treatments-xml/data/03/88/50/038850393F02FFA5FF4FFBD6FC2FFD3C.xml

<document id="1DB79F50716057D3367D7F99BB5D1440" ID-CLB-Dataset="31294" ID-DOI="10.11646/zootaxa.4358.1.6" ID-GBIF-Dataset="1320c88d-0b17-433b-ac7b-0c7002855497" ID-ISSN="1175-5326" ID-Zenodo-Dep="1067516" ID-ZooBank="3403A972-99A2-4FC8-8917-8F3E0A42F903" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1511863087629" checkinUser="plazi" docAuthor="Felippe-Bauer, Maria Luiza, Cardoso, Erick Aragão &amp; Trindade, Rosimeire Lopes Da" docDate="2017" docId="038850393F02FFA5FF4FFBD6FC2FFD3C" docLanguage="en" docName="zootaxa.4358.1.6.pdf" docOrigin="Zootaxa 4358 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Monohelea cornuta Felippe-Bauer &amp; Cardoso &amp; Trindade 2017, sp. nov." docType="treatment" docVersion="7" masterDocId="FFB128413F06FFA3FFD8FFA9FF95FFCC" masterDocTitle="New species and new records of Monohelea Kieffer from eastern Amazon, Brazil (Diptera: Ceratopogonidae)" updateTime="1698504929183" updateUser="ExternalLinkService">
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<mods:title id="8D9506E82269D33B807D50E99CF5FFDB">New species and new records of Monohelea Kieffer from eastern Amazon, Brazil (Diptera: Ceratopogonidae)</mods:title>
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<mods:namePart id="D1DEC40F6B948A935F7B5541CE2A20F0">Felippe-Bauer, Maria Luiza</mods:namePart>
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<mods:namePart id="5E35268877BD2842E2936B42B199594C">Cardoso, Erick Aragão</mods:namePart>
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<mods:namePart id="6AF98311229A432478A0BD244FF9C3D7">Trindade, Rosimeire Lopes Da</mods:namePart>
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<mods:title id="81D1F28200266105402D2EB02B1DB664">Zootaxa</mods:title>
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<mods:part id="0F2436D9C7C549C1E3EC0614F94AADFF">
<mods:date id="0022EE11D28510F9C8D98DF8A27C49C9">2017</mods:date>
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<mods:number id="88983370FA99D60E970376FF818DE646">2017-11-28</mods:number>
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<mods:number id="39430CD859FFC8BAAB1342FFF6C62B1B">4358</mods:number>
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<treatment id="038850393F02FFA5FF4FFBD6FC2FFD3C" ID-DOI="http://doi.org/10.5281/zenodo.6033290" ID-GBIF-Taxon="137580704" ID-Zenodo-Dep="6033290" LSID="urn:lsid:plazi:treatment:038850393F02FFA5FF4FFBD6FC2FFD3C" httpUri="http://treatment.plazi.org/id/038850393F02FFA5FF4FFBD6FC2FFD3C" lastPageId="6" pageId="4">
<subSubSection id="C33BB2A43F02FFA7FF4FFBD6FE3CFB77" pageId="4" type="nomenclature">
<paragraph id="8B9EE12F3F02FFA7FF4FFBD6FCABFB56" blockId="4.[151,830,1151,1211]" box="[151,830,1151,1178]" pageId="4">
<heading id="D0D656433F02FFA7FF4FFBD6FCABFB56" bold="true" box="[151,830,1151,1178]" fontSize="11" level="1" pageId="4" reason="1">
<emphasis id="B9553D3D3F02FFA7FF4FFBD6FCABFB56" bold="true" box="[151,830,1151,1178]" pageId="4">
<taxonomicName id="4C219AAC3F02FFA7FF4FFBD6FD4CFB56" ID-CoL="9HYK9" authority="Felippe-Bauer &amp; Trindade" authorityName="Felippe-Bauer &amp; Cardoso &amp; Trindade" authorityYear="2017" box="[151,729,1151,1178]" class="Insecta" family="Ceratopogonidae" genus="Monohelea" kingdom="Animalia" order="Diptera" pageId="4" phylum="Arthropoda" rank="species" species="cornuta" status="sp. nov.">
<emphasis id="B9553D3D3F02FFA7FF4FFBD6FE10FB55" bold="true" box="[151,389,1151,1177]" italics="true" pageId="4">Monohelea cornuta</emphasis>
Felippe-Bauer &amp; Trindade
</taxonomicName>
<taxonomicNameLabel id="A26680463F02FFA7FD39FB29FCABFB56" box="[737,830,1152,1178]" pageId="4" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B9EE12F3F02FFA7FF4FFB0BFE3CFB77" blockId="4.[151,830,1151,1211]" box="[151,425,1186,1211]" pageId="4">
(
<figureCitation id="131AFDAA3F02FFA7FF47FB0BFF78FB77" box="[159,237,1186,1211]" captionStart="FIGURE 1" captionStartId="1.[151,250,1937,1959]" captionTargetBox="[300,1276,230,1887]" captionTargetId="figure@1.[232,1364,193,1921]" captionTargetPageId="1" captionText="FIGURE 1. Distribution of Monohelea species in the Municipalities of Capitão Poço, Ourém, Tracuateua and Viseu from Northeast Mesoregion, and in the municipality of Santa Bárbara do Pará from Metropolitan Belém Mesoregion, Para State, Brazil." httpUri="https://zenodo.org/record/1067518/files/figure.png" pageId="4">Figs. 1</figureCitation>
; 3 A–G; 9 C–D)
</paragraph>
</subSubSection>
<subSubSection id="C33BB2A43F02FFA7FF4FFB43FBD1FA87" pageId="4" type="diagnosis">
<paragraph id="8B9EE12F3F02FFA7FF4FFB43FBD1FA87" blockId="4.[151,1436,1258,2012]" pageId="4">
<emphasis id="B9553D3D3F02FFA7FF4FFB43FE84FACF" bold="true" box="[151,273,1258,1283]" pageId="4">Diagnosis.</emphasis>
The only Neotropical species of 
<taxonomicName id="4C219AAC3F02FFA7FD56FB42FC99FACF" authorityName="Kieffer" authorityYear="1917" box="[654,780,1259,1283]" class="Insecta" family="Ceratopogonidae" genus="Monohelea" kingdom="Animalia" order="Diptera" pageId="4" phylum="Arthropoda" rank="genus">
<emphasis id="B9553D3D3F02FFA7FD56FB42FC99FACF" box="[654,780,1259,1283]" italics="true" pageId="4">Monohelea</emphasis>
</taxonomicName>
in which the males have yellowish legs, hind femur with infuscated base, mid leg without distinct mesal brown marks, paramere with a sclerotized mesal hook-shaped process, and distal portion of paramere truncated and overlapped. Female unknown.
</paragraph>
</subSubSection>
<subSubSection id="C33BB2A43F02FFA5FF1FFAF0FE0FFF7C" lastPageId="6" pageId="4" type="description">
<paragraph id="8B9EE12F3F02FFA7FF1FFAF0FE7FFA76" blockId="4.[151,1436,1258,2012]" pageId="4">
<emphasis id="B9553D3D3F02FFA7FF1FFAF0FE9EFABE" bold="true" box="[199,267,1369,1394]" pageId="4">Male.</emphasis>
Head: eyes widely separated (
<figureCitation id="131AFDAA3F02FFA7FDA1FAF0FD76FABE" box="[633,739,1369,1394]" captionStart="FIGURE 3" captionStartId="5.[151,250,1905,1927]" captionTargetBox="[203,1401,398,1871]" captionTargetId="figure@5.[181,1425,373,1895]" captionTargetPageId="5" captionText="FIGURE 3. Monohelea cornuta Felippe-Bauer &amp; Trindade sp. nov., male. A. Wing; B. Thorax showing halter; C. Head, anterior view; D. Palpus, lateral view; E. Abdomen, ventral view; F. Legs (left to right) of fore, mid-, hindlegs, lateral view; G. Genitalia, ventral view.’=" httpUri="https://zenodo.org/record/1067522/files/figure.png" pageId="4">Fig. 3 C</figureCitation>
). Antenna missing. Palpus (
<figureCitation id="131AFDAA3F02FFA7FBE8FAF0FB0DFABE" box="[1072,1176,1369,1394]" captionStart="FIGURE 3" captionStartId="5.[151,250,1905,1927]" captionTargetBox="[203,1401,398,1871]" captionTargetId="figure@5.[181,1425,373,1895]" captionTargetPageId="5" captionText="FIGURE 3. Monohelea cornuta Felippe-Bauer &amp; Trindade sp. nov., male. A. Wing; B. Thorax showing halter; C. Head, anterior view; D. Palpus, lateral view; E. Abdomen, ventral view; F. Legs (left to right) of fore, mid-, hindlegs, lateral view; G. Genitalia, ventral view.’=" httpUri="https://zenodo.org/record/1067522/files/figure.png" pageId="4">Fig. 3 D</figureCitation>
) yellowish brown; 3rd segment nearly oval with small, shallow, sensory organ on apical 1/3; lengths of segments 15- 25- 30- 23- 38; palpal ratio 1.3-1.9 (1.6, n=2).
</paragraph>
<paragraph id="8B9EE12F3F02FFA7FF1FFA6CFC1FF887" blockId="4.[151,1436,1258,2012]" pageId="4">
Thorax. Brown; scutum without definite pattern on slide mounted specimens; scutellum yellowish laterally, with 4 bristles. Legs (
<figureCitation id="131AFDAA3F02FFA7FE55FA40FE7CF9CE" box="[397,489,1513,1538]" captionStart="FIGURE 3" captionStartId="5.[151,250,1905,1927]" captionTargetBox="[203,1401,398,1871]" captionTargetId="figure@5.[181,1425,373,1895]" captionTargetPageId="5" captionText="FIGURE 3. Monohelea cornuta Felippe-Bauer &amp; Trindade sp. nov., male. A. Wing; B. Thorax showing halter; C. Head, anterior view; D. Palpus, lateral view; E. Abdomen, ventral view; F. Legs (left to right) of fore, mid-, hindlegs, lateral view; G. Genitalia, ventral view.’=" httpUri="https://zenodo.org/record/1067522/files/figure.png" pageId="4">Fig. 3 F</figureCitation>
) yellow; coxae and trochanters brown; fore femur with basal and mesal pale brown bands, mid femur with basal pale brown band, hind femur with infuscated basal band, two brown stripes in middle and a subapical ventral brown spot; apices of tibiae brown; hind tibia with additional dark irregular brown band on 1/2 subbasal; hind tibial comb with 6 bristles. Tarsi: pale; fore-, hind tarsomere 1 with one basal and one apical spine; midtarsomere 1 with 2 basal, 2 apical spines; apical spines of fore-, mid-, hind tarsomeres 2–4: 1-1-1 or 1-2- 2, 2-2-2, 1-1-1 or 1-1-2, basal spines absent; fore-, mid-, hind tarsal ratios 2.08–2.23 (2.17, n=4), 2.39–2.56 (2.47, n=4), 2.00 (n=3); claws small, paired, equal-sized, 0.3 to 0.5X as long as 5th tarsomeres. Wing (
<figureCitation id="131AFDAA3F02FFA7FAFBF96DFA1BF911" box="[1315,1422,1732,1757]" captionStart="FIGURE 3" captionStartId="5.[151,250,1905,1927]" captionTargetBox="[203,1401,398,1871]" captionTargetId="figure@5.[181,1425,373,1895]" captionTargetPageId="5" captionText="FIGURE 3. Monohelea cornuta Felippe-Bauer &amp; Trindade sp. nov., male. A. Wing; B. Thorax showing halter; C. Head, anterior view; D. Palpus, lateral view; E. Abdomen, ventral view; F. Legs (left to right) of fore, mid-, hindlegs, lateral view; G. Genitalia, ventral view.’=" httpUri="https://zenodo.org/record/1067522/files/figure.png" pageId="4">Fig. 3 A</figureCitation>
): macrotrichia restricted to vein costa; microtrichia absent; 2nd radial cell nearly 2.1X longer than 1st; wing length 0.78–0.85 (0.82, n= 5) mm, width 0.28–0.31 (0.30, n=5) mm; costal ratio 0.73–0.77 (0.75, n=5). Halter stem pale; knob pale brown in distal 1/2 a pale brown spot laterally (
<figureCitation id="131AFDAA3F02FFA7FCC6F89AFCE9F887" box="[798,892,1843,1868]" captionStart="FIGURE 3" captionStartId="5.[151,250,1905,1927]" captionTargetBox="[203,1401,398,1871]" captionTargetId="figure@5.[181,1425,373,1895]" captionTargetPageId="5" captionText="FIGURE 3. Monohelea cornuta Felippe-Bauer &amp; Trindade sp. nov., male. A. Wing; B. Thorax showing halter; C. Head, anterior view; D. Palpus, lateral view; E. Abdomen, ventral view; F. Legs (left to right) of fore, mid-, hindlegs, lateral view; G. Genitalia, ventral view.’=" httpUri="https://zenodo.org/record/1067522/files/figure.png" pageId="4">Fig. 3 B</figureCitation>
).
</paragraph>
<paragraph id="8B9EE12F3F02FFA6FF1FF8FEFB4BFE8C" blockId="4.[151,1436,1258,2012]" lastBlockId="5.[151,1437,151,320]" lastPageId="5" pageId="4">
Abdomen. Yellowish brown (
<figureCitation id="131AFDAA3F02FFA7FDC3F8FEFDEBF8A3" box="[539,638,1879,1904]" captionStart="FIGURE 3" captionStartId="5.[151,250,1905,1927]" captionTargetBox="[203,1401,398,1871]" captionTargetId="figure@5.[181,1425,373,1895]" captionTargetPageId="5" captionText="FIGURE 3. Monohelea cornuta Felippe-Bauer &amp; Trindade sp. nov., male. A. Wing; B. Thorax showing halter; C. Head, anterior view; D. Palpus, lateral view; E. Abdomen, ventral view; F. Legs (left to right) of fore, mid-, hindlegs, lateral view; G. Genitalia, ventral view.’=" httpUri="https://zenodo.org/record/1067522/files/figure.png" pageId="4">Fig. 3 E</figureCitation>
). Genitalia (
<figureCitation id="131AFDAA3F02FFA7FCD7F8FEFC15F8A3" box="[783,896,1879,1904]" captionStart="FIGURE 3" captionStartId="5.[151,250,1905,1927]" captionTargetBox="[203,1401,398,1871]" captionTargetId="figure@5.[181,1425,373,1895]" captionTargetPageId="5" captionText="FIGURE 3. Monohelea cornuta Felippe-Bauer &amp; Trindade sp. nov., male. A. Wing; B. Thorax showing halter; C. Head, anterior view; D. Palpus, lateral view; E. Abdomen, ventral view; F. Legs (left to right) of fore, mid-, hindlegs, lateral view; G. Genitalia, ventral view.’=" httpUri="https://zenodo.org/record/1067522/files/figure.png" pageId="4">Figs. 3 G</figureCitation>
