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<document id="7BFD360767E69CB1F414FA81F4B3D43B" ID-DOI="10.11646/zootaxa.5123.1.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="6399827" ID-ZooBank="036F6B4D-CDCC-4CD7-A914-9A1D8C7A097A" IM.bibliography_approvedBy="karina" IM.illustrations_approvedBy="karina" IM.materialsCitations_approvedBy="karina" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="karina" IM.taxonomicNames_approvedBy="valdenar" IM.treatments_approvedBy="karina" checkinTime="1648673534065" checkinUser="plazi" docAuthor="Harzhauser, Mathias, Landau, Bernard &amp; Janssen, Ronald" docDate="2022" docId="039487D1FF9BFFB2FFBAF9416CE6FA95" docLanguage="en" docName="zootaxa.5123.1.1.pdf" docOrigin="Zootaxa 5123 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5123.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.18:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="18" docTitle="Clavatula atatuerki Lamarck 1801" docType="treatment" docVersion="4" lastPageNumber="27" masterDocId="FFADFFA9FF82FFA8FF2DFFAB6872FFC0" masterDocTitle="The Clavatulidae (Gastropoda, Conoidea) of the Miocene Paratethys Sea with considerations on fossil and extant Clavatulidae genera" masterLastPageNumber="172" masterPageNumber="1" pageNumber="26" updateTime="1709220431393" updateUser="valdenar">
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<mods:title id="91C4DA4F0DE6EB37246BE11F7242414F">The Clavatulidae (Gastropoda, Conoidea) of the Miocene Paratethys Sea with considerations on fossil and extant Clavatulidae genera</mods:title>
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<subSubSection id="C327654CFF9BFFB1FFBAF941698AF8C5" box="[151,504,1770,1797]" pageId="25" pageNumber="26" type="nomenclature">
<paragraph id="8B8236C7FF9BFFB1FFBAF941698AF8C5" blockId="25.[151,504,1770,1839]" box="[151,504,1770,1797]" pageId="25" pageNumber="26">
<emphasis id="B949EAD5FF9BFFB1FFBAF941698AF8C5" bold="true" box="[151,504,1770,1797]" pageId="25" pageNumber="26">
<taxonomicName id="4C3D4D44FF9BFFB1FF8CF94169E0F8C5" ID-CoL="7NZRZ" authorityName="Lamarck" authorityYear="1801" box="[161,402,1770,1797]" class="Gastropoda" family="Clavatulidae" genus="Clavatula" kingdom="Animalia" order="Neogastropoda" pageId="25" pageNumber="26" phylum="Mollusca" rank="species" species="atatuerki">
<emphasis id="B949EAD5FF9BFFB1FF8CF9416964F8C5" bold="true" box="[161,278,1770,1797]" italics="true" pageId="25" pageNumber="26">Clavatula</emphasis>
<emphasis id="B949EAD5FF9BFFB1FE08F94169E0F8C5" bold="true" box="[293,402,1770,1797]" italics="true" pageId="25" pageNumber="26">atatuerki</emphasis>
</taxonomicName>
nov. sp.
</emphasis>
</paragraph>
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<subSubSection id="C327654CFF9BFFB1FFBAF8A469E3F8E9" box="[151,401,1806,1839]" pageId="25" pageNumber="26" type="description">
<paragraph id="8B8236C7FF9BFFB1FFBAF8A469E3F8E9" blockId="25.[151,504,1770,1839]" box="[151,401,1806,1839]" pageId="25" pageNumber="26">
<figureCitation id="13062A42FF9BFFB1FFBAF8A46889F8E9" box="[151,251,1807,1833]" captionStart="FIGURE 11" captionStartId="24.[151,250,1801,1825]" captionTargetBox="[164,1423,273,1778]" captionTargetId="figure-56@24.[164,1423,273,1778]" captionTargetPageId="24" captionText="FIGURE 11 A1A3. Clavatula sorini nov. sp., holotype, NHMW 1865/0001/0224, Lăpugiu de Sus (Romania). B1B3. Clavatula sorini nov. sp., paratype, NHMW 1874/0025/0046, Lăpugiu de Sus (Romania).C1C3. Clavatula camillae (Hoernes &amp;Auinger, 1891), lectotype, NHMW 1949/0005/0042, Lăpugiu de Sus (Romania).D1D3. Clavatula camillae (Hoernes &amp;Auinger, 1891), paralectotype, NHMW 1949/0005/0042b, Lăpugiu de Sus (Romania), E1E2. Clavatula atatuerki nov. sp., holotype,. NHMW 1847/0058/1216, Seyithasan (Turkey). F1F2. Clavatula atatuerki nov. sp., paratype, NHMW 1847/0058/1217, Seyithasan (Turkey). G1G2. Clavatula santhai Kovács &amp; Vicián, 2021, holotype, HNHM, PAL 2021.20.1, Letkés (Hungary)." figureDoi="http://doi.org/10.5281/zenodo.6399855" httpUri="https://zenodo.org/record/6399855/files/figure.png" pageId="25" pageNumber="26">Figs 11E</figureCitation>
<figureCitation id="13062A42FF9BFFB1FFD7F88A6950F8E9" box="[250,290,1807,1839]" captionStart="FIGURE 1" captionStartId="6.[151,250,1903,1927]" captionTargetBox="[171,1417,176,1880]" captionTargetId="figure-26@6.[171,1417,176,1880]" captionTargetPageId="6" captionText="FIGURE 1. A. Map of central and southeastern Europe, representing the area that was partly covered by the Central Paratethys Sea. The white insert corresponds to the area that is restored in the palaeogeographic map below. B. Paleogeography of the Middle Miocene Paratethys Sea showing the most important sedimentary basins (NAFB: North Alpine Foreland Basin, E.-Sopr B: EisenstadtSopron Basin)." figureDoi="http://doi.org/10.5281/zenodo.6399829" httpUri="https://zenodo.org/record/6399829/files/figure.png" pageId="25" pageNumber="26">
<subScript id="17B93482FF9BFFB1FFD7F88A6970F8EF" attach="left" box="[250,258,1825,1839]" fontSize="6" pageId="25" pageNumber="26">1</subScript>
E
</figureCitation>
<figureCitation id="13062A42FF9BFFB1FE0FF88A6958F8EF" box="[290,298,1825,1839]" captionStart="FIGURE 2" captionStartId="7.[151,250,1264,1288]" captionTargetBox="[151,1436,597,1241]" captionTargetId="figure-187@7.[151,1436,597,1241]" captionTargetPageId="7" captionText="FIGURE 2. Descriptive terms used in descriptions and measurements to shell morphology, whorl profiles and anal sinus. SL: shell length, MD: maximum diameter, AA: apical angle, LWH: last whorl height, AH: aperture height.AL: aperture length.AW: aperture width." figureDoi="http://doi.org/10.5281/zenodo.6399831" httpUri="https://zenodo.org/record/6399831/files/figure.png" pageId="25" pageNumber="26">
<subScript id="17B93482FF9BFFB1FE0FF88A6958F8EF" attach="left" box="[290,298,1825,1839]" fontSize="6" pageId="25" pageNumber="26">2</subScript>
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, F
<figureCitation id="13062A42FF9BFFB1FE6AF88A691FF8E9" box="[327,365,1807,1839]" captionStart="FIGURE 1" captionStartId="6.[151,250,1903,1927]" captionTargetBox="[171,1417,176,1880]" captionTargetId="figure-26@6.[171,1417,176,1880]" captionTargetPageId="6" captionText="FIGURE 1. A. Map of central and southeastern Europe, representing the area that was partly covered by the Central Paratethys Sea. The white insert corresponds to the area that is restored in the palaeogeographic map below. B. Paleogeography of the Middle Miocene Paratethys Sea showing the most important sedimentary basins (NAFB: North Alpine Foreland Basin, E.-Sopr B: EisenstadtSopron Basin)." figureDoi="http://doi.org/10.5281/zenodo.6399829" httpUri="https://zenodo.org/record/6399829/files/figure.png" pageId="25" pageNumber="26">
<subScript id="17B93482FF9BFFB1FE6AF88A693DF8EF" attach="left" box="[327,335,1825,1839]" fontSize="6" pageId="25" pageNumber="26">1</subScript>
F
</figureCitation>
<figureCitation id="13062A42FF9BFFB1FE40F88A6907F8EF" box="[365,373,1825,1839]" captionStart="FIGURE 2" captionStartId="7.[151,250,1264,1288]" captionTargetBox="[151,1436,597,1241]" captionTargetId="figure-187@7.[151,1436,597,1241]" captionTargetPageId="7" captionText="FIGURE 2. Descriptive terms used in descriptions and measurements to shell morphology, whorl profiles and anal sinus. SL: shell length, MD: maximum diameter, AA: apical angle, LWH: last whorl height, AH: aperture height.AL: aperture length.AW: aperture width." figureDoi="http://doi.org/10.5281/zenodo.6399831" httpUri="https://zenodo.org/record/6399831/files/figure.png" pageId="25" pageNumber="26">
<subScript id="17B93482FF9BFFB1FE40F88A6907F8EF" attach="left" box="[365,373,1825,1839]" fontSize="6" pageId="25" pageNumber="26">2</subScript>
</figureCitation>
,
<figureCitation id="13062A42FF9BFFB1FEAEF8A569E3F8E9" box="[387,401,1806,1833]" captionStart="FIGURE 5" captionStartId="15.[151,250,1999,2023]" captionTargetBox="[151,1436,631,1976]" captionTargetId="figure-214@15.[151,1436,631,1976]" captionTargetPageId="15" captionText="FIGURE 5. Anal sinus morphology of Paratethyan Clavatulidae." figureDoi="http://doi.org/10.5281/zenodo.6399839" httpUri="https://zenodo.org/record/6399839/files/figure.png" pageId="25" pageNumber="26">5</figureCitation>
</paragraph>
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<paragraph id="8B8236C7FF9BFFB1FFBAF8F86D42F849" blockId="25.[151,1437,1875,1930]" pageId="25" pageNumber="26">
<taxonomicName id="4C3D4D44FF9BFFB1FFBAF8F86A21F8AB" authority="(Hoernes &amp; Auinger, 1891)" baseAuthorityName="R. Hoernes &amp; Auinger" baseAuthorityYear="1891" box="[151,595,1875,1899]" class="Gastropoda" family="Clavatulidae" genus="Clavatula" kingdom="Animalia" order="Neogastropoda" pageId="25" pageNumber="26" phylum="Mollusca" rank="species" species="olgae">
<emphasis id="B949EAD5FF9BFFB1FFBAF8F86948F8AB" box="[151,314,1875,1899]" italics="true" pageId="25" pageNumber="26">Clavatula olgae</emphasis>
(
<bibRefCitation id="EFAC4B36FF9BFFB1FE6AF8F86A3EF8AB" author="Hoernes, R. &amp; Auinger, M." box="[327,588,1875,1899]" pageId="25" pageNumber="26" pagination="283 - 382" refId="ref135318" refString="Hoernes, R. &amp; Auinger, M. (1891) Die Gasteropoden der Meeres-Ablagerungen der ersten und zweiten Miocanen Mediterran- Stufe in der Osterreichisch-Ungarischen Monarchie. Abhandlungen der kaiserlich-koniglichen Geologischen Reichsanstalt, 12, 283 - 382, pls 37 - 50. [https: // opac. geologie. ac. at / ais 312 / dokumente / Hoernes % 20 und % 20 Auinger % 20 _ 1879 _ Gasteropoden. pdf]" type="journal article" year="1891">Hoernes &amp; Auinger, 1891</bibRefCitation>
)
</taxonomicName>
<bibRefCitation id="EFAC4B36FF9BFFB1FD40F8F86B5DF8AB" author="Landau, B. M. &amp; Harzhauser, M. &amp; Yslamodlu, Y. &amp; Marques da Silva, C." box="[621,815,1875,1899]" pageId="25" pageNumber="26" pagination="3 - 576" refId="ref137659" refString="Landau, B. M., Harzhauser, M., Yslamodlu, Y. &amp; Marques da Silva, C. (2013) Systematics and palaeobiogeography of the gastropods of the middle Miocene (Serravallian) Karaman Basin of Turkey. Cainozoic Research, 11 - 13, 3 - 576" type="journal article" year="2013">
Landau
<emphasis id="B949EAD5FF9BFFB1FDEEF8FF6A84F8AB" box="[707,758,1875,1899]" italics="true" pageId="25" pageNumber="26">et al.</emphasis>
2013
</bibRefCitation>
(
<emphasis id="B949EAD5FF9BFFB1FC13F8FF6BF0F8AB" box="[830,898,1876,1899]" italics="true" pageId="25" pageNumber="26">partim</emphasis>
): 286, pl. 48, figs 24, pl. 50, fig. 8 [
<emphasis id="B949EAD5FF9BFFB1FBD0F8FF6D51F8AB" box="[1277,1315,1876,1899]" italics="true" pageId="25" pageNumber="26">non</emphasis>
<taxonomicName id="4C3D4D44FF9BFFB1FA1FF8F86A6AF84A" authority="(Hoernes &amp; Auinger 1891)" authorityName="Lamarck" authorityYear="1801" baseAuthorityName="R. Hoernes &amp; Auinger" baseAuthorityYear="1891" class="Gastropoda" family="Clavatulidae" genus="Clavatula" kingdom="Animalia" order="Neogastropoda" pageId="25" pageNumber="26" phylum="Mollusca" rank="genus">
<emphasis id="B949EAD5FF9BFFB1FA1FF8F86DE4F8AB" box="[1330,1430,1875,1899]" italics="true" pageId="25" pageNumber="26">Clavatula</emphasis>
<emphasis id="B949EAD5FF9BFFB1FFEAF8D9688DF84A" box="[199,255,1906,1930]" italics="true" pageId="25" pageNumber="26">olgae</emphasis>
(
<bibRefCitation id="EFAC4B36FF9BFFB1FE20F8D96A63F84A" author="Hoernes, R. &amp; Auinger, M." box="[269,529,1906,1930]" pageId="25" pageNumber="26" pagination="283 - 382" refId="ref135318" refString="Hoernes, R. &amp; Auinger, M. (1891) Die Gasteropoden der Meeres-Ablagerungen der ersten und zweiten Miocanen Mediterran- Stufe in der Osterreichisch-Ungarischen Monarchie. Abhandlungen der kaiserlich-koniglichen Geologischen Reichsanstalt, 12, 283 - 382, pls 37 - 50. [https: // opac. geologie. ac. at / ais 312 / dokumente / Hoernes % 20 und % 20 Auinger % 20 _ 1879 _ Gasteropoden. pdf]" type="journal article" year="1891">Hoernes &amp; Auinger 1891</bibRefCitation>
)
</taxonomicName>
] [
<emphasis id="B949EAD5FF9BFFB1FD00F8D86A21F84A" box="[557,595,1907,1930]" italics="true" pageId="25" pageNumber="26">non</emphasis>
<bibRefCitation id="EFAC4B36FF9BFFB1FD77F8D96B68F84A" author="Landau, B. M. &amp; Harzhauser, M. &amp; Yslamodlu, Y. &amp; Marques da Silva, C." box="[602,794,1906,1930]" pageId="25" pageNumber="26" pagination="3 - 576" refId="ref137659" refString="Landau, B. M., Harzhauser, M., Yslamodlu, Y. &amp; Marques da Silva, C. (2013) Systematics and palaeobiogeography of the gastropods of the middle Miocene (Serravallian) Karaman Basin of Turkey. Cainozoic Research, 11 - 13, 3 - 576" type="journal article" year="2013">
Landau
<emphasis id="B949EAD5FF9BFFB1FD81F8D86AADF84A" box="[684,735,1906,1930]" italics="true" pageId="25" pageNumber="26">et al.</emphasis>
2013
</bibRefCitation>
, pl. 48, figs 56 =
<taxonomicName id="4C3D4D44FF9BFFB1FCCDF8D96D5EF849" authorityName="Lamarck" authorityYear="1801" box="[992,1324,1906,1930]" class="Gastropoda" family="Clavatulidae" genus="Clavatula" kingdom="Animalia" order="Neogastropoda" pageId="25" pageNumber="26" phylum="Mollusca" rank="genus">
<emphasis id="B949EAD5FF9BFFB1FCCDF8D96C36F84A" box="[992,1092,1906,1930]" italics="true" pageId="25" pageNumber="26">Clavatula</emphasis>
<emphasis id="B949EAD5FF9BFFB1FB7CF8D96CBFF849" box="[1105,1229,1906,1929]" italics="true" pageId="25" pageNumber="26">ariejansseni</emphasis>
nov. sp.]
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C327654CFF9BFFB2FFBAF8136AF8FE81" lastPageId="26" lastPageNumber="27" pageId="25" pageNumber="26" type="materials_examined">
<paragraph id="8B8236C7FF9BFFB2FFBAF8136CBEFF15" blockId="25.[151,1436,1976,2003]" lastBlockId="26.[151,1437,150,1366]" lastPageId="26" lastPageNumber="27" pageId="25" pageNumber="26">
<emphasis id="B949EAD5FF9BFFB1FFBAF8136937F812" bold="true" box="[151,325,1976,2002]" pageId="25" pageNumber="26">Type material.</emphasis>
<materialsCitation id="3B553C9AFF9BFFB2FE61F8136B7BFF77" collectionCode="MD" country="Turkey" lastPageId="26" lastPageNumber="27" location="Seyithasan" pageId="25" pageNumber="26" specimenCode="NHMW 1847" specimenCount="1" typeStatus="holotype">
<typeStatus id="54868865FF9BFFB1FE61F81369CBF812" box="[332,441,1976,2002]" pageId="25" pageNumber="26" type="holotype">Holotype</typeStatus>
:
<specimenCode id="DB9B9EBCFF9BFFB1FEEEF8126A2FF813" box="[451,605,1976,2003]" collectionCode="NHMW" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="25" pageNumber="26" type="Museum">NHMW 1847</specimenCode>
/0058/1216,
<location id="8EE2601CFF9BFFB1FDC1F8136B17F812" LSID="urn:lsid:plazi:treatment:039487D1FF9BFFB2FFBAF9416CE6FA95:8EE2601CFF9BFFB1FDC1F8136B17F812" box="[748,869,1976,2002]" country="Turkey" name="Seyithasan" pageId="25" pageNumber="26">Seyithasan</location>
(
<collectingCountry id="F32A7657FF9BFFB1FC59F8136BB8F813" box="[884,970,1976,2003]" name="Turkey" pageId="25" pageNumber="26">Turkey</collectingCountry>
), SL:
<quantity id="4CC59B22FF9BFFB1FB21F8136C00F813" box="[1036,1138,1976,2003]" metricMagnitude="-2" metricUnit="m" metricValue="4.14" pageId="25" pageNumber="26" unit="mm" value="41.4">41.4 mm</quantity>
,
<collectionCode id="ED2CAE02FF9BFFB1FB53F8126CDDF813" box="[1150,1199,1977,2003]" country="Germany" name="Museum Donaueschingen" pageId="25" pageNumber="26">MD</collectionCode>
:
<quantity id="4CC59B22FF9BFFB1FB97F8136D52F813" box="[1210,1312,1976,2003]" metricMagnitude="-2" metricUnit="m" metricValue="1.48" pageId="25" pageNumber="26" unit="mm" value="14.8">14.8 mm</quantity>
, illustrated in
<bibRefCitation id="EFAC4B36FF98FFB2FF99FF3C69E6FF71" author="Landau, B. M. &amp; Harzhauser, M. &amp; Yslamodlu, Y. &amp; Marques da Silva, C." box="[180,404,151,177]" pageId="26" pageNumber="27" pagination="3 - 576" refId="ref137659" refString="Landau, B. M., Harzhauser, M., Yslamodlu, Y. &amp; Marques da Silva, C. (2013) Systematics and palaeobiogeography of the gastropods of the middle Miocene (Serravallian) Karaman Basin of Turkey. Cainozoic Research, 11 - 13, 3 - 576" type="journal article" year="2013">
Landau
<emphasis id="B949EAD5FF98FFB2FE3DFF3C693BFF71" box="[272,329,151,177]" italics="true" pageId="26" pageNumber="27">et al.</emphasis>
(2013
</bibRefCitation>
, pl. 48, figs 3a3b), figs 11E
<subScript id="17B93482FF98FFB2FDF7FF026A90FF77" attach="left" box="[730,738,169,183]" fontSize="6" pageId="26" pageNumber="27">1</subScript>
E
<subScript id="17B93482FF98FFB2FC2CFF026B7BFF77" attach="left" box="[769,777,169,183]" fontSize="6" pageId="26" pageNumber="27">2</subScript>
</materialsCitation>
.
<materialsCitation id="3B553C9AFF98FFB2FC35FF3C695FFF15" collectionCode="NHMW" country="Turkey" location="Seyithasan" pageId="26" pageNumber="27" specimenCode="NHMW 1847" specimenCount="1" typeStatus="paratype">
<typeStatus id="54868865FF98FFB2FC35FF3C6B0CFF71" box="[792,894,151,177]" pageId="26" pageNumber="27" type="paratype">Paratype</typeStatus>
:
<specimenCode id="DB9B9EBCFF98FFB2FCA4FF3C6C51FF71" box="[905,1059,150,177]" collectionCode="NHMW" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="26" pageNumber="27" type="Museum">NHMW 1847</specimenCode>
/0058/1217,
<location id="8EE2601CFF98FFB2FB9FFF3C6D59FF71" LSID="urn:lsid:plazi:treatment:039487D1FF9BFFB2FFBAF9416CE6FA95:8EE2601CFF98FFB2FB9FFF3C6D59FF71" box="[1202,1323,151,177]" country="Turkey" name="Seyithasan" pageId="26" pageNumber="27">Seyithasan</location>
(
<collectingCountry id="F32A7657FF98FFB2FA17FF3C6DE2FF72" box="[1338,1424,151,178]" name="Turkey" pageId="26" pageNumber="27">Turkey</collectingCountry>
), SL:
<quantity id="4CC59B22FF98FFB2FFEAFF10695FFF15" box="[199,301,187,213]" metricMagnitude="-2" metricUnit="m" metricValue="3.7399999999999998" pageId="26" pageNumber="27" unit="mm" value="37.4">37.4 mm</quantity>
</materialsCitation>
,
<collectionCode id="ED2CAE02FF98FFB2FE15FF10691BFF15" box="[312,361,187,213]" country="Germany" name="Museum Donaueschingen" pageId="26" pageNumber="27">MD</collectionCode>
:
<quantity id="4CC59B22FF98FFB2FE59FF1169A8FF15" box="[372,474,186,213]" metricMagnitude="-2" metricUnit="m" metricValue="1.35" pageId="26" pageNumber="27" unit="mm" value="13.5">13.5 mm</quantity>
, illustrated in
<bibRefCitation id="EFAC4B36FF98FFB2FD54FF106B24FF15" author="Landau, B. M. &amp; Harzhauser, M. &amp; Yslamodlu, Y. &amp; Marques da Silva, C." box="[633,854,187,213]" pageId="26" pageNumber="27" pagination="3 - 576" refId="ref137659" refString="Landau, B. M., Harzhauser, M., Yslamodlu, Y. &amp; Marques da Silva, C. (2013) Systematics and palaeobiogeography of the gastropods of the middle Miocene (Serravallian) Karaman Basin of Turkey. Cainozoic Research, 11 - 13, 3 - 576" type="journal article" year="2013">Landau et al. (2013</bibRefCitation>
, pl. 48, figs 4a4b), figs 11F
<subScript id="17B93482FF98FFB2FBBAFF666CEDFF1B" attach="left" box="[1175,1183,205,219]" fontSize="6" pageId="26" pageNumber="27">1</subScript>
F
<subScript id="17B93482FF98FFB2FB90FF666CB7FF1B" attach="left" box="[1213,1221,205,219]" fontSize="6" pageId="26" pageNumber="27">2</subScript>
.
</paragraph>
<paragraph id="8B8236C7FF98FFB2FFEAFF746B60FF39" blockId="26.[151,1437,150,1366]" box="[199,786,222,250]" pageId="26" pageNumber="27">
<materialsCitation id="3B553C9AFF98FFB2FFEAFF746B7CFF39" box="[199,782,222,250]" country="Turkey" location="Basin" municipality="Seyithasan" pageId="26" pageNumber="27" specimenCount="1" stateProvince="Karaman" typeStatus="holotype">
<emphasis id="B949EAD5FF98FFB2FFEAFF746914FF38" bold="true" box="[199,358,222,249]" pageId="26" pageNumber="27">
<typeStatus id="54868865FF98FFB2FFEAFF746970FF39" box="[199,258,223,249]" pageId="26" pageNumber="27">Type</typeStatus>
locality.
</emphasis>
<collectingMunicipality id="6BE6ACBDFF98FFB2FE41FF746997FF39" box="[364,485,223,249]" pageId="26" pageNumber="27">Seyithasan</collectingMunicipality>
(
<collectingCountry id="F32A7657FF98FFB2FED9FF746A38FF3A" box="[500,586,223,250]" name="Turkey" pageId="26" pageNumber="27">Turkey</collectingCountry>
),
<collectingRegion id="49F9F825FF98FFB2FD70FF746AB6FF39" box="[605,708,223,249]" country="Turkey" name="Karaman" pageId="26" pageNumber="27">Karaman</collectingRegion>
<location id="8EE2601CFF98FFB2FDE6FF746B7CFF39" LSID="urn:lsid:plazi:treatment:039487D1FF9BFFB2FFBAF9416CE6FA95:8EE2601CFF98FFB2FDE6FF746B7CFF39" box="[715,782,223,249]" country="Turkey" municipality="Seyithasan" name="Basin" pageId="26" pageNumber="27" stateProvince="Karaman">Basin</location>
</materialsCitation>
.
</paragraph>
<paragraph id="8B8236C7FF98FFB2FFEAFEA86B6BFEDD" blockId="26.[151,1437,150,1366]" box="[199,793,259,285]" pageId="26" pageNumber="27">
<emphasis id="B949EAD5FF98FFB2FFEAFEA8691DFEDD" bold="true" box="[199,367,259,285]" pageId="26" pageNumber="27">
<typeStatus id="54868865FF98FFB2FFEAFEA86970FEDD" box="[199,258,259,285]" pageId="26" pageNumber="27">Type</typeStatus>
stratum.
</emphasis>
Silt and clay of the Týrtar Formation.
</paragraph>
<paragraph id="8B8236C7FF98FFB2FFEAFE8D6AF8FE81" blockId="26.[151,1437,150,1366]" box="[199,650,294,321]" pageId="26" pageNumber="27">
<emphasis id="B949EAD5FF98FFB2FFEAFE8D688FFE80" bold="true" box="[199,253,294,320]" pageId="26" pageNumber="27">Age.</emphasis>
Middle Miocene, late Serravallian.
</paragraph>
</subSubSection>
<subSubSection id="C327654CFF98FFB2FFEAFEE06C37FEA5" box="[199,1093,331,357]" pageId="26" pageNumber="27" type="etymology">
<paragraph id="8B8236C7FF98FFB2FFEAFEE06C37FEA5" blockId="26.[151,1437,150,1366]" box="[199,1093,331,357]" pageId="26" pageNumber="27">
<emphasis id="B949EAD5FF98FFB2FFEAFEE0693CFEA5" bold="true" box="[199,334,331,357]" pageId="26" pageNumber="27">Etymology.</emphasis>
Referring to Kemal Atatürk (18711938), first president of
<collectingCountry id="F32A7657FF98FFB2FCC2FEE06C33FEA5" box="[1007,1089,331,357]" name="Turkey" pageId="26" pageNumber="27">Turkey</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C327654CFF98FFB2FFEAFEC568A2FE11" pageId="26" pageNumber="27" type="diagnosis">
<paragraph id="8B8236C7FF98FFB2FFEAFEC568A2FE11" blockId="26.[151,1437,150,1366]" pageId="26" pageNumber="27">
<emphasis id="B949EAD5FF98FFB2FFEAFEC56930FE48" bold="true" box="[199,322,366,392]" pageId="26" pageNumber="27">Diagnosis.</emphasis>
Medium-sized, slender to moderately slender fusiform shell with elevated tuberculate adsutural cords separated by smooth concave mid-portion. Base with weak perifasciolar cords. Siphonal canal moderately long.
</paragraph>
</subSubSection>
<subSubSection id="C327654CFF98FFB2FFEAFE716D41FC9D" pageId="26" pageNumber="27" type="description">
<paragraph id="8B8236C7FF98FFB2FFEAFE716D41FC9D" blockId="26.[151,1437,150,1366]" pageId="26" pageNumber="27">
<emphasis id="B949EAD5FF98FFB2FFEAFE71692BFE34" bold="true" box="[199,345,474,500]" pageId="26" pageNumber="27">Description.</emphasis>
Shell medium-sized, solid, slender to moderately slender fusiform; apical angle 37°. Protoconch not preserved. Early teleoconch spire whorls slightly concave, with bipartite sculpture of weak subsutural cord and beaded suprasutural cord, midsection deeply concave. On later whorls subsutural collar somewhat elevated, weakly delimited, bearing short, widely-spaced spines and one or two weak spiral threads. Mid-whorl portion smooth, broadening abapically, suprasutural cord bearing close-set, axially elongated tubercles. Last whorl 63 66% total height, subsutural collar as described above, subsutural ramp broad, strongly concave, delimited by roundly angled shoulder bearing small, close-set, comma-shaped tubercles. Convex below, moderately constricted at base. Sculpture below shoulder of two to three primary cords of irregular strength, uppermost cord nodular, very irregular weak axial sculpture extends abapically from nodules beading spiral cords. Aperture small, ovate. Anal sinus moderately narrow, very deep, asymmetrically U-shaped. Weak inner lirae deep inside aperture. Columella moderately excavated in adapical third, weakly twisted. Siphonal canal moderately long, slightly twisted.
</paragraph>
</subSubSection>
<subSubSection id="C327654CFF98FFB2FFEAFCCD6A77FB29" pageId="26" pageNumber="27" type="discussion">
<paragraph id="8B8236C7FF98FFB2FFEAFCCD68A9FB99" blockId="26.[151,1437,150,1366]" pageId="26" pageNumber="27">
<emphasis id="B949EAD5FF98FFB2FFEAFCCD693EFC40" bold="true" box="[199,332,870,896]" pageId="26" pageNumber="27">Discussion.</emphasis>
This species was identified by
<bibRefCitation id="EFAC4B36FF98FFB2FD8AFCCC6BFBFC41" author="Landau, B. M. &amp; Harzhauser, M. &amp; Yslamodlu, Y. &amp; Marques da Silva, C." box="[679,905,871,897]" pageId="26" pageNumber="27" pagination="3 - 576" refId="ref137659" refString="Landau, B. M., Harzhauser, M., Yslamodlu, Y. &amp; Marques da Silva, C. (2013) Systematics and palaeobiogeography of the gastropods of the middle Miocene (Serravallian) Karaman Basin of Turkey. Cainozoic Research, 11 - 13, 3 - 576" type="journal article" year="2013">
Landau
<emphasis id="B949EAD5FF98FFB2FC2CFCCC6B4AFC41" box="[769,824,871,897]" italics="true" pageId="26" pageNumber="27">et al.</emphasis>
(2013)
</bibRefCitation>
as
<taxonomicName id="4C3D4D44FF98FFB2FC81FCCC6DE5FC41" authority="(Hoernes &amp; Auinger, 1891)" baseAuthorityName="R. Hoernes &amp; Auinger" baseAuthorityYear="1891" box="[940,1431,871,897]" class="Gastropoda" family="Clavatulidae" genus="Clavatula" kingdom="Animalia" order="Neogastropoda" pageId="26" pageNumber="27" phylum="Mollusca" rank="species" species="olgae">
<emphasis id="B949EAD5FF98FFB2FC81FCCC6C2DFC41" box="[940,1119,871,897]" italics="true" pageId="26" pageNumber="27">Clavatula olgae</emphasis>
(
<bibRefCitation id="EFAC4B36FF98FFB2FB40FCCC6DFDFC41" author="Hoernes, R. &amp; Auinger, M." box="[1133,1423,871,897]" pageId="26" pageNumber="27" pagination="283 - 382" refId="ref135318" refString="Hoernes, R. &amp; Auinger, M. (1891) Die Gasteropoden der Meeres-Ablagerungen der ersten und zweiten Miocanen Mediterran- Stufe in der Osterreichisch-Ungarischen Monarchie. Abhandlungen der kaiserlich-koniglichen Geologischen Reichsanstalt, 12, 283 - 382, pls 37 - 50. [https: // opac. geologie. ac. at / ais 312 / dokumente / Hoernes % 20 und % 20 Auinger % 20 _ 1879 _ Gasteropoden. pdf]" type="journal article" year="1891">Hoernes &amp; Auinger, 1891</bibRefCitation>
)
</taxonomicName>
, but
<taxonomicName id="4C3D4D44FF98FFB2FFE6FC20694BFC65" authorityName="Lamarck" authorityYear="1801" box="[203,313,907,933]" class="Gastropoda" family="Clavatulidae" genus="Clavatula" kingdom="Animalia" order="Neogastropoda" pageId="26" pageNumber="27" phylum="Mollusca" rank="genus">
<emphasis id="B949EAD5FF98FFB2FFE6FC20694BFC65" box="[203,313,907,933]" italics="true" pageId="26" pageNumber="27">Clavatula</emphasis>
</taxonomicName>
<taxonomicName id="4C3D4D44FF98FFB2FE6AFC2069F7FC65" baseAuthorityName="R. Hoernes &amp; Auinger" baseAuthorityYear="1891" box="[327,389,907,933]" class="Gastropoda" family="Clavatulidae" genus="Clavatula" kingdom="Animalia" order="Neogastropoda" pageId="26" pageNumber="27" phylum="Mollusca" rank="species" species="olgae">
<emphasis id="B949EAD5FF98FFB2FE6AFC2069F7FC65" box="[327,389,907,933]" italics="true" pageId="26" pageNumber="27">olgae</emphasis>
</taxonomicName>
differs in the presence of spiral sculpture on the teleoconch whorls, the more prominent beads on the adsutural cords and the more prominent inner lirae. The Turkish species seems to be closer to the Paratethyan
<taxonomicName id="4C3D4D44FF98FFB2FF8CFC78697DFC2D" authorityName="Lamarck" authorityYear="1801" box="[161,271,979,1005]" class="Gastropoda" family="Clavatulidae" genus="Clavatula" kingdom="Animalia" order="Neogastropoda" pageId="26" pageNumber="27" phylum="Mollusca" rank="genus">
<emphasis id="B949EAD5FF98FFB2FF8CFC78697DFC2D" box="[161,271,979,1005]" italics="true" pageId="26" pageNumber="27">Clavatula</emphasis>
</taxonomicName>
<taxonomicName id="4C3D4D44FF98FFB2FE32FC786ACFFC2D" authority="(Hoernes &amp; Auinger, 1891)" baseAuthorityName="Hoernes &amp; Auinger" baseAuthorityYear="1891" box="[287,701,979,1005]" class="Gastropoda" family="Turridae" genus="Pleurotoma" kingdom="Animalia" order="Neogastropoda" pageId="26" pageNumber="27" phylum="Mollusca" rank="species" species="camillae">
<emphasis id="B949EAD5FF98FFB2FE32FC78690DFC2D" box="[287,383,979,1005]" italics="true" pageId="26" pageNumber="27">camillae</emphasis>
(
<bibRefCitation id="EFAC4B36FF98FFB2FEA3FC786AC6FC2D" author="Hoernes, R. &amp; Auinger, M." box="[398,692,979,1005]" pageId="26" pageNumber="27" pagination="283 - 382" refId="ref135318" refString="Hoernes, R. &amp; Auinger, M. (1891) Die Gasteropoden der Meeres-Ablagerungen der ersten und zweiten Miocanen Mediterran- Stufe in der Osterreichisch-Ungarischen Monarchie. Abhandlungen der kaiserlich-koniglichen Geologischen Reichsanstalt, 12, 283 - 382, pls 37 - 50. [https: // opac. geologie. ac. at / ais 312 / dokumente / Hoernes % 20 und % 20 Auinger % 20 _ 1879 _ Gasteropoden. pdf]" type="journal article" year="1891">Hoernes &amp; Auinger, 1891</bibRefCitation>
)
</taxonomicName>
. Both species develop a marked and smooth mid-whorl concavity and comparatively small and reduced beads on the adsutural cords. Nevertheless,
<taxonomicName id="4C3D4D44FF98FFB2FB61FC5C6CC8FBD1" authorityName="Lamarck" authorityYear="1801" box="[1100,1210,1015,1041]" class="Gastropoda" family="Clavatulidae" genus="Clavatula" kingdom="Animalia" order="Neogastropoda" pageId="26" pageNumber="27" phylum="Mollusca" rank="genus">
<emphasis id="B949EAD5FF98FFB2FB61FC5C6CC8FBD1" box="[1100,1210,1015,1041]" italics="true" pageId="26" pageNumber="27">Clavatula</emphasis>
</taxonomicName>
<taxonomicName id="4C3D4D44FF98FFB2FBE6FC5C6D59FBD1" baseAuthorityName="Hoernes &amp; Auinger" baseAuthorityYear="1891" box="[1227,1323,1015,1041]" class="Gastropoda" family="Turridae" genus="Pleurotoma" kingdom="Animalia" order="Neogastropoda" pageId="26" pageNumber="27" phylum="Mollusca" rank="species" species="camillae">
<emphasis id="B949EAD5FF98FFB2FBE6FC5C6D59FBD1" box="[1227,1323,1015,1041]" italics="true" pageId="26" pageNumber="27">camillae</emphasis>
</taxonomicName>
is clearly distinguished from
<emphasis id="B949EAD5FF98FFB2FE57FBB069E6FBF5" box="[378,404,1051,1077]" italics="true" pageId="26" pageNumber="27">C.</emphasis>
<emphasis id="B949EAD5FF98FFB2FE89FBB06A7BFBF5" box="[420,521,1051,1077]" italics="true" pageId="26" pageNumber="27">atatuerki</emphasis>
nov. sp. by its shorter siphonal canal and very prominent, tubercular perifasciolar cords.
</paragraph>
<paragraph id="8B8236C7FF98FFB2FFEAFBC86A77FB29" blockId="26.[151,1437,150,1366]" pageId="26" pageNumber="27">
<taxonomicName id="4C3D4D44FF98FFB2FFFCFBC8694DFBBD" authorityName="Lamarck" authorityYear="1801" box="[209,319,1123,1149]" class="Gastropoda" family="Clavatulidae" genus="Clavatula" kingdom="Animalia" order="Neogastropoda" pageId="26" pageNumber="27" phylum="Mollusca" rank="genus">
<emphasis id="B949EAD5FF98FFB2FFFCFBC8694DFBBD" box="[209,319,1123,1149]" italics="true" pageId="26" pageNumber="27">Clavatula</emphasis>
</taxonomicName>
<taxonomicName id="4C3D4D44FF98FFB2FE7DFBC86AC9FBBD" authority="Kovacs &amp; Vician, 2021" authorityName="Kovacs &amp; Vician" authorityYear="2021" box="[336,699,1123,1149]" class="Gastropoda" family="Clavatulidae" genus="Clavatula" kingdom="Animalia" order="Neogastropoda" pageId="26" pageNumber="27" phylum="Mollusca" rank="species" species="santhai">
<emphasis id="B949EAD5FF98FFB2FE7DFBC869D0FBBD" box="[336,418,1123,1149]" italics="true" pageId="26" pageNumber="27">santhai</emphasis>
<bibRefCitation id="EFAC4B36FF98FFB2FE81FBC86AC9FBBD" author="Kovacs, Z. &amp; Vician, Z." box="[428,699,1123,1149]" pageId="26" pageNumber="27" pagination="137 - 158" refId="ref137046" refString="Kovacs, Z. &amp; Vician, Z. (2021) Middle Miocene Conoidea (Neogastropoda) assemblage of Letkes (Hungary), Part II. (Borsoniidae, Cochlespiridae, Clavatulidae, Turridae, Fusiturridae). Foldtani Kozlony, 151, 137 - 158. https: // doi. org / 10.23928 / foldt. kozl. 2021.151.2.137" type="journal article" year="2021">Kovács &amp; Vicián, 2021</bibRefCitation>
</taxonomicName>
differs in it much higher spire and the slowly contracting base. The specimens illustrated by
<bibRefCitation id="EFAC4B36FF98FFB2FECCFB2C6AB3FB61" author="Landau, B. M. &amp; Harzhauser, M. &amp; Yslamodlu, Y. &amp; Marques da Silva, C." box="[481,705,1159,1185]" pageId="26" pageNumber="27" pagination="3 - 576" refId="ref137659" refString="Landau, B. M., Harzhauser, M., Yslamodlu, Y. &amp; Marques da Silva, C. (2013) Systematics and palaeobiogeography of the gastropods of the middle Miocene (Serravallian) Karaman Basin of Turkey. Cainozoic Research, 11 - 13, 3 - 576" type="journal article" year="2013">
Landau
<emphasis id="B949EAD5FF98FFB2FD10FB2C6A07FB61" box="[573,629,1159,1185]" italics="true" pageId="26" pageNumber="27">et al.</emphasis>
(2013
</bibRefCitation>
, pl. 48, figs 56) are excluded from
<taxonomicName id="4C3D4D44FF98FFB2FB48FB2C6CA1FB61" authorityName="Lamarck" authorityYear="1801" box="[1125,1235,1159,1185]" class="Gastropoda" family="Clavatulidae" genus="Clavatula" kingdom="Animalia" order="Neogastropoda" pageId="26" pageNumber="27" phylum="Mollusca" rank="genus">
<emphasis id="B949EAD5FF98FFB2FB48FB2C6CA1FB61" box="[1125,1235,1159,1185]" italics="true" pageId="26" pageNumber="27">Clavatula</emphasis>
</taxonomicName>
<emphasis id="B949EAD5FF98FFB2FBC9FB2C6D3BFB61" box="[1252,1353,1159,1185]" italics="true" pageId="26" pageNumber="27">atatuerki</emphasis>
. Based on its tripartite early whorls it represents a species of the
<taxonomicName id="4C3D4D44FF98FFB2FC1EFB006BD3FB05" authorityName="Lamarck" authorityYear="1801" box="[819,929,1195,1221]" class="Gastropoda" family="Clavatulidae" genus="Clavatula" kingdom="Animalia" order="Neogastropoda" pageId="26" pageNumber="27" phylum="Mollusca" rank="genus">
<emphasis id="B949EAD5FF98FFB2FC1EFB006BD3FB05" box="[819,929,1195,1221]" italics="true" pageId="26" pageNumber="27">Clavatula</emphasis>
</taxonomicName>
<taxonomicName id="4C3D4D44FF98FFB2FC9CFB006C51FB04" baseAuthorityName="Brocchi" baseAuthorityYear="1814" box="[945,1059,1195,1220]" class="Gastropoda" family="Turridae" genus="Pleurotoma" kingdom="Animalia" order="Neogastropoda" pageId="26" pageNumber="27" phylum="Mollusca" rank="species" species="interrupta">
<emphasis id="B949EAD5FF98FFB2FC9CFB006C51FB04" box="[945,1059,1195,1220]" italics="true" pageId="26" pageNumber="27">interrupta</emphasis>
</taxonomicName>
group and is described herein as
<taxonomicName id="4C3D4D44FF98FFB2FF8CFB64697DFB29" authorityName="Lamarck" authorityYear="1801" box="[161,271,1231,1257]" class="Gastropoda" family="Clavatulidae" genus="Clavatula" kingdom="Animalia" order="Neogastropoda" pageId="26" pageNumber="27" phylum="Mollusca" rank="genus">
<emphasis id="B949EAD5FF98FFB2FF8CFB64697DFB29" box="[161,271,1231,1257]" italics="true" pageId="26" pageNumber="27">Clavatula</emphasis>
</taxonomicName>
<emphasis id="B949EAD5FF98FFB2FE32FB6469DAFB28" box="[287,424,1231,1256]" italics="true" pageId="26" pageNumber="27">ariejansseni</emphasis>
nov. sp.
</paragraph>
</subSubSection>
<subSubSection id="C327654CFF98FFB2FFEAFB596BB8FACD" box="[199,970,1266,1293]" pageId="26" pageNumber="27" type="biology_ecology">
<paragraph id="8B8236C7FF98FFB2FFEAFB596BB8FACD" blockId="26.[151,1437,150,1366]" box="[199,970,1266,1293]" pageId="26" pageNumber="27">
<emphasis id="B949EAD5FF98FFB2FFEAFB5969DAFACC" bold="true" box="[199,424,1266,1292]" pageId="26" pageNumber="27">Paleoenvironment.</emphasis>
Inner neritic environments (
<bibRefCitation id="EFAC4B36FF98FFB2FDCBFB586BCDFACD" author="Landau, B. M. &amp; Harzhauser, M. &amp; Yslamodlu, Y. &amp; Marques da Silva, C." box="[742,959,1267,1293]" pageId="26" pageNumber="27" pagination="3 - 576" refId="ref137659" refString="Landau, B. M., Harzhauser, M., Yslamodlu, Y. &amp; Marques da Silva, C. (2013) Systematics and palaeobiogeography of the gastropods of the middle Miocene (Serravallian) Karaman Basin of Turkey. Cainozoic Research, 11 - 13, 3 - 576" type="journal article" year="2013">
Landau
<emphasis id="B949EAD5FF98FFB2FC6EFB586B09FACD" box="[835,891,1267,1293]" italics="true" pageId="26" pageNumber="27">et al.</emphasis>
2013
</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C327654CFF98FFB2FFEAFABD6CE6FA95" pageId="26" pageNumber="27" type="distribution">
<paragraph id="8B8236C7FF98FFB2FFEAFABD6CE6FA95" blockId="26.[151,1437,150,1366]" pageId="26" pageNumber="27">
<emphasis id="B949EAD5FF98FFB2FFEAFABD698DFAF0" bold="true" box="[199,511,1302,1328]" pageId="26" pageNumber="27">Proto-Mediterranean Sea:</emphasis>
Serravallian (middle Miocene):
<emphasis id="B949EAD5FF98FFB2FC49FABC6C61FAF0" box="[868,1043,1303,1328]" italics="true" pageId="26" pageNumber="27">
<collectingRegion id="49F9F825FF98FFB2FC49FABC6BBCFAF0" box="[868,974,1303,1328]" country="Turkey" name="Karaman" pageId="26" pageNumber="27">Karaman</collectingRegion>
Basin
</emphasis>
: Lale,Akboðazi, Akpýnar-Pýnarlar Yaylasý, Seyithasan, Road to Mut,
<quantity id="4CC59B22FF98FFB2FD0CFA906A1AFA95" box="[545,616,1339,1365]" metricMagnitude="4" metricUnit="m" metricValue="2.7" pageId="26" pageNumber="27" unit="km" value="27.0">27 km</quantity>
south of
<collectingRegion id="49F9F825FF98FFB2FDFCFA906B4AFA95" box="[721,824,1339,1365]" country="Turkey" name="Karaman" pageId="26" pageNumber="27">Karaman</collectingRegion>
(
<collectingCountry id="F32A7657FF98FFB2FC6BFA906BEBFA96" box="[838,921,1339,1366]" name="Turkey" pageId="26" pageNumber="27">Turkey</collectingCountry>
) (
<bibRefCitation id="EFAC4B36FF98FFB2FC82FA906CFAFA95" author="Landau, B. M. &amp; Harzhauser, M. &amp; Yslamodlu, Y. &amp; Marques da Silva, C." box="[943,1160,1339,1365]" pageId="26" pageNumber="27" pagination="3 - 576" refId="ref137659" refString="Landau, B. M., Harzhauser, M., Yslamodlu, Y. &amp; Marques da Silva, C. (2013) Systematics and palaeobiogeography of the gastropods of the middle Miocene (Serravallian) Karaman Basin of Turkey. Cainozoic Research, 11 - 13, 3 - 576" type="journal article" year="2013">
Landau
<emphasis id="B949EAD5FF98FFB2FB21FA906C36FA95" box="[1036,1092,1339,1365]" italics="true" pageId="26" pageNumber="27">et al.</emphasis>
2013
</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>