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<document id="21BC8C8D8D4890285F94D2F335201F82" ID-CLB-Dataset="55547" ID-DOI="10.11646/phytotaxa.545.3.2" ID-GBIF-Dataset="d2fbb8cd-35d6-4797-8533-0c88416371fa" ID-ISSN="1179-3163" ID-Zenodo-Dep="6541464" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1652341273392" checkinUser="plazi" docAuthor="Vijver, Bart Van De, Lange-Bertalot, Horst, Goeyers, Charlotte, Mertens, Adrienne, Schuster, Tanja M. &amp; Ector, Luc" docDate="2022" docId="03F287F77D0B801ACEFBFF7DFD23E2F3" docLanguage="en" docName="phytotaxa.545.3.2.pdf" docOrigin="Phytotaxa 545 (3)" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.7:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="7" docTitle="Eunotia zackenbergensis Goeyers, Van de Vijver &amp; Lange-Bertalot 2022, sp. nov." docType="treatment" docVersion="7" lastPageNumber="272" masterDocId="FFCBFF8F7D028011CE73FFE3FFC1E00B" masterDocTitle="The identity of Eunotia paludosa Grunow 1862 (Eunotiaceae, Bacillariophyta), a revision, and the description of three new species of Eunotia Ehrenberg" masterLastPageNumber="277" masterPageNumber="261" pageNumber="270" updateTime="1699330211755" updateUser="ExternalLinkService" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="64710CCB24E6CD5295A02604B336D7E7">The identity of Eunotia paludosa Grunow 1862 (Eunotiaceae, Bacillariophyta), a revision, and the description of three new species of Eunotia Ehrenberg</mods:title>
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<mods:namePart id="0F3916F988D2D0BDD106C7B492CBF71D">Vijver, Bart Van De</mods:namePart>
<mods:affiliation id="CDDE85A0AADEAE7BFF9399F6A3C9E65B">Meise Botanic Garden Meise, Research Department, Nieuwelaan 38, 1860 Meise, Belgium &amp; University of Antwerp, Department of Biology - ECOSPHERE, Universiteitsplein 1, B- 2610 Wilrijk, Belgium</mods:affiliation>
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<mods:namePart id="01283F6A0268262E02459FB165CAFD5C">Lange-Bertalot, Horst</mods:namePart>
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<mods:affiliation id="82FCB969CBC95E1BEF68E1F8DC5DBED4">Botanisches Institut, Goethe-Universität, Frankfurt am Main, Germany &amp; retired; https: // orcid. org / 0000 - 0002 - 9513 - 3310</mods:affiliation>
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<mods:namePart id="0AB54C67FD3347C93EF6F87984D9F332">Goeyers, Charlotte</mods:namePart>
<mods:nameIdentifier id="128D067D9B50C0480F2E1B572287D6FA" type="ORCID">0000-0002-1573-3923</mods:nameIdentifier>
<mods:affiliation id="7FB2DD3DBB5800DE0E711DFB01BD2281">Meise Botanic Garden Meise, Research Department, Nieuwelaan 38, 1860 Meise, Belgium &amp; charlotte. goeyers @ plantentuinmeise. be; https: // orcid. org / 0000 - 0002 - 1573 - 3923</mods:affiliation>
<mods:nameIdentifier id="04A414B495046D46DE8B00063F0F352A" type="email">charlotte.goeyers@plantentuinmeise.be</mods:nameIdentifier>
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<mods:namePart id="15C3D5EE4D75B0C8713ABAA94DE36CA1">Mertens, Adrienne</mods:namePart>
<mods:nameIdentifier id="CF9FF108AF253C7336D0972199F0A4CB" type="ORCID">0000-0003-2362-461X</mods:nameIdentifier>
<mods:affiliation id="F3853CC353727EC4293C1553B84CB451">Diatomella, IJkelaarstraat 3, 6611 KN Overasselt, the Netherlands &amp; adrienne @ diatomella. nl; https: // orcid. org / 0000 - 0003 - 2362 - 461 X</mods:affiliation>
<mods:nameIdentifier id="3E3B0459FC77C5BCA60B24FECDCA9686" type="email">adrienne@diatomella.nl</mods:nameIdentifier>
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<mods:namePart id="28CA38D7B2FAEC908590B97EE81B03AA">Schuster, Tanja M.</mods:namePart>
<mods:nameIdentifier id="BF27F7A45AEA02EA9C78715919EFB014" type="ORCID">0000-0003-0851-3372</mods:nameIdentifier>
<mods:affiliation id="BF3B0DDA933ABCD7B2C70A05402BEB78">Natural History Museum, Department of Botany, Herbarium, Burgring 7, 1010 Vienna, Austria &amp; Tanja. Schuster @ NHM-WIEN. AC. AT; https: // orcid. org / 0000 - 0003 - 0851 - 3372</mods:affiliation>
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<mods:namePart id="C57A5F727EC35240AAD990D62F0BD6E8">Ector, Luc</mods:namePart>
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<mods:affiliation id="09CF776CB090B1E34B06969945D7B06C">Luxembourg Institute of Science and Technology (LIST), Environmental Research &amp; Innovation (ERIN) Department, Observatory for Climate, Environment and Biodiversity (OCEB), 41 rue du Brill, L- 4422 Belvaux, Luxembourg &amp; DECEASED; https: // orcid. org / 0000 - 0002 - 4573 - 9445</mods:affiliation>
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<mods:title id="411879A9B1297507E978FD517EAF3A8B">Phytotaxa</mods:title>
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<mods:date id="468D374BDF1400C893581BC9F082909B">2022</mods:date>
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<mods:number id="07CC8D9591A8B01BB025986D3014B14D">2022-05-12</mods:number>
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<mods:identifier id="FACA5E77DB9F52F698E6BED72064FE28" type="DOI">10.11646/phytotaxa.545.3.2</mods:identifier>
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<mods:identifier id="34FD1352FD4B37221B75F1AAB0202DB0" type="ISSN">1179-3163</mods:identifier>
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<treatment id="03F287F77D0B801ACEFBFF7DFD23E2F3" ID-DOI="http://doi.org/10.5281/zenodo.6543102" ID-GBIF-Taxon="195304021" ID-Zenodo-Dep="6543102" LSID="urn:lsid:plazi:treatment:03F287F77D0B801ACEFBFF7DFD23E2F3" httpUri="http://treatment.plazi.org/id/03F287F77D0B801ACEFBFF7DFD23E2F3" lastPageId="11" lastPageNumber="272" pageId="9" pageNumber="270">
<subSubSection id="C341656A7D0B8018CEFBFF7DFB74E0B2" box="[136,1205,158,185]" pageId="9" pageNumber="270" type="nomenclature">
<paragraph id="8BE436E17D0B8018CEFBFF7DFB74E0B2" blockId="9.[136,1205,158,185]" box="[136,1205,158,185]" pageId="9" pageNumber="270">
<heading id="D0AC818D7D0B8018CEFBFF7DFB74E0B2" box="[136,1205,158,185]" fontSize="11" level="2" pageId="9" pageNumber="270" reason="3">
<emphasis id="B92FEAF37D0B8018CEFBFF7DFBC3E0B2" bold="true" box="[136,1026,158,185]" pageId="9" pageNumber="270">
<taxonomicName id="4C5B4D627D0B8018CEFBFF7DFC61E0B3" authority="Goeyers, Van de Vijver &amp; Lange-Bertalot" authorityName="Goeyers, Van de Vijver &amp; Lange-Bertalot" authorityYear="2022" box="[136,928,158,185]" class="Bacillariophyceae" family="Eunotiaceae" genus="Eunotia" higherTaxonomySource="GBIF" kingdom="Chromista" order="Eunotiales" pageId="9" pageNumber="270" phylum="Bacillariophyta" rank="species" species="zackenbergensis" status="sp. nov.">
<emphasis id="B92FEAF37D0B8018CEFBFF7DFE69E0B2" bold="true" box="[136,424,158,185]" italics="true" pageId="9" pageNumber="270">Eunotia zackenbergensis</emphasis>
Goeyers, Van de Vijver &amp; Lange-Bertalot
</taxonomicName>
,
<emphasis id="B92FEAF37D0B8018CDDFFF7CFBC3E0B2" bold="true" box="[940,1026,159,185]" italics="true" pageId="9" pageNumber="270">
<taxonomicNameLabel id="A21C57887D0B8018CDDFFF7CFBC3E0B2" box="[940,1026,159,185]" pageId="9" pageNumber="270" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</emphasis>
(
<figureCitation id="13602A647D0B8018CA62FF7DFB6FE0B2" box="[1041,1198,158,185]" captionStart-0="FIGURES 127–161" captionStart-1="FIGURES 162–168" captionStartId-0="9.[136,243,1820,1842]" captionStartId-1="10.[136,243,1769,1791]" captionTargetBox-0="[151,1435,569,1796]" captionTargetBox-1="[151,1435,190,1744]" captionTargetId-0="figure-158@9.[151,1435,569,1796]" captionTargetId-1="figure-21@10.[151,1435,190,1744]" captionTargetPageId-0="9" captionTargetPageId-1="10" captionText-0="FIGURES 127–161. Eunotia zackenbergensis Goeyers, Van de Vijver &amp; Lange-Bertalot, sp. nov. LM images taken from the holotype material (sample M446, Zackenberg, Greenland, BR-4718). Fig. 127. Single valve in girdle view. Figs 128–161. LM views of the population arranged in decreasing length. Scale bar represents 10 µm." captionText-1="FIGURES 162–168. Eunotia zackenbergensis Goeyers, Van de Vijver &amp; Lange-Bertalot, sp. nov. SEM images taken from the holotype material (sample M446, Zackenberg, Greenland). Fig. 162. External view of a valve in girdle view. Note the course of the raphe branches on the mantle. Fig. 163. SEM external view in valve face view showing the short terminal raphe fissures and the absence of spines. Fig. 164. External detail of a valve apex focusing on the terminal raphe fissures. The external rimoportula opening is not discernible. Fig. 165. SEM external detail of a valve apex in girdle view, focusing on one of the raphe branches and the areolae on the apex. Fig. 166. Internal view in valve face view. Fig. 167. SEM internal detail of a valve apex showing the helictoglossa and the rimoportula. Fig. 168. SEM internal detail of a valve apex showing the helictoglossa. Scale bars indicate 10 µm for figs 162, 163 &amp; 166, 1 µm for figs 164, 165, 167 &amp; 168." figureDoi-0="http://doi.org/10.5281/zenodo.6541480" figureDoi-1="http://doi.org/10.5281/zenodo.6541482" httpUri-0="https://zenodo.org/record/6541480/files/figure.png" httpUri-1="https://zenodo.org/record/6541482/files/figure.png" pageId="9" pageNumber="270">Figs 127–168</figureCitation>
)
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C341656A7D0B801ACEFBFF25FE77E19A" lastPageId="11" lastPageNumber="272" pageId="9" pageNumber="270" type="materials_examined">
<paragraph id="8BE436E17D0B8018CEFBFF25FBE5E0F4" blockId="9.[136,1452,198,292]" pageId="9" pageNumber="270">
<emphasis id="B92FEAF37D0B8018CEFBFF25FF7BE0D0" bold="true" box="[136,186,198,219]" pageId="9" pageNumber="270">Type</emphasis>
:—
<materialsCitation id="3B333CBC7D0B8018CEABFF25FBE1E0F4" ID-GBIF-Occurrence="3766726301" collectingDate="1998-08" collectorName="L. Beyens" country="Greenland" location="Zackenberg" municipality="University of Antwerp" pageId="9" pageNumber="270" specimenCount="1" typeStatus="holotype">
<collectingCountry id="F34C76717D0B8018CEABFF25FE84E0D2" box="[216,325,198,219]" name="Greenland" pageId="9" pageNumber="270">GREENLAND</collectingCountry>
,
<location id="8E84603A7D0B8018CF23FF25FE00E0D7" LSID="urn:lsid:plazi:treatment:03F287F77D0B801ACEFBFF7DFD23E2F3:8E84603A7D0B8018CF23FF25FE00E0D7" box="[336,449,198,220]" country="Greenland" municipality="University of Antwerp" name="Zackenberg" pageId="9" pageNumber="270">Zackenberg</location>
(sample M446, coll. date
<date id="FFE510217D0B8018CCC9FF25FCDDE0D0" box="[698,796,198,219]" pageId="9" pageNumber="270" value="1998-08">
<collectingDate id="EFA1E9C97D0B8018CCC9FF25FCDDE0D0" box="[698,796,198,219]" pageId="9" pageNumber="270" value="1998-08">VIII.1998</collectingDate>
</date>
, leg.
<collectorName id="26AE53377D0B8018CD3DFF25FC73E0D0" box="[846,946,198,219]" pageId="9" pageNumber="270">L. Beyens</collectorName>
), holo- BR-4718! (
<collectorName id="26AE53377D0B8018CA15FF25FA80E0D7" box="[1126,1345,198,220]" pageId="9" pageNumber="270">Meise Botanic Garden</collectorName>
,
<collectorName id="26AE53377D0B8018CB38FF25FA60E0D7" box="[1355,1441,198,220]" pageId="9" pageNumber="270">Belgium</collectorName>
), iso- slide 406! (
<collectingMunicipality id="6B80AC9B7D0B8018CF20FF09FDEEE0F4" box="[339,559,234,256]" pageId="9" pageNumber="270">University of Antwerp</collectingMunicipality>
,
<collectorName id="26AE53377D0B8018CC4AFF09FD51E10B" box="[569,656,234,256]" pageId="9" pageNumber="270">Belgium</collectorName>
).
<collectorName id="26AE53377D0B8018CCECFF09FD05E10B" box="[671,708,234,256]" pageId="9" pageNumber="270">The</collectorName>
<typeStatus id="54E088437D0B8018CCB9FF09FCDFE10B" box="[714,798,234,256]" pageId="9" pageNumber="270" type="holotype">holotype</typeStatus>
is represented by
<figureCitation id="13602A647D0B8018CDBEFF09FBE1E0F4" box="[973,1056,234,256]" captionStart="FIGURES 127–161" captionStartId="9.[136,243,1820,1842]" captionTargetBox="[151,1435,569,1796]" captionTargetId="figure-158@9.[151,1435,569,1796]" captionTargetPageId="9" captionText="FIGURES 127–161. Eunotia zackenbergensis Goeyers, Van de Vijver &amp; Lange-Bertalot, sp. nov. LM images taken from the holotype material (sample M446, Zackenberg, Greenland, BR-4718). Fig. 127. Single valve in girdle view. Figs 128–161. LM views of the population arranged in decreasing length. Scale bar represents 10 µm." figureDoi="http://doi.org/10.5281/zenodo.6541480" httpUri="https://zenodo.org/record/6541480/files/figure.png" pageId="9" pageNumber="270">Fig. 133</figureCitation>
</materialsCitation>
.
</paragraph>
<paragraph id="8BE436E17D0B8018CEFBFEEDFD70E12F" blockId="9.[136,1452,198,292]" box="[136,689,270,292]" pageId="9" pageNumber="270">
<emphasis id="B92FEAF37D0B8018CEFBFEEDFEBAE12F" bold="true" box="[136,379,270,292]" pageId="9" pageNumber="270">PhycoBank registration</emphasis>
:—http://phycobank.org/103148
</paragraph>
<paragraph id="8BE436E17D0B8018CEFBFEB0FE22E22A" blockId="9.[136,1452,338,545]" pageId="9" pageNumber="270">
LM (
<figureCitation id="13602A647D0B8018CEB7FEB1FEA6E166" box="[196,359,338,365]" captionStart="FIGURES 127–161" captionStartId="9.[136,243,1820,1842]" captionTargetBox="[151,1435,569,1796]" captionTargetId="figure-158@9.[151,1435,569,1796]" captionTargetPageId="9" captionText="FIGURES 127–161. Eunotia zackenbergensis Goeyers, Van de Vijver &amp; Lange-Bertalot, sp. nov. LM images taken from the holotype material (sample M446, Zackenberg, Greenland, BR-4718). Fig. 127. Single valve in girdle view. Figs 128–161. LM views of the population arranged in decreasing length. Scale bar represents 10 µm." figureDoi="http://doi.org/10.5281/zenodo.6541480" httpUri="https://zenodo.org/record/6541480/files/figure.png" pageId="9" pageNumber="270">Figs 127–161</figureCitation>
): Frustules rectangular in girdle view (
<figureCitation id="13602A647D0B8018CD5DFEB1FC55E167" box="[814,916,338,364]" captionStart="FIGURES 127–161" captionStartId="9.[136,243,1820,1842]" captionTargetBox="[151,1435,569,1796]" captionTargetId="figure-158@9.[151,1435,569,1796]" captionTargetPageId="9" captionText="FIGURES 127–161. Eunotia zackenbergensis Goeyers, Van de Vijver &amp; Lange-Bertalot, sp. nov. LM images taken from the holotype material (sample M446, Zackenberg, Greenland, BR-4718). Fig. 127. Single valve in girdle view. Figs 128–161. LM views of the population arranged in decreasing length. Scale bar represents 10 µm." figureDoi="http://doi.org/10.5281/zenodo.6541480" httpUri="https://zenodo.org/record/6541480/files/figure.png" pageId="9" pageNumber="270">Fig. 127</figureCitation>
). Valves weakly arched with parallel margins. Dorsal margins clearly convex. Ventral margins moderately concave in almost all cell cycle stages, smaller valves becoming more or less straight. Apices protracted, obliquely subcapitate and usually dorsally reflexed in the entire cell diminution series. Valve dimensions (n=50): length 16–62 µm, width 2.5–3.0 µm, length-to-width ratio 6.8–20. Terminal raphe nodules close to the poles. Striae
<quantity id="4CA39B047D0B8018CCBAFE01FCEEE1F7" box="[713,815,482,508]" metricMagnitude="-1" metricUnit="m" metricValue="5.461" metricValueMax="5.588" metricValueMin="5.334" pageId="9" pageNumber="270" unit="in" value="21.5" valueMax="22.0" valueMin="21.0">21–22 in</quantity>
10 µm, equidistant throughout the entire valve length. Areolae not discernible in LM.
</paragraph>
<caption id="DF2466697D0B8018CEFBF8FFFCE6E771" ID-DOI="http://doi.org/10.5281/zenodo.6541480" ID-Zenodo-Dep="6541480" httpUri="https://zenodo.org/record/6541480/files/figure.png" pageId="9" pageNumber="270" startId="9.[136,243,1820,1842]" targetBox="[151,1435,569,1796]" targetPageId="9">
<paragraph id="8BE436E17D0B8018CEFBF8FFFCE6E771" blockId="9.[136,1451,1820,1915]" pageId="9" pageNumber="270">
<emphasis id="B92FEAF37D0B8018CEFBF8FFFE95E739" bold="true" box="[136,340,1820,1842]" pageId="9" pageNumber="270">FIGURES 127–161.</emphasis>
<taxonomicName id="4C5B4D627D0B8018CF2FF8FEFC2CE738" authority="Goeyers, Van de Vijver &amp; Lange-Bertalot" authorityName="Goeyers, Van de Vijver &amp; Lange-Bertalot" authorityYear="2022" box="[348,1005,1821,1843]" class="Bacillariophyceae" family="Eunotiaceae" genus="Eunotia" higherTaxonomySource="GBIF" kingdom="Chromista" order="Eunotiales" pageId="9" pageNumber="270" phylum="Bacillariophyta" rank="species" species="zackenbergensis" status="sp. nov.">
<emphasis id="B92FEAF37D0B8018CF2FF8FEFD8DE739" box="[348,588,1821,1842]" italics="true" pageId="9" pageNumber="270">Eunotia zackenbergensis</emphasis>
Goeyers, Van de Vijver &amp; Lange-Bertalot
</taxonomicName>
,
<emphasis id="B92FEAF37D0B8018CD8BF8FEFB82E739" box="[1016,1091,1821,1842]" italics="true" pageId="9" pageNumber="270">
<taxonomicNameLabel id="A21C57887D0B8018CD8BF8FEFB82E739" box="[1016,1091,1821,1842]" pageId="9" pageNumber="270" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
LM images taken from the holotype material (sample M446, Zackenberg, Greenland, BR-4718). Fig. 127. Single valve in girdle view. Figs 128–161. LM views of the population arranged in decreasing length. Scale bar represents 10 µm.
</paragraph>
</caption>
<caption id="DF2466697D08801BCEFBF90AFF0DE7F0" ID-DOI="http://doi.org/10.5281/zenodo.6541482" ID-Zenodo-Dep="6541482" httpUri="https://zenodo.org/record/6541482/files/figure.png" pageId="10" pageNumber="271" startId="10.[136,243,1769,1791]" targetBox="[151,1435,190,1744]" targetPageId="10">
<paragraph id="8BE436E17D08801BCEFBF90AFF0DE7F0" blockId="10.[136,1452,1769,2043]" pageId="10" pageNumber="271">
<emphasis id="B92FEAF37D08801BCEFBF90AFE92E6F5" bold="true" box="[136,339,1769,1791]" pageId="10" pageNumber="271">FIGURES 162–168.</emphasis>
<taxonomicName id="4C5B4D627D08801BCF29F90AFC27E6F4" authority="Goeyers, Van de Vijver &amp; Lange-Bertalot" authorityName="Goeyers, Van de Vijver &amp; Lange-Bertalot" authorityYear="2022" box="[346,998,1769,1791]" class="Bacillariophyceae" family="Eunotiaceae" genus="Eunotia" higherTaxonomySource="GBIF" kingdom="Chromista" order="Eunotiales" pageId="10" pageNumber="271" phylum="Bacillariophyta" rank="species" species="zackenbergensis" status="sp. nov.">
<emphasis id="B92FEAF37D08801BCF29F90AFD88E6F5" box="[346,585,1769,1790]" italics="true" pageId="10" pageNumber="271">Eunotia zackenbergensis</emphasis>
Goeyers, Van de Vijver &amp; Lange-Bertalot
</taxonomicName>
,
<emphasis id="B92FEAF37D08801BCD83F909FBFBE6F4" box="[1008,1082,1770,1791]" italics="true" pageId="10" pageNumber="271">
<taxonomicNameLabel id="A21C57887D08801BCD83F909FBFBE6F4" box="[1008,1082,1770,1791]" pageId="10" pageNumber="271" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
SEM images taken from the holotype material (sample M446, Zackenberg, Greenland). Fig. 162. External view of a valve in girdle view. Note the course of the raphe branches on the mantle. Fig. 163. SEM external view in valve face view showing the short terminal raphe fissures and the absence of spines. Fig. 164. External detail of a valve apex focusing on the terminal raphe fissures. The external rimoportula opening is not discernible. Fig. 165. SEM external detail of a valve apex in girdle view, focusing on one of the raphe branches and the areolae on the apex. Fig. 166. Internal view in valve face view. Fig. 167. SEM internal detail of a valve apex showing the helictoglossa and the rimoportula. Fig. 168. SEM internal detail of a valve apex showing the helictoglossa. Scale bars indicate 10 µm for figs 162, 163 &amp; 166, 1 µm for figs 164, 165, 167 &amp; 168.
</paragraph>
</caption>
<paragraph id="8BE436E17D09801ACECEFF7DFE77E19A" blockId="11.[136,1452,158,761]" pageId="11" pageNumber="272">
SEM (
<figureCitation id="13602A647D09801ACF74FF7DFE68E0B2" box="[263,425,158,185]" captionStart="FIGURES 162–168" captionStartId="10.[136,243,1769,1791]" captionTargetBox="[151,1435,190,1744]" captionTargetId="figure-21@10.[151,1435,190,1744]" captionTargetPageId="10" captionText="FIGURES 162–168. Eunotia zackenbergensis Goeyers, Van de Vijver &amp; Lange-Bertalot, sp. nov. SEM images taken from the holotype material (sample M446, Zackenberg, Greenland). Fig. 162. External view of a valve in girdle view. Note the course of the raphe branches on the mantle. Fig. 163. SEM external view in valve face view showing the short terminal raphe fissures and the absence of spines. Fig. 164. External detail of a valve apex focusing on the terminal raphe fissures. The external rimoportula opening is not discernible. Fig. 165. SEM external detail of a valve apex in girdle view, focusing on one of the raphe branches and the areolae on the apex. Fig. 166. Internal view in valve face view. Fig. 167. SEM internal detail of a valve apex showing the helictoglossa and the rimoportula. Fig. 168. SEM internal detail of a valve apex showing the helictoglossa. Scale bars indicate 10 µm for figs 162, 163 &amp; 166, 1 µm for figs 164, 165, 167 &amp; 168." figureDoi="http://doi.org/10.5281/zenodo.6541482" httpUri="https://zenodo.org/record/6541482/files/figure.png" pageId="11" pageNumber="272">Figs 162–168</figureCitation>
): Striae uniseriate throughout, composed of small, rounded areolae,
<quantity id="4CA39B047D09801ACAC1FF7DFAD7E0B3" box="[1202,1302,158,184]" metricMagnitude="0" metricUnit="m" metricValue="1.0922" metricValueMax="1.1176" metricValueMin="1.0668" pageId="11" pageNumber="272" unit="in" value="43.0" valueMax="44.0" valueMin="42.0">42–44 in</quantity>
10 µm (
<figureCitation id="13602A647D09801ACB09FF7DFF2CE0D6" captionStart="FIGURES 162–168" captionStartId="10.[136,243,1769,1791]" captionTargetBox="[151,1435,190,1744]" captionTargetId="figure-21@10.[151,1435,190,1744]" captionTargetPageId="10" captionText="FIGURES 162–168. Eunotia zackenbergensis Goeyers, Van de Vijver &amp; Lange-Bertalot, sp. nov. SEM images taken from the holotype material (sample M446, Zackenberg, Greenland). Fig. 162. External view of a valve in girdle view. Note the course of the raphe branches on the mantle. Fig. 163. SEM external view in valve face view showing the short terminal raphe fissures and the absence of spines. Fig. 164. External detail of a valve apex focusing on the terminal raphe fissures. The external rimoportula opening is not discernible. Fig. 165. SEM external detail of a valve apex in girdle view, focusing on one of the raphe branches and the areolae on the apex. Fig. 166. Internal view in valve face view. Fig. 167. SEM internal detail of a valve apex showing the helictoglossa and the rimoportula. Fig. 168. SEM internal detail of a valve apex showing the helictoglossa. Scale bars indicate 10 µm for figs 162, 163 &amp; 166, 1 µm for figs 164, 165, 167 &amp; 168." figureDoi="http://doi.org/10.5281/zenodo.6541482" httpUri="https://zenodo.org/record/6541482/files/figure.png" pageId="11" pageNumber="272">Figs 163, 164</figureCitation>
). Mantle striae ventrally composed of up to 7 small, rounded areolae near the valve middle (
<figureCitation id="13602A647D09801ACA98FF21FA8CE0D6" box="[1259,1357,194,221]" captionStart="FIGURES 162–168" captionStartId="10.[136,243,1769,1791]" captionTargetBox="[151,1435,190,1744]" captionTargetId="figure-21@10.[151,1435,190,1744]" captionTargetPageId="10" captionText="FIGURES 162–168. Eunotia zackenbergensis Goeyers, Van de Vijver &amp; Lange-Bertalot, sp. nov. SEM images taken from the holotype material (sample M446, Zackenberg, Greenland). Fig. 162. External view of a valve in girdle view. Note the course of the raphe branches on the mantle. Fig. 163. SEM external view in valve face view showing the short terminal raphe fissures and the absence of spines. Fig. 164. External detail of a valve apex focusing on the terminal raphe fissures. The external rimoportula opening is not discernible. Fig. 165. SEM external detail of a valve apex in girdle view, focusing on one of the raphe branches and the areolae on the apex. Fig. 166. Internal view in valve face view. Fig. 167. SEM internal detail of a valve apex showing the helictoglossa and the rimoportula. Fig. 168. SEM internal detail of a valve apex showing the helictoglossa. Scale bars indicate 10 µm for figs 162, 163 &amp; 166, 1 µm for figs 164, 165, 167 &amp; 168." figureDoi="http://doi.org/10.5281/zenodo.6541482" httpUri="https://zenodo.org/record/6541482/files/figure.png" pageId="11" pageNumber="272">Fig. 162</figureCitation>
). Apices with two rows of small, circular areolae (
<figureCitation id="13602A647D09801ACC22FF05FD73E10A" box="[593,690,230,257]" captionStart="FIGURES 162–168" captionStartId="10.[136,243,1769,1791]" captionTargetBox="[151,1435,190,1744]" captionTargetId="figure-21@10.[151,1435,190,1744]" captionTargetPageId="10" captionText="FIGURES 162–168. Eunotia zackenbergensis Goeyers, Van de Vijver &amp; Lange-Bertalot, sp. nov. SEM images taken from the holotype material (sample M446, Zackenberg, Greenland). Fig. 162. External view of a valve in girdle view. Note the course of the raphe branches on the mantle. Fig. 163. SEM external view in valve face view showing the short terminal raphe fissures and the absence of spines. Fig. 164. External detail of a valve apex focusing on the terminal raphe fissures. The external rimoportula opening is not discernible. Fig. 165. SEM external detail of a valve apex in girdle view, focusing on one of the raphe branches and the areolae on the apex. Fig. 166. Internal view in valve face view. Fig. 167. SEM internal detail of a valve apex showing the helictoglossa and the rimoportula. Fig. 168. SEM internal detail of a valve apex showing the helictoglossa. Scale bars indicate 10 µm for figs 162, 163 &amp; 166, 1 µm for figs 164, 165, 167 &amp; 168." figureDoi="http://doi.org/10.5281/zenodo.6541482" httpUri="https://zenodo.org/record/6541482/files/figure.png" pageId="11" pageNumber="272">Fig. 162</figureCitation>
). Spines lacking (
<figureCitation id="13602A647D09801ACD04FF05FBD5E10A" box="[887,1044,230,257]" captionStart="FIGURES 162–168" captionStartId="10.[136,243,1769,1791]" captionTargetBox="[151,1435,190,1744]" captionTargetId="figure-21@10.[151,1435,190,1744]" captionTargetPageId="10" captionText="FIGURES 162–168. Eunotia zackenbergensis Goeyers, Van de Vijver &amp; Lange-Bertalot, sp. nov. SEM images taken from the holotype material (sample M446, Zackenberg, Greenland). Fig. 162. External view of a valve in girdle view. Note the course of the raphe branches on the mantle. Fig. 163. SEM external view in valve face view showing the short terminal raphe fissures and the absence of spines. Fig. 164. External detail of a valve apex focusing on the terminal raphe fissures. The external rimoportula opening is not discernible. Fig. 165. SEM external detail of a valve apex in girdle view, focusing on one of the raphe branches and the areolae on the apex. Fig. 166. Internal view in valve face view. Fig. 167. SEM internal detail of a valve apex showing the helictoglossa and the rimoportula. Fig. 168. SEM internal detail of a valve apex showing the helictoglossa. Scale bars indicate 10 µm for figs 162, 163 &amp; 166, 1 µm for figs 164, 165, 167 &amp; 168." figureDoi="http://doi.org/10.5281/zenodo.6541482" httpUri="https://zenodo.org/record/6541482/files/figure.png" pageId="11" pageNumber="272">Figs 162, 163</figureCitation>
). Externally, raphe branches curving from almost halfway on the valve mantle onto the valve face (
<figureCitation id="13602A647D09801ACD3DFEE9FC2EE12E" box="[846,1007,266,293]" captionStart="FIGURES 162–168" captionStartId="10.[136,243,1769,1791]" captionTargetBox="[151,1435,190,1744]" captionTargetId="figure-21@10.[151,1435,190,1744]" captionTargetPageId="10" captionText="FIGURES 162–168. Eunotia zackenbergensis Goeyers, Van de Vijver &amp; Lange-Bertalot, sp. nov. SEM images taken from the holotype material (sample M446, Zackenberg, Greenland). Fig. 162. External view of a valve in girdle view. Note the course of the raphe branches on the mantle. Fig. 163. SEM external view in valve face view showing the short terminal raphe fissures and the absence of spines. Fig. 164. External detail of a valve apex focusing on the terminal raphe fissures. The external rimoportula opening is not discernible. Fig. 165. SEM external detail of a valve apex in girdle view, focusing on one of the raphe branches and the areolae on the apex. Fig. 166. Internal view in valve face view. Fig. 167. SEM internal detail of a valve apex showing the helictoglossa and the rimoportula. Fig. 168. SEM internal detail of a valve apex showing the helictoglossa. Scale bars indicate 10 µm for figs 162, 163 &amp; 166, 1 µm for figs 164, 165, 167 &amp; 168." figureDoi="http://doi.org/10.5281/zenodo.6541482" httpUri="https://zenodo.org/record/6541482/files/figure.png" pageId="11" pageNumber="272">Figs 162, 165</figureCitation>
). Terminal raphe fissures extending ca. halfway towards the dorsal side, visible on the valve face, ending in a distinct pore (
<figureCitation id="13602A647D09801ACA28FECDFB01E142" box="[1115,1216,302,329]" captionStart="FIGURES 162–168" captionStartId="10.[136,243,1769,1791]" captionTargetBox="[151,1435,190,1744]" captionTargetId="figure-21@10.[151,1435,190,1744]" captionTargetPageId="10" captionText="FIGURES 162–168. Eunotia zackenbergensis Goeyers, Van de Vijver &amp; Lange-Bertalot, sp. nov. SEM images taken from the holotype material (sample M446, Zackenberg, Greenland). Fig. 162. External view of a valve in girdle view. Note the course of the raphe branches on the mantle. Fig. 163. SEM external view in valve face view showing the short terminal raphe fissures and the absence of spines. Fig. 164. External detail of a valve apex focusing on the terminal raphe fissures. The external rimoportula opening is not discernible. Fig. 165. SEM external detail of a valve apex in girdle view, focusing on one of the raphe branches and the areolae on the apex. Fig. 166. Internal view in valve face view. Fig. 167. SEM internal detail of a valve apex showing the helictoglossa and the rimoportula. Fig. 168. SEM internal detail of a valve apex showing the helictoglossa. Scale bars indicate 10 µm for figs 162, 163 &amp; 166, 1 µm for figs 164, 165, 167 &amp; 168." figureDoi="http://doi.org/10.5281/zenodo.6541482" httpUri="https://zenodo.org/record/6541482/files/figure.png" pageId="11" pageNumber="272">Fig. 164</figureCitation>
). Single rimoportula present at one of both poles, internally located close to the helictoglossa (
<figureCitation id="13602A647D09801ACDB4FEB1FBA9E166" box="[967,1128,338,365]" captionStart="FIGURES 162–168" captionStartId="10.[136,243,1769,1791]" captionTargetBox="[151,1435,190,1744]" captionTargetId="figure-21@10.[151,1435,190,1744]" captionTargetPageId="10" captionText="FIGURES 162–168. Eunotia zackenbergensis Goeyers, Van de Vijver &amp; Lange-Bertalot, sp. nov. SEM images taken from the holotype material (sample M446, Zackenberg, Greenland). Fig. 162. External view of a valve in girdle view. Note the course of the raphe branches on the mantle. Fig. 163. SEM external view in valve face view showing the short terminal raphe fissures and the absence of spines. Fig. 164. External detail of a valve apex focusing on the terminal raphe fissures. The external rimoportula opening is not discernible. Fig. 165. SEM external detail of a valve apex in girdle view, focusing on one of the raphe branches and the areolae on the apex. Fig. 166. Internal view in valve face view. Fig. 167. SEM internal detail of a valve apex showing the helictoglossa and the rimoportula. Fig. 168. SEM internal detail of a valve apex showing the helictoglossa. Scale bars indicate 10 µm for figs 162, 163 &amp; 166, 1 µm for figs 164, 165, 167 &amp; 168." figureDoi="http://doi.org/10.5281/zenodo.6541482" httpUri="https://zenodo.org/record/6541482/files/figure.png" pageId="11" pageNumber="272">Figs 166, 167</figureCitation>
). Helictoglossa prominent at both poles (
<figureCitation id="13602A647D09801ACF78FE95FE6BE19A" box="[267,426,374,401]" captionStart="FIGURES 162–168" captionStartId="10.[136,243,1769,1791]" captionTargetBox="[151,1435,190,1744]" captionTargetId="figure-21@10.[151,1435,190,1744]" captionTargetPageId="10" captionText="FIGURES 162–168. Eunotia zackenbergensis Goeyers, Van de Vijver &amp; Lange-Bertalot, sp. nov. SEM images taken from the holotype material (sample M446, Zackenberg, Greenland). Fig. 162. External view of a valve in girdle view. Note the course of the raphe branches on the mantle. Fig. 163. SEM external view in valve face view showing the short terminal raphe fissures and the absence of spines. Fig. 164. External detail of a valve apex focusing on the terminal raphe fissures. The external rimoportula opening is not discernible. Fig. 165. SEM external detail of a valve apex in girdle view, focusing on one of the raphe branches and the areolae on the apex. Fig. 166. Internal view in valve face view. Fig. 167. SEM internal detail of a valve apex showing the helictoglossa and the rimoportula. Fig. 168. SEM internal detail of a valve apex showing the helictoglossa. Scale bars indicate 10 µm for figs 162, 163 &amp; 166, 1 µm for figs 164, 165, 167 &amp; 168." figureDoi="http://doi.org/10.5281/zenodo.6541482" httpUri="https://zenodo.org/record/6541482/files/figure.png" pageId="11" pageNumber="272">Figs 167, 168</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C341656A7D09801ACECEFE79FBBDE1D3" pageId="11" pageNumber="272" type="etymology">
<paragraph id="8BE436E17D09801ACECEFE79FBBDE1D3" blockId="11.[136,1452,158,761]" pageId="11" pageNumber="272">
<emphasis id="B92FEAF37D09801ACECEFE79FEFFE1BF" bold="true" box="[189,318,410,436]" pageId="11" pageNumber="272">Etymology</emphasis>
:—The specific epithet
<taxonomicName id="4C5B4D627D09801ACC4CFE79FD34E1BF" authorityName="Goeyers, Van de Vijver &amp; Lange-Bertalot" authorityYear="2022" box="[575,757,410,436]" class="Bacillariophyceae" family="Eunotiaceae" genus="Eunotia" higherTaxonomySource="GBIF" kingdom="Chromista" order="Eunotiales" pageId="11" pageNumber="272" phylum="Bacillariophyta" rank="species" species="zackenbergensis">
<emphasis id="B92FEAF37D09801ACC4CFE79FD34E1BF" box="[575,757,410,436]" italics="true" pageId="11" pageNumber="272">zackenbergensis</emphasis>
</taxonomicName>
refers to Zackenberg, a Danish research station in the Northeast
<collectingCountry id="F34C76717D09801ACEFBFE5DFF3CE1D3" box="[136,253,446,472]" name="Greenland" pageId="11" pageNumber="272">Greenland</collectingCountry>
National Park in northeastern
<collectingCountry id="F34C76717D09801ACC27FE5DFD0DE1D3" box="[596,716,446,472]" name="Greenland" pageId="11" pageNumber="272">Greenland</collectingCountry>
, where the species was first identified.
</paragraph>
</subSubSection>
<subSubSection id="C341656A7D09801ACECEFE01FD23E2F3" pageId="11" pageNumber="272" type="distribution">
<paragraph id="8BE436E17D09801ACECEFE01FD23E2F3" blockId="11.[136,1452,158,761]" pageId="11" pageNumber="272">
<emphasis id="B92FEAF37D09801ACECEFE01FE31E1F7" bold="true" box="[189,496,482,508]" pageId="11" pageNumber="272">Distribution and ecology</emphasis>
:—
<taxonomicName id="4C5B4D627D09801ACC60FE00FCEEE1F7" authorityName="Goeyers, Van de Vijver &amp; Lange-Bertalot" authorityYear="2022" box="[531,815,482,508]" class="Bacillariophyceae" family="Eunotiaceae" genus="Eunotia" higherTaxonomySource="GBIF" kingdom="Chromista" order="Eunotiales" pageId="11" pageNumber="272" phylum="Bacillariophyta" rank="species" species="zackenbergensis">
<emphasis id="B92FEAF37D09801ACC60FE00FCEEE1F7" box="[531,815,482,508]" italics="true" pageId="11" pageNumber="272">Eunotia zackenbergensis</emphasis>
</taxonomicName>
was collected from very wet mosses, growing in an Arctic bog pond in northeastern
<collectingCountry id="F34C76717D09801ACF86FDE5FDACE22B" box="[501,621,518,544]" name="Greenland" pageId="11" pageNumber="272">Greenland</collectingCountry>
.
<bibRefCitation id="EFCA4B107D09801ACC0BFDE4FC0EE22B" author="Van Kerckvoorde, A. &amp; Trappeniers, K. &amp; Nijs, I. &amp; Beyens, L." box="[632,975,518,545]" pageId="11" pageNumber="272" pagination="301 - 314" refId="ref11000" refString="Van Kerckvoorde, A., Trappeniers, K., Nijs, I. &amp; Beyens, L. (2000) The epiphytic diatom assemblages from terrestrial mosses in Zackenberg (Northeast Greenland). Systematics and Geography of Plants 70 (2): 301 - 314. https: // doi. org / 10.2307 / 3668649" type="journal article" year="2000">
Van Kerckvoorde
<emphasis id="B92FEAF37D09801ACD37FDE4FCBCE22B" box="[836,893,518,544]" italics="true" pageId="11" pageNumber="272">et al.</emphasis>
(2000)
</bibRefCitation>
identified the species as
<taxonomicName id="4C5B4D627D09801ACA99FDE4FE84E24F" authority="A. Cleve" authorityName="A. Cleve" class="Bacillariophyceae" family="Eunotiaceae" genus="Eunotia" higherTaxonomySource="GBIF" kingdom="Chromista" order="Eunotiales" pageId="11" pageNumber="272" phylum="Bacillariophyta" rank="species" species="fallax">
<emphasis id="B92FEAF37D09801ACA99FDE4FB3AE22B" box="[1258,1275,519,544]" italics="true" pageId="11" pageNumber="272">E</emphasis>
.
<emphasis id="B92FEAF37D09801ACB7AFDE5FA88E22B" box="[1289,1353,518,544]" italics="true" pageId="11" pageNumber="272">fallax</emphasis>
A.Cleve (
<bibRefCitation id="EFCA4B107D09801ACEE3FDC9FEFCE24F" author="Cleve, A." box="[144,317,554,581]" pageId="11" pageNumber="272" pagination="1 - 44" refId="ref8791" refString="Cleve, A. (1895) On recent freshwater diatoms from Lule Lappmark in Sweden. Bihang till Kongliga Svenska Vetenskaps-Akademiens Handlingar 21 (Afd. III, 2): 1 - 44." type="journal article" year="1895">Cleve,1895: 33</bibRefCitation>
)
</taxonomicName>
and grouped the sample (M446) in an assemblage characterised by high frequencies of
<taxonomicName id="4C5B4D627D09801ACB6AFDC9FD47E263" authority="(Hustedt) Round &amp; Bukhtiyarova" authorityName="Round &amp; Bukhtiyarova" baseAuthorityName="Hustedt" class="Bacillariophyceae" family="Achnanthidiaceae" genus="Rossithidium" higherTaxonomySource="GBIF" kingdom="Chromista" order="Cocconeidales" pageId="11" pageNumber="272" phylum="Bacillariophyta" rank="species" species="petersenii">
<emphasis id="B92FEAF37D09801ACB6AFDC9FF37E263" italics="true" pageId="11" pageNumber="272">Rossithidium petersenii</emphasis>
(Hustedt) Round &amp; Bukhtiyarova
</taxonomicName>
(
<bibRefCitation id="EFCA4B107D09801ACCEAFDADFCB5E263" author="Hustedt, F." box="[665,884,590,616]" pageId="11" pageNumber="272" pagination="152 - 207" refId="ref9448" refString="Hustedt, F. (1937) Susswasserdiatomeen von Island, Spitzbergen und den Faroer-Inseln. Botanisches Archiv 38: 152 - 207." type="journal article" year="1937">Hustedt 1937: 179</bibRefCitation>
,
<bibRefCitation id="EFCA4B107D09801ACDF7FDADFAD4E263" author="Round, F. E. &amp; Bukhtiyarova, L." box="[900,1301,590,616]" pageId="11" pageNumber="272" pagination="345 - 361" refId="ref10106" refString="Round, F. E. &amp; Bukhtiyarova, L. (1996) Four new genera based on Achnanthes (Achnanthidium) together with a re-definition of Achnanthidium. Diatom Research 11 (2): 345 - 361. http: // dx. doi. org / 10.1080 / 0269249 x. 1996.9705389" type="journal article" year="1996">Round &amp; Bukhtiyarova 1996: 178</bibRefCitation>
) and several
<taxonomicName id="4C5B4D627D09801ACEFBFD91FEA4E287" box="[136,357,626,652]" class="Bacillariophyceae" family="Pinnulariaceae" genus="Pinnularia" kingdom="Chromista" order="Naviculales" pageId="11" pageNumber="272" phylum="Bacillariophyta" rank="species" species="undetermined">
<emphasis id="B92FEAF37D09801ACEFBFD91FEC0E287" box="[136,257,626,652]" italics="true" pageId="11" pageNumber="272">Pinnularia</emphasis>
species.
</taxonomicName>
The new species is one of the most frequently observed
<taxonomicName id="4C5B4D627D09801ACA78FD90FB01E287" box="[1035,1216,626,652]" pageId="11" pageNumber="272">
<emphasis id="B92FEAF37D09801ACA78FD90FBA5E287" box="[1035,1124,627,652]" italics="true" pageId="11" pageNumber="272">Eunotia</emphasis>
species
</taxonomicName>
in
<collectingCountry id="F34C76717D09801ACA98FD91FAA2E287" box="[1259,1379,626,652]" name="Greenland" pageId="11" pageNumber="272">Greenland</collectingCountry>
, often reaching relative abundances of up to 70% of the total diatom composition and usually present in bog ponds or fens (Goeyers, unpubl. res.). Due to confusion with
<taxonomicName id="4C5B4D627D09801ACCA7FD58FC9AE2DF" authorityName="Grunow LM" authorityYear="1862" box="[724,859,698,724]" class="Bacillariophyceae" family="Eunotiaceae" genus="Eunotia" higherTaxonomySource="GBIF" kingdom="Chromista" order="Eunotiales" pageId="11" pageNumber="272" phylum="Bacillariophyta" rank="species" species="paludosa">
<emphasis id="B92FEAF37D09801ACCA7FD58FC9AE2DF" box="[724,859,698,724]" italics="true" pageId="11" pageNumber="272">E. paludosa</emphasis>
</taxonomicName>
, regularly reported from Arctic localities, its precise ecology and distribution is at present not well known.
</paragraph>
</subSubSection>
</treatment>
</document>
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