; 9 C–D): sternite IX spiculate except on basal portion, posterior margin with a short, convex, median lobe with 4 long setae; tergite IX tapering with a pair of apicolateral processes. Gonocoxite moderately stout, nearly 
<date id="FF9FC7EF3F02FFA7FCEFF836FCCCF87B" box="[823,857,1951,1975]" pageId="4">2X</date>
longer than basal width; gonostylus curved distally, 0.62–0.70 (0.66, n= 5)X as long as gonocoxite, moderately pilose on basal 3/4. Parameres (
<figureCitation id="131AFDAA3F02FFA7FB60F86AFA8EF810" box="[1208,1307,1987,2012]" captionStart="FIGURE9" captionStartId="14.[151,250,1928,1950]" captionTargetBox="[278,1266,201,1863]" captionTargetId="figure@14.[207,1344,189,1889]" captionTargetPageId="14" captionText="FIGURE9. Parameres and aedeagus. A–B.Monohelea barbara Felippe-Bauer&amp; Trindade sp. nov.; C–D.Monohelea cornuta Felippe-Bauer &amp; Trindade sp. nov.; E–F. Monohelea ema Felippe-Bauer &amp; Trindade sp. nov.; G–H. Monohelea forceps Felippe-Bauer &amp; Trindade sp. nov." httpUri="https://zenodo.org/record/1067534/files/figure.png" pageId="4">Fig. 9 C</figureCitation>
) 1.19–1.78 (1.44, n= 5)X as long as aedeagus, fused at base; each with an external curved, strongly sclerotized, mesal hornshaped process, 0.48–0.56 (0.52, n=5)X as long as distal portion of the paramere, that tapering to blunt apex, laterally truncated and overlapped. Aedeagus (
<figureCitation id="131AFDAA3F03FFA6FD6BFF76FC8CFF34" box="[691,793,223,248]" captionStart="FIGURE9" captionStartId="14.[151,250,1928,1950]" captionTargetBox="[278,1266,201,1863]" captionTargetId="figure@14.[207,1344,189,1889]" captionTargetPageId="14" captionText="FIGURE9. Parameres and aedeagus. A–B.Monohelea barbara Felippe-Bauer&amp; Trindade sp. nov.; C–D.Monohelea cornuta Felippe-Bauer &amp; Trindade sp. nov.; E–F. Monohelea ema Felippe-Bauer &amp; Trindade sp. nov.; G–H. Monohelea forceps Felippe-Bauer &amp; Trindade sp. nov." httpUri="https://zenodo.org/record/1067534/files/figure.png" pageId="5">Fig. 9 D</figureCitation>
) triangular, composed of 2 pointed ventral plates, with slightly sclerotized dorsal structure, which arises in the middle way to aedeagus base and produced beyond the apices of ventral plates, ending as an apical projection; basal arms slender, not expanded laterally.
</paragraph>
<caption id="DF5EB1A73F03FFA6FF4FF8D8FE0FF808" httpUri="https://zenodo.org/record/1067522/files/figure.png" pageId="5" targetBox="[203,1401,398,1871]" targetPageId="5">
<paragraph id="8B9EE12F3F03FFA6FF4FF8D8FE0FF808" blockId="5.[151,1437,1905,1988]" pageId="5">
<emphasis id="B9553D3D3F03FFA6FF4FF8D8FE83F84A" bold="true" box="[151,278,1905,1927]" pageId="5">FIGURE 3.</emphasis>
<taxonomicName id="4C219AAC3F03FFA6FEF8F8D8FC92F84B" authority="Felippe-Bauer &amp; Trindade" authorityName="Felippe-Bauer &amp; Cardoso &amp; Trindade" authorityYear="2017" box="[288,775,1905,1927]" class="Insecta" family="Ceratopogonidae" genus="Monohelea" kingdom="Animalia" order="Diptera" pageId="5" phylum="Arthropoda" rank="species" species="cornuta" status="sp. nov.">
<emphasis id="B9553D3D3F03FFA6FEF8F8D8FE7FF84B" box="[288,490,1905,1927]" italics="true" pageId="5">Monohelea cornuta</emphasis>
Felippe-Bauer &amp; Trindade
</taxonomicName>
<taxonomicNameLabel id="A26680463F03FFA6FCC9F8D8FCF1F84A" box="[785,868,1905,1926]" pageId="5" rank="species">
<emphasis id="B9553D3D3F03FFA6FCC9F8D8FCF1F84A" bold="true" box="[785,868,1905,1926]" pageId="5">sp. nov.</emphasis>
</taxonomicNameLabel>
, male. A. Wing; B. Thorax showing halter; C. Head, anterior view; D. Palpus, lateral view; E. Abdomen, ventral view; F. Legs (left to right) of fore, mid-, hindlegs, lateral view; G. Genitalia, ventral view.’=
</paragraph>
</caption>
<paragraph id="8B9EE12F3F00FFA5FF1FFF3EFE0FFF7C" blockId="6.[151,1437,151,752]" box="[199,410,151,176]" pageId="6">
<emphasis id="B9553D3D3F00FFA5FF1FFF3EFE8BFF7C" bold="true" box="[199,286,151,176]" pageId="6">Female</emphasis>
. Unknown
</paragraph>
</subSubSection>
<subSubSection id="C33BB2A43F00FFA5FF1FFF15FA0EFE8C" pageId="6" type="distribution">
<paragraph id="8B9EE12F3F00FFA5FF1FFF15FDA4FF18" blockId="6.[151,1437,151,752]" box="[199,561,188,213]" pageId="6">
<emphasis id="B9553D3D3F00FFA5FF1FFF15FECCFF19" bold="true" box="[199,345,188,213]" pageId="6">Distribution</emphasis>
. 
<collectingCountry id="F336A1BF3F00FFA5FEBFFF15FE3BFF19" box="[359,430,188,213]" name="Brazil" pageId="6">Brazil</collectingCountry>
, 
<collectingRegion id="49E52FCD3F00FFA5FE61FF15FE7EFF18" box="[441,491,188,212]" country="Brazil" name="Para" pageId="6">Pará</collectingRegion>
State.
</paragraph>
<paragraph id="8B9EE12F3F00FFA5FF1FFF76FA0EFE8C" blockId="6.[151,1437,151,752]" pageId="6">
<materialsCitation id="3B49EB723F00FFA5FF1FFF76FB2AFED0" ID-GBIF-Occurrence="1702562774" collectingDate="2007-06-20" collectingDateMax="2007-06-21" collectingDateMin="2007-06-20" collectionCode="CDC, MPEG" collectorName="Trindade R. L. &amp; Guimaraes, D." country="Brazil" latitude="-1.4322221" location="Vila de Curupaiti" longLatPrecision="21" longitude="-46.47111" municipality="Viseu" pageId="6" specimenCount="2" stateProvince="Para" typeStatus="holotype">
<typeStatus id="549A5F8D3F00FFA5FF1FFF76FE99FF34" box="[199,268,223,248]" pageId="6">
<emphasis id="B9553D3D3F00FFA5FF1FFF76FE99FF34" bold="true" box="[199,268,223,248]" pageId="6">Types</emphasis>
</typeStatus>
. 
<typeStatus id="549A5F8D3F00FFA5FEC6FF76FE13FF34" box="[286,390,223,248]" pageId="6" type="holotype">Holotype</typeStatus>
male, on microscope slide labeled “ 
<location id="8EFEB7F43F00FFA5FCE9FF76FC68FF34" LSID="urn:lsid:plazi:treatment:038850393F02FFA5FF4FFBD6FC2FFD3C:8EFEB7F43F00FFA5FCE9FF76FC68FF34" box="[817,1021,223,248]" country="Brazil" latitude="-1.4322221" longLatPrecision="21" longitude="-46.47111" municipality="Viseu" name="Vila de Curupaiti" pageId="6" stateProvince="Para">Vila de Curupaiti</location>
” (
<geoCoordinate id="EE1587E83F00FFA5FBC3FF76FB35FF34" box="[1051,1184,223,248]" direction="south" orientation="latitude" pageId="6" precision="15" value="-1.4322221">01°25’56”S</geoCoordinate>
<geoCoordinate id="EE1587E83F00FFA5FB73FF76FAABFF34" box="[1195,1342,223,248]" direction="west" orientation="longitude" pageId="6" precision="15" value="-46.47111">46°28’16”W</geoCoordinate>
), 
<collectingMunicipality id="6BFA7B553F00FFA5FA8DFF76FA0DFF34" box="[1365,1432,223,248]" pageId="6">Viseu</collectingMunicipality>
, 
<collectingRegion id="49E52FCD3F00FFA5FF4FFEADFF59FED0" box="[151,204,260,284]" country="Brazil" name="Para" pageId="6">Pará</collectingRegion>
, 
<collectingCountry id="F336A1BF3F00FFA5FF0FFEADFEA8FED0" box="[215,317,260,284]" name="Brazil" pageId="6">BRASIL</collectingCountry>
, 
<collectingDate id="EFDB3E073F00FFA5FE90FEADFE60FED0" box="[328,501,260,284]" pageId="6" value="2007-06-20" valueMax="2007-06-21" valueMin="2007-06-20">
<date id="FF9FC7EF3F00FFA5FE90FEADFE60FED0" box="[328,501,260,284]" pageId="6" value="2007-06-20" valueMax="2007-06-21" valueMin="2007-06-20">20–21.VI.2007</date>
</collectingDate>
, 
<collectionCode id="ED3079EA3F00FFA5FDD8FEADFDAFFED0" box="[512,570,260,284]" country="China" httpUri="http://biocol.org/urn:lsid:biocol.org:col:12786" name="Changdu Institute for Drug Control" pageId="6">CDC</collectionCode>
light trap, 
<collectorName id="26D484F93F00FFA5FD6DFEADFCC7FED0" box="[693,850,260,285]" pageId="6">Trindade R.L.</collectorName>
&amp; 
<collectorName id="26D484F93F00FFA5FCAEFEADFB8CFED0" box="[886,1049,260,285]" pageId="6">Guimarães, D.</collectorName>
cols. (
<collectionCode id="ED3079EA3F00FFA5FBBBFEADFB22FED0" box="[1123,1207,260,284]" country="Brazil" httpUri="http://grbio.org/cool/90vz-swjg" name="Museu Paraense Emilio Goeldi" pageId="6">MPEG</collectionCode>
)
</materialsCitation>
. Paratypes 4 males: 
<materialsCitation id="3B49EB723F00FFA5FF4FFE81FD42FE8C" ID-GBIF-Occurrence="1702562779" box="[151,727,295,320]" collectingDate="2007-06-20" collectingDateMax="2007-06-21" collectingDateMin="2007-06-20" collectionCode="CCER, MPEG" collectorName="Trindade R. L. &amp; Guimaraes, D." country="Brazil" latitude="-1.4322221" location="Vila de Curupaiti" longLatPrecision="21" longitude="-46.47111" municipality="Viseu" pageId="6" specimenCount="3" specimenCount-male="3" stateProvince="Para" typeStatus="holotype">
<specimenCount id="9D272AA63F00FFA5FF4FFE81FF78FE8C" box="[151,237,295,320]" pageId="6" type="male">3 males</specimenCount>
same data as holotype (2 
<collectionCode id="ED3079EA3F00FFA5FDC8FE81FDC8FE8C" box="[528,605,296,320]" pageId="6">CCER</collectionCode>
; 1 
<collectionCode id="ED3079EA3F00FFA5FDA5FE81FD5AFE8C" box="[637,719,296,320]" country="Brazil" httpUri="http://grbio.org/cool/90vz-swjg" name="Museu Paraense Emilio Goeldi" pageId="6">MPEG</collectionCode>
)
</materialsCitation>
, 
<materialsCitation id="3B49EB723F00FFA5FD3AFE81FA02FE8C" ID-GBIF-Occurrence="1702562776" box="[738,1431,295,320]" collectingDate="2007-06-22" collectingDateMax="2007-06-23" collectingDateMin="2007-06-22" collectionCode="MPEG" collectorName="Trindade R. L. &amp; Guimaraes, D." country="Brazil" latitude="-1.4322221" location="Vila de Curupaiti" longLatPrecision="21" longitude="-46.47111" municipality="Viseu" pageId="6" specimenCount="1" specimenCount-male="1" stateProvince="Para" typeStatus="holotype">
<specimenCount id="9D272AA63F00FFA5FD3AFE81FCB8FE8C" box="[738,813,295,320]" pageId="6" type="male">1 male</specimenCount>
same data as holotype except 
<collectingDate id="EFDB3E073F00FFA5FB5AFE81FAB9FE8C" box="[1154,1324,296,320]" pageId="6" value="2007-06-22" valueMax="2007-06-23" valueMin="2007-06-22">
<date id="FF9FC7EF3F00FFA5FB5AFE81FAB9FE8C" box="[1154,1324,296,320]" pageId="6" value="2007-06-22" valueMax="2007-06-23" valueMin="2007-06-22">22–23.VI.2007</date>
</collectingDate>
(
<collectionCode id="ED3079EA3F00FFA5FAE3FE81FA05FE8C" box="[1339,1424,296,320]" country="Brazil" httpUri="http://grbio.org/cool/90vz-swjg" name="Museu Paraense Emilio Goeldi" pageId="6">MPEG</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C33BB2A43F00FFA5FF1FFEE5FC2FFD3C" pageId="6" type="etymology">
<paragraph id="8B9EE12F3F00FFA5FF1FFEE5FD8BFE44" blockId="6.[151,1437,151,752]" pageId="6">
<emphasis id="B9553D3D3F00FFA5FF1FFEE5FED3FEA9" bold="true" box="[199,326,332,357]" pageId="6">Etymology</emphasis>
. The specific epithet is from the Latin: 
<emphasis id="B9553D3D3F00FFA5FCD8FEE4FCD4FEA8" box="[768,833,333,356]" italics="true" pageId="6">cornu</emphasis>
=horn, referring the mesal horn-shaped process on the parameres of males of this species.
</paragraph>
<paragraph id="8B9EE12F3F00FFA5FF1FFE3DFC2FFD3C" blockId="6.[151,1437,151,752]" pageId="6">
<emphasis id="B9553D3D3F00FFA5FF1FFE3DFE4FFE61" bold="true" box="[199,474,404,429]" pageId="6">Taxonomic discussion.</emphasis>
<taxonomicName id="4C219AAC3F00FFA5FE31FE3DFD58FE60" authorityName="Felippe-Bauer &amp; Cardoso &amp; Trindade" authorityYear="2017" box="[489,717,404,428]" class="Insecta" family="Ceratopogonidae" genus="Monohelea" kingdom="Animalia" order="Diptera" pageId="6" phylum="Arthropoda" rank="species" species="cornuta">
<emphasis id="B9553D3D3F00FFA5FE31FE3DFD58FE60" box="[489,717,404,428]" italics="true" pageId="6">Monohelea cornuta</emphasis>
</taxonomicName>
most closely resembles 
<taxonomicName id="4C219AAC3F00FFA5FBD8FE3CFAFFFE60" authority="Tavares &amp; Souza" authorityName="Tavares &amp; Souza" box="[1024,1386,404,428]" class="Insecta" family="Ceratopogonidae" genus="Monohelea" kingdom="Animalia" order="Diptera" pageId="6" phylum="Arthropoda" rank="species" species="aguirrei">
<emphasis id="B9553D3D3F00FFA5FBD8FE3CFB1DFE60" box="[1024,1160,405,428]" italics="true" pageId="6">M. aguirrei</emphasis>
Tavares &amp; Souza
</taxonomicName>
, 
<taxonomicName id="4C219AAC3F00FFA5FAA5FE3CFE77FE1C" authority="Tavares &amp; Souza" authorityName="Tavares &amp; Souza" class="Insecta" family="Ceratopogonidae" genus="Monohelea" kingdom="Animalia" order="Diptera" pageId="6" phylum="Arthropoda" rank="species" species="archibaldoi">
<emphasis id="B9553D3D3F00FFA5FAA5FE3CFE8CFE1C" italics="true" pageId="6">M. archibaldoi</emphasis>
Tavares &amp; Souza
</taxonomicName>
, 
<taxonomicName id="4C219AAC3F00FFA5FE36FE10FDD7FE1C" authority="Felippe-Bauer &amp; Cardoso &amp; Trindade, 2017" authorityName="Felippe-Bauer &amp; Cardoso &amp; Trindade" authorityYear="2017" box="[494,578,441,464]" class="Insecta" family="Ceratopogonidae" genus="Monohelea" kingdom="Animalia" order="Diptera" pageId="6" phylum="Arthropoda" rank="species" species="ema" status="sp. nov.">
<emphasis id="B9553D3D3F00FFA5FE36FE10FDD7FE1C" box="[494,578,441,464]" italics="true" pageId="6">M. ema</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26680463F00FFA5FD91FE1EFD36FE1C" box="[585,675,439,464]" pageId="6" rank="species">
<emphasis id="B9553D3D3F00FFA5FD91FE1EFD36FE1C" bold="true" box="[585,675,439,464]" pageId="6">sp. nov.</emphasis>
</taxonomicNameLabel>
, 
<taxonomicName id="4C219AAC3F00FFA5FD6AFE10FC5EFE1C" authority="Kieffer" authorityName="Kieffer" authorityYear="1917" box="[690,971,439,464]" class="Insecta" family="Ceratopogonidae" genus="Monohelea" kingdom="Animalia" order="Diptera" pageId="6" phylum="Arthropoda" rank="species" species="hieroglyphica">
<emphasis id="B9553D3D3F00FFA5FD6AFE10FCE7FE1C" box="[690,882,440,464]" italics="true" pageId="6">M. hieroglyphica</emphasis>
Kieffer
</taxonomicName>
, 
<taxonomicName id="4C219AAC3F00FFA5FC0FFE10FA8AFE1C" authority="(Coquillett)" authorityName="" authorityYear="1905" baseAuthorityName="Coquillett" baseAuthorityYear="1905" box="[983,1311,439,464]" class="Insecta" family="Ceratopogonidae" genus="Monohelea" kingdom="Animalia" order="Diptera" pageId="6" phylum="Arthropoda" rank="species" species="maculipennis">
<emphasis id="B9553D3D3F00FFA5FC0FFE10FB04FE1C" box="[983,1169,440,464]" italics="true" pageId="6">M. maculipennis</emphasis>
(Coquillett)
</taxonomicName>
, 
<taxonomicName id="4C219AAC3F00FFA5FAF2FE10FEA1FE39" authority="Lane &amp; Wirth" authorityName="Lane &amp; Wirth" authorityYear="1964" class="Insecta" family="Ceratopogonidae" genus="Monohelea" kingdom="Animalia" order="Diptera" pageId="6" phylum="Arthropoda" rank="species" species="poncai">
<emphasis id="B9553D3D3F00FFA5FAF2FE10FA09FE03" box="[1322,1436,440,464]" italics="true" pageId="6">M. poncai</emphasis>
Lane &amp; Wirth
</taxonomicName>
and 
<taxonomicName id="4C219AAC3F00FFA5FEB5FE74FCDBFE39" authority="Felippe-Bauer &amp; Spinelli" authorityName="Felippe-Bauer &amp; Spinelli" authorityYear="1998" box="[365,846,476,501]" class="Insecta" family="Ceratopogonidae" genus="Monohelea" kingdom="Animalia" order="Diptera" pageId="6" phylum="Arthropoda" rank="species" species="uruguayensis">
<emphasis id="B9553D3D3F00FFA5FEB5FE74FDBCFE38" box="[365,553,477,500]" italics="true" pageId="6">M. uruguayensis</emphasis>
Felippe-Bauer &amp; Spinelli
</taxonomicName>
by the presence of a sclerotized mesal hook-shaped process. The new species can be easily separated by the infuscated base of the hind femur (hind femur with brown base in 
<taxonomicName id="4C219AAC3F00FFA5FF35FD8CFEFAFDF0" box="[237,367,549,572]" class="Insecta" family="Ceratopogonidae" genus="Monohelea" kingdom="Animalia" order="Diptera" pageId="6" phylum="Arthropoda" rank="species" species="aguirrei">
<emphasis id="B9553D3D3F00FFA5FF35FD8CFEFAFDF0" box="[237,367,549,572]" italics="true" pageId="6">M. aguirrei</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C219AAC3F00FFA5FEA5FD8CFE47FDF0" authorityName="Felippe-Bauer &amp; Cardoso &amp; Trindade" authorityYear="2017" box="[381,466,549,572]" class="Insecta" family="Ceratopogonidae" genus="Monohelea" kingdom="Animalia" order="Diptera" pageId="6" phylum="Arthropoda" rank="species" species="ema">
<emphasis id="B9553D3D3F00FFA5FEA5FD8CFE47FDF0" box="[381,466,549,572]" italics="true" pageId="6">M. ema</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C219AAC3F00FFA5FE38FD8CFDC7FDF0" authorityName="Lane &amp; Wirth" authorityYear="1964" box="[480,594,549,572]" class="Insecta" family="Ceratopogonidae" genus="Monohelea" kingdom="Animalia" order="Diptera" pageId="6" phylum="Arthropoda" rank="species" species="poncai">
<emphasis id="B9553D3D3F00FFA5FE38FD8CFDC7FDF0" box="[480,594,549,572]" italics="true" pageId="6">M. poncai</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C219AAC3F00FFA5FD52FD8CFCD0FDF0" authorityName="Felippe-Bauer &amp; Spinelli" authorityYear="1998" box="[650,837,549,572]" class="Insecta" family="Ceratopogonidae" genus="Monohelea" kingdom="Animalia" order="Diptera" pageId="6" phylum="Arthropoda" rank="species" species="uruguayensis">
<emphasis id="B9553D3D3F00FFA5FD52FD8CFCD0FDF0" box="[650,837,549,572]" italics="true" pageId="6">M. uruguayensis</emphasis>
</taxonomicName>
; hind femur with pale base in 
<taxonomicName id="4C219AAC3F00FFA5FB46FD8CFAD0FDF0" box="[1182,1349,548,572]" class="Insecta" family="Ceratopogonidae" genus="Monohelea" kingdom="Animalia" order="Diptera" pageId="6" phylum="Arthropoda" rank="species" species="archibaldoi">
<emphasis id="B9553D3D3F00FFA5FB46FD8CFAD0FDF0" box="[1182,1349,548,572]" italics="true" pageId="6">M. archibaldoi</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C219AAC3F00FFA5FAA5FD8CFEA7FDAC" authorityName="Kieffer" authorityYear="1917" class="Insecta" family="Ceratopogonidae" genus="Monohelea" kingdom="Animalia" order="Diptera" pageId="6" phylum="Arthropoda" rank="species" species="hieroglyphica">
<emphasis id="B9553D3D3F00FFA5FAA5FD8CFEA7FDAC" italics="true" pageId="6">M. hieroglyphica</emphasis>
</taxonomicName>
), by the mid leg without distinct mesal brown marks as in 
<taxonomicName id="4C219AAC3F00FFA5FC19FDE0FBF2FDAC" box="[961,1127,584,608]" class="Insecta" family="Ceratopogonidae" genus="Monohelea" kingdom="Animalia" order="Diptera" pageId="6" phylum="Arthropoda" rank="species" species="archibaldoi">
<emphasis id="B9553D3D3F00FFA5FC19FDE0FBF2FDAC" box="[961,1127,584,608]" italics="true" pageId="6">M. archibaldoi</emphasis>
</taxonomicName>
(mid leg with mesal brown bands and spots in the other species), by the distal portion of the paramere, tapering to blunt apex, laterally truncated and overlapped (distal portion of the paramere progressively narrowing distally to rounded apex in 
<emphasis id="B9553D3D3F00FFA5FAA5FD38FE3FFD00" italics="true" pageId="6">
<taxonomicName id="4C219AAC3F00FFA5FAA5FD38FF60FD00" class="Insecta" family="Ceratopogonidae" genus="Monohelea" kingdom="Animalia" order="Diptera" pageId="6" phylum="Arthropoda" rank="species" species="aguirrei">M. aguirrei</taxonomicName>
, 
<taxonomicName id="4C219AAC3F00FFA5FED9FD1CFE3FFD00" box="[257,426,692,716]" class="Insecta" family="Ceratopogonidae" genus="Monohelea" kingdom="Animalia" order="Diptera" pageId="6" phylum="Arthropoda" rank="species" species="archibaldoi">M. archibaldoi</taxonomicName>
</emphasis>
, 
<taxonomicName id="4C219AAC3F00FFA5FE61FD1CFD9BFD00" authorityName="Felippe-Bauer &amp; Cardoso &amp; Trindade" authorityYear="2017" box="[441,526,693,716]" class="Insecta" family="Ceratopogonidae" genus="Monohelea" kingdom="Animalia" order="Diptera" pageId="6" phylum="Arthropoda" rank="species" species="ema">
<emphasis id="B9553D3D3F00FFA5FE61FD1CFD9BFD00" box="[441,526,693,716]" italics="true" pageId="6">M. ema</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C219AAC3F00FFA5FDC5FD1CFD4CFD00" authorityName="" authorityYear="1905" baseAuthorityName="Coquillett" baseAuthorityYear="1905" box="[541,729,692,716]" class="Insecta" family="Ceratopogonidae" genus="Monohelea" kingdom="Animalia" order="Diptera" pageId="6" phylum="Arthropoda" rank="species" species="maculipennis">
<emphasis id="B9553D3D3F00FFA5FDC5FD1CFD4CFD00" box="[541,729,692,716]" italics="true" pageId="6">M. maculipennis</emphasis>
</taxonomicName>
; distal portion of the paramere swollen, tapering and abruptly curved internally in 
<taxonomicName id="4C219AAC3F00FFA5FEA1FD70FDADFD3C" authorityName="Kieffer" authorityYear="1917" box="[377,568,728,752]" class="Insecta" family="Ceratopogonidae" genus="Monohelea" kingdom="Animalia" order="Diptera" pageId="6" phylum="Arthropoda" rank="species" species="hieroglyphica">
<emphasis id="B9553D3D3F00FFA5FEA1FD70FDADFD3C" box="[377,568,728,752]" italics="true" pageId="6">M. hieroglyphica</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C219AAC3F00FFA5FD9FFD70FD2DFD23" authorityName="Lane &amp; Wirth" authorityYear="1964" box="[583,696,728,752]" class="Insecta" family="Ceratopogonidae" genus="Monohelea" kingdom="Animalia" order="Diptera" pageId="6" phylum="Arthropoda" rank="species" species="poncai">
<emphasis id="B9553D3D3F00FFA5FD9FFD70FD2DFD23" box="[583,696,728,752]" italics="true" pageId="6">M. poncai</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C219AAC3F00FFA5FD37FD70FC3CFD23" authorityName="Felippe-Bauer &amp; Spinelli" authorityYear="1998" box="[751,937,728,752]" class="Insecta" family="Ceratopogonidae" genus="Monohelea" kingdom="Animalia" order="Diptera" pageId="6" phylum="Arthropoda" rank="species" species="uruguayensis">
<emphasis id="B9553D3D3F00FFA5FD37FD70FC3CFD23" box="[751,937,728,752]" italics="true" pageId="6">M. uruguayensis</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
</treatment>
</document>