253 lines
30 KiB
XML
253 lines
30 KiB
XML
<document id="8B3645F80AB80F4F94C3614CD6FA6E09" ID-CLB-Dataset="20492" ID-DOI="10.5169/seals-787049" ID-GBIF-Dataset="ce0756e2-5184-4b74-8472-ab69fed7951c" ID-ISSN="2624-9170" ID-Zenodo-Dep="6285683" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1637935326602" checkinUser="carolina" docAuthor="Huber, Charles & Schmidt, Joachim" docDate="2017" docId="03948799FFFBFFE1FF41FAC8FC78F9BE" docLanguage="en" docName="ContribNatHist.36.1-85.pdf.imf" docOrigin="Contributions to Natural History 36" docStyle="DocumentStyle:D319E9290ED630111D2A11900DAFC11F.4:ContribNatHist.2011-.journal_article" docStyleId="D319E9290ED630111D2A11900DAFC11F" docStyleName="ContribNatHist.2011-.journal_article" docStyleVersion="4" docTitle="Nebria (Epinebriola) zayula ANDREWES 1936" docType="treatment" docVersion="12" lastPageNumber="34" masterDocId="FFADFFE1FFDBFFC0FF9CFFE9FFB0FFA7" masterDocTitle="Notes on the Nebria subgenus Epinebriola K. DANIEL, 1904 with the description of Barbonebriola subgen. nov. and 13 new species from the Himalaya-Tibet orogen (Coleoptera, Carabidae, Nebriini)" masterLastPageNumber="85" masterPageNumber="1" pageNumber="33" updateTime="1699470525451" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:title id="5F8B63CA2E47FCD03C4A3F2E1B3B167D">Notes on the Nebria subgenus Epinebriola K. DANIEL, 1904 with the description of Barbonebriola subgen. nov. and 13 new species from the Himalaya-Tibet orogen (Coleoptera, Carabidae, Nebriini)</mods:title>
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<mods:namePart id="DCE8E9DD6362D11182A9413BB5CE102D">Huber, Charles</mods:namePart>
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<mods:namePart id="5364E49F3A7C659ECE9453941B0055FC">Schmidt, Joachim</mods:namePart>
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<mods:title id="06806E29667ED55944C71A08C4DE0C00">Contributions to Natural History</mods:title>
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<mods:date id="53A0B1A394F4E86B2FC8FCA684E0FB03">2017</mods:date>
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<mods:number id="FEF6736D3258C0913FD6C5D54E8D3779">2017-07-17</mods:number>
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<mods:number id="6EFC1904E443E30B4770C53897A0B965">36</mods:number>
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<treatment id="03948799FFFBFFE1FF41FAC8FC78F9BE" ID-DOI="http://doi.org/10.5281/zenodo.5729942" ID-GBIF-Taxon="193820257" ID-Zenodo-Dep="5729942" LSID="urn:lsid:plazi:treatment:03948799FFFBFFE1FF41FAC8FC78F9BE" httpUri="http://treatment.plazi.org/id/03948799FFFBFFE1FF41FAC8FC78F9BE" lastPageId="33" lastPageNumber="34" pageId="32" pageNumber="33">
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<subSubSection id="C3276504FFFBFFE0FF41FAC8FD6CFA9E" box="[221,732,1313,1338]" pageId="32" pageNumber="33" type="nomenclature">
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<paragraph id="8B82368FFFFBFFE0FF41FAC8FD6CFA9E" blockId="32.[221,732,1313,1338]" box="[221,732,1313,1338]" pageId="32" pageNumber="33">
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<heading id="D0CA81E3FFFBFFE0FF41FAC8FD6CFA9E" box="[221,732,1313,1338]" centered="true" fontSize="12" level="2" pageId="32" pageNumber="33" reason="2">
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<taxonomicName id="4C3D4D0CFFFBFFE0FF41FAC8FD6CFA9E" ID-CoL="45YTK" authority="ANDREWES, 1936" authorityName="ANDREWES" authorityYear="1936" box="[221,732,1313,1338]" class="Insecta" family="Carabidae" genus="Nebria" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="zayula" subGenus="Epinebriola">
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<emphasis id="B949EA9DFFFBFFE0FF41FAC8FD6CFA9E" bold="true" box="[221,732,1313,1338]" pageId="32" pageNumber="33">
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<emphasis id="B949EA9DFFFBFFE0FF41FAC8FE99FA9D" bold="true" box="[221,297,1313,1338]" italics="true" pageId="32" pageNumber="33">Nebria</emphasis>
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(
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<emphasis id="B949EA9DFFFBFFE0FEA5FAC8FE0BFA9D" bold="true" box="[313,443,1313,1338]" italics="true" pageId="32" pageNumber="33">Epinebriola</emphasis>
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)
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<emphasis id="B949EA9DFFFBFFE0FE57FAC8FDA6FA9D" bold="true" box="[459,534,1313,1338]" italics="true" pageId="32" pageNumber="33">zayula</emphasis>
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<bibRefCitation id="EFAC4B7EFFFBFFE0FD80FAC8FD6CFA9E" author="Andrewes, H. E." box="[540,732,1313,1337]" pageId="32" pageNumber="33" pagination="54 - 65" refId="ref32738" refString="Andrewes, H. E. (1936): Paper on Oriental Carabidae, 30. - The annals and magazine of natural history, Serie 10, 18: 54 - 65." type="journal article" year="1936">ANDREWES, 1936</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3276504FFFBFFE1FF41FA85FEF0FBFE" lastPageId="33" lastPageNumber="34" pageId="32" pageNumber="33" type="discussion">
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<paragraph id="8B82368FFFFBFFE0FF41FA85FE83F9BE" blockId="32.[221,1106,1388,1636]" pageId="32" pageNumber="33">
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Remarks on the
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<typeStatus id="5486882DFFFBFFE0FE07FA85FE7EFA23" box="[411,462,1388,1412]" pageId="32" pageNumber="33">type</typeStatus>
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locality: When describing the species
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<bibRefCitation id="EFAC4B7EFFFBFFE0FC12FA85FBE2FA23" author="Andrewes, H. E." box="[910,1106,1388,1412]" pageId="32" pageNumber="33" pagination="54 - 65" refId="ref32738" refString="Andrewes, H. E. (1936): Paper on Oriental Carabidae, 30. - The annals and magazine of natural history, Serie 10, 18: 54 - 65." type="journal article" year="1936">Andrewes (1936)</bibRefCitation>
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noted the
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<typeStatus id="5486882DFFFBFFE0FECBFA78FE3BFA0E" box="[343,395,1425,1449]" pageId="32" pageNumber="33">type</typeStatus>
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locality "Kawy La", a locality obviously unknow to him, and which he interpreted as follows: "
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<emphasis id="B949EA9DFFFBFFE0FDF9FA5FFEADF9BE" italics="true" pageId="32" pageNumber="33">
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The locality is apparently the
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<taxonomicName id="4C3D4D0CFFFBFFE0FC23FA5FFB83FA69" authority="Pass" authorityName="Pass" box="[959,1075,1462,1486]" class="Gastropoda" family="Viviparidae" genus="Kaya" kingdom="Animalia" order="Architaenioglossa" pageId="32" pageNumber="33" phylum="Mollusca" rank="genus">Kaya Pass</taxonomicName>
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of the Indian Ordnance Survey map, connecting the valley of the Dibang and Zayul rivers
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</emphasis>
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."
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</paragraph>
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<paragraph id="8B82368FFFFBFFE1FE9FF9CFFEF0FBFE" blockId="32.[221,1106,1388,1636]" lastBlockId="33.[184,1069,641,1561]" lastPageId="33" lastPageNumber="34" pageId="32" pageNumber="33">
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It was obviously a reading error by Andrewes: On the original label of the type specimen "Kang La" is legibly handwritten, but neither "Kawy La" nor "
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<taxonomicName id="4C3D4D0CFFFAFFE1FF23FD68FE95FD3E" authority="La" authorityName="La" box="[191,293,641,665]" class="Gastropoda" family="Viviparidae" genus="Kaya" kingdom="Animalia" order="Architaenioglossa" pageId="33" pageNumber="34" phylum="Mollusca" rank="genus">Kaya La</taxonomicName>
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". Due to the reading error, Andrewes made an interpretation error: The misinterpreted locality (
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<taxonomicName id="4C3D4D0CFFFAFFE1FE68FD4FFDE3FD19" authority="La" authorityName="La" box="[500,595,678,702]" class="Gastropoda" family="Viviparidae" genus="Kaya" kingdom="Animalia" order="Architaenioglossa" pageId="33" pageNumber="34" phylum="Mollusca" rank="genus">Kaya La</taxonomicName>
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) at the India-China border indeed crosses the Mishmi Mountains westwards to the Dibang river valley, but the correct type locality Kang La (= Ata Kang La,
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<figureCitation id="13062A0AFFFAFFE1FDEEFD18FD0FFCAE" box="[626,703,753,777]" captionStart="Fig" captionStartId="32.[221,250,160,178]" captionTargetBox="[463,1093,143,1241]" captionTargetId="figure-104@32.[463,1106,143,1251]" captionTargetPageId="32" captionText="Fig. 18: Copy of Kaul- back’s map (Kaulback 1934, p. 87) showing the more northerly part of Kingdon Ward’s and Ronald Kaulback’s expedition route in 1933 to the Ata Kang La (= type locality of Nebria (Epinebriola) zayula ANDREWES, 1936; black arrow). Red dot = approximate position of the misin- terpreted type locality Kaya La (= Kawy La) in Andrewes (1936)." figureDoi="http://doi.org/10.5281/zenodo.6285758" httpUri="https://zenodo.org/record/6285758/files/figure.png" pageId="33" pageNumber="34">Fig. 18</figureCitation>
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) crosses in a linear distance of approx.
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<quantity id="4CC59B6AFFFAFFE1FE88FCFFFEEBFC89" box="[276,347,790,814]" metricMagnitude="4" metricUnit="m" metricValue="5.0" pageId="33" pageNumber="34" unit="km" value="50.0">50 km</quantity>
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the Kangri Garpo Mountains northwards, connecting the Valley of the Kangrigarpo Qu and the valley of Rawok Lake and Mang Lake (
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<bibRefCitation id="EFAC4B7EFFFAFFE1FC5DFCD5FF42FCDE" author="Kaulback, R." pageId="33" pageNumber="34" refId="ref33333" refString="Kaulback, R. (1934): Tibetan Trek. - 319 pp., London." type="book" year="1934">Kaulback 1934</bibRefCitation>
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,
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<bibRefCitation id="EFAC4B7EFFFAFFE1FE9EFC88FE0AFCDE" author="Nakamura, T." box="[258,442,865,889]" pageId="33" pageNumber="34" pagination="1 - 7" refId="ref33705" refString="Nakamura, T. (2001): Kangri Garpo range in Southeast Tibet. - Japanese Alpine News 1: 1 - 7." type="journal article" year="2001">Nakamura 2001</bibRefCitation>
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).
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<bibRefCitation id="EFAC4B7EFFFAFFE1FE4FFC88FD21FCDE" author="Kaulback, R." box="[467,657,865,889]" pageId="33" pageNumber="34" refId="ref33333" refString="Kaulback, R. (1934): Tibetan Trek. - 319 pp., London." type="book" year="1934">Kaulback (1934)</bibRefCitation>
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, member of the expedition in 1933 and one of the two beetle collectors, noted in his travel report
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<emphasis id="B949EA9DFFFAFFE1FC14FC6FFF63FC64" italics="true" pageId="33" pageNumber="34">
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"… on
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<emphasis id="B949EA9DFFFAFFE1FC48FC6FFF63FC64" bold="true" italics="true" pageId="33" pageNumber="34">July the 15</emphasis>
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</emphasis>
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<emphasis id="B949EA9DFFFAFFE1FF4FFC41FF56FC1B" bold="true" box="[211,230,936,956]" italics="true" pageId="33" pageNumber="34">
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<superScript id="7C489BC7FFFAFFE1FF4FFC41FF56FC1B" attach="left" box="[211,230,936,956]" fontSize="8" pageId="33" pageNumber="34">th</superScript>
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</emphasis>
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[1933]
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<emphasis id="B949EA9DFFFAFFE1FEA6FC45FC40FC63" box="[314,1008,939,964]" italics="true" pageId="33" pageNumber="34">
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… I went with Kingdon Ward to the top of the
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<emphasis id="B949EA9DFFFAFFE1FCA1FC42FC75FC63" bold="true" box="[829,965,939,964]" italics="true" pageId="33" pageNumber="34">Ata Kang La</emphasis>
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…."
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</emphasis>
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, providing a travelling map (
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<figureCitation id="13062A0AFFFAFFE1FE54FC38FDA6FC4E" box="[456,534,977,1001]" captionStart="Fig" captionStartId="32.[221,250,160,178]" captionTargetBox="[463,1093,143,1241]" captionTargetId="figure-104@32.[463,1106,143,1251]" captionTargetPageId="32" captionText="Fig. 18: Copy of Kaul- back’s map (Kaulback 1934, p. 87) showing the more northerly part of Kingdon Ward’s and Ronald Kaulback’s expedition route in 1933 to the Ata Kang La (= type locality of Nebria (Epinebriola) zayula ANDREWES, 1936; black arrow). Red dot = approximate position of the misin- terpreted type locality Kaya La (= Kawy La) in Andrewes (1936)." figureDoi="http://doi.org/10.5281/zenodo.6285758" httpUri="https://zenodo.org/record/6285758/files/figure.png" pageId="33" pageNumber="34">Fig. 18</figureCitation>
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) and thus confirming the collecting date noted on the label of the
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<typeStatus id="5486882DFFFAFFE1FE16FC1FFE5FFBA9" box="[394,495,1014,1038]" pageId="33" pageNumber="34" type="holotype">holotype</typeStatus>
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specimen. Obviously Andrewes had no knowledge of Kaulback's map which was published two years before Andrewes described the species.
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</paragraph>
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</subSubSection>
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<caption id="DF426607FFFAFFE1FF24FF49FD79FF6A" ID-Table-UUID="DF426607FFFAFFE1FF24FF49FD79FF6A" httpUri="http://table.plazi.org/id/DF426607FFFAFFE1FF24FF49FD79FF6A" pageId="33" pageNumber="34" startId="33.[184,218,160,178]" targetBox="[192,1045,262,517]" targetIsTable="true" targetPageId="33">
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<paragraph id="8B82368FFFFAFFE1FF24FF49FD79FF6A" blockId="33.[184,1069,160,205]" pageId="33" pageNumber="34">
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Tab. 2: Relevant characters of the
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<taxonomicName id="4C3D4D0CFFFAFFE1FE43FF49FD7BFF15" authorityName="ANDREWES" authorityYear="1936" box="[479,715,160,178]" class="Insecta" family="Carabidae" genus="Nebria" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="zayula" subGenus="Epinebriola">
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<emphasis id="B949EA9DFFFAFFE1FE43FF49FDA8FF15" bold="true" box="[479,536,160,178]" italics="true" pageId="33" pageNumber="34">Nebria</emphasis>
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(
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<emphasis id="B949EA9DFFFAFFE1FDB9FF49FD37FF15" bold="true" box="[549,647,160,178]" italics="true" pageId="33" pageNumber="34">Epinebriola</emphasis>
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)
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<emphasis id="B949EA9DFFFAFFE1FD0FFF49FD7BFF15" bold="true" box="[659,715,160,178]" italics="true" pageId="33" pageNumber="34">zayula</emphasis>
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</taxonomicName>
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type specimen which are not or wrongly mentioned in
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<bibRefCitation id="EFAC4B7EFFFAFFE1FEB2FF52FE73FF6A" author="Andrewes, H. E." box="[302,451,187,205]" pageId="33" pageNumber="34" pagination="54 - 65" refId="ref32738" refString="Andrewes, H. E. (1936): Paper on Oriental Carabidae, 30. - The annals and magazine of natural history, Serie 10, 18: 54 - 65." type="journal article" year="1936">Andrewes (1936)</bibRefCitation>
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and in
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<bibRefCitation id="EFAC4B7EFFFAFFE1FD9DFF52FD74FF6A" author="Ledoux, G. & Roux, P." box="[513,708,187,205]" pageId="33" pageNumber="34" refId="ref33536" refString="Ledoux, G. & Roux, P. (2005): Nebria (Coleoptera, Nebriidae) Faune mondiale. - 976 pp., Saint-Just-la-Pendue." type="book" year="2005">Ledoux & Roux (2005)</bibRefCitation>
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||
.
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||
</paragraph>
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||
</caption>
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||
<paragraph id="8B82368FFFFAFFE1FF5FFEEFFCF8FDA2" pageId="33" pageNumber="34">
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||
<table id="F93DC42FFFFA003FFF5CFEEFFBA5FDA2" box="[192,1045,262,517]" gridcols="3" gridrows="6" pageId="33" pageNumber="34">
|
||
<tr id="350D34CDFFFA003FFF5CFEEFFBA5FE97" box="[192,1045,262,304]" gridrow="0" pageId="33" pageNumber="34">
|
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<th id="76DC5DB1FFFA003FFF5CFEEFFE6FFE97" box="[192,479,262,304]" gridcol="0" gridrow="0" pageId="33" pageNumber="34">
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||
<emphasis id="B949EA9DFFFAFFE1FF5FFEEFFE24FE97" bold="true" pageId="33" pageNumber="34">
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||
Characters in the
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||
<taxonomicName id="4C3D4D0CFFFAFFE1FEC5FEEFFE18FEBF" authorityName="ANDREWES" authorityYear="1936" box="[345,424,262,280]" class="Insecta" family="Carabidae" genus="Nebria" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="zayula">
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||
<emphasis id="B949EA9DFFFAFFE1FEC5FEEFFE18FEBF" bold="true" box="[345,424,262,280]" italics="true" pageId="33" pageNumber="34">N. zayula</emphasis>
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</taxonomicName>
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male holotype specimen
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</emphasis>
|
||
</th>
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<th id="76DC5DB1FFFA003FFD8DFEEFFD50FE97" box="[529,736,262,304]" gridcol="1" gridrow="0" pageId="33" pageNumber="34">
|
||
<bibRefCitation id="EFAC4B7EFFFAFFE1FD89FEEFFD16FEBF" author="Andrewes, H. E." box="[533,678,262,280]" pageId="33" pageNumber="34" pagination="54 - 65" refId="ref32738" refString="Andrewes, H. E. (1936): Paper on Oriental Carabidae, 30. - The annals and magazine of natural history, Serie 10, 18: 54 - 65." type="journal article" year="1936">
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||
<emphasis id="B949EA9DFFFAFFE1FD89FEEFFD16FEBF" bold="true" box="[533,678,262,280]" pageId="33" pageNumber="34">Andrewes (1936)</emphasis>
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||
</bibRefCitation>
|
||
</th>
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||
<th id="76DC5DB1FFFA003FFC97FEEFFBA5FE97" box="[779,1045,262,304]" gridcol="2" gridrow="0" pageId="33" pageNumber="34">
|
||
<bibRefCitation id="EFAC4B7EFFFAFFE1FC92FEEFFC7CFEBF" author="Ledoux, G. & Roux, P." box="[782,972,262,280]" pageId="33" pageNumber="34" refId="ref33536" refString="Ledoux, G. & Roux, P. (2005): Nebria (Coleoptera, Nebriidae) Faune mondiale. - 976 pp., Saint-Just-la-Pendue." type="book" year="2005">
|
||
<emphasis id="B949EA9DFFFAFFE1FC92FEEFFC7CFEBF" bold="true" box="[782,972,262,280]" pageId="33" pageNumber="34">Ledoux & Roux (2005)</emphasis>
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</bibRefCitation>
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||
</th>
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||
</tr>
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||
<tr id="350D34CDFFFA003FFF5CFEAEFBA5FEFE" box="[192,1045,327,345]" gridrow="1" pageId="33" pageNumber="34">
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<th id="76DC5DB1FFFA003FFF5CFEAEFE6FFEFE" box="[192,479,327,345]" gridcol="0" gridrow="1" pageId="33" pageNumber="34">Head:supraorbitally bisetose</th>
|
||
<td id="76DC5DB1FFFA003FFD8DFEAEFD50FEFE" box="[529,736,327,345]" gridcol="1" gridrow="1" pageId="33" pageNumber="34">Not mentioned</td>
|
||
<td id="76DC5DB1FFFA003FFC97FEAEFBA5FEFE" box="[779,1045,327,345]" gridcol="2" gridrow="1" pageId="33" pageNumber="34">Not mentioned</td>
|
||
</tr>
|
||
<tr id="350D34CDFFFA003FFF5CFE8FFBA5FE37" box="[192,1045,358,400]" gridrow="2" pageId="33" pageNumber="34">
|
||
<th id="76DC5DB1FFFA003FFF5CFE8FFE6FFE37" box="[192,479,358,400]" gridcol="0" gridrow="2" pageId="33" pageNumber="34">Submentum: 16 setae (9+7)</th>
|
||
<td id="76DC5DB1FFFA003FFD8DFE8FFD50FE37" box="[529,736,358,400]" gridcol="1" gridrow="2" pageId="33" pageNumber="34">More than a dozen setae</td>
|
||
<td id="76DC5DB1FFFA003FFC97FE8FFBA5FE37" box="[779,1045,358,400]" gridcol="2" gridrow="2" pageId="33" pageNumber="34">A dozen setae (14 setae mentioned in the key)</td>
|
||
</tr>
|
||
<tr id="350D34CDFFFA003FFF5CFE74FBA5FE08" box="[192,1045,413,431]" gridrow="3" pageId="33" pageNumber="34">
|
||
<th id="76DC5DB1FFFA003FFF5CFE74FE6FFE08" box="[192,479,413,431]" gridcol="0" gridrow="3" pageId="33" pageNumber="34">Sterna 4–6: 4–5 setae</th>
|
||
<td id="76DC5DB1FFFA003FFD8DFE74FD50FE08" box="[529,736,413,431]" gridcol="1" gridrow="3" pageId="33" pageNumber="34">3–4 setae</td>
|
||
<td id="76DC5DB1FFFA003FFC97FE74FBA5FE08" box="[779,1045,413,431]" gridcol="2" gridrow="3" pageId="33" pageNumber="34">3–4 setae</td>
|
||
</tr>
|
||
<tr id="350D34CDFFFA003FFF5CFE55FBA5FE41" box="[192,1045,444,486]" gridrow="4" pageId="33" pageNumber="34">
|
||
<th id="76DC5DB1FFFA003FFF5CFE55FE6FFE41" box="[192,479,444,486]" gridcol="0" gridrow="4" pageId="33" pageNumber="34">Anal sternum: 4–5 setae each side in male</th>
|
||
<td id="76DC5DB1FFFA003FFD8DFE55FD50FE41" box="[529,736,444,486]" gridcol="1" gridrow="4" pageId="33" pageNumber="34">Not mentioned</td>
|
||
<td id="76DC5DB1FFFA003FFC97FE55FBA5FE41" box="[779,1045,444,486]" gridcol="2" gridrow="4" pageId="33" pageNumber="34">1 seta each side in male (2 setae mentioned in the key)</td>
|
||
</tr>
|
||
<tr id="350D34CDFFFA003FFF5CFE1AFBA5FDA2" box="[192,1045,499,517]" gridrow="5" pageId="33" pageNumber="34">
|
||
<th id="76DC5DB1FFFA003FFF5CFE1AFE6FFDA2" box="[192,479,499,517]" gridcol="0" gridrow="5" pageId="33" pageNumber="34">Metacoxae: 3–5 setae</th>
|
||
<td id="76DC5DB1FFFA003FFD8DFE1AFD50FDA2" box="[529,736,499,517]" gridcol="1" gridrow="5" pageId="33" pageNumber="34">3 setae</td>
|
||
<td id="76DC5DB1FFFA003FFC97FE1AFBA5FDA2" box="[779,1045,499,517]" gridcol="2" gridrow="5" pageId="33" pageNumber="34">3 setae</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<subSubSection id="C3276504FFFAFFE1FF41FB8FFCB4FB6E" pageId="33" pageNumber="34" type="distribution">
|
||
<paragraph id="8B82368FFFFAFFE1FF41FB8FFCB4FB6E" blockId="33.[184,1069,641,1561]" pageId="33" pageNumber="34">
|
||
Distribution:
|
||
<taxonomicName id="4C3D4D0CFFFAFFE1FEE4FB8FFE54FBD9" authorityName="ANDREWES" authorityYear="1936" box="[376,484,1126,1150]" class="Insecta" family="Carabidae" genus="Nebria" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="zayula">
|
||
<emphasis id="B949EA9DFFFAFFE1FEE4FB8FFE54FBD9" box="[376,484,1126,1150]" italics="true" pageId="33" pageNumber="34">N. zayula</emphasis>
|
||
</taxonomicName>
|
||
is known only from the
|
||
<typeStatus id="5486882DFFFAFFE1FD63FB8FFC83FBD9" box="[767,819,1126,1150]" pageId="33" pageNumber="34">type</typeStatus>
|
||
locality, but the
|
||
<typeStatus id="5486882DFFFAFFE1FC65FB8FFB9CFBD9" box="[1017,1068,1126,1150]" pageId="33" pageNumber="34">type</typeStatus>
|
||
locality lies east of the deeply incised Kangrigarpo Qu river valley (= Rong To Chu in
|
||
<figureCitation id="13062A0AFFFAFFE1FE98FB58FEFFFB6E" box="[260,335,1201,1225]" captionStart="Fig" captionStartId="32.[221,250,160,178]" captionTargetBox="[463,1093,143,1241]" captionTargetId="figure-104@32.[463,1106,143,1251]" captionTargetPageId="32" captionText="Fig. 18: Copy of Kaul- back’s map (Kaulback 1934, p. 87) showing the more northerly part of Kingdon Ward’s and Ronald Kaulback’s expedition route in 1933 to the Ata Kang La (= type locality of Nebria (Epinebriola) zayula ANDREWES, 1936; black arrow). Red dot = approximate position of the misin- terpreted type locality Kaya La (= Kawy La) in Andrewes (1936)." figureDoi="http://doi.org/10.5281/zenodo.6285758" httpUri="https://zenodo.org/record/6285758/files/figure.png" pageId="33" pageNumber="34">Fig. 18</figureCitation>
|
||
) and not west of it as supposed so far.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3276504FFFAFFE1FF41FB3FFC78F9BE" pageId="33" pageNumber="34" type="description">
|
||
<paragraph id="8B82368FFFFAFFE1FF41FB3FFC78F9BE" blockId="33.[184,1069,641,1561]" pageId="33" pageNumber="34">
|
||
Description of additional morphological characters in
|
||
<taxonomicName id="4C3D4D0CFFFAFFE1FCA9FB3FFC2CFB49" authorityName="ANDREWES" authorityYear="1936" box="[821,924,1238,1262]" class="Insecta" family="Carabidae" genus="Nebria" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="zayula">
|
||
<emphasis id="B949EA9DFFFAFFE1FCA9FB3FFC2CFB49" box="[821,924,1238,1262]" italics="true" pageId="33" pageNumber="34">N. zayula</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<tableCitation id="C6BF0334FFFAFFE1FC35FB3FFC5FFB49" box="[937,1007,1238,1262]" captionStart="Tab" captionStartId="33.[184,218,160,178]" captionTargetPageId="33" captionText="Tab. 2: Relevant characters of the Nebria (Epinebriola) zayula type specimen which are not or wrongly mentioned in Andrewes (1936) and in Ledoux & Roux (2005)." httpUri="http://table.plazi.org/id/DF426607FFFAFFE1FF24FF49FD79FF6A" pageId="33" pageNumber="34" tableUuid="DF426607FFFAFFE1FF24FF49FD79FF6A">Tab. 2</tableCitation>
|
||
): We examined several relevant characters of the
|
||
<typeStatus id="5486882DFFFAFFE1FD22FB15FC93FAB3" box="[702,803,1276,1300]" pageId="33" pageNumber="34" type="holotype">holotype</typeStatus>
|
||
specimen of
|
||
<taxonomicName id="4C3D4D0CFFFAFFE1FC5DFB15FB9CFAB3" authorityName="ANDREWES" authorityYear="1936" box="[961,1068,1276,1300]" class="Insecta" family="Carabidae" genus="Nebria" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="zayula">
|
||
<emphasis id="B949EA9DFFFAFFE1FC5DFB15FB9CFAB3" box="[961,1068,1276,1300]" italics="true" pageId="33" pageNumber="34">N. zayula</emphasis>
|
||
</taxonomicName>
|
||
which had been overlooked and/or wrongly noted by
|
||
<bibRefCitation id="EFAC4B7EFFFAFFE1FC89FAC8FC67FA9E" author="Andrewes, H. E." box="[789,983,1313,1337]" pageId="33" pageNumber="34" pagination="54 - 65" refId="ref32738" refString="Andrewes, H. E. (1936): Paper on Oriental Carabidae, 30. - The annals and magazine of natural history, Serie 10, 18: 54 - 65." type="journal article" year="1936">Andrewes (1936)</bibRefCitation>
|
||
as well as by
|
||
<bibRefCitation id="EFAC4B7EFFFAFFE1FE9CFAAFFDBBFAF9" author="Ledoux, G. & Roux, P." box="[256,523,1350,1374]" pageId="33" pageNumber="34" refId="ref33536" refString="Ledoux, G. & Roux, P. (2005): Nebria (Coleoptera, Nebriidae) Faune mondiale. - 976 pp., Saint-Just-la-Pendue." type="book" year="2005">Ledoux & Roux (2005)</bibRefCitation>
|
||
. The head supraorbitally bisetose – true in
|
||
<taxonomicName id="4C3D4D0CFFFAFFE1FB8FFAAFFEB0FA23" authorityName="ANDREWES" authorityYear="1936" class="Insecta" family="Carabidae" genus="Nebria" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="zayula">
|
||
<emphasis id="B949EA9DFFFAFFE1FB8FFAAFFEB0FA23" italics="true" pageId="33" pageNumber="34">N. zayula</emphasis>
|
||
</taxonomicName>
|
||
, but overlooked so far – is rare in the subgenus
|
||
<taxonomicName id="4C3D4D0CFFFAFFE1FC8AFA85FC25FA23" authorityName="K. DANIEL" authorityYear="1904" box="[790,917,1388,1412]" class="Insecta" family="Carabidae" genus="Nebria" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="subGenus" subGenus="Epinebriola">
|
||
<emphasis id="B949EA9DFFFAFFE1FC8AFA85FC25FA23" box="[790,917,1388,1412]" italics="true" pageId="33" pageNumber="34">Epinebriola</emphasis>
|
||
</taxonomicName>
|
||
, only present in
|
||
<taxonomicName id="4C3D4D0CFFFAFFE1FF49FA78FE96FA0E" authorityName="ANDREWES" authorityYear="1936" box="[213,294,1425,1449]" class="Insecta" family="Carabidae" genus="Nebria" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="rasa">
|
||
<emphasis id="B949EA9DFFFAFFE1FF49FA78FE96FA0E" box="[213,294,1425,1449]" italics="true" pageId="33" pageNumber="34">N. rasa</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C3D4D0CFFFAFFE1FEFDFA78FDB0FA0E" authorityName="SHILENKOV" authorityYear="1998" box="[353,512,1425,1449]" class="Insecta" family="Carabidae" genus="Nebria" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="schawalleri">
|
||
<emphasis id="B949EA9DFFFAFFE1FEFDFA78FDB0FA0E" box="[353,512,1425,1449]" italics="true" pageId="33" pageNumber="34">N. schawalleri</emphasis>
|
||
</taxonomicName>
|
||
, and asymmetrically present in
|
||
<taxonomicName id="4C3D4D0CFFFAFFE1FCF3FA78FC49FA0E" authorityName="HUBER & SCHMIDT" authorityYear="2007" box="[879,1017,1425,1449]" class="Insecta" family="Carabidae" genus="Nebria" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="molendai">
|
||
<emphasis id="B949EA9DFFFAFFE1FCF3FA78FC49FA0E" box="[879,1017,1425,1449]" italics="true" pageId="33" pageNumber="34">N. molendai</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<emphasis id="B949EA9DFFFAFFE1FF24FA5FFE04FA69" box="[184,436,1462,1486]" italics="true" pageId="33" pageNumber="34">
|
||
<taxonomicName id="4C3D4D0CFFFAFFE1FF24FA5FFE8FFA69" authorityName="HUBER & SCHMIDT" authorityYear="2012" box="[184,319,1462,1486]" class="Insecta" family="Carabidae" genus="Nebria" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="martensi">N. martensi</taxonomicName>
|
||
.
|
||
<taxonomicName id="4C3D4D0CFFFAFFE1FED7FA5FFE04FA69" authorityName="ANDREWES" authorityYear="1936" box="[331,436,1462,1486]" class="Insecta" family="Carabidae" genus="Nebria" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="zayula">N. zayula</taxonomicName>
|
||
</emphasis>
|
||
is even the only
|
||
<taxonomicName id="4C3D4D0CFFFAFFE1FDE0FA5FFD4CFA69" authorityName="K. DANIEL" authorityYear="1904" box="[636,764,1462,1486]" class="Insecta" family="Carabidae" genus="Nebria" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="79" phylum="Arthropoda" rank="subGenus" subGenus="Epinebriola">
|
||
<emphasis id="B949EA9DFFFAFFE1FDE0FA5FFD4CFA69" box="[636,764,1462,1486]" italics="true" pageId="33" pageNumber="34">Epinebriola</emphasis>
|
||
</taxonomicName>
|
||
species with a row of 4–5 setae on each side of the male anal sternum (not mentioned in
|
||
<bibRefCitation id="EFAC4B7EFFFAFFE1FCEBFA35FB95FA53" author="Andrewes, H. E." box="[887,1061,1500,1524]" pageId="33" pageNumber="34" pagination="54 - 65" refId="ref32738" refString="Andrewes, H. E. (1936): Paper on Oriental Carabidae, 30. - The annals and magazine of natural history, Serie 10, 18: 54 - 65." type="journal article" year="1936">Andrewes 1936</bibRefCitation>
|
||
, wrongly noted in
|
||
<bibRefCitation id="EFAC4B7EFFFAFFE1FEEAF9E8FDEBF9BE" author="Ledoux, G. & Roux, P." box="[374,603,1537,1561]" pageId="33" pageNumber="34" refId="ref33536" refString="Ledoux, G. & Roux, P. (2005): Nebria (Coleoptera, Nebriidae) Faune mondiale. - 976 pp., Saint-Just-la-Pendue." type="book" year="2005">Ledoux & Roux 2005</bibRefCitation>
|
||
). For other characters see
|
||
<tableCitation id="C6BF0334FFFAFFE1FC1CF9E8FC71F9BE" box="[896,961,1537,1561]" captionStart="Tab" captionStartId="33.[184,218,160,178]" captionTargetPageId="33" captionText="Tab. 2: Relevant characters of the Nebria (Epinebriola) zayula type specimen which are not or wrongly mentioned in Andrewes (1936) and in Ledoux & Roux (2005)." httpUri="http://table.plazi.org/id/DF426607FFFAFFE1FF24FF49FD79FF6A" pageId="33" pageNumber="34" tableUuid="DF426607FFFAFFE1FF24FF49FD79FF6A">Tab. 2</tableCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |