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<document ID-DOI="10.5852/ejt.2020.728.1181" ID-GBIF-Dataset="4918514a-aecb-4fa1-a019-fb5fc9083f3c" ID-Zenodo-Dep="4316883" ID-ZooBank="F8DAFD36-2878-438D-B7C0-B8D05531EC5C" approvalRequired="77" approvalRequired_for_textStreams="77" checkinTime="1607682791823" checkinUser="plazi" docAuthor="Sendra, Alberto &amp; Reboleira, Ana Sofia P. S." docDate="2020" docId="03CD87F2FC72FF96292FA0D0D922ECC1" docLanguage="en" docName="ejt-728_sendra_reboleira.pdf" docOrigin="European Journal of Taxonomy 728" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.12:EJT.2018-.journal_article" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article" docStyleVersion="12" docTitle="Litocampa Silvestri 1933" docType="treatment" docVersion="10" lastPageNumber="98" masterDocId="FFF4FF8AFC2DFFF72B78A30FDC54ED27" masterDocTitle="Euro-Mediterranean fauna of Campodeinae (Campodeidae, Diplura)" masterLastPageNumber="130" masterPageNumber="1" pageNumber="96" updateTime="1645471330625" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:titleInfo>
<mods:title>Euro-Mediterranean fauna of Campodeinae (Campodeidae, Diplura)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Sendra, Alberto</mods:namePart>
<mods:nameIdentifier type="LSID">11636BAE-AE66-4898-A7C8-35B329E7E3A8</mods:nameIdentifier>
<mods:affiliation>Coleccions Torres-Sala i Siro de Fez, Servei de Patrimoni Històric, Ajuntament de València, València, Spain. Natural History Museum of Denmark, University of Copenhagen, Universitetsparken 15, DK- 2100 København Ø, Denmark. Centre for Ecology, Evolution and Enviromental Changes (cE 3 c), Faculdade de Ciências, Universidade de Lisboa, Lisbon, Portugal.</mods:affiliation>
<mods:nameIdentifier type="email">alberto.sendra@uv.es</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Reboleira, Ana Sofia P. S.</mods:namePart>
<mods:nameIdentifier type="LSID">338DE845-4839-4EF5-B684-587C021F076C</mods:nameIdentifier>
<mods:nameIdentifier type="email">asreboleira@fc.ul.pt</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>European Journal of Taxonomy</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2020</mods:date>
<mods:detail type="pubDate">
<mods:number>2020-12-09</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>728</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>130</mods:end>
</mods:extent>
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</mods:relatedItem>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.5852/ejt.2020.728.1181</mods:identifier>
<mods:identifier type="GBIF-Dataset">4918514a-aecb-4fa1-a019-fb5fc9083f3c</mods:identifier>
<mods:identifier type="Zenodo-Dep">4316883</mods:identifier>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.4345571" ID-GBIF-Taxon="174795637" ID-Zenodo-Dep="4345571" LSID="urn:lsid:plazi:treatment:03CD87F2FC72FF96292FA0D0D922ECC1" httpUri="http://treatment.plazi.org/id/03CD87F2FC72FF96292FA0D0D922ECC1" lastPageId="97" lastPageNumber="98" pageId="95" pageNumber="96">
<subSubSection box="[599,989,991,1017]" pageId="95" pageNumber="96" type="nomenclature">
<paragraph blockId="95.[599,989,991,1017]" box="[599,989,991,1017]" pageId="95" pageNumber="96">
<heading box="[599,989,991,1017]" centered="true" fontSize="11" level="2" pageId="95" pageNumber="96" reason="2">
Genus
<taxonomicName authority="Silvestri, 1933" authorityName="Silvestri" authorityYear="1933" box="[681,989,991,1017]" class="Entognatha" family="Campodeidae" genus="Litocampa" kingdom="Animalia" order="Diplura" pageId="95" pageNumber="96" phylum="Arthropoda" rank="genus">
<emphasis bold="true" box="[681,810,991,1017]" italics="true" pageId="95" pageNumber="96">Litocampa</emphasis>
Silvestri, 1933
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[189,572,1060,1087]" pageId="95" pageNumber="96" type="reference_group">
<paragraph blockId="95.[189,572,1060,1087]" box="[189,572,1060,1087]" pageId="95" pageNumber="96">
<treatmentCitationGroup box="[189,572,1060,1087]" pageId="95" pageNumber="96">
<treatmentCitation author="Silvestri F." box="[189,567,1060,1087]" page="117" pageId="95" pageNumber="96" year="1933">
<taxonomicName authority="Silvestri, 1933 c: 117" authorityName="Silvestri" authorityPageNumber="117" authorityYear="1933" box="[189,567,1060,1087]" class="Entognatha" family="Campodeidae" genus="Litocampa" kingdom="Animalia" order="Diplura" pageId="95" pageNumber="96" phylum="Arthropoda" rank="genus">
<emphasis box="[189,314,1061,1086]" italics="true" pageId="95" pageNumber="96">Litocampa</emphasis>
<bibRefCitation author="Silvestri F." box="[322,567,1060,1087]" pageId="95" pageNumber="96" pagination="114 - 124" refId="ref69261" refString="Silvestri F. 1933 c. Spedizione del Prof. Nello Beccari nella Guiana Britannica I. Thysanura (Insecta). Bolletino del Laboratorio di Zoologia generale e agraria della R. Scuola superiore d'agricoltura in Portici 27: 114 - 124." type="journal article" year="1933">Silvestri, 1933c: 117</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection pageId="95" pageNumber="96" type="diagnosis">
<paragraph blockId="95.[189,684,1129,1156]" box="[189,684,1129,1156]" pageId="95" pageNumber="96">
<emphasis bold="true" box="[189,310,1129,1155]" pageId="95" pageNumber="96">Diagnosis</emphasis>
(
<bibRefCitation author="Silvestri F." box="[325,507,1129,1156]" pageId="95" pageNumber="96" pagination="114 - 124" refId="ref69261" refString="Silvestri F. 1933 c. Spedizione del Prof. Nello Beccari nella Guiana Britannica I. Thysanura (Insecta). Bolletino del Laboratorio di Zoologia generale e agraria della R. Scuola superiore d'agricoltura in Portici 27: 114 - 124." type="journal article" year="1933">Silvestri 1933c</bibRefCitation>
;
<bibRefCitation author="Conde B." box="[519,676,1129,1156]" pageId="95" pageNumber="96" pagination="1 - 202" refId="ref64905" refString="Conde B. 1956 a. Materiaux pour une monographie des Diploures Campodeides. Memoires du Museum national d'histoire naturelle Serie A - Zoologie 12: 1 - 202." type="journal article" year="1956">Condé 1956a</bibRefCitation>
)
</paragraph>
<paragraph blockId="95.[187,1399,1180,1556]" pageId="95" pageNumber="96">
Notal formula with 1+1
<emphasis box="[475,511,1180,1205]" italics="true" pageId="95" pageNumber="96">ma</emphasis>
, 1+1
<emphasis box="[579,602,1180,1206]" italics="true" pageId="95" pageNumber="96">la</emphasis>
, 1+1
<emphasis box="[670,693,1180,1206]" italics="true" pageId="95" pageNumber="96">lp</emphasis>
macrosetae on pronotum; 1+1
<emphasis box="[1059,1095,1180,1205]" italics="true" pageId="95" pageNumber="96">ma</emphasis>
, 0+0 or 1+1
<emphasis box="[1248,1271,1180,1206]" italics="true" pageId="95" pageNumber="96">la</emphasis>
, 1+1
<emphasis box="[1339,1362,1180,1206]" italics="true" pageId="95" pageNumber="96">lp</emphasis>
on mesonotum, 0+0 or 1+1
<emphasis box="[470,506,1219,1244]" italics="true" pageId="95" pageNumber="96">ma</emphasis>
, 0+0 or 1+1
<emphasis box="[649,672,1218,1244]" italics="true" pageId="95" pageNumber="96">lp</emphasis>
on metanotum; with or without dorsal femoral macrosetae; one or two ventral tibial macrosetae; subequal to unequal elbow-like claws with small to large sized smooth lateral crests and smooth setiform telotarsal process; 0+0, 1+1 or 2+2
<emphasis box="[1015,1038,1296,1322]" italics="true" pageId="95" pageNumber="96">lp</emphasis>
and 0+0 or 1+1
<emphasis box="[1236,1259,1296,1322]" italics="true" pageId="95" pageNumber="96">la</emphasis>
macrosetae on IIIVII urotergites; with or without medial anterior or medial posterior macrosetae on urotergites and abdominal segment; urosternite I with 7+7-6+6 macrosetae; IIVII with 6+6-4+4, although variability within each species not yet been studied. Species of
<taxonomicName authorityName="Silvestri" authorityYear="1933" box="[822,947,1414,1439]" class="Entognatha" family="Campodeidae" genus="Litocampa" kingdom="Animalia" order="Diplura" pageId="95" pageNumber="96" phylum="Arthropoda" rank="genus">
<emphasis box="[822,947,1414,1439]" italics="true" pageId="95" pageNumber="96">Litocampa</emphasis>
</taxonomicName>
described outside North America and Europe (
<bibRefCitation author="Paclt J." box="[291,419,1451,1478]" pageId="95" pageNumber="96" refId="ref66500" refString="Paclt J. 1957. Diplura. Genera Insectorum 212 E. P. Wytsman, Crainhem." type="book" year="1957">Paclt 1957</bibRefCitation>
) should be placed in other genera such as
<taxonomicName authority="Paclt, 1957" authorityName="Paclt" authorityYear="1957" box="[915,1216,1451,1478]" class="Entognatha" family="Campodeidae" genus="Cocytocampa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diplura" pageId="95" pageNumber="96" phylum="Arthropoda" rank="genus">
<emphasis box="[915,1076,1452,1478]" italics="true" pageId="95" pageNumber="96">Cocytocampa</emphasis>
<bibRefCitation author="Paclt J." box="[1084,1216,1451,1478]" pageId="95" pageNumber="96" refId="ref66500" refString="Paclt J. 1957. Diplura. Genera Insectorum 212 E. P. Wytsman, Crainhem." type="book" year="1957">Paclt, 1957</bibRefCitation>
</taxonomicName>
(
<bibRefCitation author="Conde B." box="[1231,1390,1452,1478]" pageId="95" pageNumber="96" pagination="731 - 748" refId="ref65323" refString="Conde B. 1982 a. Diploures Campodeides de Papouasie. Revue suisse de Zoologie 89 (3): 731 - 748." type="journal article" year="1982">Condé 1982a</bibRefCitation>
) or in new ones. The similarities found between some species of
<taxonomicName authorityName="Silvestri" authorityYear="1932" box="[979,1119,1490,1516]" class="Entognatha" family="Campodeidae" genus="Podocampa" kingdom="Animalia" order="Diplura" pageId="95" pageNumber="96" phylum="Arthropoda" rank="genus">
<emphasis box="[979,1119,1490,1516]" italics="true" pageId="95" pageNumber="96">Podocampa</emphasis>
</taxonomicName>
is noticeable, the only difference seems to be small lateral claws and their similar distribution areas (
<bibRefCitation author="Conde B." box="[1103,1272,1529,1556]" pageId="95" pageNumber="96" pagination="1 - 202" refId="ref64905" refString="Conde B. 1956 a. Materiaux pour une monographie des Diploures Campodeides. Memoires du Museum national d'histoire naturelle Serie A - Zoologie 12: 1 - 202." type="journal article" year="1956">Condé, 1956a</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="95" pageNumber="96" type="distribution">
<paragraph blockId="95.[189,495,1598,1625]" box="[189,495,1598,1625]" pageId="95" pageNumber="96">
<heading bold="true" box="[189,495,1598,1625]" fontSize="11" level="3" pageId="95" pageNumber="96" reason="3">
<emphasis bold="true" box="[189,495,1598,1625]" pageId="95" pageNumber="96">Habitat and distribution</emphasis>
</heading>
</paragraph>
<paragraph blockId="95.[187,1399,1649,2025]" pageId="95" pageNumber="96">
Among the 33 described species of
<taxonomicName authorityName="Silvestri" authorityYear="1933" box="[649,774,1649,1674]" class="Entognatha" family="Campodeidae" genus="Litocampa" kingdom="Animalia" order="Diplura" pageId="95" pageNumber="96" phylum="Arthropoda" rank="genus">
<emphasis box="[649,774,1649,1674]" italics="true" pageId="95" pageNumber="96">Litocampa</emphasis>
</taxonomicName>
, including the American species, 23 inhabit cave ecosystems in their deep zones and are considered cave adapted.Although there are several soil-dwelling species in Europe, e.g.,
<taxonomicName authority="(Conde, 1948)" baseAuthorityName="Conde" baseAuthorityYear="1948" box="[464,758,1726,1752]" class="Entognatha" family="Campodeidae" genus="Litocampa" kingdom="Animalia" order="Diplura" pageId="95" pageNumber="96" phylum="Arthropoda" rank="species" species="Humilis">
<emphasis box="[464,580,1726,1752]" italics="true" pageId="95" pageNumber="96">L. humilis</emphasis>
(Condé, 1948)
</taxonomicName>
,
<taxonomicName authority="(Conde, 1948)" baseAuthorityName="Conde" baseAuthorityYear="1948" box="[770,1070,1726,1752]" class="Entognatha" family="Campodeidae" genus="Litocampa" kingdom="Animalia" order="Diplura" pageId="95" pageNumber="96" phylum="Arthropoda" rank="species" species="Cognata">
<emphasis box="[770,895,1727,1752]" italics="true" pageId="95" pageNumber="96">L. cognata</emphasis>
(Condé, 1948)
</taxonomicName>
or
<taxonomicName authority="(Conde, 1948)" baseAuthorityName="Conde" baseAuthorityYear="1948" box="[1107,1393,1726,1752]" class="Entognatha" family="Campodeidae" genus="Litocampa" kingdom="Animalia" order="Diplura" pageId="95" pageNumber="96" phylum="Arthropoda" rank="species" species="Tuzetai">
<emphasis box="[1107,1215,1727,1752]" italics="true" pageId="95" pageNumber="96">L. tuzetai</emphasis>
(Condé, 1948)
</taxonomicName>
, the main life-style tendency is to inhabit caves as well as MSS (
<bibRefCitation author="Bareth C." box="[952,1102,1765,1791]" pageId="95" pageNumber="96" pagination="67 - 71" refId="ref62399" refString="Bareth C. 1983. Diploures Campodeides du milieu souterrain superficiel de la Region Ariegeoise. Memoires de Biospeologie 10: 67 - 71." type="journal article" year="1983">Bareth 1983</bibRefCitation>
;
<bibRefCitation author="Conde B." box="[1116,1276,1765,1792]" pageId="95" pageNumber="96" pagination="1 - 202" refId="ref64905" refString="Conde B. 1956 a. Materiaux pour une monographie des Diploures Campodeides. Memoires du Museum national d'histoire naturelle Serie A - Zoologie 12: 1 - 202." type="journal article" year="1956">Condé 1956a</bibRefCitation>
;
<bibRefCitation author="Conde B. &amp; Mathieu A." pageId="95" pageNumber="96" pagination="439 - 472" refId="ref65660" refString="Conde B. &amp; Mathieu A. 1957. Campodeides endoges de la region Pyreneenne. Vie et Milieu 8 (4): 439 - 472." type="journal article" year="1957">Condé &amp; Mathieu 1957</bibRefCitation>
;
<bibRefCitation author="Sendra A. &amp; Garcia Y. &amp; Weber D." box="[374,597,1804,1830]" pageId="95" pageNumber="96" pagination="216 - 226" refId="ref68177" refString="Sendra A., Garcia Y. &amp; Weber D. 2013. Campodeidae (Hexapoda, Diplura) from caves of the Grand Duchy of Luxembourg. In: Weber D. (ed.) Die H ˆ hlenfauna Luxemburgs: 216 - 226. Ferrantia 69, Musee national d'histoire naturelle, Luxembourg." type="journal article" year="2013">
Sendra
<emphasis box="[465,526,1804,1830]" italics="true" pageId="95" pageNumber="96">et al.</emphasis>
2013
</bibRefCitation>
).
<taxonomicName authorityName="Silvestri" authorityYear="1933" box="[621,746,1805,1830]" class="Entognatha" family="Campodeidae" genus="Litocampa" kingdom="Animalia" order="Diplura" pageId="95" pageNumber="96" phylum="Arthropoda" rank="genus">
<emphasis box="[621,746,1805,1830]" italics="true" pageId="95" pageNumber="96">Litocampa</emphasis>
</taxonomicName>
has a disjunt distribution on both sides of the Atlantic Ocean, it is found in karst territories and the regions surrounding the Appalachian Mountains in the
<collectingCountry box="[189,348,1882,1908]" name="United States of America" pageId="95" pageNumber="96">United States</collectingCountry>
(
<bibRefCitation author="Conde B." box="[366,527,1882,1908]" pageId="95" pageNumber="96" pagination="125 - 137" refId="ref64317" refString="Conde B. 1949 a. Campodeides cavernicoles de la region des Appalaches. Notes biospeologiques 4: 125 - 137." type="journal article" year="1949">Condé 1949a</bibRefCitation>
) and in Europe from the Cantabrian Mountains throughout the Pyrenees towards the Central Massif in
<collectingCountry box="[553,633,1921,1947]" name="France" pageId="95" pageNumber="96">France</collectingCountry>
(
<bibRefCitation author="Conde B." box="[649,811,1920,1947]" pageId="95" pageNumber="96" pagination="1 - 202" refId="ref64905" refString="Conde B. 1956 a. Materiaux pour une monographie des Diploures Campodeides. Memoires du Museum national d'histoire naturelle Serie A - Zoologie 12: 1 - 202." type="journal article" year="1956">Condé 1956a</bibRefCitation>
;
<bibRefCitation author="Sendra A. &amp; Salgado J. M. &amp; Monedero E." box="[825,1042,1921,1947]" pageId="95" pageNumber="96" pagination="69 - 80" refId="ref67995" refString="Sendra A., Salgado J. M. &amp; Monedero E. 2003. Dos nuevas especies y una subespecie de campodeidos cavernicolas de la cornisa cantabrica (Diplura Campodeidae). Animal Biodiversity and Conservation 26.2: 69 - 80." type="book chapter" year="2003">
Sendra
<emphasis box="[915,975,1921,1947]" italics="true" pageId="95" pageNumber="96">et al.</emphasis>
2003
</bibRefCitation>
) in many caves, and reaching central and eastern Europe in moisty habitats, including natural and artificial caves (
<bibRefCitation author="Conde B." box="[1228,1393,1959,1986]" pageId="95" pageNumber="96" pagination="1 - 202" refId="ref64905" refString="Conde B. 1956 a. Materiaux pour une monographie des Diploures Campodeides. Memoires du Museum national d'histoire naturelle Serie A - Zoologie 12: 1 - 202." type="journal article" year="1956">Condé 1956a</bibRefCitation>
;
<bibRefCitation author="Sendra A. &amp; Nitzu E. &amp; Sanjuan A." box="[189,404,1998,2024]" pageId="95" pageNumber="96" pagination="37 - 66" refId="ref68128" refString="Sendra A., Nitzu E. &amp; Sanjuan A. 2012. Half a century after Ionescu's work on Romanian Diplura - A faunal contribution based on material collected from karst areas. Travaux de l'Institut de Speologie &quot; Emile Racovitza &quot; 51: 37 - 66." type="journal article" year="2012">
Sendra
<emphasis box="[278,337,1998,2024]" italics="true" pageId="95" pageNumber="96">et al.</emphasis>
2012
</bibRefCitation>
) (
<figureCitation box="[428,516,1998,2025]" captionStart="Fig" captionStartId="90.[189,232,998,1024]" captionTargetBox="[189,1396,265,941]" captionTargetId="figure-190@90.[189,1398,265,941]" captionTargetPageId="90" captionText="Fig. 62. Distribution in the Euro-Mediterranean region, including Azores and Canaries ofEutrichocampa Silvestri, 1932 (green), Remycampa Condé, 1952 (black), Podocampa Silvestri, 1932 (yellow), Litocampa Silvestri, 1933 (blue), Edriocampa Silvestri, 1933 (red), Libanocampa Condé, 1955 (black brown) and Oreocampa Condé, 1950 (light brown)." figureDoi="http://doi.org/10.5281/zenodo.4316929" httpUri="https://zenodo.org/record/4316929/files/figure.png" pageId="95" pageNumber="96">Fig. 62</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection box="[189,809,264,291]" pageId="96" pageNumber="97" type="discussion">
<paragraph blockId="96.[189,809,264,291]" box="[189,809,264,291]" pageId="96" pageNumber="97">
<heading bold="true" box="[189,809,264,291]" fontSize="11" level="3" pageId="96" pageNumber="97" reason="3">
<emphasis bold="true" box="[189,809,264,291]" pageId="96" pageNumber="97">
Taxonomic key for species of the genus
<taxonomicName authorityName="Silvestri" authorityYear="1933" box="[680,809,264,290]" class="Entognatha" family="Campodeidae" genus="Litocampa" kingdom="Animalia" order="Diplura" pageId="96" pageNumber="97" phylum="Arthropoda" rank="genus">
<emphasis bold="true" box="[680,809,264,290]" italics="true" pageId="96" pageNumber="97">Litocampa</emphasis>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection lastPageId="97" lastPageNumber="98" pageId="96" pageNumber="97" type="key">
<key lastPageId="97" lastPageNumber="98" pageId="96" pageNumber="97">
<keyStep pageId="96" pageNumber="97">
<paragraph blockId="96.[189,1399,332,2025]" box="[189,1399,332,358]" pageId="96" pageNumber="97">
<keyLead box="[189,1399,332,358]" pageId="96" pageNumber="97">1. Dorsal femoral macrosetae absent..................................................................................................... 2</keyLead>
</paragraph>
<paragraph blockId="96.[189,1399,332,2025]" box="[193,1399,368,395]" pageId="96" pageNumber="97">
<keyLead box="[193,1399,368,395]" pageId="96" pageNumber="97"> Dorsal femoral macrosetae present.................................................................................................... 8</keyLead>
</paragraph>
</keyStep>
<keyStep pageId="96" pageNumber="97">
<paragraph blockId="96.[189,1399,332,2025]" pageId="96" pageNumber="97">
<keyLead pageId="96" pageNumber="97">2. Lateral posterior metanotal macrosetae 1+1; lateral anterior macrosetae of the fifth urotergite 1+1.. ........................................................................................................................................................... 3</keyLead>
</paragraph>
<paragraph blockId="96.[189,1399,332,2025]" pageId="96" pageNumber="97">
<keyLead pageId="96" pageNumber="97"> Without lateral posterior metanotal macrosetae; whithout lateral anterior macrosetae of the fifth urotergite............................................................................................................................................ 7</keyLead>
</paragraph>
</keyStep>
<keyStep pageId="96" pageNumber="97">
<paragraph blockId="96.[189,1399,332,2025]" pageId="96" pageNumber="97">
<keyLead pageId="96" pageNumber="97">
3. Male
<emphasis box="[304,327,614,646]" italics="true" pageId="96" pageNumber="97">
g
<subScript attach="left" box="[318,327,632,646]" fontSize="6" pageId="96" pageNumber="97">1</subScript>
</emphasis>
glandular setae present in two separated areas; without lateral posterior macrosetae of the fifth urotergite................................................................................
<taxonomicName authority="(Conde, 1947)" baseAuthorityName="Conde" baseAuthorityYear="1947" box="[998,1398,650,677]" class="Entognatha" family="Campodeidae" genus="Litocampa" kingdom="Animalia" order="Diplura" pageId="96" pageNumber="97" phylum="Arthropoda" rank="species" species="Vandeli">
<emphasis bold="true" box="[998,1222,650,677]" italics="true" pageId="96" pageNumber="97">Litocampa vandeli</emphasis>
(Condé, 1947)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph blockId="96.[189,1399,332,2025]" pageId="96" pageNumber="97">
<keyLead pageId="96" pageNumber="97">
Male
<emphasis box="[300,323,688,720]" italics="true" pageId="96" pageNumber="97">
g
<subScript attach="left" box="[314,323,706,720]" fontSize="6" pageId="96" pageNumber="97">1</subScript>
</emphasis>
glandular setae present in a continuous area; lateral posterior macrosetae of the fifth urotergite 1+1 ..................................................................................................................................................... 4
</keyLead>
</paragraph>
</keyStep>
<keyStep pageId="96" pageNumber="97">
<paragraph blockId="96.[189,1399,332,2025]" box="[189,1399,791,818]" pageId="96" pageNumber="97">
<keyLead box="[189,1399,791,818]" pageId="96" pageNumber="97">4. Medial anterior macrosetae of the eighth urotergite 1+1................................................................... 5</keyLead>
</paragraph>
<paragraph blockId="96.[189,1399,332,2025]" box="[193,1399,828,854]" pageId="96" pageNumber="97">
<keyLead box="[193,1399,828,854]" pageId="96" pageNumber="97"> Without medial anterior macrosetae of the eighth urotergite ............................................................ 6</keyLead>
</paragraph>
</keyStep>
<keyStep pageId="96" pageNumber="97">
<paragraph blockId="96.[189,1399,332,2025]" pageId="96" pageNumber="97">
<keyLead pageId="96" pageNumber="97">
5. Medial posterior macrosetae of the ninth abdominal segment 1+1; cave-adapted body shape; complex sensillum of the cupuliform organ..................................................
<taxonomicName authority="(Conde, 1948)" baseAuthorityName="Conde" baseAuthorityYear="1948" box="[1006,1398,932,958]" class="Entognatha" family="Campodeidae" genus="Litocampa" kingdom="Animalia" order="Diplura" pageId="96" pageNumber="97" phylum="Arthropoda" rank="species" species="Tuzetai">
<emphasis bold="true" box="[1006,1222,932,958]" italics="true" pageId="96" pageNumber="97">Litocampa tuzetai</emphasis>
(Condé, 1948)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph blockId="96.[189,1399,332,2025]" pageId="96" pageNumber="97">
<keyLead pageId="96" pageNumber="97">
Without medial posterior macrosetae of the ninth abdominal segment; body shape non-cave-adapted; simple sensillum of the cupuliform organ ................................
<taxonomicName authority="(Ionescu, 1955)" baseAuthorityName="Ionescu" baseAuthorityYear="1955" box="[963,1398,1005,1032]" class="Entognatha" family="Campodeidae" genus="Litocampa" kingdom="Animalia" order="Diplura" pageId="96" pageNumber="97" phylum="Arthropoda" rank="species" species="Montana">
<emphasis bold="true" box="[963,1207,1006,1032]" italics="true" pageId="96" pageNumber="97">Litocampa montana</emphasis>
(
<bibRefCitation author="Ionescu M. A." box="[1222,1390,1005,1032]" pageId="96" pageNumber="97" pagination="1 - 48" refId="ref66058" refString="Ionescu M. A. 1955. Diplura. Fauna Republicii Populare Romane, Insecta VII (2): 1 - 48. Rumanian Academy, Bucharest." type="journal article" year="1955">Ionescu, 1955</bibRefCitation>
)
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep pageId="96" pageNumber="97">
<paragraph blockId="96.[189,1399,332,2025]" pageId="96" pageNumber="97">
<keyLead pageId="96" pageNumber="97">
6. Medial anterior macrosetae of the first urotergite 1+1; medial anterior macrosetae of the second urotergite 1+1; medial anterior macrosetae of the third urotergite 1+1; medial anterior macrosetae of the fourth urotergite 1+1...............................................................
<taxonomicName authority="(Conde, 1948)" baseAuthorityName="Conde" baseAuthorityYear="1948" box="[989,1398,1146,1173]" class="Entognatha" family="Campodeidae" genus="Litocampa" kingdom="Animalia" order="Diplura" pageId="96" pageNumber="97" phylum="Arthropoda" rank="species" species="Cognata">
<emphasis bold="true" box="[989,1222,1146,1173]" italics="true" pageId="96" pageNumber="97">Litocampa cognata</emphasis>
(Condé, 1948)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph blockId="96.[189,1399,332,2025]" pageId="96" pageNumber="97">
<keyLead pageId="96" pageNumber="97">
Without medial anterior macrosetae in first to fourth urotergites........................................................ ......................................................................................................
<taxonomicName authority="(Conde, 1948)" baseAuthorityName="Conde" baseAuthorityYear="1948" box="[994,1398,1220,1247]" class="Entognatha" family="Campodeidae" genus="Litocampa" kingdom="Animalia" order="Diplura" pageId="96" pageNumber="97" phylum="Arthropoda" rank="species" species="Humilis">
<emphasis bold="true" box="[994,1221,1220,1247]" italics="true" pageId="96" pageNumber="97">Litocampa humilis</emphasis>
(Condé, 1948)
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep pageId="96" pageNumber="97">
<paragraph blockId="96.[189,1399,332,2025]" pageId="96" pageNumber="97">
<keyLead pageId="96" pageNumber="97">
7. Medial anterior metanotal macrosetae 1+1; lateral anterior macrosetae of the sixth urotergite 1+1; lateral posterior macrosetae on sixth and seventh urotergites 1+1
<taxonomicName authority="(Denis, 1930)" baseAuthorityName="Denis" baseAuthorityYear="1930" box="[998,1398,1324,1351]" class="Entognatha" family="Campodeidae" genus="Litocampa" kingdom="Animalia" order="Diplura" pageId="96" pageNumber="97" phylum="Arthropoda" rank="species" species="Sollaudi">
<emphasis bold="true" box="[998,1230,1324,1351]" italics="true" pageId="96" pageNumber="97">Litocampa sollaudi</emphasis>
(
<bibRefCitation author="Denis J. R." box="[1245,1390,1324,1350]" pageId="96" pageNumber="97" pagination="19 - 41" refId="ref65797" refString="Denis J. R. 1930. Sur la faune francaise des Apterygotes XIe note: Diploures avec tableau de determination des especes francaises. Bulletin Societe zoologique de France 55: 19 - 41." type="journal article" year="1930">Denis, 1930</bibRefCitation>
)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph blockId="96.[189,1399,332,2025]" pageId="96" pageNumber="97">
<keyLead pageId="96" pageNumber="97">
Without medial anterior metanotal macrosetae; without lateral anterior macrosetae of the sixth urotergite; without lateral posterior macrosetae on sixth and seventh urotergites............................... ...................................................................................................
<taxonomicName authority="(Conde, 1948)" baseAuthorityName="Conde" baseAuthorityYear="1948" box="[967,1398,1434,1461]" class="Entognatha" family="Campodeidae" genus="Litocampa" kingdom="Animalia" order="Diplura" pageId="96" pageNumber="97" phylum="Arthropoda" rank="species" species="Bourgoini">
<emphasis bold="true" box="[967,1222,1434,1461]" italics="true" pageId="96" pageNumber="97">Litocampa bourgoini</emphasis>
(Condé, 1948)
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep pageId="96" pageNumber="97">
<paragraph blockId="96.[189,1399,332,2025]" box="[189,1399,1502,1535]" pageId="96" pageNumber="97">
<keyLead box="[189,1399,1502,1535]" pageId="96" pageNumber="97">
8. Male
<emphasis box="[302,326,1503,1535]" italics="true" pageId="96" pageNumber="97">
g
<subScript attach="left" box="[317,326,1521,1535]" fontSize="6" pageId="96" pageNumber="97">1</subScript>
</emphasis>
glandular setae absent........................................................................................................... 9
</keyLead>
</paragraph>
<paragraph blockId="96.[189,1399,332,2025]" box="[193,1398,1539,1572]" pageId="96" pageNumber="97">
<keyLead box="[193,1398,1539,1572]" pageId="96" pageNumber="97">
Male
<emphasis box="[302,317,1540,1565]" italics="true" pageId="96" pageNumber="97">g</emphasis>
<emphasis box="[317,326,1558,1572]" italics="true" pageId="96" pageNumber="97">1</emphasis>
glandular setae present in a continuous area....................................................................... 10
</keyLead>
</paragraph>
</keyStep>
<keyStep pageId="96" pageNumber="97">
<paragraph blockId="96.[189,1399,332,2025]" pageId="96" pageNumber="97">
<keyLead pageId="96" pageNumber="97">
9. Medial anterior macrosetae of the first urotergite 1+1; medial anterior macrosetae on second to fourth urotergites 1+1
<emphasis box="[497,849,1644,1669]" italics="true" pageId="96" pageNumber="97">................................................</emphasis>
<taxonomicName authority="Sendra &amp; Reboleira, 2010" authorityName="Sendra &amp; Reboleira" authorityYear="2010" box="[849,1399,1642,1669]" class="Entognatha" family="Campodeidae" genus="Litocampa" kingdom="Animalia" order="Diplura" pageId="96" pageNumber="97" phylum="Arthropoda" rank="species" species="Mendesi">
<emphasis bold="true" box="[849,1084,1642,1669]" italics="true" pageId="96" pageNumber="97">Litocampa mendesi</emphasis>
<bibRefCitation author="Reboleira A. S. &amp; Sendra A. &amp; Goncalves F. &amp; Oromi P." box="[1092,1399,1643,1669]" pageId="96" pageNumber="97" pagination="50 - 56" refId="ref67057" refString="Reboleira A. S., Sendra A., Goncalves F. &amp; Oromi P. 2010. The first hypogean dipluran from Portugal: description of a new species of the genus Litocampa (Diplura: Campodeidae). Zootaxa 2728: 50 - 56." type="journal article" year="2010">Sendra &amp; Reboleira, 2010</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph blockId="96.[189,1399,332,2025]" pageId="96" pageNumber="97">
<keyLead pageId="96" pageNumber="97">
Without medial anterior macrosetae on first to fourth urotergites.......................................................
<emphasis box="[240,981,1717,1742]" italics="true" pageId="96" pageNumber="97">.....................................................................................................</emphasis>
<taxonomicName authority="(Conde, 1949)" baseAuthorityName="Conde" baseAuthorityYear="1949" box="[985,1398,1716,1743]" class="Entognatha" family="Campodeidae" genus="Litocampa" kingdom="Animalia" order="Diplura" pageId="96" pageNumber="97" phylum="Arthropoda" rank="species" species="Espanoli">
<emphasis bold="true" box="[985,1222,1716,1743]" italics="true" pageId="96" pageNumber="97">Litocampa espanoli</emphasis>
(Condé, 1949)
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep pageId="96" pageNumber="97">
<paragraph blockId="96.[189,1399,332,2025]" box="[189,1399,1783,1810]" pageId="96" pageNumber="97">
<keyLead box="[189,1399,1783,1810]" pageId="96" pageNumber="97">
10. Lateral anterior pronotal macrosetae 2+2............................
<taxonomicName authority="Bareth, 2001" authorityName="Bareth" authorityYear="2001" box="[929,1399,1783,1810]" class="Entognatha" family="Campodeidae" genus="Litocampa" kingdom="Animalia" order="Diplura" pageId="96" pageNumber="97" phylum="Arthropoda" rank="species" species="Quadrisetigera">
<emphasis bold="true" box="[929,1239,1783,1810]" italics="true" pageId="96" pageNumber="97">Litocampa quadrisetigera</emphasis>
<bibRefCitation author="Bareth C." box="[1247,1399,1784,1810]" pageId="96" pageNumber="97" pagination="41 - 44" refId="ref62680" refString="Bareth C. 2001. Litocampa quadrisetigera n. sp. (Campodeidae Diplura) de la grotte de Bidalunia. Bulletin de l'Academie lorraine des Sciences 40 (4): 41 - 44." type="journal article" year="2001">Bareth, 2001</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph blockId="96.[189,1399,332,2025]" box="[193,1398,1820,1847]" pageId="96" pageNumber="97">
<keyLead box="[193,1398,1820,1847]" pageId="96" pageNumber="97"> Lateral anterior pronotal macrosetae 1+1 .........................................................................................11</keyLead>
</paragraph>
</keyStep>
<keyStep pageId="96" pageNumber="97">
<paragraph blockId="96.[189,1399,332,2025]" pageId="96" pageNumber="97">
<keyLead pageId="96" pageNumber="97">
11. Lateral posterior macrosetae of the fourth urotergite 2+2................................................................... ...............................................................
<taxonomicName authority="Sendra, Salgado &amp; Monedero, 2003" authorityName="Sendra, Salgado &amp; Monedero" authorityYear="2003" box="[709,1399,1924,1951]" class="Entognatha" family="Campodeidae" genus="Litocampa" kingdom="Animalia" order="Diplura" pageId="96" pageNumber="97" phylum="Arthropoda" rank="species" species="Zaldivarae">
<emphasis bold="true" box="[709,969,1924,1951]" italics="true" pageId="96" pageNumber="97">Litocampa zaldivarae</emphasis>
<bibRefCitation author="Sendra A. &amp; Salgado J. M. &amp; Monedero E." box="[976,1399,1924,1951]" pageId="96" pageNumber="97" pagination="69 - 80" refId="ref67995" refString="Sendra A., Salgado J. M. &amp; Monedero E. 2003. Dos nuevas especies y una subespecie de campodeidos cavernicolas de la cornisa cantabrica (Diplura Campodeidae). Animal Biodiversity and Conservation 26.2: 69 - 80." type="book chapter" year="2003">Sendra, Salgado &amp; Monedero, 2003</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph blockId="96.[189,1399,332,2025]" box="[189,1398,1961,1988]" pageId="96" pageNumber="97">
<keyLead box="[189,1398,1961,1988]" pageId="96" pageNumber="97"> Lateral posterior macrosetae of the fourth urotergite 1+1 ............................................................... 12</keyLead>
</paragraph>
<paragraph blockId="96.[189,1399,332,2025]" box="[193,1398,1998,2025]" pageId="96" pageNumber="97">
<keyLead box="[193,1398,1998,2025]" pageId="96" pageNumber="97"> Without lateral posterior macrosetae of the fourth urotergite.......................................................... 13</keyLead>
</paragraph>
</keyStep>
<keyStep pageId="97" pageNumber="98">
<paragraph blockId="97.[189,1399,265,486]" pageId="97" pageNumber="98">
<keyLead pageId="97" pageNumber="98">
12. Lateral posterior macrosetae of the fifth to seventh urotergites 2+2; medial posterior macrosetae of the eighth urotergite 1+1..................................................................
<taxonomicName authority="Bareth, 1999" authorityName="Bareth" authorityYear="1999" box="[1014,1399,297,324]" class="Entognatha" family="Campodeidae" genus="Litocampa" kingdom="Animalia" order="Diplura" pageId="97" pageNumber="98" phylum="Arthropoda" rank="species" species="Hubarti">
<emphasis bold="true" box="[1014,1240,297,324]" italics="true" pageId="97" pageNumber="98">Litocampa hubarti</emphasis>
<bibRefCitation author="Bareth C." box="[1247,1399,298,324]" pageId="97" pageNumber="98" pagination="9 - 13" refId="ref62582" refString="Bareth C. 1999. Une nouvelle espece de Litocampa decouverte dans une grotte de Belqique (Diploures Campodeidae). Bulletin des Chercheurs de la Wallonie 39: 9 - 13." type="journal article" year="1999">Bareth, 1999</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph blockId="97.[189,1399,265,486]" pageId="97" pageNumber="98">
<keyLead pageId="97" pageNumber="98">
Lateral posterior macrosetae of the fifth to seventh urotergites 1+1; without medial posterior macrosetae of the eighth urotergite....................................................
<taxonomicName authority="Conde, 1951" authorityName="Conde" authorityYear="1951" box="[1027,1399,363,390]" class="Entognatha" family="Campodeidae" genus="Litocampa" kingdom="Animalia" order="Diplura" pageId="97" pageNumber="98" phylum="Arthropoda" rank="species" species="Henryi">
<emphasis bold="true" box="[1027,1241,363,390]" italics="true" pageId="97" pageNumber="98">Litocampa henryi</emphasis>
Condé, 1951
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep pageId="97" pageNumber="98">
<paragraph blockId="97.[189,1399,265,486]" box="[189,1398,426,460]" pageId="97" pageNumber="98">
<keyLead box="[189,1398,426,460]" pageId="97" pageNumber="98">
13. Male
<emphasis box="[303,316,428,453]" italics="true" pageId="97" pageNumber="98">a</emphasis>
<emphasis box="[317,326,446,460]" italics="true" pageId="97" pageNumber="98">2</emphasis>
glandular setae absent.......................................................
<taxonomicName authority="(Conde, 1949)" baseAuthorityName="Conde" baseAuthorityYear="1949" box="[999,1398,426,453]" class="Entognatha" family="Campodeidae" genus="Litocampa" kingdom="Animalia" order="Diplura" pageId="97" pageNumber="98" phylum="Arthropoda" rank="species" species="Drescoi">
<emphasis bold="true" box="[999,1221,426,453]" italics="true" pageId="97" pageNumber="98">Litocampa drescoi</emphasis>
(Condé, 1949)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph blockId="97.[189,1399,265,486]" box="[193,1398,459,486]" pageId="97" pageNumber="98">
<keyLead box="[193,1398,459,486]" pageId="97" pageNumber="98">
Male
<emphasis box="[302,317,461,486]" italics="true" pageId="97" pageNumber="98">a</emphasis>
glandular setae present ....................................................
<taxonomicName authority="(Conde, 1948)" baseAuthorityName="Conde" baseAuthorityYear="1948" box="[992,1398,459,486]" class="Entognatha" family="Campodeidae" genus="Litocampa" kingdom="Animalia" order="Diplura" pageId="97" pageNumber="98" phylum="Arthropoda" rank="species" species="Coiffaiti">
<emphasis bold="true" box="[992,1222,459,486]" italics="true" pageId="97" pageNumber="98">Litocampa coiffaiti</emphasis>
(Condé, 1948)
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</key>
</subSubSection>
</treatment>
</document>