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<document id="D080CF57401D9C687A53EE0B3726C08C" ID-DOI="10.5281/zenodo.273688" ID-GBIF-Dataset="e73070a4-e67b-4856-a531-510f2722e7fc" ID-ISSN="1175-5326" ID-Zenodo-Dep="273688" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459811775148" checkinUser="plazi" docAuthor="Mclaughlin, Patsy A., Lemaitre, Rafael &amp; Ortiz, Manuel" docDate="2007" docId="038B576BFFEC6D5AA692FB9FD8B68D61" docLanguage="en" docName="zt01406p046.pdf" docOrigin="Zootaxa 1406" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Pylocheles (Bathycheles) cubensis Ortiz &amp; Gomez 1986" docType="treatment" docVersion="8" lastPageNumber="45" masterDocId="FFB22F13FFED6D5EA605FFF5D928885B" masterDocTitle="Redescription of Pylocheles (Bathycheles) cubensis Ortiz &amp; Gómez, 1986, the senior synonym of Pylocheles (B.) chacei Forest, 1987 (Decapoda, Anomura, Paguroidea, Pylochelidae)" masterLastPageNumber="46" masterPageNumber="41" pageNumber="42" updateTime="1698221084802" updateUser="plazi">
<mods:mods id="8EAA43DB6525CBA1951B156EA14FB944" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="08F1B251013458CD41EEB9948279CDA1">Redescription of Pylocheles (Bathycheles) cubensis Ortiz &amp; Gómez, 1986, the senior synonym of Pylocheles (B.) chacei Forest, 1987 (Decapoda, Anomura, Paguroidea, Pylochelidae)</mods:title>
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<mods:namePart id="514652EAC38AE8CEEC88753ABECFF0B4">Mclaughlin, Patsy A.</mods:namePart>
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<mods:namePart id="CAE1348B348DD2ED12B2437D65F65191">Lemaitre, Rafael</mods:namePart>
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<mods:name id="D5F1B264A70D67E27281DDBB5EDE9AF2" type="personal">
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<mods:namePart id="DEE8C9FE50DA249B183B8FB538A52B6D">Ortiz, Manuel</mods:namePart>
</mods:name>
<mods:typeOfResource id="FB8B58B2433627B7223B3CF77CDF032E">text</mods:typeOfResource>
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<mods:title id="1310DE19CDB1E65810BBD5DA78DB33DC">Zootaxa</mods:title>
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<mods:part id="6C621362A7A56EE375A65B45BF6A8303">
<mods:date id="AA5FB30652566C34437D01AE27C6B87A">2007</mods:date>
<mods:detail id="6D29DB10EBDA32B8E573F80EE97C9221" type="volume">
<mods:number id="D9B38ACA77A32614374EB8B14921A953">1406</mods:number>
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<mods:identifier id="7AB782A8FCA11FCFD212ADCC3D90378E" type="Zenodo-Dep">273688</mods:identifier>
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<treatment id="038B576BFFEC6D5AA692FB9FD8B68D61" ID-DOI="http://doi.org/10.5281/zenodo.6242781" ID-GBIF-Taxon="119354016" ID-Zenodo-Dep="6242781" LSID="urn:lsid:plazi:treatment:038B576BFFEC6D5AA692FB9FD8B68D61" httpUri="http://treatment.plazi.org/id/038B576BFFEC6D5AA692FB9FD8B68D61" lastPageId="4" lastPageNumber="45" pageId="1" pageNumber="42">
<subSubSection id="C338B5F6FFEC6D5FA692FB9FDA8F8CDF" box="[151,935,1130,1156]" pageId="1" pageNumber="42" type="nomenclature">
<paragraph id="8B9DE67DFFEC6D5FA692FB9FDA8F8CDF" blockId="1.[151,935,1130,1156]" box="[151,935,1130,1156]" pageId="1" pageNumber="42">
<heading id="D0D55111FFEC6D5FA692FB9FDA8F8CDF" bold="true" box="[151,935,1130,1156]" fontSize="11" level="1" pageId="1" pageNumber="42" reason="1">
<emphasis id="B9563A6FFFEC6D5FA692FB9FDA8F8CDF" bold="true" box="[151,935,1130,1156]" pageId="1" pageNumber="42">
<taxonomicName id="4C229DFEFFEC6D5FA692FB9FDA6F8CDF" ID-CoL="3d2d8404-53f6-4fb6-a66b-f37c97198ad2" authority="Ortiz &amp; Gomez, 1986" authorityName="Ortiz &amp; Gomez" authorityYear="1986" box="[151,839,1130,1156]" class="Malacostraca" family="Pylochelidae" genus="Pylocheles" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="42" phylum="Arthropoda" rank="species" species="cubensis" subGenus="Bathycheles">
<emphasis id="B9563A6FFFEC6D5FA692FB9FDB1C8CDF" bold="true" box="[151,564,1130,1156]" italics="true" pageId="1" pageNumber="42">Pylocheles (Bathycheles) cubensis</emphasis>
<bibRefCitation id="EFB39B8CFFEC6D5FA439FB9FDA6F8CDF" author="Ortiz" box="[572,839,1130,1156]" pageId="1" pageNumber="46" refString="Ortiz, M. &amp; Gomez, O. (1986). Una nueva especie del genero Pylocheles (Anomura Galatheoidea) (sic), de las aguas profundas del sur de Cuba. Revista de Investigaciones Marinas, 7, 31 - 38." type="journal article" year="1986">Ortiz &amp; Gómez, 1986</bibRefCitation>
</taxonomicName>
(
<figureCitation id="1319FAF8FFEC6D5FA553FB9FDAB68CDF" box="[854,926,1130,1156]" captionStart="FIGURE 1" captionStartId="2.[151,255,1527,1551]" captionTargetBox="[264,1323,354,1506]" captionTargetId="figure@2.[264,1323,354,1506]" captionTargetPageId="2" captionText="FIGURE 1. Pylocheles (Bathycheles) cubensis Ortiz &amp; Gómez, 1986, holotype male (19 mm) (ANC / IDO. 07.1.6.5.047). A, Dorsal view of whole animal (tied with strings to glass plate); B, chelae and carpi (dorsal view); C, anterior portion of shield, cephalic appendages, carpi of chelipeds, and portion of left second and third pereopods (dorsolateral view); D, enlarged dorsal view of carpi of chelipeds; E, anterior portion of shield, cephalic appendages, left cheliped, and carpi to dactyls of left second and third pereopods (lateral view); F, pleonal tergites 4 – 6, uropods and telson (dorsal and slightly posterior view); G, right first pleopod (internal view); H, right second pleopod (internal view). (Photographs by M. Ortiz)." httpUri="https://zenodo.org/record/273689/files/figure.png" pageId="1" pageNumber="42">Fig. 1</figureCitation>
)
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C338B5F6FFEC6D5FA692FB40DA0D8CB7" pageId="1" pageNumber="42" type="reference_group">
<paragraph id="8B9DE67DFFEC6D5FA692FB40DBDE8C97" blockId="1.[151,805,1205,1260]" box="[151,758,1205,1228]" pageId="1" pageNumber="42">
<treatmentCitationGroup id="AB32C153FFEC6D5FA692FB40DBDE8C97" box="[151,758,1205,1228]" pageId="1" pageNumber="42">
<taxonomicName id="4C229DFEFFEC6D5FA692FB40DBAF8C97" ID-CoL="3d2d8404-53f6-4fb6-a66b-f37c97198ad2" authority="Ortiz &amp; Gomez, 1986: 31" authorityName="Ortiz &amp; Gomez" authorityPageNumber="31" authorityYear="1986" box="[151,647,1205,1228]" class="Malacostraca" family="Pylochelidae" genus="Pylocheles" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="42" phylum="Arthropoda" rank="species" species="cubensis">
<emphasis id="B9563A6FFFEC6D5FA692FB40D8458C97" box="[151,365,1205,1228]" italics="true" pageId="1" pageNumber="42">Pylocheles cubensis</emphasis>
<treatmentCitation id="0A83C06CFFEC6D5FA771FB40DBAF8C97" author="Ortiz" box="[372,647,1205,1228]" page="31" pageId="1" pageNumber="42" year="1986">
<bibRefCitation id="EFB39B8CFFEC6D5FA771FB40DB488C97" author="Ortiz" box="[372,608,1205,1228]" pageId="1" pageNumber="46" refString="Ortiz, M. &amp; Gomez, O. (1986). Una nueva especie del genero Pylocheles (Anomura Galatheoidea) (sic), de las aguas profundas del sur de Cuba. Revista de Investigaciones Marinas, 7, 31 - 38." type="journal article" year="1986">Ortiz &amp; Gómez, 1986</bibRefCitation>
: 31
</treatmentCitation>
</taxonomicName>
, figs. 1–9.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B9DE67DFFEC6D5FA692FB20DA0D8CB7" blockId="1.[151,805,1205,1260]" box="[151,805,1237,1260]" pageId="1" pageNumber="42">
<treatmentCitationGroup id="AB32C153FFEC6D5FA692FB20DA0D8CB7" box="[151,805,1237,1260]" pageId="1" pageNumber="42">
<taxonomicName id="4C229DFEFFEC6D5FA692FB20DB8F8CB7" ID-CoL="bb14fbca-012c-4752-9058-0518708207ec" authority="Forest, 1987: 66" authorityName="Forest" authorityPageNumber="66" authorityYear="1987" box="[151,679,1237,1260]" class="Malacostraca" family="Pylochelidae" genus="Pylocheles" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="42" phylum="Arthropoda" rank="species" species="chacei" subGenus="Bathycheles">
<emphasis id="B9563A6FFFEC6D5FA692FB20D8238CB7" box="[151,267,1237,1260]" italics="true" pageId="1" pageNumber="42">Pylocheles</emphasis>
(
<emphasis id="B9563A6FFFEC6D5FA71FFB20D8B38CB7" box="[282,411,1237,1260]" italics="true" pageId="1" pageNumber="42">Bathycheles</emphasis>
)
<emphasis id="B9563A6FFFEC6D5FA7AFFB20D8C78CB7" box="[426,495,1237,1260]" italics="true" pageId="1" pageNumber="42">chacei</emphasis>
<treatmentCitation id="0A83C06CFFEC6D5FA7F3FB20DB8F8CB7" author="Forest" box="[502,679,1237,1260]" page="66" pageId="1" pageNumber="42" year="1987">
<bibRefCitation id="EFB39B8CFFEC6D5FA7F3FB20DB578CB7" author="Forest" box="[502,639,1237,1260]" pageId="1" pageNumber="46" refString="Forest, J. (1987). Les Pylochelidae ou &quot; Pagures symetriques &quot; (Crustacea Coenobitoidea). In: Resultats des campagnes MUSORSTOM. Memoires du Museum National d'Histoire Naturelle, Serie A, Zoologie, 137, 1 - 254." type="journal article" year="1987">Forest, 1987</bibRefCitation>
: 66
</treatmentCitation>
</taxonomicName>
, fig. 17a–d.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C338B5F6FFEC6D5FA692FAE8DA878DF4" pageId="1" pageNumber="42" type="materials_examined">
<paragraph id="8B9DE67DFFEC6D5FA692FAE8DAA68D04" blockId="1.[151,1437,1309,2015]" pageId="1" pageNumber="42">
<emphasis id="B9563A6FFFEC6D5FA692FAE8D8798D6C" bold="true" box="[151,337,1309,1335]" pageId="1" pageNumber="42">
<typeStatus id="549958DFFFEC6D5FA692FAE8D9FC8D6C" box="[151,212,1309,1335]" pageId="1" pageNumber="42">Type</typeStatus>
material.
</emphasis>
<emphasis id="B9563A6FFFEC6D5FA758FAE8DA038D6D" box="[349,811,1309,1334]" italics="true" pageId="1" pageNumber="42">
<typeStatus id="549958DFFFEC6D5FA758FAE8D8E18D6D" box="[349,457,1309,1334]" pageId="1" pageNumber="42" type="holotype">Holotype</typeStatus>
male of
<taxonomicName id="4C229DFEFFEC6D5FA439FAE8DA038D6D" box="[572,811,1309,1334]" class="Malacostraca" family="Pylochelidae" genus="Pylocheles" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="42" phylum="Arthropoda" rank="species" species="cubensis">Pylocheles cubensis</taxonomicName>
</emphasis>
(
<quantity id="4CDA4B98FFEC6D5FA545FAE8DAB58D6C" box="[832,925,1309,1335]" metricMagnitude="-2" metricUnit="m" metricValue="1.9" pageId="1" pageNumber="42" unit="mm" value="19.0">19 mm</quantity>
): Gulf of Cazones,
<collectingCountry id="F335A6EDFFEC6D5FA294FAE8DDFA8D6C" box="[1169,1234,1309,1335]" name="Cuba" pageId="1" pageNumber="42">Cuba</collectingCountry>
, mini-submarine
<taxonomicName id="4C229DFEFFEC6D5FA692FAB0D9F48D05" box="[151,220,1349,1374]" class="Insecta" family="Hesperiidae" genus="Argus" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="42" phylum="Arthropoda" rank="genus">
<emphasis id="B9563A6FFFEC6D5FA692FAB0D9F48D05" box="[151,220,1349,1374]" italics="true" pageId="1" pageNumber="42">Argus</emphasis>
</taxonomicName>
, stn
<quantity id="4CDA4B98FFEC6D5FA716FAB0D8EF8D04" box="[275,455,1349,1375]" metricMagnitude="4" metricUnit="m" metricValue="1.9905" metricValueMax="3.957" metricValueMin="0.024" pageId="1" pageNumber="42" unit="m" value="19905.0" valueMax="39570.0" valueMin="240.0">240–39, 570 m</quantity>
,
<date id="FF9CC0BDFFEC6D5FA7D6FAB0DB4E8D04" box="[467,614,1349,1375]" pageId="1" pageNumber="42" value="1983-10-25">25 Oct 1983</date>
, ANC/IDO.07.1.6.5.047.
</paragraph>
<paragraph id="8B9DE67DFFEC6D5FA6C3FA98DA878DF4" blockId="1.[151,1437,1309,2015]" pageId="1" pageNumber="42">
<emphasis id="B9563A6FFFEC6D5FA6C3FA98DA0E8DDD" box="[198,806,1389,1414]" italics="true" pageId="1" pageNumber="42">
<typeStatus id="549958DFFFEC6D5FA6C3FA98D8198DDD" box="[198,305,1389,1414]" pageId="1" pageNumber="42" type="holotype">Holotype</typeStatus>
female of
<taxonomicName id="4C229DFEFFEC6D5FA7AEFA98DA0E8DDD" box="[427,806,1389,1414]" class="Malacostraca" family="Pylochelidae" genus="Pylocheles" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="42" phylum="Arthropoda" rank="species" species="chacei" subGenus="Bathycheles">Pylocheles (Bathycheles) chacei</taxonomicName>
</emphasis>
(~
<quantity id="4CDA4B98FFEC6D5FA540FA98DAB48DDC" box="[837,924,1389,1415]" metricMagnitude="-2" metricUnit="m" metricValue="1.3" pageId="1" pageNumber="42" unit="mm" value="13.0">13 mm</quantity>
):
<collectingCountry id="F335A6EDFFEC6D5FA5B5FA98DDEC8DDC" box="[944,1220,1389,1415]" name="Turks and Caicos Islands" pageId="1" pageNumber="42">Turks &amp; Caicos Islands</collectingCountry>
, N.
<collectingCountry id="F335A6EDFFEC6D5FA2F6FA98DC188DDC" box="[1267,1328,1389,1415]" name="Haiti" pageId="1" pageNumber="42">Haiti</collectingCountry>
, R/V
<emphasis id="B9563A6FFFEC6D5FA376FA98D9D98DF5" italics="true" pageId="1" pageNumber="42">Silver Bay</emphasis>
stn 5142,
<geoCoordinate id="EE1680BAFFEC6D5FA76EFA60D8DD8DF4" box="[363,501,1429,1455]" direction="north" orientation="latitude" pageId="1" pageNumber="42" precision="9" value="19.866667">19º52.00’N</geoCoordinate>
,
<geoCoordinate id="EE1680BAFFEC6D5FA404FA60DBA78DF4" box="[513,655,1429,1455]" direction="west" orientation="longitude" pageId="1" pageNumber="42" precision="9" value="-71.975">71º58.50’W</geoCoordinate>
,
<quantity id="4CDA4B98FFEC6D5FA49FFA60DBC08DF4" box="[666,744,1429,1455]" metricMagnitude="2" metricUnit="m" metricValue="6.4" pageId="1" pageNumber="42" unit="m" value="640.0">640 m</quantity>
,
<collectionCode id="ED337EB8FFEC6D5FA4F6FA60DA608DF4" LSID="urn:lsid:biocol.org:col:34871" box="[755,840,1429,1455]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="1" pageNumber="42">USNM</collectionCode>
152529.
</paragraph>
</subSubSection>
<subSubSection id="C338B5F6FFEC6D5DA6C3FA48D8988E59" lastPageId="3" lastPageNumber="44" pageId="1" pageNumber="42" type="description">
<paragraph id="8B9DE67DFFEC6D5FA6C3FA48DB8D8EC4" blockId="1.[151,1437,1309,2015]" pageId="1" pageNumber="42">
<emphasis id="B9563A6FFFEC6D5FA6C3FA48D8778D8C" bold="true" box="[198,351,1469,1495]" pageId="1" pageNumber="42">Description.</emphasis>
Shield (
<figureCitation id="1319FAF8FFEC6D5FA7C0FA48DB208D8C" box="[453,520,1469,1495]" captionStart="FIGURE 1" captionStartId="2.[151,255,1527,1551]" captionTargetBox="[264,1323,354,1506]" captionTargetId="figure@2.[264,1323,354,1506]" captionTargetPageId="2" captionText="FIGURE 1. Pylocheles (Bathycheles) cubensis Ortiz &amp; Gómez, 1986, holotype male (19 mm) (ANC / IDO. 07.1.6.5.047). A, Dorsal view of whole animal (tied with strings to glass plate); B, chelae and carpi (dorsal view); C, anterior portion of shield, cephalic appendages, carpi of chelipeds, and portion of left second and third pereopods (dorsolateral view); D, enlarged dorsal view of carpi of chelipeds; E, anterior portion of shield, cephalic appendages, left cheliped, and carpi to dactyls of left second and third pereopods (lateral view); F, pleonal tergites 4 – 6, uropods and telson (dorsal and slightly posterior view); G, right first pleopod (internal view); H, right second pleopod (internal view). (Photographs by M. Ortiz)." httpUri="https://zenodo.org/record/273689/files/figure.png" pageId="1" pageNumber="42">Fig. 1</figureCitation>
A) slightly longer than broad (but broken and damaged in female), approximately 0.3 longer than posterior carapace; dorsal surface with shallow transverse groove subrostrally. Cervical groove faint to distinct laterally. Posterior median plate broad, moderately well calcified, at least anteriorly. Branchiostegites partially to well calcified dorsally and anteriorly, unarmed. Rostrum (
<figureCitation id="1319FAF8FFEC6D5FA2B4F9C0DDDE8E14" box="[1201,1270,1589,1615]" captionStart="FIGURE 1" captionStartId="2.[151,255,1527,1551]" captionTargetBox="[264,1323,354,1506]" captionTargetId="figure@2.[264,1323,354,1506]" captionTargetPageId="2" captionText="FIGURE 1. Pylocheles (Bathycheles) cubensis Ortiz &amp; Gómez, 1986, holotype male (19 mm) (ANC / IDO. 07.1.6.5.047). A, Dorsal view of whole animal (tied with strings to glass plate); B, chelae and carpi (dorsal view); C, anterior portion of shield, cephalic appendages, carpi of chelipeds, and portion of left second and third pereopods (dorsolateral view); D, enlarged dorsal view of carpi of chelipeds; E, anterior portion of shield, cephalic appendages, left cheliped, and carpi to dactyls of left second and third pereopods (lateral view); F, pleonal tergites 4 – 6, uropods and telson (dorsal and slightly posterior view); G, right first pleopod (internal view); H, right second pleopod (internal view). (Photographs by M. Ortiz)." httpUri="https://zenodo.org/record/273689/files/figure.png" pageId="1" pageNumber="42">Fig. 1</figureCitation>
A, C) absent, postocular projections bluntly triangular, unarmed. Lateral projections broadly rounded, produced considerably beyond levels of postocular projections.
</paragraph>
<paragraph id="8B9DE67DFFEC6D5FA6C3F958DBB68F4C" blockId="1.[151,1437,1309,2015]" pageId="1" pageNumber="42">Ocular peduncles 0.4–0.5 length of shield, each with few short setae in distal half of dorsal surface; corneal diameter 0.3–0.4 of peduncular length. Ocular acicles each as quite small, triangular plate situated slightly mesiad of basal peduncular midline.</paragraph>
<paragraph id="8B9DE67DFFEC6D5FA6C3F8D0DD698FD4" blockId="1.[151,1437,1309,2015]" pageId="1" pageNumber="42">Antennular peduncles overreaching distal corneal margins by approximately 0.5 lengths of basal segments. Ultimate segment approximately 0.7 length of subequal penultimate and basal segments; ultimate and penultimate segments each with several scattered setae, basal segment glabrous.</paragraph>
<paragraph id="8B9DE67DFFEC6D5CA6C3F868DABE8971" blockId="1.[151,1437,1309,2015]" lastBlockId="2.[151,1436,152,298]" lastPageId="2" lastPageNumber="43" pageId="1" pageNumber="42">Antennal peduncles with well calcified supernumerary segment; peduncles overreaching distal corneal margins by 0.5–0.8 lengths of fifth peduncular segments. Fifth and fourth segments each with numerous scattered fine setae; third segment unarmed and with only few short scattered setae; second segment with dorsolateral distal angle prominently produced, with terminal spine and 3 or 4 small spines on dorsolateral margin, 0–2 small spines basally and spinule on dorsal surface laterally, dorsomesial distal angle unarmed or with small spine. First segment with spine at ventrolateral distal angle.</paragraph>
<caption id="DF5DB6F5FFEF6D5CA692FA02D9F68E94" httpUri="https://zenodo.org/record/273689/files/figure.png" pageId="2" pageNumber="43" targetBox="[264,1323,354,1506]" targetPageId="2">
<paragraph id="8B9DE67DFFEF6D5CA692FA02D9F68E94" blockId="2.[151,1437,1527,1743]" pageId="2" pageNumber="43">
<emphasis id="B9563A6FFFEF6D5CA692FA02D8318E54" bold="true" box="[151,281,1527,1551]" pageId="2" pageNumber="43">FIGURE 1.</emphasis>
<taxonomicName id="4C229DFEFFEF6D5CA71AFA0DDA568E54" authority="Ortiz &amp; Gomez, 1986" authorityName="Ortiz &amp; Gomez" authorityYear="1986" box="[287,894,1528,1551]" class="Malacostraca" family="Pylochelidae" genus="Pylocheles" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="43" phylum="Arthropoda" rank="species" species="cubensis" subGenus="Bathycheles">
<emphasis id="B9563A6FFFEF6D5CA71AFA0DDBA68E54" box="[287,654,1528,1551]" italics="true" pageId="2" pageNumber="43">Pylocheles (Bathycheles) cubensis</emphasis>
Ortiz &amp; Gómez, 1986
</taxonomicName>
, holotype male (19 mm) (ANC/IDO.07.1.6.5.047). A, Dorsal view of whole animal (tied with strings to glass plate); B, chelae and carpi (dorsal view); C, anterior portion of shield, cephalic appendages, carpi of chelipeds, and portion of left second and third pereopods (dorsolateral view); D, enlarged dorsal view of carpi of chelipeds; E, anterior portion of shield, cephalic appendages, left cheliped, and carpi to dactyls of left second and third pereopods (lateral view); F, pleonal tergites 4–6, uropods and telson (dorsal and slightly posterior view); G, right first pleopod (internal view); H, right second pleopod (internal view). (Photographs by M. Ortiz).
</paragraph>
</caption>
<paragraph id="8B9DE67DFFEF6D5DA6C3F8DDD9C088E9" blockId="2.[151,1436,1832,2018]" lastBlockId="3.[151,1437,152,2018]" lastPageId="3" lastPageNumber="44" pageId="2" pageNumber="43">Mandible with incisor process well calcified, cutting margin entire. Maxillule with external lobe of endopod obsolete. Maxilla with endopod slightly overreaching distal margin of scaphognathite. First maxilliped with very slender exopod, epipod well developed. Second and third maxillipeds both pediform; second with dactyl approximately 0.7 length of propodus; third maxilliped with well developed crista dentata without accessory tooth, dactyl and propodus varying from approximately equal in length to propodus somewhat longer.</paragraph>
<paragraph id="8B9DE67DFFEE6D5DA6C3FF35DB998AE1" blockId="3.[151,1437,152,2018]" pageId="3" pageNumber="44">
Chelipeds (
<figureCitation id="1319FAF8FFEE6D5DA74BFF35D8BA8881" box="[334,402,192,218]" captionStart="FIGURE 1" captionStartId="2.[151,255,1527,1551]" captionTargetBox="[264,1323,354,1506]" captionTargetId="figure@2.[264,1323,354,1506]" captionTargetPageId="2" captionText="FIGURE 1. Pylocheles (Bathycheles) cubensis Ortiz &amp; Gómez, 1986, holotype male (19 mm) (ANC / IDO. 07.1.6.5.047). A, Dorsal view of whole animal (tied with strings to glass plate); B, chelae and carpi (dorsal view); C, anterior portion of shield, cephalic appendages, carpi of chelipeds, and portion of left second and third pereopods (dorsolateral view); D, enlarged dorsal view of carpi of chelipeds; E, anterior portion of shield, cephalic appendages, left cheliped, and carpi to dactyls of left second and third pereopods (lateral view); F, pleonal tergites 4 – 6, uropods and telson (dorsal and slightly posterior view); G, right first pleopod (internal view); H, right second pleopod (internal view). (Photographs by M. Ortiz)." httpUri="https://zenodo.org/record/273689/files/figure.png" pageId="3" pageNumber="44">Fig. 1</figureCitation>
A–D) symmetrical, subrectangular in shape. Left dactyl (right missing) slightly overlapped by fixed finger; dorsomesial margin with row of closely-spaced tubercles; dorsal, mesial and ventral surfaces all with tufts of short to moderately long setae, densest mesially; cutting edge with row of well developed calcareous teeth, terminating in calcareous claw. Dorsomesial and dorsolateral margins of palms each with row of tubercles not concealed by submarginal long setae, dorsal surfaces flat, unarmed, but with covering of sparse tufts of short setae generally forming longitudinal rows; mesial and lateral faces with tufts of moderately long and dense setae dorsally; ventral surfaces with scattered tufts of sparse setae. Carpi roundly subtriangular, each with dorsodistal margin considerably elevated and overhanging proximal margins of chelae; distal margins each cut into 2 lobes by deep incision in mesial 0.3, division continued posteriorly as distinct, arched depression, margin armed with row of small tubercles laterally and small spines dorsally, not concealed by tufts of long setae; dorsal surfaces (
<figureCitation id="1319FAF8FFEE6D5DA4E1FDA5DA0F8A31" box="[740,807,592,618]" captionStart="FIGURE 1" captionStartId="2.[151,255,1527,1551]" captionTargetBox="[264,1323,354,1506]" captionTargetId="figure@2.[264,1323,354,1506]" captionTargetPageId="2" captionText="FIGURE 1. Pylocheles (Bathycheles) cubensis Ortiz &amp; Gómez, 1986, holotype male (19 mm) (ANC / IDO. 07.1.6.5.047). A, Dorsal view of whole animal (tied with strings to glass plate); B, chelae and carpi (dorsal view); C, anterior portion of shield, cephalic appendages, carpi of chelipeds, and portion of left second and third pereopods (dorsolateral view); D, enlarged dorsal view of carpi of chelipeds; E, anterior portion of shield, cephalic appendages, left cheliped, and carpi to dactyls of left second and third pereopods (lateral view); F, pleonal tergites 4 – 6, uropods and telson (dorsal and slightly posterior view); G, right first pleopod (internal view); H, right second pleopod (internal view). (Photographs by M. Ortiz)." httpUri="https://zenodo.org/record/273689/files/figure.png" pageId="3" pageNumber="44">Fig. 1</figureCitation>
D) with short, transverse, tuberculate ridges in distal 0.2, larger and more widely-spaced, but less tuberculate, ridges in remaining distal 0.6, becoming much weaker proximally; ventral surfaces unarmed.
</paragraph>
<paragraph id="8B9DE67DFFEE6D5DA6C3FD3DD8608BF1" blockId="3.[151,1437,152,2018]" pageId="3" pageNumber="44">Ambulatory legs not overreaching tips of outstretched chelipeds. Dactyls slightly shorter to slightly longer than propodi; dorsal surfaces unarmed and generally glabrous, lateral and mesial faces each with weak longitudinal sulcus, surfaces each with row of moderately long and stiff, but slender setae dorsally and line of short rows of 3–6 or 7 stiff, moderately thick setae adjacent to ventral margin, lateral face also with row of widelyspaced tufts of sparse setae in distal half. Propodi, carpi, meri, and ischia unarmed but with sparse, fine setae on all surfaces.</paragraph>
<paragraph id="8B9DE67DFFEE6D5DA6C3FC4DDD628D49" blockId="3.[151,1437,152,2018]" pageId="3" pageNumber="44">
Pleon (
<figureCitation id="1319FAF8FFEE6D5DA71DFC4DD8718B89" box="[280,345,952,978]" captionStart="FIGURE 1" captionStartId="2.[151,255,1527,1551]" captionTargetBox="[264,1323,354,1506]" captionTargetId="figure@2.[264,1323,354,1506]" captionTargetPageId="2" captionText="FIGURE 1. Pylocheles (Bathycheles) cubensis Ortiz &amp; Gómez, 1986, holotype male (19 mm) (ANC / IDO. 07.1.6.5.047). A, Dorsal view of whole animal (tied with strings to glass plate); B, chelae and carpi (dorsal view); C, anterior portion of shield, cephalic appendages, carpi of chelipeds, and portion of left second and third pereopods (dorsolateral view); D, enlarged dorsal view of carpi of chelipeds; E, anterior portion of shield, cephalic appendages, left cheliped, and carpi to dactyls of left second and third pereopods (lateral view); F, pleonal tergites 4 – 6, uropods and telson (dorsal and slightly posterior view); G, right first pleopod (internal view); H, right second pleopod (internal view). (Photographs by M. Ortiz)." httpUri="https://zenodo.org/record/273689/files/figure.png" pageId="3" pageNumber="44">Fig. 1</figureCitation>
A) with pleura of pleomeres 2–5 distinctly delineated. Sixth pleonal tergite (
<figureCitation id="1319FAF8FFEE6D5DA2D1FC4DDC3E8B89" box="[1236,1302,952,978]" captionStart="FIGURE 1" captionStartId="2.[151,255,1527,1551]" captionTargetBox="[264,1323,354,1506]" captionTargetId="figure@2.[264,1323,354,1506]" captionTargetPageId="2" captionText="FIGURE 1. Pylocheles (Bathycheles) cubensis Ortiz &amp; Gómez, 1986, holotype male (19 mm) (ANC / IDO. 07.1.6.5.047). A, Dorsal view of whole animal (tied with strings to glass plate); B, chelae and carpi (dorsal view); C, anterior portion of shield, cephalic appendages, carpi of chelipeds, and portion of left second and third pereopods (dorsolateral view); D, enlarged dorsal view of carpi of chelipeds; E, anterior portion of shield, cephalic appendages, left cheliped, and carpi to dactyls of left second and third pereopods (lateral view); F, pleonal tergites 4 – 6, uropods and telson (dorsal and slightly posterior view); G, right first pleopod (internal view); H, right second pleopod (internal view). (Photographs by M. Ortiz)." httpUri="https://zenodo.org/record/273689/files/figure.png" pageId="3" pageNumber="44">Fig. 1</figureCitation>
F) with lateral margins each indented approximately at midlength or slightly more posteriorly and with shallow longitudinal furrow on dorsal surface medianly; terminal margin rounded, unarmed, but notched medianly by deep subquadrate concavity. Protopods of uropods each with prominent spine on posterior margin. Telson as long as broad, unequally divided by transverse suture, with anterior portion slightly narrower than posterior portion; anterior portion with weak depression on either side of midline anteriorly and with ovate area of decalcification at each posterolateral angle (
<figureCitation id="1319FAF8FFEE6D5DA442FB5DDBA18C99" box="[583,649,1192,1218]" captionStart="FIGURE 1" captionStartId="2.[151,255,1527,1551]" captionTargetBox="[264,1323,354,1506]" captionTargetId="figure@2.[264,1323,354,1506]" captionTargetPageId="2" captionText="FIGURE 1. Pylocheles (Bathycheles) cubensis Ortiz &amp; Gómez, 1986, holotype male (19 mm) (ANC / IDO. 07.1.6.5.047). A, Dorsal view of whole animal (tied with strings to glass plate); B, chelae and carpi (dorsal view); C, anterior portion of shield, cephalic appendages, carpi of chelipeds, and portion of left second and third pereopods (dorsolateral view); D, enlarged dorsal view of carpi of chelipeds; E, anterior portion of shield, cephalic appendages, left cheliped, and carpi to dactyls of left second and third pereopods (lateral view); F, pleonal tergites 4 – 6, uropods and telson (dorsal and slightly posterior view); G, right first pleopod (internal view); H, right second pleopod (internal view). (Photographs by M. Ortiz)." httpUri="https://zenodo.org/record/273689/files/figure.png" pageId="3" pageNumber="44">Fig. 1</figureCitation>
F); posterior portion with lateral margins rounded, terminal margin sometimes with very slight median indentation, and faint median concavity anteriorly, giving bilobed impression, surface with covering of long fine setae, at least posteriorly and marginally.
</paragraph>
<paragraph id="8B9DE67DFFEE6D5DA6C3FAD5D9E48DE9" blockId="3.[151,1437,152,2018]" pageId="3" pageNumber="44">
<collectingCountry id="F335A6EDFFEE6D5DA6C3FAD5D82A8D61" box="[198,258,1312,1338]" name="Malta" pageId="3" pageNumber="44">Male</collectingCountry>
first and second pleopods modified as gonopods. Pleopod 1 (
<figureCitation id="1319FAF8FFEE6D5DA5D7FAD5DD3C8D61" box="[978,1044,1312,1338]" captionStart="FIGURE 1" captionStartId="2.[151,255,1527,1551]" captionTargetBox="[264,1323,354,1506]" captionTargetId="figure@2.[264,1323,354,1506]" captionTargetPageId="2" captionText="FIGURE 1. Pylocheles (Bathycheles) cubensis Ortiz &amp; Gómez, 1986, holotype male (19 mm) (ANC / IDO. 07.1.6.5.047). A, Dorsal view of whole animal (tied with strings to glass plate); B, chelae and carpi (dorsal view); C, anterior portion of shield, cephalic appendages, carpi of chelipeds, and portion of left second and third pereopods (dorsolateral view); D, enlarged dorsal view of carpi of chelipeds; E, anterior portion of shield, cephalic appendages, left cheliped, and carpi to dactyls of left second and third pereopods (lateral view); F, pleonal tergites 4 – 6, uropods and telson (dorsal and slightly posterior view); G, right first pleopod (internal view); H, right second pleopod (internal view). (Photographs by M. Ortiz)." httpUri="https://zenodo.org/record/273689/files/figure.png" pageId="3" pageNumber="44">Fig. 1</figureCitation>
G) with long, moderately slender basal segment and much shorter subovate distal segment. Pleopod 2 (
<figureCitation id="1319FAF8FFEE6D5DA5DEFABDDD088D39" box="[987,1056,1352,1378]" captionStart="FIGURE 1" captionStartId="2.[151,255,1527,1551]" captionTargetBox="[264,1323,354,1506]" captionTargetId="figure@2.[264,1323,354,1506]" captionTargetPageId="2" captionText="FIGURE 1. Pylocheles (Bathycheles) cubensis Ortiz &amp; Gómez, 1986, holotype male (19 mm) (ANC / IDO. 07.1.6.5.047). A, Dorsal view of whole animal (tied with strings to glass plate); B, chelae and carpi (dorsal view); C, anterior portion of shield, cephalic appendages, carpi of chelipeds, and portion of left second and third pereopods (dorsolateral view); D, enlarged dorsal view of carpi of chelipeds; E, anterior portion of shield, cephalic appendages, left cheliped, and carpi to dactyls of left second and third pereopods (lateral view); F, pleonal tergites 4 – 6, uropods and telson (dorsal and slightly posterior view); G, right first pleopod (internal view); H, right second pleopod (internal view). (Photographs by M. Ortiz)." httpUri="https://zenodo.org/record/273689/files/figure.png" pageId="3" pageNumber="44">Fig. 1</figureCitation>
H) with moderately short, stout basal segment; distal segment with semi-articulated distal portion foliaceous and longer than proximal portion.
</paragraph>
<paragraph id="8B9DE67DFFEE6D5DA6C3FA4AD8BD8D81" blockId="3.[151,1437,152,2018]" box="[198,405,1471,1498]" pageId="3" pageNumber="44">
<emphasis id="B9563A6FFFEE6D5DA6C3FA4AD83B8D82" bold="true" box="[198,275,1471,1497]" pageId="3" pageNumber="44">Color.</emphasis>
Unknown.
</paragraph>
<paragraph id="8B9DE67DFFEE6D5DA6C3FA12D8988E59" blockId="3.[151,1437,152,2018]" box="[198,432,1511,1538]" pageId="3" pageNumber="44">
<emphasis id="B9563A6FFFEE6D5DA6C3FA12D8068E5A" bold="true" box="[198,302,1511,1537]" pageId="3" pageNumber="44">Habitat.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="C338B5F6FFEE6D5DA6C3F9FADDBF8E71" box="[198,1175,1551,1578]" pageId="3" pageNumber="44" type="distribution">
<paragraph id="8B9DE67DFFEE6D5DA6C3F9FADDBF8E71" blockId="3.[151,1437,152,2018]" box="[198,1175,1551,1578]" pageId="3" pageNumber="44">
<emphasis id="B9563A6FFFEE6D5DA6C3F9FAD84F8E72" bold="true" box="[198,359,1551,1577]" pageId="3" pageNumber="44">Distribution.</emphasis>
Gulf of Cazones,
<collectingCountry id="F335A6EDFFEE6D5DA439F9E5DB528E71" box="[572,634,1552,1578]" name="Cuba" pageId="3" pageNumber="44">Cuba</collectingCountry>
to
<collectingCountry id="F335A6EDFFEE6D5DA49AF9E5DA9D8E71" box="[671,949,1552,1578]" name="Turks and Caicos Islands" pageId="3" pageNumber="44">Turks &amp; Caicos Islands</collectingCountry>
,
<collectingCountry id="F335A6EDFFEE6D5DA5C4F9E5DAD78E71" box="[961,1023,1552,1578]" name="Haiti" pageId="3" pageNumber="44">Haiti</collectingCountry>
;
<quantity id="4CDA4B98FFEE6D5DA20EF9E5DDBB8E71" box="[1035,1171,1552,1578]" metricMagnitude="2" metricUnit="m" metricValue="6.05" metricValueMax="6.4" metricValueMin="5.7" pageId="3" pageNumber="44" unit="m" value="605.0" valueMax="640.0" valueMin="570.0">
570–
<specimenCount id="9D242DF4FFEE6D5DA243F9E5DDBB8E71" box="[1094,1171,1552,1578]" pageId="3" pageNumber="44" type="generic">640 m</specimenCount>
</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="C338B5F6FFEE6D5AA6C3F9C2D8B68D61" lastPageId="4" lastPageNumber="45" pageId="3" pageNumber="44" type="discussion">
<paragraph id="8B9DE67DFFEE6D5DA6C3F9C2DCBE8FC9" blockId="3.[151,1437,152,2018]" pageId="3" pageNumber="44">
<emphasis id="B9563A6FFFEE6D5DA6C3F9C2D8688E0A" bold="true" box="[198,320,1591,1617]" pageId="3" pageNumber="44">Remarks.</emphasis>
Only one other species of
<taxonomicName id="4C229DFEFFEE6D5DA473F9CDDBDD8E0A" box="[630,757,1592,1617]" class="Malacostraca" family="Pylochelidae" genus="Pylocheles" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="44" phylum="Arthropoda" rank="genus">
<emphasis id="B9563A6FFFEE6D5DA473F9CDDBDD8E0A" box="[630,757,1592,1617]" italics="true" pageId="3" pageNumber="44">Pylocheles</emphasis>
</taxonomicName>
,
<taxonomicName id="4C229DFEFFEE6D5DA506F9CDDA9D8E0A" box="[771,949,1592,1617]" class="Malacostraca" family="Pylochelidae" genus="Pylocheles" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="44" phylum="Arthropoda" rank="species" species="agassizii" subGenus="P.">
<emphasis id="B9563A6FFFEE6D5DA506F9CDDA9D8E0A" box="[771,949,1592,1617]" italics="true" pageId="3" pageNumber="44">P. (P.) agassizii</emphasis>
</taxonomicName>
, is known from the Western Atlantic, and it is immediately distinguished from
<taxonomicName id="4C229DFEFFEE6D5DA44DF995DBD68E22" box="[584,766,1632,1657]" class="Malacostraca" family="Pylochelidae" genus="Pylocheles" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="44" phylum="Arthropoda" rank="species" species="cubensis" subGenus="B .">
<emphasis id="B9563A6FFFEE6D5DA44DF995DBD68E22" box="[584,766,1632,1657]" italics="true" pageId="3" pageNumber="44">P. (B.) cubensis</emphasis>
</taxonomicName>
by the armament of the dorsal surfaces of the chelae, the posterior margin of the sixth pleonal tergite and the structure of the telson. In
<taxonomicName id="4C229DFEFFEE6D5DA236F97DDDC18EFA" box="[1075,1257,1672,1697]" class="Malacostraca" family="Pylochelidae" genus="Pylocheles" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="44" phylum="Arthropoda" rank="species" species="agassizii" subGenus="P.">
<emphasis id="B9563A6FFFEE6D5DA236F97DDDC18EFA" box="[1075,1257,1672,1697]" italics="true" pageId="3" pageNumber="44">P. (P.) agassizii</emphasis>
</taxonomicName>
, the chelae are armed on the dorsal surfaces with numerous tubercles or small spines; the posterior margin of the sixth pleonal tergite is weakly cut into three subequal lobes by small incisions; the anterior portion of the telson is completely calcified; and the posterior portion of the telson is divided into two circular lobes. In contrast, the dorsal surfaces of the chelae of
<taxonomicName id="4C229DFEFFEE6D5DA412F8DDDBFA8F1A" box="[535,722,1832,1857]" class="Malacostraca" family="Pylochelidae" genus="Pylocheles" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="44" phylum="Arthropoda" rank="species" species="cubensis" subGenus="Bathycheles">
<emphasis id="B9563A6FFFEE6D5DA412F8DDDBFA8F1A" box="[535,722,1832,1857]" italics="true" pageId="3" pageNumber="44">P. (B.) cubensis</emphasis>
</taxonomicName>
are provided only with rows of tufts of setae; the posterior margin of the sixth pleonal tergite has a prominent, subquadrate median concavity; the anterior margin of the telson has two membranous patches near the outer angles; and the posterior portion of the telson is undivided.
</paragraph>
<paragraph id="8B9DE67DFFEE6D5AA6C3F855DBB58991" blockId="3.[151,1437,152,2018]" lastBlockId="4.[151,1437,152,1338]" lastPageId="4" lastPageNumber="45" pageId="3" pageNumber="44">
<collectingCountry id="F335A6EDFFEE6D5DA6C3F855D9CF8FE1" box="[198,231,1952,1978]" name="American Samoa" pageId="3" pageNumber="44">As</collectingCountry>
stated by
<bibRefCitation id="EFB39B8CFFEE6D5DA758F855D8D48FE1" author="Forest" box="[349,508,1952,1978]" pageId="3" pageNumber="46" refString="Forest, J. (1987). Les Pylochelidae ou &quot; Pagures symetriques &quot; (Crustacea Coenobitoidea). In: Resultats des campagnes MUSORSTOM. Memoires du Museum National d'Histoire Naturelle, Serie A, Zoologie, 137, 1 - 254." type="journal article" year="1987">Forest (1987)</bibRefCitation>
for
<taxonomicName id="4C229DFEFFEE6D5DA42EF855DBEF8FE2" box="[555,711,1952,1977]" class="Malacostraca" family="Pylochelidae" genus="Pylocheles" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="44" phylum="Arthropoda" rank="species" species="chacei" subGenus="Bathycheles">
<emphasis id="B9563A6FFFEE6D5DA42EF855DBEF8FE2" box="[555,711,1952,1977]" italics="true" pageId="3" pageNumber="44">P. (B.) chacei</emphasis>
</taxonomicName>
, this taxon is morphologically most closely allied to the Indo- Pacific species
<taxonomicName id="4C229DFEFFEE6D5DA749F83DDBA38FB9" authority="Forest, 1987" authorityName="Forest" authorityYear="1987" box="[332,651,1992,2018]" class="Malacostraca" family="Pylochelidae" genus="Pylocheles" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="44" phylum="Arthropoda" rank="species" species="incisus" subGenus="Bathycheles">
<emphasis id="B9563A6FFFEE6D5DA749F83DD8C78FBA" box="[332,495,1992,2017]" italics="true" pageId="3" pageNumber="44">P. (B.) incisus</emphasis>
<bibRefCitation id="EFB39B8CFFEE6D5DA7F2F83DDBA38FB9" author="Forest" box="[503,651,1992,2018]" pageId="3" pageNumber="46" refString="Forest, J. (1987). Les Pylochelidae ou &quot; Pagures symetriques &quot; (Crustacea Coenobitoidea). In: Resultats des campagnes MUSORSTOM. Memoires du Museum National d'Histoire Naturelle, Serie A, Zoologie, 137, 1 - 254." type="journal article" year="1987">Forest, 1987</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C229DFEFFEE6D5DA4C0F83DDD368FB9" authority="Forest, 1987" authorityName="Forest" authorityYear="1987" box="[709,1054,1992,2018]" class="Malacostraca" family="Pylochelidae" genus="Pylocheles" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="44" phylum="Arthropoda" rank="species" species="crosnieri" subGenus="B .">
<emphasis id="B9563A6FFFEE6D5DA4C0F83DDAA88FBA" box="[709,896,1992,2017]" italics="true" pageId="3" pageNumber="44">P. (B.) crosnieri</emphasis>
<bibRefCitation id="EFB39B8CFFEE6D5DA582F83DDD368FB9" author="Forest" box="[903,1054,1992,2018]" pageId="3" pageNumber="46" refString="Forest, J. (1987). Les Pylochelidae ou &quot; Pagures symetriques &quot; (Crustacea Coenobitoidea). In: Resultats des campagnes MUSORSTOM. Memoires du Museum National d'Histoire Naturelle, Serie A, Zoologie, 137, 1 - 254." type="journal article" year="1987">Forest, 1987</bibRefCitation>
</taxonomicName>
. Forest noted the similarity with
<emphasis id="B9563A6FFFE96D5AA692FF6DD87B88EA" box="[151,339,152,177]" italics="true" pageId="4" pageNumber="45">P. (B.) c ro s n i e r i</emphasis>
in the development of the anterior margin of the shield, or lack thereof, and with
<taxonomicName id="4C229DFEFFE96D5AA34AFF6DD9C18882" class="Malacostraca" family="Pylochelidae" genus="Pylocheles" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="45" phylum="Arthropoda" rank="species" species="incisus" subGenus="B .">
<emphasis id="B9563A6FFFE96D5AA34AFF6DD9C18882" italics="true" pageId="4" pageNumber="45">P. (B.) incisus</emphasis>
</taxonomicName>
in the notch in the posterior margin of the sixth pleonal tergite. The dorsal surfaces of the chelae of both Indo-Pacific species are provided only with tufts of setae, as is the case in
<taxonomicName id="4C229DFEFFE96D5AA244FF1DDDD0895A" box="[1089,1272,232,257]" class="Malacostraca" family="Pylochelidae" genus="Pylocheles" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="45" phylum="Arthropoda" rank="species" species="cubensis" subGenus="Bathycheles">
<emphasis id="B9563A6FFFE96D5AA244FF1DDDD0895A" box="[1089,1272,232,257]" italics="true" pageId="4" pageNumber="45">P. (B.) cubensis</emphasis>
</taxonomicName>
, thus Forest’s (1987) assessment is confirmed. The structures of the mouthparts of
<taxonomicName id="4C229DFEFFE96D5AA5EFFEE5DDBD8972" box="[1002,1173,272,297]" class="Malacostraca" family="Pylochelidae" genus="Pylocheles" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="45" phylum="Arthropoda" rank="species" species="incisus" subGenus="Bathycheles">
<emphasis id="B9563A6FFFE96D5AA5EFFEE5DDBD8972" box="[1002,1173,272,297]" italics="true" pageId="4" pageNumber="45">P. (B.) incisus</emphasis>
</taxonomicName>
, illustrated by
<bibRefCitation id="EFB39B8CFFE96D5AA356FEE5D8148909" author="Forest" pageId="4" pageNumber="46" refString="Forest, J. (1987). Les Pylochelidae ou &quot; Pagures symetriques &quot; (Crustacea Coenobitoidea). In: Resultats des campagnes MUSORSTOM. Memoires du Museum National d'Histoire Naturelle, Serie A, Zoologie, 137, 1 - 254." type="journal article" year="1987">Forest (1987, fig 16)</bibRefCitation>
as representative of the subgenus, agree well with those illustrated by
<bibRefCitation id="EFB39B8CFFE96D5AA28EFECDDCB48909" author="Ortiz" box="[1163,1436,312,338]" pageId="4" pageNumber="46" refString="Ortiz, M. &amp; Gomez, O. (1986). Una nueva especie del genero Pylocheles (Anomura Galatheoidea) (sic), de las aguas profundas del sur de Cuba. Revista de Investigaciones Marinas, 7, 31 - 38." type="journal article" year="1986">Ortiz &amp; Gómez (1986)</bibRefCitation>
for
<taxonomicName id="4C229DFEFFE96D5AA6C5FE95D85F8922" box="[192,375,352,377]" class="Malacostraca" family="Pylochelidae" genus="Pylocheles" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="45" phylum="Arthropoda" rank="species" species="cubensis" subGenus="Bathycheles">
<emphasis id="B9563A6FFFE96D5AA6C5FE95D85F8922" box="[192,375,352,377]" italics="true" pageId="4" pageNumber="45">P. (B.) cubensis</emphasis>
</taxonomicName>
and those observed in the
<typeStatus id="549958DFFFE96D5AA4B4FE95DA308921" box="[689,792,352,378]" pageId="4" pageNumber="45" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C229DFEFFE96D5AA53AFE95DAF48922" box="[831,988,352,377]" class="Malacostraca" family="Pylochelidae" genus="Pylocheles" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="45" phylum="Arthropoda" rank="species" species="chacei" subGenus="Bathycheles">
<emphasis id="B9563A6FFFE96D5AA53AFE95DAF48922" box="[831,988,352,377]" italics="true" pageId="4" pageNumber="45">P. (B.) chacei</emphasis>
</taxonomicName>
. The male gonopods of
<taxonomicName id="4C229DFEFFE96D5AA2FEFE95D99E89FA" class="Malacostraca" family="Pylochelidae" genus="Pylocheles" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="45" phylum="Arthropoda" rank="species" species="cubensis" subGenus="Bathycheles">
<emphasis id="B9563A6FFFE96D5AA2FEFE95D99E89FA" italics="true" pageId="4" pageNumber="45">P. (B.) cubensis</emphasis>
</taxonomicName>
agree more closely with those of
<taxonomicName id="4C229DFEFFE96D5AA444FE7DDBD189FA" box="[577,761,392,417]" class="Malacostraca" family="Pylochelidae" genus="Pylocheles" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="45" phylum="Arthropoda" rank="species" species="crosnieri" subGenus="Bathycheles">
<emphasis id="B9563A6FFFE96D5AA444FE7DDBD189FA" box="[577,761,392,417]" italics="true" pageId="4" pageNumber="45">P. (B.) crosnieri</emphasis>
</taxonomicName>
, as illustrated by
<bibRefCitation id="EFB39B8CFFE96D5AA5CCFE7DDD4A89F9" author="Forest" box="[969,1122,392,418]" pageId="4" pageNumber="46" refString="Forest, J. (1987). Les Pylochelidae ou &quot; Pagures symetriques &quot; (Crustacea Coenobitoidea). In: Resultats des campagnes MUSORSTOM. Memoires du Museum National d'Histoire Naturelle, Serie A, Zoologie, 137, 1 - 254." type="journal article" year="1987">Forest (1987</bibRefCitation>
: fig. 9k, l), than those of
<taxonomicName id="4C229DFEFFE96D5AA383FE7DD8F38991" authority="Forest, 1987" authorityName="Forest" authorityYear="1987" class="Malacostraca" family="Pylochelidae" genus="Pylocheles" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="45" phylum="Arthropoda" rank="species" species="profundus" subGenus="Bathycheles">
<emphasis id="B9563A6FFFE96D5AA383FE7DD8698992" italics="true" pageId="4" pageNumber="45">P. (B.) profundus</emphasis>
<bibRefCitation id="EFB39B8CFFE96D5AA74DFE45D8F38991" author="Forest" box="[328,475,432,458]" pageId="4" pageNumber="46" refString="Forest, J. (1987). Les Pylochelidae ou &quot; Pagures symetriques &quot; (Crustacea Coenobitoidea). In: Resultats des campagnes MUSORSTOM. Memoires du Museum National d'Histoire Naturelle, Serie A, Zoologie, 137, 1 - 254." type="journal article" year="1987">Forest, 1987</bibRefCitation>
</taxonomicName>
(ibid.: fig.
<date id="FF9CC0BDFFE96D5AA45AFE45DB508991" box="[607,632,432,458]" pageId="4" pageNumber="45">9i</date>
, j).
</paragraph>
<paragraph id="8B9DE67DFFE96D5AA6C3FE2DD86A8B51" blockId="4.[151,1437,152,1338]" pageId="4" pageNumber="45">
Certain discrepancies and inaccuracies appearing in the original descriptions of
<emphasis id="B9563A6FFFE96D5AA289FE2DDC2589AA" box="[1164,1293,472,497]" italics="true" pageId="4" pageNumber="45">P. c u b e n s i s</emphasis>
and
<taxonomicName id="4C229DFEFFE96D5AA34AFE2DD9CC8A42" class="Malacostraca" family="Pylochelidae" genus="Pylocheles" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="45" phylum="Arthropoda" rank="species" species="chacei" subGenus="B .">
<emphasis id="B9563A6FFFE96D5AA34AFE2DD9CC8A42" italics="true" pageId="4" pageNumber="45">P. (B.) chacei</emphasis>
</taxonomicName>
are corrected herein. For
<taxonomicName id="4C229DFEFFE96D5AA415FDF5DBED8A42" box="[528,709,512,537]" class="Malacostraca" family="Pylochelidae" genus="Pylocheles" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="45" phylum="Arthropoda" rank="species" species="cubensis" subGenus="Bathycheles">
<emphasis id="B9563A6FFFE96D5AA415FDF5DBED8A42" box="[528,709,512,537]" italics="true" pageId="4" pageNumber="45">P. (B.) cubensis</emphasis>
</taxonomicName>
, there are some errors in the illustrations and labeling of Ortiz &amp; Gómez’s (1986) figures 8 and 9 of the mouthparts. Specifically, figure 8 is labeled first maxilla when it is actually the maxilla (or second maxilla), the coxal endite is depicted as being trilobed, and the scaphognathite is not illustrated. Figure 9 is labeled second maxilla, but is actually the maxillule (or first maxilla). Additionally, the coxal endite of the first maxilliped (
<bibRefCitation id="EFB39B8CFFE96D5AA4A8FD55DAD58AE1" author="Ortiz" box="[685,1021,672,698]" pageId="4" pageNumber="46" refString="Ortiz, M. &amp; Gomez, O. (1986). Una nueva especie del genero Pylocheles (Anomura Galatheoidea) (sic), de las aguas profundas del sur de Cuba. Revista de Investigaciones Marinas, 7, 31 - 38." type="journal article" year="1986">Ortiz &amp; Gómez 1986, fig. 7</bibRefCitation>
) was not illustrated, and while the exopod probably was not omitted, the fact that it is very slender and tends to overlap the endopod was not clearly shown.
</paragraph>
<paragraph id="8B9DE67DFFE96D5AA6C3FCEDD8608C79" blockId="4.[151,1437,152,1338]" pageId="4" pageNumber="45">
The illustration of the
<typeStatus id="549958DFFFE96D5AA7D4FCEDDB1F8B69" box="[465,567,792,818]" pageId="4" pageNumber="45" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C229DFEFFE96D5AA464FCEDDA2A8B6A" box="[609,770,792,817]" class="Malacostraca" family="Pylochelidae" genus="Pylocheles" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="45" phylum="Arthropoda" rank="species" species="chacei" subGenus="Bathycheles">
<emphasis id="B9563A6FFFE96D5AA464FCEDDA2A8B6A" box="[609,770,792,817]" italics="true" pageId="4" pageNumber="45">P. (B.) chacei</emphasis>
</taxonomicName>
(
<bibRefCitation id="EFB39B8CFFE96D5AA514FCEDDD238B69" author="Forest" box="[785,1035,792,818]" pageId="4" pageNumber="46" refString="Forest, J. (1987). Les Pylochelidae ou &quot; Pagures symetriques &quot; (Crustacea Coenobitoidea). In: Resultats des campagnes MUSORSTOM. Memoires du Museum National d'Histoire Naturelle, Serie A, Zoologie, 137, 1 - 254." type="journal article" year="1987" yearSuffix="a">Forest 1987, fig. 17a</bibRefCitation>
) although a composite, is reasonably accurate for the cephalothorax and cephalic appendages; however, the accuracy of illustrated portion of carpus of the left cheliped (fig. 17c) can not be verified as there is very little now remaining of that surface. The illustration of the right second pereopod (fig. 17d) is proportionally accurate, but the lines on the lateral face of the dactyl are not corneous spines as they appear, but indications of where the short rows of setae occur. Similarly, the median and dorsal marks on that surface also indicate the positions of the setation described here.
</paragraph>
<paragraph id="8B9DE67DFFE96D5AA6C3FBC5D8B68D61" blockId="4.[151,1437,152,1338]" pageId="4" pageNumber="45">
The ocular peduncles of the
<typeStatus id="549958DFFFE96D5AA421FBC5DBA38C11" box="[548,651,1072,1098]" pageId="4" pageNumber="45" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C229DFEFFE96D5AA4BDFBC5DA5D8C12" box="[696,885,1072,1097]" class="Malacostraca" family="Pylochelidae" genus="Pylocheles" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="45" phylum="Arthropoda" rank="species" species="cubensis" subGenus="Bathycheles">
<emphasis id="B9563A6FFFE96D5AA4BDFBC5DA5D8C12" box="[696,885,1072,1097]" italics="true" pageId="4" pageNumber="45">P. (B.) cubensis</emphasis>
</taxonomicName>
are slightly slenderer and the corneas a little smaller than seen in the
<typeStatus id="549958DFFFE96D5AA7BDFBADDB378C29" box="[440,543,1112,1138]" pageId="4" pageNumber="45" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C229DFEFFE96D5AA44DFBADDBCF8C2A" box="[584,743,1112,1137]" class="Malacostraca" family="Pylochelidae" genus="Pylocheles" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="45" phylum="Arthropoda" rank="species" species="chacei" subGenus="B .">
<emphasis id="B9563A6FFFE96D5AA44DFBADDBCF8C2A" box="[584,743,1112,1137]" italics="true" pageId="4" pageNumber="45">P. (B.) chacei</emphasis>
</taxonomicName>
. Whether these differences reflect size or sex variations is not known at present. The mouthparts are exposed in the damaged anterior cephalothorax of the
<typeStatus id="549958DFFFE96D5AA2FFFB75DC488CC1" box="[1274,1376,1152,1178]" pageId="4" pageNumber="45" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C229DFEFFE96D5AA383FB75D8308C9A" class="Malacostraca" family="Pylochelidae" genus="Pylocheles" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="45" phylum="Arthropoda" rank="species" species="chacei" subGenus="Bathycheles">
<emphasis id="B9563A6FFFE96D5AA383FB75D8308C9A" italics="true" pageId="4" pageNumber="45">P. (B.) chacei</emphasis>
</taxonomicName>
, and these agree well with the illustrated mouthparts of
<taxonomicName id="4C229DFEFFE96D5AA5B1FB5DDD448C9A" box="[948,1132,1192,1217]" class="Malacostraca" family="Pylochelidae" genus="Pylocheles" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="45" phylum="Arthropoda" rank="species" species="cubensis" subGenus="Bathycheles">
<emphasis id="B9563A6FFFE96D5AA5B1FB5DDD448C9A" box="[948,1132,1192,1217]" italics="true" pageId="4" pageNumber="45">P. (B.) cubensis</emphasis>
</taxonomicName>
. However, the dactyl and propodus of the pediform third maxilliped of the former taxon are approximately equal in length, whereas they are subequal in
<taxonomicName id="4C229DFEFFE96D5AA78EFB0DDB6A8D4A" box="[395,578,1272,1297]" class="Malacostraca" family="Pylochelidae" genus="Pylocheles" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="45" phylum="Arthropoda" rank="species" species="cubensis" subGenus="Bathycheles">
<emphasis id="B9563A6FFFE96D5AA78EFB0DDB6A8D4A" box="[395,578,1272,1297]" italics="true" pageId="4" pageNumber="45">P. (B.) cubensis</emphasis>
</taxonomicName>
. This again may reflect a size or sex variation as the male is appreciably larger than the female.
</paragraph>
</subSubSection>
</treatment>
</document>
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