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<document id="145B4C457B92C071B4D545943FD1A642" ID-CLB-Dataset="5703" ID-DOI="10.11646/phytotaxa.87.1.2" ID-GBIF-Dataset="4542f05d-992b-48a2-8b6e-c8128357d5e0" ID-ISSN="1179-3163" ID-Zenodo-Dep="5071814" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1625516158268" checkinUser="felipe" docAuthor="Němcová, Yvonne &amp; Kreidlová, Jana" docDate="2013" docId="039E690FFFA0291D3F8CC71BFA8635BC" docLanguage="en" docName="Phytotaxa.87.1.11-18.pdf" docOrigin="Phytotaxa 87 (1)" docSource="http://dx.doi.org/10.11646/phytotaxa.87.1.2" docStyle="DocumentStyle:13F189B31CB4CCCE9197361A4E100104.3:Phytotaxa.2011-2013.journal_article" docStyleId="13F189B31CB4CCCE9197361A4E100104" docStyleName="Phytotaxa.2011-2013.journal_article" docStyleVersion="3" docTitle="Mallomonas temonis Nemcova 2013, sp. nov." docType="treatment" docVersion="7" lastPageNumber="14" masterDocId="FFA71177FFA1291E3F1BC25FFFA8374A" masterDocTitle="Two new species of Mallomonas (Chrysophyceae: Synurales): Mallomonas temonis and Mallomonas divida" masterLastPageNumber="18" masterPageNumber="11" pageNumber="12" updateTime="1699025102589" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="BD912C8B78F9CE086414693B02CB22B3">Two new species of Mallomonas (Chrysophyceae: Synurales): Mallomonas temonis and Mallomonas divida</mods:title>
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<treatment id="039E690FFFA0291D3F8CC71BFA8635BC" ID-DOI="http://doi.org/10.5281/zenodo.5074080" ID-GBIF-Taxon="183326362" ID-Zenodo-Dep="5074080" LSID="urn:lsid:plazi:treatment:039E690FFFA0291D3F8CC71BFA8635BC" httpUri="http://treatment.plazi.org/id/039E690FFFA0291D3F8CC71BFA8635BC" lastPageId="3" lastPageNumber="14" pageId="1" pageNumber="12">
<subSubSection id="C32D8B92FFA0291F3F8CC71BFD533214" box="[151,763,1348,1374]" pageId="1" pageNumber="12" type="nomenclature">
<paragraph id="8B88D819FFA0291F3F8CC71BFD533214" blockId="1.[151,763,1348,1374]" box="[151,763,1348,1374]" pageId="1" pageNumber="12">
<heading id="D0C06F75FFA0291F3F8CC71BFD533214" box="[151,763,1348,1374]" fontSize="11" level="2" pageId="1" pageNumber="12" reason="3">
<taxonomicName id="4C37A39AFFA0291F3F8CC71BFDAD3214" authority="Nemcova" authorityName="Nemcova" authorityYear="2013" box="[151,517,1348,1374]" class="Chrysophyceae" family="Mallomonadaceae" genus="Mallomonas" higherTaxonomySource="GBIF" kingdom="Chromista" order="Synurales" pageId="1" pageNumber="12" phylum="Ochrophyta" rank="species" species="temonis" status="sp. nov.">
<emphasis id="B943040BFFA0291F3F8CC71BFE383214" bold="true" box="[151,400,1348,1374]" italics="true" pageId="1" pageNumber="12">Mallomonas temonis</emphasis>
Nĕmcová
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<emphasis id="B943040BFFA0291F3D17C71BFDF63217" box="[524,606,1348,1373]" italics="true" pageId="1" pageNumber="12">sp. nov</emphasis>
.
</taxonomicNameLabel>
(
<figureCitation id="130CC49CFFA0291F3D6FC71BFD5B3214" box="[628,755,1348,1374]" captionStart="FIGURE 1" captionStartId="1.[151,245,1798,1819]" captionText="FIGURE 1. Mallomonas temonis. AB, asymmetric apical scales with an eccentrically situated dome; CD, oval body scales; EF, rear scales; GH, a body scale bearing a bristle; G, a scale viewed in a transmission electron microscope; H, the same scale viewed in a scanning electron microscope (SEM); I, a body scale with a bristle anchored to the dome, a rounded rearmost scale and a less silicified scale marked with closely spaced pores; J, a slightly curved bristle with a sharp tip; K, a group of body scales including one rearmost scale; L, a group of body scales viewed in SEM. Note that the side of the scale adjacent to the plasma membrane is smooth. Bar = 0.5 µm, if not stated otherwise. AF and I are shown at the same magnification." pageId="1" pageNumber="12">Figs 1AL</figureCitation>
)
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C32D8B92FFA0291D3F8CC7CEFD8435E2" lastPageId="3" lastPageNumber="14" pageId="1" pageNumber="12" type="materials_examined">
<paragraph id="8B88D819FFA0291F3F8CC7CEFC343282" blockId="1.[151,1436,1425,1481]" pageId="1" pageNumber="12">
<materialsCitation id="3B5FD244FFA0291F3F8CC7CEFC303282" ID-GBIF-Occurrence="3314055302" collectingDate="2010-08-20" collectorName="Y. Nemcova" country="South Africa" elevation="1064" latitude="-33.958317" location="Table Mountain" longLatPrecision="1" longitude="18.4047" pageId="1" pageNumber="12" specimenCount="1" stateProvince="Western Cape" typeStatus="holotype">
<emphasis id="B943040BFFA0291F3F8CC7CEFF5A32E3" bold="true" box="[151,242,1425,1449]" pageId="1" pageNumber="12">
<typeStatus id="548C66BBFFA0291F3F8CC7CEFF7932E3" box="[151,209,1425,1449]" pageId="1" pageNumber="12">Type</typeStatus>
:—
</emphasis>
<collectingCountry id="F3209889FFA0291F3FE9C7CEFE1E32E2" box="[242,438,1425,1448]" name="South Africa" pageId="1" pageNumber="12">SOUTH AFRICA</collectingCountry>
.
<collectingRegion id="49F316FBFFA0291F3EA4C7CEFDFF32E2" box="[447,599,1425,1448]" country="South Africa" name="Western Cape" pageId="1" pageNumber="12">Western Cape</collectingRegion>
: unnamed small pool at
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,
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,
<geoCoordinate id="EE03BEDEFFA0291F3BB7C7CEFAEA32E2" box="[1196,1346,1425,1448]" degrees="18" direction="east" minutes="24.282" orientation="longitude" pageId="1" pageNumber="12" precision="1" value="18.4047">18° 24.282´E</geoCoordinate>
,
<elevation id="001A3F2AFFA0291F3A57C7CEFF513282" metricMagnitude="3" metricUnit="m" metricValue="1.064" pageId="1" pageNumber="12" unit="m" value="1064.0">
<quantity id="4CCF75FCFFA0291F3A57C7CEFA3432E2" box="[1356,1436,1425,1448]" metricMagnitude="3" metricUnit="m" metricValue="1.064" pageId="1" pageNumber="12" unit="m" value="1064.0">1064 m</quantity>
a.s.l.
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, sample collected by
<collectorName id="26C2BDCFFFA0291F3EFBC7EEFDCE3282" box="[480,614,1457,1481]" pageId="1" pageNumber="12">Y. Nĕmcová</collectorName>
,
<date id="FF89FED9FFA0291F3D6AC7EEFCB63282" box="[625,798,1457,1480]" pageId="1" pageNumber="12" value="2010-08-20">
<collectingDate id="EFCD0731FFA0291F3D6AC7EEFCB63282" box="[625,798,1457,1480]" pageId="1" pageNumber="12" value="2010-08-20">20 August 2010</collectingDate>
</date>
.
<figureCitation id="130CC49CFFA0291F3C33C7EEFC303282" box="[808,920,1457,1480]" captionStart="FIGURE 1" captionStartId="1.[151,245,1798,1819]" captionText="FIGURE 1. Mallomonas temonis. AB, asymmetric apical scales with an eccentrically situated dome; CD, oval body scales; EF, rear scales; GH, a body scale bearing a bristle; G, a scale viewed in a transmission electron microscope; H, the same scale viewed in a scanning electron microscope (SEM); I, a body scale with a bristle anchored to the dome, a rounded rearmost scale and a less silicified scale marked with closely spaced pores; J, a slightly curved bristle with a sharp tip; K, a group of body scales including one rearmost scale; L, a group of body scales viewed in SEM. Note that the side of the scale adjacent to the plasma membrane is smooth. Bar = 0.5 µm, if not stated otherwise. AF and I are shown at the same magnification." pageId="1" pageNumber="12">Figure 1D</figureCitation>
</materialsCitation>
.
</paragraph>
<paragraph id="8B88D819FFA0291D3F8CC7A7FE3035C8" blockId="1.[151,1436,1528,1708]" lastBlockId="3.[151,1436,152,758]" lastPageId="3" lastPageNumber="14" pageId="1" pageNumber="12">
Three
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of scale may be discerned including apical, body, and rear scales; however, a continuous transition between body and rear scales exists. Apical scales (2.6
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3.2 ×
<quantity id="4CCF75FCFFA0291F3CC0C441FBCA3172" box="[987,1122,1566,1592]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" metricValueMax="2.5" metricValueMin="1.9" pageId="1" pageNumber="12" unit="mm" value="2.2" valueMax="2.5" valueMin="1.9">
1.9
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2.5 µm
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) are broad and remarkably asymmetric, with one lateral edge shorter than the other. A large, broad, flat dome is eccentrically situated and its posterior part is extended, which forms a lip. The base plate is marked with closely spaced pores, which are clearly visible on the less silicified scale (
<figureCitation id="130CC49CFFA0291F3D65C4CDFD6631E6" box="[638,718,1682,1708]" captionStart="FIGURE 1" captionStartId="1.[151,245,1798,1819]" captionText="FIGURE 1. Mallomonas temonis. AB, asymmetric apical scales with an eccentrically situated dome; CD, oval body scales; EF, rear scales; GH, a body scale bearing a bristle; G, a scale viewed in a transmission electron microscope; H, the same scale viewed in a scanning electron microscope (SEM); I, a body scale with a bristle anchored to the dome, a rounded rearmost scale and a less silicified scale marked with closely spaced pores; J, a slightly curved bristle with a sharp tip; K, a group of body scales including one rearmost scale; L, a group of body scales viewed in SEM. Note that the side of the scale adjacent to the plasma membrane is smooth. Bar = 0.5 µm, if not stated otherwise. AF and I are shown at the same magnification." pageId="1" pageNumber="12">Fig. 1I</figureCitation>
). A scale is surrounded by a submarginal rib curving toward the dome. At the posterior part of the scale, a strong, more or less transversal rib connects the lateral parts of the submarginal rib. A secondary reticulum of smaller meshes with more delicate ribs may be developed under the transversal rib (
<figureCitation id="130CC49CFFA2291D3ED1C2BAFD8237B5" box="[458,554,229,255]" captionStart="FIGURE 1" captionStartId="1.[151,245,1798,1819]" captionText="FIGURE 1. Mallomonas temonis. AB, asymmetric apical scales with an eccentrically situated dome; CD, oval body scales; EF, rear scales; GH, a body scale bearing a bristle; G, a scale viewed in a transmission electron microscope; H, the same scale viewed in a scanning electron microscope (SEM); I, a body scale with a bristle anchored to the dome, a rounded rearmost scale and a less silicified scale marked with closely spaced pores; J, a slightly curved bristle with a sharp tip; K, a group of body scales including one rearmost scale; L, a group of body scales viewed in SEM. Note that the side of the scale adjacent to the plasma membrane is smooth. Bar = 0.5 µm, if not stated otherwise. AF and I are shown at the same magnification." pageId="3" pageNumber="14">Fig. 1A</figureCitation>
). The submarginal areas possess struts (occasionally bifurcated) radiating from the submarginal rib; the struts do not reach the proximal border. The proximal border surrounds approximately half of the scale outline (
<figureCitation id="130CC49CFFA2291D3D94C36DFCB63606" box="[655,798,306,332]" captionStart="FIGURE 1" captionStartId="1.[151,245,1798,1819]" captionText="FIGURE 1. Mallomonas temonis. AB, asymmetric apical scales with an eccentrically situated dome; CD, oval body scales; EF, rear scales; GH, a body scale bearing a bristle; G, a scale viewed in a transmission electron microscope; H, the same scale viewed in a scanning electron microscope (SEM); I, a body scale with a bristle anchored to the dome, a rounded rearmost scale and a less silicified scale marked with closely spaced pores; J, a slightly curved bristle with a sharp tip; K, a group of body scales including one rearmost scale; L, a group of body scales viewed in SEM. Note that the side of the scale adjacent to the plasma membrane is smooth. Bar = 0.5 µm, if not stated otherwise. AF and I are shown at the same magnification." pageId="3" pageNumber="14">Figs 1AB</figureCitation>
). Body scales (3.2
<emphasis id="B943040BFFA2291D3B1DC36DFBBD3606" bold="true" box="[1030,1045,306,332]" pageId="3" pageNumber="14"></emphasis>
3.9 ×
<quantity id="4CCF75FCFFA2291D3B7BC36DFB583606" box="[1120,1264,306,332]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" metricValueMax="2.7" metricValueMin="2.1" pageId="3" pageNumber="14" unit="mm" value="2.4" valueMax="2.7" valueMin="2.1">
2.1
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2.7 µm
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) are oval and essentially bilaterally symmetrical (
<figureCitation id="130CC49CFFA2291D3D43C306FD4E3639" box="[600,742,345,371]" captionStart="FIGURE 1" captionStartId="1.[151,245,1798,1819]" captionText="FIGURE 1. Mallomonas temonis. AB, asymmetric apical scales with an eccentrically situated dome; CD, oval body scales; EF, rear scales; GH, a body scale bearing a bristle; G, a scale viewed in a transmission electron microscope; H, the same scale viewed in a scanning electron microscope (SEM); I, a body scale with a bristle anchored to the dome, a rounded rearmost scale and a less silicified scale marked with closely spaced pores; J, a slightly curved bristle with a sharp tip; K, a group of body scales including one rearmost scale; L, a group of body scales viewed in SEM. Note that the side of the scale adjacent to the plasma membrane is smooth. Bar = 0.5 µm, if not stated otherwise. AF and I are shown at the same magnification." pageId="3" pageNumber="14">Figs 1CD</figureCitation>
). The submarginal rib fuses with the dome forming the continuous entity (
<figureCitation id="130CC49CFFA2291D3E69C3DFFDAC36D0" box="[370,516,384,410]" captionStart="FIGURE 1" captionStartId="1.[151,245,1798,1819]" captionText="FIGURE 1. Mallomonas temonis. AB, asymmetric apical scales with an eccentrically situated dome; CD, oval body scales; EF, rear scales; GH, a body scale bearing a bristle; G, a scale viewed in a transmission electron microscope; H, the same scale viewed in a scanning electron microscope (SEM); I, a body scale with a bristle anchored to the dome, a rounded rearmost scale and a less silicified scale marked with closely spaced pores; J, a slightly curved bristle with a sharp tip; K, a group of body scales including one rearmost scale; L, a group of body scales viewed in SEM. Note that the side of the scale adjacent to the plasma membrane is smooth. Bar = 0.5 µm, if not stated otherwise. AF and I are shown at the same magnification." pageId="3" pageNumber="14">Figs 1H, 1L</figureCitation>
). The strong transversal rib crosses the shield in approximately the lower half to third (
<figureCitation id="130CC49CFFA2291D3FE5C3F9FE2B368A" box="[254,387,422,448]" captionStart="FIGURE 1" captionStartId="1.[151,245,1798,1819]" captionText="FIGURE 1. Mallomonas temonis. AB, asymmetric apical scales with an eccentrically situated dome; CD, oval body scales; EF, rear scales; GH, a body scale bearing a bristle; G, a scale viewed in a transmission electron microscope; H, the same scale viewed in a scanning electron microscope (SEM); I, a body scale with a bristle anchored to the dome, a rounded rearmost scale and a less silicified scale marked with closely spaced pores; J, a slightly curved bristle with a sharp tip; K, a group of body scales including one rearmost scale; L, a group of body scales viewed in SEM. Note that the side of the scale adjacent to the plasma membrane is smooth. Bar = 0.5 µm, if not stated otherwise. AF and I are shown at the same magnification." pageId="3" pageNumber="14">Figs 1KL</figureCitation>
). Remnants of the reticulum of smaller meshes are sometimes visible on the shield (
<figureCitation id="130CC49CFFA2291D3A77C3F9FF1636AD" captionStart="FIGURE 1" captionStartId="1.[151,245,1798,1819]" captionText="FIGURE 1. Mallomonas temonis. AB, asymmetric apical scales with an eccentrically situated dome; CD, oval body scales; EF, rear scales; GH, a body scale bearing a bristle; G, a scale viewed in a transmission electron microscope; H, the same scale viewed in a scanning electron microscope (SEM); I, a body scale with a bristle anchored to the dome, a rounded rearmost scale and a less silicified scale marked with closely spaced pores; J, a slightly curved bristle with a sharp tip; K, a group of body scales including one rearmost scale; L, a group of body scales viewed in SEM. Note that the side of the scale adjacent to the plasma membrane is smooth. Bar = 0.5 µm, if not stated otherwise. AF and I are shown at the same magnification." pageId="3" pageNumber="14">Fig. 1G</figureCitation>
). Rear scales (2.1
<emphasis id="B943040BFFA2291D3E83C392FE0F36AD" bold="true" box="[408,423,461,487]" pageId="3" pageNumber="14"></emphasis>
2.7 ×
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1.4
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1.9 µm
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) are smaller and more rhomboidal compared to body scales (
<figureCitation id="130CC49CFFA2291D3A7CC392FF733544" captionStart="FIGURE 1" captionStartId="1.[151,245,1798,1819]" captionText="FIGURE 1. Mallomonas temonis. AB, asymmetric apical scales with an eccentrically situated dome; CD, oval body scales; EF, rear scales; GH, a body scale bearing a bristle; G, a scale viewed in a transmission electron microscope; H, the same scale viewed in a scanning electron microscope (SEM); I, a body scale with a bristle anchored to the dome, a rounded rearmost scale and a less silicified scale marked with closely spaced pores; J, a slightly curved bristle with a sharp tip; K, a group of body scales including one rearmost scale; L, a group of body scales viewed in SEM. Note that the side of the scale adjacent to the plasma membrane is smooth. Bar = 0.5 µm, if not stated otherwise. AF and I are shown at the same magnification." pageId="3" pageNumber="14">Figs 1EF</figureCitation>
). The rearmost scales (1.4
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1.6 ×
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0.8
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1.1 µm
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) are oval, lack a dome, and have a rudimentary reticulum formed by delicate ribs. Bristles (
<quantity id="4CCF75FCFFA2291D3D01C045FD0A357E" box="[538,674,538,564]" metricMagnitude="-3" metricUnit="m" metricValue="3.35" metricValueMax="3.5" metricValueMin="3.2" pageId="3" pageNumber="14" unit="mm" value="3.35" valueMax="3.5" valueMin="3.2">
3.2
<emphasis id="B943040BFFA2291D3D5BC045FDE7357E" bold="true" box="[576,591,538,564]" pageId="3" pageNumber="14"></emphasis>
3.5 µm
</quantity>
) are smooth, slightly curved and attenuated, with a sharp tip. The foot of the bristle is thin (
<figureCitation id="130CC49CFFA2291D3ED3C01EFDEE3511" box="[456,582,576,603]" captionStart="FIGURE 1" captionStartId="1.[151,245,1798,1819]" captionText="FIGURE 1. Mallomonas temonis. AB, asymmetric apical scales with an eccentrically situated dome; CD, oval body scales; EF, rear scales; GH, a body scale bearing a bristle; G, a scale viewed in a transmission electron microscope; H, the same scale viewed in a scanning electron microscope (SEM); I, a body scale with a bristle anchored to the dome, a rounded rearmost scale and a less silicified scale marked with closely spaced pores; J, a slightly curved bristle with a sharp tip; K, a group of body scales including one rearmost scale; L, a group of body scales viewed in SEM. Note that the side of the scale adjacent to the plasma membrane is smooth. Bar = 0.5 µm, if not stated otherwise. AF and I are shown at the same magnification." pageId="3" pageNumber="14">
Figs 1G
<emphasis id="B943040BFFA2291D3D32C01FFD903510" bold="true" box="[553,568,576,602]" pageId="3" pageNumber="14"></emphasis>
J
</figureCitation>
). Only
<specimenCount id="9D311390FFA2291D3DBBC01EFCAE3511" box="[672,774,577,603]" pageId="3" pageNumber="14" type="generic" typeStatus="type">one type</specimenCount>
of bristle was observed. Cyst unknown. Cell shape and dimensions unknown.
</paragraph>
<caption id="DF488891FFA0291F3F8CC559FBF1308F" pageId="1" pageNumber="12" startId="1.[151,245,1798,1819]">
<paragraph id="8B88D819FFA0291F3F8CC559FBF1308F" blockId="1.[151,1438,1798,1989]" pageId="1" pageNumber="12">
<emphasis id="B943040BFFA0291F3F8CC559FEA63051" bold="true" box="[151,270,1798,1819]" pageId="1" pageNumber="12">FIGURE 1.</emphasis>
<emphasis id="B943040BFFA0291F3E0FC559FE4E3051" box="[276,486,1798,1819]" italics="true" pageId="1" pageNumber="12">
<taxonomicName id="4C37A39AFFA0291F3E0FC559FE4A3051" authorityName="Nemcova" authorityYear="2013" box="[276,482,1798,1819]" class="Chrysophyceae" family="Mallomonadaceae" genus="Mallomonas" higherTaxonomySource="GBIF" kingdom="Chromista" order="Synurales" pageId="1" pageNumber="12" phylum="Ochrophyta" rank="species" species="temonis">Mallomonas temonis</taxonomicName>
.
</emphasis>
A
<emphasis id="B943040BFFA0291F3EE6C559FDA13051" bold="true" box="[509,521,1798,1819]" pageId="1" pageNumber="12"></emphasis>
B, asymmetric apical scales with an eccentrically situated dome; C
<emphasis id="B943040BFFA0291F3B8DC559FB0A3051" bold="true" box="[1174,1186,1798,1819]" pageId="1" pageNumber="12"></emphasis>
D, oval body scales; E
<emphasis id="B943040BFFA0291F3A65C559FA223051" bold="true" box="[1406,1418,1798,1819]" pageId="1" pageNumber="12"></emphasis>
F, rear scales; G
<emphasis id="B943040BFFA0291F3E01C578FE8E3076" bold="true" box="[282,294,1831,1852]" pageId="1" pageNumber="12"></emphasis>
H, a body scale bearing a bristle; G, a scale viewed in a transmission electron microscope; H, the same scale viewed in a scanning electron microscope (SEM); I, a body scale with a bristle anchored to the dome, a rounded rearmost scale and a less silicified scale marked with closely spaced pores; J, a slightly curved bristle with a sharp tip; K, a group of body scales including one rearmost scale; L, a group of body scales viewed in SEM. Note that the side of the scale adjacent to the plasma membrane is smooth. Bar = 0.5 µm, if not stated otherwise. A
<emphasis id="B943040BFFA0291F3D98C5EFFD27308F" bold="true" box="[643,655,1968,1989]" pageId="1" pageNumber="12"></emphasis>
F and I are shown at the same magnification.
</paragraph>
</caption>
<paragraph id="8B88D819FFA2291D3FDDC0D1FD8435E2" blockId="3.[151,1436,152,758]" box="[198,556,654,680]" pageId="3" pageNumber="14">
<emphasis id="B943040BFFA2291D3FDDC0D1FEE535E2" bold="true" box="[198,333,654,680]" pageId="3" pageNumber="14">Habitat:—</emphasis>
Small, humic pool.
</paragraph>
</subSubSection>
<subSubSection id="C32D8B92FFA2291D3FDDC0EAFBE73585" box="[198,1103,693,719]" pageId="3" pageNumber="14" type="distribution">
<paragraph id="8B88D819FFA2291D3FDDC0EAFBE73585" blockId="3.[151,1436,152,758]" box="[198,1103,693,719]" pageId="3" pageNumber="14">
<emphasis id="B943040BFFA2291D3FDDC0EAFE2E3585" bold="true" box="[198,390,693,719]" pageId="3" pageNumber="14">Distribution:—</emphasis>
<collectingCountry id="F3209889FFA2291D3E9DC0EAFD883585" box="[390,544,693,719]" name="South Africa" pageId="3" pageNumber="14">South Africa</collectingCountry>
, the species was only found in the
<typeStatus id="548C66BBFFA2291D3CA3C0EAFC423585" box="[952,1002,693,719]" pageId="3" pageNumber="14">type</typeStatus>
locality.
</paragraph>
</subSubSection>
<subSubSection id="C32D8B92FFA2291D3FDDC084FA8635BC" box="[198,1326,731,758]" pageId="3" pageNumber="14" type="etymology">
<paragraph id="8B88D819FFA2291D3FDDC084FA8635BC" blockId="3.[151,1436,152,758]" box="[198,1326,731,758]" pageId="3" pageNumber="14">
<emphasis id="B943040BFFA2291D3FDDC084FEDC35BF" bold="true" box="[198,372,731,757]" pageId="3" pageNumber="14">Etymology:—</emphasis>
The epithet refers to the strong transversal rib, from Latin “temo” (pole or beam).
</paragraph>
</subSubSection>
</treatment>
</document>