376 lines
47 KiB
XML
376 lines
47 KiB
XML
<document id="34F7A91FE30C7E7A96C3AAB401D69896" ID-DOI="10.5281/zenodo.195219" ID-GBIF-Dataset="a48deddc-2ca2-4098-aad1-3ed96238ce76" ID-ISSN="1175-5326" ID-Zenodo-Dep="195219" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460224595303" checkinUser="plazi" docAuthor="Prieto-Márquez, Albert" docDate="2010" docId="3945615AFF8EFFD7BCA7FF41FBA137C6" docLanguage="en" docName="zt02452p017.pdf" docOrigin="Zootaxa 2452" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Glishades Prieto-Márquez, 2010, gen. nov." docType="treatment" docVersion="9" lastPageNumber="7" masterDocId="C57C1922FF8CFFD1BC30FFD6FF8D325A" masterDocTitle="Glishades ericksoni, a new hadrosauroid (Dinosauria: Ornithopoda) from the Late Cretaceous of North America" masterLastPageNumber="17" masterPageNumber="1" pageNumber="3" updateTime="1698238089319" updateUser="plazi">
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<mods:titleInfo id="8C7F11B6563357B3FDD911F734FC432F">
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<mods:title id="0AEBE7691CB1D6AE1F7E57AFEC015120">Glishades ericksoni, a new hadrosauroid (Dinosauria: Ornithopoda) from the Late Cretaceous of North America</mods:title>
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</mods:titleInfo>
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<mods:name id="113ACD315BB7FEC79839F8401BFF3BE4" type="personal">
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<mods:role id="2682BDF77BA90853DA7D7D16231DEA79">
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<mods:roleTerm id="F2A8CD342D7AF8E3D30794D715AE861E">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="89101ED33664EC2BCD7073D2A97E0AD3">Prieto-Márquez, Albert</mods:namePart>
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<mods:typeOfResource id="E60FE98B022B0FF5EAEF9AE2FA125132">text</mods:typeOfResource>
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<mods:titleInfo id="705542F10DC9447043223F809DD163AD">
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<mods:title id="1E244B51615F2FCDC729273D9C883A69">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="A538128FE993037DB2E634E6404C2D44">
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<mods:date id="6791FAB2012BB84A0893B0744E12C6AC">2010</mods:date>
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<mods:detail id="58D19F42287D828B96BEE378F9707060" type="volume">
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<mods:number id="9814E4AC45E5C3BFDCBDB89549873361">2452</mods:number>
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<mods:start id="8A22EBB2759D9C74D2B3B6FB80D2AD07">1</mods:start>
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<mods:end id="BFCD1FEA1599F2E32EA52D96C2E8F153">17</mods:end>
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<mods:classification id="DC6059E785ECFF86020A6FE844F6653C">journal article</mods:classification>
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<mods:identifier id="5F86CA56943FCFF8462F034179F926B0" type="DOI">10.5281/zenodo.195219</mods:identifier>
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<mods:identifier id="07AA8772A752A33902EB218D643594F2" type="GBIF-Dataset">a48deddc-2ca2-4098-aad1-3ed96238ce76</mods:identifier>
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<mods:identifier id="29FBA1AD46EF918464103AB43EB8CD40" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="51F8FF14AB902F273C314F9D5E68AAC2" type="Zenodo-Dep">195219</mods:identifier>
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</mods:mods>
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<treatment id="3945615AFF8EFFD7BCA7FF41FBA137C6" ID-DOI="http://doi.org/10.5281/zenodo.5671939" ID-GBIF-Taxon="119385968" ID-Zenodo-Dep="5671939" LSID="urn:lsid:plazi:treatment:3945615AFF8EFFD7BCA7FF41FBA137C6" httpUri="http://treatment.plazi.org/id/3945615AFF8EFFD7BCA7FF41FBA137C6" lastPageId="6" lastPageNumber="7" pageId="2" pageNumber="3">
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<subSubSection id="F9F683C7FF8EFFD3BCA7FF41FE0F32EB" box="[151,386,151,177]" pageId="2" pageNumber="3" type="nomenclature">
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<paragraph id="B153D04CFF8EFFD3BCA7FF41FE0F32EB" blockId="2.[151,386,151,177]" box="[151,386,151,177]" pageId="2" pageNumber="3">
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<heading id="EA1B6720FF8EFFD3BCA7FF41FE0F32EB" bold="true" box="[151,386,151,177]" fontSize="11" level="1" pageId="2" pageNumber="3" reason="1">
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<emphasis id="83980C5EFF8EFFD3BCA7FF41FE0F32EB" bold="true" box="[151,386,151,177]" pageId="2" pageNumber="3">
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<taxonomicName id="76ECABCFFF8EFFD3BCA7FF41FE8332EB" box="[151,270,151,177]" class="Reptilia" family="Hadrosauridae" genus="Glishades" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="2" pageNumber="3" phylum="Chordata" rank="genus" status="gen. nov.">
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<emphasis id="83980C5EFF8EFFD3BCA7FF41FE8332EB" bold="true" box="[151,270,151,177]" italics="true" pageId="2" pageNumber="3">Glishades</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="98ABB125FF8EFFD3BD24FF41FE0F32EB" box="[276,386,151,177]" pageId="2" pageNumber="3" rank="genus">gen. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="F9F683C7FF8EFFD3BCA7FF09FE563330" pageId="2" pageNumber="3" type="etymology">
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<paragraph id="B153D04CFF8EFFD3BCA7FF09FE563330" blockId="2.[151,1436,223,436]" pageId="2" pageNumber="3">
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<emphasis id="83980C5EFF8EFFD3BCA7FF09FE9332A3" bold="true" box="[151,286,223,249]" pageId="2" pageNumber="3">Etymology</emphasis>
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: “Glis” is the Latin for “mud” and “hades” means “unseen” in Greek; thus, the generic name may be translated as “concealed in mud”, in reference to its being found in sedimentary strata. Also, “Hades” was the dark lord of the underworld in Greek mythology, here metaphorically referring to the “world” beneath the surface where fossils occur.
