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<document ID-DOI="http://doi.org/10.5281/zenodo.3803506" ID-GBIF-Dataset="fe76f860-39b1-4520-a13c-d01a47dfc130" ID-GBIF-Taxon="163738928" ID-ISSN="1175-5334" ID-Zenodo-Dep="3803506" ID-ZooBank="urn:lsid:zoobank.org:pub:BA570D47-FE77-446C-B000-9208C9A6E957" checkinTime="1587753273724" checkinUser="carolina" docAuthor="Ríos-Tamayo, Duniesky &amp; Lyle, Robin" docDate="2020" docId="03F11B20FFFBFFAF8EE83EBFFCA8D0C1" docLanguage="en" docName="zootaxa.4766.2.2.pdf.imf" docOrigin="Zootaxa 4766 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Lepthercus sofiae Ríos-Tamayo &amp; Lyle 2020, sp. nov." docType="treatment" docVersion="5" lastPageId="39" lastPageNumber="300" masterDocId="FFC86358FFDEFF888E7F3E26FFD2D232" masterDocTitle="The South African genus Lepthercus Purcell, 1902 (Araneae: Mygalomorphae) phylogeny and taxonomy" masterLastPageNumber="305" masterPageNumber="261" pageId="37" pageNumber="298" updateTime="1636538462881" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>The South African genus Lepthercus Purcell, 1902 (Araneae: Mygalomorphae) phylogeny and taxonomy</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Ríos-Tamayo, Duniesky</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Lyle, Robin</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Zootaxa</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2020</mods:date>
<mods:detail type="pubDate">
<mods:number>2020-04-20</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>4766</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>2</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>261</mods:start>
<mods:end>305</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.11646/zootaxa.4766.2.2</mods:identifier>
<mods:identifier type="GBIF-Dataset">fe76f860-39b1-4520-a13c-d01a47dfc130</mods:identifier>
<mods:identifier type="ISSN">1175-5334</mods:identifier>
<mods:identifier type="Zenodo-Dep">3765467</mods:identifier>
<mods:identifier type="ZooBank">urn:lsid:zoobank.org:pub:BA570D47-FE77-446C-B000-9208C9A6E957</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.3803506" ID-GBIF-Taxon="163738928" ID-Zenodo-Dep="3803506" LSID="urn:lsid:plazi:treatment:03F11B20FFFBFFAF8EE83EBFFCA8D0C1" httpUri="http://treatment.plazi.org/id/03F11B20FFFBFFAF8EE83EBFFCA8D0C1" lastPageId="39" lastPageNumber="300" pageId="37" pageNumber="298">
<subSubSection box="[151,466,153,180]" pageId="37" pageNumber="298" type="nomenclature">
<paragraph blockId="37.[151,1139,153,251]" box="[151,466,153,180]" pageId="37" pageNumber="298">
<heading bold="true" box="[151,466,153,180]" fontSize="10" level="2" pageId="37" pageNumber="298" reason="2">
<emphasis bold="true" box="[151,466,153,180]" pageId="37" pageNumber="298">
<taxonomicName authority="Ríos-Tamayo &amp; Lyle, 2020" authorityName="Ríos-Tamayo &amp; Lyle" authorityYear="2020" box="[151,366,153,180]" class="Arachnida" family="Nemesiidae" genus="Lepthercus" kingdom="Animalia" order="Araneae" pageId="37" pageNumber="298" phylum="Arthropoda" rank="species" species="sofiae" status="sp. nov.">
<emphasis bold="true" box="[151,366,153,180]" italics="true" pageId="37" pageNumber="298">Lepthercus sofiae</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[373,466,154,180]" pageId="37" pageNumber="298" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection pageId="37" pageNumber="298" type="description">
<paragraph blockId="37.[151,1139,153,251]" box="[151,395,189,215]" pageId="37" pageNumber="298">
(
<figureCitation box="[159,259,189,215]" captionStart="FIGURE" captionStartId="7.[151,251,1647,1672]" captionTargetBox="[151,1436,381,1622]" captionTargetId="figure@7.[151,1436,381,1622]" captionTargetPageId="7" captionText="FIGURE. 2. Preening comb. A Lepthercus haddadi sp. nov. (female paratype, NCA 2007/3847). Maxillary cuspules (males). B L. dippenaarae. C L. kwazuluensis. Serrula. D L. mandelai sp. nov. (female, NCA 2014/415A); E L. filmeri sp. nov. (female, NCA 91/1437); F L. haddadi sp. nov. (male, NCA 2007/1486); G L. haddadi sp. nov. (female, NCA 2007/3847); H L. engelbrechti sp. nov. (female, NCA 2018/385); I L. engelbrechti sp. nov. (male, NCA 2018/384); J L. dregei Purcell, 1902 (female, NCA 2012/2333); K L. sofiae sp. nov. (male, NCA 97/395)." figureDoi="http://doi.org/10.5281/zenodo.3765471" httpUri="https://zenodo.org/record/3765471/files/figure.png" pageId="37" pageNumber="298">Figs. 2K</figureCitation>
,
<figureCitation box="[271,299,189,215]" captionStart="FIGURE" captionStartId="37.[151,251,1673,1698]" captionTargetBox="[151,1436,669,1648]" captionTargetId="figure@37.[151,1436,669,1648]" captionTargetPageId="37" captionText="FIGURE. 23. Lepthercus sofiae sp. nov. (male holotype, NCA 97/395). A habitus, dorsal view; B sternum, ventral view; C chelicerae, arrow indicates the pallid intercheliceral tumesence, prolateral view; D right pedipalp, arrow indicates the palpal tibia base dorsally very convex and the development of the anterior side of the retrolateral side of the ventral excavation, retrolateral view; E right pedipalp, arrows indicates the few proventral setae and the development of the anterior side of the retrolateral side of the ventral excavation, prolateral view; FH copulatory bulb, F retrolateral view, G ventral view, H prolateral view, arrows indicate the two keels in the embolus; I tibia-metatarsus I, showing the curved metatarsus, arrow indicate the numerous ventral short and strong setae, prolateral view; J tibia I, arrow indicates the strong curved spine near of the base of the megaspine, ventral view." figureDoi="http://doi.org/10.5281/zenodo.3765517" httpUri="https://zenodo.org/record/3765517/files/figure.png" pageId="37" pageNumber="298">23</figureCitation>
AJ, 24)
</paragraph>
<paragraph blockId="37.[151,1139,153,251]" box="[151,1139,227,251]" pageId="37" pageNumber="298">
http://zoobank.org/
<uri box="[360,1139,227,251]" pageId="37" pageNumber="298">
urn:lsid:zoobank.org:act:
<uuid box="[620,1139,227,251]" pageId="37" pageNumber="298">D717E390-D4BE-4507-A1FC-7C567BEC818B</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection pageId="37" pageNumber="298" type="materials_examined">
<paragraph blockId="37.[151,1437,297,648]" pageId="37" pageNumber="298">
<emphasis bold="true" box="[151,554,297,324]" pageId="37" pageNumber="298">Material examined. Type material</emphasis>
:
<materialsCitation ID-GBIF-Occurrence="2608670261" collectingDate="1984-03" collectionCode="NCA" collectorName="Koen, J. H." country="South Africa" latitude="-33.916668" location="Gouna State Forest" longLatPrecision="1202" longitude="23.033333" municipality="Lily Vlei Nature Reserve" pageId="37" pageNumber="298" specimenCode="NCA 97" specimenCount="1" specimenCount-male="1" stateProvince="Western Cape Province" typeStatus="holotype">
<typeStatus box="[568,672,297,323]" pageId="37" pageNumber="298">Holotype</typeStatus>
<specimenCount box="[678,699,298,324]" pageId="37" pageNumber="298" type="male"></specimenCount>
.
<collectingCountry box="[712,865,297,324]" name="South Africa" pageId="37" pageNumber="298">SOUTH AFRICA</collectingCountry>
:
<collectingRegion box="[878,1140,297,324]" country="South Africa" name="Western Cape" pageId="37" pageNumber="298">Western Cape Province</collectingRegion>
,
<collectingMunicipality box="[1151,1432,297,324]" pageId="37" pageNumber="298">Lily Vlei Nature Reserve</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:03F11B20FFFBFFAF8EE83EBFFCA8D0C1:8E87FCEDFFFBFFAD8EE83F6BFEA6D35A" box="[151,372,333,360]" country="South Africa" latitude="-33.916668" longLatPrecision="1202" longitude="23.033333" municipality="Lily Vlei Nature Reserve" name="Gouna State Forest" pageId="37" pageNumber="298" stateProvince="Western Cape Province">Gouna State Forest</location>
,
<locationDeviation box="[385,531,333,360]" location="Knysna" pageId="37" pageNumber="298">N of Knysna</locationDeviation>
(
<geoCoordinate box="[549,641,333,360]" degrees="33" direction="south" minutes="55" orientation="latitude" pageId="37" pageNumber="298" precision="925" value="-33.916668">33°55S</geoCoordinate>
<geoCoordinate box="[650,731,333,359]" degrees="23" direction="east" minutes="2" orientation="longitude" pageId="37" pageNumber="298" precision="925" value="23.033333">23°2E</geoCoordinate>
),
<date box="[754,843,333,359]" pageId="37" pageNumber="298" value="1984-03">
<collectingDate box="[754,843,333,359]" pageId="37" pageNumber="298" value="1984-03">iii.1984</collectingDate>
</date>
,
<collectorName box="[856,986,334,360]" pageId="37" pageNumber="298">Koen, J. H.</collectorName>
<emphasis box="[995,1036,333,359]" italics="true" pageId="37" pageNumber="298">leg.</emphasis>
, (
<specimenCode box="[1059,1156,333,360]" collectionCode="NCA" pageId="37" pageNumber="298">NCA 97</specimenCode>
/395)
</materialsCitation>
.
<typeStatus box="[1253,1355,334,360]" pageId="37" pageNumber="298">Paratype</typeStatus>
.
<collectingCountry name="South Africa" pageId="37" pageNumber="298">SOUTH AFRICA</collectingCountry>
: same data as
<typeStatus box="[392,493,369,395]" pageId="37" pageNumber="298">holotype</typeStatus>
,
<specimenCount box="[504,567,369,396]" pageId="37" pageNumber="298" type="male">2 ♂♂</specimenCount>
(
<collectionCode box="[582,642,369,395]" pageId="37" pageNumber="298">NCA</collectionCode>
97/395).
</paragraph>
<paragraph blockId="37.[151,1437,297,648]" pageId="37" pageNumber="298">
<emphasis bold="true" box="[199,499,405,432]" pageId="37" pageNumber="298">Other material examined</emphasis>
:
<collectingCountry box="[514,667,405,432]" name="South Africa" pageId="37" pageNumber="298">SOUTH AFRICA</collectingCountry>
:
<collectingRegion box="[680,946,405,432]" country="South Africa" name="Western Cape" pageId="37" pageNumber="298">Western Cape Province</collectingRegion>
:
<collectingCounty box="[956,1270,405,432]" pageId="37" pageNumber="298">Groeneweide Forest Station</collectingCounty>
,
<locationDeviation box="[1281,1436,405,432]" location="George" pageId="37" pageNumber="298">NE of George</locationDeviation>
(
<geoCoordinate box="[160,250,441,468]" degrees="33" direction="south" minutes="57" orientation="latitude" pageId="37" pageNumber="298" precision="925" value="-33.95">33°57S</geoCoordinate>
<geoCoordinate box="[256,351,441,468]" degrees="22" direction="east" minutes="27" orientation="longitude" pageId="37" pageNumber="298" precision="925" value="22.45">22°27E</geoCoordinate>
),
<date box="[371,458,441,467]" pageId="37" pageNumber="298" value="1987-06">
<collectingDate box="[371,458,441,467]" pageId="37" pageNumber="298" value="1987-06">vi.1987</collectingDate>
</date>
,
<collectorName box="[471,599,442,468]" pageId="37" pageNumber="298">Koen, J. H.</collectorName>
<emphasis box="[607,648,441,467]" italics="true" pageId="37" pageNumber="298">leg.</emphasis>
,
<specimenCount box="[663,727,441,468]" pageId="37" pageNumber="298" type="male">2 ♂♂</specimenCount>
(
<specimenCode box="[742,838,441,467]" collectionCode="NCA" pageId="37" pageNumber="298">NCA 97</specimenCode>
/291);
<collectingMunicipality box="[912,1229,441,468]" pageId="37" pageNumber="298">Groeneweide Forest Station</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:03F11B20FFFBFFAF8EE83EBFFCA8D0C1:8E87FCEDFFFBFFAD8AA63F9FFA9ED3E1" box="[1241,1356,441,467]" country="South Africa" county="Groeneweide Forest Station" latitude="-33.916668" longLatPrecision="1202" longitude="22.733334" municipality="Groeneweide Forest Station" name="Groenkop" pageId="37" pageNumber="298" stateProvince="Western Cape">Groenkop</location>
,
<locationDeviation location="George" pageId="37" pageNumber="298">NE of George</locationDeviation>
(
<geoCoordinate box="[249,339,477,504]" degrees="33" direction="south" minutes="57" orientation="latitude" pageId="37" pageNumber="298" precision="925" value="-33.95">33°57S</geoCoordinate>
<geoCoordinate box="[345,440,477,504]" degrees="22" direction="east" minutes="27" orientation="longitude" pageId="37" pageNumber="298" precision="925" value="22.45">22°27E</geoCoordinate>
),
<date box="[459,555,477,503]" pageId="37" pageNumber="298" value="1986-07">
<collectingDate box="[459,555,477,503]" pageId="37" pageNumber="298" value="1986-07">vii.1986</collectingDate>
</date>
,
<collectorName box="[566,693,478,504]" pageId="37" pageNumber="298">Koen, J. H.</collectorName>
<emphasis box="[700,742,477,503]" italics="true" pageId="37" pageNumber="298">leg.</emphasis>
,
<specimenCount box="[756,819,478,504]" pageId="37" pageNumber="298" type="male">7 ♂♂</specimenCount>
(
<specimenCode box="[833,929,477,503]" collectionCode="NCA" pageId="37" pageNumber="298">NCA 97</specimenCode>
/331);
<location LSID="urn:lsid:plazi:treatment:03F11B20FFFBFFAF8EE83EBFFCA8D0C1:8E87FCEDFFFBFFAD8D953FFBFAD5D3CA" box="[1002,1287,477,504]" country="South Africa" county="Groeneweide Forest Station" latitude="-33.916668" longLatPrecision="1202" longitude="22.733334" municipality="Groeneweide Forest Station" name="Lily Vlei Nature Reserve" pageId="37" pageNumber="298" stateProvince="Western Cape">Lily Vlei Nature Reserve</location>
,
<location LSID="urn:lsid:plazi:treatment:03F11B20FFFBFFAF8EE83EBFFCA8D0C1:8E87FCEDFFFBFFAD8B6D3FFBFF32D02E" country="South Africa" county="Groeneweide Forest Station" latitude="-33.916668" longLatPrecision="1202" longitude="22.733334" municipality="Groeneweide Forest Station" name="Gouna State Forest" pageId="37" pageNumber="298" stateProvince="Western Cape">Gouna State Forest</location>
,
<locationDeviation box="[236,381,513,540]" location="Knysna" pageId="37" pageNumber="298">N of Knysna</locationDeviation>
(
<geoCoordinate box="[397,490,513,540]" degrees="33" direction="south" minutes="55" orientation="latitude" pageId="37" pageNumber="298" precision="925" value="-33.916668">33°55S</geoCoordinate>
<geoCoordinate box="[498,580,513,539]" degrees="23" direction="east" minutes="2" orientation="longitude" pageId="37" pageNumber="298" precision="925" value="23.033333">23°2E</geoCoordinate>
),
<date box="[601,683,513,539]" pageId="37" pageNumber="298" value="1984-02">
<collectingDate box="[601,683,513,539]" pageId="37" pageNumber="298" value="1984-02">ii.1984</collectingDate>
</date>
,
<collectorName box="[695,823,514,540]" pageId="37" pageNumber="298">Koen, J. H.</collectorName>
<emphasis box="[831,872,513,539]" italics="true" pageId="37" pageNumber="298">leg.</emphasis>
,
<specimenCount box="[887,930,513,540]" pageId="37" pageNumber="298" type="male">1 ♂</specimenCount>
(
<specimenCode box="[945,1041,513,539]" collectionCode="NCA" pageId="37" pageNumber="298">NCA 97</specimenCode>
/283);
<location LSID="urn:lsid:plazi:treatment:03F11B20FFFBFFAF8EE83EBFFCA8D0C1:8E87FCEDFFFBFFAD8A243C27FA4AD029" box="[1115,1432,513,540]" country="South Africa" county="Groeneweide Forest Station" latitude="-33.916668" longLatPrecision="1202" longitude="22.733334" municipality="Groeneweide Forest Station" name="Groeneweide Forest Station" pageId="37" pageNumber="298" stateProvince="Western Cape">Groeneweide Forest Station</location>
,
<location LSID="urn:lsid:plazi:treatment:03F11B20FFFBFFAF8EE83EBFFCA8D0C1:8E87FCEDFFFBFFAD8EE83C03FED8D00D" box="[151,266,549,575]" country="South Africa" county="Groeneweide Forest Station" latitude="-33.916668" longLatPrecision="1202" longitude="22.733334" municipality="Groeneweide Forest Station" name="Groenkop" pageId="37" pageNumber="298" stateProvince="Western Cape">Groenkop</location>
,
<locationDeviation box="[281,445,549,576]" location="George" pageId="37" pageNumber="298">NE of George</locationDeviation>
(
<geoCoordinate box="[464,555,549,576]" degrees="33" direction="south" minutes="57" orientation="latitude" pageId="37" pageNumber="298" precision="925" value="-33.95">33°57S</geoCoordinate>
<geoCoordinate box="[561,656,549,576]" degrees="22" direction="east" minutes="27" orientation="longitude" pageId="37" pageNumber="298" precision="925" value="22.45">22°27E</geoCoordinate>
),
<date box="[679,756,549,575]" pageId="37" pageNumber="298" value="1987-05">
<collectingDate box="[679,756,549,575]" pageId="37" pageNumber="298" value="1987-05">v.1987</collectingDate>
</date>
,
<collectorName box="[772,906,550,576]" pageId="37" pageNumber="298">Koen, J. H.</collectorName>
<emphasis box="[917,958,549,575]" italics="true" pageId="37" pageNumber="298">leg.</emphasis>
,
<specimenCount box="[976,1043,549,576]" pageId="37" pageNumber="298" type="male">2 ♂♂</specimenCount>
(
<specimenCode box="[1061,1161,549,575]" collectionCode="NCA" pageId="37" pageNumber="298">NCA 97</specimenCode>
/244);
<location LSID="urn:lsid:plazi:treatment:03F11B20FFFBFFAF8EE83EBFFCA8D0C1:8E87FCEDFFFBFFAD8AAA3C00FAF9D072" box="[1237,1323,550,576]" country="South Africa" county="Groeneweide Forest Station" latitude="-33.916668" longLatPrecision="1202" longitude="22.733334" municipality="Groeneweide Forest Station" name="Knysna" pageId="37" pageNumber="298" stateProvince="Western Cape">Knysna</location>
(Ara019) (
<geoCoordinate box="[160,250,585,612]" degrees="33" direction="south" minutes="57" orientation="latitude" pageId="37" pageNumber="298" precision="925" value="-33.95">33°57S</geoCoordinate>
<geoCoordinate box="[256,336,585,612]" degrees="23" direction="east" minutes="3" orientation="longitude" pageId="37" pageNumber="298" precision="925" value="23.05">23°3E</geoCoordinate>
),
<date box="[357,465,585,611]" pageId="37" pageNumber="298" value="2014-11-01">
<collectingDate box="[357,465,585,611]" pageId="37" pageNumber="298" value="2014-11-01">1.xi.2014</collectingDate>
</date>
,
<location LSID="urn:lsid:plazi:treatment:03F11B20FFFBFFAF8EE83EBFFCA8D0C1:8E87FCEDFFFBFFAD8FA03C6FFDF1D051" box="[479,547,585,611]" country="South Africa" county="Groeneweide Forest Station" latitude="-33.916668" longLatPrecision="1202" longitude="22.733334" municipality="Groeneweide Forest Station" name="Swart" pageId="37" pageNumber="298" stateProvince="Western Cape">Swart</location>
,
<collectionCode box="[559,581,586,612]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="37" pageNumber="298">R</collectionCode>
. C.
<emphasis box="[627,668,585,611]" italics="true" pageId="37" pageNumber="298">leg.</emphasis>
,
<specimenCount box="[684,748,585,612]" pageId="37" pageNumber="298" type="male">8 ♂♂</specimenCount>
(
<specimenCode box="[764,891,585,612]" collectionCode="NCA" pageId="37" pageNumber="298">NCA 2017</specimenCode>
/1609); same data (
<geoCoordinate box="[1106,1196,585,612]" degrees="33" direction="south" minutes="55" orientation="latitude" pageId="37" pageNumber="298" precision="925" value="-33.916668">33°55S</geoCoordinate>
<geoCoordinate box="[1203,1296,585,612]" degrees="22" direction="east" minutes="44" orientation="longitude" pageId="37" pageNumber="298" precision="925" value="22.733334">22°44E</geoCoordinate>
),
<specimenCount box="[1318,1361,585,612]" pageId="37" pageNumber="298" type="male">4 ♂</specimenCount>
(
<specimenCode collectionCode="NCA" pageId="37" pageNumber="298">NCA 2017</specimenCode>
/1611).
</paragraph>
</subSubSection>
<caption ID-DOI="http://doi.org/10.5281/zenodo.3765517" ID-Zenodo-Dep="3765517" httpUri="https://zenodo.org/record/3765517/files/figure.png" pageId="37" pageNumber="298" startId="37.[151,251,1673,1698]" targetBox="[151,1436,669,1648]" targetPageId="37">
<paragraph blockId="37.[151,1437,1673,1950]" pageId="37" pageNumber="298">
<emphasis bold="true" box="[151,293,1673,1698]" pageId="37" pageNumber="298">FIGURE. 23.</emphasis>
<taxonomicName authority="Ríos-Tamayo &amp; Lyle, 2020" authorityName="Ríos-Tamayo &amp; Lyle" authorityYear="2020" box="[298,477,1674,1698]" class="Arachnida" family="Nemesiidae" genus="Lepthercus" kingdom="Animalia" order="Araneae" pageId="37" pageNumber="298" phylum="Arthropoda" rank="species" species="sofiae" status="sp. nov.">
<emphasis box="[298,477,1674,1698]" italics="true" pageId="37" pageNumber="298">Lepthercus sofiae</emphasis>
</taxonomicName>
<emphasis bold="true" box="[481,561,1674,1698]" pageId="37" pageNumber="298">
<taxonomicNameLabel box="[481,561,1674,1698]" pageId="37" pageNumber="298" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(male
<typeStatus box="[630,721,1673,1698]" pageId="37" pageNumber="298">holotype</typeStatus>
,
<collectionCode box="[730,782,1674,1698]" pageId="37" pageNumber="298">NCA</collectionCode>
97/395).
<emphasis bold="true" box="[878,896,1673,1698]" pageId="37" pageNumber="298">A</emphasis>
habitus, dorsal view;
<emphasis bold="true" box="[1116,1133,1674,1698]" pageId="37" pageNumber="298">B</emphasis>
sternum, ventral view;
<emphasis bold="true" box="[1369,1387,1673,1698]" pageId="37" pageNumber="298">C</emphasis>
chelicerae, arrow indicates the pallid intercheliceral tumesence, prolateral view;
<emphasis bold="true" box="[939,957,1710,1734]" pageId="37" pageNumber="298">D</emphasis>
right pedipalp, arrow indicates the palpal tibia base dorsally very convex and the development of the anterior side of the retrolateral side of the ventral excavation, retrolateral view;
<emphasis bold="true" box="[213,230,1782,1806]" pageId="37" pageNumber="298">E</emphasis>
right pedipalp, arrows indicates the few proventral setae and the development of the anterior side of the retrolateral side of the ventral excavation, prolateral view;
<emphasis bold="true" box="[583,632,1818,1842]" pageId="37" pageNumber="298">FH</emphasis>
copulatory bulb,
<emphasis bold="true" box="[811,826,1818,1842]" pageId="37" pageNumber="298">F</emphasis>
retrolateral view,
<emphasis bold="true" box="[1012,1032,1817,1842]" pageId="37" pageNumber="298">G</emphasis>
ventral view,
<emphasis bold="true" box="[1176,1196,1818,1842]" pageId="37" pageNumber="298">H</emphasis>
prolateral view, arrows indicate the two keels in the embolus;
<emphasis bold="true" box="[540,550,1854,1878]" pageId="37" pageNumber="298">I</emphasis>
tibia-metatarsus I, showing the curved metatarsus, arrow indicate the numerous ventral short and strong setae, prolateral view;
<emphasis bold="true" box="[563,576,1890,1914]" pageId="37" pageNumber="298">J</emphasis>
tibia I, arrow indicates the strong curved spine near of the base of the megaspine, ventral view.
</paragraph>
</caption>
<subSubSection pageId="38" pageNumber="299" type="etymology">
<paragraph blockId="38.[151,1437,153,863]" pageId="38" pageNumber="299">
<emphasis bold="true" box="[199,334,153,179]" pageId="38" pageNumber="299">Etymology.</emphasis>
The specific epithet is a patronym in honour of Sofia Casadey Bollati, the first authors partner, for her continuing support.
</paragraph>
</subSubSection>
<subSubSection pageId="38" pageNumber="299" type="diagnosis">
<paragraph blockId="38.[151,1437,153,863]" pageId="38" pageNumber="299">
<emphasis bold="true" box="[199,322,224,251]" pageId="38" pageNumber="299">Diagnosis.</emphasis>
Males of
<taxonomicName authority="Ríos-Tamayo &amp; Lyle, 2020" authorityName="Ríos-Tamayo &amp; Lyle" authorityYear="2020" box="[430,525,225,251]" class="Arachnida" family="Nemesiidae" genus="Lepthercus" kingdom="Animalia" order="Araneae" pageId="38" pageNumber="299" phylum="Arthropoda" rank="species" species="sofiae" status="sp. nov.">
<emphasis box="[430,525,225,251]" italics="true" pageId="38" pageNumber="299">L. sofiae</emphasis>
</taxonomicName>
<emphasis bold="true" box="[531,619,225,251]" pageId="38" pageNumber="299">
<taxonomicNameLabel box="[531,619,225,251]" pageId="38" pageNumber="299" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, can be distinguished from species present in “Group haddadi” by a metatarsus I with the presence of numerous short, strong setae ventrally 1:2A of the segment (
<figureCitation box="[1159,1252,261,287]" captionStart="FIGURE" captionStartId="37.[151,251,1673,1698]" captionTargetBox="[151,1436,669,1648]" captionTargetId="figure@37.[151,1436,669,1648]" captionTargetPageId="37" captionText="FIGURE. 23. Lepthercus sofiae sp. nov. (male holotype, NCA 97/395). A habitus, dorsal view; B sternum, ventral view; C chelicerae, arrow indicates the pallid intercheliceral tumesence, prolateral view; D right pedipalp, arrow indicates the palpal tibia base dorsally very convex and the development of the anterior side of the retrolateral side of the ventral excavation, retrolateral view; E right pedipalp, arrows indicates the few proventral setae and the development of the anterior side of the retrolateral side of the ventral excavation, prolateral view; FH copulatory bulb, F retrolateral view, G ventral view, H prolateral view, arrows indicate the two keels in the embolus; I tibia-metatarsus I, showing the curved metatarsus, arrow indicate the numerous ventral short and strong setae, prolateral view; J tibia I, arrow indicates the strong curved spine near of the base of the megaspine, ventral view." figureDoi="http://doi.org/10.5281/zenodo.3765517" httpUri="https://zenodo.org/record/3765517/files/figure.png" pageId="38" pageNumber="299">Fig. 23I</figureCitation>
; cf. blunt spines in others), and by the higher number of maxillae cuspules (140). It can also be recognized by a very developed, almost square palp tibia (
<figureCitation box="[430,561,333,359]" captionStart="FIGURE" captionStartId="37.[151,251,1673,1698]" captionTargetBox="[151,1436,669,1648]" captionTargetId="figure@37.[151,1436,669,1648]" captionTargetPageId="37" captionText="FIGURE. 23. Lepthercus sofiae sp. nov. (male holotype, NCA 97/395). A habitus, dorsal view; B sternum, ventral view; C chelicerae, arrow indicates the pallid intercheliceral tumesence, prolateral view; D right pedipalp, arrow indicates the palpal tibia base dorsally very convex and the development of the anterior side of the retrolateral side of the ventral excavation, retrolateral view; E right pedipalp, arrows indicates the few proventral setae and the development of the anterior side of the retrolateral side of the ventral excavation, prolateral view; FH copulatory bulb, F retrolateral view, G ventral view, H prolateral view, arrows indicate the two keels in the embolus; I tibia-metatarsus I, showing the curved metatarsus, arrow indicate the numerous ventral short and strong setae, prolateral view; J tibia I, arrow indicates the strong curved spine near of the base of the megaspine, ventral view." figureDoi="http://doi.org/10.5281/zenodo.3765517" httpUri="https://zenodo.org/record/3765517/files/figure.png" pageId="38" pageNumber="299">Fig. 23D, E</figureCitation>
) and by a more rounded bulb with two well marked keels on the embolus (
<figureCitation captionStart="FIGURE" captionStartId="37.[151,251,1673,1698]" captionTargetBox="[151,1436,669,1648]" captionTargetId="figure@37.[151,1436,669,1648]" captionTargetPageId="37" captionText="FIGURE. 23. Lepthercus sofiae sp. nov. (male holotype, NCA 97/395). A habitus, dorsal view; B sternum, ventral view; C chelicerae, arrow indicates the pallid intercheliceral tumesence, prolateral view; D right pedipalp, arrow indicates the palpal tibia base dorsally very convex and the development of the anterior side of the retrolateral side of the ventral excavation, retrolateral view; E right pedipalp, arrows indicates the few proventral setae and the development of the anterior side of the retrolateral side of the ventral excavation, prolateral view; FH copulatory bulb, F retrolateral view, G ventral view, H prolateral view, arrows indicate the two keels in the embolus; I tibia-metatarsus I, showing the curved metatarsus, arrow indicate the numerous ventral short and strong setae, prolateral view; J tibia I, arrow indicates the strong curved spine near of the base of the megaspine, ventral view." figureDoi="http://doi.org/10.5281/zenodo.3765517" httpUri="https://zenodo.org/record/3765517/files/figure.png" pageId="38" pageNumber="299">Fig. 23</figureCitation>
FH). Females are unknown.
</paragraph>
</subSubSection>
<subSubSection lastPageId="39" lastPageNumber="300" pageId="38" pageNumber="299" type="description">
<paragraph blockId="38.[151,1437,153,863]" pageId="38" pageNumber="299">
<emphasis bold="true" box="[199,345,404,431]" pageId="38" pageNumber="299">Description.</emphasis>
Male
<typeStatus box="[416,514,405,431]" pageId="38" pageNumber="299">holotype</typeStatus>
(NCA 97/395,
<figureCitation box="[687,766,405,431]" captionStart="FIGURE" captionStartId="37.[151,251,1673,1698]" captionTargetBox="[151,1436,669,1648]" captionTargetId="figure@37.[151,1436,669,1648]" captionTargetPageId="37" captionText="FIGURE. 23. Lepthercus sofiae sp. nov. (male holotype, NCA 97/395). A habitus, dorsal view; B sternum, ventral view; C chelicerae, arrow indicates the pallid intercheliceral tumesence, prolateral view; D right pedipalp, arrow indicates the palpal tibia base dorsally very convex and the development of the anterior side of the retrolateral side of the ventral excavation, retrolateral view; E right pedipalp, arrows indicates the few proventral setae and the development of the anterior side of the retrolateral side of the ventral excavation, prolateral view; FH copulatory bulb, F retrolateral view, G ventral view, H prolateral view, arrows indicate the two keels in the embolus; I tibia-metatarsus I, showing the curved metatarsus, arrow indicate the numerous ventral short and strong setae, prolateral view; J tibia I, arrow indicates the strong curved spine near of the base of the megaspine, ventral view." figureDoi="http://doi.org/10.5281/zenodo.3765517" httpUri="https://zenodo.org/record/3765517/files/figure.png" pageId="38" pageNumber="299">Fig. 23</figureCitation>
AJ). Total length 13.90.
<emphasis box="[1053,1163,405,431]" italics="true" pageId="38" pageNumber="299">Carapace</emphasis>
(
<figureCitation box="[1178,1284,405,431]" captionStart="FIGURE" captionStartId="37.[151,251,1673,1698]" captionTargetBox="[151,1436,669,1648]" captionTargetId="figure@37.[151,1436,669,1648]" captionTargetPageId="37" captionText="FIGURE. 23. Lepthercus sofiae sp. nov. (male holotype, NCA 97/395). A habitus, dorsal view; B sternum, ventral view; C chelicerae, arrow indicates the pallid intercheliceral tumesence, prolateral view; D right pedipalp, arrow indicates the palpal tibia base dorsally very convex and the development of the anterior side of the retrolateral side of the ventral excavation, retrolateral view; E right pedipalp, arrows indicates the few proventral setae and the development of the anterior side of the retrolateral side of the ventral excavation, prolateral view; FH copulatory bulb, F retrolateral view, G ventral view, H prolateral view, arrows indicate the two keels in the embolus; I tibia-metatarsus I, showing the curved metatarsus, arrow indicate the numerous ventral short and strong setae, prolateral view; J tibia I, arrow indicates the strong curved spine near of the base of the megaspine, ventral view." figureDoi="http://doi.org/10.5281/zenodo.3765517" httpUri="https://zenodo.org/record/3765517/files/figure.png" pageId="38" pageNumber="299">Fig. 23A</figureCitation>
): length 5.87, width 4.40, with lateral dark marginal bristles, with a little pubescence.
<emphasis box="[971,1152,441,467]" italics="true" pageId="38" pageNumber="299">Cephalic region</emphasis>
: length 3.59; clypeus almost absent, with 7 bristles on the edge, 6 bristles in front of the OQ and 6 between the PME.
<emphasis box="[1223,1293,478,503]" italics="true" pageId="38" pageNumber="299">Fovea</emphasis>
: width 0.45, short, almost straight (slightly recurved).
<emphasis box="[622,868,513,539]" italics="true" pageId="38" pageNumber="299">Ocular measurements</emphasis>
: AME 0.17, ALE 0.31, PME 0.19, PLE 0.24, OQ length 0.47, 0.88 width; AMEALE 0.04, PMEPLE 0.00, AMEPME 0.07, ALEPLE 0.05, AMEAME 0.08, OMPOMP 0.40.
<emphasis box="[353,476,585,611]" italics="true" pageId="38" pageNumber="299">Chelicerae</emphasis>
: length 2.55, width 1.55, with dark dorsalanterior bristles, intercheliceral tumescence small, little marked with setae (
<figureCitation box="[503,606,621,647]" captionStart="FIGURE" captionStartId="37.[151,251,1673,1698]" captionTargetBox="[151,1436,669,1648]" captionTargetId="figure@37.[151,1436,669,1648]" captionTargetPageId="37" captionText="FIGURE. 23. Lepthercus sofiae sp. nov. (male holotype, NCA 97/395). A habitus, dorsal view; B sternum, ventral view; C chelicerae, arrow indicates the pallid intercheliceral tumesence, prolateral view; D right pedipalp, arrow indicates the palpal tibia base dorsally very convex and the development of the anterior side of the retrolateral side of the ventral excavation, retrolateral view; E right pedipalp, arrows indicates the few proventral setae and the development of the anterior side of the retrolateral side of the ventral excavation, prolateral view; FH copulatory bulb, F retrolateral view, G ventral view, H prolateral view, arrows indicate the two keels in the embolus; I tibia-metatarsus I, showing the curved metatarsus, arrow indicate the numerous ventral short and strong setae, prolateral view; J tibia I, arrow indicates the strong curved spine near of the base of the megaspine, ventral view." figureDoi="http://doi.org/10.5281/zenodo.3765517" httpUri="https://zenodo.org/record/3765517/files/figure.png" pageId="38" pageNumber="299">Fig. 23C</figureCitation>
). Fang groove with 7 or 8 promarginal teeth and&gt;30 mesobasal denticles.
<emphasis box="[151,237,657,683]" italics="true" pageId="38" pageNumber="299">Labium</emphasis>
: length 0.39, width 0.86.
<emphasis box="[529,638,657,683]" italics="true" pageId="38" pageNumber="299">Maxillae:</emphasis>
length 1.77, width 0.65, with about 140 cuspules on inner corner, prolateral face slightly curved, soft area long, with long uniformly distributed hairs, serrula present (
<figureCitation box="[1233,1323,693,719]" captionStart="FIGURE" captionStartId="7.[151,251,1647,1672]" captionTargetBox="[151,1436,381,1622]" captionTargetId="figure@7.[151,1436,381,1622]" captionTargetPageId="7" captionText="FIGURE. 2. Preening comb. A Lepthercus haddadi sp. nov. (female paratype, NCA 2007/3847). Maxillary cuspules (males). B L. dippenaarae. C L. kwazuluensis. Serrula. D L. mandelai sp. nov. (female, NCA 2014/415A); E L. filmeri sp. nov. (female, NCA 91/1437); F L. haddadi sp. nov. (male, NCA 2007/1486); G L. haddadi sp. nov. (female, NCA 2007/3847); H L. engelbrechti sp. nov. (female, NCA 2018/385); I L. engelbrechti sp. nov. (male, NCA 2018/384); J L. dregei Purcell, 1902 (female, NCA 2012/2333); K L. sofiae sp. nov. (male, NCA 97/395)." figureDoi="http://doi.org/10.5281/zenodo.3765471" httpUri="https://zenodo.org/record/3765471/files/figure.png" pageId="38" pageNumber="299">Fig. 2K</figureCitation>
).
<emphasis box="[1343,1436,693,719]" italics="true" pageId="38" pageNumber="299">Sternum</emphasis>
(
<figureCitation box="[160,264,729,755]" captionStart="FIGURE" captionStartId="37.[151,251,1673,1698]" captionTargetBox="[151,1436,669,1648]" captionTargetId="figure@37.[151,1436,669,1648]" captionTargetPageId="37" captionText="FIGURE. 23. Lepthercus sofiae sp. nov. (male holotype, NCA 97/395). A habitus, dorsal view; B sternum, ventral view; C chelicerae, arrow indicates the pallid intercheliceral tumesence, prolateral view; D right pedipalp, arrow indicates the palpal tibia base dorsally very convex and the development of the anterior side of the retrolateral side of the ventral excavation, retrolateral view; E right pedipalp, arrows indicates the few proventral setae and the development of the anterior side of the retrolateral side of the ventral excavation, prolateral view; FH copulatory bulb, F retrolateral view, G ventral view, H prolateral view, arrows indicate the two keels in the embolus; I tibia-metatarsus I, showing the curved metatarsus, arrow indicate the numerous ventral short and strong setae, prolateral view; J tibia I, arrow indicates the strong curved spine near of the base of the megaspine, ventral view." figureDoi="http://doi.org/10.5281/zenodo.3765517" httpUri="https://zenodo.org/record/3765517/files/figure.png" pageId="38" pageNumber="299">Fig. 23B</figureCitation>
): length 2.93, maximum width 2.31.
<emphasis box="[687,793,729,755]" italics="true" pageId="38" pageNumber="299">Abdomen</emphasis>
: length 5.42, covered with short hair. PMS: length 0.78; PLS: length of basal:medial:apical segments 1.22:0.98:1.51; total length 3.71.
<emphasis box="[1052,1337,765,791]" italics="true" pageId="38" pageNumber="299">Lengths of legs and palp</emphasis>
: I: 3.46, 2.20, 2.97 (width/length: 0.47), 2.87, 1.79, 13.29. II: 3.77, 2.10, 2.46, 2.48, 1.80, 12.61. III: 3.60, 1.93, 2.15, 2.97, 1.69, 12.34. IV: 4.21, 2.22, 3.03, 4.04, 2.00, 15.50. Palp: 2.20, 1.07, 1.56, —, 0.90, 5.73.
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.3765519" ID-Zenodo-Dep="3765519" httpUri="https://zenodo.org/record/3765519/files/figure.png" pageId="38" pageNumber="299" startId="38.[151,251,1816,1841]" targetBox="[185,1405,888,1784]" targetPageId="38">
<paragraph blockId="38.[151,1437,1812,1953]" pageId="38" pageNumber="299">
<emphasis bold="true" box="[151,293,1816,1841]" pageId="38" pageNumber="299">FIGURE. 24.</emphasis>
Distribution map of
<taxonomicName authorityName="Purcell" authorityYear="1902" box="[502,614,1817,1841]" class="Arachnida" family="Nemesiidae" genus="Lepthercus" kingdom="Animalia" order="Araneae" pageId="38" pageNumber="299" phylum="Arthropoda" rank="genus">
<emphasis box="[502,614,1817,1841]" italics="true" pageId="38" pageNumber="299">Lepthercus</emphasis>
</taxonomicName>
spp: ●
<taxonomicName authority="Ríos-Tamayo &amp; Lyle, 2020" authorityName="Ríos-Tamayo &amp; Lyle" authorityYear="2020" box="[691,897,1817,1841]" class="Arachnida" family="Nemesiidae" genus="Lepthercus" kingdom="Animalia" order="Araneae" pageId="38" pageNumber="299" phylum="Arthropoda" rank="species" species="confusus" status="sp. nov.">
<emphasis box="[691,897,1817,1841]" italics="true" pageId="38" pageNumber="299">Lepthercus confusus</emphasis>
</taxonomicName>
<emphasis bold="true" box="[902,982,1817,1841]" pageId="38" pageNumber="299">
<taxonomicNameLabel box="[902,982,1817,1841]" pageId="38" pageNumber="299" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, ○
<taxonomicName authority="Ríos-Tamayo &amp; Lyle, 2020" authorityName="Ríos-Tamayo &amp; Lyle" authorityYear="2020" box="[1019,1264,1817,1841]" class="Arachnida" family="Nemesiidae" genus="Lepthercus" kingdom="Animalia" order="Araneae" pageId="38" pageNumber="299" phylum="Arthropoda" rank="species" species="dippenaarae" status="sp. nov.">
<emphasis box="[1019,1264,1817,1841]" italics="true" pageId="38" pageNumber="299">Lepthercus dippenaarae</emphasis>
</taxonomicName>
<emphasis bold="true" box="[1269,1349,1817,1841]" pageId="38" pageNumber="299">
<taxonomicNameLabel box="[1269,1349,1817,1841]" pageId="38" pageNumber="299" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
,
<emphasis bold="true" box="[1361,1385,1816,1839]" pageId="38" pageNumber="299">Ɨ</emphasis>
<taxonomicName authority="Purcell, 1902" authorityName="Purcell" authorityYear="1902" class="Arachnida" family="Nemesiidae" genus="Lepthercus" kingdom="Animalia" order="Araneae" pageId="38" pageNumber="299" phylum="Arthropoda" rank="species" species="dregei">
<emphasis italics="true" pageId="38" pageNumber="299">Lepthercus dregei</emphasis>
<bibRefCitation author="Purcell, W. F." box="[303,441,1853,1878]" pageId="38" pageNumber="299" pagination="348 - 382" refId="ref39288" refString="Purcell, W. F. (1902) New South African trap-door spiders of the family Ctenizidae in the collection of the South African Museum. Transactions of the South African Philosophical Society, 11, 348 - 382. https: // doi. org / 10.1080 / 21560382.1900.9525972" type="journal article" year="1902">Purcell, 1902</bibRefCitation>
</taxonomicName>
,
<emphasis bold="true" box="[452,482,1847,1876]" pageId="38" pageNumber="299">O</emphasis>
<taxonomicName authority="Ríos-Tamayo &amp; Lyle, 2020" authorityName="Ríos-Tamayo &amp; Lyle" authorityYear="2020" box="[490,734,1854,1878]" class="Arachnida" family="Nemesiidae" genus="Lepthercus" kingdom="Animalia" order="Araneae" pageId="38" pageNumber="299" phylum="Arthropoda" rank="species" species="engelbrechti" status="sp. nov.">
<emphasis box="[490,734,1854,1878]" italics="true" pageId="38" pageNumber="299">Lepthercus engelbrechti</emphasis>
</taxonomicName>
<emphasis bold="true" box="[741,823,1854,1878]" pageId="38" pageNumber="299">
<taxonomicNameLabel box="[741,823,1854,1878]" pageId="38" pageNumber="299" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, A
<taxonomicName authority="Ríos-Tamayo &amp; Lyle, 2020" authorityName="Ríos-Tamayo &amp; Lyle" authorityYear="2020" box="[874,1061,1854,1878]" class="Arachnida" family="Nemesiidae" genus="Lepthercus" kingdom="Animalia" order="Araneae" pageId="38" pageNumber="299" phylum="Arthropoda" rank="species" species="filmeri" status="sp. nov.">
<emphasis box="[874,1061,1854,1878]" italics="true" pageId="38" pageNumber="299">Lepthercus filmeri</emphasis>
</taxonomicName>
<emphasis bold="true" box="[1067,1149,1854,1878]" pageId="38" pageNumber="299">
<taxonomicNameLabel box="[1067,1149,1854,1878]" pageId="38" pageNumber="299" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
,
<emphasis bold="true" box="[1162,1189,1846,1876]" pageId="38" pageNumber="299">*</emphasis>
<taxonomicName authority="Ríos-Tamayo &amp; Lyle, 2020" authorityName="Ríos-Tamayo &amp; Lyle" authorityYear="2020" box="[1197,1399,1854,1878]" class="Arachnida" family="Nemesiidae" genus="Lepthercus" kingdom="Animalia" order="Araneae" pageId="38" pageNumber="299" phylum="Arthropoda" rank="species" species="haddadi" status="sp. nov.">
<emphasis box="[1197,1399,1854,1878]" italics="true" pageId="38" pageNumber="299">Lepthercus haddadi</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="38" pageNumber="299">
<taxonomicNameLabel pageId="38" pageNumber="299" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
,
<emphasis bold="true" box="[208,235,1883,1913]" pageId="38" pageNumber="299">D</emphasis>
<taxonomicName authority="Ríos-Tamayo &amp; Lyle, 2020" authorityName="Ríos-Tamayo &amp; Lyle" authorityYear="2020" box="[242,494,1890,1914]" class="Arachnida" family="Nemesiidae" genus="Lepthercus" kingdom="Animalia" order="Araneae" pageId="38" pageNumber="299" phylum="Arthropoda" rank="species" species="kwazuluensis" status="sp. nov.">
<emphasis box="[242,494,1890,1914]" italics="true" pageId="38" pageNumber="299">Lepthercus kwazuluensis</emphasis>
</taxonomicName>
<emphasis bold="true" box="[501,581,1890,1914]" pageId="38" pageNumber="299">
<taxonomicNameLabel box="[501,581,1890,1914]" pageId="38" pageNumber="299" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, ■
<taxonomicName authority="Ríos-Tamayo &amp; Lyle, 2020" authorityName="Ríos-Tamayo &amp; Lyle" authorityYear="2020" box="[619,837,1890,1914]" class="Arachnida" family="Nemesiidae" genus="Lepthercus" kingdom="Animalia" order="Araneae" pageId="38" pageNumber="299" phylum="Arthropoda" rank="species" species="lawrencei" status="sp. nov.">
<emphasis box="[619,837,1890,1914]" italics="true" pageId="38" pageNumber="299">Lepthercus lawrencei</emphasis>
</taxonomicName>
<emphasis bold="true" box="[843,924,1890,1914]" pageId="38" pageNumber="299">
<taxonomicNameLabel box="[843,924,1890,1914]" pageId="38" pageNumber="299" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, v
<taxonomicName authority="Ríos-Tamayo &amp; Lyle, 2020" authorityName="Ríos-Tamayo &amp; Lyle" authorityYear="2020" box="[974,1186,1890,1914]" class="Arachnida" family="Nemesiidae" genus="Lepthercus" kingdom="Animalia" order="Araneae" pageId="38" pageNumber="299" phylum="Arthropoda" rank="species" species="mandelai" status="sp. nov.">
<emphasis box="[974,1186,1890,1914]" italics="true" pageId="38" pageNumber="299">Lepthercus mandelai</emphasis>
</taxonomicName>
<emphasis bold="true" box="[1192,1273,1890,1914]" pageId="38" pageNumber="299">
<taxonomicNameLabel box="[1192,1273,1890,1914]" pageId="38" pageNumber="299" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, ▼
<taxonomicName authority="Hewitt, 1917" authorityName="Hewitt" authorityYear="1917" class="Arachnida" family="Nemesiidae" genus="Lepthercus" kingdom="Animalia" order="Araneae" pageId="38" pageNumber="299" phylum="Arthropoda" rank="species" species="rattrayi">
<emphasis italics="true" pageId="38" pageNumber="299">Lepthercus rattrayi</emphasis>
<bibRefCitation author="Hewitt, J." box="[235,370,1926,1951]" pageId="38" pageNumber="299" pagination="687 - 711" refId="ref38568" refString="Hewitt, J. (1917) Descriptions of new South African Arachnida. Annals of the Natal Museum, 3, 687 - 711." type="journal article" year="1917">Hewitt, 1917</bibRefCitation>
</taxonomicName>
, ▲
<taxonomicName authority="Ríos-Tamayo &amp; Lyle, 2020" authorityName="Ríos-Tamayo &amp; Lyle" authorityYear="2020" box="[417,596,1927,1951]" class="Arachnida" family="Nemesiidae" genus="Lepthercus" kingdom="Animalia" order="Araneae" pageId="38" pageNumber="299" phylum="Arthropoda" rank="species" species="sofiae" status="sp. nov.">
<emphasis box="[417,596,1927,1951]" italics="true" pageId="38" pageNumber="299">Lepthercus sofiae</emphasis>
</taxonomicName>
<emphasis bold="true" box="[601,682,1927,1951]" pageId="38" pageNumber="299">
<taxonomicNameLabel box="[601,682,1927,1951]" pageId="38" pageNumber="299" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Solid black line indicates the two species overlapped.
</paragraph>
</caption>
<paragraph blockId="39.[151,1437,153,755]" pageId="39" pageNumber="300">
<emphasis box="[199,327,153,179]" italics="true" pageId="39" pageNumber="300">Chaetotaxy</emphasis>
: Leg I: femur, 1-1-1-1 d (centrals), 1-1 PA; patella, 1 PA; tibia, 1-1 P, 2-3-1 V, a short and robust cuticular spur, with an apical megaspine with a strong curved spine at the base of the spur (
<figureCitation box="[1192,1288,189,215]" captionStart="FIGURE" captionStartId="37.[151,251,1673,1698]" captionTargetBox="[151,1436,669,1648]" captionTargetId="figure@37.[151,1436,669,1648]" captionTargetPageId="37" captionText="FIGURE. 23. Lepthercus sofiae sp. nov. (male holotype, NCA 97/395). A habitus, dorsal view; B sternum, ventral view; C chelicerae, arrow indicates the pallid intercheliceral tumesence, prolateral view; D right pedipalp, arrow indicates the palpal tibia base dorsally very convex and the development of the anterior side of the retrolateral side of the ventral excavation, retrolateral view; E right pedipalp, arrows indicates the few proventral setae and the development of the anterior side of the retrolateral side of the ventral excavation, prolateral view; FH copulatory bulb, F retrolateral view, G ventral view, H prolateral view, arrows indicate the two keels in the embolus; I tibia-metatarsus I, showing the curved metatarsus, arrow indicate the numerous ventral short and strong setae, prolateral view; J tibia I, arrow indicates the strong curved spine near of the base of the megaspine, ventral view." figureDoi="http://doi.org/10.5281/zenodo.3765517" httpUri="https://zenodo.org/record/3765517/files/figure.png" pageId="39" pageNumber="300">Fig. 23J</figureCitation>
); metatarsus, 1-1-2 V, presence of numerous short and strong setae (1:2A,
<figureCitation box="[828,920,225,251]" captionStart="FIGURE" captionStartId="37.[151,251,1673,1698]" captionTargetBox="[151,1436,669,1648]" captionTargetId="figure@37.[151,1436,669,1648]" captionTargetPageId="37" captionText="FIGURE. 23. Lepthercus sofiae sp. nov. (male holotype, NCA 97/395). A habitus, dorsal view; B sternum, ventral view; C chelicerae, arrow indicates the pallid intercheliceral tumesence, prolateral view; D right pedipalp, arrow indicates the palpal tibia base dorsally very convex and the development of the anterior side of the retrolateral side of the ventral excavation, retrolateral view; E right pedipalp, arrows indicates the few proventral setae and the development of the anterior side of the retrolateral side of the ventral excavation, prolateral view; FH copulatory bulb, F retrolateral view, G ventral view, H prolateral view, arrows indicate the two keels in the embolus; I tibia-metatarsus I, showing the curved metatarsus, arrow indicate the numerous ventral short and strong setae, prolateral view; J tibia I, arrow indicates the strong curved spine near of the base of the megaspine, ventral view." figureDoi="http://doi.org/10.5281/zenodo.3765517" httpUri="https://zenodo.org/record/3765517/files/figure.png" pageId="39" pageNumber="300">Fig. 23I</figureCitation>
); tarsus, 0. Leg II: femur, 1-1-1-1 d, 1-1-1 PA; patella, 1-1 PA; tibia, 1-1 P, 2-2-3 V; metatarsus, 1 P, 2-2-2 V; tarsus, 0. Leg III: femur, 1-1-1-1 d (thick), 1-1-1 D-P, 1-1-1-1-1 D-R; patella, 1-1 P, 1 R; tibia, 1-1 P, 1-1 D, 1-1 R, 2-2-3 V; metatarsus, 1-1-1-1 P, 1-1-1 D-P, 1-1-1-1 D-R, 2-3-3 V; tarsus, 0. Leg IV: femur, 1-1-1-1-1 d, 1 P A, 1-1 D-R A; patella, 1 R; tibia, 1-1 P, 1-1 R, 2-2-2 V; metatarsus, 1-1-1 P, 1-1-1-1 D-P, 1-1-1-1 D-R, 2-3-3 V; tarsus, 0. Palp (
<figureCitation box="[808,942,369,395]" captionStart="FIGURE" captionStartId="37.[151,251,1673,1698]" captionTargetBox="[151,1436,669,1648]" captionTargetId="figure@37.[151,1436,669,1648]" captionTargetPageId="37" captionText="FIGURE. 23. Lepthercus sofiae sp. nov. (male holotype, NCA 97/395). A habitus, dorsal view; B sternum, ventral view; C chelicerae, arrow indicates the pallid intercheliceral tumesence, prolateral view; D right pedipalp, arrow indicates the palpal tibia base dorsally very convex and the development of the anterior side of the retrolateral side of the ventral excavation, retrolateral view; E right pedipalp, arrows indicates the few proventral setae and the development of the anterior side of the retrolateral side of the ventral excavation, prolateral view; FH copulatory bulb, F retrolateral view, G ventral view, H prolateral view, arrows indicate the two keels in the embolus; I tibia-metatarsus I, showing the curved metatarsus, arrow indicate the numerous ventral short and strong setae, prolateral view; J tibia I, arrow indicates the strong curved spine near of the base of the megaspine, ventral view." figureDoi="http://doi.org/10.5281/zenodo.3765517" httpUri="https://zenodo.org/record/3765517/files/figure.png" pageId="39" pageNumber="300">Fig. 23D, E</figureCitation>
): femur, 1-1-1-1 d, 1 PA; patella, 1 PA; tibia, 2 P, (width/length: 0.79); tarsus, 0.
</paragraph>
<paragraph blockId="39.[151,1437,153,755]" pageId="39" pageNumber="300">
<emphasis box="[199,302,441,467]" italics="true" pageId="39" pageNumber="300">Scopulae</emphasis>
: Metatarsi: I, sparse, uniformly distributed, more abundant around the strong setae; II, sparse (3:4 A), more abundant in the anterior part; IIIIV, absent. Tarsi: I-II, dense, uniformly distributed throughout the segment; III, dense, uniformly distributed and divided by a narrow band of setae; IV, sparse, uniformly distributed and divided by a wide band of setae.
<emphasis box="[422,578,549,575]" italics="true" pageId="39" pageNumber="300">Trichobothria</emphasis>
: Tibiae: I 9-10; II 11-12; III 10-11; IV 11-10. Metatarsi: I (3)1(1)1(2)1(5)1; II (4)1(2)1(5)1; III (5)1(0)1(2)1(5)1; IV (6)1(2)1(2)1(5)1. Tarsi: I 13; II 12; III 12; IV 13.
<emphasis box="[1116,1296,585,611]" italics="true" pageId="39" pageNumber="300">Preening combs</emphasis>
: metatarsus: II, 4 PV; III, 4 PV4 RV; IV, 3 PV5 RV.
<emphasis box="[623,808,621,647]" italics="true" pageId="39" pageNumber="300">Copulatory bulb</emphasis>
: long, with a curved embolus tapering gradually with 2 keels (
<figureCitation box="[226,367,657,683]" captionStart="FIGURE" captionStartId="37.[151,251,1673,1698]" captionTargetBox="[151,1436,669,1648]" captionTargetId="figure@37.[151,1436,669,1648]" captionTargetPageId="37" captionText="FIGURE. 23. Lepthercus sofiae sp. nov. (male holotype, NCA 97/395). A habitus, dorsal view; B sternum, ventral view; C chelicerae, arrow indicates the pallid intercheliceral tumesence, prolateral view; D right pedipalp, arrow indicates the palpal tibia base dorsally very convex and the development of the anterior side of the retrolateral side of the ventral excavation, retrolateral view; E right pedipalp, arrows indicates the few proventral setae and the development of the anterior side of the retrolateral side of the ventral excavation, prolateral view; FH copulatory bulb, F retrolateral view, G ventral view, H prolateral view, arrows indicate the two keels in the embolus; I tibia-metatarsus I, showing the curved metatarsus, arrow indicate the numerous ventral short and strong setae, prolateral view; J tibia I, arrow indicates the strong curved spine near of the base of the megaspine, ventral view." figureDoi="http://doi.org/10.5281/zenodo.3765517" httpUri="https://zenodo.org/record/3765517/files/figure.png" pageId="39" pageNumber="300">Fig. 23G, H</figureCitation>
), spermatic duct curved.
<emphasis box="[656,721,657,683]" italics="true" pageId="39" pageNumber="300">Color</emphasis>
: Overall orangeyellowish (chelicerae darker). Abdomen light brown, with spots, spinnerets dorsally light yellowish.
</paragraph>
</subSubSection>
<subSubSection box="[199,890,728,755]" pageId="39" pageNumber="300" type="distribution">
<paragraph blockId="39.[151,1437,153,755]" box="[199,890,728,755]" pageId="39" pageNumber="300">
<emphasis bold="true" box="[199,353,728,755]" pageId="39" pageNumber="300">Distribution.</emphasis>
<collectingRegion box="[359,624,729,755]" country="South Africa" name="Western Cape" pageId="39" pageNumber="300">Western Cape Province</collectingRegion>
,
<collectingCountry box="[636,778,729,755]" name="South Africa" pageId="39" pageNumber="300">South Africa</collectingCountry>
(
<figureCitation box="[793,878,729,755]" captionStart="FIGURE" captionStartId="38.[151,251,1816,1841]" captionTargetBox="[185,1405,888,1784]" captionTargetId="figure@38.[176,1411,885,1790]" captionTargetPageId="38" captionText="FIGURE. 24. Distribution map of Lepthercus spp: ● Lepthercus confusus sp. nov., ○ Lepthercus dippenaarae sp. nov., Ɨ Lepthercus dregei Purcell, 1902, O Lepthercus engelbrechti sp. nov., A Lepthercus filmeri sp. nov., * Lepthercus haddadi sp. nov., D Lepthercus kwazuluensis sp. nov., ■ Lepthercus lawrencei sp. nov., v Lepthercus mandelai sp. nov., ▼ Lepthercus rattrayi Hewitt, 1917, ▲ Lepthercus sofiae sp. nov. Solid black line indicates the two species overlapped." figureDoi="http://doi.org/10.5281/zenodo.3765519" httpUri="https://zenodo.org/record/3765519/files/figure.png" pageId="39" pageNumber="300">Fig. 24</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>