197 lines
19 KiB
XML
197 lines
19 KiB
XML
<document id="8C4FFD15729B4A58525EE8EC4027057E" ID-DOI="10.11646/zootaxa.3926.4.4" ID-GBIF-Dataset="7224227d-a792-4a1b-9868-d1266571d2d8" ID-ISSN="1175-5326" ID-Zenodo-Dep="241687" ID-ZooBank="6CCC3C5B-0DF1-4677-B9E9-BD49941921D4" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461008353189" checkinUser="plazi" docAuthor="Teynié, Alexandre, David, Patrick, Lottier, Anne, Le, Minh Duc, Vidal, Nicolas & Nguyen, Truong Quang" docDate="2015" docId="038BC44BFFFCFFB9FF4442EAFF7FFA0E" docLanguage="en" docName="zt03926p540.pdf" docOrigin="Zootaxa 3926 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Parafimbrios Teynié, David, Lottier, Le, Vidal & Nguyen, 2015, gen. nov." docType="treatment" docVersion="8" lastPageNumber="527" masterDocId="FFB2BC33FFF8FFBDFFD34315FFC9FF94" masterDocTitle="A new genus and species of xenodermatid snake (Squamata: Caenophidia: Xenodermatidae) from northern Lao People’s Democratic Republic" masterLastPageNumber="540" masterPageNumber="523" pageNumber="527" updateTime="1698566107079" updateUser="plazi">
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<mods:title id="4346392F0A62B2D8E9F56A8A5E800F4E">A new genus and species of xenodermatid snake (Squamata: Caenophidia: Xenodermatidae) from northern Lao People’s Democratic Republic</mods:title>
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<mods:name id="45511689F747FE48A86FF4EEFBE1468B" type="personal">
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<mods:namePart id="CF2B41F8351406EEA4A8E8D85C96BE7C">Teynié, Alexandre</mods:namePart>
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<mods:roleTerm id="8F6370406F20A225D5BB5559AFE01025">Author</mods:roleTerm>
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<mods:namePart id="688182D0B6ED0BF0E3420523CDD20755">David, Patrick</mods:namePart>
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<mods:name id="91BB39B6E7821674436B309BE9761ADB" type="personal">
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<mods:roleTerm id="8E8A97F2AD4A41B95B3F21FDE3B7FA1F">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="E6C7A7D1549D4116BDAF22A42EBBEE60">Lottier, Anne</mods:namePart>
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</mods:name>
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<mods:name id="D8EBA8D848313905EA943734129793E4" type="personal">
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<mods:roleTerm id="67E1E45CAF5E72EB9D120B7ADD9F99E9">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="E141FA89F265BE6A902D331FBF6AD198">Le, Minh Duc</mods:namePart>
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</mods:name>
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<mods:namePart id="22EEF89B4316514183D7D5A3E81D9A04">Vidal, Nicolas</mods:namePart>
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</mods:role>
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<mods:namePart id="5B6AE3627C0AB9AB242FFE4DF2D61E42">Nguyen, Truong Quang</mods:namePart>
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<mods:title id="8CDCC8938E1E88E181A7A70A99F23620">Zootaxa</mods:title>
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<mods:part id="1881058006053D644863B5953DDC23E1">
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<mods:date id="586DED782C75EA3ED9EB78B873D1173B">2015</mods:date>
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<mods:detail id="832D2FD7ABFC89A17279F1F2D4316D14" type="volume">
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<mods:number id="B6DD83DF63AB20BE612D0959EBC5FF28">3926</mods:number>
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</mods:detail>
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<mods:detail id="3D4A70F97F30993989E7B15E2404F291" type="issue">
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<mods:number id="26E44F2B5D24C029474B669F4C223366">4</mods:number>
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<mods:start id="3E663993A382542BF8B4E10D0332D235">523</mods:start>
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<mods:classification id="A3071191ECFF9328C8FF1387A3A3B6E6">journal article</mods:classification>
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<mods:identifier id="2CFED5BAB031D0ABA83F463BF317133B" type="DOI">10.11646/zootaxa.3926.4.4</mods:identifier>
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<mods:identifier id="92B5CA3D78E32893B8BD380E15476A05" type="GBIF-Dataset">7224227d-a792-4a1b-9868-d1266571d2d8</mods:identifier>
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<mods:identifier id="302682D582C6682FAF6E89767D70C595" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="E124BF84ED0BFFC38B775A9BB3219631" type="Zenodo-Dep">241687</mods:identifier>
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<treatment id="038BC44BFFFCFFB9FF4442EAFF7FFA0E" ID-DOI="http://doi.org/10.5281/zenodo.5687312" ID-GBIF-Taxon="119599414" ID-Zenodo-Dep="5687312" LSID="urn:lsid:plazi:treatment:038BC44BFFFCFFB9FF4442EAFF7FFA0E" httpUri="http://treatment.plazi.org/id/038BC44BFFFCFFB9FF4442EAFF7FFA0E" lastPageNumber="527" pageId="4" pageNumber="527">
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<subSubSection id="C33826D6FFFCFFB9FF4442EAFE62FD8E" box="[151,427,511,538]" pageId="4" pageNumber="527" type="nomenclature">
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<paragraph id="8B9D755DFFFCFFB9FF4442EAFE62FD8E" blockId="4.[151,427,511,538]" box="[151,427,511,538]" pageId="4" pageNumber="527">
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<heading id="D0D5C231FFFCFFB9FF4442EAFE62FD8E" bold="true" box="[151,427,511,538]" fontSize="11" level="1" pageId="4" pageNumber="527" reason="1">
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<emphasis id="B956A94FFFFCFFB9FF4442EAFE62FD8E" bold="true" box="[151,427,511,538]" pageId="4" pageNumber="527">
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<taxonomicName id="4C220EDEFFFCFFB9FF4442EAFEFEFD8D" box="[151,311,511,537]" class="Reptilia" family="Xenodermatidae" genus="Parafimbrios" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="527" phylum="Chordata" rank="genus" status="gen. nov.">
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||
<emphasis id="B956A94FFFFCFFB9FF4442EAFEFEFD8D" bold="true" box="[151,311,511,537]" italics="true" pageId="4" pageNumber="527">Parafimbrios</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A2651434FFFCFFB9FEEE4115FE62FD8E" box="[317,427,512,538]" pageId="4" pageNumber="527" rank="genus">gen. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C33826D6FFFCFFB9FF444153FC89FDCB" box="[151,832,582,607]" pageId="4" pageNumber="527" type="materials_examined">
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<paragraph id="8B9D755DFFFCFFB9FF444153FC89FDCB" blockId="4.[151,1436,582,1434]" box="[151,832,582,607]" pageId="4" pageNumber="527">
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<emphasis id="B956A94FFFFCFFB9FF444153FEFBFDCB" bold="true" box="[151,306,582,607]" pageId="4" pageNumber="527">
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<typeStatus id="5499CBFFFFFCFFB9FF444153FF1BFDCB" box="[151,210,582,607]" pageId="4" pageNumber="527">Type</typeStatus>
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species.
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</emphasis>
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<taxonomicName id="4C220EDEFFFCFFB9FEEB4153FE30FDCB" box="[312,505,582,607]" class="Reptilia" family="Xenodermatidae" genus="Parafimbrios" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="527" phylum="Chordata" rank="species" species="lao" status="sp. nov.">
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<emphasis id="B956A94FFFFCFFB9FEEB4153FE30FDCB" box="[312,505,582,607]" italics="true" pageId="4" pageNumber="527">Parafimbrios lao</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A2651434FFFCFFB9FDD34153FDBBFDCB" box="[512,626,582,607]" pageId="4" pageNumber="527" rank="species">
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<emphasis id="B956A94FFFFCFFB9FDD34153FDBBFDCB" bold="true" box="[512,626,582,607]" pageId="4" pageNumber="527">spec. nov.</emphasis>
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</taxonomicNameLabel>
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, herein described.
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</paragraph>
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</subSubSection>
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<subSubSection id="C33826D6FFFCFFB9FF14417FFBC4FBEF" pageId="4" pageNumber="527" type="diagnosis">
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<paragraph id="8B9D755DFFFCFFB9FF14417FFC1DFC37" blockId="4.[151,1436,582,1434]" pageId="4" pageNumber="527">
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<emphasis id="B956A94FFFFCFFB9FF14417FFE88FD17" bold="true" box="[199,321,618,643]" pageId="4" pageNumber="527">Diagnosis.</emphasis>
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A genus of Caenophidia, family
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<taxonomicName id="4C220EDEFFFCFFB9FD3B417FFC6DFD17" box="[744,932,618,643]" class="Reptilia" family="Xenodermatidae" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="527" phylum="Chordata" rank="family">Xenodermatidae</taxonomicName>
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, characterized by (1) maxillary teeth 27, progressively slightly enlarged posteriorly; (2) head not distinct from neck; (3) dorsal scales small, cycloid or subrectangular, with the correspondence of two dorsal scale rows per ventral plate throughout the body; (4) two consecutive scales of the first dorsal scale row enlarged; (5) rostral separated from internasals by a fold of skin, larger on the sides than above the middle of the rostral; (6) nostril piercing the anterior part of a large, concave nasal; (7) mental and both anterior supralabials and infralabials with strong, raised, everted edges; (8) eye with vertically elliptic pupil; (9) loreal single, large, extending from nasal to eye; (10) chin shields in a single pair, enlarged; (11) body moderately elongate; (12) ventrals large, rounded; (13) subcaudals undivided; (14) cloacal plate entire; and (15) hemipenis short, forked, distal half strongly spinose.
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</paragraph>
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<paragraph id="8B9D755DFFFCFFB9FF1440BBFBC4FBEF" blockId="4.[151,1436,582,1434]" pageId="4" pageNumber="527">
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Although
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<taxonomicName id="4C220EDEFFFCFFB9FE9340BBFE1CFC53" box="[320,469,942,967]" class="Reptilia" family="Xenodermatidae" genus="Parafimbrios" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="527" phylum="Chordata" rank="genus" status="gen. nov.">
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<emphasis id="B956A94FFFFCFFB9FE9340BBFE1CFC53" box="[320,469,942,967]" italics="true" pageId="4" pageNumber="527">Parafimbrios</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A2651434FFFCFFB9FE3640BBFD9FFC53" box="[485,598,942,967]" pageId="4" pageNumber="527" rank="genus">
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<emphasis id="B956A94FFFFCFFB9FE3640BBFD9FFC53" bold="true" box="[485,598,942,967]" pageId="4" pageNumber="527">gen. nov.</emphasis>
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</taxonomicNameLabel>
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shares morphological similarities with other members of the family
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<taxonomicName id="4C220EDEFFFCFFB9FF4440C7FE9AFC7F" box="[151,339,978,1003]" class="Reptilia" family="Xenodermatidae" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="527" phylum="Chordata" rank="family">Xenodermatidae</taxonomicName>
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, it is morphologically most similar to
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<taxonomicName id="4C220EDEFFFCFFB9FCD240C7FCA1FC7F" box="[769,872,978,1003]" class="Reptilia" family="Xenodermatidae" genus="Fimbrios" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="527" phylum="Chordata" rank="genus">
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<emphasis id="B956A94FFFFCFFB9FCD240C7FCA1FC7F" box="[769,872,978,1003]" italics="true" pageId="4" pageNumber="527">Fimbrios</emphasis>
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</taxonomicName>
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.
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<taxonomicName id="4C220EDEFFFCFFB9FCA540C7FBC5FC7F" box="[886,1036,978,1003]" class="Reptilia" family="Xenodermatidae" genus="Parafimbrios" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="527" phylum="Chordata" rank="genus" status="gen. nov.">
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<emphasis id="B956A94FFFFCFFB9FCA540C7FBC5FC7F" box="[886,1036,978,1003]" italics="true" pageId="4" pageNumber="527">Parafimbrios</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A2651434FFFCFFB9FBC040C7FBB4FC7F" box="[1043,1149,978,1003]" pageId="4" pageNumber="527" rank="genus">
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<emphasis id="B956A94FFFFCFFB9FBC040C7FBB4FC7F" bold="true" box="[1043,1149,978,1003]" pageId="4" pageNumber="527">gen. nov.</emphasis>
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</taxonomicNameLabel>
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differs from
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<taxonomicName id="4C220EDEFFFCFFB9FAC140C7FAB0FC7F" box="[1298,1401,978,1003]" class="Reptilia" family="Xenodermatidae" genus="Fimbrios" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="527" phylum="Chordata" rank="genus">
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<emphasis id="B956A94FFFFCFFB9FAC140C7FAB0FC7F" box="[1298,1401,978,1003]" italics="true" pageId="4" pageNumber="527">Fimbrios</emphasis>
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</taxonomicName>
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by a combination of the five characters listed above, the most important being (1) the correspondence of two consecutive dorsal scale rows per ventral plate throughout the body, (2) fewer dorsal scale rows, and (3) posterior teeth progressively and slightly enlarged (vs. equal). Additional comparisons between the new genus and species and members of the genus
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<taxonomicName id="4C220EDEFFFCFFB9FE174777FDE3FBEF" box="[452,554,1122,1147]" class="Reptilia" family="Xenodermatidae" genus="Fimbrios" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="527" phylum="Chordata" rank="genus">
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<emphasis id="B956A94FFFFCFFB9FE174777FDE3FBEF" box="[452,554,1122,1147]" italics="true" pageId="4" pageNumber="527">Fimbrios</emphasis>
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</taxonomicName>
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are given below in the species description.
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</paragraph>
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</subSubSection>
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<subSubSection id="C33826D6FFFCFFB9FF144793FAB1FB0B" box="[199,1400,1158,1183]" pageId="4" pageNumber="527" type="distribution">
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<paragraph id="8B9D755DFFFCFFB9FF144793FAB1FB0B" blockId="4.[151,1436,582,1434]" box="[199,1400,1158,1183]" pageId="4" pageNumber="527">
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||
<emphasis id="B956A94FFFFCFFB9FF144793FEA9FB0B" bold="true" box="[199,352,1158,1183]" pageId="4" pageNumber="527">Distribution.</emphasis>
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<taxonomicName id="4C220EDEFFFCFFB9FEB44792FE68FB0A" box="[359,417,1159,1182]" class="Reptilia" family="Xenodermatidae" genus="Parafimbrios" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="523" phylum="Chordata" rank="species" species="laos" status="sp. nov.">
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<collectingCountry id="F33535CDFFFCFFB9FEB44792FE68FB0A" box="[359,417,1159,1182]" name="Laos" pageId="4" pageNumber="527">Laos</collectingCountry>
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</taxonomicName>
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. Currently recorded from karst formations in Louangphabang and Houaphan provinces.
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</paragraph>
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</subSubSection>
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<subSubSection id="C33826D6FFFCFFB9FF1447BFFF7FFA0E" pageId="4" pageNumber="527" type="etymology">
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<paragraph id="8B9D755DFFFCFFB9FF1447BFFB6CFAC7" blockId="4.[151,1436,582,1434]" pageId="4" pageNumber="527">
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<emphasis id="B956A94FFFFCFFB9FF1447BFFE87FB57" bold="true" box="[199,334,1194,1219]" pageId="4" pageNumber="527">Etymology.</emphasis>
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The generic nomen
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<taxonomicName id="4C220EDEFFFCFFB9FDEB47BFFD04FB57" box="[568,717,1194,1219]" class="Reptilia" family="Xenodermatidae" genus="Parafimbrios" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="527" phylum="Chordata" rank="genus">
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<emphasis id="B956A94FFFFCFFB9FDEB47BFFD04FB57" box="[568,717,1194,1219]" italics="true" pageId="4" pageNumber="527">Parafimbrios</emphasis>
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</taxonomicName>
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is composed of the modern Latin generic nomen
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<taxonomicName id="4C220EDEFFFCFFB9FAD747BFFAA2FB57" box="[1284,1387,1194,1219]" class="Reptilia" family="Xenodermatidae" genus="Fimbrios" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="527" phylum="Chordata" rank="genus">
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<emphasis id="B956A94FFFFCFFB9FAD747BFFAA2FB57" box="[1284,1387,1194,1219]" italics="true" pageId="4" pageNumber="527">Fimbrios</emphasis>
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</taxonomicName>
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and the Latin adjective
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<emphasis id="B956A94FFFFCFFB9FEA247DAFE51FB72" box="[369,408,1231,1254]" italics="true" pageId="4" pageNumber="527">par</emphasis>
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(
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<emphasis id="B956A94FFFFCFFB9FE7A47DAFE2AFB72" box="[425,483,1231,1254]" italics="true" pageId="4" pageNumber="527">paris</emphasis>
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), meaning, among other possibilities, “similar to”. The nomen
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<taxonomicName id="4C220EDEFFFCFFB9FB7D47DBFADCFB73" box="[1198,1301,1230,1255]" class="Reptilia" family="Xenodermatidae" genus="Fimbrios" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="527" phylum="Chordata" rank="genus">
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<emphasis id="B956A94FFFFCFFB9FB7D47DBFADCFB73" box="[1198,1301,1230,1255]" italics="true" pageId="4" pageNumber="527">Fimbrios</emphasis>
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</taxonomicName>
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itself stems from the Latin noun
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<emphasis id="B956A94FFFFCFFB9FEAC47E7FE1BFA9F" box="[383,466,1266,1291]" italics="true" pageId="4" pageNumber="527">fimbria</emphasis>
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(-
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<emphasis id="B956A94FFFFCFFB9FE3847E1FDCCFA9F" box="[491,517,1268,1291]" italics="true" pageId="4" pageNumber="527">ae</emphasis>
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), meaning “a fringe”, by allusion to the edges of labial scales producing a kind of fringe around the snout. So, the generic nomen
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<taxonomicName id="4C220EDEFFFCFFB9FD7A4603FCF7FABB" box="[681,830,1302,1327]" class="Reptilia" family="Xenodermatidae" genus="Parafimbrios" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="527" phylum="Chordata" rank="genus">
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<emphasis id="B956A94FFFFCFFB9FD7A4603FCF7FABB" box="[681,830,1302,1327]" italics="true" pageId="4" pageNumber="527">Parafimbrios</emphasis>
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</taxonomicName>
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refers to one of the diagnostic characters of
|
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<taxonomicName id="4C220EDEFFFCFFB9FAFD4603FA5CFABB" box="[1326,1429,1302,1327]" class="Reptilia" family="Xenodermatidae" genus="Fimbrios" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="527" phylum="Chordata" rank="genus">
|
||
<emphasis id="B956A94FFFFCFFB9FAFD4603FA5CFABB" box="[1326,1429,1302,1327]" italics="true" pageId="4" pageNumber="527">Fimbrios</emphasis>
|
||
</taxonomicName>
|
||
, to which
|
||
<taxonomicName id="4C220EDEFFFCFFB9FF2C462FFE5DFAC7" box="[255,404,1338,1363]" class="Reptilia" family="Xenodermatidae" genus="Parafimbrios" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="527" phylum="Chordata" rank="genus" status="gen. nov.">
|
||
<emphasis id="B956A94FFFFCFFB9FF2C462FFE5DFAC7" box="[255,404,1338,1363]" italics="true" pageId="4" pageNumber="527">Parafimbrios</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2651434FFFCFFB9FE48462FFDCCFAC7" box="[411,517,1338,1363]" pageId="4" pageNumber="527" rank="genus">
|
||
<emphasis id="B956A94FFFFCFFB9FE48462FFDCCFAC7" bold="true" box="[411,517,1338,1363]" pageId="4" pageNumber="527">gen. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
is morphologically quite similar but taxonomically distinct.
|
||
</paragraph>
|
||
<paragraph id="8B9D755DFFFCFFB9FF14464BFF7FFA0E" blockId="4.[151,1436,582,1434]" pageId="4" pageNumber="527">
|
||
<emphasis id="B956A94FFFFCFFB9FF14464BFEF0FAE3" bold="true" box="[199,313,1374,1399]" pageId="4" pageNumber="527">Contents.</emphasis>
|
||
The new genus currently contains a single species,
|
||
<taxonomicName id="4C220EDEFFFCFFB9FC52464BFB8BFAE3" box="[897,1090,1374,1399]" class="Reptilia" family="Xenodermatidae" genus="Parafimbrios" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="527" phylum="Chordata" rank="species" species="lao" status="sp. nov.">
|
||
<emphasis id="B956A94FFFFCFFB9FC52464BFB8BFAE3" box="[897,1090,1374,1399]" italics="true" pageId="4" pageNumber="527">Parafimbrios lao</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2651434FFFCFFB9FB9A464BFB75FAE3" box="[1097,1212,1374,1399]" pageId="4" pageNumber="527" rank="species">
|
||
<emphasis id="B956A94FFFFCFFB9FB9A464BFB75FAE3" bold="true" box="[1097,1212,1374,1399]" pageId="4" pageNumber="527">spec. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, which we describe as:
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |