treatments-xml/data/03/D2/47/03D24702391CFFE2FF69FD7330D8FE2A.xml

<document id="0C258043759A8B1B31ED75C80001C8D4" ID-CLB-Dataset="24604" ID-DOI="10.11646/zootaxa.4711.2.6" ID-GBIF-Dataset="57cfb289-89a3-48b0-901c-c76fb029d2c7" ID-ISSN="1175-5326" ID-Zenodo-Dep="3577059" ID-ZooBank="0A02D6A2-9DF4-415D-9A1C-51B313F59998" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1576479411414" checkinUser="plazi" docAuthor="Li, Xu, Che, Xing-Jin &amp; Zhou, Wei" docDate="2019" docId="03D24702391CFFE2FF69FD7330D8FE2A" docLanguage="en" docName="zootaxa.4711.2.6.pdf" docOrigin="Zootaxa 4711 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Homatula anteridorsalis Li &amp; Che &amp; Zhou 2019, sp. nov." docType="treatment" docVersion="8" lastPageNumber="340" masterDocId="FFEB3F7A391BFFE8FFFEFFA4311DFF85" masterDocTitle="Loaches of Homatula (Teleostei: Nemacheilidae) from the upper Salween River in Yunnan, China with description of three new species" masterLastPageNumber="348" masterPageNumber="330" pageNumber="337" updateTime="1698774435811" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="86294C30A7CAF5502F0F3062617090CF">Loaches of Homatula (Teleostei: Nemacheilidae) from the upper Salween River in Yunnan, China with description of three new species</mods:title>
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<mods:namePart id="DD53DFC4E2ED19AB7E6D1023034792B3">Li, Xu</mods:namePart>
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<mods:namePart id="2DD511381A5CB7E6FF18F9CD9C579F71">Che, Xing-Jin</mods:namePart>
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<taxonomicName id="4C7B8D97391CFFEFFF69FD7330A2FD77" authority="Li &amp; Che &amp; Zhou, 2019" authorityName="Li &amp; Che &amp; Zhou" authorityYear="2019" box="[151,447,727,754]" class="Actinopterygii" family="Nemacheilidae" genus="Homatula" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="337" phylum="Chordata" rank="species" species="anteridorsalis" status="sp. nov.">
<emphasis id="B90F2A06391CFFEFFF69FD7330A2FD77" bold="true" box="[151,447,727,754]" italics="true" pageId="7" pageNumber="337">Homatula anteridorsalis</emphasis>
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<taxonomicNameLabel id="A23C977D391CFFEFFE38FD7C3339FD77" box="[454,548,728,754]" pageId="7" pageNumber="337" rank="species">sp. nov.</taxonomicNameLabel>
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<subSubSection id="C361A59F391CFFEFFF69FD583052FC93" box="[151,335,763,790]" pageId="7" pageNumber="337" type="description">
<paragraph id="8BC4F614391CFFEFFF69FD583052FC93" blockId="7.[151,548,727,790]" box="[151,335,763,790]" pageId="7" pageNumber="337">
(
<figureCitation id="1340EA91391CFFEFFF5EFD5831E5FC93" box="[160,248,764,790]" captionStart="FIGURE 2" captionStartId="5.[151,250,932,957]" captionTargetBox="[155,1432,320,847]" captionTargetId="figure@5.[151,1436,271,892]" captionTargetPageId="5" captionText="FIGURE 2. Relationship between anterior end of adipose crest and anal-fin base of Homatula. (A) H. anteridorsalis, SWFC 1610019, paratype, 108.1 mm SL; (B) H. cryptoclathrata, SWFC 1002001, holotype, 138.1 mm SL; (C) H. nigra, SWFC 1901005, holotype, 99.8 mm SL; (D) H. pycnolepis, SWFC 0612085, 99.6 mm SL; (E) H. laxiclathra, SWFC 1810010, 97.0 mm SL); (F) H. nanpanjiangensis, SWFC 1903001, 97.1 mm SL." figureDoi="http://doi.org/10.5281/zenodo.3577063" httpUri="https://zenodo.org/record/3577063/files/figure.png" pageId="7" pageNumber="337">Fig. 2A</figureCitation>
, 
<figureCitation id="1340EA91391CFFEFFEFAFD58305BFC93" box="[260,326,763,790]" captionStart="FIGURE 6" captionStartId="8.[151,250,763,788]" captionTargetBox="[151,1435,174,722]" captionTargetId="figure@8.[151,1435,174,722]" captionTargetPageId="8" captionText="FIGURE 6. Homatula anteriordorsalis, SWFC 1610006, holotype, 121.6 mm SL; China: Yunnan: Baoshan City: Longyang District: Lujiang Town: Bingmen Village." figureDoi="http://doi.org/10.5281/zenodo.3577073" httpUri="https://zenodo.org/record/3577073/files/figure.png" pageId="7" pageNumber="337">Fig. 6</figureCitation>
)
</paragraph>
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<paragraph id="8BC4F614391CFFEFFF69FC9A35C8FCF9" blockId="7.[151,1437,830,2045]" pageId="7" pageNumber="337">
<materialsCitation id="3B13FC49391CFFEFFF69FC9A35C8FCF9" ID-GBIF-Occurrence="2512877915" collectingDate="2016-10-08" collectorName="W. Dao &amp; Wang" country="China" county="Baoshan City" latitude="25.069786" location="Lujiang Town" longLatPrecision="1" longitude="98.83271" municipality="Longyang District" pageId="7" pageNumber="337" specimenCount="1" stateProvince="Yunnan" typeStatus="holotype">
<emphasis id="B90F2A06391CFFEFFF69FC9A3016FCDD" bold="true" box="[151,267,830,856]" pageId="7" pageNumber="337">
<typeStatus id="54C048B6391CFFEFFF69FC9A301BFCDD" box="[151,262,830,856]" pageId="7" pageNumber="337" type="holotype">Holotype</typeStatus>
.
</emphasis>
SWFC 
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SL; 
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: 
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: 
<collectingCounty id="62A58E98391CFFEFFCA2FC9A32E3FCDD" box="[860,1022,830,856]" pageId="7" pageNumber="337">Baoshan City</collectingCounty>
: 
<collectingMunicipality id="6BA06C6E391CFFEFFBF3FC9B35FEFCDD" box="[1037,1251,830,857]" pageId="7" pageNumber="337">Longyang District</collectingMunicipality>
: 
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, 
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(
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<geoCoordinate id="EE4F90D3391CFFEFFDFBFCC533B3FCF9" box="[517,686,865,892]" degrees="98" direction="east" minutes="49" orientation="longitude" pageId="7" pageNumber="337" precision="1" seconds="57.77" value="98.83271">98°49'57.77&quot;E</geoCoordinate>
); collected by 
<collectorName id="268E93C2391CFFEFFCAEFCC732B8FCF8" box="[848,933,867,893]" pageId="7" pageNumber="337">W. Dao</collectorName>
&amp; YC. 
<collectorName id="268E93C2391CFFEFFC05FCC73522FCF8" box="[1019,1087,867,893]" pageId="7" pageNumber="337">Wang</collectorName>
, 
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<collectingDate id="EF81293C391CFFEFFBB4FCC635CCFCF9" box="[1098,1233,866,892]" pageId="7" pageNumber="337" value="2016-10-08">8 Oct. 2016</collectingDate>
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.
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</paragraph>
<paragraph id="8BC4F614391CFFEFFF39FC2235EDFC6D" blockId="7.[151,1437,830,2045]" pageId="7" pageNumber="337">
<materialsCitation id="3B13FC49391CFFEFFF39FC2235F1FC6D" ID-GBIF-Occurrence="2512877914" collectingDate="2015-10-19" collectorName="C. Sun" country="China" county="Baoshan City" latitude="25.08158" location="Lujiang Town" longLatPrecision="1" longitude="98.840034" municipality="Longyang District" pageId="7" pageNumber="337" specimenCount="32" stateProvince="Yunnan" typeStatus="paratype">
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<typeStatus id="54C048B6391CFFEFFF39FC22305CFC25" box="[199,321,902,928]" pageId="7" pageNumber="337" type="paratype">Paratypes</typeStatus>
.
</emphasis>
SWFC 1610001-1610005, 1610007-1610031, 
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, 
<quantity id="4C835BF1391CFFEFFC4DFC223578FC24" box="[947,1125,902,929]" metricMagnitude="-2" metricUnit="m" metricValue="9.790000000000001" metricValueMax="12.940000000000001" metricValueMin="6.640000000000001" pageId="7" pageNumber="337" unit="mm" value="97.9" valueMax="129.4" valueMin="66.4">66.4–129.4 mm</quantity>
SL; other data same as the holotype. SWFC 1510001-1510002, 
<specimenCount id="9D7D3D9D391CFFEFFDCBFC0E3370FC40" box="[565,621,938,965]" pageId="7" pageNumber="337" type="generic">2 ex.</specimenCount>
, 108.0– 
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SL; 
<collectingCountry id="F36CB684391CFFEFFC90FC0E32A8FC41" box="[878,949,938,964]" name="China" pageId="7" pageNumber="337">China</collectingCountry>
: 
<collectingRegion id="49BF38F6391CFFEFFC40FC0F3505FC40" box="[958,1048,939,965]" country="China" name="Yunnan" pageId="7" pageNumber="337">Yunnan</collectingRegion>
: 
<collectingCounty id="62A58E98391CFFEFFBDCFC0E35A3FC41" box="[1058,1214,938,964]" pageId="7" pageNumber="337">Baoshan City</collectingCounty>
: 
<collectingMunicipality id="6BA06C6E391CFFEFFB36FC0F3485FC41" box="[1224,1432,938,965]" pageId="7" pageNumber="337">Longyang District</collectingMunicipality>
: 
<location id="8EA4A0CF391CFFEFFF69FC6A3024FC6C" LSID="urn:lsid:plazi:treatment:03D24702391CFFE2FF69FD7330D8FE2A:8EA4A0CF391CFFEFFF69FC6A3024FC6C" box="[151,313,974,1001]" country="China" county="Baoshan City" latitude="25.08158" longLatPrecision="1" longitude="98.840034" municipality="Longyang District" name="Lujiang Town" pageId="7" pageNumber="337" stateProvince="Yunnan">Lujiang Town</location>
: 
<location id="8EA4A0CF391CFFEFFEBDFC6B331EFC6D" LSID="urn:lsid:plazi:treatment:03D24702391CFFE2FF69FD7330D8FE2A:8EA4A0CF391CFFEFFEBDFC6B331EFC6D" box="[323,515,974,1001]" country="China" county="Baoshan City" latitude="25.08158" longLatPrecision="1" longitude="98.840034" municipality="Longyang District" name="Kungong Village" pageId="7" pageNumber="337" stateProvince="Yunnan">Kungong Village</location>
(
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); collected by 
<collectorName id="268E93C2391CFFEFFC02FC6A3556FC6D" box="[1020,1099,974,1000]" pageId="7" pageNumber="337">C. Sun</collectorName>
, 
<date id="FFC5D0D4391CFFEFFBA9FC6A35F1FC6D" box="[1111,1260,973,1000]" pageId="7" pageNumber="337" value="2015-10-19">
<collectingDate id="EF81293C391CFFEFFBA9FC6A35F1FC6D" box="[1111,1260,973,1000]" pageId="7" pageNumber="337" value="2015-10-19">19 Oct. 2015</collectingDate>
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.
</paragraph>
</subSubSection>
<subSubSection id="C361A59F391CFFEFFF39FC55305DFA39" pageId="7" pageNumber="337" type="diagnosis">
<paragraph id="8BC4F614391CFFEFFF39FC55305DFA39" blockId="7.[151,1437,830,2045]" pageId="7" pageNumber="337">
<emphasis id="B90F2A06391CFFEFFF39FC55305FFB89" bold="true" box="[199,322,1009,1036]" pageId="7" pageNumber="337">Diagnosis.</emphasis>
<taxonomicName id="4C7B8D97391CFFEFFEB6FC563341FB89" authorityName="Li &amp; Che &amp; Zhou" authorityYear="2019" box="[328,604,1010,1036]" class="Actinopterygii" family="Nemacheilidae" genus="Homatula" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="337" phylum="Chordata" rank="species" species="anteridorsalis">
<emphasis id="B90F2A06391CFFEFFEB6FC563341FB89" box="[328,604,1010,1036]" italics="true" pageId="7" pageNumber="337">Homatula anteridorsalis</emphasis>
</taxonomicName>
differs from other species of the densely-scaled group in 
<taxonomicName id="4C7B8D97391CFFEFFB1EFC56344DFB89" authorityName="Nichols" authorityYear="1925" box="[1248,1360,1010,1036]" class="Actinopterygii" family="Nemacheilidae" genus="Homatula" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="337" phylum="Chordata" rank="genus">
<emphasis id="B90F2A06391CFFEFFB1EFC56344DFB89" box="[1248,1360,1010,1036]" italics="true" pageId="7" pageNumber="337">Homatula</emphasis>
</taxonomicName>
by the following combination of characters: bars on flank across lateral line more than or equal to 25 (vs. less than or equal to 
<quantity id="4C835BF1391CFFEFFF4AFB9E31F1FBD1" box="[180,236,1082,1108]" metricMagnitude="-1" metricUnit="m" metricValue="5.08" pageId="7" pageNumber="337" unit="in" value="20.0">20 in</quantity>
<taxonomicName id="4C7B8D97391CFFEFFF0DFB9F30B8FBD1" baseAuthorityName="Zhou &amp; He" baseAuthorityYear="1993" box="[243,421,1082,1108]" class="Actinopterygii" family="Nemacheilidae" genus="Homatula" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="337" phylum="Chordata" rank="species" species="acuticephala">
<emphasis id="B90F2A06391CFFEFFF0DFB9F30B8FBD1" box="[243,421,1082,1108]" italics="true" pageId="7" pageNumber="337">H. acuticephala</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C7B8D97391CFFEFFE25FB9F336DFBD1" baseAuthorityName="Zhu &amp; Cao" baseAuthorityYear="1988" box="[475,624,1082,1108]" class="Actinopterygii" family="Nemacheilidae" genus="Homatula" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="337" phylum="Chordata" rank="species" species="erhaiensis">
<emphasis id="B90F2A06391CFFEFFE25FB9F336DFBD1" box="[475,624,1082,1108]" italics="true" pageId="7" pageNumber="337">H. erhaiensis</emphasis>
</taxonomicName>
); bars on flank vertical along lateral line (vs. no marks on the flank in 
<taxonomicName id="4C7B8D97391CFFEFFA7FFB9F31C9FBFD" authorityName="Li &amp; Che &amp; Zhou" authorityYear="2019" class="Actinopterygii" family="Nemacheilidae" genus="Homatula" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="337" phylum="Chordata" rank="species" species="nigra">
<emphasis id="B90F2A06391CFFEFFA7FFB9F31C9FBFD" italics="true" pageId="7" pageNumber="337">H. nigra</emphasis>
</taxonomicName>
); no free protrusions between the pelvic fins (vs. pair of free protrusions between pelvic fins in 
<taxonomicName id="4C7B8D97391CFFEFFB0CFBFB3496FBFD" authorityName="Hu &amp; Zhang" authorityYear="2010" box="[1266,1419,1118,1144]" class="Actinopterygii" family="Nemacheilidae" genus="Homatula" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="337" phylum="Chordata" rank="species" species="pycnolepis">
<emphasis id="B90F2A06391CFFEFFB0CFBFB3496FBFD" box="[1266,1419,1118,1144]" italics="true" pageId="7" pageNumber="337">H. pycnolepis</emphasis>
</taxonomicName>
); gill opening smaller, its upper angle level with lower edge of eye (vs. gill opening larger, its upper angle aligned with middle point of eye in 
<taxonomicName id="4C7B8D97391CFFEFFE27FB033393FB45" baseAuthorityName="Zhou &amp; He" baseAuthorityYear="1993" box="[473,654,1190,1216]" class="Actinopterygii" family="Nemacheilidae" genus="Homatula" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="337" phylum="Chordata" rank="species" species="acuticephala">
<emphasis id="B90F2A06391CFFEFFE27FB033393FB45" box="[473,654,1190,1216]" italics="true" pageId="7" pageNumber="337">H. acuticephala</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C7B8D97391CFFEFFD60FB033249FB45" baseAuthorityName="Zhu &amp; Wang" baseAuthorityYear="1985" box="[670,852,1190,1216]" class="Actinopterygii" family="Nemacheilidae" genus="Homatula" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="337" phylum="Chordata" rank="species" species="anguillioides">
<emphasis id="B90F2A06391CFFEFFD60FB033249FB45" box="[670,852,1190,1216]" italics="true" pageId="7" pageNumber="337">H. anguillioides</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C7B8D97391CFFEFFC9AFB033512FB45" authorityName="Endruweit, Min &amp; Yang" authorityYear="2018" box="[868,1039,1190,1216]" class="Actinopterygii" family="Nemacheilidae" genus="Homatula" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="337" phylum="Chordata" rank="species" species="coccinocola">
<emphasis id="B90F2A06391CFFEFFC9AFB033512FB45" box="[868,1039,1190,1216]" italics="true" pageId="7" pageNumber="337">H. coccinocola</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C7B8D97391CFFEFFBE1FB0335ABFB45" baseAuthorityName="Zhu &amp; Cao" baseAuthorityYear="1988" box="[1055,1206,1190,1216]" class="Actinopterygii" family="Nemacheilidae" genus="Homatula" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="337" phylum="Chordata" rank="species" species="erhaiensis">
<emphasis id="B90F2A06391CFFEFFBE1FB0335ABFB45" box="[1055,1206,1190,1216]" italics="true" pageId="7" pageNumber="337">H. erhaiensis</emphasis>
</taxonomicName>
, and 
<taxonomicName id="4C7B8D97391CFFEFFB09FB03348EFB45" authorityName="Hu &amp; Zhang" authorityYear="2010" box="[1271,1427,1190,1216]" class="Actinopterygii" family="Nemacheilidae" genus="Homatula" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="337" phylum="Chordata" rank="species" species="pycnolepis">
<emphasis id="B90F2A06391CFFEFFB09FB03348EFB45" box="[1271,1427,1190,1216]" italics="true" pageId="7" pageNumber="337">H. pycnolepis</emphasis>
</taxonomicName>
) (
<figureCitation id="1340EA91391CFFEFFF5EFB6E31FEFB61" box="[160,227,1225,1252]" captionStart="FIGURE 5" captionStartId="7.[151,250,559,584]" captionTargetBox="[166,1421,174,540]" captionTargetId="figure@7.[166,1421,174,540]" captionTargetPageId="7" captionText="FIGURE 5. Gill openings of Homatula. (A) Larger gill opening; upper angle aligned with middle point of eye (H. pycnolepis, SWFC 0612085, 99.6 mm SL); (B) smaller gill opening; upper angle level with lower edge of eye (H. cryptoclathrata, SWFC 1610006, holotype, 121.6 mm SL)." figureDoi="http://doi.org/10.5281/zenodo.3577071" httpUri="https://zenodo.org/record/3577071/files/figure.png" pageId="7" pageNumber="337">Fig. 5</figureCitation>
); with pelvic axillary lobe (vs. without pelvic axillary lobe in 
<taxonomicName id="4C7B8D97391CFFEFFC6BFB6F3554FB61" baseAuthorityName="Zhu &amp; Wang" baseAuthorityYear="1985" box="[917,1097,1226,1252]" class="Actinopterygii" family="Nemacheilidae" genus="Homatula" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="337" phylum="Chordata" rank="species" species="anguillioides">
<emphasis id="B90F2A06391CFFEFFC6BFB6F3554FB61" box="[917,1097,1226,1252]" italics="true" pageId="7" pageNumber="337">H. anguillioides</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C7B8D97391CFFEFFB80FB6F342CFB61" baseAuthorityName="Zhou &amp; He" baseAuthorityYear="1993" box="[1150,1329,1226,1252]" class="Actinopterygii" family="Nemacheilidae" genus="Homatula" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="337" phylum="Chordata" rank="species" species="acuticephala">
<emphasis id="B90F2A06391CFFEFFB80FB6F342CFB61" box="[1150,1329,1226,1252]" italics="true" pageId="7" pageNumber="337">H. acuticephala</emphasis>
</taxonomicName>
); marked notch on lower jaw (vs. lower jaw spoon-shaped and lacking notch in 
<taxonomicName id="4C7B8D97391CFFEFFC62FB4B352CFA8D" baseAuthorityName="Zhu &amp; Cao" baseAuthorityYear="1988" box="[924,1073,1262,1288]" class="Actinopterygii" family="Nemacheilidae" genus="Homatula" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="337" phylum="Chordata" rank="species" species="erhaiensis">
<emphasis id="B90F2A06391CFFEFFC62FB4B352CFA8D" box="[924,1073,1262,1288]" italics="true" pageId="7" pageNumber="337">H. erhaiensis</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C7B8D97391CFFEFFB9BFB4B3405FA8D" authorityName="Min, Yang &amp; Chen" authorityYear="2012" box="[1125,1304,1262,1288]" class="Actinopterygii" family="Nemacheilidae" genus="Homatula" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="337" phylum="Chordata" rank="species" species="wuliangensis">
<emphasis id="B90F2A06391CFFEFFB9BFB4B3405FA8D" box="[1125,1304,1262,1288]" italics="true" pageId="7" pageNumber="337">H. wuliangensis</emphasis>
</taxonomicName>
); dorsal-fin base shorter than longest branched dorsal-fin ray (vs. longer than longest branched dorsal-fin ray in 
<taxonomicName id="4C7B8D97391CFFEFFB1AFAB7348BFAA9" baseAuthorityName="Zhou &amp; He" baseAuthorityYear="1993" box="[1252,1430,1298,1324]" class="Actinopterygii" family="Nemacheilidae" genus="Homatula" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="337" phylum="Chordata" rank="species" species="acuticephala">
<emphasis id="B90F2A06391CFFEFFB1AFAB7348BFAA9" box="[1252,1430,1298,1324]" italics="true" pageId="7" pageNumber="337">H. acuticephala</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C7B8D97391CFFEFFF69FA933051FAD5" baseAuthorityName="Zhu &amp; Wang" baseAuthorityYear="1985" box="[151,332,1334,1360]" class="Actinopterygii" family="Nemacheilidae" genus="Homatula" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="337" phylum="Chordata" rank="species" species="anguillioides">
<emphasis id="B90F2A06391CFFEFFF69FA933051FAD5" box="[151,332,1334,1360]" italics="true" pageId="7" pageNumber="337">H. anguillioides</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C7B8D97391CFFEFFEA4FA933336FAD5" authorityName="Li &amp; Che &amp; Zhou" authorityYear="2019" box="[346,555,1334,1360]" class="Actinopterygii" family="Nemacheilidae" genus="Homatula" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="337" phylum="Chordata" rank="species" species="cryptoclathrata">
<emphasis id="B90F2A06391CFFEFFEA4FA933336FAD5" box="[346,555,1334,1360]" italics="true" pageId="7" pageNumber="337">H. cryptoclathrata</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C7B8D97391CFFEFFDC7FA9333D2FAD5" baseAuthorityName="Zhu &amp; Cao" baseAuthorityYear="1988" box="[569,719,1334,1360]" class="Actinopterygii" family="Nemacheilidae" genus="Homatula" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="337" phylum="Chordata" rank="species" species="erhaiensis">
<emphasis id="B90F2A06391CFFEFFDC7FA9333D2FAD5" box="[569,719,1334,1360]" italics="true" pageId="7" pageNumber="337">H. erhaiensis</emphasis>
</taxonomicName>
, and 
<taxonomicName id="4C7B8D97391CFFEFFCF2FA9332DCFAD5" authorityName="Min, Yang &amp; Chen" authorityYear="2012" box="[780,961,1334,1360]" class="Actinopterygii" family="Nemacheilidae" genus="Homatula" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="337" phylum="Chordata" rank="species" species="wuliangensis">
<emphasis id="B90F2A06391CFFEFFCF2FA9332DCFAD5" box="[780,961,1334,1360]" italics="true" pageId="7" pageNumber="337">H. wuliangensis</emphasis>
</taxonomicName>
); dorsal and pelvic fins located anteriorly, predorsal length 44.4% (42.8–46.3%) SL, prepelvic length 45% (43.3–46.7 %) SL (vs. predorsal length 47.7% (45.7–49.6 %) SL, prepelvic length 48.7% (47.0–49.9 % SL) in all congeners of densely-scaled group of 
<taxonomicName id="4C7B8D97391CFFEFFADDFADA348EFA1D" authorityName="Nichols" authorityYear="1925" box="[1315,1427,1406,1432]" class="Actinopterygii" family="Nemacheilidae" genus="Homatula" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="337" phylum="Chordata" rank="genus">
<emphasis id="B90F2A06391CFFEFFADDFADA348EFA1D" box="[1315,1427,1406,1432]" italics="true" pageId="7" pageNumber="337">Homatula</emphasis>
</taxonomicName>
) (
<tableCitation id="C6F9C3AF391CFFEFFF61FA0631E3FA39" box="[159,254,1442,1468]" captionStart="TABLE 1" captionStartId="2.[151,239,579,605]" captionTargetBox="[165,2015,624,1571]" captionText="TABLE 1. Comparisons of characteristic among the species of the densely-scaled group of Homatula" httpUri="http://table.plazi.org/id/DF04A69C3919FFEAFF69FDE735C8FDD8" pageId="7" pageNumber="337" tableUuid="DF04A69C3919FFEAFF69FDE735C8FDD8">Tables 1</tableCitation>
&amp; 
<tableCitation id="C6F9C3AF391CFFEFFEDCFA06302EFA39" box="[290,307,1442,1468]" captionStart="TABLE 2" captionStartId="14.[151,239,470,496]" captionTargetBox="[165,1975,507,1700]" captionText="TABLE 2. Morphometric data of three new Homatula species and H. pycnolepis" httpUri="http://table.plazi.org/id/DF04A69C3915FFE6FF69FE7232E8FE75" pageId="7" pageNumber="337" tableUuid="DF04A69C3915FFE6FF69FE7232E8FE75">2</tableCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C361A59F391CFFE1FF39FA613315FD2E" lastPageId="9" lastPageNumber="339" pageId="7" pageNumber="337" type="description">
<paragraph id="8BC4F614391CFFEFFF39FA613519F9AC" blockId="7.[151,1437,830,2045]" pageId="7" pageNumber="337">
<emphasis id="B90F2A06391CFFEFFF39FA613044FA65" bold="true" box="[199,345,1477,1504]" pageId="7" pageNumber="337">Description.</emphasis>
Based on 
<typeStatus id="54C048B6391CFFEFFE33FA623332FA65" box="[461,559,1478,1504]" pageId="7" pageNumber="337" type="holotype">holotype</typeStatus>
and 
<specimenCount id="9D7D3D9D391CFFEFFD98FA6233EBFA64" box="[614,758,1478,1505]" pageId="7" pageNumber="337" type="generic" typeStatus="paratypes">32 paratypes</specimenCount>
, maximum standard length is 
<quantity id="4C835BF1391CFFEFFBB8FA6235A6FA64" box="[1094,1211,1478,1505]" metricMagnitude="-1" metricUnit="m" metricValue="1.408" pageId="7" pageNumber="337" unit="mm" value="140.8">140.8 mm</quantity>
. Morphometric data are shown in 
<tableCitation id="C6F9C3AF391CFFEFFECDFA4E3090F981" box="[307,397,1514,1540]" captionStart="TABLE 2" captionStartId="14.[151,239,470,496]" captionTargetBox="[165,1975,507,1700]" captionText="TABLE 2. Morphometric data of three new Homatula species and H. pycnolepis" httpUri="http://table.plazi.org/id/DF04A69C3915FFE6FF69FE7232E8FE75" pageId="7" pageNumber="337" tableUuid="DF04A69C3915FFE6FF69FE7232E8FE75">Table 2</tableCitation>
. Dorsal-fin rays iii, 8½; anal-fin rays iii, 5½; pectoral-fin rays i, 9–10; pelvic-fin rays i, 7; caudal-fin branched rays 9+8; vertebrae 4+41 (
<specimenCount id="9D7D3D9D391CFFEFFD58F9AA322FF9AC" box="[678,818,1550,1577]" pageId="7" pageNumber="337" type="generic">2 specimens</specimenCount>
)–42 (
<specimenCount id="9D7D3D9D391CFFEFFC88F9AA32E5F9AC" box="[886,1016,1550,1577]" pageId="7" pageNumber="337" type="generic">1 specimen</specimenCount>
).
</paragraph>
<paragraph id="8BC4F614391CFFEFFF39F99632D6F93D" blockId="7.[151,1437,830,2045]" pageId="7" pageNumber="337">Body elongate, cylindrical, anteriorly slightly depressed, posteriorly compressed laterally. Maximum body depth immediately in front of dorsal fin. Dorsal and ventral profiles almost straight. Except for scaleless head and thorax, body is entirely covered by small scales, with scattered scales on abdomen. Caudal peduncle compressed laterally. Lateral line complete, extending directly along midline of body.</paragraph>
<paragraph id="8BC4F614391CFFEFFF39F9663223F8CD" blockId="7.[151,1437,830,2045]" pageId="7" pageNumber="337">Head short and depressed in frontal view, wider than high, roughly triangular in dorsal view. Snout blunt, slightly shorter than postorbital length of head. Eye small, close to dorsal profile of head, invisible from ventral view. Interorbital space wide and flat. Nostrils closely set, nearer to anterior margin of eye than to snout tip; anterior nare pierced in front side of a flap-like tube with pointed tip.</paragraph>
<paragraph id="8BC4F614391CFFEFFF39F8F635ECF879" blockId="7.[151,1437,830,2045]" pageId="7" pageNumber="337">
Mouth inferior and arched. Lips thick, slightly furrowed, but not papillated; upper lip with small median incision and lower lip with marked median incision. Jaws covered by lips; upper jaw with well developed processus dentiformis (described by 
<bibRefCitation id="EFEA8BE5391CFFEFFE3DF83E3342F831" author="Kottelat, M." box="[451,607,1946,1972]" pageId="7" pageNumber="337" refId="ref10665" refString="Kottelat, M. (1990) Indochinese Nemacheilines. A Revision of Nemacheiline Loaches (Pisces: Cypriniformes) of Thailand, Burma, Laos, Cambodia and southern Viet Nam. Pfeil, Munchen, 262 pp." type="book" year="1990">Kottelat 1990</bibRefCitation>
) corresponding with marked notch on lower jaw. Three pairs of barbels; two rostral pairs, inner one not reaching corner of mouth and outer one reaching vertical line of anterior nostril; one maxillary pair long, reaching to vertical line between middle and posterior margin of eye (
<figureCitation id="1340EA91391CFFEFFB74F84635F9F879" box="[1162,1252,2018,2044]" captionStart="FIGURE 7" captionStartId="8.[151,250,1933,1958]" captionTargetBox="[217,1391,853,1887]" captionTargetId="figure@8.[151,1436,838,1899]" captionTargetPageId="8" captionText="FIGURE 7. Mouths of Homatula. (A) H. anteridorsalis, SWFC 1610006, holotype, 121.6 mm SL; (B) H. cryptoclathrata, SWFC 1002001, holotype, 138.1 mm SL; (C) H. nigra, SWFC 1901005, holotype, 99.8 mm SL; (D) H. pycnolepis, SWFC 0612085, 99.6 mm SL." figureDoi="http://doi.org/10.5281/zenodo.3577075" httpUri="https://zenodo.org/record/3577075/files/figure.png" pageId="7" pageNumber="337">Fig. 7A</figureCitation>
).
</paragraph>
<caption id="DF04A69C3913FFE0FF69FD5F3322FCBD" ID-DOI="http://doi.org/10.5281/zenodo.3577073" ID-Zenodo-Dep="3577073" httpUri="https://zenodo.org/record/3577073/files/figure.png" pageId="8" pageNumber="338" startId="8.[151,250,763,788]" targetBox="[151,1435,174,722]" targetPageId="8">
<paragraph id="8BC4F6143913FFE0FF69FD5F3322FCBD" blockId="8.[151,1436,763,824]" pageId="8" pageNumber="338">
<emphasis id="B90F2A063913FFE0FF69FD5F3008FC91" bold="true" box="[151,277,763,788]" pageId="8" pageNumber="338">FIGURE 6.</emphasis>
<taxonomicName id="4C7B8D973913FFE0FEE3FD5F3333FC96" box="[285,558,763,787]" class="Actinopterygii" family="Nemacheilidae" genus="Homatula" kingdom="Animalia" order="Cypriniformes" pageId="8" pageNumber="338" phylum="Chordata" rank="species" species="anteriordorsalis">
<emphasis id="B90F2A063913FFE0FEE3FD5F3333FC96" box="[285,558,763,787]" italics="true" pageId="8" pageNumber="338">Homatula anteriordorsalis</emphasis>
</taxonomicName>
, SWFC 1610006, holotype, 121.6 mm SL; China: Yunnan: Baoshan City: Longyang District: Lujiang Town: Bingmen Village.
</paragraph>
</caption>
<caption id="DF04A69C3913FFE0FF69F829309FF86B" ID-DOI="http://doi.org/10.5281/zenodo.3577075" ID-Zenodo-Dep="3577075" httpUri="https://zenodo.org/record/3577075/files/figure.png" pageId="8" pageNumber="338" startId="8.[151,250,1933,1958]" targetBox="[217,1391,853,1887]" targetPageId="8">
<paragraph id="8BC4F6143913FFE0FF69F829309FF86B" blockId="8.[151,1436,1933,2030]" pageId="8" pageNumber="338">
<emphasis id="B90F2A063913FFE0FF69F8293008F823" bold="true" box="[151,277,1933,1958]" pageId="8" pageNumber="338">FIGURE 7.</emphasis>
Mouths of 
<taxonomicName id="4C7B8D973913FFE0FE6CF82A30EAF823" authorityName="Nichols" authorityYear="1925" box="[402,503,1934,1958]" class="Actinopterygii" family="Nemacheilidae" genus="Homatula" kingdom="Animalia" order="Cypriniformes" pageId="8" pageNumber="338" phylum="Chordata" rank="genus">
<emphasis id="B90F2A063913FFE0FE6CF82A30EAF823" box="[402,503,1934,1958]" italics="true" pageId="8" pageNumber="338">Homatula</emphasis>
</taxonomicName>
. (A) 
<taxonomicName id="4C7B8D973913FFE0FDCCF82B33FCF823" authorityName="Li &amp; Che &amp; Zhou" authorityYear="2019" box="[562,737,1934,1958]" class="Actinopterygii" family="Nemacheilidae" genus="Homatula" kingdom="Animalia" order="Cypriniformes" pageId="8" pageNumber="338" phylum="Chordata" rank="species" species="anteridorsalis">
<emphasis id="B90F2A063913FFE0FDCCF82B33FCF823" box="[562,737,1934,1958]" italics="true" pageId="8" pageNumber="338">H. anteridorsalis</emphasis>
</taxonomicName>
, SWFC 1610006, holotype, 121.6 mm SL; (B) 
<taxonomicName id="4C7B8D973913FFE0FB29F82B348BF823" authorityName="Li &amp; Che &amp; Zhou" authorityYear="2019" box="[1239,1430,1934,1958]" class="Actinopterygii" family="Nemacheilidae" genus="Homatula" kingdom="Animalia" order="Cypriniformes" pageId="8" pageNumber="338" phylum="Chordata" rank="species" species="cryptoclathrata">
<emphasis id="B90F2A063913FFE0FB29F82B348BF823" box="[1239,1430,1934,1958]" italics="true" pageId="8" pageNumber="338">H. cryptoclathrata</emphasis>
</taxonomicName>
, SWFC 1002001, holotype, 138.1 mm SL; (C) 
<taxonomicName id="4C7B8D973913FFE0FD82F81733C9F84F" authorityName="Li &amp; Che &amp; Zhou" authorityYear="2019" box="[636,724,1971,1994]" class="Actinopterygii" family="Nemacheilidae" genus="Homatula" kingdom="Animalia" order="Cypriniformes" pageId="8" pageNumber="338" phylum="Chordata" rank="species" species="nigra">
<emphasis id="B90F2A063913FFE0FD82F81733C9F84F" box="[636,724,1971,1994]" italics="true" pageId="8" pageNumber="338">H. nigra</emphasis>
</taxonomicName>
, SWFC 1901005, holotype, 99.8 mm SL; (D) 
<taxonomicName id="4C7B8D973913FFE0FB45F8173455F84F" authorityName="Hu &amp; Zhang" authorityYear="2010" box="[1211,1352,1970,1994]" class="Actinopterygii" family="Nemacheilidae" genus="Homatula" kingdom="Animalia" order="Cypriniformes" pageId="8" pageNumber="338" phylum="Chordata" rank="species" species="pycnolepis">
<emphasis id="B90F2A063913FFE0FB45F8173455F84F" box="[1211,1352,1970,1994]" italics="true" pageId="8" pageNumber="338">H. pycnolepis</emphasis>
</taxonomicName>
, SWFC 0612085, 99.6 mm SL.
</paragraph>
</caption>
<paragraph id="8BC4F6143912FFE1FF39FF3D3273FE72" blockId="9.[151,1437,153,683]" pageId="9" pageNumber="339">Dorsal fin with 3 simple and 8–9 branched rays; distal margin convex; origin nearer to snout tip than to caudalfin base. Pectoral fin with 1 simple and 9–10 branched rays, inserted slightly posterior to vertical through posteriormost point of operculum, tip of longest fin ray not extending beyond halfway to insertion of pelvic fin. Pelvic fin with 1 simple and 6–7 branched rays, inserted below first or second branched rays of dorsal fin; tip of longest fin ray not extending beyond half of distance between pelvic-fin insertion and anal-fin origin. Pelvic axillary lobe with pointed tip. Anal fin with 3 simple and 5 branched rays, with convex distal edge; origin closer to pelvic-fin insertion than to caudal-fin base. Posterior margin of caudal fin oblique or micro-concave. Caudal peduncle uniformly deep; with adipose crests along dorsal and ventral midlines. Adipose crest along dorsal midline of caudal peduncle not extending through posterior end of anal-fin base. Anus located posterior approximately 1/3 the distance between pelvic-fin insertion and anal-fin origin, nearer to caudal-fin base.</paragraph>
<paragraph id="8BC4F6143912FFE1FF39FDA43315FD2E" blockId="9.[151,1437,153,683]" pageId="9" pageNumber="339">
<emphasis id="B90F2A063912FFE1FF39FDA4304FFD9E" bold="true" box="[199,338,512,539]" pageId="9" pageNumber="339">Coloration:</emphasis>
Flank of fresh specimen dark yellow, abdomen pale yellow. After fixation in 95% alcohol, flank is brown, abdomen grayish yellow. Flank with 27 to 34 brown vertical bars; width of each bar approximately equal to space between bars. Bars in front of dorsal-fin base narrow and dense, gradually become wider towards caudal-fin base. Dark brown vertical bar on caudal-fin base. Dorsal surface of pectoral, pelvic, and anal fins reddish orange; dorsal and caudal fins yellowish.
</paragraph>
</subSubSection>
<caption id="DF04A69C3912FFE1FF69F86F3462F861" ID-DOI="http://doi.org/10.5281/zenodo.3577077" ID-Zenodo-Dep="3577077" box="[151,1407,1995,2020]" httpUri="https://zenodo.org/record/3577077/files/figure.png" pageId="9" pageNumber="339" startId="9.[151,250,1995,2020]" targetBox="[159,1425,702,1967]" targetPageId="9">
<paragraph id="8BC4F6143912FFE1FF69F86F3462F861" blockId="9.[151,1407,1995,2020]" box="[151,1407,1995,2020]" pageId="9" pageNumber="339">
<emphasis id="B90F2A063912FFE1FF69F86F300EF861" bold="true" box="[151,275,1995,2020]" pageId="9" pageNumber="339">FIGURE 8.</emphasis>
Distribution map of the densely-scaled group of 
<taxonomicName id="4C7B8D973912FFE1FCFBF86F3277F866" authorityName="Nichols" authorityYear="1925" box="[773,874,1995,2019]" class="Actinopterygii" family="Nemacheilidae" genus="Homatula" kingdom="Animalia" order="Cypriniformes" pageId="9" pageNumber="339" phylum="Chordata" rank="genus">
<emphasis id="B90F2A063912FFE1FCFBF86F3277F866" box="[773,874,1995,2019]" italics="true" pageId="9" pageNumber="339">Homatula</emphasis>
</taxonomicName>
. A symbol may represent several adjacent localities.
</paragraph>
</caption>
<subSubSection id="C361A59F3911FFE2FF39FF3C3207FEC7" pageId="10" pageNumber="340" type="distribution">
<paragraph id="8BC4F6143911FFE2FF39FF3C31EFFF52" blockId="10.[151,1437,152,431]" pageId="10" pageNumber="340">
<emphasis id="B90F2A063911FFE2FF39FF3C307EFF36" bold="true" box="[199,355,152,179]" pageId="10" pageNumber="340">Distribution:</emphasis>
Known only from a stream connected to the trunk of the Nu-jiang (the upper Salween River) (
<figureCitation id="1340EA913911FFE2FF5EFF1831F9FF52" box="[160,228,188,215]" captionStart="FIGURE 8" captionStartId="9.[151,250,1995,2020]" captionTargetBox="[159,1425,702,1967]" captionTargetId="figure@9.[151,1436,695,1973]" captionTargetPageId="9" captionText="FIGURE 8. Distribution map of the densely-scaled group of Homatula. A symbol may represent several adjacent localities." figureDoi="http://doi.org/10.5281/zenodo.3577077" httpUri="https://zenodo.org/record/3577077/files/figure.png" pageId="10" pageNumber="340">Fig. 8</figureCitation>
).
</paragraph>
<paragraph id="8BC4F6143911FFE2FF39FF443207FEC7" blockId="10.[151,1437,152,431]" pageId="10" pageNumber="340">
<emphasis id="B90F2A063911FFE2FF39FF4430D0FF7E" bold="true" box="[199,461,224,251]" pageId="10" pageNumber="340">Habitat and Ecology:</emphasis>
<taxonomicName id="4C7B8D973911FFE2FE26FF4433F2FF7F" authorityName="Li &amp; Che &amp; Zhou" authorityYear="2019" box="[472,751,224,250]" class="Actinopterygii" family="Nemacheilidae" genus="Homatula" kingdom="Animalia" order="Cypriniformes" pageId="10" pageNumber="340" phylum="Chordata" rank="species" species="anteridorsalis">
<emphasis id="B90F2A063911FFE2FE26FF4433F2FF7F" box="[472,751,224,250]" italics="true" pageId="10" pageNumber="340">Homatula anteridorsalis</emphasis>
</taxonomicName>
is omnivorous, mainly feeding on algae attached to rocks, organic residues, and small aquatic insects. It inhabits flowing and clear stream environments and dwells on the bottom, shuttling back and forth among rocks and gravel.
</paragraph>
</subSubSection>
<subSubSection id="C361A59F3911FFE2FF39FEE930D8FE2A" pageId="10" pageNumber="340" type="etymology">
<paragraph id="8BC4F6143911FFE2FF39FEE930D8FE2A" blockId="10.[151,1437,152,431]" pageId="10" pageNumber="340">
<emphasis id="B90F2A063911FFE2FF39FEE9304FFEE2" bold="true" box="[199,338,333,359]" pageId="10" pageNumber="340">Etymology:</emphasis>
The specific epithet, 
<emphasis id="B90F2A063911FFE2FDB6FEE833F1FEE3" box="[584,748,332,358]" italics="true" pageId="10" pageNumber="340">
<taxonomicName id="4C7B8D973911FFE2FDB6FEE833F5FEE3" authorityName="Li &amp; Che &amp; Zhou" authorityYear="2019" box="[584,744,332,358]" class="Actinopterygii" family="Nemacheilidae" genus="Homatula" kingdom="Animalia" order="Cypriniformes" pageId="10" pageNumber="340" phylum="Chordata" rank="species" species="anteridorsalis">anteridorsalis</taxonomicName>
,
</emphasis>
an adjective, is a combination of the Latin 
<emphasis id="B90F2A063911FFE2FB22FEE93402FEE3" box="[1244,1311,333,358]" italics="true" pageId="10" pageNumber="340">anteri</emphasis>
– (in front) and – 
<emphasis id="B90F2A063911FFE2FF28FED43003FE0F" box="[214,286,368,394]" italics="true" pageId="10" pageNumber="340">dorsal</emphasis>
(in reference to the dorsal fin), alluding to the more anterior origin of dorsal- and pelvic-fins than in other species of 
<taxonomicName id="4C7B8D973911FFE2FEB0FE3030A3FE2B" authorityName="Nichols" authorityYear="1925" box="[334,446,404,430]" class="Actinopterygii" family="Nemacheilidae" genus="Homatula" kingdom="Animalia" order="Cypriniformes" pageId="10" pageNumber="340" phylum="Chordata" rank="genus">
<emphasis id="B90F2A063911FFE2FEB0FE3030A3FE2B" box="[334,446,404,430]" italics="true" pageId="10" pageNumber="340">Homatula</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>