225 lines
33 KiB
XML
225 lines
33 KiB
XML
<document id="0AB75EEF4672FB58B9EC8741B0589F88" ID-DOI="10.37520/aemnp.2021.009" ID-ISSN="1804-6487" ID-Zenodo-Dep="5037330" ID-ZooBank="27EA8466-6B29-40F4-8236-DD49061C523B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1624895377019" checkinUser="carolina" docAuthor="Liu, Tian-tian, Ono, Hiroki & Maruyama, Munetoshi" docDate="2021" docId="03CB023BAC1AFF924E54F94F13F1F918" docLanguage="en" docName="ActaEntMusNatPra.61.1.163-201.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 61 (1)" docSource="http://dx.doi.org/10.37520/aemnp.2021.009" docStyle="DocumentStyle:BA2C5C8540539042E893A65A182CEA85.8:ActaEntMusNatPra.2018-.journal_article" docStyleId="BA2C5C8540539042E893A65A182CEA85" docStyleName="ActaEntMusNatPra.2018-.journal_article" docStyleVersion="8" docTitle="Bryothinusa constricta Liu & Ono & Maruyama 2021, sp. nov." docType="treatment" docVersion="3" lastPageNumber="179" masterDocId="FFF27A43AC0AFF824F48FFFB175FFFCD" masterDocTitle="Revision of the intertidal rove beetle genus Bryothinusa from Japan (Coleoptera: Staphylinidae: Aleocharinae)" masterLastPageNumber="201" masterPageNumber="163" pageNumber="179" updateTime="1698977316832" updateUser="plazi" zenodo-license-document="CC-BY-NC-ND-3.0">
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<mods:title id="E82D39A00C0D75EF65CCCB09EC0C96BB">Revision of the intertidal rove beetle genus Bryothinusa from Japan (Coleoptera: Staphylinidae: Aleocharinae)</mods:title>
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<mods:namePart id="F7AFC66F1472AB63B8FD353556B75CA2">Liu, Tian-tian</mods:namePart>
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<mods:affiliation id="B530BA2057B31E0ADC8BF8B0B002D46B">Entomological Laboratory, Graduate School of Bioresource and Bioenvironmental Sciences, Kyushu University, Fukuoka, 819 - 0395, Japan & The Kyushu University Museum, Hakozaki</mods:affiliation>
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<mods:namePart id="02EF302A14024083194D43CDFD645878">Ono, Hiroki</mods:namePart>
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<mods:affiliation id="41949100E65436D99CE72570D8C70980">Sounan, Minami-ku, Sagamihara-shi, Kanagawa, 252 - 0312, Japan</mods:affiliation>
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<mods:namePart id="7806AC5F2D8A70402A89B7FC2AAC5CB2">Maruyama, Munetoshi</mods:namePart>
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<mods:affiliation id="59704EE2788E11DD6E459378AF6A2C22">The Kyushu University Museum, Hakozaki</mods:affiliation>
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<mods:nameIdentifier id="5153DAD70012FC8266E604C5F84A910F" type="email">dendrolasius@gmail.com</mods:nameIdentifier>
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<mods:title id="28EE4E4D59B866798070167E59121A5A">Acta Entomologica Musei Nationalis Pragae</mods:title>
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<mods:date id="A23B5A6AAD4677296D3D35933635A67B">2021</mods:date>
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<mods:number id="2CCEA4C2D3B3672DDFA5FA37BDCE5E0B">2021-05-07</mods:number>
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<treatment id="03CB023BAC1AFF924E54F94F13F1F918" LSID="urn:lsid:plazi:treatment:03CB023BAC1AFF924E54F94F13F1F918" httpUri="http://treatment.plazi.org/id/03CB023BAC1AFF924E54F94F13F1F918" lastPageNumber="179" pageId="16" pageNumber="179">
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<subSubSection id="C378E0A6AC1AFF924E54F94F152CF901" box="[284,627,1716,1740]" pageId="16" pageNumber="179" type="nomenclature">
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<paragraph id="8BDDB32DAC1AFF924E54F94F152CF901" blockId="16.[284,627,1716,1740]" box="[284,627,1716,1740]" pageId="16" pageNumber="179">
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<heading id="D0950441AC1AFF924E54F94F152CF901" box="[284,627,1716,1740]" centered="true" fontSize="10" level="2" pageId="16" pageNumber="179" reason="3">
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<emphasis id="B9166F3FAC1AFF924E54F94F152CF901" bold="true" box="[284,627,1716,1740]" pageId="16" pageNumber="179">
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<taxonomicName id="4C62C8AEAC1AFF924E54F94F1548F901" ID-CoL="8XW4G" authority="Liu & Ono & Maruyama, 2021" authorityName="Liu & Ono & Maruyama" authorityYear="2021" box="[284,535,1716,1740]" class="Insecta" family="Staphylinidae" genus="Bryothinusa" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="179" phylum="Arthropoda" rank="species" species="constricta" status="sp. nov.">
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<emphasis id="B9166F3FAC1AFF924E54F94F1548F901" bold="true" box="[284,535,1716,1740]" italics="true" pageId="16" pageNumber="179">Bryothinusa constricta</emphasis>
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</taxonomicName>
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||
<taxonomicNameLabel id="A225D244AC1AFF924D56F94E152CF901" box="[542,627,1717,1740]" pageId="16" pageNumber="179" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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||
</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C378E0A6AC1AFF924E23F92815D5F8CC" pageId="16" pageNumber="179" type="description">
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<paragraph id="8BDDB32DAC1AFF924E23F928157BF92B" blockId="16.[151,761,1747,1968]" box="[363,548,1747,1766]" pageId="16" pageNumber="179">
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(
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<figureCitation id="1359AFA8AC1AFF924E39F92816E7F92B" box="[369,440,1747,1766]" captionStart="Fig" captionStartId="2.[151,182,1972,1991]" captionTargetBox="[189,1405,205,1944]" captionTargetId="figure-13@2.[175,1411,189,1947]" captionTargetPageId="2" captionText="Fig. 1. Habitus of Bryothinusa spp. A – B. algarum Sawada, 1971; B – B. minuta (Sawada, 1955); C – B. tsutsuii (Sawada, 1955); D – B. koreana Ahn & Jeon, 2004; E – B. yoshigoui sp. nov.; F – B. japonica Liu, Ono & Maruyama, 2020; G – B. sakishimana Sawada, 1991; H – B. constricta sp. nov.; I – B. hokkaidensis sp. nov.; J – B. fulvipennis sp. nov.; K – B. itsuroi sp. nov.; L – B. aikoae sp. nov.; M – B. okinawana sp. nov.; N – B. nigra sp. nov.; O – B. nakanei (Sawada, 1955); P – B. moriguchii sp. nov.; Q – B. gangjinensis Ahn & Jeon, 2004. Scale bars: 1.00 mm." figureDoi="http://doi.org/10.5281/zenodo.5037332" httpUri="https://zenodo.org/record/5037332/files/figure.png" pageId="16" pageNumber="179">Figs 1H</figureCitation>
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||
;
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<figureCitation id="1359AFA8AC1AFF924E89F9281681F92B" box="[449,478,1747,1766]" captionStart="Fig" captionStartId="4.[151,182,1978,1997]" captionTargetBox="[196,1391,190,1955]" captionTargetId="figure-13@4.[193,1394,189,1958]" captionTargetPageId="4" captionText="Fig. 2. Compound eyes of Bryothinusa spp.A – B. algarum Sawada, 1971; B – B. minuta (Sawada, 1955); C – B. tsutsuii (Sawada, 1955); D – B. koreana Ahn & Jeon, 2004; E – B. yoshigoui sp. nov.; F – B. japonica Liu, Ono & Maruyama, 2020; G – B. sakishimana Sawada, 1991; H – B. constricta sp. nov.; I – B. hokkaidensis sp. nov.; J – B. fulvipennis sp. nov.; K – B. itsuroi sp. nov.; L – B. aikoae sp. nov.; M – B. okinawana sp. nov.; N – B. nigra sp. nov.; O – B. nakanei (Sawada, 1955); P – B. moriguchii sp. nov.; Q – B. gangjinensis Ahn & Jeon, 2004. Scale bars: 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.5037334" httpUri="https://zenodo.org/record/5037334/files/figure.png" pageId="16" pageNumber="179">2H</figureCitation>
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||
;
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<figureCitation id="1359AFA8AC1AFF924EAFF92816A1F92B" box="[487,510,1747,1766]" captionStart="Fig" captionStartId="17.[150,181,2034,2053]" captionTargetBox="[153,1427,191,1935]" captionTargetId="figure-14@17.[150,1435,191,1953]" captionTargetPageId="17" captionText="Fig. 10. Bryothinusa constricta sp. nov.A – labrum; B – mandible, dorsal view; C – lacinia, dorsal view; D – labium; E – mentum; F – male sternite VIII; G – female sternite VIII; H – median lobe, ventral view; I – median lobe, lateral view; J – paramere; K – spermatheca. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5037352" httpUri="https://zenodo.org/record/5037352/files/figure.png" pageId="16" pageNumber="179">10</figureCitation>
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;
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<figureCitation id="1359AFA8AC1AFF924D4FF9281542F92B" box="[519,541,1747,1766]" captionStart="Fig" captionStartId="35.[150,181,1388,1407]" captionTargetBox="[150,1434,191,1370]" captionTargetId="figure-236@35.[150,1435,191,1371]" captionTargetPageId="35" captionText="Fig. 22. Map of examined specimens of Bryothinusa spp. from Japan: white inverted triangle – B. algarum Sawada, 1971; black inverted triangle – B. minuta (Sawada, 1955); white star – B. tsutsuii (Sawada, 1955); black star – B. koreana Ahn & Jeon, 2004; black square – B. yoshigoui sp. nov.; white triangle – B. japonica Liu, Ono & Maruyama, 2020; black diamond – B. sakishimana Sawada, 1991; white diamond – B. constricta sp. nov.; white square – B. hokkaidensis sp. nov.; black triangle – B. fulvipennis sp. nov.; double circle – B. itsuroi sp. nov.; arrow – B. aikoae sp. nov.; black hart – B. okinawana sp. nov.; circled dot – B. nigra sp. nov.; white circle – B. nakanei (Sawada, 1955); white hart – B. moriguchii sp. nov.; black circle – B. gangjinensis Ahn & Jeon, 2004." figureDoi="http://doi.org/10.5281/zenodo.5037376" httpUri="https://zenodo.org/record/5037376/files/figure.png" pageId="16" pageNumber="179">22</figureCitation>
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)
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</paragraph>
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||
<paragraph id="8BDDB32DAC1AFF924E4EF91515D5F8CC" blockId="16.[151,761,1747,1968]" box="[262,650,1774,1793]" pageId="16" pageNumber="179">(Japanese name: Kubire-nagisa-hanekakushi)</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C378E0A6AC1AFF924FDFF8EC16C4F87D" pageId="16" pageNumber="179" type="materials_examined">
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<paragraph id="8BDDB32DAC1AFF924FDFF8EC16C4F87D" blockId="16.[151,761,1747,1968]" pageId="16" pageNumber="179">
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<emphasis id="B9166F3FAC1AFF924FDFF8EC1679F8E7" bold="true" box="[151,294,1815,1834]" pageId="16" pageNumber="179">Type material.</emphasis>
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Hඈඅඈඍඒඉൾ: J,
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<emphasis id="B9166F3FAC1AFF924EF4F8EC1532F8E7" bold="true" box="[444,621,1815,1834]" pageId="16" pageNumber="179">
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<collectingCountry id="F375F3BDAC1AFF924EF4F8EC1558F8E7" box="[444,519,1815,1834]" name="Japan" pageId="16" pageNumber="179">JAPAN</collectingCountry>
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: HඈඇඌΗǕ:
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</emphasis>
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Futtsu-misaki, Futtsu-shi, Chiba-ken,
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<date id="FFDC95EDAC1AFF924E13F8C916F9F888" box="[347,422,1842,1861]" pageId="16" pageNumber="179" value="2014-05-01">1.v.2014</date>
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, H. Ono (
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<collectionCode id="ED732BE8AC1AFF924EB3F8C9156EF888" box="[507,561,1842,1861]" country="Ghana" lsid="urn:lsid:biocol.org:col:13924" name="Resource Management Support Center" pageId="16" pageNumber="179" type="Herbarium">KUM</collectionCode>
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). Pൺ*©ൺඍඒඉൾඌ:
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<emphasis id="B9166F3FAC1AFF924DECF8C915A6F888" bold="true" box="[676,761,1842,1861]" pageId="16" pageNumber="179">HඈඇඌΗǕ:</emphasis>
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<specimenCount id="9D6478A4AC1AFF924FDFF8B61797F892" box="[151,200,1862,1888]" pageId="16" pageNumber="179" type="female">3 ♀♀</specimenCount>
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, 6 unsexed spec., same locality as
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<typeStatus id="54D90D8FAC1AFF924EB7F8B61513F8AD" box="[511,588,1869,1888]" pageId="16" pageNumber="179" type="holotype">holotype</typeStatus>
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||
,
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<date id="FFDC95EDAC1AFF924D1EF8B615F3F8AD" box="[598,684,1869,1888]" pageId="16" pageNumber="179" value="2010-05-28">28.v.2010</date>
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, H. Ono (
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<collectionCode id="ED732BE8AC1AFF924FD5F89C178CF8B7" box="[157,211,1895,1914]" country="Ghana" lsid="urn:lsid:biocol.org:col:13924" name="Resource Management Support Center" pageId="16" pageNumber="179" type="Herbarium">KUM</collectionCode>
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||
); 2 JJ
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||
<specimenCount id="9D6478A4AC1AFF924E52F89C1615F8B7" box="[282,330,1889,1914]" pageId="16" pageNumber="179" type="female">5 ♀♀</specimenCount>
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||
, 10 unsexed spec., same data as
|
||
<typeStatus id="54D90D8FAC1AFF924D20F89C15ECF8B7" box="[616,691,1895,1914]" pageId="16" pageNumber="179" type="holotype">holotype</typeStatus>
|
||
(
|
||
<collectionCode id="ED732BE8AC1AFF924DF7F89C15ABF8B7" box="[703,756,1895,1914]" country="Ghana" lsid="urn:lsid:biocol.org:col:13924" name="Resource Management Support Center" pageId="16" pageNumber="179" type="Herbarium">KUM</collectionCode>
|
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,
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||
<collectionCode id="ED732BE8AC1AFF924FDFF879178AF858" box="[151,213,1922,1941]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="16" pageNumber="179">NMPC</collectionCode>
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||
,
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<collectionCode id="ED732BE8AC1AFF924F94F8791640F858" box="[220,287,1922,1941]" pageId="16" pageNumber="179">BMNH</collectionCode>
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||
);
|
||
<specimenCount id="9D6478A4AC1AFF924E65F8791614F858" box="[301,331,1916,1941]" pageId="16" pageNumber="179" type="female">1 ♀</specimenCount>
|
||
, 4 unsexed spec., Izumo, Koryô-chô, Shimane-ken,
|
||
<date id="FFDC95EDAC1AFF924FDFF86617B6F87D" box="[151,233,1949,1968]" pageId="16" pageNumber="179" value="2007-09-03">3.ix.2007</date>
|
||
,
|
||
<collectionCode id="ED732BE8AC1AFF924FBAF866165FF87D" box="[242,256,1949,1968]" name="Tavera, Department of Geology and Geophysics" pageId="16" pageNumber="179">T</collectionCode>
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||
. Hayama (
|
||
<collectionCode id="ED732BE8AC1AFF924E13F86616CEF87D" box="[347,401,1949,1968]" country="Ghana" lsid="urn:lsid:biocol.org:col:13924" name="Resource Management Support Center" pageId="16" pageNumber="179" type="Herbarium">KUM</collectionCode>
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||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C378E0A6AC1AFF924FDFF833123BFA98" pageId="16" pageNumber="179" type="description">
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<paragraph id="8BDDB32DAC1AFF924FDFF83313C2FF38" blockId="16.[151,760,1992,2047]" lastBlockId="16.[828,1437,190,1749]" pageId="16" pageNumber="179">
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<emphasis id="B9166F3FAC1AFF924FDFF833167DF812" bold="true" box="[151,290,1992,2015]" pageId="16" pageNumber="179">Description.</emphasis>
|
||
Body (
|
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<figureCitation id="1359AFA8AC1AFF924E39F833169CF812" box="[369,451,1992,2015]" captionStart="Fig" captionStartId="2.[151,182,1972,1991]" captionTargetBox="[189,1405,205,1944]" captionTargetId="figure-13@2.[175,1411,189,1947]" captionTargetPageId="2" captionText="Fig. 1. Habitus of Bryothinusa spp. A – B. algarum Sawada, 1971; B – B. minuta (Sawada, 1955); C – B. tsutsuii (Sawada, 1955); D – B. koreana Ahn & Jeon, 2004; E – B. yoshigoui sp. nov.; F – B. japonica Liu, Ono & Maruyama, 2020; G – B. sakishimana Sawada, 1991; H – B. constricta sp. nov.; I – B. hokkaidensis sp. nov.; J – B. fulvipennis sp. nov.; K – B. itsuroi sp. nov.; L – B. aikoae sp. nov.; M – B. okinawana sp. nov.; N – B. nigra sp. nov.; O – B. nakanei (Sawada, 1955); P – B. moriguchii sp. nov.; Q – B. gangjinensis Ahn & Jeon, 2004. Scale bars: 1.00 mm." figureDoi="http://doi.org/10.5281/zenodo.5037332" httpUri="https://zenodo.org/record/5037332/files/figure.png" pageId="16" pageNumber="179">Fig. 1H</figureCitation>
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) small, about 1.7–2.0 mm in length, covered with minute setae. Abdomen almost black and darker than head and pronotum; elytra lighter than pronotum; legs yellowish brown.
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</paragraph>
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<paragraph id="8BDDB32DAC1AFF924C12FF0514BBFDD8" blockId="16.[828,1437,190,1749]" pageId="16" pageNumber="179">
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Head evenly rounded, more or less produced anteriorly, wider than long, shorter than pronotum and as long as elytra; eyes (
|
||
<figureCitation id="1359AFA8AC1AFF924C8FFEC51344FE98" box="[967,1051,318,341]" captionStart="Fig" captionStartId="4.[151,182,1978,1997]" captionTargetBox="[196,1391,190,1955]" captionTargetId="figure-13@4.[193,1394,189,1958]" captionTargetPageId="4" captionText="Fig. 2. Compound eyes of Bryothinusa spp.A – B. algarum Sawada, 1971; B – B. minuta (Sawada, 1955); C – B. tsutsuii (Sawada, 1955); D – B. koreana Ahn & Jeon, 2004; E – B. yoshigoui sp. nov.; F – B. japonica Liu, Ono & Maruyama, 2020; G – B. sakishimana Sawada, 1991; H – B. constricta sp. nov.; I – B. hokkaidensis sp. nov.; J – B. fulvipennis sp. nov.; K – B. itsuroi sp. nov.; L – B. aikoae sp. nov.; M – B. okinawana sp. nov.; N – B. nigra sp. nov.; O – B. nakanei (Sawada, 1955); P – B. moriguchii sp. nov.; Q – B. gangjinensis Ahn & Jeon, 2004. Scale bars: 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.5037334" httpUri="https://zenodo.org/record/5037334/files/figure.png" pageId="16" pageNumber="179">Fig. 2H</figureCitation>
|
||
) large, setae present between about 20–22 facets, slightly longer than tempora length; antenna yellowish brown, equal to total length of head and pronotum; segment I 1.6 times longer than II, almost 3.5 times longer than III, and longer than XI; antennal segments III–X almost wider than long, and VI–VII wider than width of III–IV.
|
||
</paragraph>
|
||
<paragraph id="8BDDB32DAC1AFF924C12FDE51202FC58" blockId="16.[828,1437,190,1749]" pageId="16" pageNumber="179">
|
||
Mouthparts (
|
||
<figureCitation id="1359AFA8AC1AFF924CAFFDE51331FDF8" box="[999,1134,542,565]" captionStart="Fig" captionStartId="17.[150,181,2034,2053]" captionTargetBox="[153,1427,191,1935]" captionTargetId="figure-14@17.[150,1435,191,1953]" captionTargetPageId="17" captionText="Fig. 10. Bryothinusa constricta sp. nov.A – labrum; B – mandible, dorsal view; C – lacinia, dorsal view; D – labium; E – mentum; F – male sternite VIII; G – female sternite VIII; H – median lobe, ventral view; I – median lobe, lateral view; J – paramere; K – spermatheca. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5037352" httpUri="https://zenodo.org/record/5037352/files/figure.png" pageId="16" pageNumber="179">Figs 10A–E</figureCitation>
|
||
) with distribution of labral setae as in
|
||
<figureCitation id="1359AFA8AC1AFF924CE4FDC51378FD98" box="[940,1063,574,597]" captionStart="Fig" captionStartId="17.[150,181,2034,2053]" captionTargetBox="[153,1427,191,1935]" captionTargetId="figure-14@17.[150,1435,191,1953]" captionTargetPageId="17" captionText="Fig. 10. Bryothinusa constricta sp. nov.A – labrum; B – mandible, dorsal view; C – lacinia, dorsal view; D – labium; E – mentum; F – male sternite VIII; G – female sternite VIII; H – median lobe, ventral view; I – median lobe, lateral view; J – paramere; K – spermatheca. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5037352" httpUri="https://zenodo.org/record/5037352/files/figure.png" pageId="16" pageNumber="179">Figure 10A</figureCitation>
|
||
, with large and thick spiniform seta at antero-laterally; mandibles (
|
||
<figureCitation id="1359AFA8AC1AFF924B34FDA51386FDB8" box="[1148,1241,606,629]" captionStart="Fig" captionStartId="17.[150,181,2034,2053]" captionTargetBox="[153,1427,191,1935]" captionTargetId="figure-14@17.[150,1435,191,1953]" captionTargetPageId="17" captionText="Fig. 10. Bryothinusa constricta sp. nov.A – labrum; B – mandible, dorsal view; C – lacinia, dorsal view; D – labium; E – mentum; F – male sternite VIII; G – female sternite VIII; H – median lobe, ventral view; I – median lobe, lateral view; J – paramere; K – spermatheca. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5037352" httpUri="https://zenodo.org/record/5037352/files/figure.png" pageId="16" pageNumber="179">Fig. 10B</figureCitation>
|
||
) elongate triangle, asymmetrical, right mandible with median tooth; median tooth shorter than 1/4 of mandibles; left mandible without tooth, blunt at apex; maxilla with apical margin of lacinia (
|
||
<figureCitation id="1359AFA8AC1AFF924C0CFD2514FDFD38" box="[836,930,734,757]" captionStart="Fig" captionStartId="17.[150,181,2034,2053]" captionTargetBox="[153,1427,191,1935]" captionTargetId="figure-14@17.[150,1435,191,1953]" captionTargetPageId="17" captionText="Fig. 10. Bryothinusa constricta sp. nov.A – labrum; B – mandible, dorsal view; C – lacinia, dorsal view; D – labium; E – mentum; F – male sternite VIII; G – female sternite VIII; H – median lobe, ventral view; I – median lobe, lateral view; J – paramere; K – spermatheca. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5037352" httpUri="https://zenodo.org/record/5037352/files/figure.png" pageId="16" pageNumber="179">Fig. 10C</figureCitation>
|
||
) at right angle; at tip with two spines very small and not closely interdigitating; 3 setae on inner margin of lacinia; labial palpus (
|
||
<figureCitation id="1359AFA8AC1AFF924B64FCE513D2FCF8" box="[1068,1165,798,821]" captionStart="Fig" captionStartId="17.[150,181,2034,2053]" captionTargetBox="[153,1427,191,1935]" captionTargetId="figure-14@17.[150,1435,191,1953]" captionTargetPageId="17" captionText="Fig. 10. Bryothinusa constricta sp. nov.A – labrum; B – mandible, dorsal view; C – lacinia, dorsal view; D – labium; E – mentum; F – male sternite VIII; G – female sternite VIII; H – median lobe, ventral view; I – median lobe, lateral view; J – paramere; K – spermatheca. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5037352" httpUri="https://zenodo.org/record/5037352/files/figure.png" pageId="16" pageNumber="179">Fig. 10D</figureCitation>
|
||
) with most setae located basally; mentum (
|
||
<figureCitation id="1359AFA8AC1AFF924B40FCC51334FC98" box="[1032,1131,830,853]" captionStart="Fig" captionStartId="17.[150,181,2034,2053]" captionTargetBox="[153,1427,191,1935]" captionTargetId="figure-14@17.[150,1435,191,1953]" captionTargetPageId="17" captionText="Fig. 10. Bryothinusa constricta sp. nov.A – labrum; B – mandible, dorsal view; C – lacinia, dorsal view; D – labium; E – mentum; F – male sternite VIII; G – female sternite VIII; H – median lobe, ventral view; I – median lobe, lateral view; J – paramere; K – spermatheca. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5037352" httpUri="https://zenodo.org/record/5037352/files/figure.png" pageId="16" pageNumber="179">Fig. 10E</figureCitation>
|
||
) with antero-lateral angles prolonged into spinose processes; anterior margin deeply truncate; small “v seta” far away from apical setae.
|
||
</paragraph>
|
||
<paragraph id="8BDDB32DAC1AFF924C12FC65121AFC78" blockId="16.[828,1437,190,1749]" box="[858,1349,926,949]" pageId="16" pageNumber="179">Pronotum longer than elytra, wider than head.</paragraph>
|
||
<paragraph id="8BDDB32DAC1AFF924C12FC451344FBD8" blockId="16.[828,1437,190,1749]" pageId="16" pageNumber="179">Elytra widened apically, wider than long, almost as wide as pronotum; hind wings present but very small, apparently flightless.</paragraph>
|
||
<paragraph id="8BDDB32DAC1AFF924C12FBE51240FBB8" blockId="16.[828,1437,190,1749]" pageId="16" pageNumber="179">
|
||
Male abdominal sternite VIII (
|
||
<figureCitation id="1359AFA8AC1AFF924BD6FBE513A5FBF8" box="[1182,1274,1054,1077]" captionStart="Fig" captionStartId="17.[150,181,2034,2053]" captionTargetBox="[153,1427,191,1935]" captionTargetId="figure-14@17.[150,1435,191,1953]" captionTargetPageId="17" captionText="Fig. 10. Bryothinusa constricta sp. nov.A – labrum; B – mandible, dorsal view; C – lacinia, dorsal view; D – labium; E – mentum; F – male sternite VIII; G – female sternite VIII; H – median lobe, ventral view; I – median lobe, lateral view; J – paramere; K – spermatheca. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5037352" httpUri="https://zenodo.org/record/5037352/files/figure.png" pageId="16" pageNumber="179">Fig. 10F</figureCitation>
|
||
) with posterior margin slightly serrate. Median lobe of aedeagus as in
|
||
<figureCitation id="1359AFA8AC1AFF924C74FBA51486FBB8" box="[828,985,1118,1141]" captionStart="Fig" captionStartId="17.[150,181,2034,2053]" captionTargetBox="[153,1427,191,1935]" captionTargetId="figure-14@17.[150,1435,191,1953]" captionTargetPageId="17" captionText="Fig. 10. Bryothinusa constricta sp. nov.A – labrum; B – mandible, dorsal view; C – lacinia, dorsal view; D – labium; E – mentum; F – male sternite VIII; G – female sternite VIII; H – median lobe, ventral view; I – median lobe, lateral view; J – paramere; K – spermatheca. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5037352" httpUri="https://zenodo.org/record/5037352/files/figure.png" pageId="16" pageNumber="179">Figures 10H, I</figureCitation>
|
||
; paramere (
|
||
<figureCitation id="1359AFA8AC1AFF924B1EFBA513F1FBB8" box="[1110,1198,1118,1141]" captionStart="Fig" captionStartId="17.[150,181,2034,2053]" captionTargetBox="[153,1427,191,1935]" captionTargetId="figure-14@17.[150,1435,191,1953]" captionTargetPageId="17" captionText="Fig. 10. Bryothinusa constricta sp. nov.A – labrum; B – mandible, dorsal view; C – lacinia, dorsal view; D – labium; E – mentum; F – male sternite VIII; G – female sternite VIII; H – median lobe, ventral view; I – median lobe, lateral view; J – paramere; K – spermatheca. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5037352" httpUri="https://zenodo.org/record/5037352/files/figure.png" pageId="16" pageNumber="179">Fig. 10J</figureCitation>
|
||
) elongate.
|
||
</paragraph>
|
||
<paragraph id="8BDDB32DAC1AFF924C12FB8512C2FB18" blockId="16.[828,1437,190,1749]" pageId="16" pageNumber="179">
|
||
Female abdominal sternite VIII (
|
||
<figureCitation id="1359AFA8AC1AFF924B98FB851268FB58" box="[1232,1335,1150,1173]" captionStart="Fig" captionStartId="17.[150,181,2034,2053]" captionTargetBox="[153,1427,191,1935]" captionTargetId="figure-14@17.[150,1435,191,1953]" captionTargetPageId="17" captionText="Fig. 10. Bryothinusa constricta sp. nov.A – labrum; B – mandible, dorsal view; C – lacinia, dorsal view; D – labium; E – mentum; F – male sternite VIII; G – female sternite VIII; H – median lobe, ventral view; I – median lobe, lateral view; J – paramere; K – spermatheca. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5037352" httpUri="https://zenodo.org/record/5037352/files/figure.png" pageId="16" pageNumber="179">Fig. 10G</figureCitation>
|
||
) slightly serrate alike male on posterior margin. Spermatheca (
|
||
<figureCitation id="1359AFA8AC1AFF924A39FB651434FB18" captionStart="Fig" captionStartId="17.[150,181,2034,2053]" captionTargetBox="[153,1427,191,1935]" captionTargetId="figure-14@17.[150,1435,191,1953]" captionTargetPageId="17" captionText="Fig. 10. Bryothinusa constricta sp. nov.A – labrum; B – mandible, dorsal view; C – lacinia, dorsal view; D – labium; E – mentum; F – male sternite VIII; G – female sternite VIII; H – median lobe, ventral view; I – median lobe, lateral view; J – paramere; K – spermatheca. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5037352" httpUri="https://zenodo.org/record/5037352/files/figure.png" pageId="16" pageNumber="179">Fig. 10K</figureCitation>
|
||
) simplified in shape, about 1.5 times coiled near base.
|
||
</paragraph>
|
||
<paragraph id="8BDDB32DAC1AFF924C74FB25123BFA98" blockId="16.[828,1437,190,1749]" pageId="16" pageNumber="179">
|
||
<emphasis id="B9166F3FAC1AFF924C74FB2514B5FB38" bold="true" box="[828,1002,1246,1269]" pageId="16" pageNumber="179">Measurements.</emphasis>
|
||
BL 1.70–2.00, FBL 0.80–1.00, HL 0.30– 0.31, HW 0.35–0.39, PL 0.30–0.33, PW 0.38–0.40, EL 0.29–0.30, EW 0.36–0.40, HTL 0.30–0.31, AL 0.61–0.67, AS-I 0.11–0.14, AS-II 0.06–0.09, AS-III 0.03–0.04.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C378E0A6AC1AFF924C74FAA5130FFA18" pageId="16" pageNumber="179" type="diagnosis">
|
||
<paragraph id="8BDDB32DAC1AFF924C74FAA5130FFA18" blockId="16.[828,1437,190,1749]" pageId="16" pageNumber="179">
|
||
<emphasis id="B9166F3FAC1AFF924C74FAA5136DFAB8" bold="true" box="[828,1074,1374,1397]" pageId="16" pageNumber="179">Differential diagnosis.</emphasis>
|
||
In general appearance, the species is similar to
|
||
<taxonomicName id="4C62C8AEAC1AFF924CF4FA851301FA58" authorityName="Sawada" authorityYear="1991" box="[956,1118,1405,1429]" class="Insecta" family="Staphylinidae" genus="Bryothinusa" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="179" phylum="Arthropoda" rank="species" species="sakishimana">
|
||
<emphasis id="B9166F3FAC1AFF924CF4FA851301FA58" box="[956,1118,1405,1429]" italics="true" pageId="16" pageNumber="179">B. sakishimana</emphasis>
|
||
</taxonomicName>
|
||
, but easily distinguished from it by the elytral shape being narrowed anteriorly and the shape of the spermatheca.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C378E0A6AC1AFF924C74FA2514B1F9D8" pageId="16" pageNumber="179" type="etymology">
|
||
<paragraph id="8BDDB32DAC1AFF924C74FA2514B1F9D8" blockId="16.[828,1437,190,1749]" pageId="16" pageNumber="179">
|
||
<emphasis id="B9166F3FAC1AFF924C74FA2514E1FA38" bold="true" box="[828,958,1502,1525]" pageId="16" pageNumber="179">Etymology.</emphasis>
|
||
Bringing attention to the constricted body shape; adjective.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C378E0A6AC1AFF924C74F9E512C2F9F8" box="[828,1437,1566,1589]" pageId="16" pageNumber="179" type="distribution">
|
||
<paragraph id="8BDDB32DAC1AFF924C74F9E512C2F9F8" blockId="16.[828,1437,190,1749]" box="[828,1437,1566,1589]" pageId="16" pageNumber="179">
|
||
<emphasis id="B9166F3FAC1AFF924C74F9E51491F9F8" bold="true" box="[828,974,1566,1589]" pageId="16" pageNumber="179">Distribution.</emphasis>
|
||
<collectingCountry id="F375F3BDAC1AFF924C9BF9E5134FF9F8" box="[979,1040,1566,1589]" name="Japan" pageId="16" pageNumber="179">Japan</collectingCountry>
|
||
(
|
||
<emphasis id="B9166F3FAC1AFF924B56F9E513DFF9F8" bold="true" box="[1054,1152,1566,1589]" pageId="16" pageNumber="179">Honshû:</emphasis>
|
||
Chiba-ken, Shimane-ken).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C378E0A6AC1AFF924C74F9C513F1F918" pageId="16" pageNumber="179" type="discussion">
|
||
<paragraph id="8BDDB32DAC1AFF924C74F9C513F1F918" blockId="16.[828,1437,190,1749]" pageId="16" pageNumber="179">
|
||
<emphasis id="B9166F3FAC1AFF924C74F9C514F5F998" bold="true" box="[828,938,1598,1621]" pageId="16" pageNumber="179">Remarks.</emphasis>
|
||
This species is found mainly on the
|
||
<collectingCountry id="F375F3BDAC1AFF924A7AF9C51230F998" box="[1330,1391,1598,1621]" name="Japan" pageId="16" pageNumber="179">Japan</collectingCountry>
|
||
Sea coast of western
|
||
<collectingCountry id="F375F3BDAC1AFF924CBFF9A5136BF9B8" box="[1015,1076,1630,1653]" name="Japan" pageId="16" pageNumber="179">Japan</collectingCountry>
|
||
and east of
|
||
<collectingRegion id="49A67DCFAC1AFF924BF7F9A5125AF9B8" box="[1215,1285,1630,1653]" country="Japan" name="Tokyo" pageId="16" pageNumber="179">Tôkyô</collectingRegion>
|
||
Bay, but it is probably occurring in other areas of mainland
|
||
<collectingCountry id="F375F3BDAC1AFF924A77F9851220F958" box="[1343,1407,1662,1685]" name="Japan" pageId="16" pageNumber="179">Japan</collectingCountry>
|
||
. A distribution map of examined specimens of this species in
|
||
<collectingCountry id="F375F3BDAC1AFF924C1FF94514CBF918" box="[855,916,1726,1749]" name="Japan" pageId="16" pageNumber="179">Japan</collectingCountry>
|
||
is illustrated in
|
||
<figureCitation id="1359AFA8AC1AFF924B08F94513F5F918" box="[1088,1194,1726,1749]" captionStart="Fig" captionStartId="35.[150,181,1388,1407]" captionTargetBox="[150,1434,191,1370]" captionTargetId="figure-236@35.[150,1435,191,1371]" captionTargetPageId="35" captionText="Fig. 22. Map of examined specimens of Bryothinusa spp. from Japan: white inverted triangle – B. algarum Sawada, 1971; black inverted triangle – B. minuta (Sawada, 1955); white star – B. tsutsuii (Sawada, 1955); black star – B. koreana Ahn & Jeon, 2004; black square – B. yoshigoui sp. nov.; white triangle – B. japonica Liu, Ono & Maruyama, 2020; black diamond – B. sakishimana Sawada, 1991; white diamond – B. constricta sp. nov.; white square – B. hokkaidensis sp. nov.; black triangle – B. fulvipennis sp. nov.; double circle – B. itsuroi sp. nov.; arrow – B. aikoae sp. nov.; black hart – B. okinawana sp. nov.; circled dot – B. nigra sp. nov.; white circle – B. nakanei (Sawada, 1955); white hart – B. moriguchii sp. nov.; black circle – B. gangjinensis Ahn & Jeon, 2004." figureDoi="http://doi.org/10.5281/zenodo.5037376" httpUri="https://zenodo.org/record/5037376/files/figure.png" pageId="16" pageNumber="179">Figure 22</figureCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |