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<document id="22EC0E75D033C4193B00C662020C433D" ID-DOI="10.11646/zootaxa.4040.3.5" ID-GBIF-Dataset="163528a9-7792-480c-83ed-4fecf33f13ff" ID-ISSN="1175-5326" ID-Zenodo-Dep="241584" ID-ZooBank="D6344A6C-DF7A-490B-9EDB-20456618A049" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461335752160" checkinUser="plazi" docAuthor="Yin, Zi-Wei &amp; Li, Li-Zhen" docDate="2015" docId="0388136F690B896FFF03C82364469FE7" docLanguage="en" docName="zt04040p344.pdf" docOrigin="Zootaxa 4040 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pselaphodes spinosus Champion 1925" docType="treatment" docVersion="9" lastPageNumber="340" masterDocId="FFB16B1769028966FF94CA2361209836" masterDocTitle="Review of some species in the genera Pselaphodes Westwood and Labomimus Sharp (Coleoptera: Staphylinidae: Pselaphinae)" masterLastPageNumber="344" masterPageNumber="331" pageNumber="340" updateTime="1698619188225" updateUser="plazi">
<mods:mods id="3F5866C44BB4714C2CDB533500C71FE1" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="4DE0F921914FB05FE3E494D297EE47D1">
<mods:title id="1F2D9C6225DFB415A54734939F753AB8">Review of some species in the genera Pselaphodes Westwood and Labomimus Sharp (Coleoptera: Staphylinidae: Pselaphinae)</mods:title>
</mods:titleInfo>
<mods:name id="0875CAC9590FBB57AF99B5D876A468D4" type="personal">
<mods:role id="781F488185B89F05E40E98E0E4D214AE">
<mods:roleTerm id="E0B07A55918AD843055FB949C09CA8DE">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="B18CAC7D5EAE98CD3BEA241A86C9C3D6">Yin, Zi-Wei</mods:namePart>
</mods:name>
<mods:name id="B62EC5D63128CD7BD86B05BC0441E320" type="personal">
<mods:role id="DFD24E33C6EAF76B6DB61EB287AFAF81">
<mods:roleTerm id="50AB209E06D724872166841748A2CBF8">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="7B159CBAB82DF451825F4F0DD34FB18C">Li, Li-Zhen</mods:namePart>
</mods:name>
<mods:typeOfResource id="4A6EAC90A914822D4B9544BF51499249">text</mods:typeOfResource>
<mods:relatedItem id="967C7E890845429838826EA078CA75FF" type="host">
<mods:titleInfo id="959465537B0B5D00782E10ECC567C391">
<mods:title id="9CDCE9F9A613D0305ABF6BBD5E4C31E5">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="4161BE6A74C6BB798CEB391FB752ABCC">
<mods:date id="D7E1E93068297B1B93C7E8F98B960A6F">2015</mods:date>
<mods:detail id="6479352B9B78E28E42CD31C14959AA14" type="volume">
<mods:number id="46A1AA6DF9FAC3A79512AB3C6859C555">4040</mods:number>
</mods:detail>
<mods:detail id="511597A656CF1DCFAA0CCEDF6CE83309" type="issue">
<mods:number id="88A97DB9B49BC4AEA552E7718B699B85">3</mods:number>
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<mods:extent id="7822016010690A0CD8483014F90679FB" unit="page">
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<mods:classification id="A08CFB779A79A3E30C2B891AE456A53C">journal article</mods:classification>
<mods:identifier id="567509CF999AD8481F9E5731353A56DD" type="DOI">10.11646/zootaxa.4040.3.5</mods:identifier>
<mods:identifier id="5D9229A1210EBEF4EC1412B56BD3863B" type="GBIF-Dataset">163528a9-7792-480c-83ed-4fecf33f13ff</mods:identifier>
<mods:identifier id="BAD90C3D320240FBD146D6FF24465214" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="F1787A65F9B6F6DCB061A76FC10782A1" type="Zenodo-Dep">241584</mods:identifier>
<mods:identifier id="12E3601EEFB49EEFCECCC771DA12C785" type="ZooBank">D6344A6C-DF7A-490B-9EDB-20456618A049</mods:identifier>
</mods:mods>
<treatment id="0388136F690B896FFF03C82364469FE7" ID-DOI="http://doi.org/10.5281/zenodo.6107980" ID-GBIF-Taxon="119644488" ID-Zenodo-Dep="6107980" LSID="urn:lsid:plazi:treatment:0388136F690B896FFF03C82364469FE7" httpUri="http://treatment.plazi.org/id/0388136F690B896FFF03C82364469FE7" lastPageNumber="340" pageId="9" pageNumber="340">
<subSubSection id="C33BF1F2690B896FFF03C82363AE9A2C" box="[151,654,511,538]" pageId="9" pageNumber="340" type="nomenclature">
<paragraph id="8B9EA279690B896FFF03C82363AE9A2C" blockId="9.[151,654,511,571]" box="[151,654,511,538]" pageId="9" pageNumber="340">
<heading id="D0D61515690B896FFF03C82363AE9A2C" bold="true" box="[151,654,511,538]" fontSize="11" level="1" pageId="9" pageNumber="340" reason="1">
<emphasis id="B9557E6B690B896FFF03C82363AE9A2C" bold="true" box="[151,654,511,538]" pageId="9" pageNumber="340">
6.
<taxonomicName id="4C21D9FA690B896FFF20CBDC63AE9A2C" ID-CoL="4NF8J" authority="Champion, 1925" authorityName="Champion" authorityYear="1925" box="[180,654,511,538]" class="Insecta" family="Staphylinidae" genus="Pselaphodes" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="340" phylum="Arthropoda" rank="species" species="spinosus">
<emphasis id="B9557E6B690B896FFF20CBDC60999A2F" bold="true" box="[180,441,511,537]" italics="true" pageId="9" pageNumber="340">Pselaphodes spinosus</emphasis>
<bibRefCitation id="EFB0DF88690B896FFE55C82363AE9A2C" author="Champion" box="[449,654,512,538]" pageId="9" pageNumber="340" refString="Champion, G. C. (1925) Some Indian (and Tibetan) Coleoptera (18). Entomologist's Monthly Magazine, 61, 260 - 273." type="journal article" year="1925">Champion, 1925</bibRefCitation>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C33BF1F2690B896FFF03C801602B9A0D" box="[151,267,546,571]" pageId="9" pageNumber="340" type="description">
<paragraph id="8B9EA279690B896FFF03C801602B9A0D" blockId="9.[151,654,511,571]" box="[151,267,546,571]" pageId="9" pageNumber="340">
(
<figureCitation id="131ABEFC690B896FFF0BC80161C79A0D" box="[159,231,546,571]" captionStart="FIGURE 6" captionStartId="10.[151,250,1505,1527]" captionTargetBox="[154,1420,201,1480]" captionTargetId="figure@10.[151,1435,195,1483]" captionTargetPageId="10" captionText="FIGURE 6. Lectotype of Pselaphodes spinosus, male. A. Dorsal habitus. B. Labels of the lectotype. C. Antennal club. D. Metaventral process. E. Protrochanter and profemur. F. Mesotrochanter and mesofemur. G. Aedeagus, in dorsal view. H. Same, in lateral view. Scales: A = 1.0 mm; C – F = 0.3 mm; G, H = 0.2 mm." httpUri="https://zenodo.org/record/241590/files/figure.png" pageId="9" pageNumber="340">Figs 6</figureCitation>
,
<figureCitation id="131ABEFC690B896FFF67C80060229A0D" box="[243,258,547,571]" captionStart="FIGURE 7" captionStartId="11.[151,250,1584,1606]" captionTargetBox="[159,1425,592,1559]" captionTargetId="figure@11.[151,1435,589,1563]" captionTargetPageId="11" captionText="FIGURE 7. Holotype of Labomimus championi, male. A. Dorsal habitus. B. Labels of the holotype. C, D. Antennal club. E. Metaventral process. F. Protrochanter and profemur. G. Mesotrochanter and mesofemur. H. Aedeagus, in dorsal view. I. Same, in lateral view. Scales: A = 1.0 mm; C – G = 0.3 mm; H. I = 0.2 mm." httpUri="https://zenodo.org/record/241591/files/figure.png" pageId="9" pageNumber="340">7</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C33BF1F2690B896FFF03C849642C9AA9" pageId="9" pageNumber="340" type="reference_group">
<paragraph id="8B9EA279690B896FFF03C849637E9AB6" blockId="9.[151,1292,618,671]" box="[151,606,618,640]" pageId="9" pageNumber="340">
<treatmentCitationGroup id="AB318557690B896FFF03C849637E9AB6" box="[151,606,618,640]" pageId="9" pageNumber="340">
<taxonomicName id="4C21D9FA690B896FFF03C849637A9AB6" ID-CoL="4NF8J" authority="Champion, 1925: 261" authorityName="Champion" authorityPageNumber="261" authorityYear="1925" box="[151,602,618,640]" class="Insecta" family="Staphylinidae" genus="Pselaphodes" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="340" phylum="Arthropoda" rank="species" species="spinosus">
<emphasis id="B9557E6B690B896FFF03C84960549AB6" box="[151,372,618,640]" italics="true" pageId="9" pageNumber="340">Pselaphodes spinosus</emphasis>
<treatmentCitation id="0A808468690B896FFEEFC849637A9AB6" author="Champion" box="[379,602,618,640]" page="261" pageId="9" pageNumber="340" year="1925">
<bibRefCitation id="EFB0DF88690B896FFEEFC84963079AB6" author="Champion" box="[379,551,618,640]" pageId="9" pageNumber="340" refString="Champion, G. C. (1925) Some Indian (and Tibetan) Coleoptera (18). Entomologist's Monthly Magazine, 61, 260 - 273." type="journal article" year="1925">Champion, 1925</bibRefCitation>
: 261
</treatmentCitation>
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B9EA279690B896FFF03C8AA642C9AA9" blockId="9.[151,1292,618,671]" box="[151,1292,649,671]" pageId="9" pageNumber="340">
<treatmentCitationGroup id="AB318557690B896FFF03C8AA642C9AA9" box="[151,1292,649,671]" pageId="9" pageNumber="340">
<taxonomicName id="4C21D9FA690B896FFF03C8AA63409AA9" ID-CoL="6NPNQ" authority="Jeannel, 1960: 454" authorityName="Jeannel" authorityPageNumber="454" authorityYear="1960" box="[151,608,649,671]" class="Insecta" family="Staphylinidae" genus="Pselaphodes" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="340" phylum="Arthropoda" rank="species" species="championi" status="comb. nov.">
<emphasis id="B9557E6B690B896FFF03C8AA60A89AA9" box="[151,392,649,671]" italics="true" pageId="9" pageNumber="340">Pselaphodes championi</emphasis>
(
<treatmentCitation id="0A808468690B896FFE01C8AA63789AA9" author="Jeannel" box="[405,600,649,671]" page="454" pageId="9" pageNumber="340" year="1960">
<bibRefCitation id="EFB0DF88690B896FFE01C8AA63789AA9" author="Jeannel" box="[405,600,649,671]" pageId="9" pageNumber="340" refString="Jeannel, R. (1960) Sur les Pselaphides (Coleoptera) de l'Inde septentrionale. Bulletin of the British Museum (Natural History). Entomology, 9, 403 - 456." type="journal article" year="1960">Jeannel, 1960: 454</bibRefCitation>
</treatmentCitation>
)
</taxonomicName>
,
<taxonomicNameLabel id="A266C310690B896FFDFEC8AA62039AA9" box="[618,803,649,671]" pageId="9" pageNumber="340">
<emphasis id="B9557E6B690B896FFDFEC8AA62039AA9" bold="true" box="[618,803,649,671]" pageId="9" pageNumber="340">new combination</emphasis>
</taxonomicNameLabel>
(transferred from
<taxonomicName id="4C21D9FA690B896FFC49C8AA65739AA9" ID-CoL="8VYNS" box="[989,1107,649,671]" class="Insecta" family="Staphylinidae" genus="Labomimus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="340" phylum="Arthropoda" rank="genus">
<emphasis id="B9557E6B690B896FFC49C8AA65739AA9" box="[989,1107,649,671]" italics="true" pageId="9" pageNumber="340">Labomimus</emphasis>
</taxonomicName>
),
<emphasis id="B9557E6B690B896FFBFDC8AA64279AA8" bold="true" box="[1129,1287,649,670]" pageId="9" pageNumber="340">new synonymy</emphasis>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C33BF1F2690B896FFF03C8EF62189D36" pageId="9" pageNumber="340" type="materials_examined">
<paragraph id="8B9EA279690B896FFF03C8EF65089C33" blockId="9.[151,1437,715,2001]" pageId="9" pageNumber="340">
<emphasis id="B9557E6B690B896FFF03C8EF63E89AD2" bold="true" box="[151,712,715,741]" pageId="9" pageNumber="340">
<typeStatus id="549A1CDB690B896FFF03C8EF61F29AD3" box="[151,210,716,741]" pageId="9" pageNumber="340">Type</typeStatus>
material examined.
<taxonomicName id="4C21D9FA690B896FFE58C8E863E89AD2" box="[460,712,715,740]" class="Insecta" family="Staphylinidae" genus="Pselaphodes" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="340" phylum="Arthropoda" rank="species" species="spinosus">
<emphasis id="B9557E6B690B896FFE58C8E863E89AD2" bold="true" box="[460,712,715,740]" italics="true" pageId="9" pageNumber="340">Pselaphodes spinosus</emphasis>
</taxonomicName>
</emphasis>
:
<typeStatus id="549A1CDB690B896FFD4CC8EF626E9AD3" box="[728,846,716,741]" pageId="9" pageNumber="340" type="lectotype">
<emphasis id="B9557E6B690B896FFD4CC8EF626E9AD3" bold="true" box="[728,846,716,741]" pageId="9" pageNumber="340">Lectotype</emphasis>
</typeStatus>
(fixed by present designation; in
<collectionCode id="ED303ABC690B896FFB40C8EF640E9AD2" box="[1236,1326,716,740]" pageId="9" pageNumber="340">BMNH</collectionCode>
):
<collectingCountry id="F336E2E9690B896FFAD6C8EF64B59AD3" box="[1346,1429,716,741]" name="India" pageId="9" pageNumber="340">
<emphasis id="B9557E6B690B896FFAD6C8EF64B59AD3" bold="true" box="[1346,1429,716,741]" pageId="9" pageNumber="340">INDIA</emphasis>
</collectingCountry>
: ♂, with nine labels (
<figureCitation id="131ABEFC690B896FFEEBC8CC60E59B3E" box="[383,453,751,776]" captionStart="FIGURE 6" captionStartId="10.[151,250,1505,1527]" captionTargetBox="[154,1420,201,1480]" captionTargetId="figure@10.[151,1435,195,1483]" captionTargetPageId="10" captionText="FIGURE 6. Lectotype of Pselaphodes spinosus, male. A. Dorsal habitus. B. Labels of the lectotype. C. Antennal club. D. Metaventral process. E. Protrochanter and profemur. F. Mesotrochanter and mesofemur. G. Aedeagus, in dorsal view. H. Same, in lateral view. Scales: A = 1.0 mm; C – F = 0.3 mm; G, H = 0.2 mm." httpUri="https://zenodo.org/record/241590/files/figure.png" pageId="9" pageNumber="340">Fig. 6</figureCitation>
B): ‘Haldwani Dist., Kumaon,
<collectingCountry id="F336E2E9690B896FFCB5C8CC627E9B3E" box="[801,862,751,776]" name="India" pageId="9" pageNumber="340">India</collectingCountry>
. H. G. Champion (printed) / G. C. Champion. Brit. Mus.
<date id="FF9F84B9690B896FFF4FC93760609B1B" box="[219,320,788,813]" pageId="9" pageNumber="340" value="1925" valueMax="1942">1925–42</date>
. (printed) / ♂ (printed) /
<taxonomicName id="4C21D9FA690B896FFDFBC93763DC9B1B" box="[623,764,788,813]" class="Insecta" family="Staphylinidae" genus="Pselaphodes" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="340" phylum="Arthropoda" rank="genus">
<emphasis id="B9557E6B690B896FFDFBC93763DC9B1B" box="[623,764,788,813]" italics="true" pageId="9" pageNumber="340">Pselaphodes</emphasis>
</taxonomicName>
<taxonomicName id="4C21D9FA690B896FFC9CC93762489B1B" box="[776,872,788,813]" class="Insecta" family="Staphylinidae" genus="Pselaphodes" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="340" phylum="Arthropoda" rank="species" species="spinosus">spinosus</taxonomicName>
C
<typeStatus id="549A1CDB690B896FFC05C93662E19B1B" box="[913,961,789,813]" pageId="9" pageNumber="340">type</typeStatus>
♂ (handwritten) /
<taxonomicName id="4C21D9FA690B896FFB37C93761CF9B66" authority="Champ." authorityName="Champ." class="Insecta" family="Staphylinidae" genus="Pselaphodes" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="340" phylum="Arthropoda" rank="species" species="spinosus">
<emphasis id="B9557E6B690B896FFB37C93764BC9B1A" box="[1187,1436,788,813]" italics="true" pageId="9" pageNumber="340">Pselaphodes spinosus</emphasis>
Champ.
</taxonomicName>
(printed) / E. M. M. 1925. det. G. C. C. /
<typeStatus id="549A1CDB690B896FFD74C91B62389B66" box="[736,792,824,848]" pageId="9" pageNumber="340">Type</typeStatus>
H. T. (round printed label with a red rim) / sand banks Gauniaran Haldwani
<date id="FF9F84B9690B896FFE09C97F63359B42" box="[413,533,860,884]" pageId="9" pageNumber="340" value="1923-04-29">29. 4. 23</date>
/
<typeStatus id="549A1CDB690B896FFDD5C97E63919B43" box="[577,689,861,885]" pageId="9" pageNumber="340" type="lectotype">Lectotype</typeStatus>
♂,
<taxonomicName id="4C21D9FA690B896FFD65C97F655C9B43" authority="Champion" authorityName="Champion" box="[753,1148,860,885]" class="Insecta" family="Staphylinidae" genus="Pselaphodes" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="340" phylum="Arthropoda" rank="species" species="spinosus">
<emphasis id="B9557E6B690B896FFD65C97F62D19B42" box="[753,1009,860,885]" italics="true" pageId="9" pageNumber="340">Pselaphodes spinosus</emphasis>
Champion
</taxonomicName>
, des. Z.-W. Yin, 2015 (handwritten)’.
<taxonomicName id="4C21D9FA690B896FFED3C95C63729BAE" box="[327,594,895,920]" class="Insecta" family="Staphylinidae" genus="Labomimus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="340" phylum="Arthropoda" rank="species" species="championi">
<emphasis id="B9557E6B690B896FFED3C95C63729BAE" bold="true" box="[327,594,895,920]" italics="true" pageId="9" pageNumber="340">Labomimus championi</emphasis>
</taxonomicName>
:
<typeStatus id="549A1CDB690B896FFDF6C95C63EF9BAE" box="[610,719,895,920]" pageId="9" pageNumber="340" type="holotype">
<emphasis id="B9557E6B690B896FFDF6C95C63EF9BAE" bold="true" box="[610,719,895,920]" pageId="9" pageNumber="340">Holotype</emphasis>
</typeStatus>
(fixed by original designation; in
<collectionCode id="ED303ABC690B896FFBC4C9A3658A9BAE" box="[1104,1194,896,920]" pageId="9" pageNumber="340">BMNH</collectionCode>
):
<collectingCountry id="F336E2E9690B896FFB29C95C642F9BAE" box="[1213,1295,895,920]" name="India" pageId="9" pageNumber="340">
<emphasis id="B9557E6B690B896FFB29C95C642F9BAE" bold="true" box="[1213,1295,895,920]" pageId="9" pageNumber="340">INDIA</emphasis>
</collectingCountry>
: ♂, with six labels (
<figureCitation id="131ABEFC690B896FFF7CC987600C9B8A" box="[232,300,932,957]" captionStart="FIGURE 7" captionStartId="11.[151,250,1584,1606]" captionTargetBox="[159,1425,592,1559]" captionTargetId="figure@11.[151,1435,589,1563]" captionTargetPageId="11" captionText="FIGURE 7. Holotype of Labomimus championi, male. A. Dorsal habitus. B. Labels of the holotype. C, D. Antennal club. E. Metaventral process. F. Protrochanter and profemur. G. Mesotrochanter and mesofemur. H. Aedeagus, in dorsal view. I. Same, in lateral view. Scales: A = 1.0 mm; C – G = 0.3 mm; H. I = 0.2 mm." httpUri="https://zenodo.org/record/241591/files/figure.png" pageId="9" pageNumber="340">Fig. 7</figureCitation>
B): ‘Haldwani Dist., Kumaon,
<collectingCountry id="F336E2E9690B896FFD12C98763E39B8B" box="[646,707,932,957]" name="India" pageId="9" pageNumber="340">India</collectingCountry>
. H.G.C. (printed) / G.C. Champion coll. B.M. 1927-409 (printed) /
<collectionCode id="ED303ABC690B896FFF32C9EB61CC9BE9" box="[166,236,968,991]" pageId="9" pageNumber="340">
<typeStatus id="549A1CDB690B896FFF32C9EB61CC9BE9" box="[166,236,968,991]" pageId="9" pageNumber="340">TYPE</typeStatus>
</collectionCode>
(red printed label) /
<typeStatus id="549A1CDB690B896FFE40C9EB632C9BD6" box="[468,524,968,992]" pageId="9" pageNumber="340">Type</typeStatus>
(round printed label with a red rim) /
<taxonomicName id="4C21D9FA690B896FFC22C9E464129BD6" authority="R. Jeannel" authorityName="R. Jeannel" box="[950,1330,967,992]" class="Insecta" family="Staphylinidae" genus="Labomimus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="340" phylum="Arthropoda" rank="species" species="championi">
<emphasis id="B9557E6B690B896FFC22C9E465969BD6" box="[950,1206,967,992]" italics="true" pageId="9" pageNumber="340">Labomimus championi</emphasis>
R. Jeannel
</taxonomicName>
det., 19 /
<typeStatus id="549A1CDB690B896FFF03C9CF61DF9C33" box="[151,255,1004,1029]" pageId="9" pageNumber="340" type="holotype">Holotype</typeStatus>
♂,
<taxonomicName id="4C21D9FA690B896FFEBDC9CF63A69C33" authority="Jeannel" authorityName="Jeannel" box="[297,646,1004,1029]" class="Insecta" family="Staphylinidae" genus="Labomimus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="340" phylum="Arthropoda" rank="species" species="championi">
<emphasis id="B9557E6B690B896FFEBDC9CF63099C33" box="[297,553,1004,1029]" italics="true" pageId="9" pageNumber="340">Labomimus championi</emphasis>
Jeannel
</taxonomicName>
, det. Z.-W. Yin, 2015 (handwritten)’.
</paragraph>
<paragraph id="8B9EA279690B896FFF53CE2C62189D36" blockId="9.[151,1437,715,2001]" pageId="9" pageNumber="340">
<emphasis id="B9557E6B690B896FFF53CE2C60F99C1E" bold="true" box="[199,473,1039,1064]" pageId="9" pageNumber="340">
<typeStatus id="549A1CDB690B896FFF53CE2C601D9C1E" box="[199,317,1039,1064]" pageId="9" pageNumber="340" type="lectotype">Lectotype</typeStatus>
designation.
</emphasis>
<taxonomicName id="4C21D9FA690B896FFE71CE2C63FF9C1E" box="[485,735,1039,1064]" class="Insecta" family="Staphylinidae" genus="Pselaphodes" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="340" phylum="Arthropoda" rank="species" species="spinosus">
<emphasis id="B9557E6B690B896FFE71CE2C63FF9C1E" box="[485,735,1039,1064]" italics="true" pageId="9" pageNumber="340">Pselaphodes spinosus</emphasis>
</taxonomicName>
was described by Champion based on five males and two females without designating a
<typeStatus id="549A1CDB690B896FFE7FCE17636D9C7B" box="[491,589,1076,1101]" pageId="9" pageNumber="340" type="holotype">holotype</typeStatus>
(
<bibRefCitation id="EFB0DF88690B896FFDCECE17626C9C7B" author="Champion" box="[602,844,1076,1101]" pageId="9" pageNumber="340" refString="Champion, G. C. (1925) Some Indian (and Tibetan) Coleoptera (18). Entomologist's Monthly Magazine, 61, 260 - 273." type="journal article" year="1925">Champion 1925: 262</bibRefCitation>
). All of these specimens have a status of
<typeStatus id="549A1CDB690B896FFA87CE16644E9C7B" box="[1299,1390,1077,1101]" pageId="9" pageNumber="340" type="syntype">syntype</typeStatus>
. We did examine a male from the Natural History Museum, London, and here designate this specimen as the
<typeStatus id="549A1CDB690B896FFAA1CE7464BC9C46" box="[1333,1436,1111,1136]" pageId="9" pageNumber="340" type="lectotype">lectotype</typeStatus>
to ensure the stability of nomenclature and provide a unique name-bearing
<typeStatus id="549A1CDB690B896FFB9FCE5E651B9CA3" box="[1035,1083,1149,1173]" pageId="9" pageNumber="340">type</typeStatus>
for the name. The rest of the specimens of the
<typeStatus id="549A1CDB690B896FFECDCE8360A99C8E" box="[345,393,1184,1208]" pageId="9" pageNumber="340">type</typeStatus>
series become
<typeStatus id="549A1CDB690B896FFDA0CEBC63F99C8E" box="[564,729,1183,1208]" pageId="9" pageNumber="340" type="paralectotype">paralectotypes</typeStatus>
.
<taxonomicName id="4C21D9FA690B896FFD70CEBC62C39C8E" box="[740,995,1183,1208]" class="Insecta" family="Staphylinidae" genus="Labomimus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="340" phylum="Arthropoda" rank="species" species="championi">
<emphasis id="B9557E6B690B896FFD70CEBC62C39C8E" box="[740,995,1183,1208]" italics="true" pageId="9" pageNumber="340">Labomimus championi</emphasis>
</taxonomicName>
was described on the basis of one male and one female. Jeannel used the term ‘type’ for the male specimen (
<bibRefCitation id="EFB0DF88690B896FFC2BCEE765B79CEB" author="Jeannel" box="[959,1175,1220,1245]" pageId="9" pageNumber="340" refString="Jeannel, R. (1960) Sur les Pselaphides (Coleoptera) de l'Inde septentrionale. Bulletin of the British Museum (Natural History). Entomology, 9, 403 - 456." type="journal article" year="1960">Jeannel 1960: 453</bibRefCitation>
). According to Article 73.1.1, this male is a
<typeStatus id="549A1CDB690B896FFE11CEC460C79D36" box="[389,487,1255,1280]" pageId="9" pageNumber="340" type="holotype">holotype</typeStatus>
fixed by original designation.
</paragraph>
</subSubSection>
<subSubSection id="C33BF1F2690B896FFF53CF2F62069DA6" pageId="9" pageNumber="340" type="diagnosis">
<paragraph id="8B9EA279690B896FFF53CF2F62069DA6" blockId="9.[151,1437,715,2001]" pageId="9" pageNumber="340">
<emphasis id="B9557E6B690B896FFF53CF2F60889D13" bold="true" box="[199,424,1292,1317]" pageId="9" pageNumber="340">Diagnosis of male.</emphasis>
Reddish brown; body length over
<quantity id="4CD90F9C690B896FFCD1CF2F62829D12" box="[837,930,1292,1317]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="9" pageNumber="340" unit="mm" value="3.5">3.5 mm</quantity>
; postgenae rounded laterally; antennal club composed of enlarged antennomeres IX–XI, antennomeres XI–X modified; pronotum rounded at anterolateral margins; metathorax with long metaventral processes; trochanters and femora of fore and middle legs strongly spinose ventrally; aedeagus with asymmetric median lobe.
</paragraph>
</subSubSection>
<subSubSection id="C33BF1F2690B896FFF53CFBF631F9F9A" pageId="9" pageNumber="340" type="description">
<paragraph id="8B9EA279690B896FFF53CFBF63AF9EE3" blockId="9.[151,1437,715,2001]" pageId="9" pageNumber="340">
<emphasis id="B9557E6B690B896FFF53CFBF60F69D83" bold="true" box="[199,470,1436,1461]" pageId="9" pageNumber="340">Redescription of male.</emphasis>
Habitus as in
<figureCitation id="131ABEFC690B896FFDEACFBF63E59D83" box="[638,709,1436,1461]" captionStart="FIGURE 6" captionStartId="10.[151,250,1505,1527]" captionTargetBox="[154,1420,201,1480]" captionTargetId="figure@10.[151,1435,195,1483]" captionTargetPageId="10" captionText="FIGURE 6. Lectotype of Pselaphodes spinosus, male. A. Dorsal habitus. B. Labels of the lectotype. C. Antennal club. D. Metaventral process. E. Protrochanter and profemur. F. Mesotrochanter and mesofemur. G. Aedeagus, in dorsal view. H. Same, in lateral view. Scales: A = 1.0 mm; C – F = 0.3 mm; G, H = 0.2 mm." httpUri="https://zenodo.org/record/241590/files/figure.png" pageId="9" pageNumber="340">Figs 6</figureCitation>
A, 7A. Body length
<quantity id="4CD90F9C690B896FFC3BCFBF65779D82" box="[943,1111,1436,1461]" metricMagnitude="-3" metricUnit="m" metricValue="3.78" metricValueMax="3.85" metricValueMin="3.71" pageId="9" pageNumber="340" unit="mm" value="3.78" valueMax="3.85" valueMin="3.71">3.71–3.85 mm</quantity>
. Head longer than wide, HL
<quantity id="4CD90F9C690B896FFF03CFE3601D9DEE" box="[151,317,1472,1496]" metricMagnitude="-4" metricUnit="m" metricValue="8.35" metricValueMax="8.4" metricValueMin="8.299999999999999" pageId="9" pageNumber="340" unit="mm" value="0.835" valueMax="0.84" valueMin="0.83">0.83–0.84 mm</quantity>
, HW
<quantity id="4CD90F9C690B896FFE16CF9C63099DEE" box="[386,553,1471,1496]" metricMagnitude="-4" metricUnit="m" metricValue="7.05" metricValueMax="7.199999999999999" metricValueMin="6.8999999999999995" pageId="9" pageNumber="340" unit="mm" value="0.705" valueMax="0.72" valueMin="0.69">0.69–0.72 mm</quantity>
; each eye composed of about 45–50 facets. Antennal club composed of three apical antennomeres, antennomere IX broadened laterally at apical half (
<figureCitation id="131ABEFC690B896FFC4CCFC7653D9DCA" box="[984,1053,1508,1533]" captionStart="FIGURE 7" captionStartId="11.[151,250,1584,1606]" captionTargetBox="[159,1425,592,1559]" captionTargetId="figure@11.[151,1435,589,1563]" captionTargetPageId="11" captionText="FIGURE 7. Holotype of Labomimus championi, male. A. Dorsal habitus. B. Labels of the holotype. C, D. Antennal club. E. Metaventral process. F. Protrochanter and profemur. G. Mesotrochanter and mesofemur. H. Aedeagus, in dorsal view. I. Same, in lateral view. Scales: A = 1.0 mm; C – G = 0.3 mm; H. I = 0.2 mm." httpUri="https://zenodo.org/record/241591/files/figure.png" pageId="9" pageNumber="340">Fig. 7</figureCitation>
D), X strongly concave at middle (
<figureCitation id="131ABEFC690B896FFF0BCC2461CC9E16" box="[159,236,1543,1568]" captionStart="FIGURE 6" captionStartId="10.[151,250,1505,1527]" captionTargetBox="[154,1420,201,1480]" captionTargetId="figure@10.[151,1435,195,1483]" captionTargetPageId="10" captionText="FIGURE 6. Lectotype of Pselaphodes spinosus, male. A. Dorsal habitus. B. Labels of the lectotype. C. Antennal club. D. Metaventral process. E. Protrochanter and profemur. F. Mesotrochanter and mesofemur. G. Aedeagus, in dorsal view. H. Same, in lateral view. Scales: A = 1.0 mm; C – F = 0.3 mm; G, H = 0.2 mm." httpUri="https://zenodo.org/record/241590/files/figure.png" pageId="9" pageNumber="340">Figs 6</figureCitation>
C, 7C). Pronotum longer than wide, PL
<quantity id="4CD90F9C690B896FFD42CC2462A19E16" box="[726,897,1543,1568]" metricMagnitude="-4" metricUnit="m" metricValue="7.7" metricValueMax="7.8" metricValueMin="7.6" pageId="9" pageNumber="340" unit="mm" value="0.77" valueMax="0.78" valueMin="0.76">0.76–0.78 mm</quantity>
, PW
<quantity id="4CD90F9C690B896FFC5DCC2B65559E16" box="[969,1141,1544,1568]" metricMagnitude="-4" metricUnit="m" metricValue="7.25" metricValueMax="7.3" metricValueMin="7.199999999999999" pageId="9" pageNumber="340" unit="mm" value="0.725" valueMax="0.73" valueMin="0.72">0.72–0.73 mm</quantity>
, rounded at anterolateral margins. Elytra wider than long, EL
<quantity id="4CD90F9C690B896FFDF1CC0F62339E72" box="[613,787,1580,1605]" metricMagnitude="-3" metricUnit="m" metricValue="1.05" metricValueMax="1.06" metricValueMin="1.04" pageId="9" pageNumber="340" unit="mm" value="1.05" valueMax="1.06" valueMin="1.04">1.04–1.06 mm</quantity>
, EW
<quantity id="4CD90F9C690B896FFCF6CC0F65319E72" box="[866,1041,1580,1605]" metricMagnitude="-3" metricUnit="m" metricValue="1.335" metricValueMax="1.36" metricValueMin="1.31" pageId="9" pageNumber="340" unit="mm" value="1.335" valueMax="1.36" valueMin="1.31">1.31–1.36 mm</quantity>
. Metaventral processes elongate, emarginate at apices (
<figureCitation id="131ABEFC690B896FFE06CC6C60FC9E5E" box="[402,476,1615,1640]" captionStart="FIGURE 6" captionStartId="10.[151,250,1505,1527]" captionTargetBox="[154,1420,201,1480]" captionTargetId="figure@10.[151,1435,195,1483]" captionTargetPageId="10" captionText="FIGURE 6. Lectotype of Pselaphodes spinosus, male. A. Dorsal habitus. B. Labels of the lectotype. C. Antennal club. D. Metaventral process. E. Protrochanter and profemur. F. Mesotrochanter and mesofemur. G. Aedeagus, in dorsal view. H. Same, in lateral view. Scales: A = 1.0 mm; C – F = 0.3 mm; G, H = 0.2 mm." httpUri="https://zenodo.org/record/241590/files/figure.png" pageId="9" pageNumber="340">Figs 6</figureCitation>
D, 7E). Protrochanters, profemora (
<figureCitation id="131ABEFC690B896FFCE0CC6C629D9E5E" box="[884,957,1615,1640]" captionStart="FIGURE 6" captionStartId="10.[151,250,1505,1527]" captionTargetBox="[154,1420,201,1480]" captionTargetId="figure@10.[151,1435,195,1483]" captionTargetPageId="10" captionText="FIGURE 6. Lectotype of Pselaphodes spinosus, male. A. Dorsal habitus. B. Labels of the lectotype. C. Antennal club. D. Metaventral process. E. Protrochanter and profemur. F. Mesotrochanter and mesofemur. G. Aedeagus, in dorsal view. H. Same, in lateral view. Scales: A = 1.0 mm; C – F = 0.3 mm; G, H = 0.2 mm." httpUri="https://zenodo.org/record/241590/files/figure.png" pageId="9" pageNumber="340">Figs 6</figureCitation>
E, 7F), mesotrochanters, and mesofemora (
<figureCitation id="131ABEFC690B896FFF0BCC5761C49EBB" box="[159,228,1652,1677]" captionStart="FIGURE 6" captionStartId="10.[151,250,1505,1527]" captionTargetBox="[154,1420,201,1480]" captionTargetId="figure@10.[151,1435,195,1483]" captionTargetPageId="10" captionText="FIGURE 6. Lectotype of Pselaphodes spinosus, male. A. Dorsal habitus. B. Labels of the lectotype. C. Antennal club. D. Metaventral process. E. Protrochanter and profemur. F. Mesotrochanter and mesofemur. G. Aedeagus, in dorsal view. H. Same, in lateral view. Scales: A = 1.0 mm; C – F = 0.3 mm; G, H = 0.2 mm." httpUri="https://zenodo.org/record/241590/files/figure.png" pageId="9" pageNumber="340">Figs 6</figureCitation>
F, 7G) strongly spinose ventrally. Abdomen broad at base and narrowed posteriorly, AL
<quantity id="4CD90F9C690B896FFB54CC5764449EBA" box="[1216,1380,1652,1676]" metricMagnitude="-3" metricUnit="m" metricValue="1.125" metricValueMax="1.17" metricValueMin="1.08" pageId="9" pageNumber="340" unit="mm" value="1.125" valueMax="1.17" valueMin="1.08">1.08–1.17 mm</quantity>
, AW
<quantity id="4CD90F9C690B896FFF03CCB460619E86" box="[151,321,1687,1712]" metricMagnitude="-3" metricUnit="m" metricValue="1.365" metricValueMax="1.37" metricValueMin="1.36" pageId="9" pageNumber="340" unit="mm" value="1.365" valueMax="1.37" valueMin="1.36">1.36–1.37 mm</quantity>
. Length of aedeagus
<quantity id="4CD90F9C690B896FFDD5CCB4638C9E86" box="[577,684,1687,1712]" metricMagnitude="-4" metricUnit="m" metricValue="6.6" pageId="9" pageNumber="340" unit="mm" value="0.66">0.66 mm</quantity>
; aedeagus (
<figureCitation id="131ABEFC690B896FFCA2CCB462A19E86" box="[822,897,1687,1712]" captionStart="FIGURE 6" captionStartId="10.[151,250,1505,1527]" captionTargetBox="[154,1420,201,1480]" captionTargetId="figure@10.[151,1435,195,1483]" captionTargetPageId="10" captionText="FIGURE 6. Lectotype of Pselaphodes spinosus, male. A. Dorsal habitus. B. Labels of the lectotype. C. Antennal club. D. Metaventral process. E. Protrochanter and profemur. F. Mesotrochanter and mesofemur. G. Aedeagus, in dorsal view. H. Same, in lateral view. Scales: A = 1.0 mm; C – F = 0.3 mm; G, H = 0.2 mm." httpUri="https://zenodo.org/record/241590/files/figure.png" pageId="9" pageNumber="340">Figs 6</figureCitation>
G, 6H, 7H,
<date id="FF9F84B9690B896FFB9BCCBB65079E86" box="[1039,1063,1688,1712]" pageId="9" pageNumber="340">7I</date>
) with asymmetric median lobe, endophallus composed of two short sclerites.
</paragraph>
<paragraph id="8B9EA279690B896FFF53CCFC631F9F9A" blockId="9.[151,1437,715,2001]" pageId="9" pageNumber="340">
<emphasis id="B9557E6B690B896FFF53CCFC60909ECE" bold="true" box="[199,432,1759,1784]" pageId="9" pageNumber="340">Comparative notes.</emphasis>
The unique form of the strongly modified antennomere X, combined with the protuberant preapical portions of the protibiae, long metaventral processes, spination of the fore and middle legs, and structure of the aedeagus readily separate
<taxonomicName id="4C21D9FA690B896FFD99CD0462249F76" box="[525,772,1831,1856]" class="Insecta" family="Staphylinidae" genus="Pselaphodes" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="340" phylum="Arthropoda" rank="species" species="spinosus">
<emphasis id="B9557E6B690B896FFD99CD0462249F76" box="[525,772,1831,1856]" italics="true" pageId="9" pageNumber="340">Pselaphodes spinosus</emphasis>
</taxonomicName>
from all other known congeners. After an examination of the
<typeStatus id="549A1CDB690B896FFF55CD6F60069F53" box="[193,294,1868,1893]" pageId="9" pageNumber="340" type="holotype">holotype</typeStatus>
,
<taxonomicName id="4C21D9FA690B896FFEA6CD6F63129F53" box="[306,562,1868,1893]" class="Insecta" family="Staphylinidae" genus="Labomimus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="340" phylum="Arthropoda" rank="species" species="championi">
<emphasis id="B9557E6B690B896FFEA6CD6F63129F53" box="[306,562,1868,1893]" italics="true" pageId="9" pageNumber="340">Labomimus championi</emphasis>
</taxonomicName>
is here transferred to the genus
<taxonomicName id="4C21D9FA690B896FFC0BCD6F650D9F53" box="[927,1069,1868,1893]" class="Insecta" family="Staphylinidae" genus="Pselaphodes" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="340" phylum="Arthropoda" rank="genus">
<emphasis id="B9557E6B690B896FFC0BCD6F650D9F53" box="[927,1069,1868,1893]" italics="true" pageId="9" pageNumber="340">Pselaphodes</emphasis>
</taxonomicName>
based on the lack of the median metaventral fovea, and this species is placed as a junior synonym of
<taxonomicName id="4C21D9FA690B896FFC2DCD4C65919FBE" box="[953,1201,1903,1928]" class="Insecta" family="Staphylinidae" genus="Pselaphodes" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="340" phylum="Arthropoda" rank="species" species="spinosus">
<emphasis id="B9557E6B690B896FFC2DCD4C65919FBE" box="[953,1201,1903,1928]" italics="true" pageId="9" pageNumber="340">Pselaphodes spinosus</emphasis>
</taxonomicName>
by possessing same diagnostic characters (
<figureCitation id="131ABEFC690B896FFE06CDB760F49F9A" box="[402,468,1940,1965]" captionStart="FIGURE 7" captionStartId="11.[151,250,1584,1606]" captionTargetBox="[159,1425,592,1559]" captionTargetId="figure@11.[151,1435,589,1563]" captionTargetPageId="11" captionText="FIGURE 7. Holotype of Labomimus championi, male. A. Dorsal habitus. B. Labels of the holotype. C, D. Antennal club. E. Metaventral process. F. Protrochanter and profemur. G. Mesotrochanter and mesofemur. H. Aedeagus, in dorsal view. I. Same, in lateral view. Scales: A = 1.0 mm; C – G = 0.3 mm; H. I = 0.2 mm." httpUri="https://zenodo.org/record/241591/files/figure.png" pageId="9" pageNumber="340">Fig. 7</figureCitation>
A, 7C–I).
</paragraph>
</subSubSection>
<subSubSection id="C33BF1F2690B896FFF53CD9B64469FE7" box="[199,1382,1976,2001]" pageId="9" pageNumber="340" type="distribution">
<paragraph id="8B9EA279690B896FFF53CD9B64469FE7" blockId="9.[151,1437,715,2001]" box="[199,1382,1976,2001]" pageId="9" pageNumber="340">
<emphasis id="B9557E6B690B896FFF53CD9B60409FE7" bold="true" box="[199,352,1976,2001]" pageId="9" pageNumber="340">Distribution.</emphasis>
<taxonomicName id="4C21D9FA690B896FFEF3CD9B637C9FE6" box="[359,604,1976,2001]" class="Insecta" family="Staphylinidae" genus="Pselaphodes" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="340" phylum="Arthropoda" rank="species" species="spinosus">
<emphasis id="B9557E6B690B896FFEF3CD9B637C9FE6" box="[359,604,1976,2001]" italics="true" pageId="9" pageNumber="340">Pselaphodes spinosus</emphasis>
</taxonomicName>
is known from Haldwani in the northern Indian state of Uttarakhand.
</paragraph>
</subSubSection>
</treatment>
</document>
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