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</paragraph>
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</subSubSection>
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<subSubSection id="F9F683C7FF8EFFD3BCF6FEA3FCA333D5" box="[198,814,373,399]" pageId="2" pageNumber="3" type="materials_examined">
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<paragraph id="B153D04CFF8EFFD3BCF6FEA3FCA333D5" blockId="2.[151,1436,223,436]" box="[198,814,373,399]" pageId="2" pageNumber="3">
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<emphasis id="83980C5EFF8EFFD3BCF6FEA3FE5933D5" bold="true" box="[198,468,373,399]" pageId="2" pageNumber="3">
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<typeStatus id="6E576EEEFF8EFFD3BCF6FEA3FE8E33D5" box="[198,259,373,399]" pageId="2" pageNumber="3">Type</typeStatus>
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and only species
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</emphasis>
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:
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<taxonomicName id="76ECABCFFF8EFFD3BDD4FEA3FD4133D4" box="[484,716,373,398]" class="Reptilia" family="Hadrosauridae" genus="Glishades" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="2" pageNumber="3" phylum="Chordata" rank="species" species="ericksoni" status="sp. nov.">
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<emphasis id="83980C5EFF8EFFD3BDD4FEA3FD4133D4" box="[484,716,373,398]" italics="true" pageId="2" pageNumber="3">Glishades ericksoni</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="98ABB125FF8EFFD3BEE4FEA3FCA333D5" box="[724,814,373,399]" pageId="2" pageNumber="3" rank="species">sp. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection id="F9F683C7FF8EFFD3BCF6FE4CFF593010" pageId="2" pageNumber="3" type="diagnosis">
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<paragraph id="B153D04CFF8EFFD3BCF6FE4CFD4E33EE" blockId="2.[151,1436,223,436]" box="[198,707,410,436]" pageId="2" pageNumber="3">
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<emphasis id="83980C5EFF8EFFD3BCF6FE4CFEB233EE" bold="true" box="[198,319,410,436]" pageId="2" pageNumber="3">Diagnosis</emphasis>
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:
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<collectingCountry id="C9FB90DCFF8EFFD3BD7EFE4CFEE233EE" box="[334,367,410,436]" name="American Samoa" pageId="2" pageNumber="3">As</collectingCountry>
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for the
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<typeStatus id="6E576EEEFF8EFFD3BDFBFE4CFE7333EE" box="[459,510,410,436]" pageId="2" pageNumber="3">type</typeStatus>
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species (below).
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</paragraph>
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<paragraph id="B153D04CFF8EFFD3BCA7FDDCFF593010" blockId="2.[151,577,522,586]" pageId="2" pageNumber="3">
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<emphasis id="83980C5EFF8EFFD3BCA7FDDCFDB7307E" bold="true" box="[151,570,522,548]" pageId="2" pageNumber="3">
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<taxonomicName id="76ECABCFFF8EFFD3BCA7FDDCFE09307E" box="[151,388,522,548]" class="Reptilia" family="Hadrosauridae" genus="Glishades" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="2" pageNumber="3" phylum="Chordata" rank="species" species="ericksoni">
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<emphasis id="83980C5EFF8EFFD3BCA7FDDCFE09307E" bold="true" box="[151,388,522,548]" italics="true" pageId="2" pageNumber="3">Glishades ericksoni</emphasis>
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</taxonomicName>
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gen. et sp. nov
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</emphasis>
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.
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<figureCitation id="29D7CCC9FF8EFFD3BCA7FDE6FF593010" box="[151,212,560,586]" captionStart="FIGURE 1" captionStartId="3.[151,255,1491,1515]" captionTargetBox="[178,1400,600,1431]" captionTargetId="figure@3.[151,1436,570,1467]" captionTargetPageId="3" captionText="FIGURE 1. Paired premaxillae from the holotype specimen of Glishades ericksoni (AMNH 27414) in dorsal (a), ventral (b), left lateral (c), and anterior (d) views." httpUri="https://zenodo.org/record/195220/files/figure.png" pageId="2" pageNumber="3">Fig 1</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="F9F683C7FF8EFFD3BCA7FDACFDDA30E0" pageId="2" pageNumber="3" type="etymology">
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<paragraph id="B153D04CFF8EFFD3BCA7FDACFDDA30E0" blockId="2.[151,1438,634,1967]" pageId="2" pageNumber="3">
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<emphasis id="83980C5EFF8EFFD3BCA7FDACFEA630CE" bold="true" box="[151,299,634,660]" pageId="2" pageNumber="3">Etymology.</emphasis>
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The specific name honors Dr. Gregory M. Erickson, for his important contributions to the knowledge of archosaur paleobiology.
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</paragraph>
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</subSubSection>
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<subSubSection id="F9F683C7FF8EFFD3BCF6FD13FBCE3115" pageId="2" pageNumber="3" type="materials_examined">
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<paragraph id="B153D04CFF8EFFD3BCF6FD13FC0F3085" blockId="2.[151,1438,634,1967]" box="[198,898,709,735]" pageId="2" pageNumber="3">
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<emphasis id="83980C5EFF8EFFD3BCF6FD13FECD3085" bold="true" box="[198,320,709,735]" pageId="2" pageNumber="3">
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<typeStatus id="6E576EEEFF8EFFD3BCF6FD13FEB63085" box="[198,315,709,735]" pageId="2" pageNumber="3" type="holotype">Holotype</typeStatus>
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.
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</emphasis>
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<collectionCode id="D7FD4889FF8EFFD3BD77FD13FE2C3085" LSID="urn:lsid:biocol.org:col:34925" box="[327,417,709,735]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="2" pageNumber="3">AMNH</collectionCode>
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27414, partial left and right premaxillae.
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</paragraph>
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<paragraph id="B153D04CFF8EFFD3BCF6FD3CFBCE3115" blockId="2.[151,1438,634,1967]" pageId="2" pageNumber="3">
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<emphasis id="83980C5EFF8EFFD3BCF6FD3CFD84315E" bold="true" box="[198,521,746,772]" pageId="2" pageNumber="3">
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<typeStatus id="6E576EEEFF8EFFD3BCF6FD3CFE8E315E" box="[198,259,746,772]" pageId="2" pageNumber="3">Type</typeStatus>
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locality and horizon.
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</emphasis>
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The specimen was collected by Brown, Kaiser, and Johnson ca.
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<quantity id="76147DA9FF8EFFD3B8C9FD3CFADC315E" box="[1273,1361,746,772]" metricMagnitude="1" metricUnit="m" metricValue="4.8641" pageId="2" pageNumber="3" unit="in" value="1915.0">1915 in</quantity>
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South Milk River,
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<quantity id="76147DA9FF8EFFD3BD19FCC6FE1C3170" box="[297,401,784,810]" metricMagnitude="4" metricUnit="m" metricValue="4.828032" pageId="2" pageNumber="3" unit="mi" value="30.0">30 miles</quantity>
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west of Sweetgrass, Glacier County, Montana (
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<collectingCountry id="C9FB90DCFF8EFFD3BFFDFCC6FA883170" box="[973,1285,784,810]" name="United States of America" pageId="2" pageNumber="3">United States of America</collectingCountry>
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), from strata corresponding to the Two Medicine Formation (Campanian, Upper Cretaceous).
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</paragraph>
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</subSubSection>
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<subSubSection id="F9F683C7FF8EFFD3BCF6FC8CFB0236B0" pageId="2" pageNumber="3" type="diagnosis">
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<paragraph id="B153D04CFF8EFFD3BCF6FC8CFE233650" blockId="2.[151,1438,634,1967]" pageId="2" pageNumber="3">
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<emphasis id="83980C5EFF8EFFD3BCF6FC8CFECB312E" bold="true" box="[198,326,858,884]" pageId="2" pageNumber="3">Diagnosis.</emphasis>
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Hadrosauroid dinosaur with the following unique combination of premaxillary characters: absence of recurvature (“reflection”) of oral margin; arcuate oral margin with wide and straight, obliquely oriented, and undeflected anterolateral corner; foramina on anteromedial surface above oral edge and adjacent to proximal end of narial bar; and grooved transversal thickening on ventral surface of premaxilla posterior to denticulate oral margin.
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</paragraph>
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<paragraph id="B153D04CFF8EFFD3BCF6FBC3FB0236B0" blockId="2.[151,1438,634,1967]" pageId="2" pageNumber="3">
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<emphasis id="83980C5EFF8EFFD3BCF6FBC3FE423675" bold="true" box="[198,463,1045,1071]" pageId="2" pageNumber="3">Nomenclatural note.</emphasis>
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Clade names and phylogenetic definitions follow the comprehensive recent revision of hadrosaurid interrelationships by Prieto-Márquez (in press) (
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<figureCitation id="29D7CCC9FF8EFFD3B86EFBECFB57360E" box="[1118,1242,1082,1108]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3" captionStartId-0="4.[151,255,1926,1950]" captionStartId-1="5.[151,255,1944,1968]" captionTargetBox-0="[183,1406,199,1887]" captionTargetBox-1="[193,1400,196,1919]" captionTargetId-0="figure@4.[183,1407,197,1889]" captionTargetId-1="figure@5.[192,1401,194,1921]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionText-0="FIGURE 2. Strict consensus cladogram of the 80 most parsimonious trees derived from maximum parsimony analysis of 56 hadrosauroid taxa, showing the phylogenetic position of Glishades ericksoni. Numbers above branches indicate bootstrap proportions, whereas those below indicate decay indices." captionText-1="FIGURE 3. Maximum agreement subtree derived from maximum parsimony analysis of 56 hadrosauroid taxa, showing the phylogenetic position of Glishades ericksoni. Numbers above branches indicate bootstrap proportions, whereas those below indicate decay indices." httpUri-0="https://zenodo.org/record/195221/files/figure.png" httpUri-1="https://zenodo.org/record/195222/files/figure.png" pageId="2" pageNumber="3">Figs 2–3</figureCitation>
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). Accordingly, Hadrosauroidea consists of
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<taxonomicName id="76ECABCFFF8EFFD3BDDCFBB6FD6F3623" box="[492,738,1120,1145]" class="Reptilia" family="Hadrosauridae" genus="Hadrosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="2" pageNumber="3" phylum="Chordata" rank="species" species="foulkii">
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<emphasis id="83980C5EFF8EFFD3BDDCFBB6FD6F3623" box="[492,738,1120,1145]" italics="true" pageId="2" pageNumber="3">Hadrosaurus foulkii</emphasis>
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</taxonomicName>
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and all taxa most closely related to it than to
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<taxonomicName id="76ECABCFFF8EFFD3B929FBB6FEC436C4" class="Reptilia" family="Iguanodontidae" genus="Iguanodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="2" pageNumber="3" phylum="Chordata" rank="species" species="bernissartensis">
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<emphasis id="83980C5EFF8EFFD3B929FBB6FEC436C4" italics="true" pageId="2" pageNumber="3">Iguanodon bernissartensis</emphasis>
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</taxonomicName>
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, including
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<taxonomicName id="76ECABCFFF8EFFD3BDE0FB53FDF136C5" box="[464,636,1157,1183]" class="Reptilia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="2" pageNumber="3" phylum="Chordata" rank="subFamily" subFamily="Hadrosaurinae">Hadrosaurinae</taxonomicName>
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(represented solely by
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<taxonomicName id="76ECABCFFF8EFFD3BFA1FB53FBA636C4" box="[913,1067,1157,1182]" class="Reptilia" family="Hadrosauridae" genus="Hadrosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="2" pageNumber="3" phylum="Chordata" rank="genus">
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<emphasis id="83980C5EFF8EFFD3BFA1FB53FBA636C4" box="[913,1067,1157,1182]" italics="true" pageId="2" pageNumber="3">Hadrosaurus</emphasis>
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</taxonomicName>
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) and
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<taxonomicName id="76ECABCFFF8EFFD3B840FB53FA9336C5" box="[1136,1310,1157,1183]" class="Reptilia" family="Saurolophidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="2" pageNumber="3" phylum="Chordata" rank="family">Saurolophidae</taxonomicName>
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. The latter is defined as the last common ancestor of
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<taxonomicName id="76ECABCFFF8EFFD3BE4DFB7DFCFC369E" box="[637,881,1195,1220]" class="Reptilia" family="Hadrosauridae" genus="Saurolophus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="2" pageNumber="3" phylum="Chordata" rank="species" species="osborni">
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<emphasis id="83980C5EFF8EFFD3BE4DFB7DFCFC369E" box="[637,881,1195,1220]" italics="true" pageId="2" pageNumber="3">Saurolophus osborni</emphasis>
|
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</taxonomicName>
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,
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<taxonomicName id="76ECABCFFF8EFFD3BF4EFB7DFB0F369E" box="[894,1154,1195,1220]" class="Reptilia" family="Hadrosauridae" genus="Lambeosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="2" pageNumber="3" phylum="Chordata" rank="species" species="lambei">
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<emphasis id="83980C5EFF8EFFD3BF4EFB7DFB0F369E" box="[894,1154,1195,1220]" italics="true" pageId="2" pageNumber="3">Lambeosaurus lambei</emphasis>
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</taxonomicName>
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, and all its descendants, which includes the two major hadrosaurid clades: Saurolophinae and
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<taxonomicName id="76ECABCFFF8EFFD3BFF5FB06FB0636B0" box="[965,1163,1232,1258]" class="Reptilia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="2" pageNumber="3" phylum="Chordata" rank="subFamily" subFamily="Lambeosaurinae">Lambeosaurinae</taxonomicName>
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.
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</paragraph>
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</subSubSection>
|
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<subSubSection id="F9F683C7FF8EFFD7BCF6FB23FBA137C6" lastPageId="6" lastPageNumber="7" pageId="2" pageNumber="3" type="description">
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<paragraph id="B153D04CFF8EFFD3BCF6FB23FC8E35F5" blockId="2.[151,1438,634,1967]" pageId="2" pageNumber="3">
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<emphasis id="83980C5EFF8EFFD3BCF6FB23FED33755" bold="true" box="[198,350,1269,1295]" pageId="2" pageNumber="3">Description.</emphasis>
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Only the anterior regions of the articulated left and right premaxillae are preserved. In dorsal view, the oral margins of both premaxillae display an arcuate contour. This contour is not perfectly smooth, but is slightly truncated by the oblique orientation and nearly straight dorsal profile of the anterolateral corners of the oral margin (
|
||
<figureCitation id="29D7CCC9FF8EFFD3BDC0FAB3FDB53725" box="[496,568,1381,1407]" captionStart="FIGURE 1" captionStartId="3.[151,255,1491,1515]" captionTargetBox="[178,1400,600,1431]" captionTargetId="figure@3.[151,1436,570,1467]" captionTargetPageId="3" captionText="FIGURE 1. Paired premaxillae from the holotype specimen of Glishades ericksoni (AMNH 27414) in dorsal (a), ventral (b), left lateral (c), and anterior (d) views." httpUri="https://zenodo.org/record/195220/files/figure.png" pageId="2" pageNumber="3">Fig. 1</figureCitation>
|
||
a). The anterior and lateral regions near the oral margin are ventrally deflected. In contrast, the anterolateral region is dorsoventrally compressed and slightly convex dorsally at the oral margin (
|
||
<figureCitation id="29D7CCC9FF8EFFD3BD1EFA66FEE23790" box="[302,367,1456,1482]" captionStart="FIGURE 1" captionStartId="3.[151,255,1491,1515]" captionTargetBox="[178,1400,600,1431]" captionTargetId="figure@3.[151,1436,570,1467]" captionTargetPageId="3" captionText="FIGURE 1. Paired premaxillae from the holotype specimen of Glishades ericksoni (AMNH 27414) in dorsal (a), ventral (b), left lateral (c), and anterior (d) views." httpUri="https://zenodo.org/record/195220/files/figure.png" pageId="2" pageNumber="3">Fig. 1</figureCitation>
|
||
c–d). There are transversal fractures across the lateral oral margin. Each facture separates a posterior and anterior portion, so that the posterior portion is slightly offset ventrally relative to the anterior one. This displacement only affects the lateral half of the mediolateral width of the oral margin. When the posterior segment is restored to its original position by having its anterior edge meet the posterior edge of the anterior portion, the oral margin remains smooth and ventrally deflected. Adjacent and lateral to the interpremaxillary joint, each element is anteroposteriorly expanded to form the anteriormost extent of the narial bar. The anterior surface of this expanded area is triangular and faces anterodorsally and slightly laterally. It is pierced in each premaxilla by two foramina (
|
||
<figureCitation id="29D7CCC9FF8EFFD3BF75F963FC053495" box="[837,904,1717,1743]" captionStart="FIGURE 1" captionStartId="3.[151,255,1491,1515]" captionTargetBox="[178,1400,600,1431]" captionTargetId="figure@3.[151,1436,570,1467]" captionTargetPageId="3" captionText="FIGURE 1. Paired premaxillae from the holotype specimen of Glishades ericksoni (AMNH 27414) in dorsal (a), ventral (b), left lateral (c), and anterior (d) views." httpUri="https://zenodo.org/record/195220/files/figure.png" pageId="2" pageNumber="3">Fig. 1</figureCitation>
|
||
a, d). A larger oval foramen is located a short distance from the narial bar; the second, much smaller foramen is located anteroventrally to the larger foramen. In addition to AMNH 27414, similarly located foramina are only unambiguously present in small (e.g., AMNH 6501 and 6575) to larger (e.g.,
|
||
<bibRefCitation id="D57DADBDFF8EFFD3BED9F8F3FB893565" author="Godefroit" box="[745,1028,1829,1855]" pageId="2" pageNumber="13" refString="Godefroit, P., Dong, Z. - M., Bultynck, P., Li, H. & Feng, L. (1998) Sino-Belgian Cooperative Program. Cretaceous Dinosaurs and Mammals from Inner Mongolia: 1) New Bactrosaurus (Dinosauria: Hadrosauroidea) material from Iren Dabasu (Inner Mongolia, P. R. China). Bulletin de l'Institute Royal des Sciences Naturelles du Belgique, 68, 1 - 70." type="journal article" year="1998">
|
||
Godefroit
|
||
<emphasis id="83980C5EFF8EFFD3BF41F8F3FC203564" box="[881,941,1829,1854]" italics="true" pageId="2" pageNumber="3">et al</emphasis>
|
||
. 1998
|
||
</bibRefCitation>
|
||
: plate 3, fig 4A) individuals of
|
||
<taxonomicName id="76ECABCFFF8EFFD3BCA7F89DFE2A353E" box="[151,423,1867,1892]" class="Reptilia" family="Lambeosauridae" genus="Bactrosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="2" pageNumber="3" phylum="Chordata" rank="species" species="johnsoni">
|
||
<emphasis id="83980C5EFF8EFFD3BCA7F89DFE2A353E" box="[151,423,1867,1892]" italics="true" pageId="2" pageNumber="3">Bactrosaurus johnsoni</emphasis>
|
||
</taxonomicName>
|
||
, a subadult specimen of
|
||
<taxonomicName id="76ECABCFFF8EFFD3BEE5F89DFBB4353E" box="[725,1081,1866,1892]" class="Reptilia" family="Hadrosauridae" genus="Edmontosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="2" pageNumber="3" phylum="Chordata" rank="species" species="annectens">
|
||
<emphasis id="83980C5EFF8EFFD3BEE5F89DFC1C353E" box="[725,913,1867,1892]" italics="true" pageId="2" pageNumber="3">Edmontosaurus</emphasis>
|
||
cf.
|
||
<emphasis id="83980C5EFF8EFFD3BFF1F89DFBB4353E" box="[961,1081,1867,1892]" italics="true" pageId="2" pageNumber="3">annectens</emphasis>
|
||
</taxonomicName>
|
||
(CMN 8509), and a subadult
|
||
<taxonomicName id="76ECABCFFF8EFFD3BCA7F8A6FEBD35D3" box="[151,304,1904,1929]" class="Reptilia" family="Hadrosauridae" genus="Gryposaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="2" pageNumber="3" phylum="Chordata" rank="genus">
|
||
<emphasis id="83980C5EFF8EFFD3BCA7F8A6FEBD35D3" box="[151,304,1904,1929]" italics="true" pageId="2" pageNumber="3">Gryposaurus</emphasis>
|
||
</taxonomicName>
|
||
sp. (CMN 8784), along with additional foramina in
|
||
<taxonomicName id="76ECABCFFF8EFFD3BFADF8A6FA8035D3" box="[925,1293,1904,1929]" class="Reptilia" family="Hadrosauridae" genus="Brachylophosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="2" pageNumber="3" phylum="Chordata" rank="species" species="canadensis">
|
||
<emphasis id="83980C5EFF8EFFD3BFADF8A6FA8035D3" box="[925,1293,1904,1929]" italics="true" pageId="2" pageNumber="3">Brachylophosaurus canadensis</emphasis>
|
||
</taxonomicName>
|
||
(e.g., MOR 794) and
|
||
<taxonomicName id="76ECABCFFF8EFFD3BD35F843FDAE35F4" box="[261,547,1941,1966]" class="Reptilia" family="Hadrosauridae" genus="Maiasaura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="2" pageNumber="3" phylum="Chordata" rank="species" species="peeblesorum">
|
||
<emphasis id="83980C5EFF8EFFD3BD35F843FDAE35F4" box="[261,547,1941,1966]" italics="true" pageId="2" pageNumber="3">Maiasaura peeblesorum</emphasis>
|
||
</taxonomicName>
|
||
(e.g., OTM F138).
|
||
</paragraph>
|
||
<paragraph id="B153D04CFF8FFFD2BCF6FF4EFEB83058" blockId="3.[151,1437,152,514]" pageId="3" pageNumber="4">
|
||
The anteromedial border of each premaxilla forms an angle of 65 degrees with the ventral surface. Posterodorsally, this border curves gently, forming a less steep angle with the ventral surface of the premaxilla. Posteriorly and dorsally, the anteromedial region of each premaxilla is extremely compressed mediolaterally to form the anterior terminus of the narial bar and the medial wall of the circumnarial fossa. Only a very small portion of the latter is preserved in each premaxilla, marked by a slightly and medially recessed surface (
|
||
<figureCitation id="29D7CCC9FF8FFFD2BD59FE84FE203336" box="[361,429,338,364]" captionStart="FIGURE 1" captionStartId="3.[151,255,1491,1515]" captionTargetBox="[178,1400,600,1431]" captionTargetId="figure@3.[151,1436,570,1467]" captionTargetPageId="3" captionText="FIGURE 1. Paired premaxillae from the holotype specimen of Glishades ericksoni (AMNH 27414) in dorsal (a), ventral (b), left lateral (c), and anterior (d) views." httpUri="https://zenodo.org/record/195220/files/figure.png" pageId="3" pageNumber="4">Fig. 1</figureCitation>
|
||
c). Posterodorsally, that surface is bounded by a crescentic edge, which constitutes a small part of the anteroventral terminus of the narial fenestra. The dorsal surface of each premaxilla, posterior to the oral margin and lateral to the narial fenestra, is only preserved anteriorly. There is no evidence of the presence of premaxillary foramina in this surface, as in non-hadrosaurid hadrosauroids and lambeosaurine hadrosaurids.
|
||
</paragraph>
|
||
<caption id="E59380C4FF8FFFD2BCA7FA05FDD93451" httpUri="https://zenodo.org/record/195220/files/figure.png" pageId="3" pageNumber="4" targetBox="[178,1400,600,1431]" targetPageId="3">
|
||
<paragraph id="B153D04CFF8FFFD2BCA7FA05FDD93451" blockId="3.[151,1436,1491,1547]" pageId="3" pageNumber="4">
|
||
<emphasis id="83980C5EFF8FFFD2BCA7FA05FE9437B1" bold="true" box="[151,281,1491,1515]" pageId="3" pageNumber="4">FIGURE 1.</emphasis>
|
||
Paired premaxillae from the holotype specimen of
|
||
<taxonomicName id="76ECABCFFF8FFFD2BF0AFA02FB8137B1" box="[826,1036,1492,1515]" class="Reptilia" family="Hadrosauridae" genus="Glishades" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="3" pageNumber="4" phylum="Chordata" rank="species" species="ericksoni">
|
||
<emphasis id="83980C5EFF8FFFD2BF0AFA02FB8137B1" box="[826,1036,1492,1515]" italics="true" pageId="3" pageNumber="4">Glishades ericksoni</emphasis>
|
||
</taxonomicName>
|
||
(AMNH 27414) in dorsal (a), ventral (b), left lateral (c), and anterior (d) views.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="B153D04CFF8FFFD2BCF6F9E0FE2035FA" blockId="3.[151,1438,1590,1952]" pageId="3" pageNumber="4">
|
||
The ventral surface of each premaxilla is carved with deep anteroposteriorly-oriented grooves (
|
||
<figureCitation id="29D7CCC9FF8FFFD2B905F9E0FAF4340A" box="[1333,1401,1590,1616]" captionStart="FIGURE 1" captionStartId="3.[151,255,1491,1515]" captionTargetBox="[178,1400,600,1431]" captionTargetId="figure@3.[151,1436,570,1467]" captionTargetPageId="3" captionText="FIGURE 1. Paired premaxillae from the holotype specimen of Glishades ericksoni (AMNH 27414) in dorsal (a), ventral (b), left lateral (c), and anterior (d) views." httpUri="https://zenodo.org/record/195220/files/figure.png" pageId="3" pageNumber="4">Fig. 1</figureCitation>
|
||
b). The prominent ridges that lie between the grooves extend anteriorly into the oral margin of each premaxilla, forming denticle-shaped structures (
|
||
<figureCitation id="29D7CCC9FF8FFFD2BE77F957FD0634C1" box="[583,651,1665,1691]" captionStart="FIGURE 1" captionStartId="3.[151,255,1491,1515]" captionTargetBox="[178,1400,600,1431]" captionTargetId="figure@3.[151,1436,570,1467]" captionTargetPageId="3" captionText="FIGURE 1. Paired premaxillae from the holotype specimen of Glishades ericksoni (AMNH 27414) in dorsal (a), ventral (b), left lateral (c), and anterior (d) views." httpUri="https://zenodo.org/record/195220/files/figure.png" pageId="3" pageNumber="4">Fig. 1</figureCitation>
|
||
d). Posterior to this denticulate margin, there is another thick and transversal layer of bone. This thickening protrudes ventrally and contains the posterior extent of the series of grooves and ridges that cross most the ventral surface of each premaxilla (
|
||
<figureCitation id="29D7CCC9FF8FFFD2B817F91AFBE134BC" box="[1063,1132,1740,1766]" captionStart="FIGURE 1" captionStartId="3.[151,255,1491,1515]" captionTargetBox="[178,1400,600,1431]" captionTargetId="figure@3.[151,1436,570,1467]" captionTargetPageId="3" captionText="FIGURE 1. Paired premaxillae from the holotype specimen of Glishades ericksoni (AMNH 27414) in dorsal (a), ventral (b), left lateral (c), and anterior (d) views." httpUri="https://zenodo.org/record/195220/files/figure.png" pageId="3" pageNumber="4">Fig. 1</figureCitation>
|
||
b–d). A deep transversal sulcus separates the oral denticulate layer from this posterior thickening. This configuration of sulcus and “double denticulate layering” of the premaxilla is typically found in hadrosaurids; among non-hadrosaurid hadrosauroids it is also present in
|
||
<taxonomicName id="76ECABCFFF8FFFD2BE55F8EAFC1C350F" box="[613,913,1852,1877]" class="Reptilia" family="Lambeosauridae" genus="Bactrosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="3" pageNumber="4" phylum="Chordata" rank="species" species="johnsoni">
|
||
<emphasis id="83980C5EFF8FFFD2BE55F8EAFC1C350F" box="[613,913,1852,1877]" italics="true" pageId="3" pageNumber="4">Bactrosaurus johnsoni</emphasis>
|
||
</taxonomicName>
|
||
(e.g., AMNH 6501 and 6575). Outside Hadrosauroidea, a posterior thickening also occurs in the basal iguanodontoidean
|
||
<taxonomicName id="76ECABCFFF8FFFD2B857F8B7FA113520" box="[1127,1436,1889,1914]" class="Reptilia" family="Iguanodontidae" genus="Ouranosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="3" pageNumber="4" phylum="Chordata" rank="species" species="nigeriensis">
|
||
<emphasis id="83980C5EFF8FFFD2B857F8B7FA113520" box="[1127,1436,1889,1914]" italics="true" pageId="3" pageNumber="4">Ouranosaurus nigeriensis</emphasis>
|
||
</taxonomicName>
|
||
(e.g., cast of GDF 300).
|
||
</paragraph>
|
||
<caption id="E59380C4FF88FFD5BCA7F850FCEC3587" httpUri="https://zenodo.org/record/195221/files/figure.png" pageId="4" pageNumber="5" targetBox="[183,1406,199,1887]" targetPageId="4">
|
||
<paragraph id="B153D04CFF88FFD5BCA7F850FCEC3587" blockId="4.[151,1436,1926,2013]" pageId="4" pageNumber="5">
|
||
<emphasis id="83980C5EFF88FFD5BCA7F850FE9735C4" bold="true" box="[151,282,1926,1950]" pageId="4" pageNumber="5">FIGURE 2.</emphasis>
|
||
Strict consensus cladogram of the 80 most parsimonious trees derived from maximum parsimony analysis of 56 hadrosauroid taxa, showing the phylogenetic position of
|
||
<taxonomicName id="76ECABCFFF88FFD5BF75F870FB9135E7" box="[837,1052,1958,1981]" class="Reptilia" family="Hadrosauridae" genus="Glishades" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="ericksoni">
|
||
<emphasis id="83980C5EFF88FFD5BF75F870FB9135E7" box="[837,1052,1958,1981]" italics="true" pageId="4" pageNumber="5">Glishades ericksoni</emphasis>
|
||
</taxonomicName>
|
||
. Numbers above branches indicate bootstrap proportions, whereas those below indicate decay indices.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="E59380C4FF89FFD4BCA7F84EFE5C35B5" httpUri="https://zenodo.org/record/195222/files/figure.png" pageId="5" pageNumber="6" targetBox="[193,1400,196,1919]" targetPageId="5">
|
||
<paragraph id="B153D04CFF89FFD4BCA7F84EFE5C35B5" blockId="5.[151,1436,1944,2031]" pageId="5" pageNumber="6">
|
||
<emphasis id="83980C5EFF89FFD4BCA7F84EFE9435EA" bold="true" box="[151,281,1944,1968]" pageId="5" pageNumber="6">FIGURE 3.</emphasis>
|
||
Maximum agreement subtree derived from maximum parsimony analysis of 56 hadrosauroid taxa, showing the phylogenetic position of
|
||
<taxonomicName id="76ECABCFFF89FFD4BDF7F86FFD17358A" box="[455,666,1977,2000]" class="Reptilia" family="Hadrosauridae" genus="Glishades" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="ericksoni">
|
||
<emphasis id="83980C5EFF89FFD4BDF7F86FFD17358A" box="[455,666,1977,2000]" italics="true" pageId="5" pageNumber="6">Glishades ericksoni</emphasis>
|
||
</taxonomicName>
|
||
. Numbers above branches indicate bootstrap proportions, whereas those below indicate decay indices.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="B153D04CFF8AFFD7BCA7FC49FBA137C6" blockId="6.[151,1438,927,1436]" pageId="6" pageNumber="7">
|
||
<emphasis id="83980C5EFF8AFFD7BCA7FC49FE9331ED" bold="true" box="[151,286,927,951]" pageId="6" pageNumber="7">FIGURE 4.</emphasis>
|
||
Boxplot showing the known range of sizes for the premaxilla for the majority of hadrosauroid taxa for which this element is known. Premaxillary width is used as an indicator of the size of this bone. It is measured perpendicular to the proximal region of the narial bar, as indicated in the inserted photograph of the left premaxilla of
|
||
<taxonomicName id="76ECABCFFF8AFFD7BCA7FBD6FE2C364D" box="[151,417,1024,1047]" class="Reptilia" family="Hadrosauridae" genus="Maiasaura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="peeblesorum">
|
||
<emphasis id="83980C5EFF8AFFD7BCA7FBD6FE2C364D" box="[151,417,1024,1047]" italics="true" pageId="6" pageNumber="7">Maiasaura peeblesorum</emphasis>
|
||
</taxonomicName>
|
||
(OTM F138). For each taxon, the box plot displays up to five statistical summary measures: smallest value of the sample, lower quartile, median (darkest horizontal line), upper quartile, and largest value. Each box contains the middle 50% of the values; its upper edge indicates the 75th percentile (upper quartile) and the lower edge represents the 25th percentile (lower quartile). The darkest horizontal line inside each box represents the median value of the sample. The ends of the vertical lines (“whiskers”) indicate the minimum and maximum values of the sample for a given taxon. Outliers are indicated by small circles. Taxon abbreviations: Ba,
|
||
<taxonomicName id="76ECABCFFF8AFFD7B864FB73FAEA36E6" box="[1108,1383,1189,1212]" class="Reptilia" family="Lambeosauridae" genus="Bactrosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="johnsoni">
|
||
<emphasis id="83980C5EFF8AFFD7B864FB73FAEA36E6" box="[1108,1383,1189,1212]" italics="true" pageId="6" pageNumber="7">Bactrosaurus johnsoni</emphasis>
|
||
</taxonomicName>
|
||
; Br,
|
||
<taxonomicName id="76ECABCFFF8AFFD7BCA7FB13FE693686" box="[151,484,1221,1244]" class="Reptilia" family="Hadrosauridae" genus="Brachylophosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="canadensis">
|
||
<emphasis id="83980C5EFF8AFFD7BCA7FB13FE693686" box="[151,484,1221,1244]" italics="true" pageId="6" pageNumber="7">Brachylophosaurus canadensis</emphasis>
|
||
</taxonomicName>
|
||
; Co,
|
||
<taxonomicName id="76ECABCFFF8AFFD7BE10FB13FD333686" box="[544,702,1221,1244]" class="Reptilia" family="Hadrosauridae" genus="Corythosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="6" pageNumber="7" phylum="Chordata" rank="genus">
|
||
<emphasis id="83980C5EFF8AFFD7BE10FB13FD333686" box="[544,702,1221,1244]" italics="true" pageId="6" pageNumber="7">Corythosaurus</emphasis>
|
||
</taxonomicName>
|
||
sp., Coi,
|
||
<taxonomicName id="76ECABCFFF8AFFD7BF15FB13FC4F3686" box="[805,962,1221,1244]" class="Reptilia" family="Hadrosauridae" genus="Corythosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="intermedius">
|
||
<emphasis id="83980C5EFF8AFFD7BF15FB13FC4F3686" box="[805,962,1221,1244]" italics="true" pageId="6" pageNumber="7">C. intermedius</emphasis>
|
||
</taxonomicName>
|
||
; Cos,
|
||
<taxonomicName id="76ECABCFFF8AFFD7B837FB13FB023686" box="[1031,1167,1221,1244]" class="Reptilia" family="Hadrosauridae" genus="Corythosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="casuarius">
|
||
<emphasis id="83980C5EFF8AFFD7B837FB13FB023686" box="[1031,1167,1221,1244]" italics="true" pageId="6" pageNumber="7">C. casuarius</emphasis>
|
||
</taxonomicName>
|
||
; Ed,
|
||
<taxonomicName id="76ECABCFFF8AFFD7B8F8FB13FAE23686" box="[1224,1391,1221,1244]" class="Reptilia" family="Hadrosauridae" genus="Edmontosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="6" pageNumber="7" phylum="Chordata" rank="genus">
|
||
<emphasis id="83980C5EFF8AFFD7B8F8FB13FAE23686" box="[1224,1391,1221,1244]" italics="true" pageId="6" pageNumber="7">Edmontosaurus</emphasis>
|
||
</taxonomicName>
|
||
sp.; Eda,
|
||
<taxonomicName id="76ECABCFFF8AFFD7BCE0FB33FED336A6" box="[208,350,1253,1276]" class="Reptilia" family="Hadrosauridae" genus="Edmontosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="annectens">
|
||
<emphasis id="83980C5EFF8AFFD7BCE0FB33FED336A6" box="[208,350,1253,1276]" italics="true" pageId="6" pageNumber="7">E. annectens</emphasis>
|
||
</taxonomicName>
|
||
; Edr,
|
||
<taxonomicName id="76ECABCFFF8AFFD7BD94FB33FD9D36A6" box="[420,528,1253,1276]" class="Reptilia" family="Hadrosauridae" genus="Edmontosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="regalis">
|
||
<emphasis id="83980C5EFF8AFFD7BD94FB33FD9D36A6" box="[420,528,1253,1276]" italics="true" pageId="6" pageNumber="7">E. regalis</emphasis>
|
||
</taxonomicName>
|
||
; Eo,
|
||
<emphasis id="83980C5EFF8AFFD7BE7CFB33FCCC36A6" box="[588,833,1253,1276]" italics="true" pageId="6" pageNumber="7">Eolambia caroljonesa</emphasis>
|
||
; Eq,
|
||
<emphasis id="83980C5EFF8AFFD7BF4EFB33FBD936A6" box="[894,1108,1253,1276]" italics="true" pageId="6" pageNumber="7">Equijubus normani</emphasis>
|
||
; Gl,
|
||
<taxonomicName id="76ECABCFFF8AFFD7B8BDFB33FAE736A6" box="[1165,1386,1253,1276]" class="Reptilia" family="Hadrosauridae" genus="Glishades" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="ericksoni">
|
||
<emphasis id="83980C5EFF8AFFD7B8BDFB33FAE736A6" box="[1165,1386,1253,1276]" italics="true" pageId="6" pageNumber="7">Glishades ericksoni</emphasis>
|
||
</taxonomicName>
|
||
; Gr,
|
||
<taxonomicName id="76ECABCFFF8AFFD7BCA7FAD3FEA93746" box="[151,292,1285,1308]" class="Reptilia" family="Hadrosauridae" genus="Gryposaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="6" pageNumber="7" phylum="Chordata" rank="genus">
|
||
<emphasis id="83980C5EFF8AFFD7BCA7FAD3FEA93746" box="[151,292,1285,1308]" italics="true" pageId="6" pageNumber="7">Gryposaurus</emphasis>
|
||
</taxonomicName>
|
||
sp.; Grl,
|
||
<taxonomicName id="76ECABCFFF8AFFD7BDBCFAD3FD8D3746" box="[396,512,1285,1308]" class="Reptilia" family="Hadrosauridae" genus="Gryposaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="latidens">
|
||
<emphasis id="83980C5EFF8AFFD7BDBCFAD3FD8D3746" box="[396,512,1285,1308]" italics="true" pageId="6" pageNumber="7">G. latidens</emphasis>
|
||
</taxonomicName>
|
||
; Grm,
|
||
<taxonomicName id="76ECABCFFF8AFFD7BE7EFAD3FC9E3746" box="[590,787,1285,1308]" class="Reptilia" family="Hadrosauridae" genus="Gryposaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="monumentensis">
|
||
<emphasis id="83980C5EFF8AFFD7BE7EFAD3FC9E3746" box="[590,787,1285,1308]" italics="true" pageId="6" pageNumber="7">G. monumentensis</emphasis>
|
||
</taxonomicName>
|
||
; Grn,
|
||
<taxonomicName id="76ECABCFFF8AFFD7BF6BFAD3FC553746" box="[859,984,1285,1308]" class="Reptilia" family="Hadrosauridae" genus="Gryposaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="notabilis">
|
||
<emphasis id="83980C5EFF8AFFD7BF6BFAD3FC553746" box="[859,984,1285,1308]" italics="true" pageId="6" pageNumber="7">G. notabilis</emphasis>
|
||
</taxonomicName>
|
||
; Hya,
|
||
<taxonomicName id="76ECABCFFF8AFFD7B812FAD3FAB53746" box="[1058,1336,1285,1308]" class="Reptilia" family="Hadrosauridae" genus="Hypacrosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="altispinus">
|
||
<emphasis id="83980C5EFF8AFFD7B812FAD3FAB53746" box="[1058,1336,1285,1308]" italics="true" pageId="6" pageNumber="7">Hypacrosaurus altispinus</emphasis>
|
||
</taxonomicName>
|
||
; Hys,
|
||
<taxonomicName id="76ECABCFFF8AFFD7B9B2FAD3FE9D3766" class="Reptilia" family="Hadrosauridae" genus="Hypacrosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="stebingeri">
|
||
<emphasis id="83980C5EFF8AFFD7B9B2FAD3FE9D3766" italics="true" pageId="6" pageNumber="7">H. stebingeri</emphasis>
|
||
</taxonomicName>
|
||
; Jz,
|
||
<emphasis id="83980C5EFF8AFFD7BD7CFAF3FDCE3766" box="[332,579,1317,1340]" italics="true" pageId="6" pageNumber="7">Jinzhousaurus yangi</emphasis>
|
||
; Lm,
|
||
<taxonomicName id="76ECABCFFF8AFFD7BEBEFAF3FCB13766" box="[654,828,1317,1340]" class="Reptilia" family="Hadrosauridae" genus="Lambeosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="6" pageNumber="7" phylum="Chordata" rank="genus">
|
||
<emphasis id="83980C5EFF8AFFD7BEBEFAF3FCB13766" box="[654,828,1317,1340]" italics="true" pageId="6" pageNumber="7">Lambeosaurus</emphasis>
|
||
</taxonomicName>
|
||
sp.; Lmc,
|
||
<taxonomicName id="76ECABCFFF8AFFD7BFF0FAF3FBD23766" box="[960,1119,1317,1340]" class="Reptilia" family="Hadrosauridae" genus="Lambeosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="laticaudus">
|
||
<emphasis id="83980C5EFF8AFFD7BFF0FAF3FBD23766" box="[960,1119,1317,1340]" italics="true" pageId="6" pageNumber="7">L. laticaudus</emphasis>
|
||
</taxonomicName>
|
||
; Lml,
|
||
<taxonomicName id="76ECABCFFF8AFFD7B882FAF3FAA93766" box="[1202,1316,1317,1340]" class="Reptilia" family="Hadrosauridae" genus="Lambeosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="lambei">
|
||
<emphasis id="83980C5EFF8AFFD7B882FAF3FAA93766" box="[1202,1316,1317,1340]" italics="true" pageId="6" pageNumber="7">L. lambei</emphasis>
|
||
</taxonomicName>
|
||
; Lmm,
|
||
<taxonomicName id="76ECABCFFF8AFFD7B9B5FAF3FEB53706" class="Reptilia" family="Hadrosauridae" genus="Lambeosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="magnicristatus">
|
||
<emphasis id="83980C5EFF8AFFD7B9B5FAF3FEB53706" italics="true" pageId="6" pageNumber="7">L. magnicristatus</emphasis>
|
||
</taxonomicName>
|
||
; Ma,
|
||
<taxonomicName id="76ECABCFFF8AFFD7BD4AFA93FDA53706" box="[378,552,1349,1372]" class="Reptilia" family="Hadrosauridae" genus="Maiasaura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="peeblesorum">
|
||
<emphasis id="83980C5EFF8AFFD7BD4AFA93FDA53706" box="[378,552,1349,1372]" italics="true" pageId="6" pageNumber="7">M. peeblesorum</emphasis>
|
||
</taxonomicName>
|
||
; Ol,
|
||
<taxonomicName id="76ECABCFFF8AFFD7BE51FA93FCC33706" box="[609,846,1349,1372]" class="Reptilia" family="Hadrosauridae" genus="Olorotitan" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="ararhensis">
|
||
<emphasis id="83980C5EFF8AFFD7BE51FA93FCC33706" box="[609,846,1349,1372]" italics="true" pageId="6" pageNumber="7">Olorotitan ararhensis</emphasis>
|
||
</taxonomicName>
|
||
; Paw,
|
||
<taxonomicName id="76ECABCFFF8AFFD7BFA8FA93FB213706" box="[920,1196,1349,1372]" class="Reptilia" family="Hadrosauridae" genus="Parasaurolophus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="walkeri">
|
||
<emphasis id="83980C5EFF8AFFD7BFA8FA93FB213706" box="[920,1196,1349,1372]" italics="true" pageId="6" pageNumber="7">Parasaurolophus walkeri</emphasis>
|
||
</taxonomicName>
|
||
; Pb,
|
||
<taxonomicName id="76ECABCFFF8AFFD7B8D6FA93FF703726" class="Reptilia" family="Iguanodontidae" genus="Probactrosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="gobiensis">
|
||
<emphasis id="83980C5EFF8AFFD7B8D6FA93FF703726" italics="true" pageId="6" pageNumber="7">Probactrosaurus gobiensis</emphasis>
|
||
</taxonomicName>
|
||
; Pt,
|
||
<emphasis id="83980C5EFF8AFFD7BD00FAB3FE783726" box="[304,501,1381,1404]" italics="true" pageId="6" pageNumber="7">Protohadros byrdi</emphasis>
|
||
; Saa,
|
||
<taxonomicName id="76ECABCFFF8AFFD7BE0AFAB3FCD03726" box="[570,861,1381,1404]" class="Reptilia" family="Hadrosauridae" genus="Saurolophus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="angustirostris">
|
||
<emphasis id="83980C5EFF8AFFD7BE0AFAB3FCD03726" box="[570,861,1381,1404]" italics="true" pageId="6" pageNumber="7">Saurolophus angustirostris</emphasis>
|
||
</taxonomicName>
|
||
; Sao,
|
||
<taxonomicName id="76ECABCFFF8AFFD7BF92FAB3FB9D3726" box="[930,1040,1381,1404]" class="Reptilia" family="Hadrosauridae" genus="Saurolophus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="osborni">
|
||
<emphasis id="83980C5EFF8AFFD7BF92FAB3FB9D3726" box="[930,1040,1381,1404]" italics="true" pageId="6" pageNumber="7">S. osborni</emphasis>
|
||
</taxonomicName>
|
||
; Te,
|
||
<taxonomicName id="76ECABCFFF8AFFD7B878FAB3FA183726" box="[1096,1429,1381,1404]" class="Reptilia" family="Hadrosauridae" genus="Telmatosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="transsylvanicus">
|
||
<emphasis id="83980C5EFF8AFFD7B878FAB3FA183726" box="[1096,1429,1381,1404]" italics="true" pageId="6" pageNumber="7">Telmatosaurus transsylvanicus</emphasis>
|
||
</taxonomicName>
|
||
; Th,
|
||
<emphasis id="83980C5EFF8AFFD7BCF2FA53FE3F37C6" box="[194,434,1413,1436]" italics="true" pageId="6" pageNumber="7">Tethyshadros insularis</emphasis>
|
||
; Ts,
|
||
<taxonomicName id="76ECABCFFF8AFFD7BDD6FA53FD7337C6" box="[486,766,1413,1436]" class="Reptilia" family="Hadrosauridae" genus="Tsintaosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="spinorhinus">
|
||
<emphasis id="83980C5EFF8AFFD7BDD6FA53FD7337C6" box="[486,766,1413,1436]" italics="true" pageId="6" pageNumber="7">Tsintaosaurus spinorhinus</emphasis>
|
||
</taxonomicName>
|
||
; Ve,
|
||
<taxonomicName id="76ECABCFFF8AFFD7BF06FA53FBA837C6" box="[822,1061,1413,1436]" class="Reptilia" family="Hadrosauridae" genus="Velafrons" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="coahuilensis">
|
||
<emphasis id="83980C5EFF8AFFD7BF06FA53FBA837C6" box="[822,1061,1413,1436]" italics="true" pageId="6" pageNumber="7">Velafrons coahuilensis</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |