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<mods:title id="2EE2924E1CEE5963ED5FDB0B31E831EF">A new, to date endemic, family of dragonfly in the mid-Cretaceous fossil fish Konservat-Lagerstätte of Haqel, Lebanon (Odonata: Anisoptera)</mods:title>
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<mods:namePart id="0A69D2D7ADB39870C8EE1A77A5A0278C">Azar, Dany</mods:namePart>
<mods:nameIdentifier id="F05191E2A3FEAAAB339EBCDF67B523B6" type="ORCID">0000-0002-4485-197X</mods:nameIdentifier>
<mods:affiliation id="9711B5FE25CAA13C8EEEBA768121B11D">State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences, Nanjing, Jiangsu, 210008, China &amp; Department of Natural Sciences, Faculty of Sciences II, Lebanese University, P. O. Box: 26110217, Fanar, Matn, Lebanon</mods:affiliation>
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<mods:namePart id="52A80112EFE6C169272B7618418B6F6C">Maksoud, Sibelle</mods:namePart>
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<mods:affiliation id="97A249FA6BD85F5521F16A2365B83863">State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences, Nanjing, Jiangsu, 210008, China &amp; Department of Natural Sciences, Faculty of Sciences II, Lebanese University, P. O. Box: 26110217, Fanar, Matn, Lebanon</mods:affiliation>
<mods:nameIdentifier id="4E1A7D82FF416931915708184D1D66DA" type="email">sibelle.maksoud@ul.edu.lb</mods:nameIdentifier>
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<mods:namePart id="D4852D1435AD6E612EB538CEA6AD6ABA">Abi-Saad, Pierre</mods:namePart>
<mods:affiliation id="D734625CB7240F43D6B311A5497854A6">Memory of Time, Byblos old sook, Byblos, Lebanon</mods:affiliation>
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<mods:namePart id="FD35204F87646380CCF1C7C30A599463">Nel, Andre</mods:namePart>
<mods:affiliation id="57E29F998475DB679D4EAB050048A474">Institut de Systématique, Evolution, Biodiversité, ISYEB-UMR 7205 - CNRS, MNHN, UPMC, EPHE, Muséum National dHistoire Naturelle, Sorbonne Universités, 57 Rue Cuvier, CP 50, Entomologie, F- 75005, Paris, France</mods:affiliation>
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<mods:date id="6B0038DCB34F68736CB4178215DE0301">2024</mods:date>
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<paragraph id="8BDB36FAD2391902FF1EF096FDD13F0E" blockId="2.[151,964,1519,1581]" box="[151,582,1519,1545]" pageId="2" pageNumber="144">
<heading id="D0938196D2391902FF1EF096FDD13F0E" bold="true" box="[151,582,1519,1545]" fontSize="11" level="1" pageId="2" pageNumber="144" reason="1">
<emphasis id="B910EAE8D2391902FF1EF096FDD13F0E" bold="true" box="[151,582,1519,1545]" pageId="2" pageNumber="144">
Family
<taxonomicName id="4C644D79D2391902FF7EF096FE5A3F0E" authorityName="Azar &amp; Maksoud &amp; Abi-Saad &amp; Nel" authorityYear="2024" box="[247,461,1519,1545]" class="Insecta" family="Libanoaeshnidae" higherTaxonomySource="CoL" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="144" phylum="Arthropoda" rank="family" status="fam. nov.">Libanoaeshnidae</taxonomicName>
<taxonomicNameLabel id="A2235793D2391902FE5AF096FDD13F0E" box="[467,582,1519,1545]" pageId="2" pageNumber="144" rank="family">fam. nov.</taxonomicNameLabel>
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urn:lsid:zoobank.org:act:
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<subSubSection id="C37E6571D2391902FF1EF322FD473F9E" pageId="2" pageNumber="144" type="type_taxon">
<paragraph id="8BDB36FAD2391902FF1EF322FCDA3F72" blockId="2.[151,1437,1627,1977]" box="[151,845,1627,1653]" pageId="2" pageNumber="144">
<emphasis id="B910EAE8D2391902FF1EF322FEB33F72" bold="true" box="[151,292,1627,1653]" pageId="2" pageNumber="144">
<typeStatus id="54DF8858D2391902FF1EF322FF453F72" box="[151,210,1627,1653]" pageId="2" pageNumber="144">Type</typeStatus>
genus.
</emphasis>
<taxonomicName id="4C644D79D2391902FEA2F322FE5D3F72" authorityName="Azar &amp; Maksoud &amp; Abi-Saad &amp; Nel" authorityYear="2024" box="[299,458,1627,1653]" class="Insecta" family="Aeshnidae" genus="Libanoaeshna" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="144" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B910EAE8D2391902FEA2F322FE5D3F72" box="[299,458,1627,1653]" italics="true" pageId="2" pageNumber="144">Libanoaeshna</emphasis>
</taxonomicName>
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<emphasis id="B910EAE8D2391902FE58F322FDA23F72" bold="true" box="[465,565,1627,1653]" pageId="2" pageNumber="144">gen. nov</emphasis>
.
</taxonomicNameLabel>
, by present designation.
</paragraph>
<paragraph id="8BDB36FAD2391902FF4EF306FD473F9E" blockId="2.[151,1437,1627,1977]" box="[199,720,1663,1689]" pageId="2" pageNumber="144">
<emphasis id="B910EAE8D2391902FF4EF306FEF53F9E" bold="true" box="[199,354,1663,1689]" pageId="2" pageNumber="144">
<typeStatus id="54DF8858D2391902FF4EF306FE953F9E" box="[199,258,1663,1689]" pageId="2" pageNumber="144">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="4C644D79D2391902FEE0F306FDF83F9E" authorityName="Azar &amp; Maksoud &amp; Abi-Saad &amp; Nel" authorityYear="2024" box="[361,623,1663,1689]" class="Insecta" family="Aeshnidae" genus="Libanoaeshna" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="144" phylum="Arthropoda" rank="species" species="mikhaeli" status="sp. nov.">
<emphasis id="B910EAE8D2391902FEE0F306FDF83F9E" box="[361,623,1663,1689]" italics="true" pageId="2" pageNumber="144">Libanoaeshna mikhaeli</emphasis>
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<emphasis id="B910EAE8D2391902FDFFF306FD473F9E" bold="true" box="[630,720,1663,1689]" pageId="2" pageNumber="144">
<taxonomicNameLabel id="A2235793D2391902FDFFF306FD473F9E" box="[630,720,1663,1689]" pageId="2" pageNumber="144" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
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<subSubSection id="C37E6571D2391902FF4EF3DAFAF83FBA" box="[199,1391,1699,1727]" pageId="2" pageNumber="144" type="etymology">
<paragraph id="8BDB36FAD2391902FF4EF3DAFAF83FBA" blockId="2.[151,1437,1627,1977]" box="[199,1391,1699,1727]" pageId="2" pageNumber="144">
<emphasis id="B910EAE8D2391902FF4EF3DAFED93FBA" bold="true" box="[199,334,1699,1725]" pageId="2" pageNumber="144">Etymology.</emphasis>
Named after
<emphasis id="B910EAE8D2391902FE6EF3DAFDD63FBA" box="[487,577,1699,1725]" italics="true" pageId="2" pageNumber="144">Libanos</emphasis>
=
<collectingCountry id="F373766AD2391902FDD6F3DAFD563FBA" box="[607,705,1699,1725]" name="Lebanon" pageId="2" pageNumber="144">Lebanon</collectingCountry>
in Greek ‘Λίβανος’ and the suffix aeshna. Gender feminine.
</paragraph>
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<subSubSection id="C37E6571D2391902FF4EF3BEFD1F3EBE" pageId="2" pageNumber="144" type="diagnosis">
<paragraph id="8BDB36FAD2391902FF4EF3BEFD1F3EBE" blockId="2.[151,1437,1627,1977]" pageId="2" pageNumber="144">
<emphasis id="B910EAE8D2391902FF4EF3BEFED53FE6" bold="true" box="[199,322,1735,1761]" pageId="2" pageNumber="144">Diagnosis.</emphasis>
Forewing characters only. Putative autapomorphic characters of the new fossil are as follows: forewing discoidal triangle very long and narrow, with vein MAb having a very strong angle and a strong curvature; a very strong trigonal planate (convergences with the
<taxonomicName id="4C644D79D2391902FD71F276FB033E2E" authority="Bechly et al., 2001" authorityName="Bechly" authorityYear="2001" box="[760,1172,1807,1833]" class="Insecta" family="Liupanshaniidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="144" phylum="Arthropoda" rank="family">
Liupanshaniidae
<bibRefCitation id="EFF54B0BD2391902FC30F276FB033E2E" author="Bechly, G. &amp; Nel, A. &amp; Martinez-Delclos, X. &amp; Jarzembowski, E. A. &amp; Coram, R. &amp; Martill, D. &amp; Fleck, G. &amp; Escuillie, F. &amp; Wisshak, M. M. &amp; Maisch, M." box="[953,1172,1807,1833]" pageId="2" pageNumber="144" pagination="1 - 230" refId="ref4400" refString="Bechly, G., Nel, A., Martinez-Delclos, X., Jarzembowski, E. A., Coram, R., Martill, D., Fleck, G., Escuillie, F., Wisshak, M. M. &amp; Maisch, M. (2001) Revision and phylogenetic study of Mesozoic Aeshnoptera, with description of numerous new taxa (Insecta: Odonata: Anisoptera). Neue Palaontologische Abhandlungen, 4, 1 - 230." type="journal article" year="2001">
Bechly
<emphasis id="B910EAE8D2391902FB99F269FBD53E2E" box="[1040,1090,1807,1833]" italics="true" pageId="2" pageNumber="144">et al</emphasis>
., 2001
</bibRefCitation>
</taxonomicName>
); area between MP and posterior margin of wing broad; CuA short, strongly curved and without distinct posterior branches; an intercalary longitudinal vein between AA and AP; antenodal crossveins very numerous between Ax1 and Ax2 and distad Ax2; cells and intercalary crossveins in area below IR2 very numerous; area between RP1 and RP2 very narrow up to pterostigma; pterostigma in a basal position.
</paragraph>
</subSubSection>
</treatment>
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