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<document ID-DOI="http://dx.doi.org/10.3897/dez.69.83840" ID-Pensoft-Pub="1860-1324-1-71" ID-Pensoft-UUID="DF59A16543B05799A4FD7B66E721FC76" ID-ZooBank="7DEB0587A3734B06BAADCA6102EC6581" ModsDocID="1860-1324-69-1-71" checkinTime="1655205803791" checkinUser="pensoft" docAuthor="Gueorguiev, Borislav, Wrase, David W., Assmann, Thorsten, Muilwijk, Jan &amp; Machard, Patrice" docDate="2022" docId="5964744C623D5BDFAC9729475336017E" docLanguage="en" docName="DeutEntomolZeit 69(1): 71-122" docOrigin="Deutsche Entomologische Zeitschrift 69 (1)" docPubDate="2022-06-13" docSource="http://dx.doi.org/10.3897/dez.69.83840" docTitle="Platyderus (Eremoderus) languidus" docType="treatment" docVersion="2" id="DF59A16543B05799A4FD7B66E721FC76" lastPageNumber="71" masterDocId="DF59A16543B05799A4FD7B66E721FC76" masterDocTitle="Taxonomic revision of the African and Southwest Asian species of Platyderus Stephens, subg. Eremoderus Jeanne (Coleoptera, Carabidae, Sphodrini)" masterLastPageNumber="122" masterPageNumber="71" pageNumber="71" updateTime="1655206106596" updateUser="ExternalLinkService">
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<mods:title>Taxonomic revision of the African and Southwest Asian species of Platyderus Stephens, subg. Eremoderus Jeanne (Coleoptera, Carabidae, Sphodrini)</mods:title>
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<mods:namePart>Gueorguiev, Borislav</mods:namePart>
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<mods:affiliation>National Museum of Natural History, Bulgarian Academy of Sciences, 1 Blvd. Tsar Osvoboditel, 1000 Sofia, Bulgaria</mods:affiliation>
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<mods:namePart>Wrase, David W.</mods:namePart>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Assmann, Thorsten</mods:namePart>
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<mods:affiliation>Leuphana University Lueneburg, Institute of Ecology, Universitaetsallee 1, 21335 Lueneburg, Germany</mods:affiliation>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Muilwijk, Jan</mods:namePart>
<mods:affiliation>Department of Entomology, Naturalis Biodiversity Centre, Darwinweg 2, 2333 CR Leiden, Netherlands</mods:affiliation>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Machard, Patrice</mods:namePart>
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<treatment LSID="urn:lsid:plazi:treatment:5964744C623D5BDFAC9729475336017E" httpUri="http://treatment.plazi.org/id/5964744C623D5BDFAC9729475336017E" lastPageNumber="71" pageId="0" pageNumber="71">
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19. 
<taxonomicName LSID="5964744C-623D-5BDF-AC97-29475336017E" authority="(Reiche &amp; Saulcy, 1855)" baseAuthorityName="Reiche &amp; Saulcy" baseAuthorityYear="1855" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) languidus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="languidus" subGenus="Eremoderus">Platyderus (Eremoderus) languidus (Reiche &amp; Saulcy, 1855)</taxonomicName>
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<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Dorsal habitus. A. Platyderus (Eremoderus) insignitus insignitus Bedel, 1902, male specimen, SW Tiznit, Sous-Massa Region, Morocco; B. P. (E.) irakensis, sp. nov., holotype; C. P. (E.) jordanensis, sp. nov., holotype; D. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype and its labels; E. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male specimen, &quot; Jerusalem Syria &quot;, Jerusalem District, Israel. Scale bars: 1 mm." figureDoi="10.3897/dez.69.83840.figure4" httpUri="https://binary.pensoft.net/fig/695641" pageId="0" pageNumber="71">Figs 4D, E</figureCitation>
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Mesofemora, anterior side (A-D: left mesofemur, E-I: right mesofemur; white arrows and dots indicate position of setiferous punctures). A. Platyderus (Eremoderus) weiratheri Maran, 1940, female specimen, Boz dag koey env., Izmir District, Turkey; B. P. (E.) felixi, sp. nov., topotype female paratype; C. P. (E.) iranicus, sp. nov., topotype female paratype; D. P. (E.) lassallei, sp. nov., female paratype, Iran, Mazandaran Province, E Qolqol; E. P. (E.) klapperichi, sp. nov., holotype; F. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; G. P. (E.) afghanistanicus, sp. nov., holotype; H. P. (E.) brunneus ferrantei Reitter, 1909, lectotype; I. P. (E.) languidus (Reiche &amp; Saulcy, 1855), female specimen, Netanya 1.3.97, Central District, Israel. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure5" httpUri="https://binary.pensoft.net/fig/695642" pageId="0" pageNumber="71">, 5I</figureCitation>
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Metatarsus, dorsal view. A. Platyderus (Eremoderus) brunneus brunneus Karsch, 1881, left metatarsus, female specimen, Aziziyah, Jafara District, Libya; B. P. (E.) brunneus ferrantei Reitter, 1909, left metatarsus, female specimen, Holot Haluza, Southern District, Israel; C. P. (E.) irakensis, sp. nov., left metatarsus (black arrows indicate longitudinal grooves on metatarsomeres 2, 3 and 4), holotype; D. P. (E.) jordanensis, sp. nov., right metatarsus, holotype; E. P. (E.) languidus (Reiche &amp; Saulcy, 1855), right metatarsus, male specimen, Na ẖal Prat, Judea and Samaria Area, Israel. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure6" httpUri="https://binary.pensoft.net/fig/695643" pageId="0" pageNumber="71">, 6E</figureCitation>
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Urite, ventral view. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, E: topotype male specimen; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) vrabeci, sp. nov., holotype; E. P. (E.) lassallei, sp. nov., holotype; F. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; G. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; H. P. (E.) afghanistanicus, sp. nov., holotype; I. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; J. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; K. P. (E.) jordanensis, sp. nov., holotype; L. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; M. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male specimen, Na ẖal Prat, Judea and Samaria Area, Israel; N. P. (Platyderus) sp., male specimen, Boz Daglar Mtn., Turkey; O. P. (P.) reticulatus (Chaudoir), male specimen, Tashehzeh, Nowshahr County, Mazandaran Province, Iran. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure7" httpUri="https://binary.pensoft.net/fig/695644" pageId="0" pageNumber="71">, 7L, M</figureCitation>
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Median lobe of aedeagus, left lateral view (black arrows on D, F-H indicate convexity of ventral margin of apex of median lobe). A. Platyderus (Platyderus) sp., male specimen, Boz Daglar Mtn., Turkey; B. P. (Eremoderus) afghanistanicus, sp. nov., holotype; C. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; D. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; E. P. (E.) insignitus insignitus Bedel, 1902, male specimen, SW Tiznit, Sous-Massa Region, Morocco; F. P. (E.) jordanensis, sp. nov., holotype; G. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; H. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male specimen, Na ẖal Prat, Judea and Samaria Area, Israel. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure9" httpUri="https://binary.pensoft.net/fig/695646" pageId="0" pageNumber="71">, 9G, H</figureCitation>
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Median lobe of aedeagus, ventral view. A. Platyderus (Eremoderus) afghanistanicus, sp. nov., holotype; B. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; C. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; D. P. (E.) insignitus insignitus Bedel, 1902, male specimen, SW Tiznit, Sous-Massa Region, Morocco; E. P. (E.) jordanensis, sp. nov., holotype; F. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; G. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male specimen, Na ẖal Prat, Judea and Samaria Area, Israel. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure11" httpUri="https://binary.pensoft.net/fig/695648" pageId="0" pageNumber="71">, 11F, G</figureCitation>
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Right paramere, external face. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., holotype; E. P. (E.) vrabeci, sp. nov., holotype; F. P. (E.) lassallei, sp. nov., holotype; G. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; H. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; I. P. (E.) afghanistanicus, sp. nov., holotype; J. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; K. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; L. P. (E.) jordanensis, sp. nov., holotype; M. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; N. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male paralectotype, &quot; Syria &quot;. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure12" httpUri="https://binary.pensoft.net/fig/695649" pageId="0" pageNumber="71">, 12M, N</figureCitation>
<figureCitation captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Spermathecal complex and gonocoxites, ventral view. A. Platyderus (Eremoderus) brunneus brunneus Karsch, 1881, female specimen, Bukamash, Nuqat al Khams District, Libya; B. P. (E.) brunneus ferrantei Reitter, 1909, female specimen, Holot Haluza, Southern District, Israel; C. P. (E.) insignitus insignitus Bedel, 1902, female specimen, Plage Aglou, Sous-Massa Region, Morocco; D. P. (E.) irakensis, sp. nov., holotype; E. P. (E.) jordanensis, sp. nov., female paratype, N Wadi Musa, Ma'an Governorate, Jordan; F. P. (E.) languidus (Reiche &amp; Saulcy, 1855), female specimens, Israel, Netanya 22. xii. 1996, Central District, Israel. Scale bars: 0.5 mm. For abbreviations see captions on Fig. 13." figureDoi="10.3897/dez.69.83840.figure15" httpUri="https://binary.pensoft.net/fig/695652" pageId="0" pageNumber="71">, 15F</figureCitation>
<figureCitation captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Distribution of species from subgenus Platyderus Eremoderus in the Levant and the Arabian Peninsula: Platyderus arabicus sp. nov. (yellow question marks), P. brunki sp. nov. (blue circle), P. irakensis sp. nov. (green circles), P. jordanensis sp. nov. (brown circles), and P. languidus (red circles)." figureDoi="10.3897/dez.69.83840.figure19" httpUri="https://binary.pensoft.net/fig/695656" pageId="0" pageNumber="71">, 19</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="reference_group">
<paragraph pageId="0" pageNumber="71">
<taxonomicName authorityName="Reiche &amp; Saulcy" authorityYear="1855" class="Insecta" family="Carabidae" genus="Feronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Feronia (Argutor) languida" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="languida" subGenus="Argutor">Feronia (Argutor) languida</taxonomicName>
Reiche &amp; Saulcy, 1855: 610 (type locality: 
<normalizedToken originalValue="“Jordanis”">&quot;Jordanis&quot;</normalizedToken>
, based on lectotype designation).
</paragraph>
<paragraph pageId="0" pageNumber="71">
<taxonomicName authorityName="Fairmaire" authorityYear="1872" class="Insecta" family="Carabidae" genus="Sphodrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sphodrus parumstriatus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="parumstriatus">Sphodrus parumstriatus</taxonomicName>
= 
<taxonomicName authorityName="Fairmaire" authorityYear="1872" class="Insecta" family="Carabidae" genus="Sphodrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sphodrus parumstriatus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="parumstriatus">Sphodrus parumstriatus</taxonomicName>
Fairmaire, 1872: 47 (type locality: 
<normalizedToken originalValue="“inconnue…">&quot;inconnue...</normalizedToken>
probably 
<normalizedToken originalValue="qu’il">qu'il</normalizedToken>
appartient 
<normalizedToken originalValue="à">a</normalizedToken>
la faune 
<normalizedToken originalValue="méditerranéenne”">mediterraneenne&quot;</normalizedToken>
). Synonymy established by 
<bibRefCitation DOI="https://doi.org/10.3406/bsef.1906.23895" author="Bedel, L" journalOrPublisher="Bulletin de la Societe Entomologique de France" pageId="0" pageNumber="71" pagination="91 - 93" refId="B12" refString="Bedel, L, 1906. Communications. Synonymies de Coleopteres palearctiques. Bulletin de la Societe Entomologique de France 1906 (8): 91 - 93, DOI: https://doi.org/10.3406/bsef.1906.23895" title="Communications. Synonymies de Coleopteres palearctiques." url="https://doi.org/10.3406/bsef.1906.23895" volume="1906" year="1906">Bedel (1906</bibRefCitation>
: 91).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="note on type locality">
<paragraph pageId="0" pageNumber="71">Note on type locality.</paragraph>
<paragraph pageId="0" pageNumber="71">
The specimens were collected from: &quot;Des bords du Jourdain et de la mer Morte&quot; (
<bibRefCitation author="Reiche, L" journalOrPublisher="Annales de la Societe Entomologique de France" pageId="0" pageNumber="71" pagination="561 - 645" refId="B44" refString="Reiche, L, Saulcy, F, 1855. Especes nouvelles ou peu connues de coleopteres, recueillies par M. F. de Saulcy, membre de l'Institut, dans son voyage en Orient, et decrites par M.M. L. Reiche et Felicien de Saulcy. Annales de la Societe Entomologique de France 3: 561 - 645" title="Especes nouvelles ou peu connues de coleopteres, recueillies par M. F. de Saulcy, membre de l'Institut, dans son voyage en Orient, et decrites par M. M. L. Reiche et Felicien de Saulcy." volume="3" year="1855">Reiche and Saulcy 1855</bibRefCitation>
: 610).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="references">
<paragraph pageId="0" pageNumber="71">References.</paragraph>
<paragraph pageId="0" pageNumber="71">
<taxonomicName baseAuthorityName="Reiche &amp; Saulcy" baseAuthorityYear="1855" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus languidus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="languidus">
<emphasis italics="true" pageId="0" pageNumber="71">Platyderus languidus</emphasis>
</taxonomicName>
: Chaudoir 1866: 108-109; 
<bibRefCitation author="Piochard de la Brulerie, CJ" journalOrPublisher="Entomologische Blaetter" pageId="0" pageNumber="71" refId="B43" refString="Piochard de la Brulerie, CJ, 1876. Catalogue raisonne des coleopteres de la Syrie et de l'ile de Chypre. 1re partie (suite). Annales de la Societe Entomologique de France [5] 5(1875): 395-448." title="Catalogue raisonne des coleopteres de la Syrie et de l'ile de Chypre. 1 re partie (suite). Annales de la Societe Entomologique de France [5] 5 (1875): 395 - 448." year="1876">
Piochard de la 
<normalizedToken originalValue="Brûlerie">Brulerie</normalizedToken>
1876
</bibRefCitation>
: 428, 430; 
<bibRefCitation author="Bedel, L" journalOrPublisher="Zootaxa" pageId="0" pageNumber="71" refId="B11" refString="Bedel, L, 1902. Catalogue raisonne des Coleopteres du nord de l'Afrique (Maroc, Algerie, Tunisie et Tripolitaine) avec notes sur la faune des iles Canaries et de Madere, premiere partie. Societe Entomologique de France, Paris, 209-220." title="Catalogue raisonne des Coleopteres du nord de l'Afrique (Maroc, Algerie, Tunisie et Tripolitaine) avec notes sur la faune des iles Canaries et de Madere, premiere partie. Societe Entomologique de France, Paris, 209 - 220." year="1902">Bedel 1902</bibRefCitation>
: 211; 
<bibRefCitation author="Sahlberg, JR" journalOrPublisher="Baltic Journal of Coleopterology" pageId="0" pageNumber="71" refId="B48" refString="Sahlberg, JR, 1903. Coleoptera Levantina mensibus Februario et Martio 1896 in Palaestina et Aegypto inferiore collecta. Oefversigt af Finska Vetenskaps-Societetens Foerhandlingar 45[1902-1903](18): 1-36." title="Coleoptera Levantina mensibus Februario et Martio 1896 in Palaestina et Aegypto inferiore collecta. Oefversigt af Finska Vetenskaps-Societetens Foerhandlingar 45 [1902 - 1903] (18): 1 - 36." year="1903">Sahlberg 1903</bibRefCitation>
: 3-4; 
<bibRefCitation DOI="https://doi.org/10.3406/bsef.1906.23895" author="Bedel, L" journalOrPublisher="Bulletin de la Societe Entomologique de France" pageId="0" pageNumber="71" pagination="91 - 93" refId="B12" refString="Bedel, L, 1906. Communications. Synonymies de Coleopteres palearctiques. Bulletin de la Societe Entomologique de France 1906 (8): 91 - 93, DOI: https://doi.org/10.3406/bsef.1906.23895" title="Communications. Synonymies de Coleopteres palearctiques." url="https://doi.org/10.3406/bsef.1906.23895" volume="1906" year="1906">Bedel 1906</bibRefCitation>
: 91; 
<bibRefCitation author="Csiki, E" editor="Junk, W" journalOrPublisher="Volumen II. Carabidae II. W. Junk, Berlin" pageId="0" pageNumber="71" pagination="739 - 1022" refId="B16" refString="Csiki, E, 1931. Carabidae: Harpalinae V (Pars 115). In: Junk, W, Schenkling, S, Eds., Coleopterorum catalogus. Volumen II. Carabidae II. W. Junk, Berlin: 739 - 1022" title="Carabidae: Harpalinae V (Pars 115)." volumeTitle="Coleopterorum catalogus." year="1931">Csiki 1931</bibRefCitation>
: 769; 
<bibRefCitation author="Jedlicka, A" journalOrPublisher="Bulletin de la Societe Entomologique de France" pageId="0" pageNumber="71" refId="B26" refString="Jedlicka, A, 1963. Neue Carabiden aus Anatolien und vom Balkan. Koleopterologische Rundschau 40-41(1962-1963): 16-22." title="Neue Carabiden aus Anatolien und vom Balkan. Koleopterologische Rundschau 40 - 41 (1962 - 1963): 16 - 22." year="1963">
<normalizedToken originalValue="Jedlička">Jedlicka</normalizedToken>
1963
</bibRefCitation>
: 20, 22; Israelso 1990: 165; 
<bibRefCitation DOI="https://doi.org/10.3406/bsef.1996.17271" author="Jeanne, C" journalOrPublisher="Bulletin de la Societe Entomologique de France" pageId="0" pageNumber="71" pagination="397 - 412" refId="B25" refString="Jeanne, C, 1996. Le genre Platyderus Stephens: I. - Especes nouvelles de la peninsule Iberique (Coleoptera, Pterostichidae). Bulletin de la Societe Entomologique de France 101 (4): 397 - 412, DOI: https://doi.org/10.3406/bsef.1996.17271" title="Le genre Platyderus Stephens: I. - Especes nouvelles de la peninsule Iberique (Coleoptera, Pterostichidae)." url="https://doi.org/10.3406/bsef.1996.17271" volume="101" year="1996">Jeanne 1996</bibRefCitation>
: 398; 
<bibRefCitation author="Lorenz, W" journalOrPublisher="Acta Societatis Zoologicae Bohemicae" pageId="0" pageNumber="71" refId="B33" refString="Lorenz, W, 1998. Nomina Carabidarum - a directory of the scientific names of ground beetles (Insecta, Coleoptera &quot;Geadephaga&quot;: Trachypachidae and Carabidae incl. Paussinae, Cicindelinae, Rhysodidae). First edition. Wolfgang Lorenz, Tutzing, [ii +] 502 pp." title="Nomina Carabidarum - a directory of the scientific names of ground beetles (Insecta, Coleoptera &quot; Geadephaga &quot;: Trachypachidae and Carabidae incl. Paussinae, Cicindelinae, Rhysodidae). First edition. Wolfgang Lorenz, Tutzing, [ii +] 502 pp." year="1998">Lorenz 1998</bibRefCitation>
: 375; 
<bibRefCitation author="Hovorka, O" editor="Loebl, L" journalOrPublisher="Volume 1. Archostemata - Myxophaga - Adephaga. Apollo Books, Stenstrup" pageId="0" pageNumber="71" pagination="521 - 524" refId="B22" refString="Hovorka, O, Sciaky, R, 2003. Atranopsina Baehr, 1982. In: Loebl, L, Smetana, A, Eds., Catalogue of Palaearctic Coleoptera. Volume 1. Archostemata - Myxophaga - Adephaga. Apollo Books, Stenstrup: 521 - 524" title="Atranopsina Baehr, 1982." volumeTitle="Catalogue of Palaearctic Coleoptera." year="2003">Hovorka and Sciaky 2003</bibRefCitation>
: 521; 
<bibRefCitation author="Lorenz, W" journalOrPublisher="Acta Societatis Zoologicae Bohemicae" pageId="0" pageNumber="71" refId="B34" refString="Lorenz, W, 2005. Systematic list of extant ground beetles of the world (InsectaColeoptera &quot;Geadephaga&quot;: Trachypachidae and Carabidae incl. Paussinae, Cicindelinae, Rhysodinae). Second Edition. Wolfgang Lorenz, Tutzing, [ii +] 530 pp." title="Systematic list of extant ground beetles of the world (InsectaColeoptera &quot; Geadephaga &quot;: Trachypachidae and Carabidae incl. Paussinae, Cicindelinae, Rhysodinae). Second Edition. Wolfgang Lorenz, Tutzing, [ii +] 530 pp." year="2005">Lorenz 2005</bibRefCitation>
: 396; 
<bibRefCitation author="Hovorka, O" editor="Loebl, I" journalOrPublisher="Volume 1 Revised and Updated Edition. Archostemata - Myxophaga - Adephaga. Brill, Leiden - Boston" pageId="0" pageNumber="71" pagination="755 - 760" refId="B21" refString="Hovorka, O, 2017. Atranopsina Baehr, 1982. In: Loebl, I, Loebl, D, Eds., Catalogue of Palaearctic Coleoptera. Volume 1 Revised and Updated Edition. Archostemata - Myxophaga - Adephaga. Brill, Leiden - Boston: 755 - 760" title="Atranopsina Baehr, 1982." volumeTitle="Catalogue of Palaearctic Coleoptera." year="2017">Hovorka 2017</bibRefCitation>
: 756.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="type material">
<paragraph pageId="0" pageNumber="71">Type material.</paragraph>
<paragraph pageId="0" pageNumber="71">
<taxonomicName authorityName="Reiche &amp; Saulcy" authorityYear="1855" class="Insecta" family="Carabidae" genus="Feronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Feronia languida" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="languida">
<emphasis italics="true" pageId="0" pageNumber="71">Feronia languida</emphasis>
</taxonomicName>
Reiche &amp; Saulcy, 1855. Consists of four syntypes, 2♂♂, 2♀♀ in MHNG, all conspecific. All the specimens quite dirty and damaged. Three of them were preserved in pinned condition, whereas a male below designated as paralectotype, together with the extracted genitalia (without the urite preserved), was glued on a card; originally this specimen was pinned. The last specimen has been labelled as holotype by the late Claude Jeanne, but such a treatment was incorrect because at least four type specimens exist belonging to one species, all of them bearing the well-known yellow handwritten labels of Louis 
<normalizedToken originalValue="Jérome">Jerome</normalizedToken>
Reiche (1799-1890); all specimens have the status of syntypes.
</paragraph>
<caption doi="10.3897/dez.69.83840.figure17" httpUri="https://binary.pensoft.net/fig/695654" pageId="0" pageNumber="71" start="Figure 17" startId="F17">
<paragraph pageId="0" pageNumber="71">
<emphasis bold="true" pageId="0" pageNumber="71">Figure 17.</emphasis>
Distribution of species from subgenus 
<taxonomicName authorityName="Jeanne" authorityYear="1996" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus)" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="subGenus" subGenus="Eremoderus">Platyderus Eremoderus</taxonomicName>
in Iran and Afghanistan: 
<taxonomicName authorityName="Guéorguiev &amp; Wrase &amp; Assmann &amp; Muilwijk &amp; Machard" authorityYear="2022" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus felixi" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="felixi">
<emphasis italics="true" pageId="0" pageNumber="71">Platyderus felixi</emphasis>
</taxonomicName>
sp. nov. (red circle), 
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. iranicus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="iranicus">
<emphasis italics="true" pageId="0" pageNumber="71">P. iranicus</emphasis>
</taxonomicName>
sp. nov. (blue circles), 
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. lassallei" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="lassallei">
<emphasis italics="true" pageId="0" pageNumber="71">P. lassallei</emphasis>
</taxonomicName>
sp. nov. (yellow circles), 
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. davatchii" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="davatchii">
<emphasis italics="true" pageId="0" pageNumber="71">P. davatchii</emphasis>
</taxonomicName>
(green triangle), 
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. ledouxi" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="ledouxi">
<emphasis italics="true" pageId="0" pageNumber="71">P. ledouxi</emphasis>
</taxonomicName>
(red triangle), 
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. taghizadehi" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="taghizadehi">
<emphasis italics="true" pageId="0" pageNumber="71">P. taghizadehi</emphasis>
</taxonomicName>
(blue triangle), 
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. klapperichi" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="klapperichi">
<emphasis italics="true" pageId="0" pageNumber="71">P. klapperichi</emphasis>
</taxonomicName>
sp. nov. (black triangle), and 
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. afghanistanicus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="afghanistanicus">
<emphasis italics="true" pageId="0" pageNumber="71">P. afghanistanicus</emphasis>
</taxonomicName>
sp. nov. (red question mark).
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="71">
We chose for lectotype designation a male specimen with the label 
<normalizedToken originalValue="“Jordanis”">&quot;Jordanis&quot;</normalizedToken>
in order to retain the type locality 
<normalizedToken originalValue="“Jordanis”">&quot;Jordanis&quot;</normalizedToken>
that is in accordance with the locality that was given in the description (
<bibRefCitation author="Reiche, L" journalOrPublisher="Annales de la Societe Entomologique de France" pageId="0" pageNumber="71" pagination="561 - 645" refId="B44" refString="Reiche, L, Saulcy, F, 1855. Especes nouvelles ou peu connues de coleopteres, recueillies par M. F. de Saulcy, membre de l'Institut, dans son voyage en Orient, et decrites par M.M. L. Reiche et Felicien de Saulcy. Annales de la Societe Entomologique de France 3: 561 - 645" title="Especes nouvelles ou peu connues de coleopteres, recueillies par M. F. de Saulcy, membre de l'Institut, dans son voyage en Orient, et decrites par M. M. L. Reiche et Felicien de Saulcy." volume="3" year="1855">Reiche and Saulcy 1855</bibRefCitation>
). Although rather damaged, we took a picture of its habitus and examined its genitalia (Fig. 
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Dorsal habitus. A. Platyderus (Eremoderus) insignitus insignitus Bedel, 1902, male specimen, SW Tiznit, Sous-Massa Region, Morocco; B. P. (E.) irakensis, sp. nov., holotype; C. P. (E.) jordanensis, sp. nov., holotype; D. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype and its labels; E. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male specimen, &quot; Jerusalem Syria &quot;, Jerusalem District, Israel. Scale bars: 1 mm." figureDoi="10.3897/dez.69.83840.figure4" httpUri="https://binary.pensoft.net/fig/695641" pageId="0" pageNumber="71">4D</figureCitation>
). Afterwards, the lectotype specimen was glued onto a white card, and its genitalia were embedded in Euparal on a second smaller white card pinned below the specimen.
</paragraph>
<paragraph pageId="0" pageNumber="71">
The four specimens are labelled, as follows: lectotype ♂, 'Jordanis [h, y] // Coll. Reiche [p, w] // PARATYPE 
<taxonomicName class="Insecta" family="Carabidae" genus="Feronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Feronia" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="genus">Feronia</taxonomicName>
/ 
<taxonomicName class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus languida" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="languida">Platyderus languida</taxonomicName>
R. &amp; S / C. JEANNE des. 1990 [p, h, r] // 
<taxonomicName authorityName="Stephens" authorityYear="1827" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="genus">Platyderus</taxonomicName>
/ 
<taxonomicName baseAuthorityName="Reiche &amp; Saulcy" baseAuthorityYear="1855" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) languidus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="languidus" subGenus="Eremoderus">Platyderus Eremoderus languidus</taxonomicName>
R. &amp; S. / JEANNE det. 1990 [p, w]'; paralectotype ♂, ' 
<taxonomicName authorityName="Stephens" authorityYear="1827" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="genus">Platyderus</taxonomicName>
/ 
<taxonomicName baseAuthorityName="Reiche &amp; Saulcy" baseAuthorityYear="1855" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) languidus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="languidus" subGenus="Eremoderus">Platyderus Eremoderus languidus</taxonomicName>
. Reiche / Soc. Ent. 1855. 610 / Syria [h, y] // Coll. Reiche [p, w] // HOLOTYPE 
<taxonomicName class="Insecta" family="Carabidae" genus="Feronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Feronia" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="genus">Feronia</taxonomicName>
/ 
<taxonomicName class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus languida" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="languida">Platyderus languida</taxonomicName>
R. &amp; S. / C. JEANNE des. 1990 [p, h, r] // 
<taxonomicName authorityName="Stephens" authorityYear="1827" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="genus">Platyderus</taxonomicName>
/ 
<taxonomicName baseAuthorityName="Reiche &amp; Saulcy" baseAuthorityYear="1855" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) languidus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="languidus" subGenus="Eremoderus">Platyderus Eremoderus languidus</taxonomicName>
R. &amp; S. / JEANNE det. 1990 [p, w]'; paralectotype ♀, 'Nazareth [h, y] // Coll. Reiche [p, w] // PARATYPE 
<taxonomicName class="Insecta" family="Carabidae" genus="Feronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Feronia" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="genus">Feronia</taxonomicName>
/ 
<taxonomicName class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus languida" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="languida">Platyderus languida</taxonomicName>
R. &amp; S / C. JEANNE des. 1990 [p, h, r] // 
<taxonomicName authorityName="Stephens" authorityYear="1827" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="genus">Platyderus</taxonomicName>
/ 
<taxonomicName baseAuthorityName="Reiche &amp; Saulcy" baseAuthorityYear="1855" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) languidus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="languidus" subGenus="Eremoderus">Platyderus Eremoderus languidus</taxonomicName>
R. &amp; S. JEANNE det. 1990 [p, w]'; paralectotype ♀, 
<normalizedToken originalValue="‘Palæstina">'Palaestina</normalizedToken>
[h, y] // Coll. Reiche [p, w] // // PARATYPE 
<taxonomicName class="Insecta" family="Carabidae" genus="Feronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Feronia" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="genus">Feronia</taxonomicName>
/ 
<taxonomicName class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus languida" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="languida">Platyderus languida</taxonomicName>
R. &amp; S / C. JEANNE des. 1990 [p, h, r] // 
<taxonomicName authorityName="Stephens" authorityYear="1827" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="genus">Platyderus</taxonomicName>
/ 
<taxonomicName baseAuthorityName="Reiche &amp; Saulcy" baseAuthorityYear="1855" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) languidus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="languidus" subGenus="Eremoderus">Platyderus Eremoderus languidus</taxonomicName>
R. &amp; S. JEANNE det. 1990 [p, w]'.
</paragraph>
<caption doi="10.3897/dez.69.83840.figure18" httpUri="https://binary.pensoft.net/fig/695655" pageId="0" pageNumber="71" start="Figure 18" startId="F18">
<paragraph pageId="0" pageNumber="71">
<emphasis bold="true" pageId="0" pageNumber="71">Figure 18.</emphasis>
Distribution of species from subgenus 
<taxonomicName authorityName="Jeanne" authorityYear="1996" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus)" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="subGenus" subGenus="Eremoderus">Platyderus Eremoderus</taxonomicName>
in continental North Africa and the southern Levant: 
<taxonomicName authority="brunneus" authorityName="brunneus Karsch" authorityYear="1881" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus brunneus subsp. brunneus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="subSpecies" species="brunneus" subSpecies="brunneus">
<emphasis italics="true" pageId="0" pageNumber="71">Platyderus brunneus brunneus</emphasis>
</taxonomicName>
(blue circles), 
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. brunneus subsp. ferrantei" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="subSpecies" species="brunneus" subSpecies="ferrantei">
<emphasis italics="true" pageId="0" pageNumber="71">P. brunneus ferrantei</emphasis>
</taxonomicName>
(yellow circles), 
<taxonomicName authority="insignitus" class="Insecta" genus="P." kingdom="Animalia" lsidName="P. insignitus subsp. insignitus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="subSpecies" species="insignitus" subSpecies="insignitus">
<emphasis italics="true" pageId="0" pageNumber="71">P. insignitus insignitus</emphasis>
</taxonomicName>
(red circles), and 
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. insignitus subsp. presaharensis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="subSpecies" species="insignitus" subSpecies="presaharensis">
<emphasis italics="true" pageId="0" pageNumber="71">P. insignitus presaharensis</emphasis>
</taxonomicName>
(brown circle).
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="71">
Besides, all the four specimens have two supplementary labels subsequently added (Fig. 
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Dorsal habitus. A. Platyderus (Eremoderus) insignitus insignitus Bedel, 1902, male specimen, SW Tiznit, Sous-Massa Region, Morocco; B. P. (E.) irakensis, sp. nov., holotype; C. P. (E.) jordanensis, sp. nov., holotype; D. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype and its labels; E. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male specimen, &quot; Jerusalem Syria &quot;, Jerusalem District, Israel. Scale bars: 1 mm." figureDoi="10.3897/dez.69.83840.figure4" httpUri="https://binary.pensoft.net/fig/695641" pageId="0" pageNumber="71">4D</figureCitation>
), one printed in MHNG, ' 
<taxonomicName authorityName="Stephens" authorityYear="1827" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="genus">Platyderus</taxonomicName>
/ langidus [sic] Reiche / 
<normalizedToken originalValue="‘Syrie’">'Syrie'</normalizedToken>
[pencil crossed off] / Label MHNG 2010 [p, w]', and, one lectotype [or paralectotype] label added by the first revising authors in the context of this work.
</paragraph>
<paragraph pageId="0" pageNumber="71">A further fifth syntype exists in MNHN, but in view of its too damaged body and imprecise location, it was not subject of lectotype designation (see &quot;Imprecise locality&quot;).</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="materials_examined">
<paragraph pageId="0" pageNumber="71">Other material examined.</paragraph>
<paragraph pageId="0" pageNumber="71">
Imprecise locality: single specimen of uncertain sex, with missing left antennae, fore tarsi, middle and hind legs ' 
<taxonomicName class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus languida" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="languida">
<emphasis italics="true" pageId="0" pageNumber="71">Platyderus languida</emphasis>
</taxonomicName>
Reiche [h, w] // cotype de Reiche cite par Chaudoir [h, w] // Ex 
<normalizedToken originalValue="Musæo">Musaeo</normalizedToken>
Mniszech [p, w] // L. Bedel Vidit 1902. [p, w]', (MNHN; ex-collection Chaudoir, box 
<normalizedToken originalValue="Nº">N°</normalizedToken>
218 &quot; 
<emphasis italics="true" pageId="0" pageNumber="71">
<taxonomicName authorityName="Stephens" authorityYear="1827" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="genus">Platyderus</taxonomicName>
Oxycrepis Stolonis Loxandrus
</emphasis>
&quot;).
</paragraph>
<paragraph pageId="0" pageNumber="71">
<materialsCitation collectingDate="2022-01-01" collectingDateMax="2022-12-31" collectingDateMin="2022-01-01" country="Israel" location="Northern District" specimenCount="1" specimenCount-male="1" stateProvince="Northern District">
<emphasis bold="true" pageId="0" pageNumber="71">
<collectingCountry name="Israel">Israel</collectingCountry>
</emphasis>
: 
<collectingRegion country="Israel" name="Northern">Northern District</collectingRegion>
: 
<specimenCount type="male">1♂</specimenCount>
</materialsCitation>
, ' 
<materialsCitation collectingDate="1997-05-08" country="Israel" location="Gader" municipality="Hammat" specimenCount="1">
ISRAEL: 
<collectingMunicipality>Hammat</collectingMunicipality>
/ 
<location LSID="urn:lsid:plazi:treatment:5964744C623D5BDFAC9729475336017E:05B8022DC069857E91D35F9C27580E54" country="Israel" municipality="Hammat" name="Gader">Gader</location>
/ 
<collectingDate value="1997-05-08">8.v.1997</collectingDate>
/ V. 
<normalizedToken originalValue="CHIKATUNOV’">CHIKATUNOV'</normalizedToken>
(SMNH-TAU)
</materialsCitation>
. 
<materialsCitation collectingDate="2022-01-01" collectingDateMax="2022-12-31" collectingDateMin="2022-01-01" country="Israel" location="Central District" specimenCount="1" specimenCount-female="1" stateProvince="Central District">
<collectingRegion country="Israel" name="Central">Central District</collectingRegion>
: 
<specimenCount type="female">1♀</specimenCount>
</materialsCitation>
, ' 
<materialsCitation collectingDate="2022-01-01" collectingDateMax="2022-12-31" collectingDateMin="2022-01-01" country="Israel" location="Beit-Berl" specimenCount="1" specimenCount-female="1">
<location LSID="urn:lsid:plazi:treatment:5964744C623D5BDFAC9729475336017E:7A21785A7BC77A392D273D307B554898" country="Israel" name="Beit-Berl">Beit-Berl</location>
/ 13.2.68 // COLLECTED BY KUTY 
<normalizedToken originalValue="YEFENOF’">YEFENOF'</normalizedToken>
(SMNH-TAU); 
<specimenCount type="female">1♀</specimenCount>
</materialsCitation>
, ' 
<materialsCitation collectingDate="2022-01-01" collectingDateMax="2022-12-31" collectingDateMin="2022-01-01" collectorName="R. Hoffman'" country="Israel" location="Israel Netanya" specimenCount="1" specimenCount-male="1">
<location LSID="urn:lsid:plazi:treatment:5964744C623D5BDFAC9729475336017E:A3B736C55E47EB463B51875AF00B7BE7" country="Israel" name="Israel Netanya">Israel Netanya</location>
/ 30.11.96 / 
<collectorName>
R. 
<normalizedToken originalValue="Hoffman’">Hoffman'</normalizedToken>
</collectorName>
(SMNH-TAU); 
<specimenCount type="male">1♂</specimenCount>
</materialsCitation>
, ' 
<materialsCitation collectingDate="2022-01-01" collectingDateMax="2022-12-31" collectingDateMin="2022-01-01" collectorName="R. Hoffman'" country="Israel" location="Israel Netanya" specimenCount="1" specimenCount-female="1">
<location LSID="urn:lsid:plazi:treatment:5964744C623D5BDFAC9729475336017E:E107B2034224080774A1DA5080C15D55" country="Israel" name="Israel Netanya">Israel Netanya</location>
/ 14.12.96 / 
<collectorName>
R. 
<normalizedToken originalValue="Hoffman’">Hoffman'</normalizedToken>
</collectorName>
(SMNH-TAU); 
<specimenCount type="female">1♀</specimenCount>
</materialsCitation>
, ' 
<materialsCitation collectingDate="2022-01-01" collectingDateMax="2022-12-31" collectingDateMin="2022-01-01" collectorName="R. Hoffman'" country="Israel" location="Israel Netanya" specimenCount="1" specimenCount-male="1">
<location LSID="urn:lsid:plazi:treatment:5964744C623D5BDFAC9729475336017E:0963AD0D2DC99E9D3C0E923F0FCE3DCA" country="Israel" name="Israel Netanya">Israel Netanya</location>
/ 21.12.96 / 
<collectorName>
R. 
<normalizedToken originalValue="Hoffman’">Hoffman'</normalizedToken>
</collectorName>
(SMNH-TAU); 
<specimenCount type="male">1♂</specimenCount>
</materialsCitation>
, ' 
<materialsCitation collectingDate="1996-12-22" collectorName="V. Chikatunov'" country="Israel" location="Netanya" specimenCount="1" specimenCount-female="1">
ISRAEL: / 
<location LSID="urn:lsid:plazi:treatment:5964744C623D5BDFAC9729475336017E:67B4457465FC3577E3513B4D8B64EB0C" country="Israel" name="Netanya">Netanya</location>
<collectingDate value="1996-12-22">22.xii.1996</collectingDate>
/ 
<collectorName>
V. 
<normalizedToken originalValue="Chikatunov’">Chikatunov'</normalizedToken>
</collectorName>
(SMNH-TAU); 
<specimenCount type="female">1♀</specimenCount>
</materialsCitation>
, ' 
<materialsCitation collectingDate="2022-01-01" collectingDateMax="2022-12-31" collectingDateMin="2022-01-01" collectorName="R. Hoffman'" country="Israel" location="Israel Netanya" specimenCount="2" specimenCount-female="2">
<location LSID="urn:lsid:plazi:treatment:5964744C623D5BDFAC9729475336017E:4D5EB81CA844FB18745B3E5E09BF2529" country="Israel" name="Israel Netanya">Israel Netanya</location>
/ 30.12.96 / 
<collectorName>
R. 
<normalizedToken originalValue="Hoffman’">Hoffman'</normalizedToken>
</collectorName>
(NMNHS); 
<specimenCount type="female">2♀♀</specimenCount>
</materialsCitation>
, ' 
<materialsCitation collectingDate="2022-01-01" collectingDateMax="2022-12-31" collectingDateMin="2022-01-01" collectorName="R. Hoffman'" country="Israel" location="Israel Netanya" specimenCount="1" specimenCount-female="1">
<location LSID="urn:lsid:plazi:treatment:5964744C623D5BDFAC9729475336017E:6A3A0779027E5CEEFD198394A1D6894A" country="Israel" name="Israel Netanya">Israel Netanya</location>
/ 18.1.97 / 
<collectorName>
R. 
<normalizedToken originalValue="Hoffman’">Hoffman'</normalizedToken>
</collectorName>
(SMNH-TAU); 
<specimenCount type="female">1♀</specimenCount>
</materialsCitation>
, ' 
<materialsCitation collectingDate="2022-01-01" collectingDateMax="2022-12-31" collectingDateMin="2022-01-01" collectorName="R. Hoffman'" country="Israel" location="Israel Netanya" specimenCount="1">
<location LSID="urn:lsid:plazi:treatment:5964744C623D5BDFAC9729475336017E:56A7EC50FCA5565E9A9203A1057A12E8" country="Israel" name="Israel Netanya">Israel Netanya</location>
/ 1.3.97 / 
<collectorName>
R. 
<normalizedToken originalValue="Hoffman’">Hoffman'</normalizedToken>
</collectorName>
(SMNH-TAU)
</materialsCitation>
. 
<materialsCitation collectingDate="2022-01-01" collectingDateMax="2022-12-31" collectingDateMin="2022-01-01" country="Israel" location="Jerusalem District" specimenCount="1" specimenCount-female="1" stateProvince="Jerusalem District">
<collectingRegion country="Israel" name="Jerusalem">Jerusalem District</collectingRegion>
: 
<specimenCount type="female">1♀</specimenCount>
</materialsCitation>
, ' 
<materialsCitation collectingDate="2022-01-01" collectingDateMax="2022-12-31" collectingDateMin="2022-01-01" country="Israel" location="Frm." municipality="Jerosalim" specimenCount="1" specimenCount-male="1">
<collectingMunicipality>Jerosalim</collectingMunicipality>
// 
<taxonomicName class="Insecta" family="Carabidae" genus="Tapinopterus" kingdom="Animalia" lsidName="Tapinopterus insidiosus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="insidiosus">
<emphasis italics="true" pageId="0" pageNumber="71">Tapinopterus insidiosus</emphasis>
</taxonomicName>
<location LSID="urn:lsid:plazi:treatment:5964744C623D5BDFAC9729475336017E:4E007FF9669FD0A091AAA2BE5D8B7626" country="Israel" municipality="Jerosalim" name="Frm.">Frm.</location>
' (MFNB); 
<specimenCount type="male">1♂</specimenCount>
</materialsCitation>
, ' 
<materialsCitation collectingDate="2022-01-01" collectingDateMax="2022-12-31" collectingDateMin="2022-01-01" collectorName="A. Jedlicka" country="Israel" county="Syria" determinerName="Ing. Jedlicka'" location="Prague" municipality="National Museum" specimenCount="1" specimenCount-female="1" stateProvince="Jerusalem">
<collectingRegion country="Israel" name="Jerusalem">Jerusalem</collectingRegion>
/ 
<collectingCounty>Syria</collectingCounty>
// ex coll. 
<collectorName>
A. 
<normalizedToken originalValue="Jedlička">Jedlicka</normalizedToken>
</collectorName>
/ 
<collectingMunicipality>National Museum</collectingMunicipality>
/ 
<location LSID="urn:lsid:plazi:treatment:5964744C623D5BDFAC9729475336017E:70AB4AD40D16129A958943C542159871" country="Israel" county="Syria" municipality="National Museum" name="Prague" stateProvince="Jerusalem">Prague</location>
, 
<location LSID="urn:lsid:plazi:treatment:5964744C623D5BDFAC9729475336017E:CD1B1F3D75786E6A7930E0DB4BCBC80A" country="Israel" county="Syria" municipality="National Museum" name="Czech Republic" stateProvince="Jerusalem">Czech Republic</location>
// 
<taxonomicName baseAuthorityName="Reiche &amp; Saulcy" baseAuthorityYear="1855" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) languidus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="languidus" subGenus="Eremoderus">Platyderus Eremoderus languidus</taxonomicName>
/ 
<location LSID="urn:lsid:plazi:treatment:5964744C623D5BDFAC9729475336017E:5723A6A3E24F2FD47AE880224A5BA1E6" country="Israel" county="Syria" municipality="National Museum" name="Reiche" stateProvince="Jerusalem">Reiche</location>
/ det. 
<determinerName>
Ing. 
<normalizedToken originalValue="Jedlička’">Jedlicka'</normalizedToken>
</determinerName>
(NMPC); 
<specimenCount type="female">1♀</specimenCount>
</materialsCitation>
, ' 
<materialsCitation collectingDate="2022-01-01" collectingDateMax="2022-12-31" collectingDateMin="2022-01-01" country="Israel" location="Jerosalim" specimenCount="1" specimenCount-male="1">
<location LSID="urn:lsid:plazi:treatment:5964744C623D5BDFAC9729475336017E:CEE748A0ABF320C941EF19D6604F8F11" country="Israel" name="Jerosalim">Jerosalim</location>
30.XII.25' (MIZ); 
<specimenCount type="male">1♂</specimenCount>
</materialsCitation>
, 'PALESTINE. / F.S. Bodenheimer. // Press by / Imp. Inst. Ent. / B.M. 1944-97. // 
<materialsCitation collectingDate="2022-01-01" collectingDateMax="2022-12-31" collectingDateMin="2022-01-01" country="Israel" location="Samaria Area" municipality="Judea" specimenCount="1" specimenCount-female="1" stateProvince="Jerusalem">
<collectingRegion country="Israel" name="Jerusalem">Jerusalem</collectingRegion>
/ scopus / 26.XII.33' (NHMUK). 
<collectingMunicipality>Judea</collectingMunicipality>
and 
<location LSID="urn:lsid:plazi:treatment:5964744C623D5BDFAC9729475336017E:CF845E6DC36690B2724627D81C2636B9" country="Israel" municipality="Judea" name="Samaria Area" stateProvince="Jerusalem">Samaria Area</location>
: 
<specimenCount type="female">1♀</specimenCount>
</materialsCitation>
, 'PALESTINE / 
<materialsCitation collectingDate="1940-02-14" country="Israel" location="Bytinski-Salz" municipality="Jericho Road" specimenCount="1" specimenCount-male="1" stateProvince="Jerusalem">
<collectingRegion country="Israel" name="Jerusalem">Jerusalem</collectingRegion>
- / 
<collectingMunicipality>Jericho Road</collectingMunicipality>
/ 
<location LSID="urn:lsid:plazi:treatment:5964744C623D5BDFAC9729475336017E:0A34C0C81DD63B400D075167EC9037DC" country="Israel" municipality="Jericho Road" name="Bytinski-Salz" stateProvince="Jerusalem">Bytinski-Salz</location>
/ km 18. / 
<collectingDate value="1940-02-14">14.ii.1940</collectingDate>
' (SMNH-TAU); 
<specimenCount type="male">1♂</specimenCount>
</materialsCitation>
, ' 
<materialsCitation collectingDate="1940-02-17" collectorName="Bytinski-Salz'" country="Israel" county="Jericho" location="Palestine" municipality="Road" specimenCount="1" specimenCount-female="1" stateProvince="Jerusalem">
<collectingRegion country="Israel" name="Jerusalem">Jerusalem</collectingRegion>
<collectingCounty>Jericho</collectingCounty>
/ 
<collectingMunicipality>Road</collectingMunicipality>
km 18 / 
<location LSID="urn:lsid:plazi:treatment:5964744C623D5BDFAC9729475336017E:714E559515D3DA5CAB25272E1C1328F2" country="Israel" county="Jericho" municipality="Road" name="Palestine" stateProvince="Jerusalem">Palestine</location>
<collectingDate value="1940-02-17">17.ii.1940</collectingDate>
/ leg. 
<normalizedToken originalValue="Bytinski-Salz’">
<collectorName>Bytinski-Salz'</collectorName>
</normalizedToken>
(SMNH-TAU); 
<specimenCount type="female">1♀</specimenCount>
</materialsCitation>
, ' 
<materialsCitation collectingDate="2022-01-01" collectingDateMax="2022-12-31" collectingDateMin="2022-01-01" collectorName="Bytinski-Salz" country="Israel" determinerName="Ing. Jedlicka'" location="Kv. Schiller" specimenCount="3" specimenCount-female="1" specimenCount-male="2">
PALESTINE / 
<location LSID="urn:lsid:plazi:treatment:5964744C623D5BDFAC9729475336017E:D0D02F5F4D5B6000B657F33FA014ADB0" country="Israel" name="Kv. Schiller">Kv. Schiller</location>
/ 25.ii. 19 / leg. 
<collectorName>Bytinski-Salz</collectorName>
// 
<taxonomicName authorityName="Stephens" authorityYear="1827" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="genus">Platyderus</taxonomicName>
/ 
<taxonomicName baseAuthorityName="Reiche &amp; Saulcy" baseAuthorityYear="1855" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) languidus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="languidus" subGenus="Eremoderus">Platyderus Eremoderus languidus</taxonomicName>
R. / det. 
<determinerName>
Ing. 
<normalizedToken originalValue="Jedlička’">Jedlicka'</normalizedToken>
</determinerName>
(SMNH-TAU); 
<specimenCount type="male">2♂♂</specimenCount>
, 
<specimenCount type="female">1♀</specimenCount>
</materialsCitation>
, ' 
<materialsCitation collectingDate="2007-02-28" country="Israel" county="South-facing" location="Perat" municipality="Na" specimenCount="1" specimenCount-male="1">
ISRAEL: 
<collectingCounty>South-facing</collectingCounty>
/ slope of 
<collectingMunicipality>Na</collectingMunicipality>
ẖal 
<location LSID="urn:lsid:plazi:treatment:5964744C623D5BDFAC9729475336017E:70E1767D5D84BE86BB23EB64BFF4BB06" country="Israel" county="South-facing" municipality="Na" name="Perat">Perat</location>
[= Naẖal Prat] / 
<collectingDate value="2007-02-28">28.ii.2007</collectingDate>
/ L. 
<normalizedToken originalValue="FRIEDMAN’">FRIEDMAN'</normalizedToken>
(SMNH-TAU); 
<specimenCount type="male">1♂</specimenCount>
</materialsCitation>
, ' 
<materialsCitation collectingDate="2007-03-18" country="Israel" location="Eshkolot" specimenCount="1">
ISRAEL / 
<location LSID="urn:lsid:plazi:treatment:5964744C623D5BDFAC9729475336017E:6F8AF365E5690AFB00322AD0A13BE7F7" country="Israel" name="Eshkolot">Eshkolot</location>
/ 
<collectingDate value="2007-03-18">18.iii.2007</collectingDate>
/ I. 
<normalizedToken originalValue="SCHTIRBERG’">SCHTIRBERG'</normalizedToken>
(SMNH-TAU)
</materialsCitation>
. 
<materialsCitation collectingDate="2022-01-01" collectingDateMax="2022-12-31" collectingDateMin="2022-01-01" country="Israel" location="Southern District" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Southern District">
<collectingRegion country="Israel" name="Southern">Southern District</collectingRegion>
: 
<specimenCount type="male">1♂</specimenCount>
, 
<specimenCount type="female">1♀</specimenCount>
</materialsCitation>
, ' 
<materialsCitation collectingDate="2008-12-05" collectorName="Th. Assmann'" country="Israel" location="Nizzanim" specimenCount="1" specimenCount-female="1">
W-Israel / 
<location LSID="urn:lsid:plazi:treatment:5964744C623D5BDFAC9729475336017E:78B567EBCB98D5A35629E960E293FF17" country="Israel" name="Nizzanim">Nizzanim</location>
/ 
<locationDeviation location="Ashdod">S of Ashdod</locationDeviation>
// 
<collectingDate value="2008-12-05">5.XII.2008</collectingDate>
/ leg. 
<collectorName>
Th. 
<normalizedToken originalValue="Assmann’">Assmann'</normalizedToken>
</collectorName>
(cASL); 
<specimenCount type="female">1♀</specimenCount>
</materialsCitation>
, ' 
<materialsCitation collectingDate="2017-04-07" collectorName="I. Renan" country="Israel" location="Ashkelon" specimenCount="1" specimenCount-male="1">
<location LSID="urn:lsid:plazi:treatment:5964744C623D5BDFAC9729475336017E:32F1C5D204127B2D94F0FE6DC57A8559" country="Israel" name="Ashkelon">Ashkelon</location>
/ 
<collectingDate value="2017-04-07">7.4.2017</collectingDate>
/ 
<collectorName>I. Renan</collectorName>
// 
<normalizedToken originalValue="FS’">FS'</normalizedToken>
(cWR); 
<specimenCount type="male">1♂</specimenCount>
</materialsCitation>
, ' 
<materialsCitation collectingDate="2017-04-11" collectorName="I. Renan" country="Israel" location="Ashdod" specimenCount="1">
<location LSID="urn:lsid:plazi:treatment:5964744C623D5BDFAC9729475336017E:696307C296F0A9380D9DF6FC5C92A748" country="Israel" name="Ashdod">Ashdod</location>
/ 
<collectingDate value="2017-04-11">11.4.2017</collectingDate>
/ 
<collectorName>I. Renan</collectorName>
// 
<normalizedToken originalValue="GM’">GM'</normalizedToken>
(SMNH-TAU)
</materialsCitation>
.
</paragraph>
<paragraph pageId="0" pageNumber="71">
TME: 
<specimenCount type="generic">30 specimens</specimenCount>
. TGE: 
<specimenCount type="male">4♂♂</specimenCount>
, 
<specimenCount type="female">2♀♀</specimenCount>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="diagnosis">
<paragraph pageId="0" pageNumber="71">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="71">
A species of medium size for 
<taxonomicName authority="" authorityName="Jeanne" authorityYear="1996" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus)" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="subGenus" subGenus="Eremoderus">Platyderus (Eremoderus)</taxonomicName>
, with light reddish-brown integument, meso- and metatarsomeres dorsally convex and smooth (Fig. 
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Metatarsus, dorsal view. A. Platyderus (Eremoderus) brunneus brunneus Karsch, 1881, left metatarsus, female specimen, Aziziyah, Jafara District, Libya; B. P. (E.) brunneus ferrantei Reitter, 1909, left metatarsus, female specimen, Holot Haluza, Southern District, Israel; C. P. (E.) irakensis, sp. nov., left metatarsus (black arrows indicate longitudinal grooves on metatarsomeres 2, 3 and 4), holotype; D. P. (E.) jordanensis, sp. nov., right metatarsus, holotype; E. P. (E.) languidus (Reiche &amp; Saulcy, 1855), right metatarsus, male specimen, Na ẖal Prat, Judea and Samaria Area, Israel. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure6" httpUri="https://binary.pensoft.net/fig/695643" pageId="0" pageNumber="71">6E</figureCitation>
), and pronotum from widest point to apex slightly to moderately constricted (PW/PA: 1.31-1.41), with sides to base straight (rarely slightly concave). Posterior angles of pronotum subrounded, with tips not protruding laterally (most cases) or very slightly protruding (few cases - as in 
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. jordanensis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="jordanensis">
<emphasis italics="true" pageId="0" pageNumber="71">P. jordanensis</emphasis>
</taxonomicName>
sp. nov.). Median lobe of aedeagus (lateral view) with proximal part, including basal bulb, rather long and slender, forming an angle with distal part of lobe smaller compared to other members of the &quot; 
<taxonomicName baseAuthorityName="Reiche &amp; Saulcy" baseAuthorityYear="1855" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) languidus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="languidus" subGenus="Eremoderus">
<emphasis italics="true" pageId="0" pageNumber="71">Platyderus Eremoderus languidus</emphasis>
</taxonomicName>
&quot; group (Fig. 
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Median lobe of aedeagus, left lateral view (black arrows on D, F-H indicate convexity of ventral margin of apex of median lobe). A. Platyderus (Platyderus) sp., male specimen, Boz Daglar Mtn., Turkey; B. P. (Eremoderus) afghanistanicus, sp. nov., holotype; C. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; D. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; E. P. (E.) insignitus insignitus Bedel, 1902, male specimen, SW Tiznit, Sous-Massa Region, Morocco; F. P. (E.) jordanensis, sp. nov., holotype; G. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; H. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male specimen, Na ẖal Prat, Judea and Samaria Area, Israel. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure9" httpUri="https://binary.pensoft.net/fig/695646" pageId="0" pageNumber="71">9G, H</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="redescription">
<paragraph pageId="0" pageNumber="71">Redescription.</paragraph>
<paragraph pageId="0" pageNumber="71">
<emphasis bold="true" pageId="0" pageNumber="71">
<emphasis italics="true" pageId="0" pageNumber="71">Habitus</emphasis>
.
</emphasis>
Specimens of moderate to large size for 
<taxonomicName authorityName="Stephens" authorityYear="1827" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="71">Platyderus</emphasis>
</taxonomicName>
species (BL: 6.40-9.10 mm; BW: 2.20-3.15 mm), with elongate and moderate convex body (Fig. 
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Dorsal habitus. A. Platyderus (Eremoderus) insignitus insignitus Bedel, 1902, male specimen, SW Tiznit, Sous-Massa Region, Morocco; B. P. (E.) irakensis, sp. nov., holotype; C. P. (E.) jordanensis, sp. nov., holotype; D. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype and its labels; E. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male specimen, &quot; Jerusalem Syria &quot;, Jerusalem District, Israel. Scale bars: 1 mm." figureDoi="10.3897/dez.69.83840.figure4" httpUri="https://binary.pensoft.net/fig/695641" pageId="0" pageNumber="71">4D, E</figureCitation>
). 
<emphasis bold="true" pageId="0" pageNumber="71">
<emphasis italics="true" pageId="0" pageNumber="71">Measurements and ratios</emphasis>
.
</emphasis>
See Table 
<tableCitation captionStart="Table 3" captionStartId="T3" captionText="Table 3. Morphometric data for species of the ' Platyderus Eremoderus languidus ' group of Platyderus." httpUri="http://table.plazi.org/id/D5F1A345CA056C2DBA2597271787B1E8" pageId="0" pageNumber="71" tableUuid="D5F1A345CA056C2DBA2597271787B1E8">3</tableCitation>
. 
<emphasis bold="true" pageId="0" pageNumber="71">
<emphasis italics="true" pageId="0" pageNumber="71">Color and lustre</emphasis>
.
</emphasis>
Body and appendages uniformly yellow brown to reddish-brown, usually head darker than the rest of the body. Integument slightly to moderately shiny, head and pronotum as shiny as elytra. 
<emphasis bold="true" pageId="0" pageNumber="71">
<emphasis italics="true" pageId="0" pageNumber="71">Microsculpture and punctation</emphasis>
.
</emphasis>
Pronotum with evident microreticulation, sculpticells regular isodiametric. Elytra (including intervals, basal margin and lateral gutter) with distinct isodiametric sculpticells. Ventral surface with more or less well-impressed isodiametric or slight transverse sculpticells or with sculpticells less apparent or scarcely-visible. Head smooth, including area of frontal furrows, sometimes clypeus with fine and shallow longitudinal wrinkles, disc micropunctate. Pronotum surface largely smooth, only apical part medially in front of anterior transverse impression and basal area medially behind posterior transverse impression longitudinally wrinkled; basal foveae and adjacent lateral areas mostly smooth, rarely shallowly punctate. Elytral intervals with very superficial and scattered punctures, or without such ones. Abdominal ventrites 1-3 finely wrinkled. 
<emphasis bold="true" pageId="0" pageNumber="71">
<emphasis italics="true" pageId="0" pageNumber="71">Head</emphasis>
.
</emphasis>
About one-third or more as wide as pronotum (PW/HW= 1.27-1.38). Eyes long, slightly to moderately convex. Labrum subrectangular, shorter than clypeus, with anterior margin concave medially. Frontoclypeal suture distinct in middle, reduced to disappeared at sides. Frontal furrows small, punctiform. Paraorbital sulci fine, backward surpassing posterior margin of eye, almost reaching level of posterior supraorbital pore. 
<emphasis bold="true" pageId="0" pageNumber="71">
<emphasis italics="true" pageId="0" pageNumber="71">Thorax</emphasis>
.
</emphasis>
Pronotum one tenth to one fifth wider than long (PW/PL= 1.11-1.22), with widest point at second quarter. Anterior transverse impression absent, posterior transverse impression barely distinct. Sides sinuate, convex medially and anteriorly, straight to slightly concave posteriorly; anterior bead present laterally, lack or present in medial tenth; basal bead present laterally, reduced to absent in medial third to medial half. Metepisternum slightly longer than wide, MA/MI= 0.87-0.95. 
<emphasis bold="true" pageId="0" pageNumber="71">
<emphasis italics="true" pageId="0" pageNumber="71">Elytra</emphasis>
.
</emphasis>
Elongate, about one-and-a-half times or little more as long as elytra (EL/EW= 1.52-1.62), two-and-a-half-times as long as pronotum (EL/PL= 2.42-2.60), and a third to a half as wide as pronotum (EW/PW= 1.29-1.48), with widest point at medial third or third quarter. Parascutellar striole short, less impressed than other striae, not joining stria 1; striae 1-8 and striole superficially punctate; base of stria 1 ending in parascutellar pore, striae 2-6 reaching or not reaching basal bead, 7 ending before basal bead. Interval 3 with three discal setiferous punctures, as location of first varies (adjacent to stria 3, in midst of interval 3, or adjacent to stria 2). Umbilicate setiferous series mostly with 16 punctures on each elytron. 
<emphasis bold="true" pageId="0" pageNumber="71">
<emphasis italics="true" pageId="0" pageNumber="71">Legs</emphasis>
.
</emphasis>
Posterior side of profemur with one or two setae in basal third and one or two in medial third. Mesofemur mostly with 5 setiferous punctures on anterior side ventrally (Fig. 
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Mesofemora, anterior side (A-D: left mesofemur, E-I: right mesofemur; white arrows and dots indicate position of setiferous punctures). A. Platyderus (Eremoderus) weiratheri Maran, 1940, female specimen, Boz dag koey env., Izmir District, Turkey; B. P. (E.) felixi, sp. nov., topotype female paratype; C. P. (E.) iranicus, sp. nov., topotype female paratype; D. P. (E.) lassallei, sp. nov., female paratype, Iran, Mazandaran Province, E Qolqol; E. P. (E.) klapperichi, sp. nov., holotype; F. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; G. P. (E.) afghanistanicus, sp. nov., holotype; H. P. (E.) brunneus ferrantei Reitter, 1909, lectotype; I. P. (E.) languidus (Reiche &amp; Saulcy, 1855), female specimen, Netanya 1.3.97, Central District, Israel. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure5" httpUri="https://binary.pensoft.net/fig/695642" pageId="0" pageNumber="71">5I</figureCitation>
). Anterior side of metafemur ventrally with a few long setae, one in basal third and one to three in apical half. 
<emphasis bold="true" pageId="0" pageNumber="71">
<emphasis italics="true" pageId="0" pageNumber="71">Male genitalia</emphasis>
.
</emphasis>
Urite IX subovate to subtriangular, with proximal margin nearly symmetrical and subacuminate at apex (Fig. 
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Urite, ventral view. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, E: topotype male specimen; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) vrabeci, sp. nov., holotype; E. P. (E.) lassallei, sp. nov., holotype; F. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; G. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; H. P. (E.) afghanistanicus, sp. nov., holotype; I. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; J. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; K. P. (E.) jordanensis, sp. nov., holotype; L. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; M. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male specimen, Na ẖal Prat, Judea and Samaria Area, Israel; N. P. (Platyderus) sp., male specimen, Boz Daglar Mtn., Turkey; O. P. (P.) reticulatus (Chaudoir), male specimen, Tashehzeh, Nowshahr County, Mazandaran Province, Iran. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure7" httpUri="https://binary.pensoft.net/fig/695644" pageId="0" pageNumber="71">7L, M</figureCitation>
). Median lobe of aedeagus in lateral view slender, with long and narrow basal bulb, long and broader shaft, and short, almost straight apex, as angle between front and rear halves relatively small (Fig. 
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Median lobe of aedeagus, left lateral view (black arrows on D, F-H indicate convexity of ventral margin of apex of median lobe). A. Platyderus (Platyderus) sp., male specimen, Boz Daglar Mtn., Turkey; B. P. (Eremoderus) afghanistanicus, sp. nov., holotype; C. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; D. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; E. P. (E.) insignitus insignitus Bedel, 1902, male specimen, SW Tiznit, Sous-Massa Region, Morocco; F. P. (E.) jordanensis, sp. nov., holotype; G. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; H. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male specimen, Na ẖal Prat, Judea and Samaria Area, Israel. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure9" httpUri="https://binary.pensoft.net/fig/695646" pageId="0" pageNumber="71">9G, H</figureCitation>
); median lobe in ventral view straight, 3.5-3.7 times longer than wide; apical lamella (dorsal view) short, nearly symmetrical, rounded at tip, with right side straight to convex and left side barely concave. Internal sac in lateral view (Fig. 
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Median lobe of aedeagus, left lateral view (black arrows on D, F-H indicate convexity of ventral margin of apex of median lobe). A. Platyderus (Platyderus) sp., male specimen, Boz Daglar Mtn., Turkey; B. P. (Eremoderus) afghanistanicus, sp. nov., holotype; C. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; D. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; E. P. (E.) insignitus insignitus Bedel, 1902, male specimen, SW Tiznit, Sous-Massa Region, Morocco; F. P. (E.) jordanensis, sp. nov., holotype; G. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; H. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male specimen, Na ẖal Prat, Judea and Samaria Area, Israel. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure9" httpUri="https://binary.pensoft.net/fig/695646" pageId="0" pageNumber="71">9G, H</figureCitation>
) with ventral sclerite broadened and rounded distally; same in ventral view (Fig. 
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Median lobe of aedeagus, ventral view. A. Platyderus (Eremoderus) afghanistanicus, sp. nov., holotype; B. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; C. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; D. P. (E.) insignitus insignitus Bedel, 1902, male specimen, SW Tiznit, Sous-Massa Region, Morocco; E. P. (E.) jordanensis, sp. nov., holotype; F. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; G. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male specimen, Na ẖal Prat, Judea and Samaria Area, Israel. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure11" httpUri="https://binary.pensoft.net/fig/695648" pageId="0" pageNumber="71">11F, G</figureCitation>
), with dorsal sclerite large, reticulate, with lateral protuberances distinct, and ventral sclerite narrow and nearly straight. Right paramere as on Fig. 
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Right paramere, external face. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., holotype; E. P. (E.) vrabeci, sp. nov., holotype; F. P. (E.) lassallei, sp. nov., holotype; G. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; H. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; I. P. (E.) afghanistanicus, sp. nov., holotype; J. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; K. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; L. P. (E.) jordanensis, sp. nov., holotype; M. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; N. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male paralectotype, &quot; Syria &quot;. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure12" httpUri="https://binary.pensoft.net/fig/695649" pageId="0" pageNumber="71">12M, N</figureCitation>
. 
<emphasis bold="true" italics="true" pageId="0" pageNumber="71">Female genitalia</emphasis>
(Fig. 
<figureCitation captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Spermathecal complex and gonocoxites, ventral view. A. Platyderus (Eremoderus) brunneus brunneus Karsch, 1881, female specimen, Bukamash, Nuqat al Khams District, Libya; B. P. (E.) brunneus ferrantei Reitter, 1909, female specimen, Holot Haluza, Southern District, Israel; C. P. (E.) insignitus insignitus Bedel, 1902, female specimen, Plage Aglou, Sous-Massa Region, Morocco; D. P. (E.) irakensis, sp. nov., holotype; E. P. (E.) jordanensis, sp. nov., female paratype, N Wadi Musa, Ma'an Governorate, Jordan; F. P. (E.) languidus (Reiche &amp; Saulcy, 1855), female specimens, Israel, Netanya 22. xii. 1996, Central District, Israel. Scale bars: 0.5 mm. For abbreviations see captions on Fig. 13." figureDoi="10.3897/dez.69.83840.figure15" httpUri="https://binary.pensoft.net/fig/695652" pageId="0" pageNumber="71">15F</figureCitation>
). Apical gonocoxite semi-pointed apex and one dorsolateral ensiform seta. Spermathecal canal connected in basal third of receptaculum.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="comparisons">
<paragraph pageId="0" pageNumber="71">Comparisons.</paragraph>
<paragraph pageId="0" pageNumber="71">
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. languidus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="languidus">
<emphasis italics="true" pageId="0" pageNumber="71">P. languidus</emphasis>
</taxonomicName>
and 
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. jordanenis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="jordanenis">
<emphasis italics="true" pageId="0" pageNumber="71">P. jordanenis</emphasis>
</taxonomicName>
sp. nov. differ in characters noted in 
<normalizedToken originalValue="“Diagnosis”">&quot;Diagnosis&quot;</normalizedToken>
under the latter species.
</paragraph>
<paragraph pageId="0" pageNumber="71">
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. languidus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="languidus">
<emphasis italics="true" pageId="0" pageNumber="71">P. languidus</emphasis>
</taxonomicName>
distinguishes from both 
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. brunneus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="brunneus">
<emphasis italics="true" pageId="0" pageNumber="71">P. brunneus</emphasis>
</taxonomicName>
and 
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. irakensis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="irakensis">
<emphasis italics="true" pageId="0" pageNumber="71">P. irakensis</emphasis>
</taxonomicName>
sp. nov. by meso- and metatarsomeres dorsally neither flattened nor grooved (vs. meso- and metatarsomeres dorsally more or less flattened and grooved), see Fig. 
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Metatarsus, dorsal view. A. Platyderus (Eremoderus) brunneus brunneus Karsch, 1881, left metatarsus, female specimen, Aziziyah, Jafara District, Libya; B. P. (E.) brunneus ferrantei Reitter, 1909, left metatarsus, female specimen, Holot Haluza, Southern District, Israel; C. P. (E.) irakensis, sp. nov., left metatarsus (black arrows indicate longitudinal grooves on metatarsomeres 2, 3 and 4), holotype; D. P. (E.) jordanensis, sp. nov., right metatarsus, holotype; E. P. (E.) languidus (Reiche &amp; Saulcy, 1855), right metatarsus, male specimen, Na ẖal Prat, Judea and Samaria Area, Israel. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure6" httpUri="https://binary.pensoft.net/fig/695643" pageId="0" pageNumber="71">6E</figureCitation>
. Additionally, it is distinct from 
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. brunneus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="brunneus">
<emphasis italics="true" pageId="0" pageNumber="71">P. brunneus</emphasis>
</taxonomicName>
in the pronotum less cordate, with sides less concave and less constricted toward base. The species differs from 
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. irakensis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="irakensis">
<emphasis italics="true" pageId="0" pageNumber="71">P. irakensis</emphasis>
</taxonomicName>
sp. nov. by the less long elytra compared to the length of the elytra and pronotum (EL/EW: 1.52-1.62 and EL/PL: 2.42-2.60, vs. EL/EW: 1.62-1.63 and EL/PL: 2.65-2.72).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="habitat">
<paragraph pageId="0" pageNumber="71">Habitat.</paragraph>
<paragraph pageId="0" pageNumber="71">
The records from the Coastal Lowlands of Israel refer to dune habitats close to the shores of the Mediterranean Sea. 
<taxonomicName baseAuthorityName="Reiche &amp; Saulcy" baseAuthorityYear="1855" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus languidus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="languidus">
<emphasis italics="true" pageId="0" pageNumber="71">Platyderus languidus</emphasis>
</taxonomicName>
co-occurs there together with 
<taxonomicName class="Insecta" family="Carabidae" genus="Carabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carabus impressus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="impressus">
<emphasis italics="true" pageId="0" pageNumber="71">Carabus impressus</emphasis>
</taxonomicName>
Klug, 1832, 
<taxonomicName authorityName="Dejean" authorityYear="1828" class="Insecta" family="Carabidae" genus="Masoreus" kingdom="Animalia" lsidName="Masoreus aegyptiacus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="aegyptiacus">
<emphasis italics="true" pageId="0" pageNumber="71">Masoreus aegyptiacus</emphasis>
</taxonomicName>
Dejean, 1828, 
<taxonomicName authorityName="Renan &amp; Assmann" authorityYear="2018" class="Insecta" family="Carabidae" genus="Graphipterus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Graphipterus sharonae" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="sharonae">
<emphasis italics="true" pageId="0" pageNumber="71">Graphipterus sharonae</emphasis>
</taxonomicName>
Renan &amp; Assmann, 2018, and some eurytopic ground beetle species (see habitat characterization in 
<bibRefCitation DOI="https://doi.org/10.3897/zookeys.1.23" author="Assmann, T" journalOrPublisher="ZooKeys" pageId="0" pageNumber="71" pagination="9 - 22" refId="B6" refString="Assmann, T, Buse, J, Drees, C, Friedman, ALL, Levanony, T, Matern, A, Timm, A, Wrase, DW, 2008. The Carabus fauna of Israel - updated identification key, faunistics, and habitats (Coleoptera: Carabidae). ZooKeys 1: 9 - 22, DOI: https://doi.org/10.3897/zookeys.1.23" title="The Carabus fauna of Israel - updated identification key, faunistics, and habitats (Coleoptera: Carabidae)." url="https://doi.org/10.3897/zookeys.1.23" volume="1" year="2008">Assmann et al. 2008</bibRefCitation>
, 
<bibRefCitation author="Assmann, T" journalOrPublisher="Spixiana" pageId="0" pageNumber="71" pagination="49 - 69" refId="B7" refString="Assmann, T, Boutaud, E, Buse, J, Chikatunov, V, Drees, C, Friedman, ALL, Haerdtle, W, Homburg, K, Levanony, T, Marcus, T, Renan, I, Wrase, DW, 2015a. The ground beetle tribe Cyclosomini (Coleoptera, Carabidae) in Israel. Spixiana 38 (1): 49 - 69" title="The ground beetle tribe Cyclosomini (Coleoptera, Carabidae) in Israel." volume="38" year="2015 a">2015a</bibRefCitation>
; 
<bibRefCitation DOI="https://doi.org/10.3897/zookeys.753.22366" author="Renan, I" journalOrPublisher="ZooKeys" pageId="0" pageNumber="71" pagination="23 - 82" refId="B46" refString="Renan, I, Assmann, T, Friedberg, A, 2018. Taxonomic Revision of the Graphipterus serrator (Forskal) group (Coleoptera, Carabidae): An increase from five to 15 valid species. ZooKeys 753: 23 - 82, DOI: https://doi.org/10.3897/zookeys.753.22366" title="Taxonomic Revision of the Graphipterus serrator (Forskal) group (Coleoptera, Carabidae): An increase from five to 15 valid species." url="https://doi.org/10.3897/zookeys.753.22366" volume="753" year="2018">Renan et al. 2018</bibRefCitation>
). Photographs of this type of habitat are given in 
<bibRefCitation DOI="https://doi.org/10.3897/zookeys.1.23" author="Assmann, T" journalOrPublisher="ZooKeys" pageId="0" pageNumber="71" pagination="9 - 22" refId="B6" refString="Assmann, T, Buse, J, Drees, C, Friedman, ALL, Levanony, T, Matern, A, Timm, A, Wrase, DW, 2008. The Carabus fauna of Israel - updated identification key, faunistics, and habitats (Coleoptera: Carabidae). ZooKeys 1: 9 - 22, DOI: https://doi.org/10.3897/zookeys.1.23" title="The Carabus fauna of Israel - updated identification key, faunistics, and habitats (Coleoptera: Carabidae)." url="https://doi.org/10.3897/zookeys.1.23" volume="1" year="2008">Assmann et al. (2008</bibRefCitation>
, fig. 17, p. 17, 2015a, fig. 8b, p. 63) and 
<bibRefCitation DOI="https://doi.org/10.3897/zookeys.753.22366" author="Renan, I" journalOrPublisher="ZooKeys" pageId="0" pageNumber="71" pagination="23 - 82" refId="B46" refString="Renan, I, Assmann, T, Friedberg, A, 2018. Taxonomic Revision of the Graphipterus serrator (Forskal) group (Coleoptera, Carabidae): An increase from five to 15 valid species. ZooKeys 753: 23 - 82, DOI: https://doi.org/10.3897/zookeys.753.22366" title="Taxonomic Revision of the Graphipterus serrator (Forskal) group (Coleoptera, Carabidae): An increase from five to 15 valid species." url="https://doi.org/10.3897/zookeys.753.22366" volume="753" year="2018">Renan et al. (2018</bibRefCitation>
, fig. 15, p. 63). The other habitats in Nahal Prat (= Wadi Kelt) must differ from these ones as there are no typical dune habitats.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="distribution">
<paragraph pageId="0" pageNumber="71">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="71">
<taxonomicName baseAuthorityName="Reiche &amp; Saulcy" baseAuthorityYear="1855" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus languidus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="languidus">
<emphasis italics="true" pageId="0" pageNumber="71">Platyderus languidus</emphasis>
</taxonomicName>
was cited for Morocco, Libya, Egypt (incl. Sinai), Syria and Israel (
<bibRefCitation author="Jedlicka, A" journalOrPublisher="Bulletin de la Societe Entomologique de France" pageId="0" pageNumber="71" refId="B26" refString="Jedlicka, A, 1963. Neue Carabiden aus Anatolien und vom Balkan. Koleopterologische Rundschau 40-41(1962-1963): 16-22." title="Neue Carabiden aus Anatolien und vom Balkan. Koleopterologische Rundschau 40 - 41 (1962 - 1963): 16 - 22." year="1963">
<normalizedToken originalValue="Jedlička">Jedlicka</normalizedToken>
1963
</bibRefCitation>
: 22; 
<bibRefCitation author="Hovorka, O" editor="Loebl, I" journalOrPublisher="Volume 1 Revised and Updated Edition. Archostemata - Myxophaga - Adephaga. Brill, Leiden - Boston" pageId="0" pageNumber="71" pagination="755 - 760" refId="B21" refString="Hovorka, O, 2017. Atranopsina Baehr, 1982. In: Loebl, I, Loebl, D, Eds., Catalogue of Palaearctic Coleoptera. Volume 1 Revised and Updated Edition. Archostemata - Myxophaga - Adephaga. Brill, Leiden - Boston: 755 - 760" title="Atranopsina Baehr, 1982." volumeTitle="Catalogue of Palaearctic Coleoptera." year="2017">Hovorka 2017</bibRefCitation>
: 756). 
<bibRefCitation author="Machard, P" journalOrPublisher="Coleopterist" pageId="0" pageNumber="71" pagination="101 - 102" refId="B36" refString="Machard, P, 2017. Un Platyderus nouveau du Maroc (Coleoptera, Carabidae). Coleopterist 20 (2): 101 - 102" title="Un Platyderus nouveau du Maroc (Coleoptera, Carabidae)." volume="20" year="2017">Machard (2017</bibRefCitation>
: 102, 
<bibRefCitation author="Machard, P" journalOrPublisher="Coleopterist" pageId="0" pageNumber="71" pagination="183 - 203" refId="B37" refString="Machard, P, 2019. Catalogue des Coleopteres Carabiques du Maroc (Coleoptera, Carabidae). Coleopterist 21 (3): 183 - 203" title="Catalogue des Coleopteres Carabiques du Maroc (Coleoptera, Carabidae)." volume="21" year="2019">2019</bibRefCitation>
: 199) included it in the list of the Moroccan 
<taxonomicName authorityName="Latreille" authorityYear="1802" class="Insecta" family="Carabidae" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="family">Carabidae</taxonomicName>
noting that its 
<normalizedToken originalValue="“Présence">&quot;Presence</normalizedToken>
au Maroc non 
<normalizedToken originalValue="confirmée”">confirmee&quot;</normalizedToken>
. The review of published and new material showed that the species does not live in Africa. All examined material of 
<taxonomicName authority="" authorityName="Jeanne" authorityYear="1996" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus)" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="subGenus" subGenus="Eremoderus">Platyderus (Eremoderus)</taxonomicName>
from Tunisia, Libya and Egypt should be referred to 
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. brunneus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="brunneus">
<emphasis italics="true" pageId="0" pageNumber="71">P. brunneus</emphasis>
</taxonomicName>
(see above). The species records of 
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. languidus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="languidus">
<emphasis italics="true" pageId="0" pageNumber="71">P. languidus</emphasis>
</taxonomicName>
for Morocco are also based on former misinterpretations and should concern 
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. insignitus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="insignitus">
<emphasis italics="true" pageId="0" pageNumber="71">P. insignitus</emphasis>
</taxonomicName>
. Certainly occurs only in Israel (Fig. 
<figureCitation captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Distribution of species from subgenus Platyderus Eremoderus in the Levant and the Arabian Peninsula: Platyderus arabicus sp. nov. (yellow question marks), P. brunki sp. nov. (blue circle), P. irakensis sp. nov. (green circles), P. jordanensis sp. nov. (brown circles), and P. languidus (red circles)." figureDoi="10.3897/dez.69.83840.figure19" httpUri="https://binary.pensoft.net/fig/695656" pageId="0" pageNumber="71">19</figureCitation>
). May live also in the southernmost area of Syria.
</paragraph>
<caption doi="10.3897/dez.69.83840.figure19" httpUri="https://binary.pensoft.net/fig/695656" pageId="0" pageNumber="71" start="Figure 19" startId="F19">
<paragraph pageId="0" pageNumber="71">
<emphasis bold="true" pageId="0" pageNumber="71">Figure 19.</emphasis>
Distribution of species from subgenus 
<taxonomicName authorityName="Jeanne" authorityYear="1996" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus)" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="subGenus" subGenus="Eremoderus">Platyderus Eremoderus</taxonomicName>
in the Levant and the Arabian Peninsula: 
<taxonomicName authorityName="Guéorguiev &amp; Wrase &amp; Assmann &amp; Muilwijk &amp; Machard" authorityYear="2022" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus arabicus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="arabicus">
<emphasis italics="true" pageId="0" pageNumber="71">Platyderus arabicus</emphasis>
</taxonomicName>
sp. nov. (yellow question marks), 
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. brunki" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="brunki">
<emphasis italics="true" pageId="0" pageNumber="71">P. brunki</emphasis>
</taxonomicName>
sp. nov. (blue circle), 
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. irakensis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="irakensis">
<emphasis italics="true" pageId="0" pageNumber="71">P. irakensis</emphasis>
</taxonomicName>
sp. nov. (green circles), 
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. jordanensis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="jordanensis">
<emphasis italics="true" pageId="0" pageNumber="71">P. jordanensis</emphasis>
</taxonomicName>
sp. nov. (brown circles), and 
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. languidus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="languidus">
<emphasis italics="true" pageId="0" pageNumber="71">P. languidus</emphasis>
</taxonomicName>
(red circles).
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="notes">
<paragraph pageId="0" pageNumber="71">Notes.</paragraph>
<paragraph pageId="0" pageNumber="71">
<normalizedToken originalValue="Léon">Leon</normalizedToken>
Fairmaire described his 
<taxonomicName authorityName="Fairmaire" authorityYear="1872" class="Insecta" family="Carabidae" genus="Sphodrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sphodrus parumstriatus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="parumstriatus">
<emphasis italics="true" pageId="0" pageNumber="71">Sphodrus parumstriatus</emphasis>
</taxonomicName>
without indication of an exact locality. This author mentioned only that it comes from &quot; 
<emphasis italics="true" pageId="0" pageNumber="71">
La 
<normalizedToken originalValue="localité">localite</normalizedToken>
<normalizedToken originalValue="qu’il">qu'il</normalizedToken>
habite est inconnue, mais il est plus que probable 
<normalizedToken originalValue="qu’il">qu'il</normalizedToken>
appartient 
<normalizedToken originalValue="à">a</normalizedToken>
la faune 
<normalizedToken originalValue="méditerranénne">mediterranenne</normalizedToken>
</emphasis>
&quot;. Later on, 
<bibRefCitation DOI="https://doi.org/10.3406/bsef.1906.23895" author="Bedel, L" journalOrPublisher="Bulletin de la Societe Entomologique de France" pageId="0" pageNumber="71" pagination="91 - 93" refId="B12" refString="Bedel, L, 1906. Communications. Synonymies de Coleopteres palearctiques. Bulletin de la Societe Entomologique de France 1906 (8): 91 - 93, DOI: https://doi.org/10.3406/bsef.1906.23895" title="Communications. Synonymies de Coleopteres palearctiques." url="https://doi.org/10.3406/bsef.1906.23895" volume="1906" year="1906">Bedel (1906</bibRefCitation>
: 91) established the synonymy of 
<taxonomicName genus="S." lsidName="S. parumstriatus" pageId="0" pageNumber="71" rank="species" species="parumstriatus">
<emphasis italics="true" pageId="0" pageNumber="71">S. parumstriatus</emphasis>
</taxonomicName>
with 
<taxonomicName authorityName="Reiche &amp; Saulcy" authorityYear="1855" class="Insecta" family="Carabidae" genus="Feronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Feronia languida" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="languida">
<emphasis italics="true" pageId="0" pageNumber="71">Feronia languida</emphasis>
</taxonomicName>
and stated: &quot; 
<emphasis italics="true" pageId="0" pageNumber="71">
Le type, dont la provenance restait 
<normalizedToken originalValue="ignorée">ignoree</normalizedToken>
, provient manifestement des 
<normalizedToken originalValue="récoltes">recoltes</normalizedToken>
de F. de Saulcy en Palestine. 
<normalizedToken originalValue="J’ai">J'ai</normalizedToken>
pu 
<normalizedToken originalValue="l’examiner">l'examiner</normalizedToken>
<normalizedToken originalValue="grâce">grace</normalizedToken>
<normalizedToken originalValue="à">a</normalizedToken>
<normalizedToken originalValue="l’extrème">l'extreme</normalizedToken>
<normalizedToken originalValue="amabilité">amabilite</normalizedToken>
de M. J. Magnin, son possesseur actuel
</emphasis>
&quot;. In 2016, one of us (BG) looked for the type material of 
<taxonomicName genus="S." lsidName="S. parumstriatus" pageId="0" pageNumber="71" rank="species" species="parumstriatus">
<emphasis italics="true" pageId="0" pageNumber="71">S. parumstriatus</emphasis>
</taxonomicName>
in MNHN, a depository where the ex-collection of L. Fairmaire is housed now, but nothing has been found. Subsequently, in a correspondence with BG (pers. comm.), Dr. Th. Deuve said: &quot; 
<emphasis italics="true" pageId="0" pageNumber="71">
Nous 
<normalizedToken originalValue="n’avons">n'avons</normalizedToken>
pas la collection Magnin et 
<normalizedToken originalValue="j’ignore">j'ignore</normalizedToken>
<normalizedToken originalValue="où">ou</normalizedToken>
elle se trouve. Certains 
<normalizedToken originalValue="spécimens">specimens</normalizedToken>
ont pu 
<normalizedToken originalValue="être">etre</normalizedToken>
<normalizedToken originalValue="dispersés">disperses</normalizedToken>
et pourraient 
<normalizedToken originalValue="être">etre</normalizedToken>
dans la collection du 
<normalizedToken originalValue="Muséum">Museum</normalizedToken>
, mais je ne connais pas le type de 
<taxonomicName authorityName="Fairmaire" authorityYear="1872" class="Insecta" family="Carabidae" genus="Sphodrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sphodrus parumstriatus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="parumstriatus">Sphodrus parumstriatus</taxonomicName>
Fairmaire. De toute 
<normalizedToken originalValue="façon">facon</normalizedToken>
, les types de Fairmaire sont toujours 
<normalizedToken originalValue="très">tres</normalizedToken>
difficile 
<normalizedToken originalValue="à">a</normalizedToken>
trouver. En 
<normalizedToken originalValue="général">general</normalizedToken>
, nous 
<normalizedToken originalValue="n’en">n'en</normalizedToken>
retrouvons que 50
</emphasis>
%&quot;.
</paragraph>
<paragraph pageId="0" pageNumber="71">
It is worth noting that the distance between the closest known localities of 
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. brunneus subsp. ferrantei" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="subSpecies" species="brunneus" subSpecies="ferrantei">
<emphasis italics="true" pageId="0" pageNumber="71">P. brunneus ferrantei</emphasis>
</taxonomicName>
and 
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. languidus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="languidus">
<emphasis italics="true" pageId="0" pageNumber="71">P. languidus</emphasis>
</taxonomicName>
in south Israel, respectively Holot Haluza (being the northeasternmost locality of the former species) and Ashkelon (being the southwesternmost one of the latter species), is about 65 km in a straight line. By supposition, the two species occur in different types of habitats.
</paragraph>
</subSubSection>
</treatment>
</document>