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<document id="FC2C79C78759FE975F4D845CFED45B17" ID-CLB-Dataset="31085" ID-DOI="10.11646/zootaxa.4369.1.6" ID-GBIF-Dataset="9032e1fb-864e-48fc-a4b9-de670d73543a" ID-ISSN="1175-5326" ID-Zenodo-Dep="1135606" ID-ZooBank="554E4431-B96C-467E-953F-A3244239422E" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1515147733856" checkinUser="plazi" docAuthor="Naser, Murtada D., Davie, Peter J. F. &amp; Waltham, Nathan J." docDate="2018" docId="03CD87A2FFD3172AFF0CD2C7FACEFC16" docLanguage="en" docName="zootaxa.4369.1.6.pdf" docOrigin="Zootaxa 4369 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Austrothelphusa wasselli" docType="treatment" docVersion="8" lastPageNumber="119" masterDocId="FFF4FFDAFFD61720FF9BD767FFFFFE01" masterDocTitle="Redescription of Austrothelphusa wasselli (Bishop, 1963) (Crustacea: Brachyura: Gecarcinucidae), and designation of a new species from the Gilbert River, north Queensland, Australia" masterLastPageNumber="127" masterPageNumber="109" pageNumber="114" updateTime="1698512145660" updateUser="ExternalLinkService">
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<mods:title id="839E7B2D5B368FC85415A973A5655F1E">Redescription of Austrothelphusa wasselli (Bishop, 1963) (Crustacea: Brachyura: Gecarcinucidae), and designation of a new species from the Gilbert River, north Queensland, Australia</mods:title>
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<mods:namePart id="42455C32514AC3EC2D26A4F4294CB257">Naser, Murtada D.</mods:namePart>
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<mods:namePart id="228980878228D2654854773CE8EDE188">Davie, Peter J. F.</mods:namePart>
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<mods:namePart id="D5FE69D809B9DDE7CE44D66F682573A5">Waltham, Nathan J.</mods:namePart>
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<mods:date id="7305DA4C46FF02AABB61379C02444109">2018</mods:date>
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<treatment id="03CD87A2FFD3172AFF0CD2C7FACEFC16" ID-DOI="http://doi.org/10.5281/zenodo.5964610" ID-GBIF-Taxon="139582135" ID-Zenodo-Dep="5964610" LSID="urn:lsid:plazi:treatment:03CD87A2FFD3172AFF0CD2C7FACEFC16" httpUri="http://treatment.plazi.org/id/03CD87A2FFD3172AFF0CD2C7FACEFC16" lastPageId="10" lastPageNumber="119" pageId="5" pageNumber="114">
<subSubSection id="C37E653FFFD31725FF0CD2C7FDFAFBDD" pageId="5" pageNumber="114" type="nomenclature">
<paragraph id="8BDB36B4FFD31725FF0CD2C7FD80FBBA" blockId="5.[151,639,1440,1501]" box="[151,639,1440,1467]" pageId="5" pageNumber="114">
<heading id="D09381D8FFD31725FF0CD2C7FD80FBBA" bold="true" box="[151,639,1440,1467]" fontSize="11" level="1" pageId="5" pageNumber="114" reason="1">
<taxonomicName id="4C644D37FFD31725FF0CD2C7FD80FBBA" ID-CoL="JYF6" authority="(Bishop, 1963)" baseAuthorityName="Bishop" baseAuthorityYear="1963" box="[151,639,1440,1467]" class="Malacostraca" family="Gecarcinucidae" genus="Austrothelphusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="114" phylum="Arthropoda" rank="species" species="wasselli">
<emphasis id="B910EAA6FFD31725FF0CD2C7FD80FBBA" bold="true" box="[151,639,1440,1467]" pageId="5" pageNumber="114">
<emphasis id="B910EAA6FFD31725FF0CD2C7FE3BFBBB" bold="true" box="[151,452,1440,1466]" italics="true" pageId="5" pageNumber="114">Austrothelphusa wasselli</emphasis>
(
<bibRefCitation id="EFF54B45FFD31725FE4FD2C6FD88FBBA" author="Bishop" box="[468,631,1441,1467]" pageId="5" pageNumber="114" refString="Bishop, J. A. (1963) The Australian freshwater crabs of the family Potamonidae (Crustacea: Decapoda). Australian Journal of Marine and Freshwater Research, 14, 218 - 238. https: // doi. org / 10.1071 / MF 9630218" type="journal article" year="1963">Bishop, 1963</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BDB36B4FFD31725FF0CD2A3FDFAFBDD" blockId="5.[151,639,1440,1501]" box="[151,517,1476,1501]" pageId="5" pageNumber="114">
(
<figureCitation id="135F2A31FFD31725FF04D2A3FF18FBDD" box="[159,231,1476,1501]" captionStart="FIGURE 3" captionStartId="6.[151,250,1962,1984]" captionTargetBox="[167,1420,304,1924]" captionTargetId="figure@6.[167,1420,304,1924]" captionTargetPageId="6" captionText="FIGURE 3. Austrothelphusa wasselli, holotype male (11.2 × 9.2 mm), Port Stewart District, Cape York (AM-P.14524). A, dorsal view; B, frontal view; C, ventral view." httpUri="https://zenodo.org/record/1135612/files/figure.png" pageId="5" pageNumber="114">Figs 3</figureCitation>
,
<figureCitation id="135F2A31FFD31725FF68D2A3FEFCFBDD" box="[243,259,1476,1500]" captionStart="FIGURE 4" captionStartId="7.[151,250,1863,1885]" captionTargetBox="[154,1439,196,1833]" captionTargetId="figure@7.[154,1439,196,1833]" captionTargetPageId="7" captionText="FIGURE 4. Austrothelphusa wasselli, male (25.8 × 20.5 mm) Stewart River, Queensland QM-W20049. A, dorsal view; B, ventral view of sternum and pleon." httpUri="https://zenodo.org/record/1135614/files/figure.png" pageId="5" pageNumber="114">4</figureCitation>
,
<figureCitation id="135F2A31FFD31725FE94D2A3FEE0FBDC" box="[271,287,1476,1501]" captionStart="FIGURE5" captionStartId="8.[151,250,1924,1946]" captionTargetBox="[154,1439,875,1900]" captionTargetId="figure@8.[154,1439,875,1900]" captionTargetPageId="8" captionText="FIGURE5. Austrothelphusa wasselli, male (25.8 × 20.5 mm) Stewart River, Queensland QM-W20049.A, ventrofrontal view; B,outer view of major chela; C, ventrofrontal view showing chelae and third maxillipeds." httpUri="https://zenodo.org/record/1135616/files/figure.png" pageId="5" pageNumber="114">5</figureCitation>
,
<figureCitation id="135F2A31FFD31725FEB0D2A3FE8EFBDC" box="[299,369,1476,1501]" captionStart="FIGURE 6" captionStartId="9.[151,250,1705,1727]" captionTargetBox="[153,1434,890,1668]" captionTargetId="figure@9.[153,1434,877,1684]" captionTargetPageId="9" captionText="FIGURE 6. AC, Austrothelphusa wasselli, male (18.8 × 14.9 mm), Oscar Creek, 2.1 km SSE of Coen, Archer Catchment, Queensland (QM-W29187). DF, A. gilbertensis sp.nov., holotype male (11.2 × 9.2 mm), Port Stewart District, Cape York (AM-P.14524).A, D, ventrofrontal view of cephalothorax (orbit, frontal triangle, epistome and pterygostome); B, E, left major chela; C, F, pleon. Scale line 5 mm." httpUri="https://zenodo.org/record/1135618/files/figure.png" pageId="5" pageNumber="114">6AC</figureCitation>
,
<figureCitation id="135F2A31FFD31725FEE7D2A3FE02FBDD" box="[380,509,1476,1500]" captionStart="FIGURE 7" captionStartId="10.[151,250,1893,1915]" captionTargetBox="[176,1409,994,1864]" captionTargetId="figure@10.[156,1440,986,1865]" captionTargetPageId="10" captionText="FIGURE 7. A, C, E, F, Austrothelphusa wasselli, male (25.8 × 20.5 mm), from Stewart River, Queensland (QM-W20049). B, D, G, H, A. gilbertensis sp. nov., holotype male (22.3 × 17.6 mm), Venture Creek, E of Croydon, Gilbert Catchment, Queensland (QM-W28040). A, B, outline of frontal margin and orbits; C, D, fifth pereiopod; EH, left first gonopod (sternal view) with magnification of apical part." httpUri="https://zenodo.org/record/1135620/files/figure.png" pageId="5" pageNumber="114">7A, C, E, F</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C37E653FFFD31725FF0CD16BFC69F823" box="[151,918,1548,1570]" pageId="5" pageNumber="114" type="reference_group">
<paragraph id="8BDB36B4FFD31725FF0CD16BFC69F823" blockId="5.[151,918,1548,1570]" box="[151,918,1548,1570]" pageId="5" pageNumber="114">
<treatmentCitationGroup id="AB74119AFFD31725FF0CD16BFC69F823" box="[151,918,1548,1570]" pageId="5" pageNumber="114">
<taxonomicName id="4C644D37FFD31725FF0CD16BFDBAF823" ID-CoL="4DFC6" authority="Bishop, 1963: 229" authorityName="Bishop" authorityPageNumber="229" authorityYear="1963" box="[151,581,1548,1570]" class="Malacostraca" family="Gecarcinucidae" genus="Parathelphusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="114" phylum="Arthropoda" rank="species" species="wasselli">
<emphasis id="B910EAA6FFD31725FF0CD16BFE7BF823" box="[151,388,1548,1570]" italics="true" pageId="5" pageNumber="114">Parathelphusa wasselli</emphasis>
<treatmentCitation id="0AC510A5FFD31725FE11D16BFDBAF823" author="Bishop" box="[394,581,1548,1570]" page="229" pageId="5" pageNumber="114" year="1963">
<bibRefCitation id="EFF54B45FFD31725FE11D16BFDEBF823" author="Bishop" box="[394,532,1548,1570]" pageId="5" pageNumber="114" refString="Bishop, J. A. (1963) The Australian freshwater crabs of the family Potamonidae (Crustacea: Decapoda). Australian Journal of Marine and Freshwater Research, 14, 218 - 238. https: // doi. org / 10.1071 / MF 9630218" type="journal article" year="1963">Bishop, 1963</bibRefCitation>
: 229
</treatmentCitation>
</taxonomicName>
230, figs 1A, 2D, 4, pl. 1, fig. 1.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C37E653FFFD31725FF0CD129FC3BF8D2" pageId="5" pageNumber="114" type="materials_examined">
<paragraph id="8BDB36B4FFD31725FF0CD129FC3BF8D2" blockId="5.[151,1437,1614,2035]" pageId="5" pageNumber="114">
<materialsCitation id="3B0C3CE9FFD31725FF0CD129FD3AF8AE" ID-GBIF-Occurrence="1802482357" collectingDate="1993-04-21" collectionCode="QDPI, CYPLUS" collectorName="Material &amp; Cape York &amp; May-June &amp; J. L. Wassell. &amp; B. Herbert &amp; Peeters" location="Port Stewart District" municipality="Stewart River" pageId="5" pageNumber="114" specimenCount="2" specimenCount-male="2" stateProvince="Queensland" typeStatus="holotype">
<emphasis id="B910EAA6FFD31725FF0CD129FE85F866" bold="true" box="[151,378,1614,1639]" pageId="5" pageNumber="114">
<collectorName id="26915362FFD31725FF0CD129FF00F866" box="[151,255,1614,1639]" pageId="5" pageNumber="114">Material</collectorName>
examined
</emphasis>
.
<typeStatus id="54DF8816FFD31725FE12D129FE0EF866" box="[393,497,1614,1639]" pageId="5" pageNumber="114" type="holotype">Holotype</typeStatus>
(AM-P.14524), male (11.2 ×
<quantity id="4C9C9B51FFD31725FCDAD128FC63F866" box="[833,924,1615,1639]" metricMagnitude="-3" metricUnit="m" metricValue="9.2" pageId="5" pageNumber="114" unit="mm" value="9.2">9.2 mm</quantity>
),
<location id="8EBB606FFFD31725FC34D128FB66F866" LSID="urn:lsid:plazi:treatment:03CD87A2FFD3172AFF0CD2C7FACEFC16:8EBB606FFFD31725FC34D128FB66F866" box="[943,1177,1614,1639]" municipality="Stewart River" name="Port Stewart District" pageId="5" pageNumber="114" stateProvince="Queensland">Port Stewart District</location>
,
<collectorName id="26915362FFD31725FB3ED128FADEF866" box="[1189,1313,1614,1639]" pageId="5" pageNumber="114">Cape York</collectorName>
,
<collectorName id="26915362FFD31725FAB7D128FA63F866" box="[1324,1436,1615,1639]" pageId="5" pageNumber="114">May-June</collectorName>
1954,
<collectorName id="26915362FFD31725FF46D114FE8AF88A" box="[221,373,1650,1675]" pageId="5" pageNumber="114">J. L. Wassell.</collectorName>
QM-W20049,
<specimenCount id="9D62FD3DFFD31725FDBAD114FD87F88A" box="[545,632,1650,1675]" pageId="5" pageNumber="114" type="male">2 males</specimenCount>
(25.8 ×
<quantity id="4C9C9B51FFD31725FD42D115FCBFF88B" box="[729,832,1650,1675]" metricMagnitude="-2" metricUnit="m" metricValue="2.05" pageId="5" pageNumber="114" unit="mm" value="20.5">20.5 mm</quantity>
, 16.5 ×
<quantity id="4C9C9B51FFD31725FC07D114FBFDF88B" box="[924,1026,1651,1675]" metricMagnitude="-2" metricUnit="m" metricValue="1.31" pageId="5" pageNumber="114" unit="mm" value="13.1">13.1 mm</quantity>
,
<collectingMunicipality id="6BBFACCEFFD31725FB94D114FB54F88A" box="[1039,1195,1650,1675]" pageId="5" pageNumber="114">Stewart River</collectingMunicipality>
,
<collectingRegion id="49A0F856FFD31725FB2CD115FAC1F88A" box="[1207,1342,1650,1675]" country="Australia" name="Queensland" pageId="5" pageNumber="114">Queensland</collectingRegion>
, coll.
<collectorName id="26915362FFD31725FA18D114FF10F8AE" pageId="5" pageNumber="114">B. Herbert</collectorName>
, J.
<collectorName id="26915362FFD31725FE89D1F0FE9CF8AF" box="[274,355,1687,1710]" pageId="5" pageNumber="114">Peeters</collectorName>
(
<collectionCode id="ED75AE71FFD31725FEEAD1F0FE48F8AE" box="[369,439,1687,1711]" country="Australia" httpUri="http://grbio.org/cool/221v-j740" name="Queensland Department of Primary Industries" pageId="5" pageNumber="114">QDPI</collectionCode>
,
<collectionCode id="ED75AE71FFD31725FE5AD1F0FDCEF8AE" box="[449,561,1687,1711]" pageId="5" pageNumber="114">CYPLUS</collectionCode>
),
<collectingDate id="EF9EE99CFFD31725FDD8D1F0FD3AF8AE" box="[579,709,1687,1711]" pageId="5" pageNumber="114" value="1993-04-21">
<date id="FFDA1074FFD31725FDD8D1F0FD3AF8AE" box="[579,709,1687,1711]" pageId="5" pageNumber="114" value="1993-04-21">21.04.1993</date>
</collectingDate>
</materialsCitation>
.
<materialsCitation id="3B0C3CE9FFD31725FD54D1F0FC3BF8D2" ID-GBIF-Occurrence="1802482355" country="Heard Island and McDonald Islands" location="Oscar Creek" municipality="Archer Catchment" pageId="5" pageNumber="114" specimenCount="1" stateProvince="Queensland" typeStatus="holotype">
QM-W29187, male (18.8 ×
<quantity id="4C9C9B51FFD31725FB9CD1F0FB91F8AE" box="[1031,1134,1687,1711]" metricMagnitude="-2" metricUnit="m" metricValue="1.49" pageId="5" pageNumber="114" unit="mm" value="14.9">14.9 mm</quantity>
),
<location id="8EBB606FFFD31725FB1AD1F0FAF1F8AE" LSID="urn:lsid:plazi:treatment:03CD87A2FFD3172AFF0CD2C7FACEFC16:8EBB606FFFD31725FB1AD1F0FAF1F8AE" box="[1153,1294,1686,1711]" country="Heard Island and McDonald Islands" municipality="Archer Catchment" name="Oscar Creek" pageId="5" pageNumber="114" stateProvince="Queensland">Oscar Creek</location>
,
<locationDeviation id="9A692F96FFD31725FA83D1F0FF0CF8D2" pageId="5" pageNumber="114">
<quantity id="4C9C9B51FFD31725FA83D1F0FA99F8AE" box="[1304,1382,1686,1711]" metricMagnitude="3" metricUnit="m" metricValue="2.1" pageId="5" pageNumber="114" unit="km" value="2.1">2.1 km</quantity>
SSE of Coen
</locationDeviation>
,
<collectingMunicipality id="6BBFACCEFFD31725FF65D1DDFE30F8D2" box="[254,463,1722,1747]" pageId="5" pageNumber="114">Archer Catchment</collectingMunicipality>
,
<collectingRegion id="49A0F856FFD31725FE41D1DDFD9EF8D2" box="[474,609,1722,1747]" country="Australia" name="Queensland" pageId="5" pageNumber="114">Queensland</collectingRegion>
, coll. K.
<collectingCountry id="F3737624FFD31725FD5CD1DDFCBDF8D2" box="[711,834,1722,1747]" name="Heard Island and McDonald Islands" pageId="5" pageNumber="114">McDonald</collectingCountry>
, 8.05.2013.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C37E653FFFD31728FF5CD1B9FE40FCCC" lastPageId="8" lastPageNumber="117" pageId="5" pageNumber="114" type="description">
<paragraph id="8BDB36B4FFD31726FF5CD1B9FBB9FEF9" blockId="5.[151,1437,1614,2035]" lastBlockId="6.[151,1437,151,248]" lastPageId="6" lastPageNumber="115" pageId="5" pageNumber="114">
<emphasis id="B910EAA6FFD31725FF5CD1B9FEAEF8F6" bold="true" box="[199,337,1758,1783]" pageId="5" pageNumber="114">Description</emphasis>
. Small species (maximum CW of present material
<quantity id="4C9C9B51FFD31725FC3BD1B9FBF5F8F6" box="[928,1034,1758,1783]" metricMagnitude="-2" metricUnit="m" metricValue="2.58" pageId="5" pageNumber="114" unit="mm" value="25.8">25.8 mm</quantity>
). Carapace broadly oval (
<figureCitation id="135F2A31FFD31725FAA9D1B9FA68F8F6" box="[1330,1431,1758,1783]" captionStart="FIGURE 3" captionStartId="6.[151,250,1962,1984]" captionTargetBox="[167,1420,304,1924]" captionTargetId="figure@6.[167,1420,304,1924]" captionTargetPageId="6" captionText="FIGURE 3. Austrothelphusa wasselli, holotype male (11.2 × 9.2 mm), Port Stewart District, Cape York (AM-P.14524). A, dorsal view; B, frontal view; C, ventral view." httpUri="https://zenodo.org/record/1135612/files/figure.png" pageId="5" pageNumber="114">Figs. 3A</figureCitation>
,
<figureCitation id="135F2A31FFD31725FF0CD064FF42F91A" box="[151,189,1795,1819]" captionStart="FIGURE 4" captionStartId="7.[151,250,1863,1885]" captionTargetBox="[154,1439,196,1833]" captionTargetId="figure@7.[154,1439,196,1833]" captionTargetPageId="7" captionText="FIGURE 4. Austrothelphusa wasselli, male (25.8 × 20.5 mm) Stewart River, Queensland QM-W20049. A, dorsal view; B, ventral view of sternum and pleon." httpUri="https://zenodo.org/record/1135614/files/figure.png" pageId="5" pageNumber="114">4A</figureCitation>
); 1.26 times broader than long (n=3; CW&gt;
<quantity id="4C9C9B51FFD31725FD30D065FCEAF91A" box="[683,789,1794,1819]" metricMagnitude="-2" metricUnit="m" metricValue="1.66" pageId="5" pageNumber="114" unit="mm" value="16.6">16.6 mm</quantity>
); small specimens with less swollen branchial regions so typically narrower (
<typeStatus id="54DF8816FFD31725FEE4D041FE1CF93E" box="[383,483,1830,1855]" pageId="5" pageNumber="114" type="holotype">holotype</typeStatus>
1.21 times broader than long; CW
<quantity id="4C9C9B51FFD31725FC0DD040FBFDF93E" box="[918,1026,1831,1855]" metricMagnitude="-2" metricUnit="m" metricValue="1.1199999999999999" pageId="5" pageNumber="114" unit="mm" value="11.2">11.2 mm</quantity>
). Front projecting beyond level of exorbital angles, broadly bilobed, medial concavity shallow; inner part of supraorbital margin moderately welldefined, merging quite steeply with lateral slope of frontal lobe (
<figureCitation id="135F2A31FFD31725FCE8D009FC31F986" box="[883,974,1902,1927]" captionStart="FIGURE 7" captionStartId="10.[151,250,1893,1915]" captionTargetBox="[176,1409,994,1864]" captionTargetId="figure@10.[156,1440,986,1865]" captionTargetPageId="10" captionText="FIGURE 7. A, C, E, F, Austrothelphusa wasselli, male (25.8 × 20.5 mm), from Stewart River, Queensland (QM-W20049). B, D, G, H, A. gilbertensis sp. nov., holotype male (22.3 × 17.6 mm), Venture Creek, E of Croydon, Gilbert Catchment, Queensland (QM-W28040). A, B, outline of frontal margin and orbits; C, D, fifth pereiopod; EH, left first gonopod (sternal view) with magnification of apical part." httpUri="https://zenodo.org/record/1135620/files/figure.png" pageId="5" pageNumber="114">Fig. 7A</figureCitation>
). Frontal and orbital margins with raised rim. Postfrontal (epigastric) lobes rounded, not well developed, lacking striated crests; separated by deep narrow groove. Postorbital region moderately depressed, short slightly convex crest laterally adjacent to, but not quite reaching, epibranchial tooth. Anterolateral margins smoothly cristate, evenly convex, with one distinct but small epibranchial tooth. Branchial regions not swollen, bearing punctations; anteriorly lacking striations but with striated ridges posterolaterally. Cervical groove shallow, relatively poorly defined; gastro-cardiac (H-shaped) grooves not strongly marked. Posterolateral borders straight, convergent posteriorly.
</paragraph>
<caption id="DF1B663CFFD01726FF0CD0CDFD9EF9DE" httpUri="https://zenodo.org/record/1135612/files/figure.png" pageId="6" pageNumber="115" targetBox="[167,1420,304,1924]" targetPageId="6">
<paragraph id="8BDB36B4FFD01726FF0CD0CDFD9EF9DE" blockId="6.[151,1435,1962,2015]" pageId="6" pageNumber="115">
<emphasis id="B910EAA6FFD01726FF0CD0CDFEEAF9BE" bold="true" box="[151,277,1962,1984]" pageId="6" pageNumber="115">FIGURE 3.</emphasis>
<taxonomicName id="4C644D37FFD01726FE85D0CDFDE1F9C1" baseAuthorityName="Bishop" baseAuthorityYear="1963" box="[286,542,1962,1984]" class="Malacostraca" family="Gecarcinucidae" genus="Austrothelphusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="115" phylum="Arthropoda" rank="species" species="wasselli">
<emphasis id="B910EAA6FFD01726FE85D0CDFDE1F9C1" box="[286,542,1962,1984]" italics="true" pageId="6" pageNumber="115">Austrothelphusa wasselli</emphasis>
</taxonomicName>
, holotype male (11.2 × 9.2 mm), Port Stewart District, Cape York (AM-P.14524). A, dorsal view; B, frontal view; C, ventral view.
</paragraph>
</caption>
<caption id="DF1B663CFFD11727FF0CD020FE06F97D" httpUri="https://zenodo.org/record/1135614/files/figure.png" pageId="7" pageNumber="116" targetBox="[154,1439,196,1833]" targetPageId="7">
<paragraph id="8BDB36B4FFD11727FF0CD020FE06F97D" blockId="7.[151,1435,1863,1916]" pageId="7" pageNumber="116">
<emphasis id="B910EAA6FFD11727FF0CD020FEEAF95D" bold="true" box="[151,277,1863,1885]" pageId="7" pageNumber="116">FIGURE 4.</emphasis>
<taxonomicName id="4C644D37FFD11727FE85D020FDE1F95C" baseAuthorityName="Bishop" baseAuthorityYear="1963" box="[286,542,1863,1885]" class="Malacostraca" family="Gecarcinucidae" genus="Austrothelphusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="116" phylum="Arthropoda" rank="species" species="wasselli">
<emphasis id="B910EAA6FFD11727FE85D020FDE1F95C" box="[286,542,1863,1885]" italics="true" pageId="7" pageNumber="116">Austrothelphusa wasselli</emphasis>
</taxonomicName>
, male (25.8 × 20.5 mm) Stewart River, Queensland QM-W20049. A, dorsal view; B, ventral view of sternum and pleon.
</paragraph>
</caption>
<paragraph id="8BDB36B4FFD11727FF5CD0C4FA62F9E1" blockId="7.[151,1437,1955,2016]" pageId="7" pageNumber="116">
Male pleon (
<figureCitation id="135F2A31FFD11727FEC7D0C4FE3EF9BD" box="[348,449,1955,1980]" captionStart="FIGURE 4" captionStartId="7.[151,250,1863,1885]" captionTargetBox="[154,1439,196,1833]" captionTargetId="figure@7.[154,1439,196,1833]" captionTargetPageId="7" captionText="FIGURE 4. Austrothelphusa wasselli, male (25.8 × 20.5 mm) Stewart River, Queensland QM-W20049. A, dorsal view; B, ventral view of sternum and pleon." httpUri="https://zenodo.org/record/1135614/files/figure.png" pageId="7" pageNumber="116">Figs. 4B</figureCitation>
,
<figureCitation id="135F2A31FFD11727FE4BD0C4FE0CF9BD" box="[464,499,1955,1980]" captionStart="FIGURE 6" captionStartId="9.[151,250,1705,1727]" captionTargetBox="[153,1434,890,1668]" captionTargetId="figure@9.[153,1434,877,1684]" captionTargetPageId="9" captionText="FIGURE 6. AC, Austrothelphusa wasselli, male (18.8 × 14.9 mm), Oscar Creek, 2.1 km SSE of Coen, Archer Catchment, Queensland (QM-W29187). DF, A. gilbertensis sp.nov., holotype male (11.2 × 9.2 mm), Port Stewart District, Cape York (AM-P.14524).A, D, ventrofrontal view of cephalothorax (orbit, frontal triangle, epistome and pterygostome); B, E, left major chela; C, F, pleon. Scale line 5 mm." httpUri="https://zenodo.org/record/1135618/files/figure.png" pageId="7" pageNumber="116">6C</figureCitation>
) broadly triangular; telson tapering, apex rounded, length subequal to breadth at base. Somite 6 slightly tapering (proximal width 1.3 times distal width), c. 1.64 times wider at base than long.
</paragraph>
<paragraph id="8BDB36B4FFDE1728FF0CD7F0FDC6FF64" blockId="8.[151,1437,151,824]" pageId="8" pageNumber="117">
Somite 5 more strongly tapering (proximal width 1.6 times distal width), c. 2.97 times wider at base than long. Somite 4 also strongly tapering (proximal width 1.52 times distal width), c. 4.4 times wider at base than long. Somites 2 and 3 broad, with lateral margins evenly rounded, narrow longitudinally. [The figure given of the pleon of
<taxonomicName id="4C644D37FFDE1728FF2FD663FED1FF1C" baseAuthorityName="Bishop" baseAuthorityYear="1963" box="[180,302,260,285]" class="Malacostraca" family="Gecarcinucidae" genus="Austrothelphusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="117" phylum="Arthropoda" rank="species" species="wasselli">
<emphasis id="B910EAA6FFDE1728FF2FD663FED1FF1C" box="[180,302,260,285]" italics="true" pageId="8" pageNumber="117">A. wasselli</emphasis>
</taxonomicName>
by
<bibRefCitation id="EFF54B45FFDE1728FECCD663FDABFF1D" author="Bishop" box="[343,596,260,285]" pageId="8" pageNumber="117" refString="Bishop, J. A. (1963) The Australian freshwater crabs of the family Potamonidae (Crustacea: Decapoda). Australian Journal of Marine and Freshwater Research, 14, 218 - 238. https: // doi. org / 10.1071 / MF 9630218" type="journal article" year="1963">Bishop (1963: fig. 2D)</bibRefCitation>
appears to be in error by showing somite 5 to be longer than somite 6. This is not found in any other species of
<taxonomicName id="4C644D37FFDE1728FDCBD640FCF9FF41" authorityName="Bott" authorityYear="1969" box="[592,774,295,320]" class="Malacostraca" family="Gecarcinucidae" genus="Austrothelphusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="117" phylum="Arthropoda" rank="genus">
<emphasis id="B910EAA6FFDE1728FDCBD640FCF9FF41" box="[592,774,295,320]" italics="true" pageId="8" pageNumber="117">Austrothelphusa</emphasis>
</taxonomicName>
and does not represent the
<typeStatus id="54DF8816FFDE1728FBC3D640FB45FF41" box="[1112,1210,295,320]" pageId="8" pageNumber="117" type="holotype">holotype</typeStatus>
(
<figureCitation id="135F2A31FFDE1728FB54D640FAD5FF41" box="[1231,1322,295,320]" captionStart="FIGURE 3" captionStartId="6.[151,250,1962,1984]" captionTargetBox="[167,1420,304,1924]" captionTargetId="figure@6.[167,1420,304,1924]" captionTargetPageId="6" captionText="FIGURE 3. Austrothelphusa wasselli, holotype male (11.2 × 9.2 mm), Port Stewart District, Cape York (AM-P.14524). A, dorsal view; B, frontal view; C, ventral view." httpUri="https://zenodo.org/record/1135612/files/figure.png" pageId="8" pageNumber="117">Fig. 3C</figureCitation>
) or other topotypic males in the present study.]
</paragraph>
<paragraph id="8BDB36B4FFDE1728FF5CD608FE6AFFF4" blockId="8.[151,1437,151,824]" pageId="8" pageNumber="117">
Walking legs moderately long, total length of P5 (basis to tip of dactyl) c. 1.1 times maximum carapace width. P5 (
<figureCitation id="135F2A31FFDE1728FF5ED6F3FEE1FFAD" box="[197,286,404,429]" captionStart="FIGURE 7" captionStartId="10.[151,250,1893,1915]" captionTargetBox="[176,1409,994,1864]" captionTargetId="figure@10.[156,1440,986,1865]" captionTargetPageId="10" captionText="FIGURE 7. A, C, E, F, Austrothelphusa wasselli, male (25.8 × 20.5 mm), from Stewart River, Queensland (QM-W20049). B, D, G, H, A. gilbertensis sp. nov., holotype male (22.3 × 17.6 mm), Venture Creek, E of Croydon, Gilbert Catchment, Queensland (QM-W28040). A, B, outline of frontal margin and orbits; C, D, fifth pereiopod; EH, left first gonopod (sternal view) with magnification of apical part." httpUri="https://zenodo.org/record/1135620/files/figure.png" pageId="8" pageNumber="117">Fig. 7C</figureCitation>
): merus with anterior and posterior borders subparallel, slightly convex, 2.77 times longer than wide; propodus short, anterior border moderately convex, 1.94 longer than wide; dactylus short, similar in length to propodus (1.04 times).
</paragraph>
<paragraph id="8BDB36B4FFDE1728FF5CD567FCF1FC84" blockId="8.[151,1437,151,824]" pageId="8" pageNumber="117">
G1 (
<figureCitation id="135F2A31FFDE1728FF66D698FE85FC19" box="[253,378,511,536]" captionStart="FIGURE 7" captionStartId="10.[151,250,1893,1915]" captionTargetBox="[176,1409,994,1864]" captionTargetId="figure@10.[156,1440,986,1865]" captionTargetPageId="10" captionText="FIGURE 7. A, C, E, F, Austrothelphusa wasselli, male (25.8 × 20.5 mm), from Stewart River, Queensland (QM-W20049). B, D, G, H, A. gilbertensis sp. nov., holotype male (22.3 × 17.6 mm), Venture Creek, E of Croydon, Gilbert Catchment, Queensland (QM-W28040). A, B, outline of frontal margin and orbits; C, D, fifth pereiopod; EH, left first gonopod (sternal view) with magnification of apical part." httpUri="https://zenodo.org/record/1135620/files/figure.png" pageId="8" pageNumber="117">Fig. 7E, F</figureCitation>
) short, broad basally but tapering and slender over distal half, moderately curved inward apically. Outer lateral margin (in sternal view) relatively straight (slightly concave) over basal two-thirds, before curving inwards towards apex. Terminal opening very small, V-shaped, apical flanges smoothly tapering. Some sparse long simple setae along inner margin towards tip.
</paragraph>
<paragraph id="8BDB36B4FFDE1728FF5CD5E8FE40FCCC" blockId="8.[151,1437,151,824]" pageId="8" pageNumber="117">
<emphasis id="B910EAA6FFDE1728FF5CD5E8FEE5FCA9" bold="true" box="[199,282,655,680]" pageId="8" pageNumber="117">Colour</emphasis>
. Preserved material retains a smattering of tiny red dots on the anterolateral regions of the carapace. Live colours not recorded.
</paragraph>
</subSubSection>
<subSubSection id="C37E653FFFDE1728FF5CD5B0FF14FD39" pageId="8" pageNumber="117" type="distribution">
<paragraph id="8BDB36B4FFDE1728FF5CD5B0FF14FD39" blockId="8.[151,1437,151,824]" pageId="8" pageNumber="117">
<emphasis id="B910EAA6FFDE1728FF5CD5B0FE15FCF1" bold="true" box="[199,490,727,752]" pageId="8" pageNumber="117">Distribution and ecology</emphasis>
. Here recorded from the Stewart and adjacent Archer River Catchments, Cape York,
<collectingRegion id="49A0F856FFDE1728FF0CD59BFEE4FD14" box="[151,283,764,789]" country="Australia" name="Queensland" pageId="8" pageNumber="117">Queensland</collectingRegion>
(
<figureCitation id="135F2A31FFDE1728FEB5D59BFE8CFD15" box="[302,371,764,789]" captionStart="FIGURE 1" captionStartId="1.[151,250,1845,1867]" captionTargetBox="[342,1242,908,1824]" captionTargetId="figure@1.[342,1242,908,1824]" captionTargetPageId="1" captionText="FIGURE 1. Map of Cape York, Queensland showing catchments and localities that were sampled for the present study. In the GilbertCatchment,black pentagon = Venture Creek, dark green hexagon = small branchof GilbertRiver,and purple diamond= Pleasant Creek; in the Normanby Catchment, light green spot= Healy Creek; in the Jeannie catchment, blue oval = Howick River; in the Stewart Catchment, red rectangle = Stewart River; and in the Archer Catchment, yellow triangle = Oscar Creek (see Table 1 for precise localities). The crab shown is the holotype male of Austrothelphusa gilbertensis sp. nov. (refer to Fig. 8)." httpUri="https://zenodo.org/record/1135608/files/figure.png" pageId="8" pageNumber="117">Fig. 1</figureCitation>
).
<typeStatus id="54DF8816FFDE1728FE14D59BFE08FD14" box="[399,503,764,789]" pageId="8" pageNumber="117" type="holotype">Holotype</typeStatus>
was collected from under leaves on the hard bottom of a temporary freshwater lagoon.
</paragraph>
</subSubSection>
<caption id="DF1B663CFFDE1728FF0CD0E3FBD8F9B9" httpUri="https://zenodo.org/record/1135616/files/figure.png" pageId="8" pageNumber="117" targetBox="[154,1439,875,1900]" targetPageId="8">
<paragraph id="8BDB36B4FFDE1728FF0CD0E3FBD8F9B9" blockId="8.[151,1436,1924,1976]" pageId="8" pageNumber="117">
<emphasis id="B910EAA6FFDE1728FF0CD0E3FEEDF998" bold="true" box="[151,274,1924,1946]" pageId="8" pageNumber="117">FIGURE 5.</emphasis>
<taxonomicName id="4C644D37FFDE1728FE82D0E3FDE9F99B" baseAuthorityName="Bishop" baseAuthorityYear="1963" box="[281,534,1924,1946]" class="Malacostraca" family="Gecarcinucidae" genus="Austrothelphusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="117" phylum="Arthropoda" rank="species" species="wasselli">
<emphasis id="B910EAA6FFDE1728FE82D0E3FDE9F99B" box="[281,534,1924,1946]" italics="true" pageId="8" pageNumber="117">Austrothelphusa wasselli</emphasis>
</taxonomicName>
, male (25.8 × 20.5 mm) Stewart River, Queensland QM-W20049. A, ventrofrontal view; B, outer view of major chela; C, ventrofrontal view showing chelae and third maxillipeds.
</paragraph>
</caption>
<subSubSection id="C37E653FFFDF1729FF5CD7F0FA62F9EA" pageId="9" pageNumber="118" type="discussion">
<paragraph id="8BDB36B4FFDF1729FF5CD7F0FBAFFFF4" blockId="9.[151,1437,151,824]" pageId="9" pageNumber="118">
<emphasis id="B910EAA6FFDF1729FF5CD7F0FECBFEB1" bold="true" box="[199,308,151,176]" pageId="9" pageNumber="118">Remarks</emphasis>
.
<bibRefCitation id="EFF54B45FFDF1729FEDDD7F0FE14FEB1" author="Bishop" box="[326,491,151,176]" pageId="9" pageNumber="118" refString="Bishop, J. A. (1963) The Australian freshwater crabs of the family Potamonidae (Crustacea: Decapoda). Australian Journal of Marine and Freshwater Research, 14, 218 - 238. https: // doi. org / 10.1071 / MF 9630218" type="journal article" year="1963">Bishop (1963)</bibRefCitation>
described
<taxonomicName id="4C644D37FFDF1729FDF7D7FFFD15FEB1" baseAuthorityName="Bishop" baseAuthorityYear="1963" box="[620,746,151,176]" class="Malacostraca" family="Gecarcinucidae" genus="Austrothelphusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="118" phylum="Arthropoda" rank="species" species="wasselli">
<emphasis id="B910EAA6FFDF1729FDF7D7FFFD15FEB1" box="[620,746,151,176]" italics="true" pageId="9" pageNumber="118">A. wasselli</emphasis>
</taxonomicName>
based on small specimens collected from the Port Stewart District, Cape York, with carapace widths ranging from
<quantity id="4C9C9B51FFDF1729FC93D7DBFC75FED5" box="[776,906,188,212]" metricMagnitude="-2" metricUnit="m" metricValue="1.35" metricValueMax="1.9" metricValueMin="0.8" pageId="9" pageNumber="118" unit="mm" value="13.5" valueMax="19.0" valueMin="8.0">8 to 19 mm</quantity>
. Thus the biggest male we examined here (
<quantity id="4C9C9B51FFDF1729FAF1D7DBFF3CFEF6" metricMagnitude="-2" metricUnit="m" metricValue="2.58" pageId="9" pageNumber="118" unit="mm" value="25.8">25.8 mm</quantity>
CW) is considerably larger than the
<typeStatus id="54DF8816FFDF1729FDC5D787FD63FEF9" box="[606,668,224,248]" pageId="9" pageNumber="118">types</typeStatus>
, but nevertheless it and our other samples agree in all morphological respects with the
<typeStatus id="54DF8816FFDF1729FEFCD663FE36FF1C" box="[359,457,260,285]" pageId="9" pageNumber="118" type="holotype">holotype</typeStatus>
(
<figureCitation id="135F2A31FFDF1729FE46D663FDDAFF1D" box="[477,549,260,285]" captionStart="FIGURE 3" captionStartId="6.[151,250,1962,1984]" captionTargetBox="[167,1420,304,1924]" captionTargetId="figure@6.[167,1420,304,1924]" captionTargetPageId="6" captionText="FIGURE 3. Austrothelphusa wasselli, holotype male (11.2 × 9.2 mm), Port Stewart District, Cape York (AM-P.14524). A, dorsal view; B, frontal view; C, ventral view." httpUri="https://zenodo.org/record/1135612/files/figure.png" pageId="9" pageNumber="118">Fig. 3</figureCitation>
). Although not designated as part of the
<typeStatus id="54DF8816FFDF1729FB8BD662FBBFFF1C" box="[1040,1088,261,285]" pageId="9" pageNumber="118">type</typeStatus>
series, Bishop also identified samples from the Coen River, at Coen as being conspecific. While these two localities represent two separate catchments, the easterly flowing Stewart Catchment, and the westerly flowing Archer Catchment (
<figureCitation id="135F2A31FFDF1729FB7DD62BFAD8FF65" box="[1254,1319,332,357]" captionStart="FIGURE 1" captionStartId="1.[151,250,1845,1867]" captionTargetBox="[342,1242,908,1824]" captionTargetId="figure@1.[342,1242,908,1824]" captionTargetPageId="1" captionText="FIGURE 1. Map of Cape York, Queensland showing catchments and localities that were sampled for the present study. In the GilbertCatchment,black pentagon = Venture Creek, dark green hexagon = small branchof GilbertRiver,and purple diamond= Pleasant Creek; in the Normanby Catchment, light green spot= Healy Creek; in the Jeannie catchment, blue oval = Howick River; in the Stewart Catchment, red rectangle = Stewart River; and in the Archer Catchment, yellow triangle = Oscar Creek (see Table 1 for precise localities). The crab shown is the holotype male of Austrothelphusa gilbertensis sp. nov. (refer to Fig. 8)." httpUri="https://zenodo.org/record/1135608/files/figure.png" pageId="9" pageNumber="118">Fig. 1</figureCitation>
), we have not found any morphological differences, and the genetic divergence is low (1.8%), as is discussed more fully later. We therefore agree that
<taxonomicName id="4C644D37FFDF1729FE33D6F3FDDCFFAC" baseAuthorityName="Bishop" baseAuthorityYear="1963" box="[424,547,404,429]" class="Malacostraca" family="Gecarcinucidae" genus="Austrothelphusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="118" phylum="Arthropoda" rank="species" species="wasselli">
<emphasis id="B910EAA6FFDF1729FE33D6F3FDDCFFAC" box="[424,547,404,429]" italics="true" pageId="9" pageNumber="118">A. wasselli</emphasis>
</taxonomicName>
does also occur in the Archer Catchment. This seems understandable because the two catchments are separated by only a low and narrow section of the McIlwraith Range to the east of Coen, with the creeks at the headwaters of each catchment being geographically very close.
</paragraph>
<paragraph id="8BDB36B4FFDF1729FF5CD698FF07FD36" blockId="9.[151,1437,151,824]" pageId="9" pageNumber="118">
<bibRefCitation id="EFF54B45FFDF1729FF5CD698FE78FC19" author="Riek" box="[199,391,511,536]" pageId="9" pageNumber="118" refString="Riek, E. F. (1951) The Australian freshwater crabs (Potamonidae). Records of the Australian Museum, 22, 358 - 367." type="journal article" year="1951">Riek (1951: 354)</bibRefCitation>
reported
<emphasis id="B910EAA6FFDF1729FE68D698FD75FC19" box="[499,650,511,536]" italics="true" pageId="9" pageNumber="118">Paratelphusa</emphasis>
(
<taxonomicName id="4C644D37FFDF1729FD01D698FCE1FC19" authorityName="Alcock" authorityYear="1909" box="[666,798,511,536]" class="Malacostraca" family="Gecarcinucidae" genus="Liotelphusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="118" phylum="Arthropoda" rank="genus">
<emphasis id="B910EAA6FFDF1729FD01D698FCE1FC19" box="[666,798,511,536]" italics="true" pageId="9" pageNumber="118">Liotelphusa</emphasis>
</taxonomicName>
)
<emphasis id="B910EAA6FFDF1729FCB4D698FC5EFC19" box="[815,929,511,536]" italics="true" pageId="9" pageNumber="118">planifrons</emphasis>
(
<bibRefCitation id="EFF54B45FFDF1729FC2BD567FBBAFC19" author="Burger" box="[944,1093,512,536]" pageId="9" pageNumber="118" refString="Burger, O. (1894) Beitrage zur Kenntnis der Gattung Telphusa. Zoologische Jahrbucher, Abtheilung fur Systematik, Geographie und Biologie der Thiere, 8, 1 - 7, pl. 1." type="journal article" year="1894">Bürger, 1894</bibRefCitation>
) from the Mitchell Catchment (Walsh River and Mutchilba), but
<bibRefCitation id="EFF54B45FFDF1729FD83D543FCD4FC3D" author="Bishop" box="[536,811,548,573]" pageId="9" pageNumber="118" refString="Bishop, J. A. (1963) The Australian freshwater crabs of the family Potamonidae (Crustacea: Decapoda). Australian Journal of Marine and Freshwater Research, 14, 218 - 238. https: // doi. org / 10.1071 / MF 9630218" type="journal article" year="1963">Bishop (1963: 229, 230)</bibRefCitation>
considered these samples to be a misidentification, and instead re-identified Rieks samples as his own new species,
<taxonomicName id="4C644D37FFDF1729FCDFD52FFB8EFC61" authority="(Bishop, 1963)" baseAuthorityName="Bishop" baseAuthorityYear="1963" box="[836,1137,583,608]" class="Malacostraca" family="Gecarcinucidae" genus="Austrothelphusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="118" phylum="Arthropoda" rank="species" species="wasselli">
<emphasis id="B910EAA6FFDF1729FCDFD52FFC41FC61" box="[836,958,583,608]" italics="true" pageId="9" pageNumber="118">A. wasselli</emphasis>
(
<bibRefCitation id="EFF54B45FFDF1729FC55D520FB96FC61" author="Bishop" box="[974,1129,583,608]" pageId="9" pageNumber="118" refString="Bishop, J. A. (1963) The Australian freshwater crabs of the family Potamonidae (Crustacea: Decapoda). Australian Journal of Marine and Freshwater Research, 14, 218 - 238. https: // doi. org / 10.1071 / MF 9630218" type="journal article" year="1963">Bishop, 1963</bibRefCitation>
)
</taxonomicName>
. However Bishop (as with the Coen River samples), specifically did not include them as part of his
<typeStatus id="54DF8816FFDF1729FC78D50AFBECFC84" box="[995,1043,621,645]" pageId="9" pageNumber="118">type</typeStatus>
series of
<taxonomicName id="4C644D37FFDF1729FB1DD50BFAFEFC84" baseAuthorityName="Bishop" baseAuthorityYear="1963" box="[1158,1281,620,645]" class="Malacostraca" family="Gecarcinucidae" genus="Austrothelphusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="118" phylum="Arthropoda" rank="species" species="wasselli">
<emphasis id="B910EAA6FFDF1729FB1DD50BFAFEFC84" box="[1158,1281,620,645]" italics="true" pageId="9" pageNumber="118">A. wasselli</emphasis>
</taxonomicName>
. As part of a larger study of
<emphasis id="B910EAA6FFDF1729FEDAD5F7FDCEFCA9" box="[321,561,655,680]" italics="true" pageId="9" pageNumber="118">
<taxonomicName id="4C644D37FFDF1729FEDAD5F7FE45FCA9" authority="Naser &amp; Davie &amp; Waltham, 2018" authorityName="Naser &amp; Davie &amp; Waltham" authorityYear="2018" box="[321,442,655,680]" class="Malacostraca" family="Gecarcinucidae" genus="Austrothelphusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="118" phylum="Arthropoda" rank="species" sensu="lato" species="wasselli">A. wasselli</taxonomicName>
<taxonomicNameLabel id="A22357DDFFDF1729FE5AD5F6FDCEFCA9" box="[449,561,655,680]" pageId="9" pageNumber="118" sensu="lato">sensu lato</taxonomicNameLabel>
</emphasis>
, the present authors suspect that the crabs from the Mitchell River Catchment may belong to a discrete species for which the name
<taxonomicName id="4C644D37FFDF1729FC91D5D3FB10FCCD" authority="(McCulloch, 1917)" baseAuthorityName="McCulloch" baseAuthorityYear="1917" box="[778,1263,692,717]" class="Malacostraca" family="Gecarcinucidae" genus="Austrothelphusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="118" phylum="Arthropoda" rank="species" species="plana">
<emphasis id="B910EAA6FFDF1729FC91D5D3FBF4FCCC" box="[778,1035,692,717]" italics="true" pageId="9" pageNumber="118">Austrothelphusa plana</emphasis>
(McCulloch, 1917)
</taxonomicName>
is likely to be available (
<taxonomicName id="4C644D37FFDF1729FE95D5B0FC95FCF1" authority="McCulloch, 1917" authorityName="McCulloch" authorityYear="1917" box="[270,874,727,752]" class="Malacostraca" family="Potamidae" genus="Geothelphusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="118" phylum="Arthropoda" rank="variety" species="leichardti" variety="plana">
<emphasis id="B910EAA6FFDF1729FE95D5B0FDE1FCF1" box="[270,542,727,752]" italics="true" pageId="9" pageNumber="118">Geothelphusa leichardti</emphasis>
var.
<emphasis id="B910EAA6FFDF1729FDC2D5B0FD66FCF1" box="[601,665,727,752]" italics="true" pageId="9" pageNumber="118">plana</emphasis>
McCulloch, 1917
</taxonomicName>
, was described from Eureka Creek, Walsh River, within the Mitchell Catchment), and we have thus not included Rieks identification as part of the synonymy for
<taxonomicName id="4C644D37FFDF1729FA1FD59BFF0EFD39" authorityName="Naser &amp; Davie &amp; Waltham" authorityYear="2018" class="Malacostraca" family="Gecarcinucidae" genus="Austrothelphusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="118" phylum="Arthropoda" rank="species" species="wasselli">
<emphasis id="B910EAA6FFDF1729FA1FD59BFF0EFD39" italics="true" pageId="9" pageNumber="118">A. wasselli</emphasis>
</taxonomicName>
.
</paragraph>
<caption id="DF1B663CFFDF1729FF0CD1CEFDFFF91A" httpUri="https://zenodo.org/record/1135618/files/figure.png" pageId="9" pageNumber="118" targetBox="[153,1434,890,1668]" targetPageId="9">
<paragraph id="8BDB36B4FFDF1729FF0CD1CEFDFFF91A" blockId="9.[151,1436,1705,1819]" pageId="9" pageNumber="118">
<emphasis id="B910EAA6FFDF1729FF0CD1CEFEEBF8BF" bold="true" box="[151,276,1705,1727]" pageId="9" pageNumber="118">FIGURE 6.</emphasis>
AC,
<taxonomicName id="4C644D37FFDF1729FEC0D1CEFDA5F8BE" authorityName="Naser &amp; Davie &amp; Waltham" authorityYear="2018" box="[347,602,1705,1727]" class="Malacostraca" family="Gecarcinucidae" genus="Austrothelphusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="118" phylum="Arthropoda" rank="species" species="wasselli">
<emphasis id="B910EAA6FFDF1729FEC0D1CEFDA5F8BE" box="[347,602,1705,1727]" italics="true" pageId="9" pageNumber="118">Austrothelphusa wasselli</emphasis>
</taxonomicName>
, male (18.8 × 14.9 mm), Oscar Creek, 2.1 km SSE of Coen, Archer Catchment, Queensland (QM-W29187). DF,
<taxonomicName id="4C644D37FFDF1729FE61D1AEFD70F8DF" authority="Naser &amp; Davie &amp; Waltham, 2018" authorityName="Naser &amp; Davie &amp; Waltham" authorityYear="2018" box="[506,655,1736,1758]" class="Malacostraca" family="Gecarcinucidae" genus="Austrothelphusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="118" phylum="Arthropoda" rank="species" species="gilbertensis" status="sp. nov.">
<emphasis id="B910EAA6FFDF1729FE61D1AEFD70F8DF" box="[506,655,1736,1758]" italics="true" pageId="9" pageNumber="118">A. gilbertensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22357DDFFDF1729FD03D1AFFD1DF8DF" box="[664,738,1736,1758]" pageId="9" pageNumber="118" rank="species">
<emphasis id="B910EAA6FFDF1729FD03D1AFFD23F8DC" bold="true" box="[664,732,1736,1757]" pageId="9" pageNumber="118">sp.nov</emphasis>
.
</taxonomicNameLabel>
, holotype male (11.2 × 9.2 mm), Port Stewart District, Cape York (AM-P.14524). A, D, ventrofrontal view of cephalothorax (orbit, frontal triangle, epistome and pterygostome); B, E, left major chela; C, F, pleon. Scale line 5 mm.
</paragraph>
</caption>
<paragraph id="8BDB36B4FFDF1729FF5CD024FA62F9EA" blockId="9.[151,1437,1859,2028]" pageId="9" pageNumber="118">
The
<typeStatus id="54DF8816FFDF1729FF65D023FED1F95D" box="[254,302,1860,1884]" pageId="9" pageNumber="118">type</typeStatus>
locality of “Cape York” for
<emphasis id="B910EAA6FFDF1729FD17D024FC91F95D" box="[652,878,1859,1884]" italics="true" pageId="9" pageNumber="118">Telphusa planifrons</emphasis>
<bibRefCitation id="EFF54B45FFDF1729FCE1D023FBEBF95A" author="Burger" box="[890,1044,1859,1884]" pageId="9" pageNumber="118" refString="Burger, O. (1894) Beitrage zur Kenntnis der Gattung Telphusa. Zoologische Jahrbucher, Abtheilung fur Systematik, Geographie und Biologie der Thiere, 8, 1 - 7, pl. 1." type="journal article" year="1894">Bürger, 1894</bibRefCitation>
, is now considered to have been erroneous, and it seems clear that
<emphasis id="B910EAA6FFDF1729FD8ED00FFD5CF97E" box="[533,675,1894,1919]" italics="true" pageId="9" pageNumber="118">T. planifrons</emphasis>
is generically unrelated to
<taxonomicName id="4C644D37FFDF1729FC4FD001FB74F97E" authorityName="Bott" authorityYear="1969" box="[980,1163,1894,1919]" class="Malacostraca" family="Gecarcinucidae" genus="Austrothelphusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="118" phylum="Arthropoda" rank="genus">
<emphasis id="B910EAA6FFDF1729FC4FD001FB74F97E" box="[980,1163,1894,1919]" italics="true" pageId="9" pageNumber="118">Austrothelphusa</emphasis>
</taxonomicName>
(see
<bibRefCitation id="EFF54B45FFDF1729FB5CD00FFAC5F97E" author="Bott" box="[1223,1338,1895,1919]" pageId="9" pageNumber="118" refString="Bott, R. (1970) Die Susswasserkrabben von Europa, Asien, Australien und ihre Stammesgeschichte. Eine Revision der Potamoidea und der Parathelphusoidea. (Crustacea, Decapoda). Abhandlungen der Senckenbergischen Naturforschenden Gesellschaft, 526, 1 - 338." type="journal article" year="1970">Bott 1970</bibRefCitation>
: 5051).
<bibRefCitation id="EFF54B45FFDF1729FF0CD0ECFEC7F9A5" author="Bishop" box="[151,312,1931,1956]" pageId="9" pageNumber="118" refString="Bishop, J. A. (1963) The Australian freshwater crabs of the family Potamonidae (Crustacea: Decapoda). Australian Journal of Marine and Freshwater Research, 14, 218 - 238. https: // doi. org / 10.1071 / MF 9630218" type="journal article" year="1963">Bishop (1963)</bibRefCitation>
identified six female crabs collected from the Upper Naru River, New
<collectingCountry id="F3737624FFDF1729FBCDD0ECFB58F9A5" box="[1110,1191,1931,1956]" name="Guinea-Bissau" pageId="9" pageNumber="118">Guinea</collectingCountry>
as appearing to agree with Bürgers (1894) description, thus inferring that
<emphasis id="B910EAA6FFDF1729FD70D0C9FC36F9C6" box="[747,969,1966,1991]" italics="true" pageId="9" pageNumber="118">Telphusa planifrons</emphasis>
was most likely a New Guinean species. However,
<bibRefCitation id="EFF54B45FFDF1729FE97D0B3FE6FF9EA" author="Bott" box="[268,400,2003,2027]" pageId="9" pageNumber="118" refString="Bott, R. (1970) Die Susswasserkrabben von Europa, Asien, Australien und ihre Stammesgeschichte. Eine Revision der Potamoidea und der Parathelphusoidea. (Crustacea, Decapoda). Abhandlungen der Senckenbergischen Naturforschenden Gesellschaft, 526, 1 - 338." type="journal article" year="1970">Bott (1970)</bibRefCitation>
pointed out that Bürger compared his
<emphasis id="B910EAA6FFDF1729FCD6D0B3FC22F9ED" box="[845,989,2003,2028]" italics="true" pageId="9" pageNumber="118">T. planifrons</emphasis>
with specimens he had identified as
<emphasis id="B910EAA6FFDF1729FA1CD0B3FA62F9EA" box="[1415,1437,2004,2027]" italics="true" pageId="9" pageNumber="118">T.</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C37E653FFFDC172AFF0CD7FEFACEFC16" pageId="10" pageNumber="119" type="reference_group">
<paragraph id="8BDB36B4FFDC172AFF0CD7FEFACEFC16" blockId="10.[151,1436,151,536]" pageId="10" pageNumber="119">
<taxonomicName id="4C644D37FFDC172AFF0CD7FEFEF2FEB1" ID-CoL="JYF4" authorityName="von Martens" authorityYear="1868" box="[151,269,153,176]" class="Malacostraca" family="Gecarcinucidae" genus="Austrothelphusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="119" phylum="Arthropoda" rank="species" species="transversa">
<emphasis id="B910EAA6FFDC172AFF0CD7FEFEF2FEB1" box="[151,269,153,176]" italics="true" pageId="10" pageNumber="119">transversa</emphasis>
</taxonomicName>
from Calcutta, but that species is otherwise indigenous to
<collectingCountry id="F3737624FFDC172AFC4FD7F0FBC1FEB1" box="[980,1086,151,176]" name="Australia" pageId="10" pageNumber="119">Australia</collectingCountry>
. Thus the most parsimonious explanation is that the labels became mismatched, with the “Indian”
<emphasis id="B910EAA6FFDC172AFC3DD7DAFBCFFED5" box="[934,1072,189,212]" italics="true" pageId="10" pageNumber="119">T. tranversa</emphasis>
coming from
<collectingCountry id="F3737624FFDC172AFB48D7DBFAC2FED4" box="[1235,1341,188,213]" name="Australia" pageId="10" pageNumber="119">Australia</collectingCountry>
, and the “Australian”
<emphasis id="B910EAA6FFDC172AFEABD786FE3CFEF9" box="[304,451,223,248]" italics="true" pageId="10" pageNumber="119">T. planifrons</emphasis>
from Calcutta,
<collectingCountry id="F3737624FFDC172AFD18D7B8FD3FFEF9" box="[643,704,223,248]" name="India" pageId="10" pageNumber="119">India</collectingCountry>
.
<bibRefCitation id="EFF54B45FFDC172AFD54D787FCA9FEF9" author="Bott" box="[719,854,224,248]" pageId="10" pageNumber="119" refString="Bott, R. (1970) Die Susswasserkrabben von Europa, Asien, Australien und ihre Stammesgeschichte. Eine Revision der Potamoidea und der Parathelphusoidea. (Crustacea, Decapoda). Abhandlungen der Senckenbergischen Naturforschenden Gesellschaft, 526, 1 - 338." type="journal article" year="1970">Bott (1970)</bibRefCitation>
somewhat inexplicably placed
<emphasis id="B910EAA6FFDC172AFB56D786FAA0FEF9" box="[1229,1375,223,248]" italics="true" pageId="10" pageNumber="119">T. planifrons</emphasis>
as a subspecies of
<taxonomicName id="4C644D37FFDC172AFEA0D663FDE7FF1C" box="[315,536,260,285]" class="Malacostraca" family="Gecarcinucidae" genus="Liothelphusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="119" phylum="Arthropoda" rank="species" species="laevis">
<emphasis id="B910EAA6FFDC172AFEA0D663FDE7FF1C" box="[315,536,260,285]" italics="true" pageId="10" pageNumber="119">Liothelphusa laevis</emphasis>
</taxonomicName>
, while otherwise stating in his remarks that its closest relative was
<taxonomicName id="4C644D37FFDC172AFAA8D662FEBDFF41" authority="(Alcock, 1909)" baseAuthorityName="Alcock" baseAuthorityYear="1909" class="Malacostraca" family="Gecarcinucidae" genus="Liothelphusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="119" phylum="Arthropoda" rank="species" species="bakeri">
<emphasis id="B910EAA6FFDC172AFAA8D662FA64FF1C" box="[1331,1435,260,285]" italics="true" pageId="10" pageNumber="119">L. bakeri</emphasis>
(Alcock, 1909)
</taxonomicName>
[now in
<emphasis id="B910EAA6FFDC172AFE37D640FDA7FF41" box="[428,600,295,320]" italics="true" pageId="10" pageNumber="119">Globithelphusa</emphasis>
, see Ng
<emphasis id="B910EAA6FFDC172AFD5BD64EFD06FF41" box="[704,761,295,320]" italics="true" pageId="10" pageNumber="119">et al.</emphasis>
2008]. The
<typeStatus id="54DF8816FFDC172AFC13D640FC15FF41" box="[904,1002,295,320]" pageId="10" pageNumber="119" type="holotype">holotype</typeStatus>
of
<emphasis id="B910EAA6FFDC172AFB88D640FB0DFF41" box="[1043,1266,295,320]" italics="true" pageId="10" pageNumber="119">Telphusa planifrons</emphasis>
was originally deposited in the Göttingen Zoological Museum, but is now in the collections of the Senckenberg Museum, Frankfurt (SMF-ZMG699). It will need to be carefully re-examined, but from the description and figures of
<bibRefCitation id="EFF54B45FFDC172AFAD4D617FF1EFFAD" author="Burger" pageId="10" pageNumber="119" refString="Burger, O. (1894) Beitrage zur Kenntnis der Gattung Telphusa. Zoologische Jahrbucher, Abtheilung fur Systematik, Geographie und Biologie der Thiere, 8, 1 - 7, pl. 1." type="journal article" year="1894">Bürger (1894)</bibRefCitation>
and of
<bibRefCitation id="EFF54B45FFDC172AFEAED6F2FD84FFAD" author="Bott" box="[309,635,404,429]" pageId="10" pageNumber="119" refString="Bott, R. (1970) Die Susswasserkrabben von Europa, Asien, Australien und ihre Stammesgeschichte. Eine Revision der Potamoidea und der Parathelphusoidea. (Crustacea, Decapoda). Abhandlungen der Senckenbergischen Naturforschenden Gesellschaft, 526, 1 - 338." type="journal article" year="1970">Bott (1970, pl. 7, figs 7981)</bibRefCitation>
we agree with
<bibRefCitation id="EFF54B45FFDC172AFCBDD6F2FC58FFAD" author="Bott" box="[806,935,404,428]" pageId="10" pageNumber="119" refString="Bott, R. (1970) Die Susswasserkrabben von Europa, Asien, Australien und ihre Stammesgeschichte. Eine Revision der Potamoidea und der Parathelphusoidea. (Crustacea, Decapoda). Abhandlungen der Senckenbergischen Naturforschenden Gesellschaft, 526, 1 - 338." type="journal article" year="1970">Bott (1970)</bibRefCitation>
that it cannot belong to
<taxonomicName id="4C644D37FFDC172AFB2DD6F3FA92FFAC" ID-CoL="7NS5N" authorityName="Bott" authorityYear="1969" box="[1206,1389,404,429]" class="Malacostraca" family="Gecarcinucidae" genus="Austrothelphusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="119" phylum="Arthropoda" rank="genus">
<emphasis id="B910EAA6FFDC172AFB2DD6F3FA92FFAC" box="[1206,1389,404,429]" italics="true" pageId="10" pageNumber="119">Austrothelphusa</emphasis>
</taxonomicName>
and it looks most like a
<emphasis id="B910EAA6FFDC172AFEE7D6D0FDD6FFD1" box="[380,553,439,464]" italics="true" pageId="10" pageNumber="119">Globithelphusa</emphasis>
species as indicated by Ng
<emphasis id="B910EAA6FFDC172AFCF1D6DEFC5BFFD1" box="[874,932,439,464]" italics="true" pageId="10" pageNumber="119">et al.</emphasis>
(2008). In particular, “
<emphasis id="B910EAA6FFDC172AFB36D6D0FA63FFD1" box="[1197,1436,439,464]" italics="true" pageId="10" pageNumber="119">Telphusa” planifrons</emphasis>
has a strongly vaulted carapace, with the body much deeper than in
<taxonomicName id="4C644D37FFDC172AFC11D6BBFBBFFFF4" ID-CoL="7NS5N" authorityName="Bott" authorityYear="1969" box="[906,1088,476,501]" class="Malacostraca" family="Gecarcinucidae" genus="Austrothelphusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="119" phylum="Arthropoda" rank="genus">
<emphasis id="B910EAA6FFDC172AFC11D6BBFBBFFFF4" box="[906,1088,476,501]" italics="true" pageId="10" pageNumber="119">Austrothelphusa</emphasis>
</taxonomicName>
species; and the front is longer significantly more deflexed, and the lateral margins subparallel, unlike anything seen in
<taxonomicName id="4C644D37FFDC172AFBE8D698FAD5FC19" ID-CoL="7NS5N" authorityName="Bott" authorityYear="1969" box="[1139,1322,511,536]" class="Malacostraca" family="Gecarcinucidae" genus="Austrothelphusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="119" phylum="Arthropoda" rank="genus">
<emphasis id="B910EAA6FFDC172AFBE8D698FAD5FC19" box="[1139,1322,511,536]" italics="true" pageId="10" pageNumber="119">Austrothelphusa</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>