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<document id="746821AC86ECA2944A3C9475E732FEF0" ID-DOI="10.5281/zenodo.214045" ID-GBIF-Dataset="4f2b3286-d252-48f5-b634-d48921980fef" ID-ISSN="1175-5326" ID-Zenodo-Dep="214045" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460527612126" checkinUser="plazi" docAuthor="Lemaitre, Rafael &amp; Felder, Darryl L." docDate="2012" docId="03B09F17FF98FFD990C2FB080FAE665D" docLanguage="en" docName="zt03480p079.pdf" docOrigin="Zootaxa 3480" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Areopaguristes tudgei Lemaitre &amp; Felder, 2012, n. sp." docType="treatment" docVersion="9" lastPageNumber="76" masterDocId="FF89E76FFF99FFD09055FFE60C276309" masterDocTitle="A new species of the hermit crab genus Areopaguriste s Rahayu &amp; McLaughlin, 2010 (Crustacea: Decapoda: Anomura: Diogenidae) discovered in the Mesoamerican Barrier Reef of Belize, Caribbean Sea" masterLastPageNumber="79" masterPageNumber="67" pageNumber="68" updateTime="1698315309371" updateUser="plazi">
<mods:mods id="DE8808D85CCCCB200BB169798D0A1C1F" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="94FE547EC167308CA523EA9B91F95B2B">
<mods:title id="9EA1E411148CE9969FF9BA8AED6651A1">A new species of the hermit crab genus Areopaguriste s Rahayu &amp; McLaughlin, 2010 (Crustacea: Decapoda: Anomura: Diogenidae) discovered in the Mesoamerican Barrier Reef of Belize, Caribbean Sea</mods:title>
</mods:titleInfo>
<mods:name id="A4CE52F8AB049530759FF73F761071AE" type="personal">
<mods:role id="E2B7E57F954DB8AD29F8CDBCC9E1202B">
<mods:roleTerm id="0B07576E85F61D416FB70ABF560381AA">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="C06B95D816CFCDB6E2EC790547278B81">Lemaitre, Rafael</mods:namePart>
</mods:name>
<mods:name id="76DE4DF6A759CF7D733A821898E3C412" type="personal">
<mods:role id="B4D4E318321834710C6E72DF929383D8">
<mods:roleTerm id="B49DCE05C1B3ACFEE971F953BF2876FC">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="5798A05DA2135482C2D2ED1DD4CA9568">Felder, Darryl L.</mods:namePart>
</mods:name>
<mods:typeOfResource id="3612E6C504DF0C07376238FE1042C5AB">text</mods:typeOfResource>
<mods:relatedItem id="57E9ED01941059CB6298C7403F87E523" type="host">
<mods:titleInfo id="CB4FF73658716E2391B6A79E15E5FE35">
<mods:title id="E049DBFE6D48A0A66DF4414F6B9F0B95">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="48BA0B0F358C72BCBF1F482FBA02C5D7">
<mods:date id="BE27F48F2A4AE7001F6658357C3A9EE0">2012</mods:date>
<mods:detail id="17757566E531C70496608EA11C7A3CDE" type="volume">
<mods:number id="EA53651CBD644363166734A3BC377CE2">3480</mods:number>
</mods:detail>
<mods:extent id="ED18A21B27FD344D44E5935C75814346" unit="page">
<mods:start id="4F3E016310F722D7CB2D5577AACF267D">67</mods:start>
<mods:end id="27C257038C582DC6AAF06EC899B64EEF">79</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="EA1764EA92CCCEB0B67F5EA2AC66BD03">journal article</mods:classification>
<mods:identifier id="8B78DDE0F8247E915A55C5D8F4EF847D" type="DOI">10.5281/zenodo.214045</mods:identifier>
<mods:identifier id="A3B7D2B76D4F3B3C5AA1B599D49372C6" type="GBIF-Dataset">4f2b3286-d252-48f5-b634-d48921980fef</mods:identifier>
<mods:identifier id="5DE2D0122AD0F94804D0D09B10BD03D7" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="7E5EAB41D3C420589A4E54457E577FCA" type="Zenodo-Dep">214045</mods:identifier>
</mods:mods>
<treatment id="03B09F17FF98FFD990C2FB080FAE665D" ID-DOI="http://doi.org/10.5281/zenodo.6180685" ID-GBIF-Taxon="119526472" ID-Zenodo-Dep="6180685" LSID="urn:lsid:plazi:treatment:03B09F17FF98FFD990C2FB080FAE665D" httpUri="http://treatment.plazi.org/id/03B09F17FF98FFD990C2FB080FAE665D" lastPageId="9" lastPageNumber="76" pageId="1" pageNumber="68">
<subSubSection id="C3037D8AFF98FFD190C2FB080D356620" pageId="1" pageNumber="68" type="nomenclature">
<paragraph id="8BA62E01FF98FFD190C2FB080DCE6601" blockId="1.[151,489,1262,1321]" box="[151,489,1262,1288]" pageId="1" pageNumber="68">
<heading id="D0EE996DFF98FFD190C2FB080DCE6601" bold="true" box="[151,489,1262,1288]" fontSize="11" level="1" pageId="1" pageNumber="68" reason="1">
<emphasis id="B96DF213FF98FFD190C2FB080DCE6601" bold="true" box="[151,489,1262,1288]" pageId="1" pageNumber="68">
<taxonomicName id="4C195582FF98FFD190C2FB080DB86601" ID-CoL="8c9c2161-2d9b-4ce2-a375-99a2a1f1a39b" box="[151,415,1262,1288]" class="Malacostraca" family="Diogenidae" genus="Areopaguristes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="68" phylum="Arthropoda" rank="species" species="tudgei" status="sp. nov.">
<emphasis id="B96DF213FF98FFD190C2FB080DB86601" bold="true" box="[151,415,1262,1288]" italics="true" pageId="1" pageNumber="68">Areopaguristes tudgei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25E4F68FF98FFD191F2FB080DCE6601" box="[423,489,1262,1288]" pageId="1" pageNumber="68" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BA62E01FF98FFD190C2FAF70D356620" blockId="1.[151,489,1262,1321]" box="[151,274,1297,1321]" pageId="1" pageNumber="68">
(
<figureCitation id="13223284FF98FFD190CAFAF70D2E6620" box="[159,265,1297,1321]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStart-4="FIGURE 5" captionStart-5="FIGURE 6" captionStartId-0="3.[151,250,1589,1612]" captionStartId-1="4.[151,250,1683,1706]" captionStartId-2="5.[151,250,1316,1339]" captionStartId-3="6.[151,250,1819,1842]" captionStartId-4="7.[151,250,1852,1875]" captionStartId-5="8.[151,250,1256,1279]" captionTargetBox-0="[166,1428,367,1565]" captionTargetBox-1="[161,1433,205,1651]" captionTargetBox-2="[159,1428,529,1289]" captionTargetBox-3="[162,1432,331,1795]" captionTargetBox-4="[173,1424,276,1818]" captionTargetBox-5="[151,1436,553,1234]" captionTargetId-0="figure@3.[151,1436,353,1573]" captionTargetId-1="figure@4.[151,1436,193,1662]" captionTargetId-2="figure@5.[151,1436,517,1295]" captionTargetId-3="figure@6.[151,1436,317,1806]" captionTargetId-4="figure@7.[151,1436,265,1831]" captionTargetId-5="figure@8.[151,1436,553,1235]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionTargetPageId-3="6" captionTargetPageId-4="7" captionTargetPageId-5="8" captionText-0="FIGURE 1. Areopaguristes tudgei n. sp., Carrie Bow Cay, Belize: a, c – e, h, male paratype, 2.1 mm ,, February 8 (USNM 1156993); b, f, g, i, female paratype, 2.1 mm, February 4 (USNM 1156991). a, shield and cephalic appendages, dorsal view; b, anterior portion of shield, dorsal view; c, male left first pleopod, posterior view; d, left second pleopod, posterior view; e, male left third pleopod, posterior view; f, female left first pleopod, posterior view; g, female left third pleopod, posterior view; h, uropods and telson, dorsal view; i, telson, dorsal view. Scales equal 0.5 mm (a, b, e, h, i), 0.2 mm (c, d, f, g)." captionText-1="FIGURE 2. Areopaguristes tudgei n. sp.: male paratype, 2.1 mm, Carrie Bow Cay, Belize, February 8 (USNM 1156993). Left mouthparts, internal view: a, mandible; b, maxillule; c, maxilla; d, first maxilliped; e, second maxilliped; f, third maxilliped, g, ischium of same with crista dentata. Scales equal 0.1 mm (a – d, g), and 0.2 mm (e, f)." captionText-2="FIGURE 3. Areopaguristes tudgei n. sp.: male paratype, 2.1 mm, Carrie Bow Cay, Belize, February 8 (USNM 1156993). Chelipeds, all dorsal view except f (mesial), and c (ventral); a, b, showing approximate density of setation; c – f, with setae omitted. a, d, left; b, e, right; c, fingers of same; f, right. Scale equals 0.5 mm." captionText-3="FIGURE 4. Areopaguristes tudgei n. sp.: male paratype, 2.1 mm, Carrie Bow Cay, Belize, February 8 (USNM 1156993). a, second right pereopod with plumose setae, lateral view; b, dactyl of same, mesial view; c, same pereopod with setae omitted, mesial view; d, third right pereopod, lateral view; e, dactyl of same; f, propodus and dactyl of right fourth pereopod, lateral view; g, propodus and dactyl of right fifth pereopod, lateral view. Scales equal 0.5 mm." captionText-4="FIGURE 5. Areopaguristes tudgei n. sp.: male paratype, 2.1 mm, Carrie Bow Cay, Belize, February 8 (USNM 1156993). Sternites XI – XIV, coxae of chelipeds and second to fifth pereopods, and anterior portion of pleon with first and second pleopods, ventral view. Scale equals 0.5 mm." captionText-5="FIGURE 6. Areopaguristes tudgei n. sp., Carrie Bow Cay, Belize, February 4: a, female paratype 1.9 mm (ULLZ 12530); b, male paratype 3.0 mm (ULLZ 12576). Scales equal 1 mm." httpUri-0="https://zenodo.org/record/214046/files/figure.png" httpUri-1="https://zenodo.org/record/214047/files/figure.png" httpUri-2="https://zenodo.org/record/214048/files/figure.png" httpUri-3="https://zenodo.org/record/214049/files/figure.png" httpUri-4="https://zenodo.org/record/214050/files/figure.png" httpUri-5="https://zenodo.org/record/214051/files/figure.png" pageId="1" pageNumber="68">Figs. 1–6</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3037D8AFF98FFD190C2FABF0FD4652C" pageId="1" pageNumber="68" type="materials_examined">
<paragraph id="8BA62E01FF98FFD190C2FABF0FD4652C" blockId="1.[151,1437,1369,2005]" pageId="1" pageNumber="68">
<emphasis id="B96DF213FF98FFD190C2FABF0DA6667B" bold="true" box="[151,385,1369,1394]" pageId="1" pageNumber="68">Material examined.</emphasis>
<typeStatus id="54A290A3FF98FFD191DCFABF0DD26678" box="[393,501,1369,1393]" pageId="1" pageNumber="68" type="holotype">Holotype</typeStatus>
: ♂
<quantity id="4CE183E4FF98FFD19248FABF0E516678" box="[541,630,1369,1393]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="1" pageNumber="68" unit="mm" value="2.5">2.5 mm</quantity>
, Carrie Bow Cay,
<collectingCountry id="F30E6E91FF98FFD19313FABF0FB66678" box="[838,913,1369,1393]" name="Belize" pageId="1" pageNumber="68">Belize</collectingCountry>
,
<geoCoordinate id="EE2D48C6FF98FFD193C8FABF08276678" box="[925,1024,1369,1393]" direction="north" orientation="latitude" pageId="1" pageNumber="68" precision="925" value="16.8">16°48’N</geoCoordinate>
,
<geoCoordinate id="EE2D48C6FF98FFD19459FABF08546678" box="[1036,1139,1369,1393]" direction="west" orientation="longitude" pageId="1" pageNumber="68" precision="925" value="-88.083336">88°05’W</geoCoordinate>
, front reef crest (E side of island), under rocks, intertidal,
<date id="FFA708C1FF98FFD191A3FA9B0E9E669C" box="[502,697,1405,1429]" pageId="1" pageNumber="68" value="2011-02-05">February 5, 2011</date>
, colls. C. Tudge, D. Felder, J. Felder, R. Lemaitre, and S. Pecnick (
<collectionCode id="ED08B6C4FF98FFD190CBFA470CD666B0" LSID="urn:lsid:biocol.org:col:34871" box="[158,241,1441,1465]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="1" pageNumber="68">USNM</collectionCode>
1156990).
<typeStatus id="54A290A3FF98FFD1913AFA440DF866B3" box="[367,479,1442,1466]" pageId="1" pageNumber="68" type="paratype">Paratypes</typeStatus>
, all same locality and year as
<typeStatus id="54A290A3FF98FFD1937EFA470FB766B0" box="[811,912,1441,1465]" pageId="1" pageNumber="68" type="holotype">holotype</typeStatus>
,
<date id="FFA708C1FF98FFD193CEFA47083166B0" box="[923,1046,1441,1465]" pageId="1" pageNumber="68">February 4</date>
,
<specimenCount id="9D1FE588FF98FFD19477FA47086C66B0" box="[1058,1099,1441,1465]" pageId="1" pageNumber="68" type="generic">2 ♂</specimenCount>
1.9,
<quantity id="4CE183E4FF98FFD194D6FA4708FD66B0" box="[1155,1242,1441,1465]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" pageId="1" pageNumber="68" unit="mm" value="2.1">2.1 mm</quantity>
, 8 ♀
<quantity id="4CE183E4FF98FFD19543FA4709BB66B0" box="[1302,1436,1441,1465]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" metricValueMax="2.1" metricValueMin="1.5" pageId="1" pageNumber="68" unit="mm" value="1.8" valueMax="2.1" valueMin="1.5">1.5–2.1 mm</quantity>
(
<collectionCode id="ED08B6C4FF98FFD190CBFA230CD666D4" LSID="urn:lsid:biocol.org:col:34871" box="[158,241,1477,1501]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="1" pageNumber="68">USNM</collectionCode>
1156991); 2 ♀, 1.9 (color voucher),
<quantity id="4CE183E4FF98FFD192CBFA230ED066D4" box="[670,759,1477,1501]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" pageId="1" pageNumber="68" unit="mm" value="2.1">2.1 mm</quantity>
(
<collectionCode id="ED08B6C4FF98FFD19352FA230F7766D4" box="[775,848,1477,1501]" httpUri="http://grbio.org/cool/ggzt-j23h" name="University of Louisiana at Layafette, Zoological Collection" pageId="1" pageNumber="68">ULLZ</collectionCode>
12530);
<specimenCount id="9D1FE588FF98FFD193EFFA230FC166D4" box="[954,998,1477,1501]" pageId="1" pageNumber="68" type="generic">2 ♂</specimenCount>
2.9, 3.0 (color voucher), 1 ♀
<quantity id="4CE183E4FF98FFD19511FA2309BB66D4" box="[1348,1436,1477,1501]" metricMagnitude="-3" metricUnit="m" metricValue="2.6" pageId="1" pageNumber="68" unit="mm" value="2.6">2.6 mm</quantity>
(
<collectionCode id="ED08B6C4FF98FFD190CBFA0F0CC06508" box="[158,231,1513,1537]" httpUri="http://grbio.org/cool/ggzt-j23h" name="University of Louisiana at Layafette, Zoological Collection" pageId="1" pageNumber="68">ULLZ</collectionCode>
12576);
<date id="FFA708C1FF98FFD19119FA0F0DC86508" box="[332,495,1513,1537]" pageId="1" pageNumber="68" value="2010-02-05">
February 5,
<specimenCount id="9D1FE588FF98FFD19186FA0F0DC86508" box="[467,495,1513,1537]" pageId="1" pageNumber="68" type="generic">10</specimenCount>
</date>
♂ 1.0–
<quantity id="4CE183E4FF98FFD19217FA0F0EBC6508" box="[578,667,1513,1537]" metricMagnitude="-3" metricUnit="m" metricValue="2.7" pageId="1" pageNumber="68" unit="mm" value="2.7">2.7 mm</quantity>
, 33 ♀ 1.0–
<quantity id="4CE183E4FF98FFD19340FA0F0F4C6508" box="[789,875,1513,1537]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" pageId="1" pageNumber="68" unit="mm" value="2.1">2.1 mm</quantity>
, 1 ovig. ♀ 2.0 mm (
<collectionCode id="ED08B6C4FF98FFD1941FFA0F08BA6508" LSID="urn:lsid:biocol.org:col:34871" box="[1098,1181,1513,1537]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="1" pageNumber="68">USNM</collectionCode>
1156992);
<date id="FFA708C1FF98FFD1954EFA0F09B06508" box="[1307,1431,1513,1537]" pageId="1" pageNumber="68">February 8</date>
,
<specimenCount id="9D1FE588FF98FFD190C2F9EB0CE6652C" box="[151,193,1549,1573]" pageId="1" pageNumber="68" type="generic">9 ♂</specimenCount>
<quantity id="4CE183E4FF98FFD1909DF9EB0D76652C" box="[200,337,1549,1573]" metricMagnitude="-3" metricUnit="m" metricValue="2.15" metricValueMax="2.7" metricValueMin="1.6" pageId="1" pageNumber="68" unit="mm" value="2.15" valueMax="2.7" valueMin="1.6">1.6–2.7 mm</quantity>
, 9 ♀
<quantity id="4CE183E4FF98FFD191D8F9EB0E31652C" box="[397,534,1549,1573]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" metricValueMax="1.8" metricValueMin="1.4" pageId="1" pageNumber="68" unit="mm" value="1.6" valueMax="1.8" valueMin="1.4">1.4–1.8 mm</quantity>
, 8 ovig. ♀
<quantity id="4CE183E4FF98FFD192C6F9EB0F3D652C" box="[659,794,1549,1573]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" metricValueMax="2.5" metricValueMin="1.5" pageId="1" pageNumber="68" unit="mm" value="2.0" valueMax="2.5" valueMin="1.5">1.5–2.5 mm</quantity>
(
<collectionCode id="ED08B6C4FF98FFD1937DF9EB0F5D652C" LSID="urn:lsid:biocol.org:col:34871" box="[808,890,1549,1573]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="1" pageNumber="68">USNM</collectionCode>
1156993).
</paragraph>
</subSubSection>
<subSubSection id="C3037D8AFF98FFD19092F9D70DBA6428" pageId="1" pageNumber="68" type="diagnosis">
<paragraph id="8BA62E01FF98FFD19092F9D70DBA6428" blockId="1.[151,1437,1369,2005]" pageId="1" pageNumber="68">
<emphasis id="B96DF213FF98FFD19092F9D70D666543" bold="true" box="[199,321,1585,1610]" pageId="1" pageNumber="68">Diagnosis.</emphasis>
Gills biserial. Shield unarmed or at most with few minute spines on anterodorsal margins. Ocular peduncles about half length of shield, decreasing in width distally. Chelipeds with dorsal surfaces of chelae and carpi densely covered with plumose setae obscuring armature beneath. Second and third pereopods with dense fringe of plumose setae on dorsal margins of meri, carpi, propodi and dactyls. Telson with terminal margins oblique, setose, each side with 1–5 small to minute, blunt to sharp calcareous spines. Male first pleopod with distal margins of external lobe and inferior lamella entire, lacking spines, teeth or setae. Female lacking brood pouch. Color as in description.
</paragraph>
</subSubSection>
<subSubSection id="C3037D8AFF98FFD29092F8CB0FBC6215" lastPageId="2" lastPageNumber="69" pageId="1" pageNumber="68" type="description">
<paragraph id="8BA62E01FF98FFD29092F8CB0FBC6215" blockId="1.[151,1437,1369,2005]" lastBlockId="2.[151,1436,151,284]" lastPageId="2" lastPageNumber="69" pageId="1" pageNumber="68">
<emphasis id="B96DF213FF98FFD19092F8CB0D7E644F" bold="true" box="[199,345,1837,1862]" pageId="1" pageNumber="68">Description.</emphasis>
Gills biserial. Shield (
<figureCitation id="13223284FF98FFD1920FF8CB0EBC644C" box="[602,667,1837,1861]" captionStart="FIGURE 1" captionStartId="3.[151,250,1589,1612]" captionTargetBox="[166,1428,367,1565]" captionTargetId="figure@3.[151,1436,353,1573]" captionTargetPageId="3" captionText="FIGURE 1. Areopaguristes tudgei n. sp., Carrie Bow Cay, Belize: a, c – e, h, male paratype, 2.1 mm ,, February 8 (USNM 1156993); b, f, g, i, female paratype, 2.1 mm, February 4 (USNM 1156991). a, shield and cephalic appendages, dorsal view; b, anterior portion of shield, dorsal view; c, male left first pleopod, posterior view; d, left second pleopod, posterior view; e, male left third pleopod, posterior view; f, female left first pleopod, posterior view; g, female left third pleopod, posterior view; h, uropods and telson, dorsal view; i, telson, dorsal view. Scales equal 0.5 mm (a, b, e, h, i), 0.2 mm (c, d, f, g)." httpUri="https://zenodo.org/record/214046/files/figure.png" pageId="1" pageNumber="68">Fig. 1</figureCitation>
a) subtriangular, about 1.2 times as long as broad, posterior margin about 0.3 shield width; dorsal surface weakly convex, generally smooth or at most with few minute spines on anterior dorsolateral margins, with plumose setae lateromedially and scattered short setae elsewhere; ventrolateral margin with minute spine distally. Anterolateral margins sloping. Anterior margins between rostrum and lateral projections concave. Lateral projections triangular, ending in small sharp or blunt spine. Rostrum narrowly triangular, often slender (
<figureCitation id="13223284FF9BFFD291EFFF710DDA63A6" box="[442,509,151,175]" captionStart="FIGURE 1" captionStartId="3.[151,250,1589,1612]" captionTargetBox="[166,1428,367,1565]" captionTargetId="figure@3.[151,1436,353,1573]" captionTargetPageId="3" captionText="FIGURE 1. Areopaguristes tudgei n. sp., Carrie Bow Cay, Belize: a, c – e, h, male paratype, 2.1 mm ,, February 8 (USNM 1156993); b, f, g, i, female paratype, 2.1 mm, February 4 (USNM 1156991). a, shield and cephalic appendages, dorsal view; b, anterior portion of shield, dorsal view; c, male left first pleopod, posterior view; d, left second pleopod, posterior view; e, male left third pleopod, posterior view; f, female left first pleopod, posterior view; g, female left third pleopod, posterior view; h, uropods and telson, dorsal view; i, telson, dorsal view. Scales equal 0.5 mm (a, b, e, h, i), 0.2 mm (c, d, f, g)." httpUri="https://zenodo.org/record/214046/files/figure.png" pageId="2" pageNumber="69">Fig. 1</figureCitation>
a, b), extending distinctly beyond lateral projections, ending in rounded tip with several setae. Ocular peduncles (including corneas) about half length of shield, decreasing in width distally, with plumose setae dorsally; corneas slightly dilated; ocular acicles subtriangular, separated by rostrum, each ending in sharp spine and with 1–3 additional spines on oblique lateral margin.
</paragraph>
</subSubSection>
<caption id="DF667E89FF9BFFD290C2FEAD0DA762D9" pageId="2" pageNumber="69">
<paragraph id="8BA62E01FF9BFFD290C2FEAD0DA762D9" blockId="2.[151,1436,331,1822]" pageId="2" pageNumber="69">
<emphasis id="B96DF213FF9BFFD290C2FEAD0D2C626A" bold="true" box="[151,267,331,355]" pageId="2" pageNumber="69">TABLE 1</emphasis>
. Checklist of species of
<taxonomicName id="4C195582FF9BFFD2926BFEAA0F62626A" authority="Dana, 1851" authorityName="Dana" authorityYear="1851" box="[574,837,332,355]" class="Malacostraca" family="Diogenidae" genus="Paguristes" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="69" phylum="Arthropoda" rank="genus">
<emphasis id="B96DF213FF9BFFD2926BFEAA0E97626A" box="[574,688,332,355]" italics="true" pageId="2" pageNumber="69">Paguristes</emphasis>
Dana, 1851
</taxonomicName>
,
<taxonomicName id="4C195582FF9BFFD2930FFEAA0945626A" authority="Rahayu &amp; McLaughlin, 2010" authorityName="Rahayu &amp; McLaughlin" authorityYear="2010" box="[858,1378,332,355]" class="Malacostraca" family="Diogenidae" genus="Areopaguristes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="69" phylum="Arthropoda" rank="genus">
<emphasis id="B96DF213FF9BFFD2930FFEAA0FDB626A" box="[858,1020,332,355]" italics="true" pageId="2" pageNumber="69">Areopaguristes</emphasis>
Rahayu &amp; McLaughlin, 2010
</taxonomicName>
, and
<taxonomicName id="4C195582FF9BFFD290C2FE8A0E00628A" authority="McLaughlin, 2002" authorityName="McLaughlin" authorityYear="2002" box="[151,551,364,387]" class="Malacostraca" family="Diogenidae" genus="Pseudopaguristes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="69" phylum="Arthropoda" rank="genus">
<emphasis id="B96DF213FF9BFFD290C2FE8A0D72628A" box="[151,341,364,387]" italics="true" pageId="2" pageNumber="69">Pseudopaguristes</emphasis>
McLaughlin, 2002
</taxonomicName>
, from the western Atlantic, including primary synonyms (after McLaughlin
<emphasis id="B96DF213FF9BFFD29530FE8A09BB628A" box="[1381,1436,364,387]" italics="true" pageId="2" pageNumber="69">et al.</emphasis>
2010). Asterisks indicate taxa currently assigned to the
<taxonomicName id="4C195582FF9BFFD292B3FE6A0F9E62AA" box="[742,953,396,419]" class="Malacostraca" family="Diogenidae" genus="Paguristes" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="69" phylum="Arthropoda" rank="species" species="tortugae">
<emphasis id="B96DF213FF9BFFD292B3FE6A0F9E62AA" box="[742,953,396,419]" italics="true" pageId="2" pageNumber="69">Paguristes tortugae</emphasis>
</taxonomicName>
complex.
<taxonomicName id="4C195582FF9BFFD290C2FE5F0DA762D9" authority="Dana, 1851" authorityName="Dana" authorityYear="1851" box="[151,384,441,464]" class="Malacostraca" family="Diogenidae" genus="Paguristes" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="69" phylum="Arthropoda" rank="genus">
<emphasis id="B96DF213FF9BFFD290C2FE5F0D2262C6" bold="true" box="[151,261,441,463]" italics="true" pageId="2" pageNumber="69">Paguristes</emphasis>
Dana, 1851
</taxonomicName>
</paragraph>
</caption>
<subSubSection id="C3037D8AFF9BFFD890C2FE0E0D0D62FD" lastPageId="8" lastPageNumber="75" pageId="2" pageNumber="69" type="reference_group">
<paragraph id="8BA62E01FF9BFFD290C2FE0E0DAB6130" blockId="2.[151,1436,331,1822]" pageId="2" pageNumber="69">
<taxonomicName id="4C195582FF9BFFD290C2FE0E0EA962F5" ID-CoL="491f23ac-bebe-420d-a079-d0956184eeff" authority="McLaughlin &amp; Provenzano, 1974" authorityName="McLaughlin &amp; Provenzano" authorityYear="1974" box="[151,654,486,509]" class="Malacostraca" family="Diogenidae" genus="Paguristes" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="69" phylum="Arthropoda" rank="species" species="angustitheca">
<emphasis id="B96DF213FF9BFFD290C2FE0E0D1662F4" box="[151,305,487,509]" italics="true" pageId="2" pageNumber="69">P. angustitheca</emphasis>
<bibRefCitation id="EF8853F0FF9BFFD29162FE000EA962F5" author="McLaughlin" box="[311,654,486,508]" pageId="2" pageNumber="69" refString="McLaughlin, P. A. &amp; Provenzano, A. J., Jr. (1974) The genus Paguristes (Crustacea: Decapoda: Diogenidae) from the western Atlantic. Part I. The Paguristes tortugae complex, with notes on variation. Bulletin of Marine Science, 24 (1), 165 - 234, figs 1 - 27." type="journal article" year="1974">McLaughlin &amp; Provenzano, 1974</bibRefCitation>
</taxonomicName>
*
<taxonomicName id="4C195582FF9BFFD290C2FDE00D8E6113" ID-CoL="9491f3bb-dbee-4e88-a93f-10d4e4d4a3ef" authority="Bouvier, 1918" authorityName="Bouvier" authorityYear="1918" box="[151,425,517,539]" class="Malacostraca" family="Diogenidae" genus="Paguristes" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="69" phylum="Arthropoda" rank="species" species="anomalus">
<emphasis id="B96DF213FF9BFFD290C2FDE00D356112" box="[151,274,517,539]" italics="true" pageId="2" pageNumber="69">P. anomalus</emphasis>
<bibRefCitation id="EF8853F0FF9BFFD2914DFDE30D8E6113" author="Bouvier" box="[280,425,517,539]" pageId="2" pageNumber="69" refString="Bouvier, E. - L. (1918) Sur une petite collection de Crustaces de Cuba offerte au Museum par M. de Boury. Bulletin du Museum national d'Histoire naturelle, 24, 6 - 15." type="journal article" year="1918">Bouvier, 1918</bibRefCitation>
</taxonomicName>
(=
<taxonomicName id="4C195582FF9BFFD29199FDE00F3A6112" ID-CoL="8a601d4b-05e6-47ce-b55c-a4c8af7783a2" authority="Provenzano, 1965" authorityName="Provenzano" authorityYear="1965" box="[460,797,517,539]" class="Malacostraca" family="Diogenidae" genus="Paguristes" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="69" phylum="Arthropoda" rank="species" species="anaryballus">
<emphasis id="B96DF213FF9BFFD29199FDE00E7A6112" box="[460,605,517,539]" italics="true" pageId="2" pageNumber="69">P. anaryballus</emphasis>
<bibRefCitation id="EF8853F0FF9BFFD29231FDE00F3A6112" author="Provenzano" box="[612,797,517,539]" pageId="2" pageNumber="69" refString="Provenzano, A. J., Jr. (1965) Two new West Indian hermit crabs of the genus Paguristes (Crustacea: Diogenidae). Bulletin of Marine Science, 15 (3), 726 - 736." type="journal article" year="1965">Provenzano, 1965</bibRefCitation>
</taxonomicName>
)*
<taxonomicName id="4C195582FF9BFFD290C2FDC30DAB6130" ID-CoL="9ea3ea08-9fe6-4cfc-b3d5-cd8fd83f37ba" authority="Forest, 1954" authorityName="Forest" authorityYear="1954" box="[151,396,547,570]" class="Malacostraca" family="Diogenidae" genus="Paguristes" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="69" phylum="Arthropoda" rank="species" species="cadenati">
<emphasis id="B96DF213FF9BFFD290C2FDC30D206133" box="[151,263,548,570]" italics="true" pageId="2" pageNumber="69">P. cadenati</emphasis>
<bibRefCitation id="EF8853F0FF9BFFD29158FDC30DAB6130" author="Forest" box="[269,396,547,570]" pageId="2" pageNumber="69" refString="Forest, J. (1954) Sur un pagure littoral nouveau de la Martinique, Paguristes cadenati sp. nov. Bulletin du Museum national d'Histoire naturelle, (2) 26 (3), 353 - 357." type="journal article" year="1954">Forest, 1954</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="8BA62E01FF9BFFD290C2FDA50D8A617E" blockId="2.[151,1436,331,1822]" pageId="2" pageNumber="69">
<taxonomicName id="4C195582FF9BFFD290C2FDA50D9F6151" ID-CoL="05926aea-fb00-4a41-aa3f-4d6a2e458bd3" authority="Stimpson, 1859" authorityName="Stimpson" authorityYear="1859" box="[151,440,578,600]" class="Malacostraca" family="Diogenidae" genus="Paguristes" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="69" phylum="Arthropoda" rank="species" species="depressus">
<emphasis id="B96DF213FF9BFFD290C2FDA50D346151" box="[151,275,578,600]" italics="true" pageId="2" pageNumber="69">P. depressus</emphasis>
<bibRefCitation id="EF8853F0FF9BFFD2914CFDA40D9F6151" author="Stimpson" box="[281,440,578,600]" pageId="2" pageNumber="69" refString="Stimpson, W. (1859) Notes on North American Crustacea. [Preprint from] Annals of the Lyceum of Natural History in New York, 7 [1862], 49 - 93 [preprint pages 3 - 47.]" type="journal article" year="1859">Stimpson, 1859</bibRefCitation>
</taxonomicName>
[
<emphasis id="B96DF213FF9BFFD29193FDA50DF06151" box="[454,471,579,600]" italics="true" pageId="2" pageNumber="69">=</emphasis>
<taxonomicName id="4C195582FF9BFFD29188FDA50ED96151" ID-CoL="05926aea-fb00-4a41-aa3f-4d6a2e458bd3" authority="Stimpson, 1858" authorityName="Stimpson" authorityYear="1858" box="[477,766,578,600]" class="Malacostraca" family="Diogenidae" genus="Paguristes" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="69" phylum="Arthropoda" rank="species" species="depressus">
<emphasis id="B96DF213FF9BFFD29188FDA50E7E6151" box="[477,601,578,600]" italics="true" pageId="2" pageNumber="69">P. depressus</emphasis>
<bibRefCitation id="EF8853F0FF9BFFD2920AFDA40ED96151" author="Stimpson" box="[607,766,578,600]" pageId="2" pageNumber="69" refString="Stimpson, W. (1858) Prodromus descriptionis animalium evertebratorum, quae in expeditione ad oceanum Pacificum septentrionalem, a Republica Federate missa, Cadwaldaro Ringgold et Johanne Rodgers ducibus, obseravit et descripsit. VII. [Preprint (December 1858) from] Proceedings of the Academy of Natural Sciencs of Philadelphia, 1858, 225 - 252." type="book chapter" year="1858">Stimpson, 1858</bibRefCitation>
</taxonomicName>
(nomen nudum)]
<taxonomicName id="4C195582FF9BFFD290C2FD840D8A617E" ID-CoL="93271b34-b1f3-4fc0-bf53-5e8112bb9511" authority="Holthuis, 1959" authorityName="Holthuis" authorityYear="1959" box="[151,429,609,631]" class="Malacostraca" family="Diogenidae" genus="Paguristes" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="69" phylum="Arthropoda" rank="species" species="erythrops">
<emphasis id="B96DF213FF9BFFD290C2FD840D37617E" box="[151,272,609,631]" italics="true" pageId="2" pageNumber="69">P. erythrops</emphasis>
<bibRefCitation id="EF8853F0FF9BFFD29143FD870D8A617E" author="Holthuis" box="[278,429,609,631]" pageId="2" pageNumber="69" refString="Holthuis, L. B. (1959) The Crustacea Decapoda of Suriname (Dutch Guiana). Zoologische Verhandelingen, 44, 1 - 296." type="journal article" year="1959">Holthuis, 1959</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="8BA62E01FF9BFFD290C2FD670DA2619C" blockId="2.[151,1436,331,1822]" box="[151,389,640,662]" pageId="2" pageNumber="69">
<emphasis id="B96DF213FF9BFFD290C2FD670CC5619F" box="[151,226,641,662]" italics="true" pageId="2" pageNumber="69">P. g r a y i</emphasis>
<bibRefCitation id="EF8853F0FF9BFFD290BEFD660DA2619C" author="Benedict" box="[235,389,640,662]" pageId="2" pageNumber="69" refString="Benedict, J. E. (1901) The anomuran collections, made by the Fish Hawk Expedition to Porto Rico. Bulletin of the United States Fish Commission, (1900), 20, 131 - 148." type="journal article" year="1901">Benedict, 1901</bibRefCitation>
</paragraph>
<paragraph id="8BA62E01FF9BFFD290C2FD460E4E6019" blockId="2.[151,1436,331,1822]" pageId="2" pageNumber="69">
<taxonomicName id="4C195582FF9BFFD290C2FD460EBE61BD" ID-CoL="ffcde54e-1b02-4508-a3f6-2e0cf5ae206b" authority="McLaughlin &amp; Provenzano, 1974" authorityName="McLaughlin &amp; Provenzano" authorityYear="1974" box="[151,665,670,693]" class="Malacostraca" family="Diogenidae" genus="Paguristes" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="69" phylum="Arthropoda" rank="species" species="hernancortezi">
<emphasis id="B96DF213FF9BFFD290C2FD460D1B61BC" box="[151,316,671,693]" italics="true" pageId="2" pageNumber="69">P. hernancortezi</emphasis>
<bibRefCitation id="EF8853F0FF9BFFD29116FD780EBE61BD" author="McLaughlin" box="[323,665,670,692]" pageId="2" pageNumber="69" refString="McLaughlin, P. A. &amp; Provenzano, A. J., Jr. (1974) The genus Paguristes (Crustacea: Decapoda: Diogenidae) from the western Atlantic. Part I. The Paguristes tortugae complex, with notes on variation. Bulletin of Marine Science, 24 (1), 165 - 234, figs 1 - 27." type="journal article" year="1974">McLaughlin &amp; Provenzano, 1974</bibRefCitation>
</taxonomicName>
*
<taxonomicName id="4C195582FF9BFFD290C2FD580F4861DB" ID-CoL="00ab44f6-97b2-4760-b58b-a6f64e4f9c9b" authority="Manjon-Cabeza, Garcia Raso &amp; Martinez Iglesias, 2002" authorityName="Manjon-Cabeza, Garcia Raso &amp; Martinez Iglesias" authorityYear="2002" box="[151,879,701,723]" class="Malacostraca" family="Diogenidae" genus="Paguristes" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="69" phylum="Arthropoda" rank="species" species="holguinensis">
<emphasis id="B96DF213FF9BFFD290C2FD580D1761DA" box="[151,304,701,723]" italics="true" pageId="2" pageNumber="69">P. holguinensis</emphasis>
Manjón-Cabeza, García Raso &amp; Martínez Iglesias, 2002
</taxonomicName>
<taxonomicName id="4C195582FF9BFFD290C2FD3B0E5061F8" ID-CoL="5c37d3a0-9b04-457e-9e8f-4b28a2d4cc1b" authority="McLaughlin &amp; Provenzano, 1975" authorityName="McLaughlin &amp; Provenzano" authorityYear="1975" box="[151,631,731,754]" class="Malacostraca" family="Diogenidae" genus="Paguristes" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="69" phylum="Arthropoda" rank="species" species="inconstans">
<emphasis id="B96DF213FF9BFFD290C2FD3B0D3B61FB" box="[151,284,733,754]" italics="true" pageId="2" pageNumber="69">P. inconstans</emphasis>
<bibRefCitation id="EF8853F0FF9BFFD29177FD3D0E5061F8" author="McLaughlin" box="[290,631,731,754]" pageId="2" pageNumber="69" refString="McLaughlin, P. A. &amp; Provenzano, A. J., Jr. (1975) The genus Paguristes (Crustacea: Decapoda: Diogenidae) from the western Atlantic. Part II. Six new species. Bulletin of Marine Science, 24 (4) [1974], 885 - 938, figs 1 - 18." type="journal article" year="1975">McLaughlin &amp; Provenzano, 1975</bibRefCitation>
</taxonomicName>
<taxonomicName id="4C195582FF9BFFD290C2FD1D0E4E6019" ID-CoL="a0ef8c8a-d979-4cab-8c5a-1fbad8af4411" authority="McLaughlin &amp; Provenzano, 1975" authorityName="McLaughlin &amp; Provenzano" authorityYear="1975" box="[151,617,762,784]" class="Malacostraca" family="Diogenidae" genus="Paguristes" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="69" phylum="Arthropoda" rank="species" species="lapillatus">
<emphasis id="B96DF213FF9BFFD290C2FD1D0D376019" box="[151,272,762,784]" italics="true" pageId="2" pageNumber="69">P. lapillatus</emphasis>
<bibRefCitation id="EF8853F0FF9BFFD29140FD1C0E4E6019" author="McLaughlin" box="[277,617,762,784]" pageId="2" pageNumber="69" refString="McLaughlin, P. A. &amp; Provenzano, A. J., Jr. (1975) The genus Paguristes (Crustacea: Decapoda: Diogenidae) from the western Atlantic. Part II. Six new species. Bulletin of Marine Science, 24 (4) [1974], 885 - 938, figs 1 - 18." type="journal article" year="1975">McLaughlin &amp; Provenzano, 1975</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="8BA62E01FF9BFFD290C2FCFC0E4A6026" blockId="2.[151,1436,331,1822]" box="[151,621,793,815]" pageId="2" pageNumber="69">
<taxonomicName id="4C195582FF9BFFD290C2FCFC0E4A6026" ID-CoL="e8aa674b-8d66-452d-a0be-a328b664db0a" authority="McLaughlin &amp; Provenzano, 1975" authorityName="McLaughlin &amp; Provenzano" authorityYear="1975" box="[151,621,793,815]" class="Malacostraca" family="Diogenidae" genus="Paguristes" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="69" phylum="Arthropoda" rank="species" species="laticlavus">
<emphasis id="B96DF213FF9BFFD290C2FCFC0D356026" box="[151,274,793,815]" italics="true" pageId="2" pageNumber="69">P. laticlavus</emphasis>
<bibRefCitation id="EF8853F0FF9BFFD2914DFCFF0E4A6026" author="McLaughlin" box="[280,621,793,815]" pageId="2" pageNumber="69" refString="McLaughlin, P. A. &amp; Provenzano, A. J., Jr. (1975) The genus Paguristes (Crustacea: Decapoda: Diogenidae) from the western Atlantic. Part II. Six new species. Bulletin of Marine Science, 24 (4) [1974], 885 - 938, figs 1 - 18." type="journal article" year="1975">McLaughlin &amp; Provenzano, 1975</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="8BA62E01FF9BFFD290C2FCDF0E4F6065" blockId="2.[151,1436,331,1822]" pageId="2" pageNumber="69">
<taxonomicName id="4C195582FF9BFFD290C2FCDF0E506047" ID-CoL="627136e3-ef21-4f35-b869-1151fa4b59e0" authority="McLaughlin &amp; Provenzano, 1975" authorityName="McLaughlin &amp; Provenzano" authorityYear="1975" box="[151,631,824,846]" class="Malacostraca" family="Diogenidae" genus="Paguristes" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="69" phylum="Arthropoda" rank="species" species="limonensis">
<emphasis id="B96DF213FF9BFFD290C2FCDF0D3B6047" box="[151,284,824,846]" italics="true" pageId="2" pageNumber="69">P. limonensis</emphasis>
<bibRefCitation id="EF8853F0FF9BFFD29177FCDE0E506047" author="McLaughlin" box="[290,631,824,846]" pageId="2" pageNumber="69" refString="McLaughlin, P. A. &amp; Provenzano, A. J., Jr. (1975) The genus Paguristes (Crustacea: Decapoda: Diogenidae) from the western Atlantic. Part II. Six new species. Bulletin of Marine Science, 24 (4) [1974], 885 - 938, figs 1 - 18." type="journal article" year="1975">McLaughlin &amp; Provenzano, 1975</bibRefCitation>
</taxonomicName>
<emphasis id="B96DF213FF9BFFD290C2FCBE0CD66064" box="[151,241,855,877]" italics="true" pageId="2" pageNumber="69">P. l y m a n i</emphasis>
<bibRefCitation id="EF8853F0FF9BFFD290AEFCB10E4F6065" author="Milne-Edwards" box="[251,616,854,876]" pageId="2" pageNumber="69" refString="Milne-Edwards, A. &amp; Bouvier, E. - L. (1893) Reports on the results of dredging, under the supervision of Alexander Agassiz, in the Gulf of Mexico (1877 - 78), in the Caribbean Sea (1878 - 79), and along the Atlantic coast of the United States (1880), by the U. S. Coast Survey Steamer &quot; Blake &quot;, Lieut. - Commander C. D Sigsbee, U. S. N., and Commander J. R. Bartlett, U. S. N., commanding. XXXIII. Description des Crustaces de la famille des paguriens recueillis pendant l'expedition. Memoirs of the Museum of Comparative Zoology, Harvard College, 14 (3), 5 - 172." type="journal article" year="1893">A. Milne-Edwards &amp; Bouvier, 1893</bibRefCitation>
</paragraph>
<paragraph id="8BA62E01FF9BFFD290C2FC900D9C60A0" blockId="2.[151,1436,331,1822]" pageId="2" pageNumber="69">
<taxonomicName id="4C195582FF9BFFD290C2FC900E8E6083" ID-CoL="ee9a18d5-a08b-4e99-b0ee-df39f6d6a491" authority="Martinez-Iglesias &amp; Gomez, 1989" authorityName="Martinez-Iglesias &amp; Gomez" authorityYear="1989" box="[151,681,885,907]" class="Malacostraca" family="Diogenidae" genus="Paguristes" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="69" phylum="Arthropoda" rank="species" species="maclaughlinae">
<emphasis id="B96DF213FF9BFFD290C2FC900D616082" box="[151,326,885,907]" italics="true" pageId="2" pageNumber="69">P. maclaughlinae</emphasis>
<bibRefCitation id="EF8853F0FF9BFFD29119FC930E8E6083" author="Martinez-Iglesias" box="[332,681,885,907]" pageId="2" pageNumber="69" refString="Martinez-Iglesias, J. C. &amp; Gomez, O. (1989) Una especie nueva del genero Paguristes (Crustacea: Decapoda: Diogenidae) en las aguas de Cuba. Poeyana, 379, 1 - 30." type="journal article" year="1989">Martínez-Iglesias &amp; Gómez, 1989</bibRefCitation>
</taxonomicName>
*
<taxonomicName id="4C195582FF9BFFD290C2FC730D9C60A0" ID-CoL="3738a604-8e67-43ab-874f-677ffe6c1482" authority="Sandberg, 1996" authorityName="Sandberg" authorityYear="1996" box="[151,443,915,938]" class="Malacostraca" family="Diogenidae" genus="Paguristes" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="69" phylum="Arthropoda" rank="species" species="markhami">
<emphasis id="B96DF213FF9BFFD290C2FC730D3060A3" box="[151,279,916,938]" italics="true" pageId="2" pageNumber="69">P. markhami</emphasis>
<bibRefCitation id="EF8853F0FF9BFFD29149FC750D9C60A0" author="Sandberg" box="[284,443,915,937]" pageId="2" pageNumber="69" refString="Sandberg, L. (1996) Hermit crabs of the genus Paguristes (Crustacea: Decapoda: Diogenidae) from the western Atlantic. Part III. Paguristes markhami, a new species from the Bahama and Caicos Islands. Proceedings of the Biological Society of Washington, 109, 470 - 475." type="journal article" year="1996">Sandberg, 1996</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="8BA62E01FF9BFFD290C2FC550DB160C1" blockId="2.[151,1436,331,1822]" box="[151,406,946,968]" pageId="2" pageNumber="69">
<emphasis id="B96DF213FF9BFFD290C2FC550CD460C1" box="[151,243,947,968]" italics="true" pageId="2" pageNumber="69">P. m o o re i</emphasis>
<bibRefCitation id="EF8853F0FF9BFFD290A8FC540DB160C1" author="Benedict" box="[253,406,946,968]" pageId="2" pageNumber="69" refString="Benedict, J. E. (1901) The anomuran collections, made by the Fish Hawk Expedition to Porto Rico. Bulletin of the United States Fish Commission, (1900), 20, 131 - 148." type="journal article" year="1901">Benedict, 1901</bibRefCitation>
</paragraph>
<paragraph id="8BA62E01FF9BFFD290C2FC340DC260EE" blockId="2.[151,1436,331,1822]" box="[151,485,977,999]" pageId="2" pageNumber="69">
<taxonomicName id="4C195582FF9BFFD290C2FC340DC260EE" ID-CoL="a642d01e-e735-4c4a-9096-421542ff241e" authority="Holthuis, 1959" authorityName="Holthuis" authorityYear="1959" box="[151,485,977,999]" class="Malacostraca" family="Diogenidae" genus="Paguristes" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="69" phylum="Arthropoda" rank="species" species="oxyophthalmus">
<emphasis id="B96DF213FF9BFFD290C2FC340D6E60EE" box="[151,329,977,999]" italics="true" pageId="2" pageNumber="69">P. oxyophthalmus</emphasis>
<bibRefCitation id="EF8853F0FF9BFFD2911AFC370DC260EE" author="Holthuis" box="[335,485,977,999]" pageId="2" pageNumber="69" refString="Holthuis, L. B. (1959) The Crustacea Decapoda of Suriname (Dutch Guiana). Zoologische Verhandelingen, 44, 1 - 296." type="journal article" year="1959">Holthuis, 1959</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="8BA62E01FF9BFFD290C2FC170E4A672D" blockId="2.[151,1436,331,1822]" pageId="2" pageNumber="69">
<taxonomicName id="4C195582FF9BFFD290C2FC170EB8670F" ID-CoL="3361fb72-8c79-4d46-994b-a82e0f091092" authority="McLaughlin &amp; Provenzano, 1975" authorityName="McLaughlin &amp; Provenzano" authorityYear="1975" box="[151,671,1008,1030]" class="Malacostraca" family="Diogenidae" genus="Paguristes" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="69" phylum="Arthropoda" rank="species" species="paraguanensis">
<emphasis id="B96DF213FF9BFFD290C2FC170D62670F" box="[151,325,1009,1030]" italics="true" pageId="2" pageNumber="69">P. paraguanensis</emphasis>
<bibRefCitation id="EF8853F0FF9BFFD2911EFC160EB8670F" author="McLaughlin" box="[331,671,1008,1030]" pageId="2" pageNumber="69" refString="McLaughlin, P. A. &amp; Provenzano, A. J., Jr. (1975) The genus Paguristes (Crustacea: Decapoda: Diogenidae) from the western Atlantic. Part II. Six new species. Bulletin of Marine Science, 24 (4) [1974], 885 - 938, figs 1 - 18." type="journal article" year="1975">McLaughlin &amp; Provenzano, 1975</bibRefCitation>
</taxonomicName>
<taxonomicName id="4C195582FF9BFFD290C2FBF60D40672D" ID-CoL="a7135fe0-2366-4be7-ade3-101e0bcc79e6" authority="Forest" authorityName="Forest" box="[151,359,1039,1061]" class="Malacostraca" family="Diogenidae" genus="Paguristes" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="69" phylum="Arthropoda" rank="species" species="pauciparus">
<emphasis id="B96DF213FF9BFFD290C2FBF60D05672C" box="[151,290,1040,1061]" italics="true" pageId="2" pageNumber="69">P. pauciparus</emphasis>
Forest
</taxonomicName>
&amp; de
<bibRefCitation id="EF8853F0FF9BFFD291F3FBE80E4A672D" author="Saint" box="[422,621,1038,1060]" pageId="2" pageNumber="69" refString="Forest, J. &amp; de Saint Laurent, M. (1968) Resultats scientifiques des campagnes de la &quot; Calypso, Part VII. Campagne de la Calypso au large des cotes Atlantiques de l'Amerique du Sud (1961 - 1962). 6. Crustaces Dcapodes: Pagurides. Annales de l'Institut Oceanographique de Monaco, n. s., 45 (2), 45 - 172." type="journal article" year="1968">Saint Laurent, 1968</bibRefCitation>
</paragraph>
<paragraph id="8BA62E01FF9BFFD290C2FBC80D9C6789" blockId="2.[151,1436,331,1822]" pageId="2" pageNumber="69">
<taxonomicName id="4C195582FF9BFFD290C2FBC80E48674B" ID-CoL="72616444-bc8a-4f10-a9a4-eefd7161d7ba" authority="McLaughlin &amp; Provenzano, 1974" authorityName="McLaughlin &amp; Provenzano" authorityYear="1974" box="[151,623,1069,1091]" class="Malacostraca" family="Diogenidae" genus="Paguristes" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="69" phylum="Arthropoda" rank="species" species="perplexus">
<emphasis id="B96DF213FF9BFFD290C2FBC80D35674A" box="[151,274,1069,1091]" italics="true" pageId="2" pageNumber="69">P. perplexus</emphasis>
<bibRefCitation id="EF8853F0FF9BFFD2914DFBCB0E48674B" author="McLaughlin" box="[280,623,1069,1091]" pageId="2" pageNumber="69" refString="McLaughlin, P. A. &amp; Provenzano, A. J., Jr. (1974) The genus Paguristes (Crustacea: Decapoda: Diogenidae) from the western Atlantic. Part I. The Paguristes tortugae complex, with notes on variation. Bulletin of Marine Science, 24 (1), 165 - 234, figs 1 - 27." type="journal article" year="1974">McLaughlin &amp; Provenzano, 1974</bibRefCitation>
</taxonomicName>
*
<taxonomicName id="4C195582FF9BFFD290C2FBAB0E5D6768" ID-CoL="4918f813-6247-48c7-9289-fabb69cf2e4c" authority="A. Milne-Edwards &amp; Bouvier, 1893" authorityName="A. Milne-Edwards &amp; Bouvier" authorityYear="1893" box="[151,634,1099,1122]" class="Malacostraca" family="Diogenidae" genus="Paguristes" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="69" phylum="Arthropoda" rank="species" species="planatus">
<emphasis id="B96DF213FF9BFFD290C2FBAB0D20676B" box="[151,263,1100,1122]" italics="true" pageId="2" pageNumber="69">P. planatus</emphasis>
<bibRefCitation id="EF8853F0FF9BFFD29158FBAA0E5D6768" author="Milne-Edwards" box="[269,634,1099,1121]" pageId="2" pageNumber="69" refString="Milne-Edwards, A. &amp; Bouvier, E. - L. (1893) Reports on the results of dredging, under the supervision of Alexander Agassiz, in the Gulf of Mexico (1877 - 78), in the Caribbean Sea (1878 - 79), and along the Atlantic coast of the United States (1880), by the U. S. Coast Survey Steamer &quot; Blake &quot;, Lieut. - Commander C. D Sigsbee, U. S. N., and Commander J. R. Bartlett, U. S. N., commanding. XXXIII. Description des Crustaces de la famille des paguriens recueillis pendant l'expedition. Memoirs of the Museum of Comparative Zoology, Harvard College, 14 (3), 5 - 172." type="journal article" year="1893">A. Milne-Edwards &amp; Bouvier, 1893</bibRefCitation>
</taxonomicName>
<taxonomicName id="4C195582FF9BFFD290C2FB8D0D9C6789" ID-CoL="cf5f9c93-0dae-4ed9-85df-73a54c251c86" authority="Benedict, 1901" authorityName="Benedict" authorityYear="1901" box="[151,443,1130,1152]" class="Malacostraca" family="Diogenidae" genus="Paguristes" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="69" phylum="Arthropoda" rank="species" species="puncticeps">
<emphasis id="B96DF213FF9BFFD290C2FB8D0D3B6789" box="[151,284,1131,1152]" italics="true" pageId="2" pageNumber="69">P. puncticeps</emphasis>
<bibRefCitation id="EF8853F0FF9BFFD29177FB8C0D9C6789" author="Benedict" box="[290,443,1130,1152]" pageId="2" pageNumber="69" refString="Benedict, J. E. (1901) The anomuran collections, made by the Fish Hawk Expedition to Porto Rico. Bulletin of the United States Fish Commission, (1900), 20, 131 - 148." type="journal article" year="1901">Benedict, 1901</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="8BA62E01FF9BFFD290C2FB6C0E746796" blockId="2.[151,1436,331,1822]" box="[151,595,1161,1183]" pageId="2" pageNumber="69">
<taxonomicName id="4C195582FF9BFFD290C2FB6C0D6C6796" ID-CoL="18441366-7498-4c70-a200-1ad973da50f5" authority="Forest" authorityName="Forest" box="[151,331,1161,1183]" class="Malacostraca" family="Diogenidae" genus="Paguristes" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="69" phylum="Arthropoda" rank="species" species="robustus">
<emphasis id="B96DF213FF9BFFD290C2FB6C0D216796" box="[151,262,1161,1183]" italics="true" pageId="2" pageNumber="69">P. robustus</emphasis>
Forest
</taxonomicName>
&amp; de
<bibRefCitation id="EF8853F0FF9BFFD291DFFB6F0E746796" author="Saint" box="[394,595,1161,1183]" pageId="2" pageNumber="69" refString="Forest, J. &amp; de Saint Laurent, M. (1968) Resultats scientifiques des campagnes de la &quot; Calypso, Part VII. Campagne de la Calypso au large des cotes Atlantiques de l'Amerique du Sud (1961 - 1962). 6. Crustaces Dcapodes: Pagurides. Annales de l'Institut Oceanographique de Monaco, n. s., 45 (2), 45 - 172." type="journal article" year="1968">Saint Laurent, 1968</bibRefCitation>
</paragraph>
<paragraph id="8BA62E01FF9BFFD290C2FB4F0E6B67B4" blockId="2.[151,1436,331,1822]" box="[151,588,1192,1214]" pageId="2" pageNumber="69">
<emphasis id="B96DF213FF9BFFD290C2FB4F0CF267B7" box="[151,213,1193,1214]" italics="true" pageId="2" pageNumber="69">P. s a y i</emphasis>
<bibRefCitation id="EF8853F0FF9BFFD2908AFB4E0E6B67B4" author="Milne-Edwards" box="[223,588,1192,1214]" pageId="2" pageNumber="69" refString="Milne-Edwards, A. &amp; Bouvier, E. - L. (1893) Reports on the results of dredging, under the supervision of Alexander Agassiz, in the Gulf of Mexico (1877 - 78), in the Caribbean Sea (1878 - 79), and along the Atlantic coast of the United States (1880), by the U. S. Coast Survey Steamer &quot; Blake &quot;, Lieut. - Commander C. D Sigsbee, U. S. N., and Commander J. R. Bartlett, U. S. N., commanding. XXXIII. Description des Crustaces de la famille des paguriens recueillis pendant l'expedition. Memoirs of the Museum of Comparative Zoology, Harvard College, 14 (3), 5 - 172." type="journal article" year="1893">A. Milne-Edwards &amp; Bouvier, 1893</bibRefCitation>
</paragraph>
<paragraph id="8BA62E01FF9BFFD290C2FB2E0D966631" blockId="2.[151,1436,331,1822]" pageId="2" pageNumber="69">
<taxonomicName id="4C195582FF9BFFD290C2FB2E0E2467D5" ID-CoL="fc65a844-0b83-45f6-8cf5-d20fd1c8a5af" authority="A. Milne Edwards, 1880" authorityName="A. Milne Edwards" authorityYear="1880" box="[151,515,1222,1245]" class="Malacostraca" family="Diogenidae" genus="Paguristes" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="69" phylum="Arthropoda" rank="species" species="sericeus">
<emphasis id="B96DF213FF9BFFD290C2FB2E0D2467D4" box="[151,259,1224,1245]" italics="true" pageId="2" pageNumber="69">P. sericeus</emphasis>
A. Milne
<bibRefCitation id="EF8853F0FF9BFFD2913EFB200E2467D5" author="Edwards" box="[363,515,1222,1244]" pageId="2" pageNumber="69" refString="Milne-Edwards, A. (1880) Report on the results of dredging, under the supervision of Alexander Agassiz, in the Gulf of Mexico, and in the Caribbean Sea, 1877, 78, 79, by the United States Coast Survey steamer &quot; Blake &quot;, Lieut. - Commander C. D. Sigsbee, U. S. N., and Commander J. R. Bartlett, U. S. N., commanding. VIII. Etudes preliminaires sur les Crustaces. Bulletin of the Museum of Comparative Zoology, Harvard College, 8 (1), 1 - 68." type="journal article" year="1880">Edwards, 1880</bibRefCitation>
</taxonomicName>
(=
<taxonomicName id="4C195582FF9BFFD29273FB2E0F6667D5" ID-CoL="1b65f7e6-472d-4918-be4f-c45db624a997" authority="Benedict, 1901" authorityName="Benedict" authorityYear="1901" box="[550,833,1222,1245]" class="Malacostraca" family="Diogenidae" genus="Paguristes" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="69" phylum="Arthropoda" rank="species" species="rectifrons">
<emphasis id="B96DF213FF9BFFD29273FB2E0E8767D4" box="[550,672,1223,1245]" italics="true" pageId="2" pageNumber="69">P. rectifrons</emphasis>
<bibRefCitation id="EF8853F0FF9BFFD292F0FB200F6667D5" author="Benedict" box="[677,833,1222,1244]" pageId="2" pageNumber="69" refString="Benedict, J. E. (1901) The anomuran collections, made by the Fish Hawk Expedition to Porto Rico. Bulletin of the United States Fish Commission, (1900), 20, 131 - 148." type="journal article" year="1901">Benedict, 1901</bibRefCitation>
</taxonomicName>
; =
<taxonomicName id="4C195582FF9BFFD29334FB2E08AA67D5" ID-CoL="ab3f862a-f01c-42cb-9345-65bb1f852783" authority="Benedict, 1901" authorityName="Benedict" authorityYear="1901" box="[865,1165,1222,1245]" class="Malacostraca" family="Diogenidae" genus="Paguristes" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="69" phylum="Arthropoda" rank="species" species="tenuirostris">
<emphasis id="B96DF213FF9BFFD29334FB2E0FC967D4" box="[865,1006,1224,1245]" italics="true" pageId="2" pageNumber="69">P. tenuirostris</emphasis>
<bibRefCitation id="EF8853F0FF9BFFD293A1FB2008AA67D5" author="Benedict" box="[1012,1165,1222,1244]" pageId="2" pageNumber="69" refString="Benedict, J. E. (1901) The anomuran collections, made by the Fish Hawk Expedition to Porto Rico. Bulletin of the United States Fish Commission, (1900), 20, 131 - 148." type="journal article" year="1901">Benedict, 1901</bibRefCitation>
</taxonomicName>
<taxonomicName id="4C195582FF9BFFD290C2FB000E5067F2" ID-CoL="332f6610-70c4-41ad-be40-65c6728f77f1" authority="McLaughlin &amp; Provenzano, 1975" authorityName="McLaughlin &amp; Provenzano" authorityYear="1975" box="[151,631,1253,1275]" class="Malacostraca" family="Diogenidae" genus="Paguristes" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="69" phylum="Arthropoda" rank="species" species="spectabilis">
<emphasis id="B96DF213FF9BFFD290C2FB000D3B67F2" box="[151,284,1253,1275]" italics="true" pageId="2" pageNumber="69">P. spectabilis</emphasis>
<bibRefCitation id="EF8853F0FF9BFFD29177FB030E5067F2" author="McLaughlin" box="[290,631,1253,1275]" pageId="2" pageNumber="69" refString="McLaughlin, P. A. &amp; Provenzano, A. J., Jr. (1975) The genus Paguristes (Crustacea: Decapoda: Diogenidae) from the western Atlantic. Part II. Six new species. Bulletin of Marine Science, 24 (4) [1974], 885 - 938, figs 1 - 18." type="journal article" year="1975">McLaughlin &amp; Provenzano, 1975</bibRefCitation>
</taxonomicName>
<taxonomicName id="4C195582FF9BFFD290C2FAE30E226610" ID-CoL="c4168854-1d15-4139-b87d-1bb43cacc4e7" authority="A. Milne-Edwards, 1880" authorityName="A. Milne-Edwards" authorityYear="1880" box="[151,517,1283,1306]" class="Malacostraca" family="Diogenidae" genus="Paguristes" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="69" phylum="Arthropoda" rank="species" species="spinipes">
<emphasis id="B96DF213FF9BFFD290C2FAE30D246613" box="[151,259,1285,1306]" italics="true" pageId="2" pageNumber="69">P. spinipes</emphasis>
<bibRefCitation id="EF8853F0FF9BFFD2915CFAE20E226610" author="Milne-Edwards" box="[265,517,1283,1305]" pageId="2" pageNumber="69" refString="Milne-Edwards, A. (1880) Report on the results of dredging, under the supervision of Alexander Agassiz, in the Gulf of Mexico, and in the Caribbean Sea, 1877, 78, 79, by the United States Coast Survey steamer &quot; Blake &quot;, Lieut. - Commander C. D. Sigsbee, U. S. N., and Commander J. R. Bartlett, U. S. N., commanding. VIII. Etudes preliminaires sur les Crustaces. Bulletin of the Museum of Comparative Zoology, Harvard College, 8 (1), 1 - 68." type="journal article" year="1880">A. Milne-Edwards, 1880</bibRefCitation>
</taxonomicName>
(=
<emphasis id="B96DF213FF9BFFD2927DFAE30EB66613" box="[552,657,1285,1306]" italics="true" pageId="2" pageNumber="69">P. a r m a t u s</emphasis>
<bibRefCitation id="EF8853F0FF9BFFD292CFFAE30F226610" author="Hay" box="[666,773,1284,1306]" pageId="2" pageNumber="69" refString="Hay, W. P. (1917) Preliminary descriptions of five new species of crustaceans from the coast of North Carolina. Proceedings of the Biological Society of Washington, 30, 71 - 74." type="journal article" year="1917">Hay, 1917</bibRefCitation>
; =
<taxonomicName id="4C195582FF9BFFD29371FAE308036613" ID-CoL="78c42c07-dd29-420e-9eab-91bf03f6ec0e" authority="Henderson, 1888" authorityName="Henderson" authorityYear="1888" box="[804,1060,1283,1306]" class="Malacostraca" family="Diogenidae" genus="Paguristes" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="69" phylum="Arthropoda" rank="species" species="visor">
<emphasis id="B96DF213FF9BFFD29371FAE30F576613" box="[804,880,1285,1306]" italics="true" pageId="2" pageNumber="69">P. visor</emphasis>
<bibRefCitation id="EF8853F0FF9BFFD29323FAE508036613" author="Henderson" box="[886,1060,1283,1306]" pageId="2" pageNumber="69" refString="Henderson, J. R. (1888) Report on the Anomura collected by H. M. S. Challenger during the years 1873 - 76. Scientific Results of the Exploratory Voyage of HMS Challenger, (Zoology) 27, i - xi, 1 - 221. Edinburgh etc., Her Majesty's Stationary Office." type="journal article" year="1888">Henderson, 1888</bibRefCitation>
</taxonomicName>
)
<taxonomicName id="4C195582FF9BFFD290C2FAC50D966631" ID-CoL="8a47b0b8-dadb-448c-89c0-302d0156aa1f" authority="Provenzano, 1965" authorityName="Provenzano" authorityYear="1965" box="[151,433,1314,1336]" class="Malacostraca" family="Diogenidae" genus="Paguristes" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="69" phylum="Arthropoda" rank="species" species="starcki">
<emphasis id="B96DF213FF9BFFD290C2FAC50CD36631" box="[151,244,1314,1336]" italics="true" pageId="2" pageNumber="69">P. starcki</emphasis>
<bibRefCitation id="EF8853F0FF9BFFD290AFFAC50D966631" author="Provenzano" box="[250,433,1314,1336]" pageId="2" pageNumber="69" refString="Provenzano, A. J., Jr. (1965) Two new West Indian hermit crabs of the genus Paguristes (Crustacea: Diogenidae). Bulletin of Marine Science, 15 (3), 726 - 736." type="journal article" year="1965">Provenzano, 1965</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="8BA62E01FF9BFFD290C2FAA40D80665E" blockId="2.[151,1436,331,1822]" box="[151,423,1345,1367]" pageId="2" pageNumber="69">
<taxonomicName id="4C195582FF9BFFD290C2FAA40DBB665E" ID-CoL="29918680-722e-4ce7-b2eb-4680b5673468" authority="Schmitt, 1933" authorityName="Schmitt" authorityYear="1933" box="[151,412,1345,1367]" class="Malacostraca" family="Diogenidae" genus="Paguristes" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="69" phylum="Arthropoda" rank="species" species="tortugae">
<emphasis id="B96DF213FF9BFFD290C2FAA40D21665E" box="[151,262,1346,1367]" italics="true" pageId="2" pageNumber="69">P. tortugae</emphasis>
<bibRefCitation id="EF8853F0FF9BFFD29159FAA70DBB665E" author="Schmitt" box="[268,412,1345,1367]" pageId="2" pageNumber="69" refString="Schmitt, W. L. (1933) Four new species of decapod crustaceans from Porto Rico. American Museum Novitates, 662, 1 - 9." type="journal article" year="1933">Schmitt, 1933</bibRefCitation>
</taxonomicName>
*
</paragraph>
<paragraph id="8BA62E01FF9BFFD290C2FA870D8266BA" blockId="2.[151,1436,331,1822]" pageId="2" pageNumber="69">
<taxonomicName id="4C195582FF9BFFD290C2FA870D5C667F" ID-CoL="427713b2-887f-45ea-9abe-b4c1d945dee2" authority="Forest" authorityName="Forest" box="[151,379,1376,1398]" class="Malacostraca" family="Diogenidae" genus="Paguristes" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="69" phylum="Arthropoda" rank="species" species="triangulopsis">
<emphasis id="B96DF213FF9BFFD290C2FA870D12667F" box="[151,309,1376,1398]" italics="true" pageId="2" pageNumber="69">P. triangulopsis</emphasis>
Forest
</taxonomicName>
&amp; de
<bibRefCitation id="EF8853F0FF9BFFD291ECFA860EA5667F" author="Saint" box="[441,642,1376,1398]" pageId="2" pageNumber="69" refString="Forest, J. &amp; de Saint Laurent, M. (1968) Resultats scientifiques des campagnes de la &quot; Calypso, Part VII. Campagne de la Calypso au large des cotes Atlantiques de l'Amerique du Sud (1961 - 1962). 6. Crustaces Dcapodes: Pagurides. Annales de l'Institut Oceanographique de Monaco, n. s., 45 (2), 45 - 172." type="journal article" year="1968">Saint Laurent, 1968</bibRefCitation>
<taxonomicName id="4C195582FF9BFFD290C2FA660EB8669D" ID-CoL="4d55e0bb-7150-48b4-b936-d8897c98313f" authority="A. Milne-Edwards &amp; Bouvier, 1893" authorityName="A. Milne-Edwards &amp; Bouvier" authorityYear="1893" box="[151,671,1406,1429]" class="Malacostraca" family="Diogenidae" genus="Paguristes" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="69" phylum="Arthropoda" rank="species" species="triangulatus">
<emphasis id="B96DF213FF9BFFD290C2FA660D0B669C" box="[151,300,1407,1429]" italics="true" pageId="2" pageNumber="69">P. triangulatus</emphasis>
<bibRefCitation id="EF8853F0FF9BFFD29167FA990EB8669D" author="Milne-Edwards" box="[306,671,1406,1428]" pageId="2" pageNumber="69" refString="Milne-Edwards, A. &amp; Bouvier, E. - L. (1893) Reports on the results of dredging, under the supervision of Alexander Agassiz, in the Gulf of Mexico (1877 - 78), in the Caribbean Sea (1878 - 79), and along the Atlantic coast of the United States (1880), by the U. S. Coast Survey Steamer &quot; Blake &quot;, Lieut. - Commander C. D Sigsbee, U. S. N., and Commander J. R. Bartlett, U. S. N., commanding. XXXIII. Description des Crustaces de la famille des paguriens recueillis pendant l'expedition. Memoirs of the Museum of Comparative Zoology, Harvard College, 14 (3), 5 - 172." type="journal article" year="1893">A. Milne-Edwards &amp; Bouvier, 1893</bibRefCitation>
</taxonomicName>
<taxonomicName id="4C195582FF9BFFD290C2FA780D8266BA" ID-CoL="e175f004-18a8-4c81-822c-9917956c86ac" authority="Provenzano, 1961" authorityName="Provenzano" authorityYear="1961" box="[151,421,1437,1459]" class="Malacostraca" family="Diogenidae" genus="Paguristes" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="69" phylum="Arthropoda" rank="species" species="wassi">
<emphasis id="B96DF213FF9BFFD290C2FA780CCF66BA" box="[151,232,1438,1459]" italics="true" pageId="2" pageNumber="69">P. wassi</emphasis>
<bibRefCitation id="EF8853F0FF9BFFD290BBFA780D8266BA" author="Provenzano" box="[238,421,1437,1459]" pageId="2" pageNumber="69" refString="Provenzano, A. J., Jr. (1961) Pagurid crabs (Decapoda Anomura) from St. John, Virgin Islands, with descriptions of three new species. Crustaceana, 3 (2), 151 - 166." type="journal article" year="1961">Provenzano, 1961</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="8BA62E01FF9BFFD290C2FA5B0E0466D8" blockId="2.[151,1436,331,1822]" box="[151,547,1467,1490]" pageId="2" pageNumber="69">
<emphasis id="B96DF213FF9BFFD290C2FA5B0D2166DB" box="[151,262,1468,1490]" italics="true" pageId="2" pageNumber="69">P. w e rd i n g i</emphasis>
<bibRefCitation id="EF8853F0FF9BFFD2915AFA5A0E3F66D8" author="Campos" box="[271,536,1467,1489]" pageId="2" pageNumber="69" refString="Campos, N. H. &amp; Sanchez, H. (1995) Los cangrejos ermitanos del genero Paguristes Dana (Anomura: Diogenidae) de la costa norte colombiana, con la descripcion de dos nuevas especies. Caldasia, 17 (82 - 85), 569 - 586." type="journal article" year="1995">Campos &amp; Sánchez, 1995</bibRefCitation>
*
</paragraph>
<paragraph id="8BA62E01FF9BFFD290C2FA3D0E2666F9" blockId="2.[151,1436,331,1822]" box="[151,513,1498,1520]" pageId="2" pageNumber="69">
<emphasis id="B96DF213FF9BFFD290C2FA3D0CC366F9" box="[151,228,1498,1520]" italics="true" pageId="2" pageNumber="69">P. z e b r a</emphasis>
<bibRefCitation id="EF8853F0FF9BFFD290BBFA3D0DDF66F9" author="Campos" box="[238,504,1498,1520]" pageId="2" pageNumber="69" refString="Campos, N. H. &amp; Sanchez, H. (1995) Los cangrejos ermitanos del genero Paguristes Dana (Anomura: Diogenidae) de la costa norte colombiana, con la descripcion de dos nuevas especies. Caldasia, 17 (82 - 85), 569 - 586." type="journal article" year="1995">Campos &amp; Sánchez, 1995</bibRefCitation>
*
</paragraph>
<paragraph id="8BA62E01FF9BFFD290C2F9E10E426514" blockId="2.[151,1436,331,1822]" box="[151,613,1543,1565]" pageId="2" pageNumber="69">
<taxonomicName id="4C195582FF9BFFD290C2F9E10E426514" ID-CoL="ff241479-0d4e-4044-a78a-9a85e5094e50" authority="Rahayu &amp; McLaughlin, 2010" authorityName="Rahayu &amp; McLaughlin" authorityYear="2010" box="[151,613,1543,1565]" class="Malacostraca" family="Diogenidae" genus="Areopaguristes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="69" phylum="Arthropoda" rank="genus">
<emphasis id="B96DF213FF9BFFD290C2F9E10D126514" bold="true" box="[151,309,1543,1565]" italics="true" pageId="2" pageNumber="69">Areopaguristes</emphasis>
Rahayu &amp; McLaughlin, 2010
</taxonomicName>
</paragraph>
<paragraph id="8BA62E01FF9BFFD290C2F9D20DB16540" blockId="2.[151,1436,331,1822]" box="[151,406,1587,1610]" pageId="2" pageNumber="69">
<taxonomicName id="4C195582FF9BFFD290C2F9D20DAD6540" ID-CoL="b9b5bf7d-ec90-4b27-aae4-c603782ff318" authority="Wass, 1963" authorityName="Wass" authorityYear="1963" box="[151,394,1587,1610]" class="Malacostraca" family="Diogenidae" genus="Areopaguristes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="69" phylum="Arthropoda" rank="species" species="hewatti">
<emphasis id="B96DF213FF9BFFD290C2F9D20CD96543" box="[151,254,1588,1610]" italics="true" pageId="2" pageNumber="69">A. hewatti</emphasis>
(
<bibRefCitation id="EF8853F0FF9BFFD2915EF9D20DA46540" author="Wass" box="[267,387,1587,1609]" pageId="2" pageNumber="69" refString="Wass, M. L. (1963) New species of hermit crabs (Decapoda, Paguridae) from the western Atlantic. Crustaceana, 6, 133 - 157." type="journal article" year="1963">Wass, 1963</bibRefCitation>
)
</taxonomicName>
*
</paragraph>
<paragraph id="8BA62E01FF9BFFD290C2F9B50DA46561" blockId="2.[151,1436,331,1822]" box="[151,387,1618,1640]" pageId="2" pageNumber="69">
<taxonomicName id="4C195582FF9BFFD290C2F9B50DA46561" ID-CoL="99513801-460e-454a-b4c6-aca4a1da614d" authority="Wass, 1955" authorityName="Wass" authorityYear="1955" box="[151,387,1618,1640]" class="Malacostraca" family="Diogenidae" genus="Areopaguristes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="69" phylum="Arthropoda" rank="species" species="hummi">
<emphasis id="B96DF213FF9BFFD290C2F9B50CDF6561" box="[151,248,1618,1640]" italics="true" pageId="2" pageNumber="69">A. hummi</emphasis>
(
<bibRefCitation id="EF8853F0FF9BFFD29150F9B50D5B6561" author="Wass" box="[261,380,1618,1640]" pageId="2" pageNumber="69" refString="Wass, M. L. (1955) The decapod crustaceans of Alligator Harbor and adjacent inshore areas of northwestern Florida. The Quarterly Journal of the Florida Academy of Sciences, 18 (3), 129 - 176." type="journal article" year="1955">Wass, 1955</bibRefCitation>
)
</taxonomicName>
</paragraph>
<paragraph id="8BA62E01FF9BFFD290C2F9940E15658E" blockId="2.[151,1436,331,1822]" box="[151,562,1649,1671]" pageId="2" pageNumber="69">
<taxonomicName id="4C195582FF9BFFD290C2F9940CF2658E" ID-CoL="b06ce98a-edb0-4b9d-a685-0e0b7f109dc2" box="[151,213,1650,1671]" class="Malacostraca" family="Diogenidae" genus="Areopaguristes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="69" phylum="Arthropoda" rank="species" species="iris">
<emphasis id="B96DF213FF9BFFD290C2F9940CF2658E" box="[151,213,1650,1671]" italics="true" pageId="2" pageNumber="69">A. iris</emphasis>
</taxonomicName>
(Forest &amp; de
<bibRefCitation id="EF8853F0FF9BFFD29134F9970E0C658E" author="Saint" box="[353,555,1649,1671]" pageId="2" pageNumber="69" refString="Forest, J. &amp; de Saint Laurent, M. (1968) Resultats scientifiques des campagnes de la &quot; Calypso, Part VII. Campagne de la Calypso au large des cotes Atlantiques de l'Amerique du Sud (1961 - 1962). 6. Crustaces Dcapodes: Pagurides. Annales de l'Institut Oceanographique de Monaco, n. s., 45 (2), 45 - 172." type="journal article" year="1968">Saint Laurent, 1968</bibRefCitation>
)
</paragraph>
<paragraph id="8BA62E01FF9BFFD290C2F9770D0865AF" blockId="2.[151,1436,331,1822]" box="[151,303,1680,1702]" pageId="2" pageNumber="69">
<taxonomicName id="4C195582FF9BFFD290C2F9770CD465AF" ID-CoL="8c9c2161-2d9b-4ce2-a375-99a2a1f1a39b" box="[151,243,1680,1702]" class="Malacostraca" family="Diogenidae" genus="Areopaguristes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="69" phylum="Arthropoda" rank="species" species="tudgei" status="sp. nov.">
<emphasis id="B96DF213FF9BFFD290C2F9770CD465AF" box="[151,243,1680,1702]" italics="true" pageId="2" pageNumber="69">A. tudgei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25E4F68FF9BFFD290ACF9770D0865AF" box="[249,303,1681,1702]" pageId="2" pageNumber="69" rank="species">n. sp.</taxonomicNameLabel>
</paragraph>
<paragraph id="8BA62E01FF9BFFD290C2F95A0E3665DB" blockId="2.[151,1436,331,1822]" box="[151,529,1724,1746]" pageId="2" pageNumber="69">
<taxonomicName id="4C195582FF9BFFD290C2F95A0E3665DB" ID-CoL="b5d7825c-c5ac-4582-9ce9-683adc498c35" authority="McLaughlin, 2002" authorityName="McLaughlin" authorityYear="2002" box="[151,529,1724,1746]" class="Malacostraca" family="Diogenidae" genus="Pseudopaguristes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="69" phylum="Arthropoda" rank="genus">
<emphasis id="B96DF213FF9BFFD290C2F95A0D6965DB" bold="true" box="[151,334,1724,1746]" italics="true" pageId="2" pageNumber="69">Pseudopaguristes</emphasis>
<bibRefCitation id="EF8853F0FF9BFFD29101F95A0E3665DB" author="McLaughlin" box="[340,529,1724,1746]" pageId="2" pageNumber="69" refString="McLaughlin, P. A. (2002) Pseudopaguristes, a new and aberrant genus of hermit crabs (Anomura: Paguridea: Diogenidae). Micronesica, 34 (2), 185 - 199." type="journal article" year="2002">McLaughlin, 2002</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="8BA62E01FF9BFFD290C2F90C0E4965F6" blockId="2.[151,1436,331,1822]" box="[151,622,1769,1791]" pageId="2" pageNumber="69">
<taxonomicName id="4C195582FF9BFFD290C2F90C0D3665F6" ID-CoL="c22f537d-694c-49fe-bff3-168e44de9edf" box="[151,273,1769,1791]" class="Malacostraca" family="Diogenidae" genus="Pseudopaguristes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="69" phylum="Arthropoda" rank="species" species="calliopsis">
<emphasis id="B96DF213FF9BFFD290C2F90C0D3665F6" box="[151,273,1769,1791]" italics="true" pageId="2" pageNumber="69">P. calliopsis</emphasis>
</taxonomicName>
(Forest &amp; de
<bibRefCitation id="EF8853F0FF9BFFD291C8F90F0E4065F6" author="Saint" box="[413,615,1769,1791]" pageId="2" pageNumber="69" refString="Forest, J. &amp; de Saint Laurent, M. (1968) Resultats scientifiques des campagnes de la &quot; Calypso, Part VII. Campagne de la Calypso au large des cotes Atlantiques de l'Amerique du Sud (1961 - 1962). 6. Crustaces Dcapodes: Pagurides. Annales de l'Institut Oceanographique de Monaco, n. s., 45 (2), 45 - 172." type="journal article" year="1968">Saint Laurent, 1968</bibRefCitation>
)
</paragraph>
<paragraph id="8BA62E01FF9BFFD290C2F8EF0E926417" blockId="2.[151,1436,331,1822]" box="[151,693,1800,1822]" pageId="2" pageNumber="69">
<taxonomicName id="4C195582FF9BFFD290C2F8EF0E8C6417" ID-CoL="393cf6d0-1336-40c9-88c2-5c32d3ba6a79" authority="McLaughlin &amp; Provenzano, 1974" authorityName="McLaughlin &amp; Provenzano" authorityYear="1974" box="[151,683,1800,1822]" class="Malacostraca" family="Diogenidae" genus="Pseudopaguristes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="69" phylum="Arthropoda" rank="species" species="invisisacculus">
<emphasis id="B96DF213FF9BFFD290C2F8EF0D196417" box="[151,318,1800,1822]" italics="true" pageId="2" pageNumber="69">P. invisisacculus</emphasis>
(
<bibRefCitation id="EF8853F0FF9BFFD2911EF8EE0E836417" author="McLaughlin" box="[331,676,1800,1822]" pageId="2" pageNumber="69" refString="McLaughlin, P. A. &amp; Provenzano, A. J., Jr. (1974) The genus Paguristes (Crustacea: Decapoda: Diogenidae) from the western Atlantic. Part I. The Paguristes tortugae complex, with notes on variation. Bulletin of Marine Science, 24 (1), 165 - 234, figs 1 - 27." type="journal article" year="1974">McLaughlin &amp; Provenzano, 1974</bibRefCitation>
)
</taxonomicName>
*
</paragraph>
<paragraph id="8BA62E01FF9BFFD29092F8BE08EE649D" blockId="2.[151,1436,1880,2013]" pageId="2" pageNumber="69">
Antennular peduncles (
<figureCitation id="13223284FF9BFFD2918FF8BE0E066479" box="[474,545,1880,1904]" captionStart="FIGURE 1" captionStartId="3.[151,250,1589,1612]" captionTargetBox="[166,1428,367,1565]" captionTargetId="figure@3.[151,1436,353,1573]" captionTargetPageId="3" captionText="FIGURE 1. Areopaguristes tudgei n. sp., Carrie Bow Cay, Belize: a, c – e, h, male paratype, 2.1 mm ,, February 8 (USNM 1156993); b, f, g, i, female paratype, 2.1 mm, February 4 (USNM 1156991). a, shield and cephalic appendages, dorsal view; b, anterior portion of shield, dorsal view; c, male left first pleopod, posterior view; d, left second pleopod, posterior view; e, male left third pleopod, posterior view; f, female left first pleopod, posterior view; g, female left third pleopod, posterior view; h, uropods and telson, dorsal view; i, telson, dorsal view. Scales equal 0.5 mm (a, b, e, h, i), 0.2 mm (c, d, f, g)." httpUri="https://zenodo.org/record/214046/files/figure.png" pageId="2" pageNumber="69">Fig. 1</figureCitation>
a) slightly exceeding distal margins of corneas. Ultimate and penultimate segments subequal in length. Basal segment with spine on dorsolateral margin of statocyst lobe.
</paragraph>
<paragraph id="8BA62E01FF9BFFD39092F84608506236" blockId="2.[151,1436,1880,2013]" lastBlockId="3.[151,1436,151,319]" lastPageId="3" lastPageNumber="70" pageId="2" pageNumber="69">
Antennal peduncles (
<figureCitation id="13223284FF9BFFD291E3F8460DD164B1" box="[438,502,1952,1976]" captionStart="FIGURE 1" captionStartId="3.[151,250,1589,1612]" captionTargetBox="[166,1428,367,1565]" captionTargetId="figure@3.[151,1436,353,1573]" captionTargetPageId="3" captionText="FIGURE 1. Areopaguristes tudgei n. sp., Carrie Bow Cay, Belize: a, c – e, h, male paratype, 2.1 mm ,, February 8 (USNM 1156993); b, f, g, i, female paratype, 2.1 mm, February 4 (USNM 1156991). a, shield and cephalic appendages, dorsal view; b, anterior portion of shield, dorsal view; c, male left first pleopod, posterior view; d, left second pleopod, posterior view; e, male left third pleopod, posterior view; f, female left first pleopod, posterior view; g, female left third pleopod, posterior view; h, uropods and telson, dorsal view; i, telson, dorsal view. Scales equal 0.5 mm (a, b, e, h, i), 0.2 mm (c, d, f, g)." httpUri="https://zenodo.org/record/214046/files/figure.png" pageId="2" pageNumber="69">Fig. 1</figureCitation>
a) exceeding distal margin of corneas by about 0.2 length of fifth segment. Fourth segment with small dorsodistal spine. Third segment ventrodistal margin prominently produced and terminating in strong, acute spine. Second segment with plumose setae dorsally; with dorsolateral distal angle produced, terminating in simple or bifid spine and with 1–3 spines on lateral margin. First segment unarmed or sometimes with minute spine on distolateral angle. Antennal acicles reaching at most to about midportion of corneas, terminating in strong simple to bifid spine; mesial margin with 1–4 spines, lateral margin with 2–4 spines. Flagellum short, about as long as shield; all articles with short setae &lt;1 article in length.
</paragraph>
<caption id="DF667E89FF9AFFD390C2F9D308F865CF" httpUri="https://zenodo.org/record/214046/files/figure.png" pageId="3" pageNumber="70" targetBox="[166,1428,367,1565]" targetPageId="3">
<paragraph id="8BA62E01FF9AFFD390C2F9D308F865CF" blockId="3.[151,1436,1589,1734]" pageId="3" pageNumber="70">
<emphasis id="B96DF213FF9AFFD390C2F9D30D316542" bold="true" box="[151,278,1589,1612]" pageId="3" pageNumber="70">FIGURE 1.</emphasis>
<taxonomicName id="4C195582FF9AFFD39175F9D10E256545" ID-CoL="8c9c2161-2d9b-4ce2-a375-99a2a1f1a39b" box="[288,514,1590,1612]" class="Malacostraca" family="Diogenidae" genus="Areopaguristes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="70" phylum="Arthropoda" rank="species" species="tudgei" status="sp. nov.">
<emphasis id="B96DF213FF9AFFD39175F9D10E256545" box="[288,514,1590,1612]" italics="true" pageId="3" pageNumber="70">Areopaguristes tudgei</emphasis>
</taxonomicName>
<emphasis id="B96DF213FF9AFFD39259F9D00E6E6542" bold="true" box="[524,585,1590,1611]" pageId="3" pageNumber="70">
<taxonomicNameLabel id="A25E4F68FF9AFFD39259F9D00E6E6542" box="[524,585,1590,1611]" pageId="3" pageNumber="70" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, Carrie Bow Cay, Belize: a, c–e, h, male paratype, 2.1 mm,,February 8 (USNM 1156993); b, f, g, i, female paratype, 2.1 mm, February 4 (USNM 1156991). a, shield and cephalic appendages, dorsal view; b, anterior portion of shield, dorsal view; c, male left first pleopod, posterior view; d, left second pleopod, posterior view; e, male left third pleopod, posterior view; f, female left first pleopod, posterior view; g, female left third pleopod, posterior view; h, uropods and telson, dorsal view; i, telson, dorsal view. Scales equal 0.5 mm (a, b, e, h, i), 0.2 mm (c, d, f, g).
</paragraph>
</caption>
<paragraph id="8BA62E01FF9AFFD39092F9150F5764EA" blockId="3.[151,1436,1779,2020]" pageId="3" pageNumber="70">
Mandible (
<figureCitation id="13223284FF9AFFD39111F9150DAD6402" box="[324,394,1779,1803]" captionStart="FIGURE 2" captionStartId="4.[151,250,1683,1706]" captionTargetBox="[161,1433,205,1651]" captionTargetId="figure@4.[151,1436,193,1662]" captionTargetPageId="4" captionText="FIGURE 2. Areopaguristes tudgei n. sp.: male paratype, 2.1 mm, Carrie Bow Cay, Belize, February 8 (USNM 1156993). Left mouthparts, internal view: a, mandible; b, maxillule; c, maxilla; d, first maxilliped; e, second maxilliped; f, third maxilliped, g, ischium of same with crista dentata. Scales equal 0.1 mm (a – d, g), and 0.2 mm (e, f)." httpUri="https://zenodo.org/record/214047/files/figure.png" pageId="3" pageNumber="70">Fig. 2</figureCitation>
a) with upper and lower incisor edges calcareous; ultimate segment of palp broad, setose, nearly as long as combined length of penultimate and basal segments. Maxillule (
<figureCitation id="13223284FF9AFFD3941DF8F108AA6426" box="[1096,1165,1815,1839]" captionStart="FIGURE 2" captionStartId="4.[151,250,1683,1706]" captionTargetBox="[161,1433,205,1651]" captionTargetId="figure@4.[151,1436,193,1662]" captionTargetPageId="4" captionText="FIGURE 2. Areopaguristes tudgei n. sp.: male paratype, 2.1 mm, Carrie Bow Cay, Belize, February 8 (USNM 1156993). Left mouthparts, internal view: a, mandible; b, maxillule; c, maxilla; d, first maxilliped; e, second maxilliped; f, third maxilliped, g, ischium of same with crista dentata. Scales equal 0.1 mm (a – d, g), and 0.2 mm (e, f)." httpUri="https://zenodo.org/record/214047/files/figure.png" pageId="3" pageNumber="70">Fig. 2</figureCitation>
b) with proximal endite subquadrate; endopod with 2 or 3 long setae on weakly developed internal lobe, external lobe elongate, about half as long as endopod, and curving proximally. Maxilla (
<figureCitation id="13223284FF9AFFD3936FF8B90FA1647E" box="[826,902,1887,1911]" captionStart="FIGURE 2" captionStartId="4.[151,250,1683,1706]" captionTargetBox="[161,1433,205,1651]" captionTargetId="figure@4.[151,1436,193,1662]" captionTargetPageId="4" captionText="FIGURE 2. Areopaguristes tudgei n. sp.: male paratype, 2.1 mm, Carrie Bow Cay, Belize, February 8 (USNM 1156993). Left mouthparts, internal view: a, mandible; b, maxillule; c, maxilla; d, first maxilliped; e, second maxilliped; f, third maxilliped, g, ischium of same with crista dentata. Scales equal 0.1 mm (a – d, g), and 0.2 mm (e, f)." httpUri="https://zenodo.org/record/214047/files/figure.png" pageId="3" pageNumber="70">Fig. 2</figureCitation>
c) with endopod extending to distal end of scaphognathite, tapered distally. First maxilliped (
<figureCitation id="13223284FF9AFFD392B5F8650F006492" box="[736,807,1923,1947]" captionStart="FIGURE 2" captionStartId="4.[151,250,1683,1706]" captionTargetBox="[161,1433,205,1651]" captionTargetId="figure@4.[151,1436,193,1662]" captionTargetPageId="4" captionText="FIGURE 2. Areopaguristes tudgei n. sp.: male paratype, 2.1 mm, Carrie Bow Cay, Belize, February 8 (USNM 1156993). Left mouthparts, internal view: a, mandible; b, maxillule; c, maxilla; d, first maxilliped; e, second maxilliped; f, third maxilliped, g, ischium of same with crista dentata. Scales equal 0.1 mm (a – d, g), and 0.2 mm (e, f)." httpUri="https://zenodo.org/record/214047/files/figure.png" pageId="3" pageNumber="70">Fig. 2</figureCitation>
d) with endopod about 0.75 length of exopod, setose distally. Second maxilliped (
<figureCitation id="13223284FF9AFFD391BEF8410E1364B6" box="[491,564,1959,1983]" captionStart="FIGURE 2" captionStartId="4.[151,250,1683,1706]" captionTargetBox="[161,1433,205,1651]" captionTargetId="figure@4.[151,1436,193,1662]" captionTargetPageId="4" captionText="FIGURE 2. Areopaguristes tudgei n. sp.: male paratype, 2.1 mm, Carrie Bow Cay, Belize, February 8 (USNM 1156993). Left mouthparts, internal view: a, mandible; b, maxillule; c, maxilla; d, first maxilliped; e, second maxilliped; f, third maxilliped, g, ischium of same with crista dentata. Scales equal 0.1 mm (a – d, g), and 0.2 mm (e, f)." httpUri="https://zenodo.org/record/214047/files/figure.png" pageId="3" pageNumber="70">Fig. 2</figureCitation>
e) without distinctive characters. Third maxilliped (
<figureCitation id="13223284FF9AFFD394CAF84108CF64B6" box="[1183,1256,1959,1983]" captionStart="FIGURE 2" captionStartId="4.[151,250,1683,1706]" captionTargetBox="[161,1433,205,1651]" captionTargetId="figure@4.[151,1436,193,1662]" captionTargetPageId="4" captionText="FIGURE 2. Areopaguristes tudgei n. sp.: male paratype, 2.1 mm, Carrie Bow Cay, Belize, February 8 (USNM 1156993). Left mouthparts, internal view: a, mandible; b, maxillule; c, maxilla; d, first maxilliped; e, second maxilliped; f, third maxilliped, g, ischium of same with crista dentata. Scales equal 0.1 mm (a – d, g), and 0.2 mm (e, f)." httpUri="https://zenodo.org/record/214047/files/figure.png" pageId="3" pageNumber="70">Fig. 2</figureCitation>
f, g) with well developed crista dentata on ischium and without accessory tooth.
</paragraph>
<caption id="DF667E89FF9DFFD490C2F9750FD465E7" httpUri="https://zenodo.org/record/214047/files/figure.png" pageId="4" pageNumber="71" targetBox="[161,1433,205,1651]" targetPageId="4">
<paragraph id="8BA62E01FF9DFFD490C2F9750FD465E7" blockId="4.[151,1436,1683,1774]" pageId="4" pageNumber="71">
<emphasis id="B96DF213FF9DFFD490C2F9750D3465A0" bold="true" box="[151,275,1683,1706]" pageId="4" pageNumber="71">FIGURE 2.</emphasis>
<taxonomicName id="4C195582FF9DFFD4914CF9730DDF65A3" ID-CoL="8c9c2161-2d9b-4ce2-a375-99a2a1f1a39b" box="[281,504,1684,1706]" class="Malacostraca" family="Diogenidae" genus="Areopaguristes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="71" phylum="Arthropoda" rank="species" species="tudgei" status="sp. nov.">
<emphasis id="B96DF213FF9DFFD4914CF9730DDF65A3" box="[281,504,1684,1706]" italics="true" pageId="4" pageNumber="71">Areopaguristes tudgei</emphasis>
</taxonomicName>
<emphasis id="B96DF213FF9DFFD491AAF9720E1F65A0" bold="true" box="[511,568,1684,1705]" pageId="4" pageNumber="71">
<taxonomicNameLabel id="A25E4F68FF9DFFD491AAF9720E1F65A0" box="[511,568,1684,1705]" pageId="4" pageNumber="71" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
: male paratype, 2.1 mm, Carrie Bow Cay, Belize, February 8 (USNM 1156993). Left mouthparts, internal view: a, mandible; b, maxillule; c, maxilla; d, first maxilliped; e, second maxilliped; f, third maxilliped, g, ischium of same with crista dentata. Scales equal 0.1 mm (a–d, g), and 0.2 mm (e, f).
</paragraph>
</caption>
<paragraph id="8BA62E01FF9DFFD59092F8F90F6A62C6" blockId="4.[151,1436,1823,2026]" lastBlockId="5.[151,1436,151,463]" lastPageId="5" lastPageNumber="72" pageId="4" pageNumber="71">
Chelipeds (
<figureCitation id="13223284FF9DFFD49113F8F90DA0643E" box="[326,391,1823,1847]" captionStart="FIGURE 3" captionStartId="5.[151,250,1316,1339]" captionTargetBox="[159,1428,529,1289]" captionTargetId="figure@5.[151,1436,517,1295]" captionTargetPageId="5" captionText="FIGURE 3. Areopaguristes tudgei n. sp.: male paratype, 2.1 mm, Carrie Bow Cay, Belize, February 8 (USNM 1156993). Chelipeds, all dorsal view except f (mesial), and c (ventral); a, b, showing approximate density of setation; c – f, with setae omitted. a, d, left; b, e, right; c, fingers of same; f, right. Scale equals 0.5 mm." httpUri="https://zenodo.org/record/214048/files/figure.png" pageId="4" pageNumber="71">Fig. 3</figureCitation>
a–f) subequal, or left slightly larger than right, similarly armed; dorsal surfaces of chelae and carpi densely covered with tufts of plumose setae obscuring armature beneath, setae evenly present or more dense and long on dorsolateral margins of chelae, setation generally denser in males than in females. Fingers and palms with dorsal surfaces covered with irregularly placed small spines or tubercles; cutting edge of fingers each with several small calcareous teeth on proximal 0.7, and with row of fused corneous teeth forming hoof-like claw distally (
<figureCitation id="13223284FF9DFFD490AFF8340D1964E3" box="[250,318,2002,2026]" captionStart="FIGURE 3" captionStartId="5.[151,250,1316,1339]" captionTargetBox="[159,1428,529,1289]" captionTargetId="figure@5.[151,1436,517,1295]" captionTargetPageId="5" captionText="FIGURE 3. Areopaguristes tudgei n. sp.: male paratype, 2.1 mm, Carrie Bow Cay, Belize, February 8 (USNM 1156993). Chelipeds, all dorsal view except f (mesial), and c (ventral); a, b, showing approximate density of setation; c – f, with setae omitted. a, d, left; b, e, right; c, fingers of same; f, right. Scale equals 0.5 mm." httpUri="https://zenodo.org/record/214048/files/figure.png" pageId="4" pageNumber="71">Fig. 3</figureCitation>
c). Dactyl about 1.4 times as long as palm. Palms with dorsal surfaces evenly convex; each with 3 strong, corneous-tipped spines on dorsomesial margin; lateral and mesial margins rounded, unarmed or with scattered small tubercles; ventral faces mostly unarmed or with scattered small tubercles and few ventrolateral plumose setae. Carpi with dorsal surfaces with well-spaced small spines or tubercles; dorsomesial margins each with 4 or 5 corneous-tipped spines (proximal 2 spines small), dorsolateral margins with regular or irregular rows of small spines; lateral, mesial, and ventral faces unarmed or with scattered small tubercles and setae. Meri long, about 1.7 times as long as carpi, subtriangular in crossection; dorsal margins with plumose setae; lateral and mesial faces unarmed except for transverse row of minute blunt spines subdistally on each face; ventrolateral and ventromesial margins each with row of spines (smaller on ventrolateral margins); ventral faces with plumose setae. Ischia unarmed except for few setae on ventromesial margins.
</paragraph>
<caption id="DF667E89FF9CFFD590C2FAC20F8B6676" httpUri="https://zenodo.org/record/214048/files/figure.png" pageId="5" pageNumber="72" targetBox="[159,1428,529,1289]" targetPageId="5">
<paragraph id="8BA62E01FF9CFFD590C2FAC20F8B6676" blockId="5.[151,1437,1316,1407]" pageId="5" pageNumber="72">
<emphasis id="B96DF213FF9CFFD590C2FAC20D326630" bold="true" box="[151,277,1316,1339]" pageId="5" pageNumber="72">FIGURE 3.</emphasis>
<taxonomicName id="4C195582FF9CFFD5914AFAC30E266633" ID-CoL="8c9c2161-2d9b-4ce2-a375-99a2a1f1a39b" box="[287,513,1316,1338]" class="Malacostraca" family="Diogenidae" genus="Areopaguristes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="72" phylum="Arthropoda" rank="species" species="tudgei" status="sp. nov.">
<emphasis id="B96DF213FF9CFFD5914AFAC30E266633" box="[287,513,1316,1338]" italics="true" pageId="5" pageNumber="72">Areopaguristes tudgei</emphasis>
</taxonomicName>
<emphasis id="B96DF213FF9CFFD5925FFAC30E616633" bold="true" box="[522,582,1317,1338]" pageId="5" pageNumber="72">
<taxonomicNameLabel id="A25E4F68FF9CFFD5925FFAC30E616633" box="[522,582,1317,1338]" pageId="5" pageNumber="72" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
: male paratype, 2.1 mm, Carrie Bow Cay, Belize, February 8 (USNM 1156993). Chelipeds, all dorsal view except f (mesial), and c (ventral); a, b, showing approximate density of setation; c–f, with setae omitted. a, d, left; b, e, right; c, fingers of same; f, right. Scale equals 0.5 mm.
</paragraph>
</caption>
<paragraph id="8BA62E01FF9CFFD59092FA490EFD65EE" blockId="5.[151,1437,1455,2020]" pageId="5" pageNumber="72">
Second and third pereopods (
<figureCitation id="13223284FF9CFFD5925AFA490E7566CE" box="[527,594,1455,1479]" captionStart="FIGURE 4" captionStartId="6.[151,250,1819,1842]" captionTargetBox="[162,1432,331,1795]" captionTargetId="figure@6.[151,1436,317,1806]" captionTargetPageId="6" captionText="FIGURE 4. Areopaguristes tudgei n. sp.: male paratype, 2.1 mm, Carrie Bow Cay, Belize, February 8 (USNM 1156993). a, second right pereopod with plumose setae, lateral view; b, dactyl of same, mesial view; c, same pereopod with setae omitted, mesial view; d, third right pereopod, lateral view; e, dactyl of same; f, propodus and dactyl of right fourth pereopod, lateral view; g, propodus and dactyl of right fifth pereopod, lateral view. Scales equal 0.5 mm." httpUri="https://zenodo.org/record/214049/files/figure.png" pageId="5" pageNumber="72">Fig. 4</figureCitation>
a–e) generally similar except left slightly shorter than right; dorsal margins of meri, carpi, propodi and dactyls with dense fringe of plumose setae. Dactyls about same length as propodi, terminating in sharp corneous claw; ventromesial margin armed with 9–11 corneous spinules in addition to long subdistal bristle near base of claw; ventral margins with well-spaced plumose setae. Propodi each with dorsomesial row of small spines; dorsolateral faces with short low ridges and with scattered plumose setae; ventromesial faces each with short, transverse rows of small spines. Carpi each with dorsal row of spines; dorsolateral face with dense plumose setae, and short low ridges; dorsomesial face with irregular rows of small spines. Meri each with plumose setae on ventral margin and ventromesial distal row of small spines. Anterior portion of sternite XII (
<figureCitation id="13223284FF9CFFD59540F94D097E65CA" box="[1301,1369,1707,1731]" captionStart="FIGURE 5" captionStartId="7.[151,250,1852,1875]" captionTargetBox="[173,1424,276,1818]" captionTargetId="figure@7.[151,1436,265,1831]" captionTargetPageId="7" captionText="FIGURE 5. Areopaguristes tudgei n. sp.: male paratype, 2.1 mm, Carrie Bow Cay, Belize, February 8 (USNM 1156993). Sternites XI – XIV, coxae of chelipeds and second to fifth pereopods, and anterior portion of pleon with first and second pleopods, ventral view. Scale equals 0.5 mm." httpUri="https://zenodo.org/record/214050/files/figure.png" pageId="5" pageNumber="72">Fig. 5</figureCitation>
) with transverse, setose ridge slightly elevated at midline.
</paragraph>
<paragraph id="8BA62E01FF9CFFD59092F9150DB66471" blockId="5.[151,1437,1455,2020]" pageId="5" pageNumber="72">
Fourth pereopods (
<figureCitation id="13223284FF9CFFD591CEF9150DF96402" box="[411,478,1779,1803]" captionStart="FIGURE 4" captionStartId="6.[151,250,1819,1842]" captionTargetBox="[162,1432,331,1795]" captionTargetId="figure@6.[151,1436,317,1806]" captionTargetPageId="6" captionText="FIGURE 4. Areopaguristes tudgei n. sp.: male paratype, 2.1 mm, Carrie Bow Cay, Belize, February 8 (USNM 1156993). a, second right pereopod with plumose setae, lateral view; b, dactyl of same, mesial view; c, same pereopod with setae omitted, mesial view; d, third right pereopod, lateral view; e, dactyl of same; f, propodus and dactyl of right fourth pereopod, lateral view; g, propodus and dactyl of right fifth pereopod, lateral view. Scales equal 0.5 mm." httpUri="https://zenodo.org/record/214049/files/figure.png" pageId="5" pageNumber="72">Fig. 4</figureCitation>
f) with propodi distinctly longer than high, or infrequently in some large males (≥ 2.0 mm) about as long as high; propodal rasp extending about 0.7 distal length of segment, with 2 or 3 rows of ovate corneous scales distally, and 1 row proximally. Fifth pereopods (
<figureCitation id="13223284FF9CFFD59333F8DD0F8F645A" box="[870,936,1851,1875]" captionStart="FIGURE 4" captionStartId="6.[151,250,1819,1842]" captionTargetBox="[162,1432,331,1795]" captionTargetId="figure@6.[151,1436,317,1806]" captionTargetPageId="6" captionText="FIGURE 4. Areopaguristes tudgei n. sp.: male paratype, 2.1 mm, Carrie Bow Cay, Belize, February 8 (USNM 1156993). a, second right pereopod with plumose setae, lateral view; b, dactyl of same, mesial view; c, same pereopod with setae omitted, mesial view; d, third right pereopod, lateral view; e, dactyl of same; f, propodus and dactyl of right fourth pereopod, lateral view; g, propodus and dactyl of right fifth pereopod, lateral view. Scales equal 0.5 mm." httpUri="https://zenodo.org/record/214049/files/figure.png" pageId="5" pageNumber="72">
Fig.
<quantity id="4CE183E4FF9CFFD593CCF8DD0F8F645A" box="[921,936,1851,1875]" metricMagnitude="-3" metricUnit="kg" metricValue="4.0" pageId="5" pageNumber="72" unit="g" value="4.0">4</quantity>
</figureCitation>
g) chelate; propodal rasp extending for about 0.3 length of segment.
</paragraph>
<paragraph id="8BA62E01FF9CFFD69092F865088F6215" blockId="5.[151,1437,1455,2020]" lastBlockId="6.[151,1436,151,285]" lastPageId="6" lastPageNumber="73" pageId="5" pageNumber="72">
Male with paired first and second pleopods modified as gonopods, and unpaired left second to fifth; first pleopod (
<figureCitation id="13223284FF9CFFD59156F8410D6364B6" box="[259,324,1959,1983]" captionStart="FIGURE 1" captionStartId="3.[151,250,1589,1612]" captionTargetBox="[166,1428,367,1565]" captionTargetId="figure@3.[151,1436,353,1573]" captionTargetPageId="3" captionText="FIGURE 1. Areopaguristes tudgei n. sp., Carrie Bow Cay, Belize: a, c – e, h, male paratype, 2.1 mm ,, February 8 (USNM 1156993); b, f, g, i, female paratype, 2.1 mm, February 4 (USNM 1156991). a, shield and cephalic appendages, dorsal view; b, anterior portion of shield, dorsal view; c, male left first pleopod, posterior view; d, left second pleopod, posterior view; e, male left third pleopod, posterior view; f, female left first pleopod, posterior view; g, female left third pleopod, posterior view; h, uropods and telson, dorsal view; i, telson, dorsal view. Scales equal 0.5 mm (a, b, e, h, i), 0.2 mm (c, d, f, g)." httpUri="https://zenodo.org/record/214046/files/figure.png" pageId="5" pageNumber="72">Fig. 1</figureCitation>
c) with long setae on mesial margin of internal lobe, distal margins of external lobe and inferior lamella entire, lacking spines, teeth or setae; second pleopod (
<figureCitation id="13223284FF9CFFD59328F82D0FE564EA" box="[893,962,1995,2019]" captionStart="FIGURE 1" captionStartId="3.[151,250,1589,1612]" captionTargetBox="[166,1428,367,1565]" captionTargetId="figure@3.[151,1436,353,1573]" captionTargetPageId="3" captionText="FIGURE 1. Areopaguristes tudgei n. sp., Carrie Bow Cay, Belize: a, c – e, h, male paratype, 2.1 mm ,, February 8 (USNM 1156993); b, f, g, i, female paratype, 2.1 mm, February 4 (USNM 1156991). a, shield and cephalic appendages, dorsal view; b, anterior portion of shield, dorsal view; c, male left first pleopod, posterior view; d, left second pleopod, posterior view; e, male left third pleopod, posterior view; f, female left first pleopod, posterior view; g, female left third pleopod, posterior view; h, uropods and telson, dorsal view; i, telson, dorsal view. Scales equal 0.5 mm (a, b, e, h, i), 0.2 mm (c, d, f, g)." httpUri="https://zenodo.org/record/214046/files/figure.png" pageId="5" pageNumber="72">Fig. 1</figureCitation>
d) with basal segment naked except for occasional few distal setae, distal segment with setose margins; third to fifth pleopods (
<figureCitation id="13223284FF9FFFD694CCFF7108FA63A6" box="[1177,1245,151,175]" captionStart="FIGURE 1" captionStartId="3.[151,250,1589,1612]" captionTargetBox="[166,1428,367,1565]" captionTargetId="figure@3.[151,1436,353,1573]" captionTargetPageId="3" captionText="FIGURE 1. Areopaguristes tudgei n. sp., Carrie Bow Cay, Belize: a, c – e, h, male paratype, 2.1 mm ,, February 8 (USNM 1156993); b, f, g, i, female paratype, 2.1 mm, February 4 (USNM 1156991). a, shield and cephalic appendages, dorsal view; b, anterior portion of shield, dorsal view; c, male left first pleopod, posterior view; d, left second pleopod, posterior view; e, male left third pleopod, posterior view; f, female left first pleopod, posterior view; g, female left third pleopod, posterior view; h, uropods and telson, dorsal view; i, telson, dorsal view. Scales equal 0.5 mm (a, b, e, h, i), 0.2 mm (c, d, f, g)." httpUri="https://zenodo.org/record/214046/files/figure.png" pageId="6" pageNumber="73">Fig. 1</figureCitation>
e) with vestigial endopods. Female with paired gonopods, paired first pleopods (
<figureCitation id="13223284FF9FFFD69334FF5A0F8763DD" box="[865,928,188,212]" captionStart="FIGURE 1" captionStartId="3.[151,250,1589,1612]" captionTargetBox="[166,1428,367,1565]" captionTargetId="figure@3.[151,1436,353,1573]" captionTargetPageId="3" captionText="FIGURE 1. Areopaguristes tudgei n. sp., Carrie Bow Cay, Belize: a, c – e, h, male paratype, 2.1 mm ,, February 8 (USNM 1156993); b, f, g, i, female paratype, 2.1 mm, February 4 (USNM 1156991). a, shield and cephalic appendages, dorsal view; b, anterior portion of shield, dorsal view; c, male left first pleopod, posterior view; d, left second pleopod, posterior view; e, male left third pleopod, posterior view; f, female left first pleopod, posterior view; g, female left third pleopod, posterior view; h, uropods and telson, dorsal view; i, telson, dorsal view. Scales equal 0.5 mm (a, b, e, h, i), 0.2 mm (c, d, f, g)." httpUri="https://zenodo.org/record/214046/files/figure.png" pageId="6" pageNumber="73">Fig. 1</figureCitation>
f), and unpaired left second to fifth (
<figureCitation id="13223284FF9FFFD6956EFF5A095C63DD" box="[1339,1403,188,212]" captionStart="FIGURE 1" captionStartId="3.[151,250,1589,1612]" captionTargetBox="[166,1428,367,1565]" captionTargetId="figure@3.[151,1436,353,1573]" captionTargetPageId="3" captionText="FIGURE 1. Areopaguristes tudgei n. sp., Carrie Bow Cay, Belize: a, c – e, h, male paratype, 2.1 mm ,, February 8 (USNM 1156993); b, f, g, i, female paratype, 2.1 mm, February 4 (USNM 1156991). a, shield and cephalic appendages, dorsal view; b, anterior portion of shield, dorsal view; c, male left first pleopod, posterior view; d, left second pleopod, posterior view; e, male left third pleopod, posterior view; f, female left first pleopod, posterior view; g, female left third pleopod, posterior view; h, uropods and telson, dorsal view; i, telson, dorsal view. Scales equal 0.5 mm (a, b, e, h, i), 0.2 mm (c, d, f, g)." httpUri="https://zenodo.org/record/214046/files/figure.png" pageId="6" pageNumber="73">
Fig.
<quantity id="4CE183E4FF9FFFD69525FF5A095C63DD" box="[1392,1403,188,212]" metricMagnitude="-3" metricUnit="kg" metricValue="1.0" pageId="6" pageNumber="73" unit="g" value="1.0">1</quantity>
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g), lacking brood pouch; first pleopod with naked basal segment, and distal segment with long, marginal bristle-like setae; number of eggs ranging 8–21 (n = 10), relatively large (~
<quantity id="4CE183E4FF9FFFD69330FEE20F9C6215" box="[869,955,260,284]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="6" pageNumber="73" unit="mm" value="0.7">0.7 mm</quantity>
in maximum width).
</paragraph>
<caption id="DF667E89FF9FFFD690C2F8FD082C6484" httpUri="https://zenodo.org/record/214049/files/figure.png" pageId="6" pageNumber="73" targetBox="[162,1432,331,1795]" targetPageId="6">
<paragraph id="8BA62E01FF9FFFD690C2F8FD082C6484" blockId="6.[151,1436,1819,1933]" pageId="6" pageNumber="73">
<emphasis id="B96DF213FF9FFFD690C2F8FD0D336438" bold="true" box="[151,276,1819,1842]" pageId="6" pageNumber="73">FIGURE 4.</emphasis>
<taxonomicName id="4C195582FF9FFFD69149F8FA0DDB643B" ID-CoL="8c9c2161-2d9b-4ce2-a375-99a2a1f1a39b" box="[284,508,1820,1842]" class="Malacostraca" family="Diogenidae" genus="Areopaguristes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="73" phylum="Arthropoda" rank="species" species="tudgei" status="sp. nov.">
<emphasis id="B96DF213FF9FFFD69149F8FA0DDB643B" box="[284,508,1820,1842]" italics="true" pageId="6" pageNumber="73">Areopaguristes tudgei</emphasis>
</taxonomicName>
<emphasis id="B96DF213FF9FFFD69251F8FA0E186438" bold="true" box="[516,575,1820,1841]" pageId="6" pageNumber="73">
<taxonomicNameLabel id="A25E4F68FF9FFFD69251F8FA0E186438" box="[516,575,1820,1841]" pageId="6" pageNumber="73" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
: male paratype, 2.1 mm, Carrie Bow Cay, Belize, February 8 (USNM 1156993). a, second right pereopod with plumose setae, lateral view; b, dactyl of same, mesial view; c, same pereopod with setae omitted, mesial view; d, third right pereopod, lateral view; e, dactyl of same; f, propodus and dactyl of right fourth pereopod, lateral view; g, propodus and dactyl of right fifth pereopod, lateral view. Scales equal 0.5 mm.
</paragraph>
</caption>
<paragraph id="8BA62E01FF9FFFD79092F8530E5563DD" blockId="6.[151,1436,1973,2033]" lastBlockId="7.[151,1436,151,212]" lastPageId="7" lastPageNumber="74" pageId="6" pageNumber="73">
Telson and uropods (
<figureCitation id="13223284FF9FFFD691E7F8530DD564C4" box="[434,498,1973,1997]" captionStart="FIGURE 1" captionStartId="3.[151,250,1589,1612]" captionTargetBox="[166,1428,367,1565]" captionTargetId="figure@3.[151,1436,353,1573]" captionTargetPageId="3" captionText="FIGURE 1. Areopaguristes tudgei n. sp., Carrie Bow Cay, Belize: a, c – e, h, male paratype, 2.1 mm ,, February 8 (USNM 1156993); b, f, g, i, female paratype, 2.1 mm, February 4 (USNM 1156991). a, shield and cephalic appendages, dorsal view; b, anterior portion of shield, dorsal view; c, male left first pleopod, posterior view; d, left second pleopod, posterior view; e, male left third pleopod, posterior view; f, female left first pleopod, posterior view; g, female left third pleopod, posterior view; h, uropods and telson, dorsal view; i, telson, dorsal view. Scales equal 0.5 mm (a, b, e, h, i), 0.2 mm (c, d, f, g)." httpUri="https://zenodo.org/record/214046/files/figure.png" pageId="6" pageNumber="73">Fig. 1</figureCitation>
h, i) asymmetrical. Telson with distinct, deep lateral incisions dividing anterior and posterior portions; anterior lobes with long setae on posterolateral angles; posterior lobes separated by shallow median cleft; terminal margins oblique, setose, each with 1–5 small to minute, blunt or sharp calcareous spines (larger on left side) often curved ventrally.
</paragraph>
<caption id="DF667E89FF9EFFD790C2F8DA0E476499" httpUri="https://zenodo.org/record/214050/files/figure.png" pageId="7" pageNumber="74" targetBox="[173,1424,276,1818]" targetPageId="7">
<paragraph id="8BA62E01FF9EFFD790C2F8DA0E476499" blockId="7.[151,1437,1852,1936]" pageId="7" pageNumber="74">
<emphasis id="B96DF213FF9EFFD790C2F8DA0D32645B" bold="true" box="[151,277,1852,1875]" pageId="7" pageNumber="74">FIGURE 5.</emphasis>
<taxonomicName id="4C195582FF9EFFD7914AF8DB0E26645A" ID-CoL="8c9c2161-2d9b-4ce2-a375-99a2a1f1a39b" box="[287,513,1853,1875]" class="Malacostraca" family="Diogenidae" genus="Areopaguristes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="74" phylum="Arthropoda" rank="species" species="tudgei" status="sp. nov.">
<emphasis id="B96DF213FF9EFFD7914AF8DB0E26645A" box="[287,513,1853,1875]" italics="true" pageId="7" pageNumber="74">Areopaguristes tudgei</emphasis>
</taxonomicName>
<emphasis id="B96DF213FF9EFFD7925FF8DB0E61645B" bold="true" box="[522,582,1853,1874]" pageId="7" pageNumber="74">
<taxonomicNameLabel id="A25E4F68FF9EFFD7925FF8DB0E61645B" box="[522,582,1853,1874]" pageId="7" pageNumber="74" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
: male paratype, 2.1 mm, Carrie Bow Cay, Belize, February 8 (USNM 1156993). Sternites XI–XIV, coxae of chelipeds and second to fifth pereopods, and anterior portion of pleon with first and second pleopods, ventral view. Scale equals 0.5 mm.
</paragraph>
</caption>
<paragraph id="8BA62E01FF9EFFD89092F8510D0D62FD" blockId="7.[151,1435,1975,2035]" lastBlockId="8.[151,1436,151,500]" lastPageId="8" lastPageNumber="75" pageId="7" pageNumber="74">
<emphasis id="B96DF213FF9EFFD79092F8510D2C64D9" bold="true" box="[199,267,1975,2000]" pageId="7" pageNumber="74">Color</emphasis>
(
<figureCitation id="13223284FF9EFFD7914EF8510D7864C6" box="[283,351,1975,1999]" captionStart="FIGURE 6" captionStartId="8.[151,250,1256,1279]" captionTargetBox="[151,1436,553,1234]" captionTargetId="figure@8.[151,1436,553,1235]" captionTargetPageId="8" captionText="FIGURE 6. Areopaguristes tudgei n. sp., Carrie Bow Cay, Belize, February 4: a, female paratype 1.9 mm (ULLZ 12530); b, male paratype 3.0 mm (ULLZ 12576). Scales equal 1 mm." httpUri="https://zenodo.org/record/214051/files/figure.png" pageId="7" pageNumber="74">Fig. 6</figureCitation>
). In life, shield whitish or cream color, with 3 brownish-red patches anteriorly (1 directly behind each anterolateral projection, and 1 behind the rostrum). Posterior carapace semitransparent; anterolateral corner with distinct pink patch. Ocular peduncles whitish except for short distal faint, dorsomesial brown stripe; distally semitransparent. Ocular scales whitish. Antennules whitish except for light blue distal segment and flagella. Antennal acicles whitish except for acicle distally brownish, and flagella light blue. Chelipeds with dorsal surfaces of carpi and chelae brownish except for whitish distal portion of fingers; meri whitish with dorsodistal surfaces brownish. Walking legs (second and third pereopods) with dactyls brown except for whitish distal portion; carpi, and propodi brownish on entire mesial faces, on dorsal surface and on upper half of lateral faces; meri whitish except for brownish dorsodistal portion. Fourth and fifth pereopods whitish. Pleon with pair of distinct small dark spots anterodorsally near division of thorax and pleon, and several dot-like spots on each side between second and third pleonal somite (spots visible on
<figureCitation id="13223284FF91FFD8926BFE510E5E62C6" box="[574,633,439,463]" captionStart="FIGURE 6" captionStartId="8.[151,250,1256,1279]" captionTargetBox="[151,1436,553,1234]" captionTargetId="figure@8.[151,1436,553,1235]" captionTargetPageId="8" captionText="FIGURE 6. Areopaguristes tudgei n. sp., Carrie Bow Cay, Belize, February 4: a, female paratype 1.9 mm (ULLZ 12530); b, male paratype 3.0 mm (ULLZ 12576). Scales equal 1 mm." httpUri="https://zenodo.org/record/214051/files/figure.png" pageId="8" pageNumber="75">Fig 6</figureCitation>
a only). Female pleon showing dark orange gonads; male pleon whitish or yellowish.
</paragraph>
</subSubSection>
<caption id="DF667E89FF91FFD890C2FB0E0ECE6628" httpUri="https://zenodo.org/record/214051/files/figure.png" pageId="8" pageNumber="75" targetBox="[151,1436,553,1234]" targetPageId="8">
<paragraph id="8BA62E01FF91FFD890C2FB0E0ECE6628" blockId="8.[151,1435,1256,1313]" pageId="8" pageNumber="75">
<emphasis id="B96DF213FF91FFD890C2FB0E0D3467F4" bold="true" box="[151,275,1256,1279]" pageId="8" pageNumber="75">FIGURE 6.</emphasis>
<taxonomicName id="4C195582FF91FFD8914EFB0F0DDC67F7" box="[283,507,1256,1278]" class="Malacostraca" family="Diogenidae" genus="Areopaguristes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="75" phylum="Arthropoda" rank="species" species="tudgei" status="sp. nov.">
<emphasis id="B96DF213FF91FFD8914EFB0F0DDC67F7" box="[283,507,1256,1278]" italics="true" pageId="8" pageNumber="75">Areopaguristes tudgei</emphasis>
</taxonomicName>
<emphasis id="B96DF213FF91FFD89256FB0F0E1A67F7" bold="true" box="[515,573,1257,1278]" pageId="8" pageNumber="75">
<taxonomicNameLabel id="A25E4F68FF91FFD89256FB0F0E1A67F7" box="[515,573,1257,1278]" pageId="8" pageNumber="75" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, Carrie Bow Cay, Belize, February 4: a, female paratype 1.9 mm (ULLZ 12530); b, male paratype 3.0 mm (ULLZ 12576). Scales equal 1 mm.
</paragraph>
</caption>
<subSubSection id="C3037D8AFF91FFD89092FAB70EC166B8" pageId="8" pageNumber="75" type="distribution">
<paragraph id="8BA62E01FF91FFD89092FAB70EC166B8" blockId="8.[151,1436,1361,2034]" pageId="8" pageNumber="75">
<emphasis id="B96DF213FF91FFD89092FAB70DD46663" bold="true" box="[199,499,1361,1386]" pageId="8" pageNumber="75">Distribution and habitat.</emphasis>
So far known only from Carrie Bow Cay, on the Caribbean Mesoamerican Barrier Reef off the coast of
<collectingCountry id="F30E6E91FF91FFD891DCFA930DF36684" box="[393,468,1397,1421]" name="Belize" pageId="8" pageNumber="75">Belize</collectingCountry>
; intertidal under dead coral boulders heavily encrusted with calcareous red and other algae. All specimens were living in gastropod shells.
</paragraph>
</subSubSection>
<subSubSection id="C3037D8AFF91FFD89092FA5809BB64A0" pageId="8" pageNumber="75" type="etymology">
<paragraph id="8BA62E01FF91FFD89092FA580E1E6548" blockId="8.[151,1436,1361,2034]" pageId="8" pageNumber="75">
<emphasis id="B96DF213FF91FFD89092FA580D6966DF" bold="true" box="[199,334,1470,1494]" pageId="8" pageNumber="75">Etymology.</emphasis>
This species is named after our colleague Christopher C. Tudge (American University) who first noticed and collected populations of this diminutive hermit crab living under large dead coral boulders during joint field work in Carrie Bow Cay. The name also acknowledges his unique contributions to knowledge of the reproductive biology of hermit crabs.
</paragraph>
<paragraph id="8BA62E01FF91FFD89092F9AB09BB64A0" blockId="8.[151,1436,1361,2034]" pageId="8" pageNumber="75">
<emphasis id="B96DF213FF91FFD89092F9AB0E27656F" bold="true" box="[199,512,1613,1638]" pageId="8" pageNumber="75">Morphological variations.</emphasis>
In the 80+ specimens examined, morphological variations are minimal in some characters, such as spination of chelipeds and walking legs. The shield is smooth, usually lacking spines, although rarely a few minute, blunt spines are present on the anterior dorsolateral margin. In other characters, variations are significant or related to sex, such as the shape of the rostrum, degree and pattern of setation on the chelipeds, and the armature of the terminal margins of the telson. The rostrum varies from moderately slender to slender (
<figureCitation id="13223284FF91FFD89513F93B09A165FC" box="[1350,1414,1757,1781]" captionStart="FIGURE 1" captionStartId="3.[151,250,1589,1612]" captionTargetBox="[166,1428,367,1565]" captionTargetId="figure@3.[151,1436,353,1573]" captionTargetPageId="3" captionText="FIGURE 1. Areopaguristes tudgei n. sp., Carrie Bow Cay, Belize: a, c – e, h, male paratype, 2.1 mm ,, February 8 (USNM 1156993); b, f, g, i, female paratype, 2.1 mm, February 4 (USNM 1156991). a, shield and cephalic appendages, dorsal view; b, anterior portion of shield, dorsal view; c, male left first pleopod, posterior view; d, left second pleopod, posterior view; e, male left third pleopod, posterior view; f, female left first pleopod, posterior view; g, female left third pleopod, posterior view; h, uropods and telson, dorsal view; i, telson, dorsal view. Scales equal 0.5 mm (a, b, e, h, i), 0.2 mm (c, d, f, g)." httpUri="https://zenodo.org/record/214046/files/figure.png" pageId="8" pageNumber="75">Fig. 1</figureCitation>
a, b). Setation of the carpi and chelae of both cheliped is denser in males than in females. On the chelae of the latter, a dorsolateral fringe of plumose setae is typically present, whereas in males the plumose setae are not arranged as a fringe and instead are more or less evenly dense throughout the dorsal surface of the chelae (
<figureCitation id="13223284FF91FFD894B6F8AF090E6468" box="[1251,1321,1865,1889]" captionStart="FIGURE 3" captionStartId="5.[151,250,1316,1339]" captionTargetBox="[159,1428,529,1289]" captionTargetId="figure@5.[151,1436,517,1295]" captionTargetPageId="5" captionText="FIGURE 3. Areopaguristes tudgei n. sp.: male paratype, 2.1 mm, Carrie Bow Cay, Belize, February 8 (USNM 1156993). Chelipeds, all dorsal view except f (mesial), and c (ventral); a, b, showing approximate density of setation; c – f, with setae omitted. a, d, left; b, e, right; c, fingers of same; f, right. Scale equals 0.5 mm." httpUri="https://zenodo.org/record/214048/files/figure.png" pageId="8" pageNumber="75">Fig. 3</figureCitation>
a, b). The terminal margins of the telson can be armed on each side with one to five blunt, calcareous spines (
<figureCitation id="13223284FF91FFD8954DF88B097D648C" box="[1304,1370,1901,1925]" captionStart="FIGURE 1" captionStartId="3.[151,250,1589,1612]" captionTargetBox="[166,1428,367,1565]" captionTargetId="figure@3.[151,1436,353,1573]" captionTargetPageId="3" captionText="FIGURE 1. Areopaguristes tudgei n. sp., Carrie Bow Cay, Belize: a, c – e, h, male paratype, 2.1 mm ,, February 8 (USNM 1156993); b, f, g, i, female paratype, 2.1 mm, February 4 (USNM 1156991). a, shield and cephalic appendages, dorsal view; b, anterior portion of shield, dorsal view; c, male left first pleopod, posterior view; d, left second pleopod, posterior view; e, male left third pleopod, posterior view; f, female left first pleopod, posterior view; g, female left third pleopod, posterior view; h, uropods and telson, dorsal view; i, telson, dorsal view. Scales equal 0.5 mm (a, b, e, h, i), 0.2 mm (c, d, f, g)." httpUri="https://zenodo.org/record/214046/files/figure.png" pageId="8" pageNumber="75">Fig. 1</figureCitation>
h), or often with only one minute spine on each laterodistal angle (
<figureCitation id="13223284FF91FFD89363F8770F5264A0" box="[822,885,1937,1961]" captionStart="FIGURE 1" captionStartId="3.[151,250,1589,1612]" captionTargetBox="[166,1428,367,1565]" captionTargetId="figure@3.[151,1436,353,1573]" captionTargetPageId="3" captionText="FIGURE 1. Areopaguristes tudgei n. sp., Carrie Bow Cay, Belize: a, c – e, h, male paratype, 2.1 mm ,, February 8 (USNM 1156993); b, f, g, i, female paratype, 2.1 mm, February 4 (USNM 1156991). a, shield and cephalic appendages, dorsal view; b, anterior portion of shield, dorsal view; c, male left first pleopod, posterior view; d, left second pleopod, posterior view; e, male left third pleopod, posterior view; f, female left first pleopod, posterior view; g, female left third pleopod, posterior view; h, uropods and telson, dorsal view; i, telson, dorsal view. Scales equal 0.5 mm (a, b, e, h, i), 0.2 mm (c, d, f, g)." httpUri="https://zenodo.org/record/214046/files/figure.png" pageId="8" pageNumber="75">
Fig.
<date id="FFA708C1FF91FFD8933CF8770F5264A0" box="[873,885,1937,1961]" pageId="8" pageNumber="75">1</date>
</figureCitation>
i), or occasionally unarmed except for long setae.
</paragraph>
</subSubSection>
<subSubSection id="C3037D8AFF91FFD99092F8500FAE665D" lastPageId="9" lastPageNumber="76" pageId="8" pageNumber="75" type="discussion">
<paragraph id="8BA62E01FF91FFD99092F85008B3628E" blockId="8.[151,1436,1361,2034]" lastBlockId="9.[151,1436,151,1364]" lastPageId="9" lastPageNumber="76" pageId="8" pageNumber="75">
<emphasis id="B96DF213FF91FFD89092F8500D1C64C7" bold="true" box="[199,315,1974,1998]" pageId="8" pageNumber="75">Remarks.</emphasis>
As previously mentioned,
<taxonomicName id="4C195582FF91FFD892D8F8530FAA64C4" box="[653,909,1973,1997]" class="Malacostraca" family="Diogenidae" genus="Areopaguristes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="75" phylum="Arthropoda" rank="species" species="tudgei" status="sp. nov.">
<emphasis id="B96DF213FF91FFD892D8F8530FAA64C4" box="[653,909,1973,1997]" italics="true" pageId="8" pageNumber="75">Areopaguristes tudgei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25E4F68FF91FFD893C8F8500FC564C4" box="[925,994,1974,1997]" pageId="8" pageNumber="75" rank="species">n. sp.</taxonomicNameLabel>
might potentially be confused with
<taxonomicName id="4C195582FF91FFD890C2F83D0E1764F8" authority="Schmitt, 1933" authorityName="Schmitt" authorityYear="1933" box="[151,560,2009,2034]" class="Malacostraca" family="Diogenidae" genus="Paguristes" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="75" phylum="Arthropoda" rank="species" species="tortugae">
<emphasis id="B96DF213FF91FFD890C2F83D0D5C64FB" box="[151,379,2011,2034]" italics="true" pageId="8" pageNumber="75">Paguristes tortugae</emphasis>
<bibRefCitation id="EF8853F0FF91FFD891DCF83F0E1764F8" author="Schmitt" box="[393,560,2009,2033]" pageId="8" pageNumber="75" refString="Schmitt, W. L. (1933) Four new species of decapod crustaceans from Porto Rico. American Museum Novitates, 662, 1 - 9." type="journal article" year="1933">Schmitt, 1933</bibRefCitation>
</taxonomicName>
, or suspected to represent one of the recognized but as yet undescribed representatives of the
<taxonomicName id="4C195582FF90FFD991C8FF7F0E3B63B9" box="[413,540,153,176]" class="Malacostraca" family="Diogenidae" genus="Paguristes" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="76" phylum="Arthropoda" rank="species" species="tortugae">
<emphasis id="B96DF213FF90FFD991C8FF7F0E3B63B9" box="[413,540,153,176]" italics="true" pageId="9" pageNumber="76">P. tortugae</emphasis>
</taxonomicName>
complex.
<bibRefCitation id="EF8853F0FF90FFD992C9FF71080863A6" author="McLaughlin" box="[668,1071,151,175]" pageId="9" pageNumber="76" refString="McLaughlin, P. A. &amp; Provenzano, A. J., Jr. (1974) The genus Paguristes (Crustacea: Decapoda: Diogenidae) from the western Atlantic. Part I. The Paguristes tortugae complex, with notes on variation. Bulletin of Marine Science, 24 (1), 165 - 234, figs 1 - 27." type="journal article" year="1974">McLaughlin &amp; Provenzano (1974)</bibRefCitation>
included seven species in this complex, and three more were subsequently added by
<bibRefCitation id="EF8853F0FF90FFD992AEFF5A08A063DD" author="Martinez-Iglesias" box="[763,1159,188,212]" pageId="9" pageNumber="76" refString="Martinez-Iglesias, J. C. &amp; Gomez, O. (1989) Una especie nueva del genero Paguristes (Crustacea: Decapoda: Diogenidae) en las aguas de Cuba. Poeyana, 379, 1 - 30." type="journal article" year="1989">Martínez-Iglesias &amp; Gómez (1989)</bibRefCitation>
and
<bibRefCitation id="EF8853F0FF90FFD994EBFF5A0CC263FE" author="Campos" pageId="9" pageNumber="76" refString="Campos, N. H. &amp; Sanchez, H. (1995) Los cangrejos ermitanos del genero Paguristes Dana (Anomura: Diogenidae) de la costa norte colombiana, con la descripcion de dos nuevas especies. Caldasia, 17 (82 - 85), 569 - 586." type="journal article" year="1995">Campos &amp; Sánchez (1995)</bibRefCitation>
. Of the ten species that have been assigned to the complex (
<tableCitation id="C69B1BBAFF90FFD993FAFF39082063FE" box="[943,1031,223,247]" captionStart="TABLE 1" captionStartId="2.[151,239,331,355]" captionText="TABLE 1. Checklist of species of Paguristes Dana, 1851, Areopaguristes Rahayu &amp; McLaughlin, 2010, and Pseudopaguristes McLaughlin, 2002, from the western Atlantic, including primary synonyms (after McLaughlin et al. 2010). Asterisks indicate taxa currently assigned to the Paguristes tortugae complex. Paguristes Dana, 1851" pageId="9" pageNumber="76">Table 1</tableCitation>
),
<taxonomicName id="4C195582FF90FFD99476FF07090463FE" authority="Wass, 1963" authorityName="Wass" authorityYear="1963" box="[1059,1315,223,248]" class="Malacostraca" family="Diogenidae" genus="Areopaguristes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="76" phylum="Arthropoda" rank="species" species="hewatti">
<emphasis id="B96DF213FF90FFD99476FF0708B363F1" box="[1059,1172,224,248]" italics="true" pageId="9" pageNumber="76">P. hewatti</emphasis>
<bibRefCitation id="EF8853F0FF90FFD994CAFF06090463FE" author="Wass" box="[1183,1315,223,247]" pageId="9" pageNumber="76" refString="Wass, M. L. (1963) New species of hermit crabs (Decapoda, Paguridae) from the western Atlantic. Crustaceana, 6, 133 - 157." type="journal article" year="1963">Wass, 1963</bibRefCitation>
</taxonomicName>
is now in
<emphasis id="B96DF213FF90FFD990C2FEE20D666215" box="[151,321,260,284]" italics="true" pageId="9" pageNumber="76">Aeropaguristes</emphasis>
, and
<taxonomicName id="4C195582FF90FFD991D0FEE30FEA6215" authority="McLaughlin &amp; Provenzano, 1974" authorityName="McLaughlin &amp; Provenzano" authorityYear="1974" box="[389,973,260,284]" class="Malacostraca" family="Diogenidae" genus="Pseudopaguristes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="76" phylum="Arthropoda" rank="species" species="invisisacculus">
<emphasis id="B96DF213FF90FFD991D0FEE30E666215" box="[389,577,260,284]" italics="true" pageId="9" pageNumber="76">P. invisisacculus</emphasis>
<bibRefCitation id="EF8853F0FF90FFD9921FFEE20FEA6215" author="McLaughlin" box="[586,973,260,284]" pageId="9" pageNumber="76" refString="McLaughlin, P. A. &amp; Provenzano, A. J., Jr. (1974) The genus Paguristes (Crustacea: Decapoda: Diogenidae) from the western Atlantic. Part I. The Paguristes tortugae complex, with notes on variation. Bulletin of Marine Science, 24 (1), 165 - 234, figs 1 - 27." type="journal article" year="1974">
McLaughlin &amp; Provenzano,
<quantity id="4CE183E4FF90FFD993C0FEE20FEA6215" box="[917,973,260,284]" metricMagnitude="1" metricUnit="m" metricValue="5.01396" pageId="9" pageNumber="76" unit="in" value="1974.0">1974</quantity>
</bibRefCitation>
</taxonomicName>
in
<taxonomicName id="4C195582FF90FFD993A2FEE209BB6215" authority="McLaughlin, 2002" authorityName="McLaughlin" authorityYear="2002" box="[1015,1436,260,284]" class="Malacostraca" family="Diogenidae" genus="Pseudopaguristes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="76" phylum="Arthropoda" rank="genus">
<emphasis id="B96DF213FF90FFD993A2FEE208996215" box="[1015,1214,260,284]" italics="true" pageId="9" pageNumber="76">Pseudopaguristes</emphasis>
<bibRefCitation id="EF8853F0FF90FFD9949DFEE209BB6215" author="McLaughlin" box="[1224,1436,260,284]" pageId="9" pageNumber="76" refString="McLaughlin, P. A. (2002) Pseudopaguristes, a new and aberrant genus of hermit crabs (Anomura: Paguridea: Diogenidae). Micronesica, 34 (2), 185 - 199." type="journal article" year="2002">McLaughlin, 2002</bibRefCitation>
</taxonomicName>
(see
<tableCitation id="C69B1BBAFF90FFD99099FEC10D076236" box="[204,288,295,319]" captionStart="TABLE 1" captionStartId="2.[151,239,331,355]" captionText="TABLE 1. Checklist of species of Paguristes Dana, 1851, Areopaguristes Rahayu &amp; McLaughlin, 2010, and Pseudopaguristes McLaughlin, 2002, from the western Atlantic, including primary synonyms (after McLaughlin et al. 2010). Asterisks indicate taxa currently assigned to the Paguristes tortugae complex. Paguristes Dana, 1851" pageId="9" pageNumber="76">Table 1</tableCitation>
). As in
<taxonomicName id="4C195582FF90FFD991D5FECF0DDB6249" box="[384,508,297,320]" class="Malacostraca" family="Diogenidae" genus="Paguristes" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="76" phylum="Arthropoda" rank="species" species="tortugae">
<emphasis id="B96DF213FF90FFD991D5FECF0DDB6249" box="[384,508,297,320]" italics="true" pageId="9" pageNumber="76">P. tortugae</emphasis>
</taxonomicName>
and members of the complex,
<taxonomicName id="4C195582FF90FFD99334FECE0FEF6249" box="[865,968,296,320]" class="Malacostraca" family="Diogenidae" genus="Areopaguristes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="76" phylum="Arthropoda" rank="species" species="tudgei" status="sp. nov.">
<emphasis id="B96DF213FF90FFD99334FECE0FEF6249" box="[865,968,296,320]" italics="true" pageId="9" pageNumber="76">A. tudgei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25E4F68FF90FFD99385FECE08296236" box="[976,1038,296,319]" pageId="9" pageNumber="76" rank="species">n. sp.</taxonomicNameLabel>
has dense plumose setation on the chelipeds, and at least in females, there is a fringe of plumose setae on the dorsolateral margins of the chelae. A fringe of setae is also present on the dorsal margins of the walking legs of this new species.
</paragraph>
<paragraph id="8BA62E01FF90FFD99092FE720F72618D" blockId="9.[151,1436,151,1364]" pageId="9" pageNumber="76">
Aside from the fundamental difference in number of gills between
<taxonomicName id="4C195582FF90FFD993E7FE72087B62A5" box="[946,1116,404,428]" class="Malacostraca" family="Diogenidae" genus="Areopaguristes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="76" phylum="Arthropoda" rank="genus">
<emphasis id="B96DF213FF90FFD993E7FE72087B62A5" box="[946,1116,404,428]" italics="true" pageId="9" pageNumber="76">Areopaguristes</emphasis>
</taxonomicName>
and
<taxonomicName id="4C195582FF90FFD994C7FE73092D62A5" box="[1170,1290,405,428]" class="Malacostraca" family="Diogenidae" genus="Paguristes" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="76" phylum="Arthropoda" rank="genus" sensu="stricto">
<emphasis id="B96DF213FF90FFD994C7FE73092D62A5" box="[1170,1290,405,428]" italics="true" pageId="9" pageNumber="76">Paguristes</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25E4F68FF90FFD99544FE7309BB62A5" box="[1297,1436,404,428]" pageId="9" pageNumber="76" sensu="stricto">sensu stricto</taxonomicNameLabel>
(12 pairs in the former, and
<quantity id="4CE183E4FF90FFD991B5FE510E3C62C6" box="[480,539,439,463]" metricMagnitude="-1" metricUnit="m" metricValue="3.302" pageId="9" pageNumber="76" unit="in" value="13.0">13 in</quantity>
the latter), there are other important differences between
<taxonomicName id="4C195582FF90FFD994ECFE5E090662D9" box="[1209,1313,440,464]" class="Malacostraca" family="Diogenidae" genus="Areopaguristes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="76" phylum="Arthropoda" rank="species" species="tudgei" status="sp. nov.">
<emphasis id="B96DF213FF90FFD994ECFE5E090662D9" box="[1209,1313,440,464]" italics="true" pageId="9" pageNumber="76">A. tudgei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25E4F68FF90FFD9957EFE5E094D62C6" box="[1323,1386,440,463]" pageId="9" pageNumber="76" rank="species">n. sp.</taxonomicNameLabel>
and species assigned to the
<taxonomicName id="4C195582FF90FFD991CAFE3B0E3C62FD" box="[415,539,477,500]" class="Malacostraca" family="Diogenidae" genus="Paguristes" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="76" phylum="Arthropoda" rank="species" species="tortugae">
<emphasis id="B96DF213FF90FFD991CAFE3B0E3C62FD" box="[415,539,477,500]" italics="true" pageId="9" pageNumber="76">P. tortugae</emphasis>
</taxonomicName>
complex that should prevent any confusion, such as the armature of the telson (terminal margins with weak blunt spines or unarmed in
<taxonomicName id="4C195582FF90FFD99349FDE60FA46111" box="[796,899,512,536]" class="Malacostraca" family="Diogenidae" genus="Areopaguristes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="76" phylum="Arthropoda" rank="species" species="tudgei">
<emphasis id="B96DF213FF90FFD99349FDE60FA46111" box="[796,899,512,536]" italics="true" pageId="9" pageNumber="76">A. tudgei</emphasis>
</taxonomicName>
vs. armed with strong spines in the
<taxonomicName id="4C195582FF90FFD99575FDE709BB6111" box="[1312,1436,513,536]" class="Malacostraca" family="Diogenidae" genus="Paguristes" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="76" phylum="Arthropoda" rank="species" species="tortugae">
<emphasis id="B96DF213FF90FFD99575FDE709BB6111" box="[1312,1436,513,536]" italics="true" pageId="9" pageNumber="76">P. tortugae</emphasis>
</taxonomicName>
complex), the male first pleopod (distal margin of inferior lamellae unarmed in
<taxonomicName id="4C195582FF90FFD99462FDC208B86135" box="[1079,1183,548,572]" class="Malacostraca" family="Diogenidae" genus="Areopaguristes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="76" phylum="Arthropoda" rank="species" species="tudgei">
<emphasis id="B96DF213FF90FFD99462FDC208B86135" box="[1079,1183,548,572]" italics="true" pageId="9" pageNumber="76">A. tudgei</emphasis>
</taxonomicName>
vs. armed with small hooked spines in
<taxonomicName id="4C195582FF90FFD9910FFDAF0DF36169" box="[346,468,585,608]" class="Malacostraca" family="Diogenidae" genus="Paguristes" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="76" phylum="Arthropoda" rank="species" species="tortugae">
<emphasis id="B96DF213FF90FFD9910FFDAF0DF36169" box="[346,468,585,608]" italics="true" pageId="9" pageNumber="76">P. tortugae</emphasis>
</taxonomicName>
complex), and the presence or absence of brood pouch (absent in
<taxonomicName id="4C195582FF90FFD994EDFDAE093A6169" box="[1208,1309,584,608]" class="Malacostraca" family="Diogenidae" genus="Areopaguristes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="76" phylum="Arthropoda" rank="species" species="tudgei">
<emphasis id="B96DF213FF90FFD994EDFDAE093A6169" box="[1208,1309,584,608]" italics="true" pageId="9" pageNumber="76">A. tudgei</emphasis>
</taxonomicName>
vs. present in the
<taxonomicName id="4C195582FF90FFD99088FD8B0D70618D" box="[221,343,621,644]" class="Malacostraca" family="Diogenidae" genus="Paguristes" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="76" phylum="Arthropoda" rank="species" species="tortugae">
<emphasis id="B96DF213FF90FFD99088FD8B0D70618D" box="[221,343,621,644]" italics="true" pageId="9" pageNumber="76">P. tortugae</emphasis>
</taxonomicName>
complex except
<taxonomicName id="4C195582FF90FFD99240FD8B0F6F618D" authority="Bouvier, 1918" authorityName="Bouvier" authorityYear="1918" box="[533,840,620,644]" class="Malacostraca" family="Diogenidae" genus="Paguristes" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="76" phylum="Arthropoda" rank="species" species="anomalus">
<emphasis id="B96DF213FF90FFD99240FD8B0EB9618D" box="[533,670,620,644]" italics="true" pageId="9" pageNumber="76">P. anomalus</emphasis>
<bibRefCitation id="EF8853F0FF90FFD992F0FD8A0F6F618D" author="Bouvier" box="[677,840,620,644]" pageId="9" pageNumber="76" refString="Bouvier, E. - L. (1918) Sur une petite collection de Crustaces de Cuba offerte au Museum par M. de Boury. Bulletin du Museum national d'Histoire naturelle, 24, 6 - 15." type="journal article" year="1918">Bouvier, 1918</bibRefCitation>
</taxonomicName>
).
</paragraph>
<paragraph id="8BA62E01FF90FFD99092FD690FAE665D" blockId="9.[151,1436,151,1364]" pageId="9" pageNumber="76">
There are marked morphological differences among the four western Atlantic species that have so far been assigned to
<taxonomicName id="4C195582FF90FFD99172FD520DF661C5" box="[295,465,692,716]" class="Malacostraca" family="Diogenidae" genus="Areopaguristes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="76" phylum="Arthropoda" rank="genus">
<emphasis id="B96DF213FF90FFD99172FD520DF661C5" box="[295,465,692,716]" italics="true" pageId="9" pageNumber="76">Areopaguristes</emphasis>
</taxonomicName>
, and allow for their clear separation, particularly if specimens of both sexes are available.
<taxonomicName id="4C195582FF90FFD9915FFD3E0E8461E6" authority="Wass, 1963" authorityName="Wass" authorityYear="1963" box="[266,675,727,752]" class="Malacostraca" family="Diogenidae" genus="Areopaguristes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="76" phylum="Arthropoda" rank="species" species="hummi">
<emphasis id="B96DF213FF90FFD9915FFD3E0E2061F9" box="[266,519,728,752]" italics="true" pageId="9" pageNumber="76">Areopaguristes hummi</emphasis>
(
<bibRefCitation id="EF8853F0FF90FFD99242FD310EBD61E6" author="Wass" box="[535,666,727,751]" pageId="9" pageNumber="76" refString="Wass, M. L. (1963) New species of hermit crabs (Decapoda, Paguridae) from the western Atlantic. Crustaceana, 6, 133 - 157." type="journal article" year="1963">Wass, 1963</bibRefCitation>
)
</taxonomicName>
can be immediately differentiated by the adjacent ocular scales and short rostrum, whereas in the other three western Atlantic congeners the ocular scales are separated and the rostrum extends forwardly between the scales.
<taxonomicName id="4C195582FF90FFD99219FCC60F046031" box="[588,803,800,824]" class="Malacostraca" family="Diogenidae" genus="Areopaguristes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="76" phylum="Arthropoda" rank="species" species="iris">
<emphasis id="B96DF213FF90FFD99219FCC60F046031" box="[588,803,800,824]" italics="true" pageId="9" pageNumber="76">Areopaguristes iris</emphasis>
</taxonomicName>
(Forest &amp; de
<bibRefCitation id="EF8853F0FF90FFD99396FCF90883603E" author="Saint" box="[963,1188,799,824]" pageId="9" pageNumber="76" refString="Forest, J. &amp; de Saint Laurent, M. (1968) Resultats scientifiques des campagnes de la &quot; Calypso, Part VII. Campagne de la Calypso au large des cotes Atlantiques de l'Amerique du Sud (1961 - 1962). 6. Crustaces Dcapodes: Pagurides. Annales de l'Institut Oceanographique de Monaco, n. s., 45 (2), 45 - 172." type="journal article" year="1968">Saint Laurent, 1968</bibRefCitation>
) is distinguishable by the slender walking legs, in particular the dactyls which can be 10 times as long as their basal height, whereas in other congeners from the region the dactyls are at most six times as long as high.
<taxonomicName id="4C195582FF90FFD99416FC8E0CFF60AD" authority="Wass, 1963" authorityName="Wass" authorityYear="1963" class="Malacostraca" family="Diogenidae" genus="Areopaguristes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="76" phylum="Arthropoda" rank="species" species="hewatti">
<emphasis id="B96DF213FF90FFD99416FC8E096F6089" box="[1091,1352,872,896]" italics="true" pageId="9" pageNumber="76">Areopaguristes hewatti</emphasis>
(
<bibRefCitation id="EF8853F0FF90FFD9950CFC810CF760AD" author="Wass" pageId="9" pageNumber="76" refString="Wass, M. L. (1963) New species of hermit crabs (Decapoda, Paguridae) from the western Atlantic. Crustaceana, 6, 133 - 157." type="journal article" year="1963">Wass, 1963</bibRefCitation>
)
</taxonomicName>
can be separated by the shape and armature of the chelipeds. In
<taxonomicName id="4C195582FF90FFD993E1FC6A080160AD" box="[948,1062,908,932]" class="Malacostraca" family="Diogenidae" genus="Areopaguristes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="76" phylum="Arthropoda" rank="species" species="hewatti">
<emphasis id="B96DF213FF90FFD993E1FC6A080160AD" box="[948,1062,908,932]" italics="true" pageId="9" pageNumber="76">A. hewatti</emphasis>
</taxonomicName>
the dorsolateral and dorsomesial margins of the carpi and palm, and the distomedial region of the carpi, are distinctly elevated; and the dorsomedian armature of the carpi and chela consists of many low, mostly blunt spines or tubercles. Also,
<taxonomicName id="4C195582FF90FFD994F8FC32090760E5" box="[1197,1312,980,1004]" class="Malacostraca" family="Diogenidae" genus="Areopaguristes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="76" phylum="Arthropoda" rank="species" species="hewatti">
<emphasis id="B96DF213FF90FFD994F8FC32090760E5" box="[1197,1312,980,1004]" italics="true" pageId="9" pageNumber="76">A. hewatti</emphasis>
</taxonomicName>
is the only western Atlantic
<taxonomicName id="4C195582FF90FFD9910DFC1E0E256719" box="[344,514,1016,1040]" class="Malacostraca" family="Diogenidae" genus="Areopaguristes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="76" phylum="Arthropoda" rank="genus">
<emphasis id="B96DF213FF90FFD9910DFC1E0E256719" box="[344,514,1016,1040]" italics="true" pageId="9" pageNumber="76">Areopaguristes</emphasis>
</taxonomicName>
species in which females have a brood pouch, although presence or absence of a brood pouch needs to be confirmed for
<taxonomicName id="4C195582FF90FFD99232FBFA0E89673D" box="[615,686,1052,1076]" class="Malacostraca" family="Diogenidae" genus="Areopaguristes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="76" phylum="Arthropoda" rank="species" species="iris">
<emphasis id="B96DF213FF90FFD99232FBFA0E89673D" box="[615,686,1052,1076]" italics="true" pageId="9" pageNumber="76">A. iris</emphasis>
</taxonomicName>
when females of that species are discovered as
<taxonomicName id="4C195582FF90FFD994B4FBFA090D673D" box="[1249,1322,1052,1076]" class="Malacostraca" family="Diogenidae" genus="Areopaguristes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="76" phylum="Arthropoda" rank="species" species="iris">
<emphasis id="B96DF213FF90FFD994B4FBFA090D673D" box="[1249,1322,1052,1076]" italics="true" pageId="9" pageNumber="76">A. iris</emphasis>
</taxonomicName>
is known exclusively from the
<typeStatus id="54A290A3FF90FFD991C4FBA70DE66750" box="[401,449,1089,1113]" pageId="9" pageNumber="76">type</typeStatus>
material consisting of four males (male
<typeStatus id="54A290A3FF90FFD993F3FBA6082C6751" box="[934,1035,1088,1112]" pageId="9" pageNumber="76" type="holotype">holotype</typeStatus>
, MNHN Pg 2377, and three male
<typeStatus id="54A290A3FF90FFD990C2FB830D226774" box="[151,261,1125,1149]" pageId="9" pageNumber="76" type="paratype">paratypes</typeStatus>
, MNHN Pg 2378) used by
<bibRefCitation id="EF8853F0FF90FFD99215FB830FBA6775" box="[576,925,1124,1148]" pageId="9" pageNumber="76" refString="Forest, J. &amp; de Saint Laurent, M. (1968) Resultats scientifiques des campagnes de la &quot; Calypso, Part VII. Campagne de la Calypso au large des cotes Atlantiques de l'Amerique du Sud (1961 - 1962). 6. Crustaces Dcapodes: Pagurides. Annales de l'Institut Oceanographique de Monaco, n. s., 45 (2), 45 - 172." type="journal article">Forest &amp; Saint Laurent (1968)</bibRefCitation>
. The new species,
<taxonomicName id="4C195582FF90FFD99420FB8208FB6775" box="[1141,1244,1124,1148]" class="Malacostraca" family="Diogenidae" genus="Areopaguristes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="76" phylum="Arthropoda" rank="species" species="tudgei">
<emphasis id="B96DF213FF90FFD99420FB8208FB6775" box="[1141,1244,1124,1148]" italics="true" pageId="9" pageNumber="76">A. tudgei</emphasis>
</taxonomicName>
, is distinguished from all other congeners from the region primarily by the weakly armed or unarmed, terminal margins of the telson (
<figureCitation id="13223284FF90FFD990CAFB4A0CC567CD" box="[159,226,1196,1220]" captionStart="FIGURE 1" captionStartId="3.[151,250,1589,1612]" captionTargetBox="[166,1428,367,1565]" captionTargetId="figure@3.[151,1436,353,1573]" captionTargetPageId="3" captionText="FIGURE 1. Areopaguristes tudgei n. sp., Carrie Bow Cay, Belize: a, c – e, h, male paratype, 2.1 mm ,, February 8 (USNM 1156993); b, f, g, i, female paratype, 2.1 mm, February 4 (USNM 1156991). a, shield and cephalic appendages, dorsal view; b, anterior portion of shield, dorsal view; c, male left first pleopod, posterior view; d, left second pleopod, posterior view; e, male left third pleopod, posterior view; f, female left first pleopod, posterior view; g, female left third pleopod, posterior view; h, uropods and telson, dorsal view; i, telson, dorsal view. Scales equal 0.5 mm (a, b, e, h, i), 0.2 mm (c, d, f, g)." httpUri="https://zenodo.org/record/214046/files/figure.png" pageId="9" pageNumber="76">
Fig.
<date id="FFA708C1FF90FFD99083FB4A0CC567CD" box="[214,226,1196,1220]" pageId="9" pageNumber="76">1</date>
</figureCitation>
i), all others having strong spines; and males having first pleopods with the distal margin of the inferior lamella entire, unarmed (
<figureCitation id="13223284FF90FFD991E7FB360DD567E1" box="[434,498,1232,1256]" captionStart="FIGURE 1" captionStartId="3.[151,250,1589,1612]" captionTargetBox="[166,1428,367,1565]" captionTargetId="figure@3.[151,1436,353,1573]" captionTargetPageId="3" captionText="FIGURE 1. Areopaguristes tudgei n. sp., Carrie Bow Cay, Belize: a, c – e, h, male paratype, 2.1 mm ,, February 8 (USNM 1156993); b, f, g, i, female paratype, 2.1 mm, February 4 (USNM 1156991). a, shield and cephalic appendages, dorsal view; b, anterior portion of shield, dorsal view; c, male left first pleopod, posterior view; d, left second pleopod, posterior view; e, male left third pleopod, posterior view; f, female left first pleopod, posterior view; g, female left third pleopod, posterior view; h, uropods and telson, dorsal view; i, telson, dorsal view. Scales equal 0.5 mm (a, b, e, h, i), 0.2 mm (c, d, f, g)." httpUri="https://zenodo.org/record/214046/files/figure.png" pageId="9" pageNumber="76">Fig. 1</figureCitation>
c), all others having minute hooked spines. There are also differences in coloration among the four western species of
<taxonomicName id="4C195582FF90FFD9926EFB120EC26605" box="[571,741,1268,1292]" class="Malacostraca" family="Diogenidae" genus="Areopaguristes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="76" phylum="Arthropoda" rank="genus">
<emphasis id="B96DF213FF90FFD9926EFB120EC26605" box="[571,741,1268,1292]" italics="true" pageId="9" pageNumber="76">Areopaguristes</emphasis>
</taxonomicName>
; however, a detailed comparison of all four species is not possible because Forest &amp; de Saint Laurent’s (1968: 84) description of coloration in
<taxonomicName id="4C195582FF90FFD994C6FAFE08FA6639" box="[1171,1245,1304,1328]" class="Malacostraca" family="Diogenidae" genus="Areopaguristes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="76" phylum="Arthropoda" rank="species" species="iris">
<emphasis id="B96DF213FF90FFD994C6FAFE08FA6639" box="[1171,1245,1304,1328]" italics="true" pageId="9" pageNumber="76">A. iris</emphasis>
</taxonomicName>
was very brief, indicating only the presence of iridescence in preserved specimens.
</paragraph>
</subSubSection>
</treatment>
</document>
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