,1364,1390]" pageId="1" pageNumber="594">SCAU</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C33822CCFD368429FF013939A65BF92F" pageId="1" pageNumber="594" type="diagnosis">
<paragraph id="8B9D7147FD368429FF013939A65BF92F" blockId="1.[151,1437,1184,2039]" pageId="1" pageNumber="594">
<emphasis id="B956AD55FD368429FF013939A24EFA6B" bold="true" box="[199,322,1400,1426]" pageId="1" pageNumber="594">Diagnosis.</emphasis>
<taxonomicName id="4C220AC4FD368429FE813939A179FA6B" authority="Guo &amp; Huang &amp; Wang &amp; Ma, 2023" authorityName="Guo &amp; Huang &amp; Wang &amp; Ma" authorityYear="2023" box="[327,629,1400,1426]" class="Insecta" family="Nymphalidae" genus="Chonala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="594" phylum="Arthropoda" rank="subSpecies" species="laurae" status="ssp. n." subSpecies="wuyishana">
<emphasis id="B956AD55FD368429FE813939A179FA6B" box="[327,629,1400,1426]" italics="true" pageId="1" pageNumber="594">Chonala laurae wuyishana</emphasis>
</taxonomicName>
<emphasis id="B956AD55FD368429FDBD3939A1C8FA6B" bold="true" box="[635,708,1400,1426]" pageId="1" pageNumber="594">
<taxonomicNameLabel id="A265102EFD368429FDBD3939A1C8FA6B" box="[635,708,1400,1426]" pageId="1" pageNumber="594" rank="subSpecies">ssp. n.</taxonomicNameLabel>
</emphasis>
can be distinguished from the nominate subspecies (
<figureCitation id="13196DC2FD368429FACD3939A662FA6A" box="[1291,1390,1400,1427]" captionStart="FIGURES 1–8" captionStartId="2.[151,264,1943,1967]" captionTargetBox="[203,1384,181,1919]" captionTargetId="figure-21@2.[203,1384,181,1919]" captionTargetPageId="2" captionText="FIGURES 1–8. Male of Chonala sspp. 1–2, C. laurae wuyishana ssp. nov., holotype, Huanggangshan, Mt. Wuyi, Fujian (SCAU); 3–4, ditto, paratype, ditto (SCAU); 5–6, C. laurae laurae, holotype, Qinling, Shaanxi (ZFMK); 7–8, ditto, Shennongjia, Hubei, SATY0700 (CSYL). Scale=10 mm." figureDoi="http://doi.org/10.5281/zenodo.8243789" httpUri="https://zenodo.org/record/8243789/files/figure.png" pageId="1" pageNumber="594">Figs 5–8</figureCitation>
,
<figureCitation id="13196DC2FD368429FABF3939A3B9FA4F" captionStart="FIGURES 9–16" captionStartId="4.[151,264,1943,1967]" captionTargetBox="[203,1384,181,1919]" captionTargetId="figure-21@4.[203,1384,181,1919]" captionTargetPageId="4" captionText="FIGURES 9–16. Adults of Chonala spp. 9–10, C. houae, holotype, Mt. Jinfo, Chongqing, SATY1156 (CQMNH); 11–12, ditto, paratype, ditto, SATY0466 (CQMNH); 13–14, C. laurae wuyishana ssp. nov., paratype, Mt. Wuyi, Fujian (SCAU); 15–16, C. laurae laurae, Shennongjia, Hubei (CSYL); 9–12 male, 13–16 female. Scale=10 mm." figureDoi="http://doi.org/10.5281/zenodo.8243793" httpUri="https://zenodo.org/record/8243793/files/figure.png" pageId="1" pageNumber="594">15, 16</figureCitation>
,
<figureCitation id="13196DC2FD368429FF0739DDA3D2FA4F" box="[193,222,1436,1462]" captionStart="FIGURES 17–20" captionStartId="5.[151,264,1943,1967]" captionTargetBox="[197,1391,194,1907]" captionTargetId="figure-20@5.[184,1403,181,1919]" captionTargetPageId="5" captionText="FIGURES 17–20. Male genitalia of Chonala sspp. Depository of the genitalia: 17 in SCAU, 18 in CSYL, 19 &amp; 20 in CQMNH. a=genitalia capsule with left valva and aedeagus removed; b=aedeagus in dorsal view; c=aedeagus in lateral view; d=left valva in dorsal view. Scale=1 mm." figureDoi="http://doi.org/10.5281/zenodo.8243795" httpUri="https://zenodo.org/record/8243795/files/figure.png" pageId="1" pageNumber="594">18</figureCitation>
) from the central part of
<collectingCountry id="F33531D7FD368429FE3139DDA136FA4F" box="[503,570,1436,1462]" name="China" pageId="1" pageNumber="594">China</collectingCountry>
by a larger and more prominent costal ocherous yellow spot on the upperside of the forewing in both sexes, a more sharply defined and generally larger ocherous yellow patch in space 2 on the upperside of the forewing in males (such a patch is diffused and blur in the nominate subspecies), and the ground color is tinted with reddish brown on both sides of wings in females (ground color is much greyish in the nominate subspecies). The new subspecies is also reminiscent of
<taxonomicName id="4C220AC4FD368429FCCA3A6DA07EF9BF" authorityName="Lang, Li &amp; Deng" authorityYear="2017" box="[780,882,1580,1606]" class="Insecta" family="Nymphalidae" genus="Chonala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="594" phylum="Arthropoda" rank="species" species="houae">
<emphasis id="B956AD55FD368429FCCA3A6DA07EF9BF" box="[780,882,1580,1606]" italics="true" pageId="1" pageNumber="594">C. houae</emphasis>
</taxonomicName>
(
<figureCitation id="13196DC2FD368429FC453A6DA0F1F9BF" box="[899,1021,1580,1606]" captionStart="FIGURES 9–16" captionStartId="4.[151,264,1943,1967]" captionTargetBox="[203,1384,181,1919]" captionTargetId="figure-21@4.[203,1384,181,1919]" captionTargetPageId="4" captionText="FIGURES 9–16. Adults of Chonala spp. 9–10, C. houae, holotype, Mt. Jinfo, Chongqing, SATY1156 (CQMNH); 11–12, ditto, paratype, ditto, SATY0466 (CQMNH); 13–14, C. laurae wuyishana ssp. nov., paratype, Mt. Wuyi, Fujian (SCAU); 15–16, C. laurae laurae, Shennongjia, Hubei (CSYL); 9–12 male, 13–16 female. Scale=10 mm." figureDoi="http://doi.org/10.5281/zenodo.8243793" httpUri="https://zenodo.org/record/8243793/files/figure.png" pageId="1" pageNumber="594">Figs. 9–12</figureCitation>
,
<figureCitation id="13196DC2FD368429FBCC3A6DA75FF9BF" box="[1034,1107,1580,1606]" captionStart="FIGURES 17–20" captionStartId="5.[151,264,1943,1967]" captionTargetBox="[197,1391,194,1907]" captionTargetId="figure-20@5.[184,1403,181,1919]" captionTargetPageId="5" captionText="FIGURES 17–20. Male genitalia of Chonala sspp. Depository of the genitalia: 17 in SCAU, 18 in CSYL, 19 &amp; 20 in CQMNH. a=genitalia capsule with left valva and aedeagus removed; b=aedeagus in dorsal view; c=aedeagus in lateral view; d=left valva in dorsal view. Scale=1 mm." figureDoi="http://doi.org/10.5281/zenodo.8243795" httpUri="https://zenodo.org/record/8243795/files/figure.png" pageId="1" pageNumber="594">19, 20</figureCitation>
) from Mt. Jinfo,
<collectingRegion id="49E6BFA5FD368429FADF3A6DA694F9BF" box="[1305,1432,1580,1606]" country="China" name="Chongqing" pageId="1" pageNumber="594">Chongqing</collectingRegion>
, but it can be easily distinguished by the following features in males: the pattern is less tint with ocherous on the forewing upperside, the coloration of the rectangular patch in space 2 on the forewing underside is uniform throughout (coloration of such a patch in
<taxonomicName id="4C220AC4FD368429FDAF3AD9A1C2F94B" authorityName="Lang, Li &amp; Deng" authorityYear="2017" box="[617,718,1688,1714]" class="Insecta" family="Nymphalidae" genus="Chonala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="594" phylum="Arthropoda" rank="species" species="houae">
<emphasis id="B956AD55FD368429FDAF3AD9A1C2F94B" box="[617,718,1688,1714]" italics="true" pageId="1" pageNumber="594">C. houae</emphasis>
</taxonomicName>
is gradually changing from ochreous yellow in the outer half to ochreous in the inner half), and a much longer gnathos in genitalia. The female of
<taxonomicName id="4C220AC4FD368429FBF63AFDA799F92F" authorityName="Lang, Li &amp; Deng" authorityYear="2017" box="[1072,1173,1724,1750]" class="Insecta" family="Nymphalidae" genus="Chonala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="594" phylum="Arthropoda" rank="species" species="houae">
<emphasis id="B956AD55FD368429FBF63AFDA799F92F" box="[1072,1173,1724,1750]" italics="true" pageId="1" pageNumber="594">C. houae</emphasis>
</taxonomicName>
is still unknown.
</paragraph>
</subSubSection>
<subSubSection id="C33822CCFD36842BFF013AA1A248FD34" lastPageId="3" lastPageNumber="596" pageId="1" pageNumber="594" type="description">
<paragraph id="8B9D7147FD36842BFF013AA1A040FE0C" blockId="1.[151,1437,1184,2039]" lastBlockId="3.[151,1437,151,861]" lastPageId="3" lastPageNumber="596" pageId="1" pageNumber="594">
<emphasis id="B956AD55FD368429FF013AA1A2A8F903" bold="true" box="[199,420,1760,1786]" pageId="1" pageNumber="594">Description. Male.</emphasis>
Length of forewing 27.5–
<quantity id="4CDADCA2FD368429FD0C3AA1A014F902" box="[714,792,1760,1787]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" pageId="1" pageNumber="594" unit="mm" value="30.0">30 mm</quantity>
(n=7, 27.
<quantity id="4CDADCA2FD368429FC423AA1A0C8F902" box="[900,964,1760,1787]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="1" pageNumber="594" unit="mm" value="5.0">5 mm</quantity>
in
<typeStatus id="5499CFE5FD368429FC203AA1A742F903" box="[998,1102,1760,1786]" pageId="1" pageNumber="594">holotype</typeStatus>
). Antenna dorsally brown and ventrally dusted with white at base of each segment, with its tip dull yellow. Head, thorax, and abdomen dorsally brown. Upperside of forewing with ground color brown from base to discal area and pale black in submarginal and apical area, markings yellow with slight ochreous tint. Two dots present in subapical area with the lower one twice the size of the upper one. A rectangular patch present at upper angle of the cell at costa. An oblique short band dumbbell-shaped, extending from vein 6 to vein 4. A trapezoid patch present in the outer half of space 2. Cilia chequered with pale black and pale yellowish white. Upperside of hindwing with ground color brown, with ocelli sometimes present in spaces 2 and 3. Cilia as in forewing. Underside of forewing with ground color brown in basal half and black in distal half. The cell crossed by two reddish brown streaks with the inner one broken and incomplete. Costal patch and short band as in upperside. A black ocellus present in distal half of space 5, pupillated and ringed with yellow. Two white dots present beyond ocellus in spaces 6 and 7, with the lower one twice the size of the upper one. Two wavy submarginal white lines extending from costa to space 2. A rectangular patch present in space 2, uniform in color. Basal section of spaces 2 and 3 and area around cubital vein suffused with pale ochreous yellow. Hindwing underside, ground color mostly grey and become white around apical area. Basal and discal sections of wing scattered with several brownish red wavy lines. A series of ocelli extending from apex to tornus, with one in space 6 largest and one in space 1 bipupillated. Submarginal area with two white wavy lines.
<emphasis id="B956AD55FD34842BFAF83DD2A690FE54" bold="true" box="[1342,1436,403,429]" pageId="3" pageNumber="596">Female.</emphasis>
Length of forewing 26.5–
<quantity id="4CDADCA2FD34842BFE733DF7A10EFE28" box="[437,514,438,465]" metricMagnitude="-2" metricUnit="m" metricValue="2.8" pageId="3" pageNumber="596" unit="mm" value="28.0">28 mm</quantity>
(n=3). Similar to male, but forewing more rounded, forewing upperside with patch in space 2 much longer and larger, underside same as in male.
</paragraph>
<caption id="DF5D21CFFD35842AFF513BD6A140F80E" ID-DOI="http://doi.org/10.5281/zenodo.8243789" ID-Zenodo-Dep="8243789" httpUri="https://zenodo.org/record/8243789/files/figure.png" pageId="2" pageNumber="595" startId="2.[151,264,1943,1967]" targetBox="[203,1384,181,1919]" targetPageId="2" targetType="figure">
<paragraph id="8B9D7147FD35842AFF513BD6A140F80E" blockId="2.[151,1437,1943,2039]" pageId="2" pageNumber="595">
<emphasis id="B956AD55FD35842AFF513BD6A232F856" bold="true" box="[151,318,1943,1967]" pageId="2" pageNumber="595">FIGURES 1–8.</emphasis>
Male of
<taxonomicName id="4C220AC4FD35842AFE633BD6A2F0F856" authorityName="Moore" authorityYear="1893" box="[421,508,1943,1967]" class="Insecta" family="Nymphalidae" genus="Chonala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="595" phylum="Arthropoda" rank="genus">
<emphasis id="B956AD55FD35842AFE633BD6A2F0F856" box="[421,508,1943,1967]" italics="true" pageId="2" pageNumber="595">Chonala</emphasis>
</taxonomicName>
sspp. 1–2,
<taxonomicName id="4C220AC4FD35842AFDBF3BD9A05EF856" authorityName="Guo &amp; Huang &amp; Wang &amp; Ma" authorityYear="2023" box="[633,850,1943,1967]" class="Insecta" family="Nymphalidae" genus="Chonala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="595" phylum="Arthropoda" rank="subSpecies" species="laurae" subSpecies="wuyishana">
<emphasis id="B956AD55FD35842AFDBF3BD9A05EF856" box="[633,850,1943,1967]" italics="true" pageId="2" pageNumber="595">C. laurae wuyishana</emphasis>
</taxonomicName>
<emphasis id="B956AD55FD35842AFC9A3BD6A0B6F856" bold="true" box="[860,954,1943,1967]" pageId="2" pageNumber="595">ssp. nov.</emphasis>
, holotype, Huanggangshan, Mt. Wuyi, Fujian (SCAU); 3–4,
<emphasis id="B956AD55FD35842AFEEE3BFAA25AF82A" box="[296,342,1979,2003]" italics="true" pageId="2" pageNumber="595">ditto</emphasis>
, paratype,
<emphasis id="B956AD55FD35842AFE023BFAA2FEF82A" box="[452,498,1979,2003]" italics="true" pageId="2" pageNumber="595">ditto</emphasis>
(SCAU); 5–6,
<taxonomicName id="4C220AC4FD35842AFD4E3BFDA021F82A" authorityName="Bozano" authorityYear="1999" box="[648,813,1979,2003]" class="Insecta" family="Nymphalidae" genus="Chonala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="595" phylum="Arthropoda" rank="subSpecies" species="laurae" subSpecies="laurae">
<emphasis id="B956AD55FD35842AFD4E3BFDA021F82A" box="[648,813,1979,2003]" italics="true" pageId="2" pageNumber="595">C. laurae laurae</emphasis>
</taxonomicName>
, holotype, Qinling, Shaanxi (ZFMK); 7–8,
<emphasis id="B956AD55FD35842AFB183BFAA600F82A" box="[1246,1292,1979,2003]" italics="true" pageId="2" pageNumber="595">ditto</emphasis>
, Shennongjia, Hubei, SATY0700 (CSYL). Scale=10 mm.
</paragraph>
</caption>
<paragraph id="8B9D7147FD34842BFF013DBEA248FD34" blockId="3.[151,1437,151,861]" pageId="3" pageNumber="596">
<emphasis id="B956AD55FD34842BFF013DBEA274FDE1" bold="true" box="[199,376,510,537]" pageId="3" pageNumber="596">Male genitalia.</emphasis>
Tegumen simple. Uncus moderately long, almost the same length as tegumen and slightly bent downwards medially. Gnathos almost one fourth of the length of uncus, gradually narrowed towards distal end and slightly curved upwards. Saccus short, swollen at distal end. Valva twice the length of uncus, broad at base, gradually narrowed towards tip. Tip of the valva, in dorsal view, enlarged near distal end and forming a triangular process extending interiorly with its distal end dentate. Aedeagus simple, moderately long, with a small bulge at basal one third.
</paragraph>
</subSubSection>
<subSubSection id="C33822CCFD34842BFF013E97A246FCEC" pageId="3" pageNumber="596" type="distribution">
<paragraph id="8B9D7147FD34842BFF013E97A246FCEC" blockId="3.[151,1437,151,861]" pageId="3" pageNumber="596">
<emphasis id="B956AD55FD34842BFF013E97A26DFD09" bold="true" box="[199,353,726,752]" pageId="3" pageNumber="596">Distribution.</emphasis>
Northwestern
<collectingRegion id="49E6BFA5FD34842BFDCC3E96A1B2FD08" box="[522,702,727,753]" country="China" name="Fujian" pageId="3" pageNumber="596">Fujian Province</collectingRegion>
(Huanggangshan, the summit of Mt. Wuyi Range), southeastern
<collectingCountry id="F33531D7FD34842BFF513EBAA3D6FCEC" box="[151,218,763,789]" name="China" pageId="3" pageNumber="596">China</collectingCountry>
(
<figureCitation id="13196DC2FD34842BFF2F3EBAA231FCEC" box="[233,317,763,789]" captionStart="FIGURES 21" captionStartId="6.[151,264,1435,1459]" captionTargetBox="[151,1436,525,1412]" captionTargetId="figure-195@6.[151,1436,525,1412]" captionTargetPageId="6" captionText="FIGURES 21. Distribution map of Chonala spp. in the current study." figureDoi="http://doi.org/10.5281/zenodo.8243797" httpUri="https://zenodo.org/record/8243797/files/figure.png" pageId="3" pageNumber="596">Fig. 21</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C33822CCFD34842BFF013F5EA7FBFCC0" box="[199,1271,799,825]" pageId="3" pageNumber="596" type="etymology">
<paragraph id="8B9D7147FD34842BFF013F5EA7FBFCC0" blockId="3.[151,1437,151,861]" box="[199,1271,799,825]" pageId="3" pageNumber="596">
<emphasis id="B956AD55FD34842BFF013F5EA242FCC0" bold="true" box="[199,334,799,825]" pageId="3" pageNumber="596">Etymology.</emphasis>
The subspecific epithet
<emphasis id="B956AD55FD34842BFD993F5EA1DBFCC0" box="[607,727,799,825]" italics="true" pageId="3" pageNumber="596">wuyishana</emphasis>
is derived from the
<typeStatus id="5499CFE5FD34842BFC7F3F5EA0E5FCC0" box="[953,1001,799,825]" pageId="3" pageNumber="596">type</typeStatus>
locality, Mt. Wuyishan.
</paragraph>
</subSubSection>
<subSubSection id="C33822CCFD34842BFF013F02A7B2FCA4" box="[199,1214,835,861]" pageId="3" pageNumber="596" type="discussion">
<paragraph id="8B9D7147FD34842BFF013F02A7B2FCA4" blockId="3.[151,1437,151,861]" box="[199,1214,835,861]" pageId="3" pageNumber="596">
<emphasis id="B956AD55FD34842BFF013F02A237FCA4" bold="true" box="[199,315,835,861]" pageId="3" pageNumber="596">Remarks.</emphasis>
This species is found flying with
<taxonomicName id="4C220AC4FD34842BFD713F02A729FCA4" authority="(Leech, 1890)" baseAuthorityName="Leech" baseAuthorityYear="1890" box="[695,1061,835,861]" class="Insecta" family="Nymphalidae" genus="Chonala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="596" phylum="Arthropoda" rank="species" species="praeusta">
<emphasis id="B956AD55FD34842BFD713F02A08DFCA5" box="[695,897,835,861]" italics="true" pageId="3" pageNumber="596">Chonala praeusta</emphasis>
(
<bibRefCitation id="EFB30CB6FD34842BFC563F02A711FCA4" author="Leech, J. H." box="[912,1053,835,861]" pageId="3" pageNumber="596" pagination="26 - 50" refId="ref5032" refString="Leech, J. H. (1890) New species of Lepidoptera from China. Entomologist, 23, 26 - 50 + 81 - 83 + 109 - 114 + 187 - 192, pl. 1." type="journal article" year="1890">Leech, 1890</bibRefCitation>
)
</taxonomicName>
in its habitat.
</paragraph>
</subSubSection>
</treatment>
</document>
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