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<document ID-DOI="10.5852/ejt.2020.728.1181" ID-GBIF-Dataset="4918514a-aecb-4fa1-a019-fb5fc9083f3c" ID-Zenodo-Dep="4316883" ID-ZooBank="F8DAFD36-2878-438D-B7C0-B8D05531EC5C" approvalRequired="77" approvalRequired_for_textStreams="77" checkinTime="1607682791823" checkinUser="plazi" docAuthor="Sendra, Alberto &amp; Reboleira, Ana Sofia P. S." docDate="2020" docId="03CD87F2FC10FFC92A87A583D94CE95F" docLanguage="en" docName="ejt-728_sendra_reboleira.pdf" docOrigin="European Journal of Taxonomy 728" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Campodea (Campodea) zuluetai Silvestri 1932" docType="treatment" docVersion="11" lastPageNumber="63" masterDocId="FFF4FF8AFC2DFFF72B78A30FDC54ED27" masterDocTitle="Euro-Mediterranean fauna of Campodeinae (Campodeidae, Diplura)" masterLastPageNumber="130" masterPageNumber="1" pageNumber="62" updateTime="1661889289615" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:titleInfo>
<mods:title>Euro-Mediterranean fauna of Campodeinae (Campodeidae, Diplura)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Sendra, Alberto</mods:namePart>
<mods:nameIdentifier type="LSID">11636BAE-AE66-4898-A7C8-35B329E7E3A8</mods:nameIdentifier>
<mods:affiliation>Coleccions Torres-Sala i Siro de Fez, Servei de Patrimoni Històric, Ajuntament de València, València, Spain. Natural History Museum of Denmark, University of Copenhagen, Universitetsparken 15, DK- 2100 København Ø, Denmark. Centre for Ecology, Evolution and Enviromental Changes (cE 3 c), Faculdade de Ciências, Universidade de Lisboa, Lisbon, Portugal.</mods:affiliation>
<mods:nameIdentifier type="email">alberto.sendra@uv.es</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Reboleira, Ana Sofia P. S.</mods:namePart>
<mods:nameIdentifier type="LSID">338DE845-4839-4EF5-B684-587C021F076C</mods:nameIdentifier>
<mods:nameIdentifier type="email">asreboleira@fc.ul.pt</mods:nameIdentifier>
</mods:name>
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<mods:title>European Journal of Taxonomy</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2020</mods:date>
<mods:detail type="pubDate">
<mods:number>2020-12-09</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>728</mods:number>
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<mods:start>1</mods:start>
<mods:end>130</mods:end>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.5852/ejt.2020.728.1181</mods:identifier>
<mods:identifier type="GBIF-Dataset">4918514a-aecb-4fa1-a019-fb5fc9083f3c</mods:identifier>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.4345476" ID-GBIF-Taxon="174795811" ID-Zenodo-Dep="4345476" LSID="urn:lsid:plazi:treatment:03CD87F2FC10FFC92A87A583D94CE95F" httpUri="http://treatment.plazi.org/id/03CD87F2FC10FFC92A87A583D94CE95F" lastPageId="62" lastPageNumber="63" pageId="61" pageNumber="62">
<subSubSection box="[511,1077,1676,1703]" pageId="61" pageNumber="62" type="nomenclature">
<paragraph blockId="61.[511,1077,1676,1703]" box="[511,1077,1676,1703]" pageId="61" pageNumber="62">
<heading box="[511,1077,1676,1703]" centered="true" fontSize="11" level="2" pageId="61" pageNumber="62" reason="2">
<taxonomicName authority="Silvestri, 1932" authorityName="Silvestri" authorityYear="1932" box="[511,1077,1676,1703]" class="Entognatha" family="Campodeidae" genus="Campodea" kingdom="Animalia" order="Diplura" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="zuluetai" subGenus="Campodea">
<emphasis bold="true" box="[511,898,1676,1703]" pageId="61" pageNumber="62">
<emphasis bold="true" box="[511,640,1676,1703]" italics="true" pageId="61" pageNumber="62">Campodea</emphasis>
(
<emphasis bold="true" box="[656,785,1676,1703]" italics="true" pageId="61" pageNumber="62">Campodea</emphasis>
)
<emphasis bold="true" box="[802,898,1676,1703]" italics="true" pageId="61" pageNumber="62">zuluetai</emphasis>
</emphasis>
Silvestri, 1932
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[189,906,1742,1769]" pageId="61" pageNumber="62" type="reference_group">
<paragraph blockId="61.[189,906,1742,1769]" box="[189,906,1742,1769]" pageId="61" pageNumber="62">
<taxonomicName authority="Silvestri, 1932: 128" authorityName="Silvestri" authorityPageNumber="128" authorityYear="1932" box="[189,810,1742,1769]" class="Entognatha" family="Campodeidae" genus="Campodea" kingdom="Animalia" order="Diplura" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="zuluetai" subGenus="Campodea">
<emphasis box="[189,316,1742,1768]" italics="true" pageId="61" pageNumber="62">Campodea</emphasis>
(
<emphasis box="[333,460,1742,1768]" italics="true" pageId="61" pageNumber="62">Campodea</emphasis>
)
<emphasis box="[477,570,1742,1768]" italics="true" pageId="61" pageNumber="62">zuluetai</emphasis>
Silvestri, 1932: 128
</taxonomicName>
, fig. VI.
</paragraph>
</subSubSection>
<subSubSection lastPageId="62" lastPageNumber="63" pageId="61" pageNumber="62" type="diagnosis">
<paragraph blockId="61.[189,814,1808,1835]" box="[189,814,1808,1835]" pageId="61" pageNumber="62">
<emphasis bold="true" box="[189,310,1808,1834]" pageId="61" pageNumber="62">Diagnosis</emphasis>
(
<bibRefCitation author="Silvestri F." box="[325,504,1808,1834]" pageId="61" pageNumber="62" pagination="115 - 164" refId="ref69090" refString="Silvestri F. 1932 a. Campodeidae (Thysanura) de Espana (primera parte). Eos 8: 115 - 164." type="journal article" year="1932">Silvestri 1932a</bibRefCitation>
;
<bibRefCitation author="Conde B. &amp; Mathieu A." box="[526,806,1808,1835]" pageId="61" pageNumber="62" pagination="439 - 472" refId="ref65660" refString="Conde B. &amp; Mathieu A. 1957. Campodeides endoges de la region Pyreneenne. Vie et Milieu 8 (4): 439 - 472." type="journal article" year="1957">Condé &amp; Mathieu 1957</bibRefCitation>
)
</paragraph>
<paragraph blockId="61.[188,1399,1855,2025]" pageId="61" pageNumber="62">
Body length 2.55.0 mm; epicuticle without microdenticles; long, smooth clothing setae; antennae with 2029 antennomeres; sensillum of third antennomere in dorsal or ventral position; long, barbed notal macrosetae with barbs along the distal two-thirds distal; marginal setae with long apical barbs; 1+1
<emphasis box="[1363,1399,1928,1953]" italics="true" pageId="61" pageNumber="62">ma</emphasis>
macrosetae on IVII urotergites with very variable length (all
<emphasis box="[939,975,1963,1988]" italics="true" pageId="61" pageNumber="62">ma</emphasis>
macrosetae short or slightly longer form the anterior urotergites to the posterior ones, the short
<emphasis box="[926,962,1999,2024]" italics="true" pageId="61" pageNumber="62">ma</emphasis>
macrosetae less barbed than longer
</paragraph>
<paragraph blockId="62.[188,1398,265,327]" pageId="62" pageNumber="63">
ones); 1+1
<emphasis box="[320,343,265,291]" italics="true" pageId="62" pageNumber="63">la</emphasis>
, 1+1
<emphasis box="[410,433,265,291]" italics="true" pageId="62" pageNumber="63">lp</emphasis>
on VVII urotergites; 1+1
<emphasis box="[751,787,266,291]" italics="true" pageId="62" pageNumber="63">mp</emphasis>
, 3+3
<emphasis box="[854,877,265,291]" italics="true" pageId="62" pageNumber="63">lp</emphasis>
on VIII urotergite and 1+1
<emphasis box="[1199,1235,266,291]" italics="true" pageId="62" pageNumber="63">mp</emphasis>
, 5+5
<emphasis box="[1302,1325,265,291]" italics="true" pageId="62" pageNumber="63">lp</emphasis>
on IX abdominal segment; cerci covered in long macrosetae almost from basis and a few clothing setae.
</paragraph>
<paragraph blockId="62.[188,1400,366,819]" pageId="62" pageNumber="63">
Specimens from populations inhabiting cave habitats have antennae with 3041 antennomeres, and moreover six to eight complex olfactory chemoreceptors inside the cupuliform organ (
<bibRefCitation author="Conde B." box="[1229,1394,401,428]" pageId="62" pageNumber="63" pagination="51 - 62" refId="ref64533" refString="Conde B. 1951 b. Campodeides cavernicoles de Catalogne. Speleon 2: 51 - 62." type="journal article" year="1951">Condé 1951b</bibRefCitation>
;
<bibRefCitation author="Sendra A. &amp; Moreno A." box="[189,467,437,464]" pageId="62" pageNumber="63" pagination="19 - 38" refId="ref67814" refString="Sendra A. &amp; Moreno A. 2004. El subgenero Campodea s. str. en la Peninsula Iberica (Hexapoda: Diplura: Campodeidae). Boletin de la Sociedad Entomologica Aragonesa 35: 19 - 38." type="journal article" year="2004">Sendra &amp; Moreno 2004</bibRefCitation>
). Other specimens from different populations collected at several localities in the southeast of the Iberian Peninsula show the
<emphasis box="[699,722,473,499]" italics="true" pageId="62" pageNumber="63">la</emphasis>
macrosetae on VVII urotergites in a backward insertion, almost medial posterior position (
<bibRefCitation author="Sendra A." box="[588,755,508,534]" pageId="62" pageNumber="63" pagination="35 - 51" refId="ref67391" refString="Sendra A. 1989 b. Nuevas aportaciones a la fauna de Campodeidos edaficos de la Peninsula Iberica e Islas Baleares (Diplura, Campodeidae). Boletin de la Asociacion espanola de Entomologia 13: 35 - 51." type="journal article" year="1989">Sendra 1989b</bibRefCitation>
,
<bibRefCitation author="Sendra A. &amp; Moreno A." box="[769,1056,508,535]" pageId="62" pageNumber="63" pagination="19 - 38" refId="ref67814" refString="Sendra A. &amp; Moreno A. 2004. El subgenero Campodea s. str. en la Peninsula Iberica (Hexapoda: Diplura: Campodeidae). Boletin de la Sociedad Entomologica Aragonesa 35: 19 - 38." type="journal article" year="2004">Sendra &amp; Moreno 2004</bibRefCitation>
). Two
<typeStatus box="[1136,1198,509,535]" pageId="62" pageNumber="63">types</typeStatus>
of spermatozoid fascicles in two different populations of
<taxonomicName authorityName="Silvestri" authorityYear="1932" box="[679,809,544,570]" class="Entognatha" family="Campodeidae" genus="Campodea" kingdom="Animalia" order="Diplura" pageId="62" pageNumber="63" phylum="Arthropoda" rank="species" species="zuluetai">
<emphasis box="[679,809,544,570]" italics="true" pageId="62" pageNumber="63">C. zuluetai</emphasis>
</taxonomicName>
have been described: the specimens from Ariège (
<collectingCountry box="[199,282,579,605]" name="France" pageId="62" pageNumber="63">France</collectingCountry>
) with 2429 antennomeres, sensillum of third antennomere in ventral position and
<emphasis box="[1250,1286,580,605]" italics="true" pageId="62" pageNumber="63">ma</emphasis>
urotergal macrosetae reaching the basis of marginal setae from the IV urotergite, and the specimens from the southeast of the Iberian Peninsula with
<emphasis box="[668,691,650,676]" italics="true" pageId="62" pageNumber="63">la</emphasis>
macrosetae of urotergites in backward position. Both have similar spermatozoid fascicles: small spermatozoid fascicles with 3040 μm in diameter and 3 μm thick, spiral filament with 1½ to 2 spirals, 200250 μm long and round shaped with 1 μm in widest section (
<bibRefCitation author="Bareth C." box="[199,350,757,783]" pageId="62" pageNumber="63" pagination="2 - 108" refId="ref62716" refString="Bareth C. 2006. Les Campodes de France (Diploures Campodeides). Memoires de Biospeologie, numero hors serie: 2 - 108." type="book chapter" year="2006">Bareth 2006</bibRefCitation>
;
<bibRefCitation author="Sendra A." box="[365,519,757,783]" pageId="62" pageNumber="63" refId="ref67316" refString="Sendra A. 1988. Taxonomia, Filogenia y Biogeografia de la Fauna de Campodeidos Iberica, Balear y Canaria (Hexapoda, Diplura, Campodeidae). PhD thesis, Biological Science Faculty, Valencia University, Spain." type="book" year="1988">Sendra 1988</bibRefCitation>
). It is clear to conclude that such a high degree of variability in different populations in
<taxonomicName authorityName="Silvestri" authorityYear="1932" box="[363,491,792,819]" class="Entognatha" family="Campodeidae" genus="Campodea" kingdom="Animalia" order="Diplura" pageId="62" pageNumber="63" phylum="Arthropoda" rank="species" species="zuluetai">
<emphasis box="[363,491,792,819]" italics="true" pageId="62" pageNumber="63">C. zuluetai</emphasis>
</taxonomicName>
will need to be throroughly reviewed.
</paragraph>
</subSubSection>
<subSubSection pageId="62" pageNumber="63" type="distribution">
<paragraph blockId="62.[189,495,858,884]" box="[189,495,858,884]" pageId="62" pageNumber="63">
<heading bold="true" box="[189,495,858,884]" fontSize="11" level="3" pageId="62" pageNumber="63" reason="3">
<emphasis bold="true" box="[189,495,858,884]" pageId="62" pageNumber="63">Habitat and distribution</emphasis>
</heading>
</paragraph>
<paragraph blockId="62.[187,1399,905,1144]" pageId="62" pageNumber="63">
Inhabits soil and cave environments. It has been collected in caves and found under scree slopes (MSS) in Ariège, Pyrenées (
<bibRefCitation author="Bareth C." box="[427,573,940,966]" pageId="62" pageNumber="63" pagination="67 - 71" refId="ref62399" refString="Bareth C. 1983. Diploures Campodeides du milieu souterrain superficiel de la Region Ariegeoise. Memoires de Biospeologie 10: 67 - 71." type="journal article" year="1983">Bareth 1983</bibRefCitation>
) and Guadarrama National Park (
<bibRefCitation author="Sendra A. &amp; Jimenez-Valverde A. &amp; Gilgado J. D. &amp; Ledesma E. &amp; Baquero E. &amp; Perez-Suarez G. &amp; Cuesta E. &amp; Herrero-Borgonon J. J. &amp; Jordana R. &amp; Tinaut A. &amp; Barranco P. &amp; Ortuno V. M." box="[958,1186,940,966]" pageId="62" pageNumber="63" pagination="61 - 80" refId="ref68345" refString="Sendra A., Jimenez-Valverde A., Gilgado J. D., Ledesma E., Baquero E., Perez-Suarez G., Cuesta E., Herrero-Borgonon J. J., Jordana R., Tinaut A., Barranco P. &amp; Ortuno V. M. 2017 a. Diplurans of subsurface terrestrial habitats in the Iberian Peninsula, with a new species description (Diplura: Campodeidae). Zootaxa 4291 (1): 61 - 80. https: // doi. org / 10.11646 / zootaxa. 4291.1.4" type="journal article" year="2017">
Sendra
<emphasis box="[1046,1103,940,966]" italics="true" pageId="62" pageNumber="63">et al.</emphasis>
2017a
</bibRefCitation>
). Some specimens collected in caves show hyperthrophy of the cave-adapted traits. It was also found at high-altitude,
<quantity box="[189,278,1011,1038]" metricMagnitude="3" metricUnit="m" metricValue="2.785" pageId="62" pageNumber="63" unit="m" value="2785.0">2785 m</quantity>
a.s.l. in the Cangou peak, Pyrenees (
<bibRefCitation author="Conde B. &amp; Mathieu A." box="[712,997,1011,1038]" pageId="62" pageNumber="63" pagination="439 - 472" refId="ref65660" refString="Conde B. &amp; Mathieu A. 1957. Campodeides endoges de la region Pyreneenne. Vie et Milieu 8 (4): 439 - 472." type="journal article" year="1957">Condé &amp; Mathieu 1957</bibRefCitation>
). It is distributed in a large area in the eastern Iberian Peninsula, from the French Pyrenees to the Baetic Mountains, including the Central System Massif (
<bibRefCitation author="Conde B." box="[380,544,1082,1109]" pageId="62" pageNumber="63" pagination="51 - 62" refId="ref64533" refString="Conde B. 1951 b. Campodeides cavernicoles de Catalogne. Speleon 2: 51 - 62." type="journal article" year="1951">Condé 1951b</bibRefCitation>
;
<bibRefCitation author="Conde B. &amp; Mathieu A." box="[557,842,1082,1109]" pageId="62" pageNumber="63" pagination="439 - 472" refId="ref65660" refString="Conde B. &amp; Mathieu A. 1957. Campodeides endoges de la region Pyreneenne. Vie et Milieu 8 (4): 439 - 472." type="journal article" year="1957">Condé &amp; Mathieu 1957</bibRefCitation>
;
<bibRefCitation author="Sendra A. &amp; Moreno A." box="[856,1142,1082,1109]" pageId="62" pageNumber="63" pagination="19 - 38" refId="ref67814" refString="Sendra A. &amp; Moreno A. 2004. El subgenero Campodea s. str. en la Peninsula Iberica (Hexapoda: Diplura: Campodeidae). Boletin de la Sociedad Entomologica Aragonesa 35: 19 - 38." type="journal article" year="2004">Sendra &amp; Moreno 2004</bibRefCitation>
). It was also found in two caves in
<collectingCountry box="[344,451,1118,1144]" name="Spain" pageId="62" pageNumber="63">Menorca</collectingCountry>
, and in Sardinia where it seems to be abundant in the soil (
<bibRefCitation author="Bareth C." box="[1141,1291,1118,1144]" pageId="62" pageNumber="63" pagination="121 - 135" refId="ref62373" refString="Bareth C. 1980. Campodeides endoges de Sardaigne recoltes par R. Dallai (Insecta Diplura). Redia 63: 121 - 135." type="journal article" year="1980">Bareth 1980</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>