425 lines
54 KiB
XML
425 lines
54 KiB
XML
<document ID-DOI="http://doi.org/10.5281/zenodo.3803486" ID-GBIF-Dataset="fe76f860-39b1-4520-a13c-d01a47dfc130" ID-GBIF-Taxon="163738938" ID-ISSN="1175-5334" ID-Zenodo-Dep="3803486" ID-ZooBank="urn:lsid:zoobank.org:pub:BA570D47-FE77-446C-B000-9208C9A6E957" checkinTime="1587753273724" checkinUser="carolina" docAuthor="Ríos-Tamayo, Duniesky & Lyle, Robin" docDate="2020" docId="03F11B20FFC5FF968EE83B1BFBDDD5D2" docLanguage="en" docName="zootaxa.4766.2.2.pdf.imf" docOrigin="Zootaxa 4766 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Lepthercus engelbrechti Ríos-Tamayo & Lyle 2020, sp. nov." docType="treatment" docVersion="5" lastPageId="30" lastPageNumber="291" masterDocId="FFC86358FFDEFF888E7F3E26FFD2D232" masterDocTitle="The South African genus Lepthercus Purcell, 1902 (Araneae: Mygalomorphae) phylogeny and taxonomy" masterLastPageNumber="305" masterPageNumber="261" pageId="27" pageNumber="288" updateTime="1636538462881" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>The South African genus Lepthercus Purcell, 1902 (Araneae: Mygalomorphae) phylogeny and taxonomy</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Ríos-Tamayo, Duniesky</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Lyle, Robin</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part>
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||
<mods:date>2020</mods:date>
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||
<mods:detail type="pubDate">
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||
<mods:number>2020-04-20</mods:number>
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||
</mods:detail>
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||
<mods:detail type="volume">
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<mods:number>4766</mods:number>
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||
</mods:detail>
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||
<mods:detail type="issue">
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<mods:number>2</mods:number>
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||
</mods:detail>
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||
<mods:extent unit="page">
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<mods:start>261</mods:start>
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<mods:end>305</mods:end>
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</mods:extent>
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</mods:part>
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||
</mods:relatedItem>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.11646/zootaxa.4766.2.2</mods:identifier>
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<mods:identifier type="GBIF-Dataset">fe76f860-39b1-4520-a13c-d01a47dfc130</mods:identifier>
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<mods:identifier type="ISSN">1175-5334</mods:identifier>
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<mods:identifier type="Zenodo-Dep">3765467</mods:identifier>
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<mods:identifier type="ZooBank">urn:lsid:zoobank.org:pub:BA570D47-FE77-446C-B000-9208C9A6E957</mods:identifier>
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</mods:mods>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.3803486" ID-GBIF-Taxon="163738938" ID-Zenodo-Dep="3803486" LSID="urn:lsid:plazi:treatment:03F11B20FFC5FF968EE83B1BFBDDD5D2" httpUri="http://treatment.plazi.org/id/03F11B20FFC5FF968EE83B1BFBDDD5D2" lastPageId="30" lastPageNumber="291" pageId="27" pageNumber="288">
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<subSubSection box="[151,545,1341,1368]" pageId="27" pageNumber="288" type="nomenclature">
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<paragraph blockId="27.[151,1130,1341,1438]" box="[151,545,1341,1368]" pageId="27" pageNumber="288">
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<heading bold="true" box="[151,545,1341,1368]" fontSize="10" level="2" pageId="27" pageNumber="288" reason="2">
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<emphasis bold="true" box="[151,545,1341,1368]" pageId="27" pageNumber="288">
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<taxonomicName authority="Ríos-Tamayo & Lyle, 2020" authorityName="Ríos-Tamayo & Lyle" authorityYear="2020" box="[151,444,1341,1368]" class="Arachnida" family="Nemesiidae" genus="Lepthercus" kingdom="Animalia" order="Araneae" pageId="27" pageNumber="288" phylum="Arthropoda" rank="species" species="engelbrechti" status="sp. nov.">
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<emphasis bold="true" box="[151,444,1341,1368]" italics="true" pageId="27" pageNumber="288">Lepthercus engelbrechti</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[451,545,1341,1367]" pageId="27" pageNumber="288" rank="species">sp. nov.</taxonomicNameLabel>
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||
</emphasis>
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||
</heading>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection pageId="27" pageNumber="288" type="description">
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||
<paragraph blockId="27.[151,1130,1341,1438]" pageId="27" pageNumber="288">
|
||
(
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||
<figureCitation box="[159,282,1377,1403]" captionStart="FIGURE" captionStartId="7.[151,251,1647,1672]" captionTargetBox="[151,1436,381,1622]" captionTargetId="figure@7.[151,1436,381,1622]" captionTargetPageId="7" captionText="FIGURE. 2. Preening comb. A Lepthercus haddadi sp. nov. (female paratype, NCA 2007/3847). Maxillary cuspules (males). B L. dippenaarae. C L. kwazuluensis. Serrula. D L. mandelai sp. nov. (female, NCA 2014/415A); E L. filmeri sp. nov. (female, NCA 91/1437); F L. haddadi sp. nov. (male, NCA 2007/1486); G L. haddadi sp. nov. (female, NCA 2007/3847); H L. engelbrechti sp. nov. (female, NCA 2018/385); I L. engelbrechti sp. nov. (male, NCA 2018/384); J L. dregei Purcell, 1902 (female, NCA 2012/2333); K L. sofiae sp. nov. (male, NCA 97/395)." figureDoi="http://doi.org/10.5281/zenodo.3765471" httpUri="https://zenodo.org/record/3765471/files/figure.png" pageId="27" pageNumber="288">Figs. 2H, I</figureCitation>
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||
,
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<figureCitation box="[294,378,1377,1403]" captionStart="FIGURE" captionStartId="28.[151,251,585,610]" captionTargetBox="[173,1422,228,547]" captionTargetId="figure@28.[151,1437,181,559]" captionTargetPageId="28" captionText="FIGURE. 16. Lepthercus engelbrechti sp. nov., in vivo. A male; B female. (photos courtesy Ian Engelbrecht)" figureDoi="http://doi.org/10.5281/zenodo.3765501" httpUri="https://zenodo.org/record/3765501/files/figure.png" pageId="27" pageNumber="288">16A, B</figureCitation>
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,
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<figureCitation box="[389,415,1377,1403]" captionStart="FIGURE" captionStartId="28.[151,251,1761,1786]" captionTargetBox="[151,1436,631,1735]" captionTargetId="figure@28.[151,1436,631,1735]" captionTargetPageId="28" captionText="FIGURE. 17. Lepthercus engelbrechti sp. nov. (male holotype, NCA 2018/384). A habitus, dorsal view; B sternum, ventral view; C chelicerae, prolateral view, arrow indicates the intercheliceral tumesence; D right pedipalp, arrows indicates the palpal tibia base dorsally very convex and a development of the anterior part of the retrolateral side of ventral excavation; E right pedipalp, arrow indicates the development of the anterior part of the retrolateral side of ventral excavation, prolateral view; F–H copulatory bulb, F retrolateral view, G ventral view, H prolateral view; I tibia-metatarsus I, arrow indicates the curved metatarsus and the apical patch of numerous black blunt spinules, prolateral view; J tibia I, arrow indicates the strong curved spine near of the base of the megaspine and the prolateral strong spine, ventral view." figureDoi="http://doi.org/10.5281/zenodo.3765503" httpUri="https://zenodo.org/record/3765503/files/figure.png" pageId="27" pageNumber="288">17</figureCitation>
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A–K, 18A–C, 24) http://zoobank.org/
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||
<uri box="[362,1130,1414,1438]" pageId="27" pageNumber="288">
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||
urn:lsid:zoobank.org:act:
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||
<uuid box="[624,1130,1414,1438]" pageId="27" pageNumber="288">5CC73DE1-28E2-4FA2-9784-BA1C58B0F910</uuid>
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||
</uri>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection pageId="27" pageNumber="288" type="materials_examined">
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||
<paragraph blockId="27.[151,1437,1484,1979]" pageId="27" pageNumber="288">
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||
<emphasis bold="true" box="[151,557,1484,1511]" pageId="27" pageNumber="288">Material examined. Type material</emphasis>
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||
:
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<materialsCitation ID-GBIF-Occurrence="2608670262" collectingDate="2013-05-19" collectionCode="NCA" collectorName="Engelbrecht, I. & Correia, M." country="South Africa" latitude="-33.922142" location="Swellendam District" longLatPrecision="1" longitude="20.06775" pageId="27" pageNumber="288" specimenCode="NCA 2018" specimenCount="1" specimenCount-male="1" stateProvince="Western Cape Province" typeStatus="holotype">
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||
<typeStatus box="[572,676,1485,1511]" pageId="27" pageNumber="288">Holotype</typeStatus>
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||
<specimenCount box="[683,704,1485,1511]" pageId="27" pageNumber="288" type="male">♂</specimenCount>
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||
.
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||
<collectingCountry box="[718,871,1485,1511]" name="South Africa" pageId="27" pageNumber="288">
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||
<subScript attach="left" box="[733,787,1492,1509]" fontSize="7" pageId="27" pageNumber="288">SOUTH</subScript>
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||
<subScript attach="left" box="[813,871,1492,1509]" fontSize="7" pageId="27" pageNumber="288">AFRICA</subScript>
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||
</collectingCountry>
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||
:
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||
<collectingRegion box="[885,1149,1485,1511]" country="South Africa" name="Western Cape" pageId="27" pageNumber="288">Western Cape Province</collectingRegion>
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||
,
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||
<location LSID="urn:lsid:plazi:treatment:03F11B20FFC5FF968EE83B1BFBDDD5D2:8E87FCEDFFC5FF938AF63BEBFAA0D7D5" box="[1161,1394,1485,1511]" country="South Africa" latitude="-33.922142" longLatPrecision="1" longitude="20.06775" name="Swellendam District" pageId="27" pageNumber="288" stateProvince="Western Cape Province">Swellendam District</location>
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||
; ca.
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||
<locationDeviation box="[151,399,1521,1547]" location="Bonnievale" pageId="27" pageNumber="288">
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||
<quantity box="[151,201,1521,1547]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="27" pageNumber="288" unit="km" value="3.0">3km</quantity>
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||
W of Bonnievale
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||
</locationDeviation>
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||
; S slope of hill on S side of tar road (
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||
<geoCoordinate box="[804,972,1520,1547]" degrees="33" direction="south" minutes="55" orientation="latitude" pageId="27" pageNumber="288" precision="1" seconds="19.70" value="-33.922142">33°55’19.70”S</geoCoordinate>
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||
<geoCoordinate box="[978,1122,1521,1547]" degrees="20" direction="east" minutes="4" orientation="longitude" pageId="27" pageNumber="288" precision="1" seconds="3.90" value="20.06775">20°4’3.90”E</geoCoordinate>
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||
),
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||
<date box="[1141,1254,1521,1547]" pageId="27" pageNumber="288" value="2013-05-19">
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<collectingDate box="[1141,1254,1521,1547]" pageId="27" pageNumber="288" value="2013-05-19">19.v.2013</collectingDate>
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||
</date>
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||
,
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||
<collectorName box="[1265,1430,1521,1547]" pageId="27" pageNumber="288">Engelbrecht, I.</collectorName>
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||
,
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||
<collectorName box="[151,281,1557,1583]" pageId="27" pageNumber="288">Correia, M.</collectorName>
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||
<emphasis box="[288,329,1557,1583]" italics="true" pageId="27" pageNumber="288">leg.</emphasis>
|
||
, (
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||
<specimenCode box="[351,475,1557,1583]" collectionCode="NCA" pageId="27" pageNumber="288">NCA 2018</specimenCode>
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||
/384)
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||
</materialsCitation>
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||
. –
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||
<typeStatus box="[567,669,1557,1583]" pageId="27" pageNumber="288">Paratype</typeStatus>
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||
.
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||
<collectingCountry box="[679,833,1557,1583]" name="South Africa" pageId="27" pageNumber="288">
|
||
<subScript attach="left" box="[695,749,1564,1581]" fontSize="7" pageId="27" pageNumber="288">SOUTH</subScript>
|
||
<subScript attach="left" box="[775,833,1564,1581]" fontSize="7" pageId="27" pageNumber="288">AFRICA</subScript>
|
||
</collectingCountry>
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||
: same data as
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||
<typeStatus box="[996,1097,1557,1583]" pageId="27" pageNumber="288">holotype</typeStatus>
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||
,
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||
<specimenCount box="[1108,1150,1557,1583]" pageId="27" pageNumber="288" type="female">1 ♀</specimenCount>
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||
(
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||
<collectionCode box="[1164,1224,1557,1583]" pageId="27" pageNumber="288">NCA</collectionCode>
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||
2018/385).
|
||
</paragraph>
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||
<paragraph blockId="27.[151,1437,1484,1979]" pageId="27" pageNumber="288">
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||
<emphasis bold="true" box="[199,511,1592,1619]" pageId="27" pageNumber="288">Other material examined</emphasis>
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||
:
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||
<collectingCountry box="[532,691,1593,1619]" name="South Africa" pageId="27" pageNumber="288">
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||
<subScript attach="left" box="[547,601,1600,1617]" fontSize="7" pageId="27" pageNumber="288">SOUTH</subScript>
|
||
<subScript attach="left" box="[633,691,1600,1617]" fontSize="7" pageId="27" pageNumber="288">AFRICA</subScript>
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||
</collectingCountry>
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||
:
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||
<collectingRegion box="[711,988,1593,1619]" country="South Africa" name="Western Cape" pageId="27" pageNumber="288">Western Cape Province</collectingRegion>
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||
:
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<collectorName box="[1004,1145,1593,1619]" pageId="27" pageNumber="288">Swellendam</collectorName>
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||
,
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||
<collectorName box="[1162,1325,1593,1619]" pageId="27" pageNumber="288">Bontebok N.P</collectorName>
|
||
(
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||
<geoCoordinate box="[1346,1436,1592,1619]" degrees="34" direction="south" minutes="04" orientation="latitude" pageId="27" pageNumber="288" precision="925" value="-34.066666">34°04′S</geoCoordinate>
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||
<geoCoordinate box="[151,246,1628,1655]" degrees="20" direction="east" minutes="27" orientation="longitude" pageId="27" pageNumber="288" precision="925" value="20.45">20°27′E</geoCoordinate>
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||
),
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||
<date box="[264,378,1629,1655]" pageId="27" pageNumber="288" value="1987-10-27">
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<collectingDate box="[264,378,1629,1655]" pageId="27" pageNumber="288" value="1987-10-27">27.x.1987</collectingDate>
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||
</date>
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||
,
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||
<collectorName box="[389,587,1629,1655]" pageId="27" pageNumber="288">Entomology Staff</collectorName>
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||
<emphasis box="[593,634,1629,1655]" italics="true" pageId="27" pageNumber="288">leg.</emphasis>
|
||
,
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||
<specimenCount box="[647,692,1629,1655]" pageId="27" pageNumber="288" type="male">1 ♂</specimenCount>
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||
<specimenCount box="[691,708,1629,1655]" pageId="27" pageNumber="288" type="female">♀</specimenCount>
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||
(
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||
<specimenCode box="[722,887,1629,1655]" collectionCode="NMBA" country="Switzerland" httpUri="http://grbio.org/cool/js1h-4156" name="Naturhistorisches Museum, Basel" pageId="27" pageNumber="288">NMBA 17794</specimenCode>
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||
);
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||
<location LSID="urn:lsid:plazi:treatment:03F11B20FFC5FF968EE83B1BFBDDD5D2:8E87FCEDFFC5FF938DF6387BFBA3D445" box="[905,1137,1629,1655]" country="South Africa" latitude="-33.922142" longLatPrecision="1" longitude="20.06775" name="Swellendam District" pageId="27" pageNumber="288" stateProvince="Western Cape">Swellendam District</location>
|
||
; ca.
|
||
<locationDeviation box="[1185,1432,1629,1655]" location="Bonnievale" pageId="27" pageNumber="288">
|
||
<quantity box="[1185,1234,1629,1655]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="27" pageNumber="288" unit="km" value="3.0">3km</quantity>
|
||
W of Bonnievale
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||
</locationDeviation>
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; S slope of hill on S side of tar road (
|
||
<geoCoordinate box="[543,712,1664,1691]" degrees="33" direction="south" minutes="55" orientation="latitude" pageId="27" pageNumber="288" precision="1" seconds="19.70" value="-33.922142">33°55’19.70”S</geoCoordinate>
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||
<geoCoordinate box="[718,862,1665,1691]" degrees="20" direction="east" minutes="4" orientation="longitude" pageId="27" pageNumber="288" precision="1" seconds="3.90" value="20.06775">20°4’3.90”E</geoCoordinate>
|
||
),
|
||
<date box="[881,993,1665,1691]" pageId="27" pageNumber="288" value="2013-05-19">
|
||
<collectingDate box="[881,993,1665,1691]" pageId="27" pageNumber="288" value="2013-05-19">19.v.2013</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName box="[1004,1167,1665,1691]" pageId="27" pageNumber="288">Engelbrecht, I.</collectorName>
|
||
<emphasis box="[1174,1215,1665,1691]" italics="true" pageId="27" pageNumber="288">leg.</emphasis>
|
||
,
|
||
<specimenCount box="[1227,1291,1665,1691]" pageId="27" pageNumber="288" type="male">3 ♂♂</specimenCount>
|
||
,
|
||
<specimenCount box="[1302,1364,1665,1691]" pageId="27" pageNumber="288" type="female">7 ♀♀</specimenCount>
|
||
(
|
||
<specimenCode collectionCode="NCA" pageId="27" pageNumber="288">NCA 2018</specimenCode>
|
||
/383).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="27" pageNumber="288" type="etymology">
|
||
<paragraph blockId="27.[151,1437,1484,1979]" pageId="27" pageNumber="288">
|
||
<emphasis bold="true" box="[199,334,1737,1763]" pageId="27" pageNumber="288">Etymology.</emphasis>
|
||
The specific epithet is a patronymic in honor of Dr Ian Engelbrecht who collected the
|
||
<typeStatus box="[1314,1362,1737,1763]" pageId="27" pageNumber="288">type</typeStatus>
|
||
specimens and for his contribution to arachnology in
|
||
<collectingCountry box="[691,837,1773,1799]" name="South Africa" pageId="27" pageNumber="288">South Africa</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="27" pageNumber="288" type="diagnosis">
|
||
<paragraph blockId="27.[151,1437,1484,1979]" pageId="27" pageNumber="288">
|
||
<emphasis bold="true" box="[199,322,1808,1835]" pageId="27" pageNumber="288">Diagnosis.</emphasis>
|
||
Males of
|
||
<taxonomicName authority="Ríos-Tamayo & Lyle, 2020" authorityName="Ríos-Tamayo & Lyle" authorityYear="2020" box="[435,604,1809,1835]" class="Arachnida" family="Nemesiidae" genus="Lepthercus" kingdom="Animalia" order="Araneae" pageId="27" pageNumber="288" phylum="Arthropoda" rank="species" species="engelbrechti" status="sp. nov.">
|
||
<emphasis box="[435,604,1809,1835]" italics="true" pageId="27" pageNumber="288">L. engelbrechti</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" box="[611,701,1809,1835]" pageId="27" pageNumber="288">
|
||
<taxonomicNameLabel box="[611,701,1809,1835]" pageId="27" pageNumber="288" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, can be recognized from all other species in “Group haddadi), by the presence of a curved megaspine with the apex flattened laterally (
|
||
<figureCitation box="[951,1072,1845,1871]" captionStart="FIGURE" captionStartId="28.[151,251,1761,1786]" captionTargetBox="[151,1436,631,1735]" captionTargetId="figure@28.[151,1436,631,1735]" captionTargetPageId="28" captionText="FIGURE. 17. Lepthercus engelbrechti sp. nov. (male holotype, NCA 2018/384). A habitus, dorsal view; B sternum, ventral view; C chelicerae, prolateral view, arrow indicates the intercheliceral tumesence; D right pedipalp, arrows indicates the palpal tibia base dorsally very convex and a development of the anterior part of the retrolateral side of ventral excavation; E right pedipalp, arrow indicates the development of the anterior part of the retrolateral side of ventral excavation, prolateral view; F–H copulatory bulb, F retrolateral view, G ventral view, H prolateral view; I tibia-metatarsus I, arrow indicates the curved metatarsus and the apical patch of numerous black blunt spinules, prolateral view; J tibia I, arrow indicates the strong curved spine near of the base of the megaspine and the prolateral strong spine, ventral view." figureDoi="http://doi.org/10.5281/zenodo.3765503" httpUri="https://zenodo.org/record/3765503/files/figure.png" pageId="27" pageNumber="288">Fig. 17I, J</figureCitation>
|
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); the presence of one prolateral strong and curved spine, near the spur (
|
||
<figureCitation box="[596,713,1881,1907]" captionStart="FIGURE" captionStartId="28.[151,251,1761,1786]" captionTargetBox="[151,1436,631,1735]" captionTargetId="figure@28.[151,1436,631,1735]" captionTargetPageId="28" captionText="FIGURE. 17. Lepthercus engelbrechti sp. nov. (male holotype, NCA 2018/384). A habitus, dorsal view; B sternum, ventral view; C chelicerae, prolateral view, arrow indicates the intercheliceral tumesence; D right pedipalp, arrows indicates the palpal tibia base dorsally very convex and a development of the anterior part of the retrolateral side of ventral excavation; E right pedipalp, arrow indicates the development of the anterior part of the retrolateral side of ventral excavation, prolateral view; F–H copulatory bulb, F retrolateral view, G ventral view, H prolateral view; I tibia-metatarsus I, arrow indicates the curved metatarsus and the apical patch of numerous black blunt spinules, prolateral view; J tibia I, arrow indicates the strong curved spine near of the base of the megaspine and the prolateral strong spine, ventral view." figureDoi="http://doi.org/10.5281/zenodo.3765503" httpUri="https://zenodo.org/record/3765503/files/figure.png" pageId="27" pageNumber="288">Fig. 17I, J</figureCitation>
|
||
) and by the presence of a shorter copulatory bulb (
|
||
<figureCitation box="[1289,1369,1881,1907]" captionStart="FIGURE" captionStartId="28.[151,251,1761,1786]" captionTargetBox="[151,1436,631,1735]" captionTargetId="figure@28.[151,1436,631,1735]" captionTargetPageId="28" captionText="FIGURE. 17. Lepthercus engelbrechti sp. nov. (male holotype, NCA 2018/384). A habitus, dorsal view; B sternum, ventral view; C chelicerae, prolateral view, arrow indicates the intercheliceral tumesence; D right pedipalp, arrows indicates the palpal tibia base dorsally very convex and a development of the anterior part of the retrolateral side of ventral excavation; E right pedipalp, arrow indicates the development of the anterior part of the retrolateral side of ventral excavation, prolateral view; F–H copulatory bulb, F retrolateral view, G ventral view, H prolateral view; I tibia-metatarsus I, arrow indicates the curved metatarsus and the apical patch of numerous black blunt spinules, prolateral view; J tibia I, arrow indicates the strong curved spine near of the base of the megaspine and the prolateral strong spine, ventral view." figureDoi="http://doi.org/10.5281/zenodo.3765503" httpUri="https://zenodo.org/record/3765503/files/figure.png" pageId="27" pageNumber="288">Fig. 17</figureCitation>
|
||
F–H). Females can be distinguished from
|
||
<taxonomicName authority="Ríos-Tamayo & Lyle, 2020" authorityName="Ríos-Tamayo & Lyle" authorityYear="2020" box="[551,673,1917,1943]" class="Arachnida" family="Nemesiidae" genus="Lepthercus" kingdom="Animalia" order="Araneae" pageId="27" pageNumber="288" phylum="Arthropoda" rank="species" species="haddadi" status="sp. nov.">
|
||
<emphasis box="[551,673,1917,1943]" italics="true" pageId="27" pageNumber="288">L. haddadi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" box="[680,770,1917,1943]" pageId="27" pageNumber="288">
|
||
<taxonomicNameLabel box="[680,770,1917,1943]" pageId="27" pageNumber="288" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, by the shape of their spermathecae with a short stalk (
|
||
<figureCitation captionStart="FIGURE" captionStartId="29.[151,251,1870,1895]" captionTargetBox="[164,1423,181,1844]" captionTargetId="figure@29.[164,1423,181,1844]" captionTargetPageId="29" captionText="FIGURE. 18. Lepthercus engelbrechti sp. nov. (female paratype, NCA 2018/385). A habitus, dorsal view; B sternum, ventral view; C chelicerae, prolateral view; D spermathecae" figureDoi="http://doi.org/10.5281/zenodo.3765507" httpUri="https://zenodo.org/record/3765507/files/figure.png" pageId="27" pageNumber="288">Fig. 18D</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.3765501" ID-Zenodo-Dep="3765501" box="[151,1264,585,610]" httpUri="https://zenodo.org/record/3765501/files/figure.png" pageId="28" pageNumber="289" startId="28.[151,251,585,610]" targetBox="[173,1422,228,547]" targetPageId="28">
|
||
<paragraph blockId="28.[151,1264,585,610]" box="[151,1264,585,610]" pageId="28" pageNumber="289">
|
||
<emphasis bold="true" box="[151,294,585,610]" pageId="28" pageNumber="289">FIGURE. 16.</emphasis>
|
||
<taxonomicName authority="Ríos-Tamayo & Lyle, 2020" authorityName="Ríos-Tamayo & Lyle" authorityYear="2020" box="[300,544,585,609]" class="Arachnida" family="Nemesiidae" genus="Lepthercus" kingdom="Animalia" order="Araneae" pageId="28" pageNumber="289" phylum="Arthropoda" rank="species" species="engelbrechti" status="sp. nov.">
|
||
<emphasis box="[300,544,585,609]" italics="true" pageId="28" pageNumber="289">Lepthercus engelbrechti</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" box="[550,631,585,609]" pageId="28" pageNumber="289">
|
||
<taxonomicNameLabel box="[550,631,585,609]" pageId="28" pageNumber="289" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
,
|
||
<emphasis box="[644,712,586,609]" italics="true" pageId="28" pageNumber="289">in vivo</emphasis>
|
||
.
|
||
<emphasis bold="true" box="[725,743,585,610]" pageId="28" pageNumber="289">A</emphasis>
|
||
male;
|
||
<emphasis bold="true" box="[812,829,585,609]" pageId="28" pageNumber="289">B</emphasis>
|
||
female. (photos courtesy Ian Engelbrecht)
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph blockId="28.[1264,1268,585,610]" box="[1264,1268,585,610]" pageId="28" pageNumber="289">.</paragraph>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.3765503" ID-Zenodo-Dep="3765503" httpUri="https://zenodo.org/record/3765503/files/figure.png" pageId="28" pageNumber="289" startId="28.[151,251,1761,1786]" targetBox="[151,1436,631,1735]" targetPageId="28">
|
||
<paragraph blockId="28.[151,1437,1761,2002]" pageId="28" pageNumber="289">
|
||
<emphasis bold="true" box="[151,296,1761,1786]" pageId="28" pageNumber="289">FIGURE. 17.</emphasis>
|
||
<taxonomicName authority="Ríos-Tamayo & Lyle, 2020" authorityName="Ríos-Tamayo & Lyle" authorityYear="2020" box="[304,550,1761,1785]" class="Arachnida" family="Nemesiidae" genus="Lepthercus" kingdom="Animalia" order="Araneae" pageId="28" pageNumber="289" phylum="Arthropoda" rank="species" species="engelbrechti" status="sp. nov.">
|
||
<emphasis box="[304,550,1761,1785]" italics="true" pageId="28" pageNumber="289">Lepthercus engelbrechti</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" box="[558,640,1762,1786]" pageId="28" pageNumber="289">
|
||
<taxonomicNameLabel box="[558,640,1762,1786]" pageId="28" pageNumber="289" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(male
|
||
<typeStatus box="[714,805,1761,1786]" pageId="28" pageNumber="289">holotype</typeStatus>
|
||
, NCA 2018/384).
|
||
<emphasis bold="true" box="[996,1014,1761,1786]" pageId="28" pageNumber="289">A</emphasis>
|
||
habitus, dorsal view;
|
||
<emphasis bold="true" box="[1245,1262,1762,1786]" pageId="28" pageNumber="289">B</emphasis>
|
||
sternum, ventral view;
|
||
<emphasis bold="true" box="[214,232,1797,1822]" pageId="28" pageNumber="289">C</emphasis>
|
||
chelicerae, prolateral view, arrow indicates the intercheliceral tumesence;
|
||
<emphasis bold="true" box="[987,1005,1798,1822]" pageId="28" pageNumber="289">D</emphasis>
|
||
right pedipalp, arrows indicates the palpal tibia base dorsally very convex and a development of the anterior part of the retrolateral side of ventral excavation;
|
||
<emphasis bold="true" box="[1363,1380,1834,1858]" pageId="28" pageNumber="289">E</emphasis>
|
||
right pedipalp, arrow indicates the development of the anterior part of the retrolateral side of ventral excavation, prolateral view;
|
||
<emphasis bold="true" box="[1388,1437,1870,1894]" pageId="28" pageNumber="289">F–H</emphasis>
|
||
copulatory bulb,
|
||
<emphasis bold="true" box="[323,338,1906,1930]" pageId="28" pageNumber="289">F</emphasis>
|
||
retrolateral view,
|
||
<emphasis bold="true" box="[522,542,1905,1930]" pageId="28" pageNumber="289">G</emphasis>
|
||
ventral view,
|
||
<emphasis bold="true" box="[684,704,1906,1930]" pageId="28" pageNumber="289">H</emphasis>
|
||
prolateral view;
|
||
<emphasis bold="true" box="[875,885,1906,1930]" pageId="28" pageNumber="289">I</emphasis>
|
||
tibia-metatarsus I, arrow indicates the curved metatarsus and the apical patch of numerous black blunt spinules, prolateral view;
|
||
<emphasis bold="true" box="[904,917,1942,1966]" pageId="28" pageNumber="289">J</emphasis>
|
||
tibia I, arrow indicates the strong curved spine near of the base of the megaspine and the prolateral strong spine, ventral view.
|
||
</paragraph>
|
||
</caption>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.3765507" ID-Zenodo-Dep="3765507" httpUri="https://zenodo.org/record/3765507/files/figure.png" pageId="29" pageNumber="290" startId="29.[151,251,1870,1895]" targetBox="[164,1423,181,1844]" targetPageId="29">
|
||
<paragraph blockId="29.[151,1436,1870,1931]" pageId="29" pageNumber="290">
|
||
<emphasis bold="true" box="[151,295,1870,1895]" pageId="29" pageNumber="290">FIGURE. 18.</emphasis>
|
||
<taxonomicName authority="Ríos-Tamayo & Lyle, 2020" authorityName="Ríos-Tamayo & Lyle" authorityYear="2020" box="[302,546,1871,1895]" class="Arachnida" family="Nemesiidae" genus="Lepthercus" kingdom="Animalia" order="Araneae" pageId="29" pageNumber="290" phylum="Arthropoda" rank="species" species="engelbrechti" status="sp. nov.">
|
||
<emphasis box="[302,546,1871,1895]" italics="true" pageId="29" pageNumber="290">Lepthercus engelbrechti</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" box="[553,634,1871,1895]" pageId="29" pageNumber="290">
|
||
<taxonomicNameLabel box="[553,634,1871,1895]" pageId="29" pageNumber="290" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(female
|
||
<typeStatus box="[726,816,1871,1895]" pageId="29" pageNumber="290">paratype</typeStatus>
|
||
, NCA 2018/385).
|
||
<emphasis bold="true" box="[1003,1021,1870,1895]" pageId="29" pageNumber="290">A</emphasis>
|
||
habitus, dorsal view;
|
||
<emphasis bold="true" box="[1247,1264,1871,1895]" pageId="29" pageNumber="290">B</emphasis>
|
||
sternum, ventral view;
|
||
<emphasis bold="true" box="[214,232,1906,1931]" pageId="29" pageNumber="290">C</emphasis>
|
||
chelicerae, prolateral view;
|
||
<emphasis bold="true" box="[520,538,1907,1931]" pageId="29" pageNumber="290">D</emphasis>
|
||
spermathecae
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph blockId="29.[151,1436,1870,1931]" box="[686,690,1906,1931]" pageId="29" pageNumber="290">.</paragraph>
|
||
<subSubSection pageId="30" pageNumber="291" type="description">
|
||
<paragraph blockId="30.[151,1437,152,2017]" pageId="30" pageNumber="291">
|
||
<emphasis bold="true" box="[199,345,152,179]" pageId="30" pageNumber="291">Description.</emphasis>
|
||
Male
|
||
<typeStatus box="[420,518,153,179]" pageId="30" pageNumber="291">holotype</typeStatus>
|
||
(NCA 2018/384,
|
||
<figureCitation box="[724,815,153,179]" captionStart="FIGURE" captionStartId="28.[151,251,1761,1786]" captionTargetBox="[151,1436,631,1735]" captionTargetId="figure@28.[151,1436,631,1735]" captionTargetPageId="28" captionText="FIGURE. 17. Lepthercus engelbrechti sp. nov. (male holotype, NCA 2018/384). A habitus, dorsal view; B sternum, ventral view; C chelicerae, prolateral view, arrow indicates the intercheliceral tumesence; D right pedipalp, arrows indicates the palpal tibia base dorsally very convex and a development of the anterior part of the retrolateral side of ventral excavation; E right pedipalp, arrow indicates the development of the anterior part of the retrolateral side of ventral excavation, prolateral view; F–H copulatory bulb, F retrolateral view, G ventral view, H prolateral view; I tibia-metatarsus I, arrow indicates the curved metatarsus and the apical patch of numerous black blunt spinules, prolateral view; J tibia I, arrow indicates the strong curved spine near of the base of the megaspine and the prolateral strong spine, ventral view." figureDoi="http://doi.org/10.5281/zenodo.3765503" httpUri="https://zenodo.org/record/3765503/files/figure.png" pageId="30" pageNumber="291">Figs. 17</figureCitation>
|
||
A–J). Total length 12.38.
|
||
<emphasis box="[1110,1220,153,179]" italics="true" pageId="30" pageNumber="291">Carapace</emphasis>
|
||
(
|
||
<figureCitation box="[1238,1345,153,179]" captionStart="FIGURE" captionStartId="28.[151,251,1761,1786]" captionTargetBox="[151,1436,631,1735]" captionTargetId="figure@28.[151,1436,631,1735]" captionTargetPageId="28" captionText="FIGURE. 17. Lepthercus engelbrechti sp. nov. (male holotype, NCA 2018/384). A habitus, dorsal view; B sternum, ventral view; C chelicerae, prolateral view, arrow indicates the intercheliceral tumesence; D right pedipalp, arrows indicates the palpal tibia base dorsally very convex and a development of the anterior part of the retrolateral side of ventral excavation; E right pedipalp, arrow indicates the development of the anterior part of the retrolateral side of ventral excavation, prolateral view; F–H copulatory bulb, F retrolateral view, G ventral view, H prolateral view; I tibia-metatarsus I, arrow indicates the curved metatarsus and the apical patch of numerous black blunt spinules, prolateral view; J tibia I, arrow indicates the strong curved spine near of the base of the megaspine and the prolateral strong spine, ventral view." figureDoi="http://doi.org/10.5281/zenodo.3765503" httpUri="https://zenodo.org/record/3765503/files/figure.png" pageId="30" pageNumber="291">Fig. 17A</figureCitation>
|
||
): length 5.52, width 4.05, with lateral dark marginal bristles, with a little pubescence.
|
||
<emphasis box="[1015,1195,188,214]" italics="true" pageId="30" pageNumber="291">Cephalic region</emphasis>
|
||
: length 3.29; clypeus almost absent, without bristles on the edge, 4 bristles in front of the OQ and 5 between the PME.
|
||
<emphasis box="[1285,1355,223,248]" italics="true" pageId="30" pageNumber="291">Fovea</emphasis>
|
||
: width 0.39, short, slightly recurved.
|
||
<emphasis box="[489,735,257,283]" italics="true" pageId="30" pageNumber="291">Ocular measurements</emphasis>
|
||
: AME 0.14, ALE 0.24, PME 0.18, PLE 0.22, OQ length 0.45, 0.75 width; AME–ALE 0.04, PME–PLE 0.00, AME–PME 0.05, ALE–PLE 0.03, AME–AME 0.09, OMP–OMP 0.29.
|
||
<emphasis box="[213,336,326,352]" italics="true" pageId="30" pageNumber="291">Chelicerae</emphasis>
|
||
: length 2.35, width 1.36, with dark dorsal–anterior bristles, intercheliceral tumescence small, wellmarked with setae (
|
||
<figureCitation box="[375,479,361,387]" captionStart="FIGURE" captionStartId="28.[151,251,1761,1786]" captionTargetBox="[151,1436,631,1735]" captionTargetId="figure@28.[151,1436,631,1735]" captionTargetPageId="28" captionText="FIGURE. 17. Lepthercus engelbrechti sp. nov. (male holotype, NCA 2018/384). A habitus, dorsal view; B sternum, ventral view; C chelicerae, prolateral view, arrow indicates the intercheliceral tumesence; D right pedipalp, arrows indicates the palpal tibia base dorsally very convex and a development of the anterior part of the retrolateral side of ventral excavation; E right pedipalp, arrow indicates the development of the anterior part of the retrolateral side of ventral excavation, prolateral view; F–H copulatory bulb, F retrolateral view, G ventral view, H prolateral view; I tibia-metatarsus I, arrow indicates the curved metatarsus and the apical patch of numerous black blunt spinules, prolateral view; J tibia I, arrow indicates the strong curved spine near of the base of the megaspine and the prolateral strong spine, ventral view." figureDoi="http://doi.org/10.5281/zenodo.3765503" httpUri="https://zenodo.org/record/3765503/files/figure.png" pageId="30" pageNumber="291">Fig. 17C</figureCitation>
|
||
). Fang groove with 8 promarginal teeth and 18 mesobasal denticles.
|
||
<emphasis box="[1264,1350,361,387]" italics="true" pageId="30" pageNumber="291">Labium</emphasis>
|
||
: length 0.41, width 0.72.
|
||
<emphasis box="[353,453,396,422]" italics="true" pageId="30" pageNumber="291">Maxillae</emphasis>
|
||
: length 1.64, width 0.71, with about 71 short cuspules on inner corner, prolateral face slightly curved, soft area small, with long uniformly distributed hairs, serrula present (
|
||
<figureCitation box="[1113,1192,430,456]" captionStart="FIGURE" captionStartId="7.[151,251,1647,1672]" captionTargetBox="[151,1436,381,1622]" captionTargetId="figure@7.[151,1436,381,1622]" captionTargetPageId="7" captionText="FIGURE. 2. Preening comb. A Lepthercus haddadi sp. nov. (female paratype, NCA 2007/3847). Maxillary cuspules (males). B L. dippenaarae. C L. kwazuluensis. Serrula. D L. mandelai sp. nov. (female, NCA 2014/415A); E L. filmeri sp. nov. (female, NCA 91/1437); F L. haddadi sp. nov. (male, NCA 2007/1486); G L. haddadi sp. nov. (female, NCA 2007/3847); H L. engelbrechti sp. nov. (female, NCA 2018/385); I L. engelbrechti sp. nov. (male, NCA 2018/384); J L. dregei Purcell, 1902 (female, NCA 2012/2333); K L. sofiae sp. nov. (male, NCA 97/395)." figureDoi="http://doi.org/10.5281/zenodo.3765471" httpUri="https://zenodo.org/record/3765471/files/figure.png" pageId="30" pageNumber="291">Fig. 2I</figureCitation>
|
||
).
|
||
<emphasis box="[1212,1305,431,457]" italics="true" pageId="30" pageNumber="291">Sternum</emphasis>
|
||
(
|
||
<figureCitation box="[1320,1423,430,456]" captionStart="FIGURE" captionStartId="28.[151,251,1761,1786]" captionTargetBox="[151,1436,631,1735]" captionTargetId="figure@28.[151,1436,631,1735]" captionTargetPageId="28" captionText="FIGURE. 17. Lepthercus engelbrechti sp. nov. (male holotype, NCA 2018/384). A habitus, dorsal view; B sternum, ventral view; C chelicerae, prolateral view, arrow indicates the intercheliceral tumesence; D right pedipalp, arrows indicates the palpal tibia base dorsally very convex and a development of the anterior part of the retrolateral side of ventral excavation; E right pedipalp, arrow indicates the development of the anterior part of the retrolateral side of ventral excavation, prolateral view; F–H copulatory bulb, F retrolateral view, G ventral view, H prolateral view; I tibia-metatarsus I, arrow indicates the curved metatarsus and the apical patch of numerous black blunt spinules, prolateral view; J tibia I, arrow indicates the strong curved spine near of the base of the megaspine and the prolateral strong spine, ventral view." figureDoi="http://doi.org/10.5281/zenodo.3765503" httpUri="https://zenodo.org/record/3765503/files/figure.png" pageId="30" pageNumber="291">Fig. 17B</figureCitation>
|
||
): length 2.66, maximum width 2.05.
|
||
<emphasis box="[552,658,465,491]" italics="true" pageId="30" pageNumber="291">Abdomen</emphasis>
|
||
: length 5.28, covered with small dark hair and bristles. PMS: length 0.54; PLS: length of basal:medial:apical segments 1.02:0.63:1.23; total length 2.88.
|
||
<emphasis box="[1117,1402,500,526]" italics="true" pageId="30" pageNumber="291">Lengths of legs and palp</emphasis>
|
||
: I: 3.47, 2.18, 2.66 (width/length: 0.53), 2.42, 1.60, 12.33. II: 3.40, 2.06, 2.25, 2.17, 1.48, 11.36. III: 3.08, 1.79, 1.88, 2.66, 1.47, 10.88. IV: 3.88, 2.19, 2.71, 3.51, 1.68, 13.97. Palp: 1.76, 1.01, 1.41, —, 0.80, 4.98.
|
||
</paragraph>
|
||
<paragraph blockId="30.[151,1437,152,2017]" pageId="30" pageNumber="291">
|
||
<emphasis box="[199,327,604,630]" italics="true" pageId="30" pageNumber="291">Chaetotaxy</emphasis>
|
||
: Leg I: femur, 1-1-1-1 d (centrals), 1 PA; patella, 1 PA; tibia, 1-1 P, 2-3-1 V, a small spur, with an apical megaspine with the apex flattened laterally. Presence of a strong curved spine near of the base of the megaspine and with another prolateral strong curved spine, near the spur (
|
||
<figureCitation box="[973,1088,673,699]" captionStart="FIGURE" captionStartId="28.[151,251,1761,1786]" captionTargetBox="[151,1436,631,1735]" captionTargetId="figure@28.[151,1436,631,1735]" captionTargetPageId="28" captionText="FIGURE. 17. Lepthercus engelbrechti sp. nov. (male holotype, NCA 2018/384). A habitus, dorsal view; B sternum, ventral view; C chelicerae, prolateral view, arrow indicates the intercheliceral tumesence; D right pedipalp, arrows indicates the palpal tibia base dorsally very convex and a development of the anterior part of the retrolateral side of ventral excavation; E right pedipalp, arrow indicates the development of the anterior part of the retrolateral side of ventral excavation, prolateral view; F–H copulatory bulb, F retrolateral view, G ventral view, H prolateral view; I tibia-metatarsus I, arrow indicates the curved metatarsus and the apical patch of numerous black blunt spinules, prolateral view; J tibia I, arrow indicates the strong curved spine near of the base of the megaspine and the prolateral strong spine, ventral view." figureDoi="http://doi.org/10.5281/zenodo.3765503" httpUri="https://zenodo.org/record/3765503/files/figure.png" pageId="30" pageNumber="291">Fig. 17I, J</figureCitation>
|
||
); metatarsus, 1-1-2 V, presence of numerous black blunt spinules (1:3A,
|
||
<figureCitation box="[614,707,708,734]" captionStart="FIGURE" captionStartId="28.[151,251,1761,1786]" captionTargetBox="[151,1436,631,1735]" captionTargetId="figure@28.[151,1436,631,1735]" captionTargetPageId="28" captionText="FIGURE. 17. Lepthercus engelbrechti sp. nov. (male holotype, NCA 2018/384). A habitus, dorsal view; B sternum, ventral view; C chelicerae, prolateral view, arrow indicates the intercheliceral tumesence; D right pedipalp, arrows indicates the palpal tibia base dorsally very convex and a development of the anterior part of the retrolateral side of ventral excavation; E right pedipalp, arrow indicates the development of the anterior part of the retrolateral side of ventral excavation, prolateral view; F–H copulatory bulb, F retrolateral view, G ventral view, H prolateral view; I tibia-metatarsus I, arrow indicates the curved metatarsus and the apical patch of numerous black blunt spinules, prolateral view; J tibia I, arrow indicates the strong curved spine near of the base of the megaspine and the prolateral strong spine, ventral view." figureDoi="http://doi.org/10.5281/zenodo.3765503" httpUri="https://zenodo.org/record/3765503/files/figure.png" pageId="30" pageNumber="291">Fig. 17I</figureCitation>
|
||
); tarsus, 0. Leg II: femur, 1-1-1-1 d, 1-1 PA; patella, 1 PA; tibia, 1-1 P, 2-2-3 V; metatarsus, 1-1-1 P, 2-3-2 V; tarsus, 0. Leg III: femur, 1-1-1-1 d, 1-1 PA; patella, 1-1 P, 1 R; tibia, 1-1 P, 1-1 D, 1-1 R, 2-2-3 V; metatarsus, 1-1-1 P, 1-1-1 D-P, 1-1-1-1 D-R, 2-3-1-3 V; tarsus, 0. Leg IV: femur, 1-1-1-1 d, 1-1 RA; patella, 1 R; tibia, 1-1 R, 2-2-2 V; metatarsus, 2-1-1-1 P, 1-1-1 D-P, 1-1-0-1 D-R, 1-1 R, 2-3-3 V; tarsus, 0. Palp (
|
||
<figureCitation box="[216,352,846,873]" captionStart="FIGURE" captionStartId="28.[151,251,1761,1786]" captionTargetBox="[151,1436,631,1735]" captionTargetId="figure@28.[151,1436,631,1735]" captionTargetPageId="28" captionText="FIGURE. 17. Lepthercus engelbrechti sp. nov. (male holotype, NCA 2018/384). A habitus, dorsal view; B sternum, ventral view; C chelicerae, prolateral view, arrow indicates the intercheliceral tumesence; D right pedipalp, arrows indicates the palpal tibia base dorsally very convex and a development of the anterior part of the retrolateral side of ventral excavation; E right pedipalp, arrow indicates the development of the anterior part of the retrolateral side of ventral excavation, prolateral view; F–H copulatory bulb, F retrolateral view, G ventral view, H prolateral view; I tibia-metatarsus I, arrow indicates the curved metatarsus and the apical patch of numerous black blunt spinules, prolateral view; J tibia I, arrow indicates the strong curved spine near of the base of the megaspine and the prolateral strong spine, ventral view." figureDoi="http://doi.org/10.5281/zenodo.3765503" httpUri="https://zenodo.org/record/3765503/files/figure.png" pageId="30" pageNumber="291">Fig. 17D, E</figureCitation>
|
||
): femur, 1-1-1 d (1:2A), 1 PA; patella, 1 pa; tibia, 1 P, (width/length: 0.80); tarsus, 0.
|
||
</paragraph>
|
||
<paragraph blockId="30.[151,1437,152,2017]" pageId="30" pageNumber="291">
|
||
<emphasis box="[199,302,881,907]" italics="true" pageId="30" pageNumber="291">Scopulae</emphasis>
|
||
: Metatarsi: I, sparse, around the black blunt spinules; II, sparse (3–4 A), more abundant in the anterior part, with some long and fine setae; III, sparse (A); IV, absent. Tarsi: I–II, dense, uniformly distributed throughout the segment; III, sparse, uniformly distributed and divided by a narrow band of setae; IV, sparse, uniformly distributed and divided by a wide band of setae.
|
||
<emphasis box="[635,791,985,1011]" italics="true" pageId="30" pageNumber="291">Trichobothria</emphasis>
|
||
: Tibiae: I 11-11; II 10-10; III 9-9; IV 11-11. Metatarsi: I (3)1(2)1(4)1; II (5)1(2)1(3)1; III (4)1(2)1(5)1; IV (6)1(2)1(3)1(6)1. Tarsi: I 13; II 12; III-IV 13.
|
||
<emphasis box="[1247,1429,1020,1046]" italics="true" pageId="30" pageNumber="291">Preening combs</emphasis>
|
||
: metatarsus: II, 3 PV; III, 3 PV–2 RV; IV, 2 PV–3 RV.
|
||
<emphasis box="[763,949,1054,1080]" italics="true" pageId="30" pageNumber="291">Copulatory bulb</emphasis>
|
||
: with a curved embolus tapering gradually (
|
||
<figureCitation box="[160,237,1089,1115]" captionStart="FIGURE" captionStartId="28.[151,251,1761,1786]" captionTargetBox="[151,1436,631,1735]" captionTargetId="figure@28.[151,1436,631,1735]" captionTargetPageId="28" captionText="FIGURE. 17. Lepthercus engelbrechti sp. nov. (male holotype, NCA 2018/384). A habitus, dorsal view; B sternum, ventral view; C chelicerae, prolateral view, arrow indicates the intercheliceral tumesence; D right pedipalp, arrows indicates the palpal tibia base dorsally very convex and a development of the anterior part of the retrolateral side of ventral excavation; E right pedipalp, arrow indicates the development of the anterior part of the retrolateral side of ventral excavation, prolateral view; F–H copulatory bulb, F retrolateral view, G ventral view, H prolateral view; I tibia-metatarsus I, arrow indicates the curved metatarsus and the apical patch of numerous black blunt spinules, prolateral view; J tibia I, arrow indicates the strong curved spine near of the base of the megaspine and the prolateral strong spine, ventral view." figureDoi="http://doi.org/10.5281/zenodo.3765503" httpUri="https://zenodo.org/record/3765503/files/figure.png" pageId="30" pageNumber="291">Fig. 17</figureCitation>
|
||
F–H), spermatic duct slightly curved.
|
||
<emphasis box="[659,724,1089,1115]" italics="true" pageId="30" pageNumber="291">Color</emphasis>
|
||
: Overall brown-reddish (Mt-T I darker). Abdomen mottled with light spots forming wide bands; spinnerets light yellow with ventral dark longitudinal bands.
|
||
</paragraph>
|
||
<paragraph blockId="30.[151,1437,152,2017]" pageId="30" pageNumber="291">
|
||
Female
|
||
<typeStatus box="[289,385,1159,1185]" pageId="30" pageNumber="291">paratype</typeStatus>
|
||
(NCA 2018/385,
|
||
<figureCitation box="[589,679,1158,1184]" captionStart="FIGURE" captionStartId="29.[151,251,1870,1895]" captionTargetBox="[164,1423,181,1844]" captionTargetId="figure@29.[164,1423,181,1844]" captionTargetPageId="29" captionText="FIGURE. 18. Lepthercus engelbrechti sp. nov. (female paratype, NCA 2018/385). A habitus, dorsal view; B sternum, ventral view; C chelicerae, prolateral view; D spermathecae" figureDoi="http://doi.org/10.5281/zenodo.3765507" httpUri="https://zenodo.org/record/3765507/files/figure.png" pageId="30" pageNumber="291">Figs. 18</figureCitation>
|
||
A–D). Total length 12.85.
|
||
<emphasis box="[978,1088,1159,1185]" italics="true" pageId="30" pageNumber="291">Carapace</emphasis>
|
||
(
|
||
<figureCitation box="[1105,1211,1158,1184]" captionStart="FIGURE" captionStartId="29.[151,251,1870,1895]" captionTargetBox="[164,1423,181,1844]" captionTargetId="figure@29.[164,1423,181,1844]" captionTargetPageId="29" captionText="FIGURE. 18. Lepthercus engelbrechti sp. nov. (female paratype, NCA 2018/385). A habitus, dorsal view; B sternum, ventral view; C chelicerae, prolateral view; D spermathecae" figureDoi="http://doi.org/10.5281/zenodo.3765507" httpUri="https://zenodo.org/record/3765507/files/figure.png" pageId="30" pageNumber="291">Fig. 18A</figureCitation>
|
||
): length 4.88, width 3.62.
|
||
<emphasis box="[214,395,1193,1219]" italics="true" pageId="30" pageNumber="291">Cephalic region</emphasis>
|
||
: length 3.20, clypeus small (almost absent) with 5 marginal bristles; 7 bristles in front of the OQ and numerous small ones between the PME.
|
||
<emphasis box="[716,795,1228,1253]" italics="true" pageId="30" pageNumber="291">Fovea:</emphasis>
|
||
width 0.34, almost straight (slightly procurved).
|
||
<emphasis italics="true" pageId="30" pageNumber="291">Ocular measurements</emphasis>
|
||
: AME 0.13, ALE 0.28, PME 0.19, PLE 0.25, OQ length 0.45, width 0.80; AME–ALE 0.06, PME– PLE 0.00, AME–PME 0.06, ALE–PLE 0.03, AME–AME 0.12, PME–PME 0.30.
|
||
<emphasis box="[1096,1219,1297,1323]" italics="true" pageId="30" pageNumber="291">Chelicerae</emphasis>
|
||
(
|
||
<figureCitation box="[1237,1343,1297,1323]" captionStart="FIGURE" captionStartId="29.[151,251,1870,1895]" captionTargetBox="[164,1423,181,1844]" captionTargetId="figure@29.[164,1423,181,1844]" captionTargetPageId="29" captionText="FIGURE. 18. Lepthercus engelbrechti sp. nov. (female paratype, NCA 2018/385). A habitus, dorsal view; B sternum, ventral view; C chelicerae, prolateral view; D spermathecae" figureDoi="http://doi.org/10.5281/zenodo.3765507" httpUri="https://zenodo.org/record/3765507/files/figure.png" pageId="30" pageNumber="291">Fig. 18C</figureCitation>
|
||
): length 2.64, width 1.71; with dorsal-retrolateral dark bristles. Fang groove with 8 promarginal teeth (one small between 7th and 8th) and 15 mesobasal denticles.
|
||
<emphasis box="[577,663,1366,1392]" italics="true" pageId="30" pageNumber="291">Labium</emphasis>
|
||
: length 0.40, width 0.81.
|
||
<emphasis box="[957,1057,1366,1392]" italics="true" pageId="30" pageNumber="291">Maxillae</emphasis>
|
||
: length 1.71, width 0.75, with 75 cuspules on inner corner, prolateral face curved, soft area long, with long uniformly distributed hairs, serrula present on anterior inner corner (
|
||
<figureCitation box="[435,527,1436,1462]" captionStart="FIGURE" captionStartId="7.[151,251,1647,1672]" captionTargetBox="[151,1436,381,1622]" captionTargetId="figure@7.[151,1436,381,1622]" captionTargetPageId="7" captionText="FIGURE. 2. Preening comb. A Lepthercus haddadi sp. nov. (female paratype, NCA 2007/3847). Maxillary cuspules (males). B L. dippenaarae. C L. kwazuluensis. Serrula. D L. mandelai sp. nov. (female, NCA 2014/415A); E L. filmeri sp. nov. (female, NCA 91/1437); F L. haddadi sp. nov. (male, NCA 2007/1486); G L. haddadi sp. nov. (female, NCA 2007/3847); H L. engelbrechti sp. nov. (female, NCA 2018/385); I L. engelbrechti sp. nov. (male, NCA 2018/384); J L. dregei Purcell, 1902 (female, NCA 2012/2333); K L. sofiae sp. nov. (male, NCA 97/395)." figureDoi="http://doi.org/10.5281/zenodo.3765471" httpUri="https://zenodo.org/record/3765471/files/figure.png" pageId="30" pageNumber="291">Fig. 2H</figureCitation>
|
||
).
|
||
<emphasis box="[547,640,1436,1462]" italics="true" pageId="30" pageNumber="291">Sternum</emphasis>
|
||
(
|
||
<figureCitation box="[657,761,1436,1462]" captionStart="FIGURE" captionStartId="29.[151,251,1870,1895]" captionTargetBox="[164,1423,181,1844]" captionTargetId="figure@29.[164,1423,181,1844]" captionTargetPageId="29" captionText="FIGURE. 18. Lepthercus engelbrechti sp. nov. (female paratype, NCA 2018/385). A habitus, dorsal view; B sternum, ventral view; C chelicerae, prolateral view; D spermathecae" figureDoi="http://doi.org/10.5281/zenodo.3765507" httpUri="https://zenodo.org/record/3765507/files/figure.png" pageId="30" pageNumber="291">Fig. 18B</figureCitation>
|
||
): length 2.36, maximum width 1.99.
|
||
<emphasis box="[1181,1287,1436,1462]" italics="true" pageId="30" pageNumber="291">Abdomen</emphasis>
|
||
: length 5.60, with small bristles in the dorsum. PMS: length 0.64; PLS: length of basal:medial:apical segments 1.00:0.58:1.00; total length 2.58.
|
||
<emphasis box="[347,621,1505,1531]" italics="true" pageId="30" pageNumber="291">Lengths of legs and palp</emphasis>
|
||
: I: 3.24, 2.06, 2.06, 1.48, 1.03, 9.87. II: 2.82, 1.79, 1.68, 1,43, 1.06, 8.78. III: 2.05, 1.44, 1.36, 1.92, 1.15, 7.92. IV: 3.31, 1.98, 2.19, 2.76, 1.31, 11.55. Palp: 2.21, 1.36, 1.29, —, 1.31, 6.17.
|
||
</paragraph>
|
||
<paragraph blockId="30.[151,1437,152,2017]" pageId="30" pageNumber="291">
|
||
<emphasis box="[199,327,1574,1600]" italics="true" pageId="30" pageNumber="291">Chaetotaxy</emphasis>
|
||
: Leg I: femur, 0; patella, 0; tibia, 1-2 vant; metatarsus, 2-1-2 V; tarsus, 0. Leg II: femur, 1 pa; patella, 1 pa; tibia, 1 P, 1-2 vant; metatarsus, 2-2-3 V; tarsus, 0. Leg III: femur, 1-1-1 d, 1-1 ra; patella, 1-1 P, 1 R; tibia, 1-1 P, 1-1 D, 1 R, 2-2-3 V; metatarsus, 2-1-1 P, 1-1-1 D-P, 1-1-1-1 D-R, 2-3-3 V; tarsus, 0. Leg IV: femur, 1-1-1-1 d, 1 ra; patella, 1 R; tibia, 1-1 R, 2-2-2 V; metatarsus, 1-1-1 P, 1-1-1 D-P, 1-1-1-0-1 D-R, 1 R (1:3A), 2-3-3 V; tarsus, 0. Palp: femur, 1-1-1-1 d (1:2A), 1 pa; patella, 0; tibia, 1-3 V; tarsus, 0.
|
||
</paragraph>
|
||
<paragraph blockId="30.[151,1437,152,2017]" pageId="30" pageNumber="291">
|
||
<emphasis box="[199,302,1748,1774]" italics="true" pageId="30" pageNumber="291">Scopulae</emphasis>
|
||
: Metatarsi: I, dense, uniformly distributed, divided by setae; II–IV, without scopula. Tarsi: I, dense, uniformly distributed, divided by setae; II dense, uniformly distributed, divided by a narrow band of setae; III–IV, sparse, uniformly distributed and divided by a wide band of setae.
|
||
<emphasis box="[911,1067,1817,1843]" italics="true" pageId="30" pageNumber="291">Trichobothria</emphasis>
|
||
: Tibiae: palp 8-7; I 9-9; II 9-10; III 9-8; IV 10-10. Metatarsi: I (3)1(2)1(4)1; II (4)1(2)1(4)1; III (4)1(2)1(5)1; IV (4)1(1)1(2)1(6)1. Tarsi: palp 11; I 12; II 11; III 12; IV 11.
|
||
<emphasis box="[417,598,1886,1912]" italics="true" pageId="30" pageNumber="291">Preening combs</emphasis>
|
||
: metatarsus: II 3 PV; III 3 PV–3 RV; IV 2 PV–3 RV.
|
||
<emphasis box="[1195,1354,1886,1912]" italics="true" pageId="30" pageNumber="291">Spermathecae</emphasis>
|
||
: small, tubulars and curved (
|
||
<figureCitation box="[390,495,1921,1947]" captionStart="FIGURE" captionStartId="29.[151,251,1870,1895]" captionTargetBox="[164,1423,181,1844]" captionTargetId="figure@29.[164,1423,181,1844]" captionTargetPageId="29" captionText="FIGURE. 18. Lepthercus engelbrechti sp. nov. (female paratype, NCA 2018/385). A habitus, dorsal view; B sternum, ventral view; C chelicerae, prolateral view; D spermathecae" figureDoi="http://doi.org/10.5281/zenodo.3765507" httpUri="https://zenodo.org/record/3765507/files/figure.png" pageId="30" pageNumber="291">Fig. 18D</figureCitation>
|
||
).
|
||
<emphasis box="[516,581,1921,1947]" italics="true" pageId="30" pageNumber="291">Color</emphasis>
|
||
: Overall orange-yellowish (chelicerae darker). Abdomen mottled with light spots forming wide bands, spinnerets light yellow with dark, longitudinal bands ventrally.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection box="[199,1039,1989,2017]" pageId="30" pageNumber="291" type="distribution">
|
||
<paragraph blockId="30.[151,1437,152,2017]" box="[199,1039,1989,2017]" pageId="30" pageNumber="291">
|
||
<emphasis bold="true" box="[199,353,1989,2016]" pageId="30" pageNumber="291">Distribution.</emphasis>
|
||
Eastern and
|
||
<collectingRegion box="[497,653,1990,2017]" country="South Africa" name="Western Cape" pageId="30" pageNumber="291">Western Cape</collectingRegion>
|
||
Provinces,
|
||
<collectingCountry box="[784,927,1990,2016]" name="South Africa" pageId="30" pageNumber="291">South Africa</collectingCountry>
|
||
(
|
||
<figureCitation box="[942,1026,1990,2016]" captionStart="FIGURE" captionStartId="38.[151,251,1816,1841]" captionTargetBox="[185,1405,888,1784]" captionTargetId="figure@38.[176,1411,885,1790]" captionTargetPageId="38" captionText="FIGURE. 24. Distribution map of Lepthercus spp: ● Lepthercus confusus sp. nov., ○ Lepthercus dippenaarae sp. nov., Ɨ Lepthercus dregei Purcell, 1902, O Lepthercus engelbrechti sp. nov., A Lepthercus filmeri sp. nov., * Lepthercus haddadi sp. nov., D Lepthercus kwazuluensis sp. nov., ■ Lepthercus lawrencei sp. nov., v Lepthercus mandelai sp. nov., ▼ Lepthercus rattrayi Hewitt, 1917, ▲ Lepthercus sofiae sp. nov. Solid black line indicates the two species overlapped." figureDoi="http://doi.org/10.5281/zenodo.3765519" httpUri="https://zenodo.org/record/3765519/files/figure.png" pageId="30" pageNumber="291">Fig. 24</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |