treatments-xml/data/03/A0/37/03A03716FFF9AF08FE86FBF626F62CC3.xml

<document ID-DOI="http://doi.org/10.5281/zenodo.4763863" ID-GBIF-Dataset="7e69e5d0-b481-4034-bc40-7cd95db92d59" ID-GBIF-Taxon="182075045" ID-ISSN="1854-0392" ID-Zenodo-Dep="4763863" ID-ZooBank="66C4E575-ABF5-4B81-9132-0D771B52D68" approvalRequired="1" approvalRequired_for_treatments="1" checkinTime="1620972287440" checkinUser="felipe" docAuthor="Grubbs, Scott A., Baumann, Richard W. &amp; Burton, David K." docDate="2018" docId="03A03716FFF9AF08FE86FBF626F62CC3" docLanguage="en" docName="Illiesia.14.3.30-43.pdf" docOrigin="Illiesia 14 (3)" docStyle="DocumentStyle:3ED4ECBC8873166DC296BE619562C54F.3:Illiesia.2009-.journal_article" docStyleId="3ED4ECBC8873166DC296BE619562C54F" docStyleName="Illiesia.2009-.journal_article" docStyleVersion="3" docTitle="Nemoura arctica Esben-Petersen 1910" docType="treatment" docVersion="3" lastPageId="8" lastPageNumber="52" masterDocId="FF994F6EFFFDAF00FF8DFFEF22272F0F" masterDocTitle="Nearctic Nemoura Trispinosa Claassen, 1923 And N. Rickeri Jewett, 1971 Are Junior Synonyms Of Holarctic Nemoura Species (Plecoptera: Nemouridae)" masterLastPageNumber="64" masterPageNumber="44" pageId="4" pageNumber="48" updateTime="1643473119139" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Nearctic Nemoura Trispinosa Claassen, 1923 And N. Rickeri Jewett, 1971 Are Junior Synonyms Of Holarctic Nemoura Species (Plecoptera: Nemouridae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Grubbs, Scott A.</mods:namePart>
<mods:affiliation>, Richard W. Baumann &amp; Department of Biology and Center for Biodiversity Studies, Western Kentucky University, Bowling Green, Kentucky 42101, U. S. A. E-mail: scott. grubbs @ wku. edu</mods:affiliation>
<mods:nameIdentifier type="email">scott.grubbs@wku.edu</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Baumann, Richard W.</mods:namePart>
<mods:affiliation>, and David K. Burton &amp; Department of Biology, Monte L. Bean Life Science Museum, Brigham Young University, Provo, Utah, U. S. A., 84602. E-mail: richard _ baumann @ byu. edu</mods:affiliation>
<mods:nameIdentifier type="email">richard_baumann@byu.edu</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Burton, David K.</mods:namePart>
<mods:affiliation>Faculty of Education, University of Ottawa, Ottawa, Ontario, Canada K 1 N 6 N 5 E-mail: dburton @ uottawa. ca &amp; Canadian National Collection, Agriculture Canada, Central Experimental Farm, Ottawa, Ontario, Canada K 1 A 0 C 6</mods:affiliation>
<mods:nameIdentifier type="email">dburton@uottawa.ca</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Illiesia</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2018</mods:date>
<mods:detail type="volume">
<mods:number>14</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>3</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>44</mods:start>
<mods:end>64</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://doi.org/10.5281/zenodo.4761204</mods:identifier>
<mods:identifier type="GBIF-Dataset">7e69e5d0-b481-4034-bc40-7cd95db92d59</mods:identifier>
<mods:identifier type="ISSN">1854-0392</mods:identifier>
<mods:identifier type="Zenodo-Dep">4761204</mods:identifier>
<mods:identifier type="ZooBank">66C4E575-ABF5-4B81-9132-0D771B52D68</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.4763863" ID-GBIF-Taxon="182075045" ID-Zenodo-Dep="4763863" LSID="urn:lsid:plazi:treatment:03A03716FFF9AF08FE86FBF626F62CC3" httpUri="http://treatment.plazi.org/id/03A03716FFF9AF08FE86FBF626F62CC3" lastPageId="8" lastPageNumber="52" pageId="4" pageNumber="48">
<subSubSection box="[267,720,1048,1073]" pageId="4" pageNumber="48" type="nomenclature">
<paragraph blockId="4.[194,793,1048,1210]" box="[267,720,1048,1073]" pageId="4" pageNumber="48">
<taxonomicName authority="Esben-Petersen, 1910 Arctic Forestfly" authorityName="Esben-Petersen" authorityYear="1910" box="[267,720,1048,1073]" class="Insecta" family="Nemouridae" genus="Nemoura" kingdom="Animalia" order="Plecoptera" pageId="4" pageNumber="48" phylum="Arthropoda" rank="species" species="arctica">
<emphasis bold="true" box="[267,466,1048,1072]" italics="true" pageId="4" pageNumber="48">Nemoura arctica</emphasis>
<bibRefCitation author="Esben-Petersen, P." box="[472,720,1048,1073]" pageId="4" pageNumber="48" pagination="75 - 89" refId="ref6669" refString="Esben-Petersen, P. 1910. Bidrag til en fortegnelse over arktisk. Norges Neuropterfauna. II. Tromso Museums Arshefter, 31 / 32: 75 - 89." type="journal article" year="1910">Esben-Petersen, 1910</bibRefCitation>
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="4" pageNumber="48" type="description">
<paragraph blockId="4.[194,793,1048,1210]" pageId="4" pageNumber="48">
Arctic Forestfly (
<figureCitation box="[435,553,1120,1144]" captionStart-0="Figs" captionStart-1="Figs" captionStart-2="Figs" captionStart-3="Figs" captionStart-4="Figs" captionStart-5="Figs" captionStart-6="Figs" captionStartId-0="2.[189,238,1788,1812]" captionStartId-1="3.[189,238,1784,1808]" captionStartId-2="5.[189,238,1746,1770]" captionStartId-3="6.[189,238,1748,1772]" captionStartId-4="7.[189,238,1748,1772]" captionStartId-5="9.[189,238,1409,1433]" captionStartId-6="10.[189,238,1416,1440]" captionTargetBox-0="[324,1307,254,1740]" captionTargetBox-1="[324,1307,254,1737]" captionTargetBox-2="[324,1307,218,1700]" captionTargetBox-3="[324,1307,218,1701]" captionTargetBox-4="[324,1307,218,1701]" captionTargetBox-5="[324,1307,254,1362]" captionTargetBox-6="[324,1307,254,1368]" captionTargetId-0="figure-125@2.[324,1307,254,1740]" captionTargetId-1="figure-116@3.[324,1307,254,1737]" captionTargetId-2="figure-138@5.[324,1307,218,1700]" captionTargetId-3="figure-149@6.[324,1307,218,1701]" captionTargetId-4="figure-137@7.[324,1307,218,1701]" captionTargetId-5="figure-247@9.[324,1307,254,1362]" captionTargetId-6="figure-236@10.[324,1307,254,1368]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="5" captionTargetPageId-3="6" captionTargetPageId-4="7" captionTargetPageId-5="9" captionTargetPageId-6="10" captionText-0="Figs. 1–8. Nemoura arctica, cerci, distal view. 1, USA, Iowa, Twin Springs, 545X; 2, USA, Illinois, Elgin, 516X; 3, USA, Ohio, John Bryan State Park, 518X; 4, USA, Wisconsin, spring, 1 mile N Irvington, 509X; 5, USA, South Dakota, Burnt Fork, 497X; 6, USA, Wyoming, Blacktail Creek, 584X; 7, Canada, Saskatchewan, Promontory Campground, 605X; 8, Canada, New Brunswick, Coombs Brook, 591X." captionText-1="Figs. 9–16. Nemoura arctica, cerci, distal view. 9, Canada, Manitoba, Warkworth Creek, 773X; 10, Canada, Northwest Territories, Thelon River, 636X; 11, Canada, Northwest Territories, Tuktoyaktuk, 612X; 12, Russia, Siberia, Magadan Oblask, 486X; 13, Norway, Buktved Garnvika, 443X; 14, Norway, Kautokeino, 431X; 15, Mongolia, Khoton Lake, 675X; 16, Mongolia, Tsagaaan Lake, 646X." captionText-2="Figs. 17–24. Nemoura arctica, epiproct, lateral view. 17, USA, Iowa, Twin Springs, 270X; 18, Canada, New Brunswick, Coombs Brook, 288X; 19, Canada, Northwest Territories, Thelon River, 285X; 20, Canada, Northwest Territories, Tuktoyaktuk, 328X; 21, Russia, Siberia, Magadan Oblask, 271X; 22, Norway, Buktved Garnvika, 286X; 23, Norway, Kautokeino, 316X; 24, Mongolia, Khoton Lake, 276X. AP = apical prong, BC = basal cushion, DS = dorsal sclerite, LA = lateral areas, VS = ventral sclerite." captionText-3="Figs. 25–32. Nemoura arctica, epiproct, dorsal view. 25, USA, Iowa, Twin Springs, 141X; 26, USA, Illinois, Elgin, 165X; 27, USA, Ohio, spring, John Bryan State Park, 144X; 28, USA, Wisconsin, spring, 1 mile N Irvington, 132X; 29, USA, South Dakota, Burnt Fork, 140X; 30, USA, South Dakota, Burnt Fork, 229X; 31, Canada, Prince Edward Island, Balsam Hollow, 163X; 32, Canada, New Brunswick, Coombs Brook, 168X. AP = apical prong, BC = basal cushion, DS = dorsal sclerite, LA = lateral area, VS = ventral sclerite." captionText-4="Figs. 33–40. Nemoura arctica, epiproct, dorsal view. 33, Canada, Saskatchewan, Promontory Campground, 187X; 34, Canada, Northwest Territories, Thelon River, 282X; 35, Canada, Northwest Territories, Tuktoyaktuk, 267X; 36, Norway, Buktved Garnvika, 321X; 37, Norway, Kautokeino, 278X; 38, Russia, Siberia, Magadan Oblask, 206X; 39, Mongolia, Khoton Lake, 277X; 40, Mongolia, Tsagaaan Lake, 281X. AP = apical prong, BC = basal cushion, DS = dorsal sclerite, LA = lateral area, VS = ventral sclerite." captionText-5="Figs. 41–46. Nemoura arctica, epiproct, dorsal view. 41, USA, Iowa, Twin Springs, 584X; 42, USA, Illinois, Elgin, 519X; 43, USA, South Dakota, Burnt Fork, 669X; 44, Canada, Saskatchewan, Promontory Campground, 1056X; 45, Canada, New Brunswick, Coombs Brook, 770X; 46, Canada, Prince Edward Island, Balsam Hollow, 695X. AP = apical prong." captionText-6="Figs. 47–52. Nemoura arctica, epiproct, dorsal view. 47, Canada, Northwest Territories, Thelon River, 1161X; 48, Russia, Siberia, Magadan Oblask, 1204X; 49, Norway, Buktved Garnvika, 751X; 50, Norway, Kautokeino, 809X; 51, Mongolia, Khoton Lake, 598X; 52, Mongolia, Tsagaaan Lake, 969X. AP = apical prong." figureDoi-0="http://doi.org/10.5281/zenodo.4761206" figureDoi-1="http://doi.org/10.5281/zenodo.4761208" figureDoi-2="http://doi.org/10.5281/zenodo.4761210" figureDoi-3="http://doi.org/10.5281/zenodo.4761214" figureDoi-4="http://doi.org/10.5281/zenodo.4761216" figureDoi-5="http://doi.org/10.5281/zenodo.4761218" figureDoi-6="http://doi.org/10.5281/zenodo.4761220" httpUri-0="https://zenodo.org/record/4761206/files/figure.png" httpUri-1="https://zenodo.org/record/4761208/files/figure.png" httpUri-2="https://zenodo.org/record/4761210/files/figure.png" httpUri-3="https://zenodo.org/record/4761214/files/figure.png" httpUri-4="https://zenodo.org/record/4761216/files/figure.png" httpUri-5="https://zenodo.org/record/4761218/files/figure.png" httpUri-6="https://zenodo.org/record/4761220/files/figure.png" pageId="4" pageNumber="48">Figs. 1–52</figureCitation>
)
</paragraph>
<paragraph blockId="4.[194,793,1048,1210]" pageId="4" pageNumber="48">
http://lsid.speciesfile.org/urn:lsid: 
<taxonomicName box="[527,639,1155,1177]" class="Insecta" kingdom="Animalia" order="Plecoptera" pageId="4" pageNumber="48" phylum="Arthropoda" rank="order">Plecoptera</taxonomicName>
.speciesfile.org: TaxonName:6232
</paragraph>
</subSubSection>
<subSubSection box="[227,656,1252,1274]" pageId="4" pageNumber="48" type="reference_group">
<paragraph blockId="4.[227,797,1252,1922]" box="[227,656,1252,1274]" pageId="4" pageNumber="48">
<treatmentCitationGroup box="[227,652,1252,1274]" pageId="4" pageNumber="48">
<treatmentCitation box="[227,652,1252,1274]" pageId="4" pageNumber="48">
<taxonomicName authority="Esben-Petersen 1910: 85" authorityName="Esben-Petersen" authorityPageNumber="85" authorityYear="1910" box="[227,652,1252,1274]" class="Insecta" family="Nemouridae" genus="Nemoura" kingdom="Animalia" order="Plecoptera" pageId="4" pageNumber="48" phylum="Arthropoda" rank="species" species="arctica">
<emphasis box="[227,391,1253,1274]" italics="true" pageId="4" pageNumber="48">Nemoura arctica</emphasis>
<bibRefCitation author="Esben-Petersen, P." box="[400,652,1252,1274]" pageId="4" pageNumber="48" pagination="75 - 89" refId="ref6669" refString="Esben-Petersen, P. 1910. Bidrag til en fortegnelse over arktisk. Norges Neuropterfauna. II. Tromso Museums Arshefter, 31 / 32: 75 - 89." type="journal article" year="1910">Esben-Petersen 1910:85</bibRefCitation>
</taxonomicName>
</treatmentCitation>
</treatmentCitationGroup>
.
</paragraph>
</subSubSection>
<subSubSection pageId="4" pageNumber="48" type="materials_examined">
<paragraph blockId="4.[227,797,1252,1922]" pageId="4" pageNumber="48">
<materialsCitation ID-GBIF-Occurrence="3111722308" country="Norway" location="Karasjok" pageId="4" pageNumber="48" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<typeStatus box="[664,764,1252,1274]" pageId="4" pageNumber="48">Holotype</typeStatus>
<specimenCount box="[773,791,1255,1273]" pageId="4" pageNumber="48" type="male">♂</specimenCount>
, Type locality – 
<location LSID="urn:lsid:plazi:treatment:03A03716FFF9AF08FE86FBF626F62CC3:8ED6D0DBFFF9AF04FE08FAEA23C32A14" box="[389,484,1285,1307]" country="Norway" name="Karasjok" pageId="4" pageNumber="48">Karasjok</location>
, 
<collectingCountry box="[494,582,1285,1306]" name="Norway" pageId="4" pageNumber="48">Norway</collectingCountry>
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection box="[227,646,1317,1339]" pageId="4" pageNumber="48" type="reference_group">
<paragraph blockId="4.[227,797,1252,1922]" box="[227,646,1317,1339]" pageId="4" pageNumber="48">
<treatmentCitationGroup box="[227,646,1317,1339]" pageId="4" pageNumber="48">
<treatmentCitation box="[227,646,1317,1339]" pageId="4" pageNumber="48">
<taxonomicName authority="Claassen 1923: 289" authorityName="Claassen" authorityPageNumber="289" authorityYear="1923" box="[227,643,1317,1339]" class="Insecta" family="Nemouridae" genus="Nemoura" kingdom="Animalia" order="Plecoptera" pageId="4" pageNumber="48" phylum="Arthropoda" rank="species" species="trispinosa">
<emphasis box="[227,427,1318,1339]" italics="true" pageId="4" pageNumber="48">Nemoura trispinosa</emphasis>
<bibRefCitation author="Claassen, P. W." box="[441,643,1317,1339]" pageId="4" pageNumber="48" pagination="281 - 292" refId="ref6415" refString="Claassen, P. W. 1923. New species of North American Plecoptera, Part II. The Canadian Entomologist, 55: 281 - 292." type="journal article" year="1923">Claassen 1923:289</bibRefCitation>
</taxonomicName>
.
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection pageId="4" pageNumber="48" type="materials_examined">
<paragraph blockId="4.[227,797,1252,1922]" pageId="4" pageNumber="48">
<materialsCitation ID-GBIF-Occurrence="3111722303" county="Tompkins Co." location="Tompkins Co." pageId="4" pageNumber="48" specimenCount="1" specimenCount-male="1" stateProvince="New York" typeStatus="holotype">
<typeStatus box="[659,759,1317,1339]" pageId="4" pageNumber="48">Holotype</typeStatus>
<specimenCount box="[773,791,1320,1338]" pageId="4" pageNumber="48" type="male">♂</specimenCount>
, Type locality – Mud Creek, 
<collectingCounty box="[566,727,1350,1372]" pageId="4" pageNumber="48">Tompkins Co.</collectingCounty>
, 
<collectingRegion country="United States of America" name="New York" pageId="4" pageNumber="48">New York</collectingRegion>
. 
<emphasis bold="true" box="[291,451,1382,1403]" pageId="4" pageNumber="48">New synonym</emphasis>
(Holotype ♂ examined)
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="4" pageNumber="48" type="reference_group">
<paragraph blockId="4.[227,797,1252,1922]" pageId="4" pageNumber="48">
<treatmentCitationGroup box="[227,763,1414,1436]" pageId="4" pageNumber="48">
<treatmentCitation box="[227,763,1414,1436]" pageId="4" pageNumber="48">
<taxonomicName authority=": Needham &amp; Claassen 1925: 213" authorityName="Needham &amp; Claassen" authorityPageNumber="213" authorityYear="1925" box="[227,757,1414,1436]" class="Insecta" family="Nemouridae" genus="Nemoura" kingdom="Animalia" order="Plecoptera" pageId="4" pageNumber="48" phylum="Arthropoda" rank="species" species="trispinosa">
<emphasis box="[227,419,1415,1436]" italics="true" pageId="4" pageNumber="48">Nemoura trispinosa</emphasis>
: 
<bibRefCitation author="Needham, J. H. &amp; P. W. Claassen" box="[431,757,1414,1436]" pageId="4" pageNumber="48" pagination="1 - 397" refId="ref7781" refString="Needham, J. H. &amp; P. W. Claassen. 1925. A monograph of the Plecoptera or stoneflies of America north of Mexico. Thomas Say Foundation, Entomological Society of America, 2: 1 - 397." type="journal article" year="1925">Needham &amp; Claassen 1925:213</bibRefCitation>
</taxonomicName>
.
</treatmentCitation>
</treatmentCitationGroup>
<treatmentCitationGroup box="[227,585,1447,1469]" pageId="4" pageNumber="48">
<treatmentCitation box="[227,585,1447,1469]" pageId="4" pageNumber="48">
<taxonomicName authority=": Claassen 1940: 50" authorityName="Claassen" authorityPageNumber="50" authorityYear="1940" box="[227,578,1447,1469]" class="Insecta" family="Nemouridae" genus="Nemoura" kingdom="Animalia" order="Plecoptera" pageId="4" pageNumber="48" phylum="Arthropoda" rank="species" species="arctica">
<emphasis box="[227,388,1448,1469]" italics="true" pageId="4" pageNumber="48">Nemoura arctica</emphasis>
: 
<bibRefCitation author="Claassen, P. W." box="[401,578,1447,1469]" pageId="4" pageNumber="48" pagination="1 - 235" refId="ref6464" refString="Claassen, P. W. 1940. A catalogue of the Plecoptera of the world. Cornell University Experimental Station Memoir, 232: 1 - 235." type="journal article" year="1940">Claassen 1940:50</bibRefCitation>
</taxonomicName>
.
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
<paragraph blockId="4.[227,797,1252,1922]" box="[227,616,1479,1501]" pageId="4" pageNumber="48">
<treatmentCitationGroup box="[227,616,1479,1501]" pageId="4" pageNumber="48">
<treatmentCitation box="[227,616,1479,1501]" pageId="4" pageNumber="48">
<taxonomicName authority=": Claassen 1940: 64" authorityName="Claassen" authorityPageNumber="64" authorityYear="1940" box="[227,609,1479,1501]" class="Insecta" family="Nemouridae" genus="Nemoura" kingdom="Animalia" order="Plecoptera" pageId="4" pageNumber="48" phylum="Arthropoda" rank="species" species="trispinosa">
<emphasis box="[227,419,1480,1501]" italics="true" pageId="4" pageNumber="48">Nemoura trispinosa</emphasis>
: 
<bibRefCitation author="Claassen, P. W." box="[431,609,1479,1501]" pageId="4" pageNumber="48" pagination="1 - 235" refId="ref6464" refString="Claassen, P. W. 1940. A catalogue of the Plecoptera of the world. Cornell University Experimental Station Memoir, 232: 1 - 235." type="journal article" year="1940">Claassen 1940:64</bibRefCitation>
</taxonomicName>
.
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
<paragraph blockId="4.[227,797,1252,1922]" box="[227,595,1511,1533]" pageId="4" pageNumber="48">
<treatmentCitationGroup box="[227,595,1511,1533]" pageId="4" pageNumber="48">
<treatmentCitation box="[227,595,1511,1533]" pageId="4" pageNumber="48">
<taxonomicName authority=": Frison 1942: 261" authorityName="Frison" authorityPageNumber="261" authorityYear="1942" box="[227,595,1511,1533]" class="Insecta" family="Nemouridae" genus="Nemoura" kingdom="Animalia" order="Plecoptera" pageId="4" pageNumber="48" phylum="Arthropoda" rank="species" species="trispinosa">
<emphasis box="[227,419,1512,1533]" italics="true" pageId="4" pageNumber="48">Nemoura trispinosa</emphasis>
: 
<bibRefCitation author="Frison, T. H." box="[431,595,1511,1533]" pageId="4" pageNumber="48" pagination="235 - 355" refId="ref6766" refString="Frison, T. H. 1942. Studies of North American Plecoptera with special reference to the fauna of Illinois. Illinois Natural History Survey Bulletin, 22: 235 - 355." type="journal article" year="1942">Frison 1942:261</bibRefCitation>
</taxonomicName>
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
<paragraph blockId="4.[227,797,1252,1922]" box="[227,676,1544,1566]" pageId="4" pageNumber="48">
<treatmentCitationGroup box="[227,676,1544,1566]" pageId="4" pageNumber="48">
<treatmentCitation box="[227,676,1544,1566]" pageId="4" pageNumber="48">
<taxonomicName authority=": Koponen &amp; Brinck 1949: 7" authorityName="Koponen &amp; Brinck" authorityPageNumber="7" authorityYear="1949" box="[227,670,1544,1566]" class="Insecta" family="Nemouridae" genus="Nemoura" kingdom="Animalia" order="Plecoptera" pageId="4" pageNumber="48" phylum="Arthropoda" rank="species" species="arctica">
<emphasis box="[227,388,1544,1566]" italics="true" pageId="4" pageNumber="48">Nemoura arctica</emphasis>
: 
<bibRefCitation author="Koponen, J. S. &amp; P. Brinck" box="[401,670,1544,1566]" pageId="4" pageNumber="48" pagination="1 - 21" refId="ref7321" refString="Koponen, J. S. &amp; P. Brinck. 1949. Neue oder wenig bekannte Plecoptera. Annales Entomologici Fennicae, 15: 1 - 21." type="journal article" year="1949">Koponen &amp; Brinck 1949:7</bibRefCitation>
</taxonomicName>
.
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
<paragraph blockId="4.[227,797,1252,1922]" box="[227,604,1576,1598]" pageId="4" pageNumber="48">
<treatmentCitationGroup box="[227,604,1576,1598]" pageId="4" pageNumber="48">
<treatmentCitation box="[227,604,1576,1598]" pageId="4" pageNumber="48">
<taxonomicName authority=": Weber 1950: 175" authorityName="Weber" authorityPageNumber="175" authorityYear="1950" box="[227,597,1576,1598]" class="Insecta" family="Nemouridae" genus="Nemoura" kingdom="Animalia" order="Plecoptera" pageId="4" pageNumber="48" phylum="Arthropoda" rank="species" species="trispinosa">
<emphasis box="[227,419,1577,1598]" italics="true" pageId="4" pageNumber="48">Nemoura trispinosa</emphasis>
: 
<bibRefCitation author="Weber, N. A." box="[431,597,1576,1598]" pageId="4" pageNumber="48" pagination="147 - 206" refId="ref8514" refString="Weber, N. A. 1950. A survey of the insects and related arthropods of Arctic Alaska, Part I. Transactions of the American Entomological Society, 76: 147 - 206. http: // www. jstor. org / stable / 25077616" type="journal article" year="1950">Weber 1950:175</bibRefCitation>
</taxonomicName>
.
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
<paragraph blockId="4.[227,797,1252,1922]" box="[227,572,1609,1631]" pageId="4" pageNumber="48">
<treatmentCitationGroup box="[227,572,1609,1631]" pageId="4" pageNumber="48">
<treatmentCitation box="[227,572,1609,1631]" pageId="4" pageNumber="48">
<taxonomicName authority=": Brinck 1952: 107" authorityName="Brinck" authorityPageNumber="107" authorityYear="1952" box="[227,565,1609,1631]" class="Insecta" family="Nemouridae" genus="Nemoura" kingdom="Animalia" order="Plecoptera" pageId="4" pageNumber="48" phylum="Arthropoda" rank="species" species="arctica">
<emphasis box="[227,388,1610,1631]" italics="true" pageId="4" pageNumber="48">Nemoura arctica</emphasis>
: 
<bibRefCitation author="Brinck, P." box="[401,565,1609,1631]" pageId="4" pageNumber="48" pagination="1 - 126" refId="ref6317" refString="Brinck, P. 1952. Backslandor, Plecoptera. Svensk Insektfauna, 15: 1 - 126." type="journal article" year="1952">Brinck 1952:107</bibRefCitation>
</taxonomicName>
.
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
<paragraph blockId="4.[227,797,1252,1922]" pageId="4" pageNumber="48">
<treatmentCitationGroup box="[227,706,1641,1663]" pageId="4" pageNumber="48">
<treatmentCitation box="[227,706,1641,1663]" pageId="4" pageNumber="48">
<taxonomicName authority=": Harden &amp; Mickel 1952: 19" authorityName="Harden &amp; Mickel" authorityPageNumber="19" authorityYear="1952" box="[227,700,1641,1663]" class="Insecta" family="Nemouridae" genus="Nemoura" kingdom="Animalia" order="Plecoptera" pageId="4" pageNumber="48" phylum="Arthropoda" rank="species" species="trispinosa">
<emphasis box="[227,419,1642,1663]" italics="true" pageId="4" pageNumber="48">Nemoura trispinosa</emphasis>
: 
<bibRefCitation author="Harden, P. H. &amp; C. E. Mickel" box="[431,700,1641,1663]" pageId="4" pageNumber="48" pagination="1 - 84" refId="ref6800" refString="Harden, P. H. &amp; C. E. Mickel. 1952. The stoneflies of Minnesota (Plecoptera). University of Minnesota Technical Bulletin, 201: 1 - 84." type="journal article" year="1952">Harden &amp; Mickel 1952:19</bibRefCitation>
</taxonomicName>
.
</treatmentCitation>
</treatmentCitationGroup>
<treatmentCitationGroup box="[227,672,1673,1695]" pageId="4" pageNumber="48">
<treatmentCitation box="[227,672,1673,1695]" pageId="4" pageNumber="48">
<taxonomicName authority=": Ricker 1952: 36" authorityName="Ricker" authorityPageNumber="36" authorityYear="1952" box="[227,665,1673,1695]" class="Insecta" family="Nemouridae" genus="Nemoura" kingdom="Animalia" order="Plecoptera" pageId="4" pageNumber="48" phylum="Arthropoda" rank="species" species="arctica" subGenus="Nemoura">
<emphasis box="[227,317,1674,1695]" italics="true" pageId="4" pageNumber="48">Nemoura</emphasis>
(
<emphasis box="[331,422,1674,1695]" italics="true" pageId="4" pageNumber="48">Nemoura</emphasis>
) 
<emphasis box="[436,501,1674,1695]" italics="true" pageId="4" pageNumber="48">arctica</emphasis>
: 
<bibRefCitation author="Ricker, W. E." box="[513,665,1673,1695]" pageId="4" pageNumber="48" pagination="1 - 200" refId="ref8013" refString="Ricker, W. E. 1952. Systematic Studies in Plecoptera. Indiana University Publications Series, 18: 1 - 200. http: // www. nativefishlab. net / library / textpdf / 16" type="journal article" year="1952">Ricker 1952:36</bibRefCitation>
</taxonomicName>
.
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
<paragraph blockId="4.[227,797,1252,1922]" box="[227,605,1706,1728]" pageId="4" pageNumber="48">
<treatmentCitationGroup box="[227,605,1706,1728]" pageId="4" pageNumber="48">
<treatmentCitation box="[227,605,1706,1728]" pageId="4" pageNumber="48">
<taxonomicName authority=": Zhiltzova 1964: 187" authorityName="Zhiltzova" authorityPageNumber="187" authorityYear="1964" box="[227,599,1706,1728]" class="Insecta" family="Nemouridae" genus="Nemoura" kingdom="Animalia" order="Plecoptera" pageId="4" pageNumber="48" phylum="Arthropoda" rank="species" species="arctica">
<emphasis box="[227,388,1706,1728]" italics="true" pageId="4" pageNumber="48">Nemoura arctica</emphasis>
: 
<bibRefCitation author="Zhiltzova, L. A." box="[401,599,1706,1728]" pageId="4" pageNumber="48" pagination="177 - 200" refId="ref8562" refString="Zhiltzova, L. A. 1964. Order Plecoptera (Stoneflies). Pages 177 - 200. In: Bei-Bienko (editor). Opredelitel' nasekomykh Evropeiskoi chasti SSSR (Identification Guide to Insects of the European Part of the USSR), Moscow, Leningrad." type="book chapter" year="1964">Zhiltzova 1964:187</bibRefCitation>
</taxonomicName>
.
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
<paragraph blockId="4.[227,797,1252,1922]" box="[227,553,1738,1760]" pageId="4" pageNumber="48">
<treatmentCitationGroup box="[227,553,1738,1760]" pageId="4" pageNumber="48">
<treatmentCitation box="[227,553,1738,1760]" pageId="4" pageNumber="48">
<taxonomicName authority=": Illies 1966: 194" authorityName="Illies" authorityPageNumber="194" authorityYear="1966" box="[227,548,1738,1760]" class="Insecta" family="Nemouridae" genus="Nemoura" kingdom="Animalia" order="Plecoptera" pageId="4" pageNumber="48" phylum="Arthropoda" rank="species" species="arctica">
<emphasis box="[227,388,1739,1760]" italics="true" pageId="4" pageNumber="48">Nemoura arctica</emphasis>
: 
<bibRefCitation author="Illies, J." box="[401,548,1738,1760]" pageId="4" pageNumber="48" refId="ref6996" refString="Illies, J. 1966. Katalog der rezenten Plecoptera. Das Tierreich. 82. Walter de Gruyter, Berlin. 631 pp." type="book" year="1966">Illies 1966:194</bibRefCitation>
</taxonomicName>
.
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
<paragraph blockId="4.[227,797,1252,1922]" box="[227,585,1770,1792]" pageId="4" pageNumber="48">
<treatmentCitationGroup box="[227,585,1770,1792]" pageId="4" pageNumber="48">
<treatmentCitation box="[227,585,1770,1792]" pageId="4" pageNumber="48">
<taxonomicName authority=": Illies 1966: 214" authorityName="Illies" authorityPageNumber="214" authorityYear="1966" box="[227,578,1770,1792]" class="Insecta" family="Nemouridae" genus="Nemoura" kingdom="Animalia" order="Plecoptera" pageId="4" pageNumber="48" phylum="Arthropoda" rank="species" species="trispinosa">
<emphasis box="[227,419,1771,1792]" italics="true" pageId="4" pageNumber="48">Nemoura trispinosa</emphasis>
: 
<bibRefCitation author="Illies, J." box="[431,578,1770,1792]" pageId="4" pageNumber="48" refId="ref6996" refString="Illies, J. 1966. Katalog der rezenten Plecoptera. Das Tierreich. 82. Walter de Gruyter, Berlin. 631 pp." type="book" year="1966">Illies 1966:214</bibRefCitation>
</taxonomicName>
.
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
<paragraph blockId="4.[227,797,1252,1922]" box="[227,651,1803,1825]" pageId="4" pageNumber="48">
<treatmentCitationGroup box="[227,651,1803,1825]" pageId="4" pageNumber="48">
<treatmentCitation box="[227,651,1803,1825]" pageId="4" pageNumber="48">
<taxonomicName authority=": Lillehammer 1972 a: 163" authorityName="Lillehammer" authorityPageNumber="163" authorityYear="1972" box="[227,644,1803,1825]" class="Insecta" family="Nemouridae" genus="Nemoura" kingdom="Animalia" order="Plecoptera" pageId="4" pageNumber="48" phylum="Arthropoda" rank="species" species="arctica">
<emphasis box="[227,388,1804,1825]" italics="true" pageId="4" pageNumber="48">Nemoura arctica</emphasis>
: 
<bibRefCitation author="Lillehammer, A." box="[401,644,1803,1825]" pageId="4" pageNumber="48" pagination="161 - 163" refId="ref7350" refString="Lillehammer, A. 1972 a. A new species of the genus Nemoura (Plecoptera) from Finnmark, North Norway. Norsk Entomologisk Tidsskrift, 19: 161 - 163. http: // www. entomologi. no / journals / nje / old / V 19" type="journal article" year="1972">Lillehammer 1972a:163</bibRefCitation>
</taxonomicName>
.
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
<paragraph blockId="4.[227,797,1252,1922]" box="[227,682,1835,1857]" pageId="4" pageNumber="48">
<treatmentCitationGroup box="[227,682,1835,1857]" pageId="4" pageNumber="48">
<treatmentCitation box="[227,682,1835,1857]" pageId="4" pageNumber="48">
<taxonomicName authority=": Lillehammer 1972 a: 163" authorityName="Lillehammer" authorityPageNumber="163" authorityYear="1972" box="[227,675,1835,1857]" class="Insecta" family="Nemouridae" genus="Nemoura" kingdom="Animalia" order="Plecoptera" pageId="4" pageNumber="48" phylum="Arthropoda" rank="species" species="trispinosa">
<emphasis box="[227,419,1836,1857]" italics="true" pageId="4" pageNumber="48">Nemoura trispinosa</emphasis>
: 
<bibRefCitation author="Lillehammer, A." box="[431,675,1835,1857]" pageId="4" pageNumber="48" pagination="161 - 163" refId="ref7350" refString="Lillehammer, A. 1972 a. A new species of the genus Nemoura (Plecoptera) from Finnmark, North Norway. Norsk Entomologisk Tidsskrift, 19: 161 - 163. http: // www. entomologi. no / journals / nje / old / V 19" type="journal article" year="1972">Lillehammer 1972a:163</bibRefCitation>
</taxonomicName>
.
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
<paragraph blockId="4.[227,797,1252,1922]" box="[227,582,1868,1890]" pageId="4" pageNumber="48">
<treatmentCitationGroup box="[227,582,1868,1890]" pageId="4" pageNumber="48">
<treatmentCitation box="[227,582,1868,1890]" pageId="4" pageNumber="48">
<taxonomicName authority=": Zwick 1973 a: 332" authorityName="Zwick" authorityPageNumber="332" authorityYear="1973" box="[227,576,1868,1890]" class="Insecta" family="Nemouridae" genus="Nemoura" kingdom="Animalia" order="Plecoptera" pageId="4" pageNumber="48" phylum="Arthropoda" rank="species" species="arctica">
<emphasis box="[227,388,1868,1889]" italics="true" pageId="4" pageNumber="48">Nemoura arctica</emphasis>
: 
<bibRefCitation author="Zwick, P." box="[401,576,1868,1890]" pageId="4" pageNumber="48" refId="ref8764" refString="Zwick, P. 1973 a. Insecta: Plecoptera. Phylogenetisches System und Katalog. Das Tierreich 94. Walter de Gruyter, Berlin. 465 pp." type="book" year="1973">Zwick 1973a:332</bibRefCitation>
</taxonomicName>
.
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
<paragraph blockId="4.[227,797,1252,1922]" box="[227,613,1900,1922]" pageId="4" pageNumber="48">
<treatmentCitationGroup box="[227,613,1900,1922]" pageId="4" pageNumber="48">
<treatmentCitation box="[227,613,1900,1922]" pageId="4" pageNumber="48">
<taxonomicName authority=": Zwick 1973 a: 342" authorityName="Zwick" authorityPageNumber="342" authorityYear="1973" box="[227,607,1900,1922]" class="Insecta" family="Nemouridae" genus="Nemoura" kingdom="Animalia" order="Plecoptera" pageId="4" pageNumber="48" phylum="Arthropoda" rank="species" species="trispinosa">
<emphasis box="[227,419,1901,1922]" italics="true" pageId="4" pageNumber="48">Nemoura trispinosa</emphasis>
: 
<bibRefCitation author="Zwick, P." box="[431,607,1900,1922]" pageId="4" pageNumber="48" refId="ref8764" refString="Zwick, P. 1973 a. Insecta: Plecoptera. Phylogenetisches System und Katalog. Das Tierreich 94. Walter de Gruyter, Berlin. 465 pp." type="book" year="1973">Zwick 1973a:342</bibRefCitation>
</taxonomicName>
.
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
<paragraph blockId="4.[873,1404,227,508]" box="[873,1285,227,249]" pageId="4" pageNumber="48">
<treatmentCitationGroup box="[873,1285,227,249]" pageId="4" pageNumber="48">
<treatmentCitation box="[873,1285,227,249]" pageId="4" pageNumber="48">
<taxonomicName authority=": Lillehammer 1974 a: 82" authorityName="Lillehammer" authorityPageNumber="82" authorityYear="1974" box="[873,1278,227,249]" class="Insecta" family="Nemouridae" genus="Nemoura" kingdom="Animalia" order="Plecoptera" pageId="4" pageNumber="48" phylum="Arthropoda" rank="species" species="arctica">
<emphasis box="[873,1034,228,249]" italics="true" pageId="4" pageNumber="48">Nemoura arctica</emphasis>
: 
<bibRefCitation author="Lillehammer, A." box="[1047,1278,227,249]" pageId="4" pageNumber="48" pagination="59 - 107" refId="ref7469" refString="Lillehammer, A. 1974 a. Norwegian stoneflies. I. Analysis of the variations in morphological and structural characters used in taxonomy. Norsk Entomologisk Tidsskrift, 21: 59 - 107." type="journal article" year="1974">Lillehammer 1974a:82</bibRefCitation>
</taxonomicName>
.
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
<paragraph blockId="4.[873,1404,227,508]" box="[873,1239,259,281]" pageId="4" pageNumber="48">
<treatmentCitationGroup box="[873,1239,259,281]" pageId="4" pageNumber="48">
<treatmentCitation box="[873,1239,259,281]" pageId="4" pageNumber="48">
<taxonomicName authority=": Baumann 1975: 21" authorityName="Baumann" authorityPageNumber="21" authorityYear="1975" box="[873,1232,259,281]" class="Insecta" family="Nemouridae" genus="Nemoura" kingdom="Animalia" order="Plecoptera" pageId="4" pageNumber="48" phylum="Arthropoda" rank="species" species="arctica">
<emphasis box="[873,1034,260,281]" italics="true" pageId="4" pageNumber="48">Nemoura arctica</emphasis>
: 
<bibRefCitation author="Baumann, R. W." box="[1047,1232,259,281]" pageId="4" pageNumber="48" refId="ref5991" refString="Baumann, R. W. 1975. Revision of the stonefly family Nemouridae (Plecoptera): a study of the" type="book" year="1975">Baumann 1975:21</bibRefCitation>
</taxonomicName>
.
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
<paragraph blockId="4.[873,1404,227,508]" box="[873,1270,292,314]" pageId="4" pageNumber="48">
<treatmentCitationGroup box="[873,1270,292,314]" pageId="4" pageNumber="48">
<treatmentCitation box="[873,1270,292,314]" pageId="4" pageNumber="48">
<taxonomicName authority=": Baumann 1975: 21" authorityName=": Baumann" authorityPageNumber="21" authorityYear="1975" box="[873,1263,292,314]" class="Insecta" family="Nemouridae" genus="Nemoura" kingdom="Animalia" order="Plecoptera" pageId="4" pageNumber="48" phylum="Arthropoda" rank="species" species="trispinosa">
<emphasis box="[873,1065,293,314]" italics="true" pageId="4" pageNumber="48">Nemoura trispinosa</emphasis>
: 
<bibRefCitation author="Baumann, R. W." box="[1077,1263,292,314]" pageId="4" pageNumber="48" refId="ref5991" refString="Baumann, R. W. 1975. Revision of the stonefly family Nemouridae (Plecoptera): a study of the" type="book" year="1975">Baumann 1975:21</bibRefCitation>
</taxonomicName>
.
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
<paragraph blockId="4.[873,1404,227,508]" box="[873,1295,324,346]" pageId="4" pageNumber="48">
<treatmentCitationGroup box="[873,1295,324,346]" pageId="4" pageNumber="48">
<treatmentCitation box="[873,1295,324,346]" pageId="4" pageNumber="48">
<taxonomicName authority=": Baumann et al. 1977: 34" authorityName="Baumann" authorityPageNumber="34" authorityYear="1977" box="[873,1289,324,346]" class="Insecta" family="Nemouridae" genus="Nemoura" kingdom="Animalia" order="Plecoptera" pageId="4" pageNumber="48" phylum="Arthropoda" rank="species" species="arctica">
<emphasis box="[873,1034,325,346]" italics="true" pageId="4" pageNumber="48">Nemoura arctica</emphasis>
: 
<bibRefCitation author="Baumann, R. W. &amp; A. R. Gaufin &amp; R. E. Surdick" box="[1047,1289,324,346]" pageId="4" pageNumber="48" pagination="1 - 207" refId="ref6031" refString="Baumann, R. W., A. R. Gaufin, &amp; R. E. Surdick. 1977. The stoneflies (Plecoptera) of the Rocky Mountains. Memoirs of the American Entomological Society, 31: 1 - 207." type="journal article" year="1977">Baumann et al. 1977:34</bibRefCitation>
</taxonomicName>
.
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
<paragraph blockId="4.[873,1404,227,508]" box="[873,1285,356,378]" pageId="4" pageNumber="48">
<treatmentCitationGroup box="[873,1285,356,378]" pageId="4" pageNumber="48">
<treatmentCitation box="[873,1285,356,378]" pageId="4" pageNumber="48">
<taxonomicName authority=": Lillehammer 1988: 113" authorityName="Lillehammer" authorityPageNumber="113" authorityYear="1988" box="[873,1278,356,378]" class="Insecta" family="Nemouridae" genus="Nemoura" kingdom="Animalia" order="Plecoptera" pageId="4" pageNumber="48" phylum="Arthropoda" rank="species" species="arctica">
<emphasis box="[873,1034,357,378]" italics="true" pageId="4" pageNumber="48">Nemoura arctica</emphasis>
: 
<bibRefCitation author="Lillehammer, A." box="[1047,1278,356,378]" pageId="4" pageNumber="48" refId="ref7671" refString="Lillehammer, A. 1988. Stoneflies (Plecoptera) of Fennoscandia and Denmark. Fauna Entomologica Scandinavica, Volume 21. E. J. Brill / Scandinavian Science Press, New York. 165 pp." type="book" year="1988">Lillehammer 1988:113</bibRefCitation>
</taxonomicName>
.
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
<paragraph blockId="4.[873,1404,227,508]" box="[873,1251,389,411]" pageId="4" pageNumber="48">
<treatmentCitationGroup box="[873,1251,389,411]" pageId="4" pageNumber="48">
<treatmentCitation box="[873,1251,389,411]" pageId="4" pageNumber="48">
<taxonomicName authority=": Zhiltzova 2003: 266" authorityName="Zhiltzova" authorityPageNumber="266" authorityYear="2003" box="[873,1245,389,411]" class="Insecta" family="Nemouridae" genus="Nemoura" kingdom="Animalia" order="Plecoptera" pageId="4" pageNumber="48" phylum="Arthropoda" rank="species" species="arctica">
<emphasis box="[873,1034,389,411]" italics="true" pageId="4" pageNumber="48">Nemoura arctica</emphasis>
: 
<bibRefCitation author="Zhiltzova, L. A." box="[1047,1245,389,411]" pageId="4" pageNumber="48" refId="ref8643" refString="Zhiltzova, L. A. 2003. Fauna of Russia and Neighboring Countries, Vol. 1, Issue 1, Insecta Plecoptera, Plecoptera Gruppe Euoholognatha, NAUKA, St. Petersburg, 537 pp." type="book" year="2003">Zhiltzova 2003:266</bibRefCitation>
</taxonomicName>
.
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
<paragraph blockId="4.[873,1404,227,508]" pageId="4" pageNumber="48">
<treatmentCitationGroup box="[873,1404,421,443]" pageId="4" pageNumber="48">
<treatmentCitation box="[873,1404,421,443]" pageId="4" pageNumber="48">
<taxonomicName authority=": Kondratieff &amp; Baumann 2004: 114" authorityName="Kondratieff &amp; Baumann" authorityPageNumber="114" authorityYear="2004" box="[873,1399,421,443]" class="Insecta" family="Nemouridae" genus="Nemoura" kingdom="Animalia" order="Plecoptera" pageId="4" pageNumber="48" phylum="Arthropoda" rank="species" species="arctica">
<emphasis box="[873,1034,422,443]" italics="true" pageId="4" pageNumber="48">Nemoura arctica</emphasis>
: 
<bibRefCitation author="Kondratieff, B. C. &amp; R. W. Baumann" box="[1047,1399,421,443]" pageId="4" pageNumber="48" pagination="113 - 115" refId="ref7227" refString="Kondratieff, B. C. &amp; R. W. Baumann. 2004. A record of the Arctic Forestfly, Nemoura arctica (Plecoptera: Nemouridae), from the contiguous United States. Entomological News, 115: 113 - 115." type="journal article" year="2004">Kondratieff &amp; Baumann 2004:114</bibRefCitation>
</taxonomicName>
.
</treatmentCitation>
</treatmentCitationGroup>
<treatmentCitationGroup box="[873,1339,454,476]" pageId="4" pageNumber="48">
<treatmentCitation box="[873,1339,454,476]" pageId="4" pageNumber="48">
<taxonomicName authority=": Stewart &amp; Oswood 2006: 78" authorityName="Stewart &amp; Oswood" authorityPageNumber="78" authorityYear="2006" box="[873,1332,454,476]" class="Insecta" family="Nemouridae" genus="Nemoura" kingdom="Animalia" order="Plecoptera" pageId="4" pageNumber="48" phylum="Arthropoda" rank="species" species="arctica">
<emphasis box="[873,1034,454,475]" italics="true" pageId="4" pageNumber="48">Nemoura arctica</emphasis>
: 
<bibRefCitation author="Stewart, K. W. &amp; M. W. Oswood" box="[1047,1332,454,476]" pageId="4" pageNumber="48" refId="ref8079" refString="Stewart, K. W. &amp; M. W. Oswood. 2006. The stoneflies (Plecoptera) of Alaska and western Canada. Caddis Press, Columbus, Ohio. 325 pp." type="book" year="2006">Stewart &amp; Oswood 2006:78</bibRefCitation>
</taxonomicName>
.
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
<paragraph blockId="4.[873,1404,227,508]" box="[873,1317,486,508]" pageId="4" pageNumber="48">
<treatmentCitationGroup box="[873,1317,486,508]" pageId="4" pageNumber="48">
<treatmentCitation box="[873,1317,486,508]" pageId="4" pageNumber="48">
<taxonomicName authority=": Judson &amp; Nelson 2012: 33" authorityName=": Judson &amp; Nelson" authorityPageNumber="33" authorityYear="2012" box="[873,1311,486,508]" class="Insecta" family="Nemouridae" genus="Nemoura" kingdom="Animalia" order="Plecoptera" pageId="4" pageNumber="48" phylum="Arthropoda" rank="species" species="arctica">
<emphasis box="[873,1034,487,508]" italics="true" pageId="4" pageNumber="48">Nemoura arctica</emphasis>
: 
<bibRefCitation author="Judson, S. W. &amp; C. R. Nelson" box="[1047,1311,486,508]" pageId="4" pageNumber="48" pagination="1 - 118" refId="ref7116" refString="Judson, S. W. &amp; C. R. Nelson. 2012. A guide to Mongolian stoneflies (Insecta: Plecoptera). Zootaxa, 3541: 1 - 118." type="journal article" year="2012">Judson &amp; Nelson 2012:33</bibRefCitation>
</taxonomicName>
.
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection pageId="4" pageNumber="48" type="distribution">
<paragraph blockId="4.[835,1443,559,727]" pageId="4" pageNumber="48">
<emphasis bold="true" box="[835,994,559,583]" pageId="4" pageNumber="48">Distribution.</emphasis>
<collectingCountry box="[1007,1101,559,583]" name="Canada" pageId="4" pageNumber="48">Canada</collectingCountry>
: AB, BC, LB, 
<collectionCode box="[1280,1324,559,582]" country="Portugal" name="Universidade de Lisboa, Museu Bocage" pageId="4" pageNumber="48">MB</collectionCode>
, NB, NS, NT, 
<collectionCode box="[893,938,596,619]" country="Thailand" name="Department of Microbiology, Faculty of Science" pageId="4" pageNumber="48">NU</collectionCode>
, ON, PE, PQ, YK. Europe: Baltic States, 
<collectingCountry box="[835,928,631,655]" name="Finland" pageId="4" pageNumber="48">Finland</collectingCountry>
, 
<collectingCountry box="[943,1043,631,655]" name="Norway" pageId="4" pageNumber="48">Norway</collectingCountry>
, 
<collectingCountry box="[1057,1154,631,655]" name="Sweden" pageId="4" pageNumber="48">Sweden</collectingCountry>
. 
<collectingCountry box="[1168,1285,631,655]" name="Mongolia" pageId="4" pageNumber="48">Mongolia</collectingCountry>
. 
<collectingCountry box="[1298,1375,631,655]" name="Russia" pageId="4" pageNumber="48">Russia</collectingCountry>
East, 
<collectingCountry box="[835,912,667,691]" name="Russia" pageId="4" pageNumber="48">Russia</collectingCountry>
North, West Siberia. 
<collectingCountry box="[1178,1237,667,691]" name="United States of America" pageId="4" pageNumber="48">USA</collectingCountry>
: AK, IA, IL, ME, MI, NY, OH, PA, SD, WI, WY (
<bibRefCitation author="DeWalt, R. E. &amp; M. D. Maehr &amp; U. Neu-Becker &amp; G. Stueber" box="[1198,1416,703,727]" pageId="4" pageNumber="48" refId="ref6493" refString="DeWalt, R. E., M. D. Maehr, U. Neu-Becker &amp; G. Stueber. 2018. Plecoptera Species File Online. Version 5.0 / 5.0. Accessed 1 March 2018. http: // Plecoptera. SpeciesFile. org" type="url" year="2018">DeWalt et al. 2018</bibRefCitation>
).
</paragraph>
<subSection lastPageId="8" lastPageNumber="52" pageId="4" pageNumber="48" type="multiple">
<paragraph blockId="4.[835,1411,778,802]" box="[835,1411,778,802]" pageId="4" pageNumber="48">
<heading bold="true" box="[835,1411,778,802]" fontSize="10" level="2" pageId="4" pageNumber="48" reason="2">
<emphasis bold="true" box="[835,1411,778,802]" pageId="4" pageNumber="48">
New Canadian province and 
<collectingCountry box="[1189,1247,778,802]" name="United States of America" pageId="4" pageNumber="48">USA</collectingCountry>
state records.
</emphasis>
</heading>
</paragraph>
<subSubSection pageId="4" pageNumber="48" type="materials_examined">
<paragraph blockId="4.[835,1444,814,1918]" pageId="4" pageNumber="48">
<materialsCitation ID-GBIF-Occurrence="3111722302" pageId="4" pageNumber="48">
<emphasis bold="true" box="[835,1134,814,838]" pageId="4" pageNumber="48">
<collectingCountry box="[835,928,814,838]" name="Canada" pageId="4" pageNumber="48">Canada</collectingCountry>
, 
<collectingRegion box="[955,1129,814,838]" country="Canada" name="Saskatchewan" pageId="4" pageNumber="48">Saskatchewan</collectingRegion>
,
</emphasis>
stream at Promontory Campground, 
<quantity box="[1009,1107,850,874]" metricMagnitude="4" metricUnit="m" metricValue="2.414016" pageId="4" pageNumber="48" unit="mi" value="15.0">15 miles</quantity>
north of La Rouge, junction Hwy 102, 
<date box="[986,1115,887,911]" pageId="4" pageNumber="48" value="1976-06-21">21-VI-1976</date>
, L.M. Dosdall, 
<specimenCount box="[1340,1374,886,911]" pageId="4" pageNumber="48" type="male">4♂</specimenCount>
, 
<specimenCount box="[1403,1437,886,911]" pageId="4" pageNumber="48" type="female">2♀</specimenCount>
;
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3111722304" pageId="4" pageNumber="48">
Puskwakau River, Hwy 106, 
<date box="[1241,1362,922,947]" pageId="4" pageNumber="48" value="1976-05-30">30-V-1976</date>
, L.M. Dosdall, 
<specimenCount box="[946,980,958,982]" pageId="4" pageNumber="48" type="male">1♂</specimenCount>
.
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3111722307" pageId="4" pageNumber="48">
<emphasis bold="true" box="[999,1205,958,982]" pageId="4" pageNumber="48">
<collectingCountry box="[999,1058,958,982]" name="United States of America" pageId="4" pageNumber="48">USA</collectingCountry>
, 
<collectingRegion box="[1075,1205,958,982]" country="United States of America" name="Minnesota" pageId="4" pageNumber="48">Minnesota</collectingRegion>
</emphasis>
(
<bibRefCitation author="Harden, P. H. &amp; C. E. Mickel" pageId="4" pageNumber="48" pagination="1 - 84" refId="ref6800" refString="Harden, P. H. &amp; C. E. Mickel. 1952. The stoneflies of Minnesota (Plecoptera). University of Minnesota Technical Bulletin, 201: 1 - 84." type="journal article" year="1952">Harden &amp; Mickel 1952</bibRefCitation>
, their pp. 19–20). 
<emphasis bold="true" box="[1109,1319,994,1018]" pageId="4" pageNumber="48">
<collectingRegion box="[1109,1311,994,1018]" country="United States of America" name="New Hampshire" pageId="4" pageNumber="48">New Hampshire</collectingRegion>
,
</emphasis>
Coös Co., Lakes of the Clouds, White Mountains, 
<date pageId="4" pageNumber="48" value="1951-06-20">20 June 1951</date>
, C.P. Alexander, 
<specimenCount box="[1118,1152,1066,1091]" pageId="4" pageNumber="48" type="male">1♂</specimenCount>
(USNM);
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3111722305" pageId="4" pageNumber="48">
Grafton Co., Franconia, A. 
<collectionCode box="[997,1015,1102,1126]" name="Tavera, Department of Geology and Geophysics" pageId="4" pageNumber="48">T</collectionCode>
. Slosson, no date information, 
<specimenCount box="[1409,1443,1102,1126]" pageId="4" pageNumber="48" type="male">1♂</specimenCount>
(USNM;
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3111722306" pageId="4" pageNumber="48">
Ac #26226). 
<emphasis bold="true" box="[1103,1279,1138,1162]" pageId="4" pageNumber="48">
<collectingRegion box="[1103,1279,1138,1162]" country="United States of America" name="West Virginia" pageId="4" pageNumber="48">West Virginia</collectingRegion>
</emphasis>
, Tucker Co., Abe Run, Canaan Valley State Park, 
<date box="[1282,1438,1174,1199]" pageId="4" pageNumber="48" value="1993-05-28">28 May 1993</date>
, S.M. Clark, 
<specimenCount box="[973,1006,1210,1235]" pageId="4" pageNumber="48" type="female">1♀</specimenCount>
(
<collectionCode box="[1021,1081,1211,1235]" country="USA" httpUri="http://grbio.org/cool/0am8-rdhc" name="Monte L. Bean Life Science Museum" pageId="4" pageNumber="48" type="University or college">BYU</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="4" pageNumber="48" type="diagnosis">
<paragraph blockId="4.[835,1444,814,1918]" box="[835,965,1246,1270]" pageId="4" pageNumber="48">
<emphasis bold="true" box="[835,958,1246,1270]" pageId="4" pageNumber="48">Diagnosis</emphasis>
.
</paragraph>
<paragraph blockId="4.[835,1444,814,1918]" pageId="4" pageNumber="48">
<emphasis bold="true" box="[835,921,1282,1306]" pageId="4" pageNumber="48">
<emphasis bold="true" box="[835,914,1282,1306]" italics="true" pageId="4" pageNumber="48">Cercus</emphasis>
.
</emphasis>
Highly variable. Adults of 
<taxonomicName authorityName=": Judson &amp; Nelson" authorityYear="2012" box="[1249,1356,1283,1306]" class="Insecta" family="Nemouridae" genus="Nemoura" kingdom="Animalia" order="Plecoptera" pageId="4" pageNumber="48" phylum="Arthropoda" rank="species" species="arctica">
<emphasis box="[1249,1356,1283,1306]" italics="true" pageId="4" pageNumber="48">N. arctica</emphasis>
</taxonomicName>
and 
<taxonomicName authorityName=": Baumann" authorityYear="1975" class="Insecta" family="Nemouridae" genus="Nemoura" kingdom="Animalia" order="Plecoptera" pageId="4" pageNumber="48" phylum="Arthropoda" rank="species" species="trispinosa">
<emphasis italics="true" pageId="4" pageNumber="48">N. trispinosa</emphasis>
</taxonomicName>
have been previously differentiated by a combination of cercal characteristics (males) and body size and distribution (females) (
<bibRefCitation author="Ricker, W. E." box="[1287,1431,1390,1414]" pageId="4" pageNumber="48" pagination="1 - 200" refId="ref8013" refString="Ricker, W. E. 1952. Systematic Studies in Plecoptera. Indiana University Publications Series, 18: 1 - 200. http: // www. nativefishlab. net / library / textpdf / 16" type="journal article" year="1952">Ricker 1952</bibRefCitation>
). Male cerci are sclerotized laterally and terminate typically in a pair of appressed spines that vary in length and degree of tapering (
<figureCitation box="[1220,1344,1498,1522]" captionStart-0="Figs" captionStart-1="Figs" captionStartId-0="2.[189,238,1788,1812]" captionStartId-1="3.[189,238,1784,1808]" captionTargetBox-0="[324,1307,254,1740]" captionTargetBox-1="[324,1307,254,1737]" captionTargetId-0="figure-125@2.[324,1307,254,1740]" captionTargetId-1="figure-116@3.[324,1307,254,1737]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionText-0="Figs. 1–8. Nemoura arctica, cerci, distal view. 1, USA, Iowa, Twin Springs, 545X; 2, USA, Illinois, Elgin, 516X; 3, USA, Ohio, John Bryan State Park, 518X; 4, USA, Wisconsin, spring, 1 mile N Irvington, 509X; 5, USA, South Dakota, Burnt Fork, 497X; 6, USA, Wyoming, Blacktail Creek, 584X; 7, Canada, Saskatchewan, Promontory Campground, 605X; 8, Canada, New Brunswick, Coombs Brook, 591X." captionText-1="Figs. 9–16. Nemoura arctica, cerci, distal view. 9, Canada, Manitoba, Warkworth Creek, 773X; 10, Canada, Northwest Territories, Thelon River, 636X; 11, Canada, Northwest Territories, Tuktoyaktuk, 612X; 12, Russia, Siberia, Magadan Oblask, 486X; 13, Norway, Buktved Garnvika, 443X; 14, Norway, Kautokeino, 431X; 15, Mongolia, Khoton Lake, 675X; 16, Mongolia, Tsagaaan Lake, 646X." figureDoi-0="http://doi.org/10.5281/zenodo.4761206" figureDoi-1="http://doi.org/10.5281/zenodo.4761208" httpUri-0="https://zenodo.org/record/4761206/files/figure.png" httpUri-1="https://zenodo.org/record/4761208/files/figure.png" pageId="4" pageNumber="48">Figs. 1–16</figureCitation>
), plus a third unit that is highly variable and has been used for the past ca. 65 years to separate males of 
<taxonomicName authorityName=": Judson &amp; Nelson" authorityYear="2012" class="Insecta" family="Nemouridae" genus="Nemoura" kingdom="Animalia" order="Plecoptera" pageId="4" pageNumber="48" phylum="Arthropoda" rank="species" species="arctica">
<emphasis italics="true" pageId="4" pageNumber="48">N. arctica</emphasis>
</taxonomicName>
and 
<taxonomicName authority="(Ricker 1952)" baseAuthorityName="Ricker" baseAuthorityYear="1952" box="[969,1279,1606,1630]" class="Insecta" family="Nemouridae" genus="Nemoura" kingdom="Animalia" order="Plecoptera" pageId="4" pageNumber="48" phylum="Arthropoda" rank="species" species="trispinosa">
<emphasis box="[969,1111,1607,1630]" italics="true" pageId="4" pageNumber="48">N. trispinosa</emphasis>
(
<bibRefCitation author="Ricker, W. E." box="[1128,1272,1606,1630]" pageId="4" pageNumber="48" pagination="1 - 200" refId="ref8013" refString="Ricker, W. E. 1952. Systematic Studies in Plecoptera. Indiana University Publications Series, 18: 1 - 200. http: // www. nativefishlab. net / library / textpdf / 16" type="journal article" year="1952">Ricker 1952</bibRefCitation>
)
</taxonomicName>
. 
<bibRefCitation author="Lillehammer, A." pageId="4" pageNumber="48" pagination="161 - 163" refId="ref7350" refString="Lillehammer, A. 1972 a. A new species of the genus Nemoura (Plecoptera) from Finnmark, North Norway. Norsk Entomologisk Tidsskrift, 19: 161 - 163. http: // www. entomologi. no / journals / nje / old / V 19" type="journal article" year="1972">Lillehammer (1972a)</bibRefCitation>
later illustrated cerci as either lacking (
<taxonomicName authorityName=": Judson &amp; Nelson" authorityYear="2012" class="Insecta" family="Nemouridae" genus="Nemoura" kingdom="Animalia" order="Plecoptera" pageId="4" pageNumber="48" phylum="Arthropoda" rank="species" species="arctica">
<emphasis italics="true" pageId="4" pageNumber="48">N. arctica</emphasis>
</taxonomicName>
, his 
<figureCitation box="[980,1067,1678,1702]" captionStart="Figs" captionStartId="2.[189,238,1788,1812]" captionTargetBox="[324,1307,254,1740]" captionTargetId="figure-125@2.[324,1307,254,1740]" captionTargetPageId="2" captionText="Figs. 1–8. Nemoura arctica, cerci, distal view. 1, USA, Iowa, Twin Springs, 545X; 2, USA, Illinois, Elgin, 516X; 3, USA, Ohio, John Bryan State Park, 518X; 4, USA, Wisconsin, spring, 1 mile N Irvington, 509X; 5, USA, South Dakota, Burnt Fork, 497X; 6, USA, Wyoming, Blacktail Creek, 584X; 7, Canada, Saskatchewan, Promontory Campground, 605X; 8, Canada, New Brunswick, Coombs Brook, 591X." figureDoi="http://doi.org/10.5281/zenodo.4761206" httpUri="https://zenodo.org/record/4761206/files/figure.png" pageId="4" pageNumber="48">Fig. 4b</figureCitation>
) or possessing a distinct (
<taxonomicName baseAuthorityName="Ricker" baseAuthorityYear="1952" class="Insecta" family="Nemouridae" genus="Nemoura" kingdom="Animalia" order="Plecoptera" pageId="4" pageNumber="48" phylum="Arthropoda" rank="species" species="trispinosa">
<emphasis italics="true" pageId="4" pageNumber="48">N. trispinosa</emphasis>
</taxonomicName>
, his 
<figureCitation box="[1005,1088,1714,1738]" captionStart="Figs" captionStartId="2.[189,238,1788,1812]" captionTargetBox="[324,1307,254,1740]" captionTargetId="figure-125@2.[324,1307,254,1740]" captionTargetPageId="2" captionText="Figs. 1–8. Nemoura arctica, cerci, distal view. 1, USA, Iowa, Twin Springs, 545X; 2, USA, Illinois, Elgin, 516X; 3, USA, Ohio, John Bryan State Park, 518X; 4, USA, Wisconsin, spring, 1 mile N Irvington, 509X; 5, USA, South Dakota, Burnt Fork, 497X; 6, USA, Wyoming, Blacktail Creek, 584X; 7, Canada, Saskatchewan, Promontory Campground, 605X; 8, Canada, New Brunswick, Coombs Brook, 591X." figureDoi="http://doi.org/10.5281/zenodo.4761206" httpUri="https://zenodo.org/record/4761206/files/figure.png" pageId="4" pageNumber="48">Fig. 4a</figureCitation>
) spine. 
<bibRefCitation author="Ricker, W. E." box="[1188,1358,1714,1738]" pageId="4" pageNumber="48" pagination="1 - 200" refId="ref8013" refString="Ricker, W. E. 1952. Systematic Studies in Plecoptera. Indiana University Publications Series, 18: 1 - 200. http: // www. nativefishlab. net / library / textpdf / 16" type="journal article" year="1952">Ricker’s (1952</bibRefCitation>
, p. 36) key to 
<taxonomicName baseAuthorityName="Ricker" baseAuthorityYear="1952" box="[921,1064,1751,1774]" class="Insecta" family="Nemouridae" genus="Nemoura" kingdom="Animalia" order="Plecoptera" pageId="4" pageNumber="48" phylum="Arthropoda" rank="species" species="trispinosa">
<emphasis box="[921,1064,1751,1774]" italics="true" pageId="4" pageNumber="48">N. trispinosa</emphasis>
</taxonomicName>
males focused on “…the outer edge of the cercus produced into a slender acute spine… may be forked once or twice at the tip”. This feature is common in North America and shown here clearly for populations from 
<collectingRegion box="[1272,1330,1895,1918]" country="United States of America" name="Iowa" pageId="4" pageNumber="48">Iowa</collectingRegion>
(
<figureCitation box="[1354,1426,1894,1918]" captionStart="Figs" captionStartId="2.[189,238,1788,1812]" captionTargetBox="[324,1307,254,1740]" captionTargetId="figure-125@2.[324,1307,254,1740]" captionTargetPageId="2" captionText="Figs. 1–8. Nemoura arctica, cerci, distal view. 1, USA, Iowa, Twin Springs, 545X; 2, USA, Illinois, Elgin, 516X; 3, USA, Ohio, John Bryan State Park, 518X; 4, USA, Wisconsin, spring, 1 mile N Irvington, 509X; 5, USA, South Dakota, Burnt Fork, 497X; 6, USA, Wyoming, Blacktail Creek, 584X; 7, Canada, Saskatchewan, Promontory Campground, 605X; 8, Canada, New Brunswick, Coombs Brook, 591X." figureDoi="http://doi.org/10.5281/zenodo.4761206" httpUri="https://zenodo.org/record/4761206/files/figure.png" pageId="4" pageNumber="48">Fig. 1</figureCitation>
),
</paragraph>
</subSubSection>
<caption ID-DOI="http://doi.org/10.5281/zenodo.4761210" ID-Zenodo-Dep="4761210" httpUri="https://zenodo.org/record/4761210/files/figure.png" pageId="5" pageNumber="49" startId="5.[189,238,1746,1770]" targetBox="[324,1307,218,1700]" targetPageId="5">
<paragraph blockId="5.[189,1442,1746,1914]" pageId="5" pageNumber="49">
Figs. 17–24. 
<taxonomicName authorityName=": Judson &amp; Nelson" authorityYear="2012" box="[335,517,1747,1770]" class="Insecta" family="Nemouridae" genus="Nemoura" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="49" phylum="Arthropoda" rank="species" species="arctica">
<emphasis box="[335,517,1747,1770]" italics="true" pageId="5" pageNumber="49">Nemoura arctica</emphasis>
</taxonomicName>
, epiproct, lateral view. 17, USA, Iowa, Twin Springs, 270X; 18, Canada, New Brunswick, Coombs Brook, 288X; 19, Canada, Northwest Territories, Thelon River, 285X; 20, Canada, Northwest Territories, Tuktoyaktuk, 328X; 21, Russia, Siberia, Magadan Oblask, 271X; 22, Norway, Buktved Garnvika, 286X; 23, Norway, Kautokeino, 316X; 24, Mongolia, Khoton Lake, 276X. AP = apical prong, BC = basal cushion, DS = dorsal sclerite, LA = lateral areas, VS = ventral sclerite.
</paragraph>
</caption>
<caption ID-DOI="http://doi.org/10.5281/zenodo.4761214" ID-Zenodo-Dep="4761214" httpUri="https://zenodo.org/record/4761214/files/figure.png" pageId="6" pageNumber="50" startId="6.[189,238,1748,1772]" targetBox="[324,1307,218,1701]" targetPageId="6">
<paragraph blockId="6.[189,1443,1748,1917]" pageId="6" pageNumber="50">
Figs. 25–32. 
<taxonomicName authorityName=": Judson &amp; Nelson" authorityYear="2012" box="[336,517,1749,1772]" class="Insecta" family="Nemouridae" genus="Nemoura" kingdom="Animalia" order="Plecoptera" pageId="6" pageNumber="50" phylum="Arthropoda" rank="species" species="arctica">
<emphasis box="[336,517,1749,1772]" italics="true" pageId="6" pageNumber="50">Nemoura arctica</emphasis>
</taxonomicName>
, epiproct, dorsal view. 25, USA, Iowa, Twin Springs, 141X; 26, USA, Illinois, Elgin, 165X; 27, USA, Ohio, spring, John Bryan State Park, 144X; 28, USA, Wisconsin, spring, 1 mile N Irvington, 132X; 29, USA, South Dakota, Burnt Fork, 140X; 30, USA, South Dakota, Burnt Fork, 229X; 31, Canada, Prince Edward Island, Balsam Hollow, 163X; 32, Canada, New Brunswick, Coombs Brook, 168X. AP = apical prong, BC = basal cushion, DS = dorsal sclerite, LA = lateral area, VS = ventral sclerite.
</paragraph>
</caption>
<caption ID-DOI="http://doi.org/10.5281/zenodo.4761216" ID-Zenodo-Dep="4761216" httpUri="https://zenodo.org/record/4761216/files/figure.png" pageId="7" pageNumber="51" startId="7.[189,238,1748,1772]" targetBox="[324,1307,218,1701]" targetPageId="7">
<paragraph blockId="7.[189,1443,1748,1917]" pageId="7" pageNumber="51">
Figs. 33–40. 
<taxonomicName authorityName=": Judson &amp; Nelson" authorityYear="2012" box="[334,515,1749,1772]" class="Insecta" family="Nemouridae" genus="Nemoura" kingdom="Animalia" order="Plecoptera" pageId="7" pageNumber="51" phylum="Arthropoda" rank="species" species="arctica">
<emphasis box="[334,515,1749,1772]" italics="true" pageId="7" pageNumber="51">Nemoura arctica</emphasis>
</taxonomicName>
, epiproct, dorsal view. 33, Canada, Saskatchewan, Promontory Campground, 187X; 34, Canada, Northwest Territories, Thelon River, 282X; 35, Canada, Northwest Territories, Tuktoyaktuk, 267X; 36, Norway, Buktved Garnvika, 321X; 37, Norway, Kautokeino, 278X; 38, Russia, Siberia, Magadan Oblask, 206X; 39, Mongolia, Khoton Lake, 277X; 40, Mongolia, Tsagaaan Lake, 281X. AP = apical prong, BC = basal cushion, DS = dorsal sclerite, LA = lateral area, VS = ventral sclerite.
</paragraph>
</caption>
<subSubSection pageId="8" pageNumber="52" type="description">
<paragraph blockId="8.[189,798,228,1907]" pageId="8" pageNumber="52">
Illinois (
<figureCitation box="[291,364,228,252]" captionStart="Figs" captionStartId="2.[189,238,1788,1812]" captionTargetBox="[324,1307,254,1740]" captionTargetId="figure-125@2.[324,1307,254,1740]" captionTargetPageId="2" captionText="Figs. 1–8. Nemoura arctica, cerci, distal view. 1, USA, Iowa, Twin Springs, 545X; 2, USA, Illinois, Elgin, 516X; 3, USA, Ohio, John Bryan State Park, 518X; 4, USA, Wisconsin, spring, 1 mile N Irvington, 509X; 5, USA, South Dakota, Burnt Fork, 497X; 6, USA, Wyoming, Blacktail Creek, 584X; 7, Canada, Saskatchewan, Promontory Campground, 605X; 8, Canada, New Brunswick, Coombs Brook, 591X." figureDoi="http://doi.org/10.5281/zenodo.4761206" httpUri="https://zenodo.org/record/4761206/files/figure.png" pageId="8" pageNumber="52">Fig. 2</figureCitation>
), Ohio (
<figureCitation box="[470,541,228,252]" captionStart="Figs" captionStartId="2.[189,238,1788,1812]" captionTargetBox="[324,1307,254,1740]" captionTargetId="figure-125@2.[324,1307,254,1740]" captionTargetPageId="2" captionText="Figs. 1–8. Nemoura arctica, cerci, distal view. 1, USA, Iowa, Twin Springs, 545X; 2, USA, Illinois, Elgin, 516X; 3, USA, Ohio, John Bryan State Park, 518X; 4, USA, Wisconsin, spring, 1 mile N Irvington, 509X; 5, USA, South Dakota, Burnt Fork, 497X; 6, USA, Wyoming, Blacktail Creek, 584X; 7, Canada, Saskatchewan, Promontory Campground, 605X; 8, Canada, New Brunswick, Coombs Brook, 591X." figureDoi="http://doi.org/10.5281/zenodo.4761206" httpUri="https://zenodo.org/record/4761206/files/figure.png" pageId="8" pageNumber="52">Fig. 3</figureCitation>
), Wisconsin (
<figureCitation box="[711,783,228,252]" captionStart="Figs" captionStartId="2.[189,238,1788,1812]" captionTargetBox="[324,1307,254,1740]" captionTargetId="figure-125@2.[324,1307,254,1740]" captionTargetPageId="2" captionText="Figs. 1–8. Nemoura arctica, cerci, distal view. 1, USA, Iowa, Twin Springs, 545X; 2, USA, Illinois, Elgin, 516X; 3, USA, Ohio, John Bryan State Park, 518X; 4, USA, Wisconsin, spring, 1 mile N Irvington, 509X; 5, USA, South Dakota, Burnt Fork, 497X; 6, USA, Wyoming, Blacktail Creek, 584X; 7, Canada, Saskatchewan, Promontory Campground, 605X; 8, Canada, New Brunswick, Coombs Brook, 591X." figureDoi="http://doi.org/10.5281/zenodo.4761206" httpUri="https://zenodo.org/record/4761206/files/figure.png" pageId="8" pageNumber="52">Fig. 4</figureCitation>
), South Dakota (
<figureCitation box="[369,437,264,288]" captionStart="Figs" captionStartId="2.[189,238,1788,1812]" captionTargetBox="[324,1307,254,1740]" captionTargetId="figure-125@2.[324,1307,254,1740]" captionTargetPageId="2" captionText="Figs. 1–8. Nemoura arctica, cerci, distal view. 1, USA, Iowa, Twin Springs, 545X; 2, USA, Illinois, Elgin, 516X; 3, USA, Ohio, John Bryan State Park, 518X; 4, USA, Wisconsin, spring, 1 mile N Irvington, 509X; 5, USA, South Dakota, Burnt Fork, 497X; 6, USA, Wyoming, Blacktail Creek, 584X; 7, Canada, Saskatchewan, Promontory Campground, 605X; 8, Canada, New Brunswick, Coombs Brook, 591X." figureDoi="http://doi.org/10.5281/zenodo.4761206" httpUri="https://zenodo.org/record/4761206/files/figure.png" pageId="8" pageNumber="52">Fig. 5</figureCitation>
), Wyoming (
<figureCitation box="[594,662,264,288]" captionStart="Figs" captionStartId="2.[189,238,1788,1812]" captionTargetBox="[324,1307,254,1740]" captionTargetId="figure-125@2.[324,1307,254,1740]" captionTargetPageId="2" captionText="Figs. 1–8. Nemoura arctica, cerci, distal view. 1, USA, Iowa, Twin Springs, 545X; 2, USA, Illinois, Elgin, 516X; 3, USA, Ohio, John Bryan State Park, 518X; 4, USA, Wisconsin, spring, 1 mile N Irvington, 509X; 5, USA, South Dakota, Burnt Fork, 497X; 6, USA, Wyoming, Blacktail Creek, 584X; 7, Canada, Saskatchewan, Promontory Campground, 605X; 8, Canada, New Brunswick, Coombs Brook, 591X." figureDoi="http://doi.org/10.5281/zenodo.4761206" httpUri="https://zenodo.org/record/4761206/files/figure.png" pageId="8" pageNumber="52">Fig. 6</figureCitation>
), and New Brunswick (
<figureCitation box="[333,401,300,324]" captionStart="Figs" captionStartId="2.[189,238,1788,1812]" captionTargetBox="[324,1307,254,1740]" captionTargetId="figure-125@2.[324,1307,254,1740]" captionTargetPageId="2" captionText="Figs. 1–8. Nemoura arctica, cerci, distal view. 1, USA, Iowa, Twin Springs, 545X; 2, USA, Illinois, Elgin, 516X; 3, USA, Ohio, John Bryan State Park, 518X; 4, USA, Wisconsin, spring, 1 mile N Irvington, 509X; 5, USA, South Dakota, Burnt Fork, 497X; 6, USA, Wyoming, Blacktail Creek, 584X; 7, Canada, Saskatchewan, Promontory Campground, 605X; 8, Canada, New Brunswick, Coombs Brook, 591X." figureDoi="http://doi.org/10.5281/zenodo.4761206" httpUri="https://zenodo.org/record/4761206/files/figure.png" pageId="8" pageNumber="52">Fig. 8</figureCitation>
), and also from Siberia (
<figureCitation box="[700,783,300,324]" captionStart="Figs" captionStartId="3.[189,238,1784,1808]" captionTargetBox="[324,1307,254,1737]" captionTargetId="figure-116@3.[324,1307,254,1737]" captionTargetPageId="3" captionText="Figs. 9–16. Nemoura arctica, cerci, distal view. 9, Canada, Manitoba, Warkworth Creek, 773X; 10, Canada, Northwest Territories, Thelon River, 636X; 11, Canada, Northwest Territories, Tuktoyaktuk, 612X; 12, Russia, Siberia, Magadan Oblask, 486X; 13, Norway, Buktved Garnvika, 443X; 14, Norway, Kautokeino, 431X; 15, Mongolia, Khoton Lake, 675X; 16, Mongolia, Tsagaaan Lake, 646X." figureDoi="http://doi.org/10.5281/zenodo.4761208" httpUri="https://zenodo.org/record/4761208/files/figure.png" pageId="8" pageNumber="52">Fig. 12</figureCitation>
), 
<collectingCountry box="[189,286,337,361]" name="Norway" pageId="8" pageNumber="52">Norway</collectingCountry>
(
<figureCitation box="[312,402,336,360]" captionStart="Figs" captionStartId="3.[189,238,1784,1808]" captionTargetBox="[324,1307,254,1737]" captionTargetId="figure-116@3.[324,1307,254,1737]" captionTargetPageId="3" captionText="Figs. 9–16. Nemoura arctica, cerci, distal view. 9, Canada, Manitoba, Warkworth Creek, 773X; 10, Canada, Northwest Territories, Thelon River, 636X; 11, Canada, Northwest Territories, Tuktoyaktuk, 612X; 12, Russia, Siberia, Magadan Oblask, 486X; 13, Norway, Buktved Garnvika, 443X; 14, Norway, Kautokeino, 431X; 15, Mongolia, Khoton Lake, 675X; 16, Mongolia, Tsagaaan Lake, 646X." figureDoi="http://doi.org/10.5281/zenodo.4761208" httpUri="https://zenodo.org/record/4761208/files/figure.png" pageId="8" pageNumber="52">Fig. 13</figureCitation>
) and 
<collectingCountry box="[490,604,336,360]" name="Mongolia" pageId="8" pageNumber="52">Mongolia</collectingCountry>
(
<figureCitation box="[630,722,336,360]" captionStart="Figs" captionStartId="3.[189,238,1784,1808]" captionTargetBox="[324,1307,254,1737]" captionTargetId="figure-116@3.[324,1307,254,1737]" captionTargetPageId="3" captionText="Figs. 9–16. Nemoura arctica, cerci, distal view. 9, Canada, Manitoba, Warkworth Creek, 773X; 10, Canada, Northwest Territories, Thelon River, 636X; 11, Canada, Northwest Territories, Tuktoyaktuk, 612X; 12, Russia, Siberia, Magadan Oblask, 486X; 13, Norway, Buktved Garnvika, 443X; 14, Norway, Kautokeino, 431X; 15, Mongolia, Khoton Lake, 675X; 16, Mongolia, Tsagaaan Lake, 646X." figureDoi="http://doi.org/10.5281/zenodo.4761208" httpUri="https://zenodo.org/record/4761208/files/figure.png" pageId="8" pageNumber="52">Fig. 16</figureCitation>
). The spine, however, varies in width and degree of tapering, For example, males from South Dakota have an outer spine that is rectangular, not “slender”, and crenulated distally (
<figureCitation box="[607,674,480,504]" captionStart="Figs" captionStartId="2.[189,238,1788,1812]" captionTargetBox="[324,1307,254,1740]" captionTargetId="figure-125@2.[324,1307,254,1740]" captionTargetPageId="2" captionText="Figs. 1–8. Nemoura arctica, cerci, distal view. 1, USA, Iowa, Twin Springs, 545X; 2, USA, Illinois, Elgin, 516X; 3, USA, Ohio, John Bryan State Park, 518X; 4, USA, Wisconsin, spring, 1 mile N Irvington, 509X; 5, USA, South Dakota, Burnt Fork, 497X; 6, USA, Wyoming, Blacktail Creek, 584X; 7, Canada, Saskatchewan, Promontory Campground, 605X; 8, Canada, New Brunswick, Coombs Brook, 591X." figureDoi="http://doi.org/10.5281/zenodo.4761206" httpUri="https://zenodo.org/record/4761206/files/figure.png" pageId="8" pageNumber="52">Fig. 5</figureCitation>
). 
<taxonomicName authorityName=": Judson &amp; Nelson" authorityYear="2012" class="Insecta" family="Nemouridae" genus="Nemoura" kingdom="Animalia" order="Plecoptera" pageId="8" pageNumber="52" phylum="Arthropoda" rank="species" species="arctica">
<emphasis italics="true" pageId="8" pageNumber="52">Nemoura arctica</emphasis>
</taxonomicName>
was separated out by 
<bibRefCitation author="Ricker, W. E." box="[552,711,516,540]" pageId="8" pageNumber="52" pagination="1 - 200" refId="ref8013" refString="Ricker, W. E. 1952. Systematic Studies in Plecoptera. Indiana University Publications Series, 18: 1 - 200. http: // www. nativefishlab. net / library / textpdf / 16" type="journal article" year="1952">Ricker (1952)</bibRefCitation>
as the “…outer edge of the cercus bordered by a crenulate shelf or ridge”. No form of a spine is evident for populations studied from North America from Saskatchewan (
<figureCitation box="[575,652,660,684]" captionStart="Figs" captionStartId="2.[189,238,1788,1812]" captionTargetBox="[324,1307,254,1740]" captionTargetId="figure-125@2.[324,1307,254,1740]" captionTargetPageId="2" captionText="Figs. 1–8. Nemoura arctica, cerci, distal view. 1, USA, Iowa, Twin Springs, 545X; 2, USA, Illinois, Elgin, 516X; 3, USA, Ohio, John Bryan State Park, 518X; 4, USA, Wisconsin, spring, 1 mile N Irvington, 509X; 5, USA, South Dakota, Burnt Fork, 497X; 6, USA, Wyoming, Blacktail Creek, 584X; 7, Canada, Saskatchewan, Promontory Campground, 605X; 8, Canada, New Brunswick, Coombs Brook, 591X." figureDoi="http://doi.org/10.5281/zenodo.4761206" httpUri="https://zenodo.org/record/4761206/files/figure.png" pageId="8" pageNumber="52">Fig. 7</figureCitation>
), Manitoba (
<figureCitation box="[197,265,696,720]" captionStart="Figs" captionStartId="3.[189,238,1784,1808]" captionTargetBox="[324,1307,254,1737]" captionTargetId="figure-116@3.[324,1307,254,1737]" captionTargetPageId="3" captionText="Figs. 9–16. Nemoura arctica, cerci, distal view. 9, Canada, Manitoba, Warkworth Creek, 773X; 10, Canada, Northwest Territories, Thelon River, 636X; 11, Canada, Northwest Territories, Tuktoyaktuk, 612X; 12, Russia, Siberia, Magadan Oblask, 486X; 13, Norway, Buktved Garnvika, 443X; 14, Norway, Kautokeino, 431X; 15, Mongolia, Khoton Lake, 675X; 16, Mongolia, Tsagaaan Lake, 646X." figureDoi="http://doi.org/10.5281/zenodo.4761208" httpUri="https://zenodo.org/record/4761208/files/figure.png" pageId="8" pageNumber="52">Fig. 9</figureCitation>
), and the Northwest Territories (
<figureCitation box="[657,789,696,720]" captionStart="Figs" captionStartId="3.[189,238,1784,1808]" captionTargetBox="[324,1307,254,1737]" captionTargetId="figure-116@3.[324,1307,254,1737]" captionTargetPageId="3" captionText="Figs. 9–16. Nemoura arctica, cerci, distal view. 9, Canada, Manitoba, Warkworth Creek, 773X; 10, Canada, Northwest Territories, Thelon River, 636X; 11, Canada, Northwest Territories, Tuktoyaktuk, 612X; 12, Russia, Siberia, Magadan Oblask, 486X; 13, Norway, Buktved Garnvika, 443X; 14, Norway, Kautokeino, 431X; 15, Mongolia, Khoton Lake, 675X; 16, Mongolia, Tsagaaan Lake, 646X." figureDoi="http://doi.org/10.5281/zenodo.4761208" httpUri="https://zenodo.org/record/4761208/files/figure.png" pageId="8" pageNumber="52">Figs. 10–11</figureCitation>
) plus 
<collectingCountry box="[257,354,732,756]" name="Norway" pageId="8" pageNumber="52">Norway</collectingCountry>
(
<figureCitation box="[379,469,732,756]" captionStart="Figs" captionStartId="3.[189,238,1784,1808]" captionTargetBox="[324,1307,254,1737]" captionTargetId="figure-116@3.[324,1307,254,1737]" captionTargetPageId="3" captionText="Figs. 9–16. Nemoura arctica, cerci, distal view. 9, Canada, Manitoba, Warkworth Creek, 773X; 10, Canada, Northwest Territories, Thelon River, 636X; 11, Canada, Northwest Territories, Tuktoyaktuk, 612X; 12, Russia, Siberia, Magadan Oblask, 486X; 13, Norway, Buktved Garnvika, 443X; 14, Norway, Kautokeino, 431X; 15, Mongolia, Khoton Lake, 675X; 16, Mongolia, Tsagaaan Lake, 646X." figureDoi="http://doi.org/10.5281/zenodo.4761208" httpUri="https://zenodo.org/record/4761208/files/figure.png" pageId="8" pageNumber="52">Fig. 14</figureCitation>
) and 
<collectingCountry box="[555,669,732,756]" name="Mongolia" pageId="8" pageNumber="52">Mongolia</collectingCountry>
(
<figureCitation box="[693,784,732,756]" captionStart="Figs" captionStartId="3.[189,238,1784,1808]" captionTargetBox="[324,1307,254,1737]" captionTargetId="figure-116@3.[324,1307,254,1737]" captionTargetPageId="3" captionText="Figs. 9–16. Nemoura arctica, cerci, distal view. 9, Canada, Manitoba, Warkworth Creek, 773X; 10, Canada, Northwest Territories, Thelon River, 636X; 11, Canada, Northwest Territories, Tuktoyaktuk, 612X; 12, Russia, Siberia, Magadan Oblask, 486X; 13, Norway, Buktved Garnvika, 443X; 14, Norway, Kautokeino, 431X; 15, Mongolia, Khoton Lake, 675X; 16, Mongolia, Tsagaaan Lake, 646X." figureDoi="http://doi.org/10.5281/zenodo.4761208" httpUri="https://zenodo.org/record/4761208/files/figure.png" pageId="8" pageNumber="52">Fig. 15</figureCitation>
). Overall, there is sufficient variability (e.g. forked vs. crenulated; tapered or not) to strongly suggest that the male cercus does not provide objective, diagnostic information to support 
<taxonomicName baseAuthorityName="Ricker" baseAuthorityYear="1952" box="[615,760,876,899]" class="Insecta" family="Nemouridae" genus="Nemoura" kingdom="Animalia" order="Plecoptera" pageId="8" pageNumber="52" phylum="Arthropoda" rank="species" species="trispinosa">
<emphasis box="[615,760,876,899]" italics="true" pageId="8" pageNumber="52">N. trispinosa</emphasis>
</taxonomicName>
as distinct from 
<taxonomicName authorityName=": Judson &amp; Nelson" authorityYear="2012" box="[348,454,912,935]" class="Insecta" family="Nemouridae" genus="Nemoura" kingdom="Animalia" order="Plecoptera" pageId="8" pageNumber="52" phylum="Arthropoda" rank="species" species="arctica">
<emphasis box="[348,454,912,935]" italics="true" pageId="8" pageNumber="52">N. arctica</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph blockId="8.[189,798,228,1907]" pageId="8" pageNumber="52">
<emphasis bold="true" box="[189,298,947,972]" pageId="8" pageNumber="52">
<emphasis bold="true" box="[189,291,947,971]" italics="true" pageId="8" pageNumber="52">Epiproct</emphasis>
.
</emphasis>
Males exhibit consistency with epiproct shape and characteristics across the Holarctic with only minor differences between individuals. In lateral aspect, the basal cushion occupies the anterior ca. ½ and is separated from the dorsal sclerite by smooth lateral areas (
<figureCitation box="[591,729,1128,1152]" captionStart="Figs" captionStartId="5.[189,238,1746,1770]" captionTargetBox="[324,1307,218,1700]" captionTargetId="figure-138@5.[324,1307,218,1700]" captionTargetPageId="5" captionText="Figs. 17–24. Nemoura arctica, epiproct, lateral view. 17, USA, Iowa, Twin Springs, 270X; 18, Canada, New Brunswick, Coombs Brook, 288X; 19, Canada, Northwest Territories, Thelon River, 285X; 20, Canada, Northwest Territories, Tuktoyaktuk, 328X; 21, Russia, Siberia, Magadan Oblask, 271X; 22, Norway, Buktved Garnvika, 286X; 23, Norway, Kautokeino, 316X; 24, Mongolia, Khoton Lake, 276X. AP = apical prong, BC = basal cushion, DS = dorsal sclerite, LA = lateral areas, VS = ventral sclerite." figureDoi="http://doi.org/10.5281/zenodo.4761210" httpUri="https://zenodo.org/record/4761210/files/figure.png" pageId="8" pageNumber="52">Figs. 17–24</figureCitation>
). The lateral areas vary in thickness but are consistently recurved slightly over the distal medial portion of the basal cushion. The dorsal sclerite appears scaly at high magnifications, especially at the distal tips (
<figureCitation box="[197,335,1307,1332]" captionStart-0="Figs" captionStart-1="Figs" captionStartId-0="9.[189,238,1409,1433]" captionStartId-1="10.[189,238,1416,1440]" captionTargetBox-0="[324,1307,254,1362]" captionTargetBox-1="[324,1307,254,1368]" captionTargetId-0="figure-247@9.[324,1307,254,1362]" captionTargetId-1="figure-236@10.[324,1307,254,1368]" captionTargetPageId-0="9" captionTargetPageId-1="10" captionText-0="Figs. 41–46. Nemoura arctica, epiproct, dorsal view. 41, USA, Iowa, Twin Springs, 584X; 42, USA, Illinois, Elgin, 519X; 43, USA, South Dakota, Burnt Fork, 669X; 44, Canada, Saskatchewan, Promontory Campground, 1056X; 45, Canada, New Brunswick, Coombs Brook, 770X; 46, Canada, Prince Edward Island, Balsam Hollow, 695X. AP = apical prong." captionText-1="Figs. 47–52. Nemoura arctica, epiproct, dorsal view. 47, Canada, Northwest Territories, Thelon River, 1161X; 48, Russia, Siberia, Magadan Oblask, 1204X; 49, Norway, Buktved Garnvika, 751X; 50, Norway, Kautokeino, 809X; 51, Mongolia, Khoton Lake, 598X; 52, Mongolia, Tsagaaan Lake, 969X. AP = apical prong." figureDoi-0="http://doi.org/10.5281/zenodo.4761218" figureDoi-1="http://doi.org/10.5281/zenodo.4761220" httpUri-0="https://zenodo.org/record/4761218/files/figure.png" httpUri-1="https://zenodo.org/record/4761220/files/figure.png" pageId="8" pageNumber="52">Figs. 41–52</figureCitation>
). The dorsal sclerite is open apically, exposing parallel, broad, hatchet-like apical prongs of the ventral sclerite (
<figureCitation box="[461,595,1379,1404]" captionStart-0="Figs" captionStart-1="Figs" captionStartId-0="6.[189,238,1748,1772]" captionStartId-1="7.[189,238,1748,1772]" captionTargetBox-0="[324,1307,218,1701]" captionTargetBox-1="[324,1307,218,1701]" captionTargetId-0="figure-149@6.[324,1307,218,1701]" captionTargetId-1="figure-137@7.[324,1307,218,1701]" captionTargetPageId-0="6" captionTargetPageId-1="7" captionText-0="Figs. 25–32. Nemoura arctica, epiproct, dorsal view. 25, USA, Iowa, Twin Springs, 141X; 26, USA, Illinois, Elgin, 165X; 27, USA, Ohio, spring, John Bryan State Park, 144X; 28, USA, Wisconsin, spring, 1 mile N Irvington, 132X; 29, USA, South Dakota, Burnt Fork, 140X; 30, USA, South Dakota, Burnt Fork, 229X; 31, Canada, Prince Edward Island, Balsam Hollow, 163X; 32, Canada, New Brunswick, Coombs Brook, 168X. AP = apical prong, BC = basal cushion, DS = dorsal sclerite, LA = lateral area, VS = ventral sclerite." captionText-1="Figs. 33–40. Nemoura arctica, epiproct, dorsal view. 33, Canada, Saskatchewan, Promontory Campground, 187X; 34, Canada, Northwest Territories, Thelon River, 282X; 35, Canada, Northwest Territories, Tuktoyaktuk, 267X; 36, Norway, Buktved Garnvika, 321X; 37, Norway, Kautokeino, 278X; 38, Russia, Siberia, Magadan Oblask, 206X; 39, Mongolia, Khoton Lake, 277X; 40, Mongolia, Tsagaaan Lake, 281X. AP = apical prong, BC = basal cushion, DS = dorsal sclerite, LA = lateral area, VS = ventral sclerite." figureDoi-0="http://doi.org/10.5281/zenodo.4761214" figureDoi-1="http://doi.org/10.5281/zenodo.4761216" httpUri-0="https://zenodo.org/record/4761214/files/figure.png" httpUri-1="https://zenodo.org/record/4761216/files/figure.png" pageId="8" pageNumber="52">Figs. 25–40</figureCitation>
) and prominent, scaly, apical prongs positioned ca. perpendicular to the ridges (
<figureCitation box="[344,489,1451,1476]" captionStart-0="Figs" captionStart-1="Figs" captionStartId-0="9.[189,238,1409,1433]" captionStartId-1="10.[189,238,1416,1440]" captionTargetBox-0="[324,1307,254,1362]" captionTargetBox-1="[324,1307,254,1368]" captionTargetId-0="figure-247@9.[324,1307,254,1362]" captionTargetId-1="figure-236@10.[324,1307,254,1368]" captionTargetPageId-0="9" captionTargetPageId-1="10" captionText-0="Figs. 41–46. Nemoura arctica, epiproct, dorsal view. 41, USA, Iowa, Twin Springs, 584X; 42, USA, Illinois, Elgin, 519X; 43, USA, South Dakota, Burnt Fork, 669X; 44, Canada, Saskatchewan, Promontory Campground, 1056X; 45, Canada, New Brunswick, Coombs Brook, 770X; 46, Canada, Prince Edward Island, Balsam Hollow, 695X. AP = apical prong." captionText-1="Figs. 47–52. Nemoura arctica, epiproct, dorsal view. 47, Canada, Northwest Territories, Thelon River, 1161X; 48, Russia, Siberia, Magadan Oblask, 1204X; 49, Norway, Buktved Garnvika, 751X; 50, Norway, Kautokeino, 809X; 51, Mongolia, Khoton Lake, 598X; 52, Mongolia, Tsagaaan Lake, 969X. AP = apical prong." figureDoi-0="http://doi.org/10.5281/zenodo.4761218" figureDoi-1="http://doi.org/10.5281/zenodo.4761220" httpUri-0="https://zenodo.org/record/4761218/files/figure.png" httpUri-1="https://zenodo.org/record/4761220/files/figure.png" pageId="8" pageNumber="52">Figs. 41–52</figureCitation>
). The prongs terminate laterally bearing two short, thick, grooved spines (e.g. 
<figureCitation box="[246,377,1523,1548]" captionStart="Figs" captionStartId="9.[189,238,1409,1433]" captionTargetBox="[324,1307,254,1362]" captionTargetId="figure-247@9.[324,1307,254,1362]" captionTargetPageId="9" captionText="Figs. 41–46. Nemoura arctica, epiproct, dorsal view. 41, USA, Iowa, Twin Springs, 584X; 42, USA, Illinois, Elgin, 519X; 43, USA, South Dakota, Burnt Fork, 669X; 44, Canada, Saskatchewan, Promontory Campground, 1056X; 45, Canada, New Brunswick, Coombs Brook, 770X; 46, Canada, Prince Edward Island, Balsam Hollow, 695X. AP = apical prong." figureDoi="http://doi.org/10.5281/zenodo.4761218" httpUri="https://zenodo.org/record/4761218/files/figure.png" pageId="8" pageNumber="52">Figs. 42, 45</figureCitation>
, 
<figureCitation box="[388,457,1523,1548]" captionStart="Figs" captionStartId="10.[189,238,1416,1440]" captionTargetBox="[324,1307,254,1368]" captionTargetId="figure-236@10.[324,1307,254,1368]" captionTargetPageId="10" captionText="Figs. 47–52. Nemoura arctica, epiproct, dorsal view. 47, Canada, Northwest Territories, Thelon River, 1161X; 48, Russia, Siberia, Magadan Oblask, 1204X; 49, Norway, Buktved Garnvika, 751X; 50, Norway, Kautokeino, 809X; 51, Mongolia, Khoton Lake, 598X; 52, Mongolia, Tsagaaan Lake, 969X. AP = apical prong." figureDoi="http://doi.org/10.5281/zenodo.4761220" httpUri="https://zenodo.org/record/4761220/files/figure.png" pageId="8" pageNumber="52">49, 52</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="8" pageNumber="52" type="discussion">
<paragraph blockId="8.[189,798,228,1907]" lastBlockId="8.[835,1444,228,972]" pageId="8" pageNumber="52">
<emphasis bold="true" box="[189,328,1559,1583]" pageId="8" pageNumber="52">Comments.</emphasis>
<taxonomicName baseAuthorityName="Ricker" baseAuthorityYear="1952" box="[356,591,1560,1583]" class="Insecta" family="Nemouridae" genus="Nemoura" kingdom="Animalia" order="Plecoptera" pageId="8" pageNumber="52" phylum="Arthropoda" rank="species" species="trispinosa">
<emphasis box="[356,591,1560,1583]" italics="true" pageId="8" pageNumber="52">Nemoura trispinosa</emphasis>
</taxonomicName>
is placed in synonymy with 
<taxonomicName authorityName=": Judson &amp; Nelson" authorityYear="2012" box="[392,504,1596,1619]" class="Insecta" family="Nemouridae" genus="Nemoura" kingdom="Animalia" order="Plecoptera" pageId="8" pageNumber="52" phylum="Arthropoda" rank="species" species="arctica">
<emphasis box="[392,504,1596,1619]" italics="true" pageId="8" pageNumber="52">N. arctica</emphasis>
</taxonomicName>
due to consistencies in epiproct characteristics across the Holarctic, particularly the paired apical prongs of the ventral sclerite (
<figureCitation box="[301,448,1703,1728]" captionStart-0="Figs" captionStart-1="Figs" captionStartId-0="9.[189,238,1409,1433]" captionStartId-1="10.[189,238,1416,1440]" captionTargetBox-0="[324,1307,254,1362]" captionTargetBox-1="[324,1307,254,1368]" captionTargetId-0="figure-247@9.[324,1307,254,1362]" captionTargetId-1="figure-236@10.[324,1307,254,1368]" captionTargetPageId-0="9" captionTargetPageId-1="10" captionText-0="Figs. 41–46. Nemoura arctica, epiproct, dorsal view. 41, USA, Iowa, Twin Springs, 584X; 42, USA, Illinois, Elgin, 519X; 43, USA, South Dakota, Burnt Fork, 669X; 44, Canada, Saskatchewan, Promontory Campground, 1056X; 45, Canada, New Brunswick, Coombs Brook, 770X; 46, Canada, Prince Edward Island, Balsam Hollow, 695X. AP = apical prong." captionText-1="Figs. 47–52. Nemoura arctica, epiproct, dorsal view. 47, Canada, Northwest Territories, Thelon River, 1161X; 48, Russia, Siberia, Magadan Oblask, 1204X; 49, Norway, Buktved Garnvika, 751X; 50, Norway, Kautokeino, 809X; 51, Mongolia, Khoton Lake, 598X; 52, Mongolia, Tsagaaan Lake, 969X. AP = apical prong." figureDoi-0="http://doi.org/10.5281/zenodo.4761218" figureDoi-1="http://doi.org/10.5281/zenodo.4761220" httpUri-0="https://zenodo.org/record/4761218/files/figure.png" httpUri-1="https://zenodo.org/record/4761220/files/figure.png" pageId="8" pageNumber="52">Figs. 41–52</figureCitation>
). In contrast, cercal spine characteristics are highly variable and do not provide objective, diagnostic information. Even males from the same geographic entity (e.g. 
<collectingCountry box="[189,305,1847,1871]" name="Mongolia" pageId="8" pageNumber="52">Mongolia</collectingCountry>
, 
<figureCitation box="[330,474,1847,1871]" captionStart="Figs" captionStartId="3.[189,238,1784,1808]" captionTargetBox="[324,1307,254,1737]" captionTargetId="figure-116@3.[324,1307,254,1737]" captionTargetPageId="3" captionText="Figs. 9–16. Nemoura arctica, cerci, distal view. 9, Canada, Manitoba, Warkworth Creek, 773X; 10, Canada, Northwest Territories, Thelon River, 636X; 11, Canada, Northwest Territories, Tuktoyaktuk, 612X; 12, Russia, Siberia, Magadan Oblask, 486X; 13, Norway, Buktved Garnvika, 443X; 14, Norway, Kautokeino, 431X; 15, Mongolia, Khoton Lake, 675X; 16, Mongolia, Tsagaaan Lake, 646X." figureDoi="http://doi.org/10.5281/zenodo.4761208" httpUri="https://zenodo.org/record/4761208/files/figure.png" pageId="8" pageNumber="52">Figs. 15–16</figureCitation>
) exhibit different cercal forms. 
<bibRefCitation author="Lillehammer, A." box="[298,560,1883,1907]" pageId="8" pageNumber="52" pagination="161 - 163" refId="ref7350" refString="Lillehammer, A. 1972 a. A new species of the genus Nemoura (Plecoptera) from Finnmark, North Norway. Norsk Entomologisk Tidsskrift, 19: 161 - 163. http: // www. entomologi. no / journals / nje / old / V 19" type="journal article" year="1972">Lillehammer (1972a</bibRefCitation>
, his 
<figureCitation box="[665,790,1883,1907]" captionStart="Figs" captionStartId="6.[189,238,1748,1772]" captionTargetBox="[324,1307,218,1701]" captionTargetId="figure-149@6.[324,1307,218,1701]" captionTargetPageId="6" captionText="Figs. 25–32. Nemoura arctica, epiproct, dorsal view. 25, USA, Iowa, Twin Springs, 141X; 26, USA, Illinois, Elgin, 165X; 27, USA, Ohio, spring, John Bryan State Park, 144X; 28, USA, Wisconsin, spring, 1 mile N Irvington, 132X; 29, USA, South Dakota, Burnt Fork, 140X; 30, USA, South Dakota, Burnt Fork, 229X; 31, Canada, Prince Edward Island, Balsam Hollow, 163X; 32, Canada, New Brunswick, Coombs Brook, 168X. AP = apical prong, BC = basal cushion, DS = dorsal sclerite, LA = lateral area, VS = ventral sclerite." figureDoi="http://doi.org/10.5281/zenodo.4761214" httpUri="https://zenodo.org/record/4761214/files/figure.png" pageId="8" pageNumber="52">Fig. 25.4</figureCitation>
) previously illustrated several different cercal forms from 
<collectingCountry box="[899,999,265,289]" name="Norway" pageId="8" pageNumber="52">Norway</collectingCountry>
.
</paragraph>
<paragraph blockId="8.[835,1444,228,972]" pageId="8" pageNumber="52">
<emphasis bold="true" box="[835,1064,299,324]" pageId="8" pageNumber="52">Geographic notes</emphasis>
. Holarctic: Scandinavia east across Asia; east to Alaska, Yukon, Northwest Territories and Nunavut. South in Europe to 
<collectingCountry box="[1368,1442,372,396]" name="Latvia" pageId="8" pageNumber="52">Latvia</collectingCountry>
and in far eastern Asia to 
<collectingCountry box="[1172,1285,408,432]" name="Mongolia" pageId="8" pageNumber="52">Mongolia</collectingCountry>
and Siberia. South in North America to Wyoming and South Dakota east across the Great Lakes region to Atlantic 
<collectingCountry box="[969,1062,516,540]" name="Canada" pageId="8" pageNumber="52">Canada</collectingCountry>
, with relictual southern populations in Ohio, Pennsylvania, and West Virginia. Additional notable references include 
<bibRefCitation author="Ricker, W. E." box="[835,984,624,648]" pageId="8" pageNumber="52" pagination="174 - 185" refId="ref7975" refString="Ricker, W. E. 1944. Some Plecoptera from the far north. The Canadian Entomologist, 76: 174 - 185. https: // doi. org / 10.4039 / Ent 76174 - 9" type="journal article" year="1944">Ricker (1944</bibRefCitation>
, 
<bibRefCitation author="Ricker, W. E." box="[996,1053,624,648]" pageId="8" pageNumber="52" pagination="50 - 71" refId="ref8054" refString="Ricker, W. E. 1964. Distribution of Canadian stoneflies. Gewasser und Abwasser, 34 / 35: 50 - 71." type="journal article" year="1964">1964</bibRefCitation>
), 
<bibRefCitation author="Brinck, P." box="[1073,1231,624,648]" pageId="8" pageNumber="52" pagination="1 - 11" refId="ref6336" refString="Brinck, P. 1958. On some stoneflies recorded from Nowaya Zemlya. Norske videnskaps-akademi, Matematisk-naturvidenskapelig klasse, 2: 1 - 11." type="journal article" year="1958">Brinck (1958)</bibRefCitation>
, 
<bibRefCitation author="Ulfstrand, S." box="[1244,1438,624,648]" pageId="8" pageNumber="52" pagination="145 - 165" refId="ref8421" refString="Ulfstrand, S. 1969. Ephemeroptera and Plecoptera from the river Vindelalven in Swedish Lapland. Entomologisk Tidskrift, 90: 145 - 165." type="journal article" year="1969">Ulfstrand (1969)</bibRefCitation>
, 
<bibRefCitation author="Jewett, S. G." box="[835,990,660,684]" pageId="8" pageNumber="52" pagination="189 - 192" refId="ref7070" refString="Jewett, S. G. 1971. Some Alaskan stoneflies (Plecoptera). The Pan-Pacific Entomologist, 47: 189 - 192." type="journal article" year="1971">Jewett (1971)</bibRefCitation>
, Lillehammer (1974, 1985, 1986, 1988), 
<bibRefCitation author="Harper, P. P." box="[835,1013,696,720]" pageId="8" pageNumber="52" pagination="309 - 356" refId="ref6835" refString="Harper, P. P. 1973. Life histories of Nemouridae and Leuctridae in southern Ontario (Plecoptera). Hydrobiologia, 41: 309 - 356." type="journal article" year="1973">Harper (1973)</bibRefCitation>
, 
<bibRefCitation author="Flannagan, P. M. &amp; E Flannagan" box="[1037,1438,696,720]" pageId="8" pageNumber="52" pagination="1 - 79" refId="ref6720" refString="Flannagan, P. M. &amp; J. E Flannagan. 1982. Present distribution and the post-glacial origin of the Ephemeroptera, Plecoptera, and Trichoptera of Manitoba. Manitoba Department of Natural Resources and Fisheries Technical Report 82: 1 - 79." type="book chapter" year="1982">Flannagan &amp; Flannagan (1982)</bibRefCitation>
, 
<bibRefCitation author="Burton, D. K." box="[835,1008,732,756]" pageId="8" pageNumber="52" pagination="39 - 51" refId="ref6362" refString="Burton, D. K. 1984. Distribution of Manitoba stoneflies (Plecoptera). Proceedings of the Entomological Society of Manitoba, 40: 39 - 51." type="journal article" year="1984">Burton (1984)</bibRefCitation>
, 
<bibRefCitation author="Stewart, K. W. &amp; R. L. Hassage &amp; S. J. Holder &amp; M. W. Oswood" box="[1030,1297,732,756]" pageId="8" pageNumber="52" pagination="207 - 214" refId="ref8195" refString="Stewart, K. W., R. L. Hassage, S. J. Holder, &amp; M. W. Oswood. 1990. Life cycles of six stonefly species (Plecoptera) in subarctic and arctic Alaska streams. Annals of the Entomological Society of America, 83: 207 - 214." type="journal article" year="1990">Stewart et al. (1990)</bibRefCitation>
, 
<bibRefCitation author="Harper, P. P. &amp; W. E. Ricker" pageId="8" pageNumber="52" pagination="43 - 66" refId="ref6864" refString="Harper, P. P. &amp; W. E. Ricker. 1994. Distribution of Ontario stoneflies (Plecoptera). Proceedings of the Entomological Society of Ontario, 125: 43 - 66." type="journal article" year="1994">Harper &amp; Ricker (1994)</bibRefCitation>
, 
<bibRefCitation author="Stewart, K. W. &amp; W. E. Ricker" box="[1010,1301,768,792]" pageId="8" pageNumber="52" pagination="201 - 222" refId="ref8115" refString="Stewart, K. W. &amp; W. E. Ricker. 1997. Stoneflies (Plecoptera) of the Yukon. Pages 201 - 222. In: Danks, H. V. &amp; J. A. Downes (editors). Insects of the Yukon. Biological Survey of Canada (Terrestrial Arthropods), Ottawa. http: // www. biology. ualberta. ca / bsc / pdf / stewart." type="book chapter" year="1997">Stewart &amp; Ricker (1997)</bibRefCitation>
, 
<bibRefCitation author="Huntsman, B. O. &amp; R. W. Baumann &amp; B. C. Kondratieff" pageId="8" pageNumber="52" pagination="1 - 17" refId="ref6922" refString="Huntsman, B. O., R. W. Baumann, &amp; B. C. Kondratieff 1999. Stoneflies (Plecoptera) of the Black Hills of South Dakota and Wyoming, USA: distribution and zoogeographic affinities. Great Basin Naturalist, 59: 1 - 17." type="journal article" year="1999">Huntsman et al. (1999)</bibRefCitation>
, 
<bibRefCitation author="Teslenko, V. &amp; N. Bazova" box="[992,1307,804,828]" pageId="8" pageNumber="52" pagination="1059 - 1068" refId="ref8326" refString="Teslenko, V. &amp; N. Bazova. 2009. On the stonefly (Plecoptera) fauna of the transfrontier Selenga River basin. Entomological Review, 89: 1059 - 1068." type="journal article" year="2009">Teslenko &amp; Bazova (2009)</bibRefCitation>
, 
<bibRefCitation author="Walters, K. R. &amp; Sformo, B. M. &amp; Barnes, J. G. &amp; Duman" pageId="8" pageNumber="52" pagination="305 - 312" refId="ref8469" refString="Walters, K. R., T, Sformo, B. M. Barnes, J. G. &amp; Duman. 2009. Freeze tolerance in an arctic Alaska stonefly. Journal of Experimental Biology, 212: 305 - 312." type="journal article" year="2009">Walters et al. (2009)</bibRefCitation>
, 
<bibRefCitation author="Zhou, X. &amp; L. M. Jacobus &amp; R. E. DeWalt &amp; S. J. Adamowicz &amp; P. D. N. Hebert" box="[976,1215,840,864]" pageId="8" pageNumber="52" pagination="814 - 837" refId="ref8681" refString="Zhou, X., L. M. Jacobus, R. E. DeWalt, S. J. Adamowicz, &amp; P. D. N. Hebert. 2010. Ephemeroptera, Plecoptera, and Trichoptera fauna of Churchill (Manitoba, Canada): insights into biodiversity patterns from DNA barcoding. Journal of the North American Benthological Society, 29, 814 - 837. https: // doi. org / 10.1899 / 09 - 121.1" type="journal article" year="2010">Zhou et al. (2010)</bibRefCitation>
, 
<bibRefCitation author="Boumans, L." box="[1237,1436,840,864]" pageId="8" pageNumber="52" pagination="280 - 290" refId="ref6149" refString="Boumans, L. 2011. The Plecoptera collection at the Natural History Museum in Oslo. Illiesia, 7: 280 - 290." type="journal article" year="2011">Boumans (2011)</bibRefCitation>
, 
<bibRefCitation author="Boumans, L. &amp; J. E. Brittain" box="[835,1180,876,900]" pageId="8" pageNumber="52" pagination="196 - 215" refId="ref6243" refString="Boumans, L. &amp; J. E. Brittain. 2012. Faunistics of stoneflies (Plecoptera) in Finnmark, northern Norway, including DNA barcoding of Nemouridae. Norwegian Journal of Entomology, 59: 196 - 215." type="journal article" year="2012">Boumans &amp; Brittain (2012)</bibRefCitation>
, 
<bibRefCitation author="Surenkhorloo, P. &amp; R. Samiya &amp; J. Slowik &amp; M. Muhlenberg" pageId="8" pageNumber="52" pagination="245 - 249" refId="ref8253" refString="Surenkhorloo, P., R. Samiya, J. Slowik, &amp; M. Muhlenberg. 2012. Some taxonomic records of aquatic insects in the Eroo River Basin (West Khentii, Northern Mongolia). Erforschung biologischer Ressourcen der Mongolei, 12: 245 - 249." type="journal article" year="2012">
Surenkhorloo 
<emphasis box="[1381,1444,876,900]" italics="true" pageId="8" pageNumber="52">et al.</emphasis>
(2012)
</bibRefCitation>
, 
<bibRefCitation author="Dosdall, L. M. &amp; D. J. Giberson" box="[930,1275,912,936]" pageId="8" pageNumber="52" pagination="201 - 229" refId="ref6591" refString="Dosdall, L. M. &amp; D. J. Giberson. 2014. Stoneflies (Plecoptera) of the Canadian Prairie Provinces. Pages 201 - 229. In: Carcamo, H. A. &amp; D. J. Giberson (editors). Arthropods of Canadian Grasslands (Volume 3): Biodiversity and Systematics Part 1. Biological Survey of Canada, Ottawa. doi / 10.3752 / 9780968932162. ch 7" type="book chapter" year="2014">Dosdall &amp; Giberson (2014)</bibRefCitation>
, 
<bibRefCitation author="Kendrick, M. R. &amp; A. D. Huryn" pageId="8" pageNumber="52" pagination="275 - 285" refId="ref7180" refString="Kendrick, M. R. &amp; A. D. Huryn. 2014. The Plecoptera and Trichoptera of the Arctic North Slope of Alaska. Western North American Naturalist 74: 275 - 285. https: // doi. org / 10.3398 / 064.074.0303" type="journal article" year="2014">Kendrick &amp; Huryn (2014)</bibRefCitation>
, and 
<bibRefCitation author="Potikha, E. V." box="[1057,1229,948,972]" pageId="8" pageNumber="52" pagination="22 - 31" refId="ref7824" refString="Potikha, E. V. 2015. A taxonomic list of the mayflies, stoneflies and caddisflies (Insecta: Ephemeroptera, Plecoptera and Trichoptera) of the Sikhote-Alin Biosphere Reserve. Achievements in the Life Sciences, 9: 22 - 31. https: // ac. els-cdn. com / S 2078152015000346 / 1 -" type="journal article" year="2015">Potikha (2015)</bibRefCitation>
.
</paragraph>
</subSubSection>
</subSection>
</subSubSection>
</treatment>
</document>