treatments-xml/data/03/CD/87/03CD87BBFF8B431AFF47F96AFE3CFDCE.xml

<document id="CCB071C2EC12424F36ECBC600463570C" ID-CLB-Dataset="27827" ID-DOI="10.11646/zootaxa.4571.3.10" ID-GBIF-Dataset="b70119b4-6b49-4e87-a1b4-ff33339ed699" ID-ISSN="1175-5326" ID-Zenodo-Dep="2612657" ID-ZooBank="3D722F87-0901-438A-AFCC-7872874F7587" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1553699415132" checkinUser="plazi" docAuthor="Gisondi, Silvia, Lenzi, Alice, Ziegler, Joachim, Giulio, Andrea Di &amp; Cerretti, Pierfilippo" docDate="2019" docId="03CD87BBFF8B431AFF47F96AFE3CFDCE" docLanguage="en" docName="zootaxa.4571.3.10.pdf" docOrigin="Zootaxa 4571 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Stevenia gilasiani Gisondi &amp; Lenzi &amp; Ziegler &amp; Giulio &amp; Cerretti 2019, sp. nov." docType="treatment" docVersion="8" lastPageNumber="429" masterDocId="FFF4FFC3FF89431CFFD0FFDDFF98FFF3" masterDocTitle="Stevenia gilasiani sp. nov. (Diptera: Rhinophoridae): the first woodlouse fly with male sexual-patches" masterLastPageNumber="431" masterPageNumber="423" pageNumber="425" updateTime="1698714610456" updateUser="ExternalLinkService">
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<mods:title id="72B11998EC7E0A96F11CEDC0FB2D0613">Stevenia gilasiani sp. nov. (Diptera: Rhinophoridae): the first woodlouse fly with male sexual-patches</mods:title>
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<taxonomicName id="4C644D2EFF8B431EFF47F96AFD5FF921" authority="Ziegler, Gisondi &amp; Cerretti" authorityName="Gisondi &amp; Lenzi &amp; Ziegler &amp; Giulio &amp; Cerretti" authorityYear="2019" box="[151,711,1719,1746]" class="Insecta" family="Tachinidae" genus="Stevenia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="425" phylum="Arthropoda" rank="species" species="gilasiani" status="sp. nov.">
<emphasis id="B910EABFFF8B431EFF47F96AFEF4F922" bold="true" box="[151,364,1719,1745]" italics="true" pageId="2" pageNumber="425">Stevenia gilasiani</emphasis>
Ziegler, Gisondi &amp; Cerretti
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, 
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<paragraph id="8BDB36ADFF8B431EFF47F906FE81F900" blockId="2.[151,818,1719,1780]" box="[151,281,1755,1780]" pageId="2" pageNumber="425">
(
<figureCitation id="135F2A28FF8B431EFF4FF906FE88F900" box="[159,272,1755,1780]" captionStart-0="FIGURES 2–5" captionStart-1="FIGURES 6–11" captionStart-2="FIGURES 12–14" captionStartId-0="3.[151,264,1976,1999]" captionStartId-1="4.[151,264,1121,1144]" captionStartId-2="5.[151,264,1777,1800]" captionTargetBox-0="[153,1435,807,1955]" captionTargetBox-1="[161,1413,209,1090]" captionTargetBox-2="[245,1340,379,1725]" captionTargetId-0="figure@3.[151,1435,807,1955]" captionTargetId-1="figure@4.[150,1436,193,1101]" captionTargetId-2="figure@5.[209,1378,348,1756]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionText-0="FIGURES 2–5. Stevenia gilasiani Ziegler, Gisondi &amp; Cerretti sp. nov. (holotype ♂). 2. Habitus in lateral view. 3. Head in lateral view. 4. Right wing in dorsal view. 5. Head in frontal view." captionText-1="FIGURES 6–11. Stevenia gilasiani sp. nov.. 6–8. Holotype ♂. 6. Detail of thorax, showing two postpronotal setae. 7. Abdomen in dorsal view. 8. Abdomen in lateral view, showing the sexual-patches on tergites 3 and 4. 9–11. Details of cuticle and setae composing the sexual-patches, using scanning electron microscopy (SEM) (paratype, MZUR). 9. Sexual-patches of tergite 3 in ventral view. 10. Seta and porous area. 11. Internal surface of cuticle at of the base of a seta composing a sexualpatch, showing the sieve-like structure at level of porous area." captionText-2="FIGURES 12–14. Stevenia gilasiani Ziegler, Gisondi &amp; Cerretti, sp. nov., terminalia (paratype, MZUR). 12. Lateral view. 13. Dorsolateral view. 14. Detail of the terminalia in ventral view. Abbreviations: aj = ejaculatory apodeme; app = phallus apodeme; c = cerci; ep = epandrium; hyp = hypandrium; pr = pregonite; ps = postgonite; s = surstylus; sd = sperm ducts." httpUri-0="https://zenodo.org/record/2612661/files/figure.png" httpUri-1="https://zenodo.org/record/2612663/files/figure.png" httpUri-2="https://zenodo.org/record/2612665/files/figure.png" pageId="2" pageNumber="425">Figs 2–14</figureCitation>
)
</paragraph>
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<emphasis id="B910EABFFF8B431EFF47F8FEFEDEF8CF" bold="true" box="[151,326,1827,1852]" pageId="2" pageNumber="425">Type material.</emphasis>
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: 
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, 
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, south of 
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data: “ 
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prov. / 
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(
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) 
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/ S of 
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, 
<location id="8EBB6076FF8B431EFB85F89BFB5AF8AC" LSID="urn:lsid:plazi:treatment:03CD87BBFF8B431AFF47F96AFE3CFDCE:8EBB6076FF8B431EFB85F89BFB5AF8AC" box="[1109,1218,1862,1887]" country="Iran" county="Ulugh Dagh mountain range" latitude="37.16439" longLatPrecision="1" longitude="44.885696" municipality="Urmia. Label" name="Mountain" pageId="2" pageNumber="425" stateProvince="Azarbayjan-e Gharbi">Mountain</location>
top / 
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/ legit 
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” (
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, same data as holotype (
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<collectionCode id="ED75AE68FF8B431EFC63F84DFB9FF854" box="[947,1031,1936,1959]" country="Italy" httpUri="http://grbio.org/cool/ek3c-66y7" name="Museo di Zoologia dell'Universita &amp;quot;La Sapienza&amp;quot;" pageId="2" pageNumber="425">MZUR</collectionCode>
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<collectionCode id="ED75AE68FF8B431EFB9FF852FB38F854" box="[1103,1184,1935,1959]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34840" lsid="urn:lsid:biocol.org:col:34840" name="Staatliches Museum fuer Naturkund Stuttgart" pageId="2" pageNumber="425">SMNS</collectionCode>
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; 
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- 
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/ Oshnavieh-Dizaj-Kulabi / 
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0 4.1 / 
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. / leg. 
<collectorName id="2691537BFF8B431EFBECF869FB20F83F" box="[1084,1208,1971,1996]" pageId="2" pageNumber="425">E. Gilasian</collectorName>
/ 
<date id="FFDA106DFF8B431EFB1DF86EFACFF838" box="[1229,1367,1971,1995]" pageId="2" pageNumber="425" value="2008-07-24">
<collectingDate id="EF9EE985FF8B431EFB1DF86EFACFF838" box="[1229,1367,1971,1995]" pageId="2" pageNumber="425" value="2008-07-24">24.VII.2008</collectingDate>
</date>
” (
<quantity id="4C9C9B48FF8B431EFAA4F86EFA04F83F" box="[1396,1436,1971,1996]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="2" pageNumber="425" unit="in" value="1.0">1 in</quantity>
<collectionCode id="ED75AE68FF8B431EFF47F805FF73F81C" box="[151,235,2008,2031]" country="Italy" httpUri="http://grbio.org/cool/ek3c-66y7" name="Museo di Zoologia dell'Universita &amp;quot;La Sapienza&amp;quot;" pageId="2" pageNumber="425">MZUR</collectionCode>
, 
<quantity id="4C9C9B48FF8B431EFF26F80AFEB9F81C" box="[246,289,2006,2031]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="2" pageNumber="425" unit="in" value="4.0">4 in</quantity>
HMIM) [
<collectingMethod id="52254EBAFF8B431EFE5EF80BFD3AF81C" box="[398,674,2006,2031]" pageId="2" pageNumber="425">all collected by hand net</collectingMethod>
on stones and rocks]
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C37E6526FF8A431FFF17FF4AFCE4FF04" pageId="3" pageNumber="426" type="etymology">
<paragraph id="8BDB36ADFF8A431FFF17FF4AFCE4FF04" blockId="3.[151,1436,151,755]" pageId="3" pageNumber="426">
<emphasis id="B910EABFFF8A431FFF17FF4AFED6FF43" bold="true" box="[199,334,151,176]" pageId="3" pageNumber="426">Etymology.</emphasis>
The specific epithet is chosen in honour of our Iranian colleague Ebrahim Gilasian, who collected 
<specimenCount id="9D62FD24FF8A431FFF47FF61FE80FF26" box="[151,280,188,213]" pageId="3" pageNumber="426" type="generic" typeStatus="paratypes">5 paratypes</specimenCount>
of this species. Furthermore, one of us (JZ) is thankful for Ebrahim’s long staying friendship, collegial cooperation and organization of wonderful collecting trips in 
<collectingCountry id="F373763DFF8A431FFC97FF3DFCE0FF04" box="[839,888,224,247]" name="Iran" pageId="3" pageNumber="426">Iran</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C37E6526FF8A431FFF17FED9FC06FEB3" pageId="3" pageNumber="426" type="diagnosis">
<paragraph id="8BDB36ADFF8A431FFF17FED9FC06FEB3" blockId="3.[151,1436,151,755]" pageId="3" pageNumber="426">
<emphasis id="B910EABFFF8A431FFF17FED9FED9FEEE" bold="true" box="[199,321,260,285]" pageId="3" pageNumber="426">Diagnosis.</emphasis>
Two postpronotal setae, instead of the usual three arranged in a distinct triangle. Male with ventral sexual-patches on abdominal tergites 3, 4 and 5 and sternites 3 and 4.
</paragraph>
</subSubSection>
<subSubSection id="C37E6526FF8A431AFF17FE91FE06FE06" lastPageId="6" lastPageNumber="429" pageId="3" pageNumber="426" type="description">
<paragraph id="8BDB36ADFF8A431FFF17FE91FB8EFE96" blockId="3.[151,1436,151,755]" box="[199,1046,332,357]" pageId="3" pageNumber="426">
<emphasis id="B910EABFFF8A431FFF17FE91FEC1FE96" bold="true" box="[199,345,332,357]" pageId="3" pageNumber="426">Description.</emphasis>
<emphasis id="B910EABFFF8A431FFE8FFE91FE01FE96" box="[351,409,332,357]" italics="true" pageId="3" pageNumber="426">Male</emphasis>
(measurements in square brackets refer to the 
<typeStatus id="54DF880FFF8A431FFC73FE91FB92FE96" box="[931,1034,332,357]" pageId="3" pageNumber="426" type="holotype">holotype</typeStatus>
).
</paragraph>
<paragraph id="8BDB36ADFF8A431FFF17FEB2FE0DFE74" blockId="3.[151,1436,151,755]" box="[199,405,367,392]" pageId="3" pageNumber="426">Length: [6.7] mm.</paragraph>
<paragraph id="8BDB36ADFF8A431FFF17FE49FB10FD00" blockId="3.[151,1436,151,755]" pageId="3" pageNumber="426">
Colour (
<figureCitation id="135F2A28FF8A431FFEF3FE49FE11FE5F" box="[291,393,404,429]" captionStart-0="FIGURES 2–5" captionStart-1="FIGURES 6–11" captionStartId-0="3.[151,264,1976,1999]" captionStartId-1="4.[151,264,1121,1144]" captionTargetBox-0="[153,1435,807,1955]" captionTargetBox-1="[161,1413,209,1090]" captionTargetId-0="figure@3.[151,1435,807,1955]" captionTargetId-1="figure@4.[150,1436,193,1101]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionText-0="FIGURES 2–5. Stevenia gilasiani Ziegler, Gisondi &amp; Cerretti sp. nov. (holotype ♂). 2. Habitus in lateral view. 3. Head in lateral view. 4. Right wing in dorsal view. 5. Head in frontal view." captionText-1="FIGURES 6–11. Stevenia gilasiani sp. nov.. 6–8. Holotype ♂. 6. Detail of thorax, showing two postpronotal setae. 7. Abdomen in dorsal view. 8. Abdomen in lateral view, showing the sexual-patches on tergites 3 and 4. 9–11. Details of cuticle and setae composing the sexual-patches, using scanning electron microscopy (SEM) (paratype, MZUR). 9. Sexual-patches of tergite 3 in ventral view. 10. Seta and porous area. 11. Internal surface of cuticle at of the base of a seta composing a sexualpatch, showing the sieve-like structure at level of porous area." httpUri-0="https://zenodo.org/record/2612661/files/figure.png" httpUri-1="https://zenodo.org/record/2612663/files/figure.png" pageId="3" pageNumber="426">Figs 2–8</figureCitation>
). Head black in ground colour, covered with silvery-grey microtomentum; frontal vitta black; area between gena and parafacial dark brown; scape and pedicel orange (same colour as abdomen laterally) to brownish and postpedicel mostly dark brown, shading to yellowish at junction with pedicel; palpus yellow. Thorax black, mostly covered with grey microtomentum, with three broad presutural dark vittae in posterior view. Legs black in ground colour. Upper and lower calypters whitish. Wing hyaline; area between vein R 
<subScript id="17E034E8FF8A431FFB68FDEEFB48FDB2" attach="left" box="[1208,1232,563,577]" fontSize="6" pageId="3" pageNumber="426">2+3</subScript>
and anterior wing margin slightly brownish infuscated; tegula reddish-brown, basicosta yellow; veins black or dark brown. Abdomen largely reddish-orange laterally, with a mediodorsal dark vitta interrupted mediodorsally; syntergite 1+2 and tergite 3 orange laterally, with orange portion not reaching posterior margin; tergite 4 black dorsally and slightly reddish anterolaterally; tergite 5 black; tergites 3–4 each with a band of whitish microtomentum on anterior 1/4 (dorsally) to 1/2 (laterally), syntergite 1+2 with diffuse whitish microtomentum; terminalia blackish.
</paragraph>
<caption id="DF1B6625FF8A431FFF47F865FCABF81E" httpUri="https://zenodo.org/record/2612661/files/figure.png" pageId="3" pageNumber="426" startId="3.[151,264,1976,1999]" targetBox="[153,1435,807,1955]" targetPageId="3">
<paragraph id="8BDB36ADFF8A431FFF47F865FCABF81E" blockId="3.[151,1436,1976,2029]" pageId="3" pageNumber="426">
<emphasis id="B910EABFFF8A431FFF47F865FEAEF83E" bold="true" box="[151,310,1976,1999]" pageId="3" pageNumber="426">FIGURES 2–5</emphasis>
. 
<taxonomicName id="4C644D2EFF8A431FFE95F864FC86F83D" authority="Ziegler, Gisondi &amp; Cerretti" authorityName="Gisondi &amp; Lenzi &amp; Ziegler &amp; Giulio &amp; Cerretti" authorityYear="2019" box="[325,798,1976,1998]" class="Insecta" family="Tachinidae" genus="Stevenia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="426" phylum="Arthropoda" rank="species" species="gilasiani" status="sp. nov.">
<emphasis id="B910EABFFF8A431FFE95F864FE64F83D" box="[325,508,1976,1998]" italics="true" pageId="3" pageNumber="426">Stevenia gilasiani</emphasis>
Ziegler, Gisondi &amp; Cerretti
</taxonomicName>
<emphasis id="B910EABFFF8A431FFCF7F864FCE2F83D" bold="true" box="[807,890,1977,1998]" pageId="3" pageNumber="426">
<taxonomicNameLabel id="A22357C4FF8A431FFCF7F864FCE2F83D" box="[807,890,1977,1998]" pageId="3" pageNumber="426" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(holotype ♂). 
<emphasis id="B910EABFFF8A431FFBC8F865FBB2F83E" bold="true" box="[1048,1066,1976,1997]" pageId="3" pageNumber="426">2.</emphasis>
Habitus in lateral view. 
<emphasis id="B910EABFFF8A431FFAFEF865FAD9F83E" bold="true" box="[1326,1345,1976,1997]" pageId="3" pageNumber="426">3.</emphasis>
Head in lateral view. 
<emphasis id="B910EABFFF8A431FFEC9F80AFEB3F81F" bold="true" box="[281,299,2007,2028]" pageId="3" pageNumber="426">4.</emphasis>
Right wing in dorsal view. 
<emphasis id="B910EABFFF8A431FFD94F80AFDCFF81F" bold="true" box="[580,599,2007,2028]" pageId="3" pageNumber="426">5.</emphasis>
Head in frontal view.
</paragraph>
</caption>
<caption id="DF1B6625FF8D4318FF47FBBCFC93FB01" httpUri="https://zenodo.org/record/2612663/files/figure.png" pageId="4" pageNumber="427" startId="4.[151,264,1121,1144]" targetBox="[161,1413,209,1090]" targetPageId="4">
<paragraph id="8BDB36ADFF8D4318FF47FBBCFC93FB01" blockId="4.[151,1436,1121,1266]" pageId="4" pageNumber="427">
<emphasis id="B910EABFFF8D4318FF47FBBCFED3FB84" bold="true" box="[151,331,1121,1144]" pageId="4" pageNumber="427">FIGURES 6–11.</emphasis>
<taxonomicName id="4C644D2EFF8D4318FE86FBBFFD88FB8B" authority="Gisondi &amp; Lenzi &amp; Ziegler &amp; Giulio &amp; Cerretti, 2019" authorityName="Gisondi &amp; Lenzi &amp; Ziegler &amp; Giulio &amp; Cerretti" authorityYear="2019" box="[342,528,1122,1144]" class="Insecta" family="Tachinidae" genus="Stevenia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="427" phylum="Arthropoda" rank="species" species="gilasiani" status="sp. nov.">
<emphasis id="B910EABFFF8D4318FE86FBBFFD88FB8B" box="[342,528,1122,1144]" italics="true" pageId="4" pageNumber="427">Stevenia gilasiani</emphasis>
</taxonomicName>
<emphasis id="B910EABFFF8D4318FDCBFBBFFDE9FB84" bold="true" box="[539,625,1122,1143]" pageId="4" pageNumber="427">
<taxonomicNameLabel id="A22357C4FF8D4318FDCBFBBFFDE9FB84" box="[539,625,1122,1143]" pageId="4" pageNumber="427" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
. 
<emphasis id="B910EABFFF8D4318FD54FBBFFD28FB84" bold="true" box="[644,688,1122,1143]" pageId="4" pageNumber="427">6–8.</emphasis>
Holotype ♂. 
<emphasis id="B910EABFFF8D4318FC9BFBBFFCC5FB84" bold="true" box="[843,861,1122,1143]" pageId="4" pageNumber="427">6.</emphasis>
Detail of thorax, showing two postpronotal setae. 
<emphasis id="B910EABFFF8D4318FA59FBBFFA04FB84" bold="true" box="[1417,1436,1122,1143]" pageId="4" pageNumber="427">7.</emphasis>
Abdomen in dorsal view. 
<emphasis id="B910EABFFF8D4318FE4FFB5DFE2AFB66" bold="true" box="[415,434,1152,1173]" pageId="4" pageNumber="427">8.</emphasis>
Abdomen in lateral view, showing the sexual-patches on tergites 3 and 4. 
<emphasis id="B910EABFFF8D4318FB7EFB5DFB7DFB66" bold="true" box="[1198,1253,1152,1173]" pageId="4" pageNumber="427">9–11.</emphasis>
Details of cuticle and setae composing the sexual-patches, using scanning electron microscopy (SEM) (paratype, MZUR). 
<emphasis id="B910EABFFF8D4318FB1CFB42FB46FB47" bold="true" box="[1228,1246,1183,1204]" pageId="4" pageNumber="427">9.</emphasis>
Sexual-patches of tergite 3 in ventral view. 
<emphasis id="B910EABFFF8D4318FE49FB63FE21FB20" bold="true" box="[409,441,1214,1235]" pageId="4" pageNumber="427">10.</emphasis>
Seta and porous area. 
<emphasis id="B910EABFFF8D4318FD74FB63FD5AFB20" bold="true" box="[676,706,1214,1235]" pageId="4" pageNumber="427">11.</emphasis>
Internal surface of cuticle at of the base of a seta composing a sexualpatch, showing the sieve-like structure at level of porous area.
</paragraph>
</caption>
<paragraph id="8BDB36ADFF8D4318FF17FAC7FF66F968" blockId="4.[151,1437,1306,2015]" pageId="4" pageNumber="427">
Head (
<figureCitation id="135F2A28FF8D4318FEC4FAC7FE19FAC0" box="[276,385,1306,1331]" captionStart="FIGURES 2–5" captionStartId="3.[151,264,1976,1999]" captionTargetBox="[153,1435,807,1955]" captionTargetId="figure@3.[151,1435,807,1955]" captionTargetPageId="3" captionText="FIGURES 2–5. Stevenia gilasiani Ziegler, Gisondi &amp; Cerretti sp. nov. (holotype ♂). 2. Habitus in lateral view. 3. Head in lateral view. 4. Right wing in dorsal view. 5. Head in frontal view." httpUri="https://zenodo.org/record/2612661/files/figure.png" pageId="4" pageNumber="427">Figs 3, 5</figureCitation>
). Frons about 0.53–0.65 [0.55] of a compound eye in dorsal view. Outer vertical seta not differentiated. Ocellar setae short, hair-like and proclinate. Frons with 6–9 frontal setae reaching upper margin of pedicel. Fronto-orbital plate with scattered setulae lateral to row of frontal setae. One upper reclinate orbital seta and no proclinate orbital setae. Parafacial variable, approximately 0.8–1.4 times as wide as postpedicel at mid length, with 2–3 setae on lower half and with a few hairs over its whole length. Facial ridge concave with robust setae above vibrissa on lower 1/5 of its length. Vibrissa inserted at level of lower facial margin. Face and lower facial margin not visible in lateral view. Occiput with black setulae only. Gena approximately 0.3–0.4 of compound eye height. Genal dilation well developed. Antenna long, 5/4 of the height of gena. Postpedicel about 1.6–2.0 times as long as pedicel. Arista bare and thickened on proximal 1/5. First aristomere very short, not longer than wide. Second aristomere about as long as wide. Prementum about twice as long as width at mid length. Palpus apically enlarged.
</paragraph>
<paragraph id="8BDB36ADFF8D4319FF17F97BFAFDFED7" blockId="4.[151,1437,1306,2015]" lastBlockId="5.[151,1437,151,297]" lastPageId="5" lastPageNumber="428" pageId="4" pageNumber="427">
Thorax. Postpronotum with two setae (
<figureCitation id="135F2A28FF8D4318FD51F97BFD5FF94C" box="[641,711,1702,1727]" captionStart="FIGURES 6–11" captionStartId="4.[151,264,1121,1144]" captionTargetBox="[161,1413,209,1090]" captionTargetId="figure@4.[150,1436,193,1101]" captionTargetPageId="4" captionText="FIGURES 6–11. Stevenia gilasiani sp. nov.. 6–8. Holotype ♂. 6. Detail of thorax, showing two postpronotal setae. 7. Abdomen in dorsal view. 8. Abdomen in lateral view, showing the sexual-patches on tergites 3 and 4. 9–11. Details of cuticle and setae composing the sexual-patches, using scanning electron microscopy (SEM) (paratype, MZUR). 9. Sexual-patches of tergite 3 in ventral view. 10. Seta and porous area. 11. Internal surface of cuticle at of the base of a seta composing a sexualpatch, showing the sieve-like structure at level of porous area." httpUri="https://zenodo.org/record/2612663/files/figure.png" pageId="4" pageNumber="427">Fig. 6</figureCitation>
). Acrostichal setae absent. Two presutural and three postsutural dorsocentral setae. Two postsutural intra-alar setae separated by a distance greater than distance between first seta and suture, or only one postsutural intra-alar seta. First postsutural supra-alar seta not differentiated. Two katepisternal setae. Katepimeron bare. Anepimeral seta present but mostly weak. Two pairs of marginal scutellar setae: apical pair crossed. One pair of discal scutellar setae. Anatergite bare. Anterior and posterior fringes of posterior spiracle of approximately the same size. Postmetacoxal area membranous. 
<emphasis id="B910EABFFF8D4318FB8CF886FB09F881" box="[1116,1169,1883,1906]" italics="true" pageId="4" pageNumber="427">Legs</emphasis>
. Preapical anterodorsal seta of fore tibia about as long as preapical dorsal seta. Mid tibia with two anterodorsal setae. Hind tibia with three preapical dorsal setae. Preapical posteroventral seta of hind tibia about as long as preapical anteroventral one. Hind tibia with two well-developed anterodorsal setae. Posterodorsal margin of coxa bare. 
<emphasis id="B910EABFFF8D4318FB80F81AFB11F82D" box="[1104,1161,1991,2014]" italics="true" pageId="4" pageNumber="427">Wing</emphasis>
(
<figureCitation id="135F2A28FF8D4318FB49F81BFB46F82C" box="[1177,1246,1990,2015]" captionStart="FIGURES 2–5" captionStartId="3.[151,264,1976,1999]" captionTargetBox="[153,1435,807,1955]" captionTargetId="figure@3.[151,1435,807,1955]" captionTargetPageId="3" captionText="FIGURES 2–5. Stevenia gilasiani Ziegler, Gisondi &amp; Cerretti sp. nov. (holotype ♂). 2. Habitus in lateral view. 3. Head in lateral view. 4. Right wing in dorsal view. 5. Head in frontal view." httpUri="https://zenodo.org/record/2612661/files/figure.png" pageId="4" pageNumber="427">Fig. 4</figureCitation>
). Lower calypter tongue-shaped. Second costal section (CS 
<subScript id="17E034E8FF8C4319FDBBFF7BFDEBFF47" attach="left" box="[619,627,166,180]" fontSize="6" pageId="5" pageNumber="428">2</subScript>
) bare ventrally. Costal spine not differentiated from other costal setulae. R 
<subScript id="17E034E8FF8C4319FF79FF10FF29FF28" attach="left" box="[169,177,205,219]" fontSize="6" pageId="5" pageNumber="428">1</subScript>
bare. Base of vein R 
<subScript id="17E034E8FF8C4319FE7AFF10FE5AFF28" attach="left" box="[426,450,205,219]" fontSize="6" pageId="5" pageNumber="428">4+5</subScript>
with 2–5 setulae dorsally and ventrally. Bend of M 
<subScript id="17E034E8FF8C4319FBC7FF10FB87FF28" attach="left" box="[1047,1055,205,219]" fontSize="6" pageId="5" pageNumber="428">1</subScript>
forming an obtuse angle. Fourth costal section (CS 
<subScript id="17E034E8FF8C4319FEB2FF29FEF2FEF1" attach="left" box="[354,362,244,258]" fontSize="6" pageId="5" pageNumber="428">4</subScript>
) longer than sixth (CS 
<subScript id="17E034E8FF8C4319FDBBFF29FDEBFEF1" attach="right" box="[619,627,244,258]" fontSize="6" pageId="5" pageNumber="428">6</subScript>
). Section of M between crossveins r-m and dm-cu 1.7–2.3 times longer than section between dm-cu and bend of M. Cell r 
<subScript id="17E034E8FF8C4319FD18FEC6FD78FEDA" attach="left" box="[712,736,283,297]" fontSize="6" pageId="5" pageNumber="428">4+5</subScript>
with a petiole 3/4 the length of postangular section of M.
</paragraph>
<caption id="DF1B6625FF8C4319FF47F92CFAC5F8B6" httpUri="https://zenodo.org/record/2612665/files/figure.png" pageId="5" pageNumber="428" startId="5.[151,264,1777,1800]" targetBox="[245,1340,379,1725]" targetPageId="5">
<paragraph id="8BDB36ADFF8C4319FF47F92CFAC5F8B6" blockId="5.[151,1436,1777,1861]" pageId="5" pageNumber="428">
<emphasis id="B910EABFFF8C4319FF47F92CFECCF8F4" bold="true" box="[151,340,1777,1800]" pageId="5" pageNumber="428">FIGURES 12–14.</emphasis>
<taxonomicName id="4C644D2EFF8C4319FE8AF92FFCB3F8FB" authority="Ziegler, Gisondi &amp; Cerretti" authorityName="Gisondi &amp; Lenzi &amp; Ziegler &amp; Giulio &amp; Cerretti" authorityYear="2019" box="[346,811,1778,1800]" class="Insecta" family="Tachinidae" genus="Stevenia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="428" phylum="Arthropoda" rank="species" species="gilasiani" status="sp. nov.">
<emphasis id="B910EABFFF8C4319FE8AF92FFE37F8F4" box="[346,431,1778,1799]" italics="true" pageId="5" pageNumber="428">Stevenia</emphasis>
<emphasis id="B910EABFFF8C4319FE65F92FFD96F8FB" box="[437,526,1778,1800]" italics="true" pageId="5" pageNumber="428">gilasiani</emphasis>
Ziegler, Gisondi &amp; Cerretti
</taxonomicName>
, 
<emphasis id="B910EABFFF8C4319FCE5F92FFC1EF8F4" bold="true" box="[821,902,1778,1799]" pageId="5" pageNumber="428">
<taxonomicNameLabel id="A22357C4FF8C4319FCE5F92FFC1EF8F4" box="[821,902,1778,1799]" pageId="5" pageNumber="428" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, terminalia (paratype, MZUR). 
<emphasis id="B910EABFFF8C4319FB1BF92FFB73F8F4" bold="true" box="[1227,1259,1778,1799]" pageId="5" pageNumber="428">12.</emphasis>
Lateral view. 
<emphasis id="B910EABFFF8C4319FAACF92FFA04F8F4" bold="true" box="[1404,1436,1778,1799]" pageId="5" pageNumber="428">13.</emphasis>
Dorsolateral view. 
<emphasis id="B910EABFFF8C4319FEB1F8CDFE18F8D6" bold="true" box="[353,384,1808,1829]" pageId="5" pageNumber="428">14.</emphasis>
Detail of the terminalia in ventral view. Abbreviations: aj = ejaculatory apodeme; app = phallus apodeme; c = cerci; ep = epandrium; hyp = hypandrium; pr = pregonite; ps = postgonite; s = surstylus; sd = sperm ducts.
</paragraph>
</caption>
<paragraph id="8BDB36ADFF8C431AFF17F8ACFEAFFE23" blockId="5.[151,1436,1905,2002]" lastBlockId="6.[151,1436,151,573]" lastPageId="6" lastPageNumber="429" pageId="5" pageNumber="428">
Abdomen (
<figureCitation id="135F2A28FF8C4319FE94F8ACFE32F879" box="[324,426,1905,1930]" captionStart="FIGURES 6–11" captionStartId="4.[151,264,1121,1144]" captionTargetBox="[161,1413,209,1090]" captionTargetId="figure@4.[150,1436,193,1101]" captionTargetPageId="4" captionText="FIGURES 6–11. Stevenia gilasiani sp. nov.. 6–8. Holotype ♂. 6. Detail of thorax, showing two postpronotal setae. 7. Abdomen in dorsal view. 8. Abdomen in lateral view, showing the sexual-patches on tergites 3 and 4. 9–11. Details of cuticle and setae composing the sexual-patches, using scanning electron microscopy (SEM) (paratype, MZUR). 9. Sexual-patches of tergite 3 in ventral view. 10. Seta and porous area. 11. Internal surface of cuticle at of the base of a seta composing a sexualpatch, showing the sieve-like structure at level of porous area." httpUri="https://zenodo.org/record/2612663/files/figure.png" pageId="5" pageNumber="428">Figs 7–8</figureCitation>
). Mid-dorsal depression of syntergite 1+2 confined to anterior 1/4 of syntergite. Syntergite 1+2 without median marginal setae, with several lateral discal and lateral marginal setae. Tergite 3 with one pair of median marginal setae, one pair of lateral discal setae and two pairs of lateral marginal setae. Tergites 4 and 5 each with a row of marginal setae and without discal setae. Tergite 5 very short, approximately 1/3 length of tergite 4. Sternite 4 exposed. Ventral sides of tergites 3, 4 and 5 with paired patches of appressed setae and cuticular microsculpture (i.e., sexual-patches) (
<figureCitation id="135F2A28FF8F431AFDEBFF02FD35FF0B" box="[571,685,223,248]" captionStart="FIGURES 6–11" captionStartId="4.[151,264,1121,1144]" captionTargetBox="[161,1413,209,1090]" captionTargetId="figure@4.[150,1436,193,1101]" captionTargetPageId="4" captionText="FIGURES 6–11. Stevenia gilasiani sp. nov.. 6–8. Holotype ♂. 6. Detail of thorax, showing two postpronotal setae. 7. Abdomen in dorsal view. 8. Abdomen in lateral view, showing the sexual-patches on tergites 3 and 4. 9–11. Details of cuticle and setae composing the sexual-patches, using scanning electron microscopy (SEM) (paratype, MZUR). 9. Sexual-patches of tergite 3 in ventral view. 10. Seta and porous area. 11. Internal surface of cuticle at of the base of a seta composing a sexualpatch, showing the sieve-like structure at level of porous area." httpUri="https://zenodo.org/record/2612663/files/figure.png" pageId="6" pageNumber="429">Figs 8–11</figureCitation>
). Sexual-patches also present on sternites 3 and 4. 
<emphasis id="B910EABFFF8F431AFB32FF02FAC4FF0B" box="[1250,1372,223,248]" italics="true" pageId="6" pageNumber="429">Terminalia</emphasis>
(
<figureCitation id="135F2A28FF8F431AFABAFF02FF79FEEF" captionStart="FIGURES 12–14" captionStartId="5.[151,264,1777,1800]" captionTargetBox="[245,1340,379,1725]" captionTargetId="figure@5.[209,1378,348,1756]" captionTargetPageId="5" captionText="FIGURES 12–14. Stevenia gilasiani Ziegler, Gisondi &amp; Cerretti, sp. nov., terminalia (paratype, MZUR). 12. Lateral view. 13. Dorsolateral view. 14. Detail of the terminalia in ventral view. Abbreviations: aj = ejaculatory apodeme; app = phallus apodeme; c = cerci; ep = epandrium; hyp = hypandrium; pr = pregonite; ps = postgonite; s = surstylus; sd = sperm ducts." httpUri="https://zenodo.org/record/2612665/files/figure.png" pageId="6" pageNumber="429">Figs 12–14</figureCitation>
). Epandrium short and convex. Cercus with curled cuticular microsculpture, long setae on lateral surface and one pair of long setae on apical dorsal surface; cerci approximately 2/3 of length of surstylus, with a dorsomedial suture separating them. Surstylus dagger-like in lateral view and rounded at apex in posterior view, twisted inward (
<figureCitation id="135F2A28FF8F431AFF4FFEB2FEBBFE7B" box="[159,291,367,392]" captionStart="FIGURES 12–14" captionStartId="5.[151,264,1777,1800]" captionTargetBox="[245,1340,379,1725]" captionTargetId="figure@5.[209,1378,348,1756]" captionTargetPageId="5" captionText="FIGURES 12–14. Stevenia gilasiani Ziegler, Gisondi &amp; Cerretti, sp. nov., terminalia (paratype, MZUR). 12. Lateral view. 13. Dorsolateral view. 14. Detail of the terminalia in ventral view. Abbreviations: aj = ejaculatory apodeme; app = phallus apodeme; c = cerci; ep = epandrium; hyp = hypandrium; pr = pregonite; ps = postgonite; s = surstylus; sd = sperm ducts." httpUri="https://zenodo.org/record/2612665/files/figure.png" pageId="6" pageNumber="429">Figs 12–13</figureCitation>
). Processus longus stick-like and firmly fused to base of surstylus. Distiphallus as long as surstylus; lateroventral face of distiphallus sclerotized and covered with scale-like spinules. Acrophallus with tripartite sperm duct (
<figureCitation id="135F2A28FF8F431AFF06FE6AFEB2FE23" box="[214,298,439,464]" captionStart="FIGURES 12–14" captionStartId="5.[151,264,1777,1800]" captionTargetBox="[245,1340,379,1725]" captionTargetId="figure@5.[209,1378,348,1756]" captionTargetPageId="5" captionText="FIGURES 12–14. Stevenia gilasiani Ziegler, Gisondi &amp; Cerretti, sp. nov., terminalia (paratype, MZUR). 12. Lateral view. 13. Dorsolateral view. 14. Detail of the terminalia in ventral view. Abbreviations: aj = ejaculatory apodeme; app = phallus apodeme; c = cerci; ep = epandrium; hyp = hypandrium; pr = pregonite; ps = postgonite; s = surstylus; sd = sperm ducts." httpUri="https://zenodo.org/record/2612665/files/figure.png" pageId="6" pageNumber="429">Fig. 14</figureCitation>
).
</paragraph>
<paragraph id="8BDB36ADFF8F431AFF17FE01FE06FE06" blockId="6.[151,1436,151,573]" box="[199,414,476,501]" pageId="6" pageNumber="429">
<emphasis id="B910EABFFF8F431AFF17FE01FE83FE06" box="[199,283,476,501]" italics="true" pageId="6" pageNumber="429">Female</emphasis>
. Unknown.
</paragraph>
</subSubSection>
<subSubSection id="C37E6526FF8F431AFF17FE22FD60FDEB" box="[199,760,511,536]" pageId="6" pageNumber="429" type="distribution">
<paragraph id="8BDB36ADFF8F431AFF17FE22FD60FDEB" blockId="6.[151,1436,151,573]" box="[199,760,511,536]" pageId="6" pageNumber="429">
<emphasis id="B910EABFFF8F431AFF17FE22FEF8FDEB" bold="true" box="[199,352,511,536]" pageId="6" pageNumber="429">Distribution.</emphasis>
Palaearctic: 
<collectingCountry id="F373763DFF8F431AFE21FDDDFD86FDE4" box="[497,542,512,535]" name="Iran" pageId="6" pageNumber="429">Iran</collectingCountry>
(
<collectingRegion id="49A0F84FFF8F431AFDFCFDDDFD74FDEB" box="[556,748,511,536]" country="Iran" name="Azarbayjan-e Gharbi" pageId="6" pageNumber="429">West Azerbaijan</collectingRegion>
).
</paragraph>
</subSubSection>
<subSubSection id="C37E6526FF8F431AFF17FDF9FE3CFDCE" box="[199,420,548,573]" pageId="6" pageNumber="429" type="biology_ecology">
<paragraph id="8BDB36ADFF8F431AFF17FDF9FE3CFDCE" blockId="6.[151,1436,151,573]" box="[199,420,548,573]" pageId="6" pageNumber="429">
<emphasis id="B910EABFFF8F431AFF17FDF9FEBFFDCE" bold="true" box="[199,295,548,573]" pageId="6" pageNumber="429">Biology.</emphasis>
Unknown.
</paragraph>
</subSubSection>
</treatment>
</document>