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<document id="3DAB5CAACC33F630198016DEF1EEC9D2" ID-CLB-Dataset="183563" ID-DOI="10.11646/zootaxa.5205.4.3" ID-GBIF-Dataset="606deb86-203b-4450-aaba-814fa4835e33" ID-ISSN="1175-5326" ID-Zenodo-Dep="7307035" ID-ZooBank="A4731737-9899-45BE-B342-E4A6F632E756" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1667981723880" checkinUser="plazi" docAuthor="Eb. Bonaldo, Ramírez, Martín J., Om. Labarque, Shimano, Yulie, Silva-Junior, Cláudio J. &amp; Haddad, Charles R." docDate="2022" docId="03FAED38F02FFF8956CEFADBF4ADFDFB" docLanguage="en" docName="zootaxa.5205.4.3.pdf" docOrigin="Zootaxa 5205 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Carteronius gentilis Eb.Bonaldo &amp; Ramírez &amp; Om.Labarque &amp; Shimano &amp; Silva-Junior &amp; Haddad 2022, comb. nov." docType="treatment" docVersion="7" lastPageNumber="367" masterDocId="FFC39540F03AFF915659FFA5F067FFE2" masterDocTitle="Switching identities: a revision of the Afrotropical spider genus Carteronius Simon 1897 (Araneae, Corinnidae), senior synonym of Mandaneta Strand, 1932, with a new genus of the Pronophaea group" masterLastPageNumber="373" masterPageNumber="343" pageNumber="364" updateTime="1699411079873" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="E9BE962BF9FBD15F0D306B00850D0D6E">Switching identities: a revision of the Afrotropical spider genus Carteronius Simon 1897 (Araneae, Corinnidae), senior synonym of Mandaneta Strand, 1932, with a new genus of the Pronophaea group</mods:title>
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<mods:namePart id="2D11A7C649973F4CC7D99215E6B7FAEB">Eb. Bonaldo</mods:namePart>
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<mods:namePart id="25509A14315AF76D3EBCBCB482151AC4">Shimano, Yulie</mods:namePart>
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<mods:affiliation id="3651855BE24DD40E9390B7A13A84DABC">Instituto Nacional de Pesquisa do Pantanal, Campus Avançado do Museu Paraense Emílio Goeldi, Cuiabá, 78060 - 900 Mato Grosso, Brazil. shimano. yulie @ gmail. com; https: // orcid. org / 0000 - 0003 - 2931 - 4719</mods:affiliation>
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<mods:namePart id="C18443BE7862EE0EABE71E81849C2CE8">Silva-Junior, Cláudio J.</mods:namePart>
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<mods:affiliation id="335E45993009D7E6AFBB11119C0937E2">Museu Paraense Emílio Goeldi-MPEG, Coordenação de Zoologia, Laboratório de Aracnologia, Avenida Perimetral, 1901, Terra Firme, CEP: 66077 - 830, Belém, Pará, Brazil &amp; Programa de Pós-Graduação em Zoologia-Universidade Federal do Pará-Museu Paraense Emílio Goeldi claudiojr. uepa @ gmail. com; https: // orcid. org / 0000 - 0003 - 1300 - 8872</mods:affiliation>
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<mods:namePart id="C904104537463A99127E54CD8EA6CBF8">Haddad, Charles R.</mods:namePart>
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<mods:affiliation id="4E852F2C4F6108A949A27F0F0D4A8751">Department of Zoology &amp; Entomology, University of the Free State, P. O. Box 339, Bloemfontein 9300, South Africa haddadCR @ ufs. ac. za; https: // orcid. org / 0000 - 0002 - 2317 - 7760</mods:affiliation>
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<heading id="D0A4EB42F02FFF8456CEFADBF2AEFA7B" bold="true" box="[151,713,1406,1433]" fontSize="11" level="1" pageId="21" pageNumber="364" reason="1">
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<emphasis id="B927803CF02FFF8456CEFADBF1EFFA7B" bold="true" box="[151,392,1406,1433]" italics="true" pageId="21" pageNumber="364">Carteronius gentilis</emphasis>
(
<bibRefCitation id="EFC221DFF02FFF8457CEFADBF254FA7A" author="Simon, E." box="[407,563,1406,1432]" pageId="21" pageNumber="364" pagination="335 - 449" refId="ref18798" refString="Simon, E. (1909) Arachnides recueillis par L. Fea sur la cote occidentale d'Afrique. 2 e partie. Annali del Museo Civico di Storia Naturale di Genova, 44, 335 - 449." type="journal article" year="1909">Simon, 1909</bibRefCitation>
)
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<figureCitation id="136840ABF02FFF8456CEFA07F08BFA5F" box="[151,236,1442,1469]" captionStart="FIGURE 18" captionStartId="22.[152,255,1304,1328]" captionTargetBox="[151,1435,181,1281]" captionTargetId="figure-19@22.[151,1435,181,1281]" captionTargetPageId="22" captionText="FIGURE 18. Carteronius gentilis (Simon, 1909) comb. nov.: A male holotype habitus, dorsal view; B left palp, prolateral view; C same, ventral view; D same, retrolateral view; E original label; F female abdomen, dorsal view; B female carapace, dorsal view. Scale: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7307090" httpUri="https://zenodo.org/record/7307090/files/figure.png" pageId="21" pageNumber="364">Figs 18</figureCitation>
, 
<figureCitation id="136840ABF02FFF8456A1FA06F170FA5F" box="[248,279,1443,1469]" captionStart="FIGURE 19" captionStartId="23.[152,255,1851,1875]" captionTargetBox="[197,1385,188,1822]" captionTargetId="figure-19@23.[192,1395,181,1828]" captionTargetPageId="23" captionText="FIGURE 19. Carteronius gentilis (Simon, 1909) comb. nov.: A male palp, ventral view; B same, retrolateral view; C epigynum, ventral view; D same, dorsal view. Abbreviations: E, embolus; C, conductor; MA, median apophysis; VL, ventral lobe; ML, medial lobe; DL, dorsal lobe; AS, spur; CDv, ventral sector of the copulatory duct; CDd, dorsal sector of the copulatory duct; ST2 secondary spermatheca; ST1 primary spermatheca. Scale: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7307096" httpUri="https://zenodo.org/record/7307096/files/figure.png" pageId="21" pageNumber="364">19</figureCitation>
; 
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<paragraph id="8BEC5C2EF02FFF8456CEFA4DF502FA1D" blockId="21.[151,1381,1511,1535]" box="[151,1381,1511,1535]" pageId="21" pageNumber="364">
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<treatmentCitation id="0AF27A3FF02FFF8456CEFA4DF26CFA1D" author="Simon, E." box="[151,523,1511,1535]" page="382" pageId="21" pageNumber="364" year="1909">
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<emphasis id="B927803CF02FFF8456CEFA4DF137FA1D" box="[151,336,1511,1535]" italics="true" pageId="21" pageNumber="364">Procopius gentilis</emphasis>
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<bibRefCitation id="EFC221DFF02FFF84570FFA42F26CFA1D" author="Simon, E." box="[342,523,1511,1535]" pageId="21" pageNumber="364" pagination="335 - 449" refId="ref18798" refString="Simon, E. (1909) Arachnides recueillis par L. Fea sur la cote occidentale d'Afrique. 2 e partie. Annali del Museo Civico di Storia Naturale di Genova, 44, 335 - 449." type="journal article" year="1909">Simon, 1909: 382</bibRefCitation>
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(
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from “ 
<collectingCountry id="F3441CBEF02FFF84548BFA42F302FA1C" box="[722,869,1511,1535]" name="Equatorial Guinea" pageId="21" pageNumber="364">Fernando Poo</collectingCountry>
: Basilé”, leg. 
<collectorName id="26A639F8F02FFF8455ABFA42F452FA1C" box="[1010,1077,1511,1534]" pageId="21" pageNumber="364">L. Fea</collectorName>
, 
<specimenCode id="DBF5F455F02FFF845219FA42F4BDFA1D" box="[1088,1242,1511,1535]" collectionCode="MNHN" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="21" pageNumber="364">MNHN 22.254</specimenCode>
—examined).
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<emphasis id="B927803CF02FFF8456CEF988F0B1F9A5" bold="true" box="[151,214,1581,1607]" pageId="21" pageNumber="364">Note.</emphasis>
The male 
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, from Bioko Island, 
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, is here tentatively associated with a female from continental Africa (
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). Bioko is a continental-shelf island that supports relatively low levels of endemism of angiosperms, bats, birds, reptiles, and amphibians when compared to oceanic islands of the Gulf of 
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(
<collectingCountry id="F3441CBEF02FFF8456C6F93CF166F951" box="[159,257,1689,1715]" name="Sao Tome and Principe" pageId="21" pageNumber="364">Príncipe</collectingCountry>
, 
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and 
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) (
<bibRefCitation id="EFC221DFF02FFF845465F93CF2A5F951" author="Jones, P. J." box="[572,706,1689,1715]" pageId="21" pageNumber="364" pagination="772 - 784" refId="ref18173" refString="Jones, P. J. (2004) Biodiversity in the Gulf of Guinea: an overview. Biodiversity &amp; Conservation, 3, 772 - 784. https: // doi. org / 10.1007 / BF 00129657" type="journal article" year="2004">Jones 2004</bibRefCitation>
). This island was separated from mainland Africa by the rise in sea level at the end of the last glacial period, approximately 10 000 years ago (
<bibRefCitation id="EFC221DFF02FFF84524EF918F52EF935" author="Schabetsberger, R. &amp; Drozdowski, G. &amp; Drozdowski, I. &amp; Jersabek, C. D. &amp; Rott, E." box="[1047,1353,1725,1751]" pageId="21" pageNumber="364" pagination="79 - 90" refId="ref18616" refString="Schabetsberger, R., Drozdowski, G., Drozdowski, I., Jersabek, C. D. &amp; Rott, E. (2004) Limmnological aspects of two tropical crater lakes (Lago Biao and Lago Loreto) on the island of Bioko (Equatorial Guinea). Hydrobiologia, 524, 79 - 90. https: // doi. org / 10.1023 / B: HYDR. 0000036121.07007. e 8" type="journal article" year="2004">
Schabetsberger 
<emphasis id="B927803CF02FFF845292F91BF562F935" box="[1227,1285,1725,1751]" italics="true" pageId="21" pageNumber="364">et al.</emphasis>
2004
</bibRefCitation>
; PérezPérez &amp; Yu 2021). Both the Bioko and 
<collectingCountry id="F3441CBEF02FFF845438F944F2B1F919" box="[609,726,1761,1787]" name="Cameroon" pageId="21" pageNumber="364">Cameroon</collectingCountry>
specimens share a strikingly similar dorsal abdominal pattern (
<figureCitation id="136840ABF02FFF8456C6F8A0F142F8FD" box="[159,293,1797,1823]" captionStart="FIGURE 18" captionStartId="22.[152,255,1304,1328]" captionTargetBox="[151,1435,181,1281]" captionTargetId="figure-19@22.[151,1435,181,1281]" captionTargetPageId="22" captionText="FIGURE 18. Carteronius gentilis (Simon, 1909) comb. nov.: A male holotype habitus, dorsal view; B left palp, prolateral view; C same, ventral view; D same, retrolateral view; E original label; F female abdomen, dorsal view; B female carapace, dorsal view. Scale: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7307090" httpUri="https://zenodo.org/record/7307090/files/figure.png" pageId="21" pageNumber="364">Fig. 18A, F</figureCitation>
), as well as similarities in leg spination. Nevertheless, this association must be tested when additional samples come to light. From a strictly nomenclatural point of view, proposing this doubtful association is preferable to making available a possibly unnecessary specific name.
</paragraph>
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<paragraph id="8BEC5C2EF02FFF84569EF8D4F210F815" blockId="21.[151,1437,1581,2040]" pageId="21" pageNumber="364">
<emphasis id="B927803CF02FFF84569EF8D4F125F869" bold="true" box="[199,322,1905,1931]" pageId="21" pageNumber="364">Diagnosis.</emphasis>
Males of 
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<emphasis id="B927803CF02FFF8457EEF8D4F24CF869" box="[439,555,1905,1931]" italics="true" pageId="21" pageNumber="364">C. gentilis</emphasis>
</taxonomicName>
<emphasis id="B927803CF02FFF84546DF8D4F2D1F869" bold="true" box="[564,694,1905,1931]" pageId="21" pageNumber="364">
<taxonomicNameLabel id="A2143D47F02FFF84546DF8D4F2D1F869" box="[564,694,1905,1931]" pageId="21" pageNumber="364" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
differ from all other 
<taxonomicName id="4C5327ADF02FFF8455F3F8D4F4EDF869" box="[938,1162,1905,1931]" pageId="21" pageNumber="364">
<emphasis id="B927803CF02FFF8455F3F8D4F457F869" box="[938,1072,1905,1931]" italics="true" pageId="21" pageNumber="364">Carteronius</emphasis>
species
</taxonomicName>
in having a bifid dorsal lobe on the RTA (
<figureCitation id="136840ABF02FFF845702F830F186F84D" box="[347,481,1941,1967]" captionStart="FIGURE 19" captionStartId="23.[152,255,1851,1875]" captionTargetBox="[197,1385,188,1822]" captionTargetId="figure-19@23.[192,1395,181,1828]" captionTargetPageId="23" captionText="FIGURE 19. Carteronius gentilis (Simon, 1909) comb. nov.: A male palp, ventral view; B same, retrolateral view; C epigynum, ventral view; D same, dorsal view. Abbreviations: E, embolus; C, conductor; MA, median apophysis; VL, ventral lobe; ML, medial lobe; DL, dorsal lobe; AS, spur; CDv, ventral sector of the copulatory duct; CDd, dorsal sector of the copulatory duct; ST2 secondary spermatheca; ST1 primary spermatheca. Scale: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7307096" httpUri="https://zenodo.org/record/7307096/files/figure.png" pageId="21" pageNumber="364">Fig. 19A, B</figureCitation>
). Females resemble those of 
<taxonomicName id="4C5327ADF02FFF84557DF830F3EAF84D" authority="Eb.Bonaldo &amp; Ramírez &amp; Om.Labarque &amp; Shimano &amp; Silva-Junior &amp; Haddad, 2022" authorityName="Eb.Bonaldo &amp; Ramírez &amp; Om.Labarque &amp; Shimano &amp; Silva-Junior &amp; Haddad" authorityYear="2022" box="[804,909,1941,1967]" class="Arachnida" family="Clubionidae" genus="Carteronius" kingdom="Animalia" order="Araneae" pageId="21" pageNumber="364" phylum="Arthropoda" rank="species" species="myene" status="sp. nov.">
<emphasis id="B927803CF02FFF84557DF830F3EAF84D" box="[804,909,1941,1967]" italics="true" pageId="21" pageNumber="364">C. myene</emphasis>
</taxonomicName>
<emphasis id="B927803CF02FFF8455CAF830F38BF84D" bold="true" box="[915,1004,1941,1967]" pageId="21" pageNumber="364">
<taxonomicNameLabel id="A2143D47F02FFF8455CAF830F38BF84D" box="[915,1004,1941,1967]" pageId="21" pageNumber="364" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
by the slightly curved transverse ridge (
<figureCitation id="136840ABF02FFF8456C6F81CF16AF831" box="[159,269,1977,2003]" captionStart="FIGURE 14" captionStartId="17.[152,255,1382,1406]" captionTargetBox="[169,1421,196,1350]" captionTargetId="figure-19@17.[159,1428,181,1358]" captionTargetPageId="17" captionText="FIGURE 14. Carteronius myene sp. nov.: A epigynum, ventral view; B same dorsal view; Carteronius teke sp. nov.: C epigynum, ventral view (arrow: epigynal transversal ridge); D same, dorsal view. Abbreviations: CO, copulatory opening; CDv, ventral sector of the copulatory duct; CDd, dorsal sector of the copulatory duct; ST2, secondary spermatheca; ST1, primary spermatheca. Scale: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7307077" httpUri="https://zenodo.org/record/7307077/files/figure.png" pageId="21" pageNumber="364">Figs 14A</figureCitation>
, 
<figureCitation id="136840ABF02FFF845742F81CF12AF831" box="[283,333,1977,2003]" captionStart="FIGURE 19" captionStartId="23.[152,255,1851,1875]" captionTargetBox="[197,1385,188,1822]" captionTargetId="figure-19@23.[192,1395,181,1828]" captionTargetPageId="23" captionText="FIGURE 19. Carteronius gentilis (Simon, 1909) comb. nov.: A male palp, ventral view; B same, retrolateral view; C epigynum, ventral view; D same, dorsal view. Abbreviations: E, embolus; C, conductor; MA, median apophysis; VL, ventral lobe; ML, medial lobe; DL, dorsal lobe; AS, spur; CDv, ventral sector of the copulatory duct; CDd, dorsal sector of the copulatory duct; ST2 secondary spermatheca; ST1 primary spermatheca. Scale: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7307096" httpUri="https://zenodo.org/record/7307096/files/figure.png" pageId="21" pageNumber="364">19C</figureCitation>
), but differ by the relatively small posterior sector and the extremely narrow copulatory duct in relation to the spermathecae (
<figureCitation id="136840ABF02FFF8457B8F878F20DF815" box="[481,618,2013,2039]" captionStart="FIGURE 19" captionStartId="23.[152,255,1851,1875]" captionTargetBox="[197,1385,188,1822]" captionTargetId="figure-19@23.[192,1395,181,1828]" captionTargetPageId="23" captionText="FIGURE 19. Carteronius gentilis (Simon, 1909) comb. nov.: A male palp, ventral view; B same, retrolateral view; C epigynum, ventral view; D same, dorsal view. Abbreviations: E, embolus; C, conductor; MA, median apophysis; VL, ventral lobe; ML, medial lobe; DL, dorsal lobe; AS, spur; CDv, ventral sector of the copulatory duct; CDd, dorsal sector of the copulatory duct; ST2 secondary spermatheca; ST1 primary spermatheca. Scale: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7307096" httpUri="https://zenodo.org/record/7307096/files/figure.png" pageId="21" pageNumber="364">Fig. 19C, D</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="DF2C0CA6F02CFF8756C1FABDF13EFA9A" ID-DOI="http://doi.org/10.5281/zenodo.7307090" ID-Zenodo-Dep="7307090" httpUri="https://zenodo.org/record/7307090/files/figure.png" pageId="22" pageNumber="365" startId="22.[152,255,1304,1328]" targetBox="[151,1435,181,1281]" targetPageId="22">
<paragraph id="8BEC5C2EF02CFF8756C1FABDF13EFA9A" blockId="22.[151,1436,1304,1401]" pageId="22" pageNumber="365">
<emphasis id="B927803CF02CFF8756C1FABDF143FAD2" bold="true" box="[152,292,1304,1328]" pageId="22" pageNumber="365">FIGURE 18.</emphasis>
<taxonomicName id="4C5327ADF02CFF875770FABCF2EEFAD3" authority="(Simon, 1909)" authorityName="Eb.Bonaldo &amp; Ramírez &amp; Om.Labarque &amp; Shimano &amp; Silva-Junior &amp; Haddad" authorityYear="2022" baseAuthorityName="Simon" baseAuthorityYear="1909" box="[297,649,1304,1329]" class="Arachnida" family="Clubionidae" genus="Carteronius" kingdom="Animalia" order="Araneae" pageId="22" pageNumber="365" phylum="Arthropoda" rank="species" species="gentilis" status="comb. nov.">
<emphasis id="B927803CF02CFF875770FABCF194FAD2" box="[297,499,1304,1328]" italics="true" pageId="22" pageNumber="365">Carteronius gentilis</emphasis>
(
<bibRefCitation id="EFC221DFF02CFF8757A6FABDF2E5FAD3" author="Simon, E." box="[511,642,1304,1329]" pageId="22" pageNumber="365" pagination="335 - 449" refId="ref18798" refString="Simon, E. (1909) Arachnides recueillis par L. Fea sur la cote occidentale d'Afrique. 2 e partie. Annali del Museo Civico di Storia Naturale di Genova, 44, 335 - 449." type="journal article" year="1909">Simon, 1909</bibRefCitation>
)
</taxonomicName>
<emphasis id="B927803CF02CFF8754D7FABCF299FAD3" bold="true" box="[654,766,1305,1329]" pageId="22" pageNumber="365">
<taxonomicNameLabel id="A2143D47F02CFF8754D7FABCF299FAD3" box="[654,766,1305,1329]" pageId="22" pageNumber="365" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
: 
<emphasis id="B927803CF02CFF875553FABDF37CFAD2" bold="true" box="[778,795,1304,1328]" pageId="22" pageNumber="365">A</emphasis>
male holotype habitus, dorsal view; 
<emphasis id="B927803CF02CFF8752D3FABCF4FCFAD3" bold="true" box="[1162,1179,1305,1329]" pageId="22" pageNumber="365">B</emphasis>
left palp, prolateral view; 
<emphasis id="B927803CF02CFF8756CEFA99F0CEFAB6" bold="true" box="[151,169,1340,1364]" pageId="22" pageNumber="365">C</emphasis>
same, ventral view; 
<emphasis id="B927803CF02CFF8757D9FA98F1F5FAB7" bold="true" box="[384,402,1341,1365]" pageId="22" pageNumber="365">D</emphasis>
same, retrolateral view; 
<emphasis id="B927803CF02CFF8754CBFA98F2C4FAB7" bold="true" box="[658,675,1341,1365]" pageId="22" pageNumber="365">E</emphasis>
original label; 
<emphasis id="B927803CF02CFF875518FA98F337FAB7" bold="true" box="[833,848,1341,1365]" pageId="22" pageNumber="365">F</emphasis>
female abdomen, dorsal view; 
<emphasis id="B927803CF02CFF8752CAFA98F4C3FAB7" bold="true" box="[1171,1188,1341,1365]" pageId="22" pageNumber="365">B</emphasis>
female carapace, dorsal view. Scale: 1 mm.
</paragraph>
</caption>
<subSubSection id="C3490FA5F02CFF89569EFA03F56AFE17" lastPageId="24" lastPageNumber="367" pageId="22" pageNumber="365" type="description">
<paragraph id="8BEC5C2EF02CFF87569EFA03F27EF893" blockId="22.[151,1437,1446,2014]" pageId="22" pageNumber="365">
<emphasis id="B927803CF02CFF87569EFA03F1F8FA23" bold="true" box="[199,415,1446,1473]" pageId="22" pageNumber="365">Description. Male</emphasis>
(
<typeStatus id="54E8E28CF02CFF8757E9FA02F27EFA23" box="[432,537,1447,1473]" pageId="22" pageNumber="365" type="holotype">holotype</typeStatus>
). Measurements: Total length 8.03, CL 3.82, CW 3.57, AL 3.95, AW 2.80, SL 1.93, SW 1.81. Eye diameters and interdistances: AME 0.32, ALE 0.21, PLE 0.24, PME 0.26, AME-AME 0.21, AME-ALE 0.24, ALE-ALE 1.66, PME-PME 0.85, PME-PLE 0.52, PLE-PLE 2.07. Length of leg segments: I 4.13+ 1.74+4.05+3.10+1.22=14.24; II 4.02+1.70+3.95+3.20+1.26=14.13; III 2.61+1.26+2.18+2.15+1.00=9.20; IV 3.36+ 1.29+2.82+3.12+1.10=11.69. Chelicerae: promargin with three spaced teeth, median tooth largest; retromargin with two spaced teeth, subequal in size. Leg spination: femora: I 
<emphasis id="B927803CF02CFF875568F9FEF32AF997" box="[817,845,1627,1653]" italics="true" pageId="22" pageNumber="365">do</emphasis>
0-1-1 
<emphasis id="B927803CF02CFF8755CEF9FEF3CAF997" box="[919,941,1627,1653]" italics="true" pageId="22" pageNumber="365">pl</emphasis>
0-1-0, II 
<emphasis id="B927803CF02CFF87524FF9FEF455F997" box="[1046,1074,1627,1653]" italics="true" pageId="22" pageNumber="365">do</emphasis>
0-1-0, III 
<emphasis id="B927803CF02CFF8752FCF9FEF4A6F997" box="[1189,1217,1627,1653]" italics="true" pageId="22" pageNumber="365">do</emphasis>
0-1-2, IV 
<emphasis id="B927803CF02CFF87536CF9FEF536F997" box="[1333,1361,1627,1653]" italics="true" pageId="22" pageNumber="365">do</emphasis>
0-1-2; tibiae: I 
<emphasis id="B927803CF02CFF8756ADF925F16AF97B" box="[244,269,1664,1689]" italics="true" pageId="22" pageNumber="365">ve</emphasis>
2-2-2-2-2-2, II 
<emphasis id="B927803CF02CFF8757E7F925F1B0F97B" box="[446,471,1664,1689]" italics="true" pageId="22" pageNumber="365">ve</emphasis>
1 
<emphasis id="B927803CF02CFF8757B5F925F19DF97B" box="[492,506,1664,1689]" italics="true" pageId="22" pageNumber="365">p</emphasis>
-1 
<emphasis id="B927803CF02CFF875448F925F27BF97B" box="[529,540,1664,1689]" italics="true" pageId="22" pageNumber="365">r</emphasis>
-2-2-2-2, III 
<emphasis id="B927803CF02CFF8754F2F925F2A3F97B" box="[683,708,1664,1689]" italics="true" pageId="22" pageNumber="365">ve</emphasis>
0-2-2-0, IV 
<emphasis id="B927803CF02CFF875509F925F30EF97B" box="[848,873,1664,1689]" italics="true" pageId="22" pageNumber="365">ve</emphasis>
0-0-0-1 
<emphasis id="B927803CF02CFF87559DF925F3A8F97B" box="[964,975,1664,1689]" italics="true" pageId="22" pageNumber="365">r</emphasis>
; metatarsi: I 
<emphasis id="B927803CF02CFF87523AF925F41BF97B" box="[1123,1148,1664,1689]" italics="true" pageId="22" pageNumber="365">ve</emphasis>
0-2-2-0, II 
<emphasis id="B927803CF02CFF8752A6F925F57FF97B" box="[1279,1304,1664,1689]" italics="true" pageId="22" pageNumber="365">ve</emphasis>
0-2-2-0, III 
<emphasis id="B927803CF02CFF8756CEF906F0CAF95F" box="[151,173,1699,1725]" italics="true" pageId="22" pageNumber="365">pl</emphasis>
0-1-1 
<emphasis id="B927803CF02CFF8756A3F906F16AF95F" box="[250,269,1699,1725]" italics="true" pageId="22" pageNumber="365">rl</emphasis>
0-1-1 
<emphasis id="B927803CF02CFF875703F901F114F95F" box="[346,371,1700,1725]" italics="true" pageId="22" pageNumber="365">ve</emphasis>
2-2-1, IV 
<emphasis id="B927803CF02CFF8757B5F906F265F95F" box="[492,514,1699,1725]" italics="true" pageId="22" pageNumber="365">pl</emphasis>
0-1-1 
<emphasis id="B927803CF02CFF875416F906F205F95F" box="[591,610,1699,1725]" italics="true" pageId="22" pageNumber="365">rl</emphasis>
0-1-1 
<emphasis id="B927803CF02CFF8754F6F901F2AFF95F" box="[687,712,1700,1725]" italics="true" pageId="22" pageNumber="365">ve</emphasis>
2-2-0. Coloration: carapace and chelicerae dark reddish-brown. Endites, labium and sternum reddish-brown. Legs dark yellow, with femora I and II reddish-brown. Abdomen dorsally with yellowish area anteriorly and five transversal follicular bands in posterior half (
<figureCitation id="136840ABF02CFF8752F5F94EF572F8E7" box="[1196,1301,1771,1797]" captionStart="FIGURE 18" captionStartId="22.[152,255,1304,1328]" captionTargetBox="[151,1435,181,1281]" captionTargetId="figure-19@22.[151,1435,181,1281]" captionTargetPageId="22" captionText="FIGURE 18. Carteronius gentilis (Simon, 1909) comb. nov.: A male holotype habitus, dorsal view; B left palp, prolateral view; C same, ventral view; D same, retrolateral view; E original label; F female abdomen, dorsal view; B female carapace, dorsal view. Scale: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7307090" httpUri="https://zenodo.org/record/7307090/files/figure.png" pageId="22" pageNumber="365">Fig. 18A</figureCitation>
). Palp: RTA with apical spur approximately same size as ventral lobe, medial lobe short and pointed; ventral lobe sub-squared and excavated; sperm duct with long loop, retrolateral apical tegular process present, blunt; embolus long and thin, without projections (
<figureCitation id="136840ABF02CFF875726F8F2F26CF893" box="[383,523,1879,1905]" captionStart="FIGURE 19" captionStartId="23.[152,255,1851,1875]" captionTargetBox="[197,1385,188,1822]" captionTargetId="figure-19@23.[192,1395,181,1828]" captionTargetPageId="23" captionText="FIGURE 19. Carteronius gentilis (Simon, 1909) comb. nov.: A male palp, ventral view; B same, retrolateral view; C epigynum, ventral view; D same, dorsal view. Abbreviations: E, embolus; C, conductor; MA, median apophysis; VL, ventral lobe; ML, medial lobe; DL, dorsal lobe; AS, spur; CDv, ventral sector of the copulatory duct; CDd, dorsal sector of the copulatory duct; ST2 secondary spermatheca; ST1 primary spermatheca. Scale: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7307096" httpUri="https://zenodo.org/record/7307096/files/figure.png" pageId="22" pageNumber="365">Figs 19A, B</figureCitation>
).
</paragraph>
<paragraph id="8BEC5C2EF02CFF89569EF8DEF3CBFE4F" blockId="22.[151,1437,1446,2014]" lastBlockId="24.[151,1437,150,537]" lastPageId="24" lastPageNumber="367" pageId="22" pageNumber="365">
<emphasis id="B927803CF02CFF87569EF8DEF179F877" bold="true" box="[199,286,1915,1941]" pageId="22" pageNumber="365">Female</emphasis>
(MRAC 162.128). Measurements: Total length 7.92, CL 3.26, CW 2.78, AL 4.62, AW 3.26, SL 1.57, SW 1.60. Eyes diameters and interdistances: AME 0.22, ALE 0.18, PLE 0.19, PME 0.19, AME-AME 0.63, AME-ALE 0.20, ALE-ALE 1.35, PME-PME 0.32, PME-PLE 0.38, PLE-PLE 1.60. Length of leg segments I 3.13+1.33+3.01+2.30+1.10=10.87; II 3.05+1.30+2.90+2.29+1.09=10.63; III 2.23+1.00+1.89+1.73+0.87=7.72; IV 2.74+1.06+2.30+2.40+0.93=9.43. Chelicerae: promargin with three spaced teeth, median tooth largest; retromargin with two teeth, inner tooth larger. Leg spination: femora: I 
<emphasis id="B927803CF022FF895573FF7AF321FF1B" box="[810,838,223,249]" italics="true" pageId="24" pageNumber="367">do</emphasis>
0-1-0 
<emphasis id="B927803CF022FF8955C8FF7AF3C0FF1B" box="[913,935,223,249]" italics="true" pageId="24" pageNumber="367">pl</emphasis>
0-1-0, II 
<emphasis id="B927803CF022FF89524BFF7AF449FF1B" box="[1042,1070,223,249]" italics="true" pageId="24" pageNumber="367">do</emphasis>
0-1-0, III 
<emphasis id="B927803CF022FF8952FAFF7AF4D8FF1B" box="[1187,1215,223,249]" italics="true" pageId="24" pageNumber="367">do</emphasis>
0-1-2, IV 
<emphasis id="B927803CF022FF89536CFF7AF536FF1B" box="[1333,1361,223,249]" italics="true" pageId="24" pageNumber="367">do</emphasis>
0-1-2; tibiae: I 
<emphasis id="B927803CF022FF8956AEFEA6F177FEFE" box="[247,272,259,284]" italics="true" pageId="24" pageNumber="367">ve</emphasis>
2-2-2-2-2-2, II 
<emphasis id="B927803CF022FF89579CFEA6F1B9FEFE" box="[453,478,259,284]" italics="true" pageId="24" pageNumber="367">ve</emphasis>
2-2-2-2-2, III 
<emphasis id="B927803CF022FF8954DCFEA6F2F9FEFE" box="[645,670,259,284]" italics="true" pageId="24" pageNumber="367">ve</emphasis>
1 
<emphasis id="B927803CF022FF8954ECFEA6F2A4FEFE" box="[693,707,259,284]" italics="true" pageId="24" pageNumber="367">p</emphasis>
-1 
<emphasis id="B927803CF022FF895483FEA6F282FEFE" box="[730,741,259,284]" italics="true" pageId="24" pageNumber="367">r</emphasis>
-2, IV 
<emphasis id="B927803CF022FF895568FEA6F320FEFF" box="[817,839,259,285]" italics="true" pageId="24" pageNumber="367">pl</emphasis>
0-1-0-1 
<emphasis id="B927803CF022FF8955F2FEA6F3D9FEFF" box="[939,958,259,285]" italics="true" pageId="24" pageNumber="367">rl</emphasis>
0-1-0-1 
<emphasis id="B927803CF022FF89527AFEA6F45BFEFE" box="[1059,1084,259,284]" italics="true" pageId="24" pageNumber="367">ve</emphasis>
1 
<emphasis id="B927803CF022FF89520BFEA6F40EFEFE" box="[1106,1129,259,284]" italics="true" pageId="24" pageNumber="367">p-</emphasis>
1 
<emphasis id="B927803CF022FF89522EFEA6F4E2FEFE" box="[1143,1157,259,284]" italics="true" pageId="24" pageNumber="367">p</emphasis>
-0-0; metatarsi: I 
<emphasis id="B927803CF022FF895315FEA6F502FEFE" box="[1356,1381,259,284]" italics="true" pageId="24" pageNumber="367">ve</emphasis>
0-2- 2-0, II 
<emphasis id="B927803CF022FF8956BFFE82F098FEA2" box="[230,255,295,320]" italics="true" pageId="24" pageNumber="367">ve</emphasis>
0-2-2-0, III 
<emphasis id="B927803CF022FF8957D4FE82F1C4FEA3" box="[397,419,295,321]" italics="true" pageId="24" pageNumber="367">pl</emphasis>
1-0-0-1 
<emphasis id="B927803CF022FF89545FFE82F27EFEA3" box="[518,537,295,321]" italics="true" pageId="24" pageNumber="367">rl</emphasis>
0-0-1 
<emphasis id="B927803CF022FF89543CFE82F219FEA2" box="[613,638,295,320]" italics="true" pageId="24" pageNumber="367">ve</emphasis>
2-2, IV 
<emphasis id="B927803CF022FF895487FE82F293FEA3" box="[734,756,295,321]" italics="true" pageId="24" pageNumber="367">pl</emphasis>
0-1-0-1 
<emphasis id="B927803CF022FF895501FE82F30CFEA3" box="[856,875,295,321]" italics="true" pageId="24" pageNumber="367">rl</emphasis>
0-1-0-1 
<emphasis id="B927803CF022FF895597FE82F380FEA2" box="[974,999,295,320]" italics="true" pageId="24" pageNumber="367">ve</emphasis>
1 
<emphasis id="B927803CF022FF8955A5FE82F474FEA2" box="[1020,1043,295,320]" italics="true" pageId="24" pageNumber="367">p-</emphasis>
1 
<emphasis id="B927803CF022FF89527BFE82F457FEA2" box="[1058,1072,295,320]" italics="true" pageId="24" pageNumber="367">p</emphasis>
-1 
<emphasis id="B927803CF022FF89521EFE82F435FEA2" box="[1095,1106,295,320]" italics="true" pageId="24" pageNumber="367">r</emphasis>
-1 
<emphasis id="B927803CF022FF895230FE82F413FEA2" box="[1129,1140,295,320]" italics="true" pageId="24" pageNumber="367">r</emphasis>
. Coloration: carapace and chelicerae dark reddish brown. Endites, labium and sternum orange red. Legs whitish with femora I and II reddish. Abdomen as in male (
<figureCitation id="136840ABF022FF8957CDFECAF19EFE6B" box="[404,505,367,393]" captionStart="FIGURE 18" captionStartId="22.[152,255,1304,1328]" captionTargetBox="[151,1435,181,1281]" captionTargetId="figure-19@22.[151,1435,181,1281]" captionTargetPageId="22" captionText="FIGURE 18. Carteronius gentilis (Simon, 1909) comb. nov.: A male holotype habitus, dorsal view; B left palp, prolateral view; C same, ventral view; D same, retrolateral view; E original label; F female abdomen, dorsal view; B female carapace, dorsal view. Scale: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7307090" httpUri="https://zenodo.org/record/7307090/files/figure.png" pageId="24" pageNumber="367">Fig. 18F</figureCitation>
). Epigynum: CDv long, straight, narrow; ST2 tappering, anteriorly located, gland ducts conspicuous, smaller than ST1; CDd large, S-shaped (
<figureCitation id="136840ABF022FF89556EFE36F3C7FE4F" box="[823,928,403,429]" captionStart="FIGURE 19" captionStartId="23.[152,255,1851,1875]" captionTargetBox="[197,1385,188,1822]" captionTargetId="figure-19@23.[192,1395,181,1828]" captionTargetPageId="23" captionText="FIGURE 19. Carteronius gentilis (Simon, 1909) comb. nov.: A male palp, ventral view; B same, retrolateral view; C epigynum, ventral view; D same, dorsal view. Abbreviations: E, embolus; C, conductor; MA, median apophysis; VL, ventral lobe; ML, medial lobe; DL, dorsal lobe; AS, spur; CDv, ventral sector of the copulatory duct; CDd, dorsal sector of the copulatory duct; ST2 secondary spermatheca; ST1 primary spermatheca. Scale: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7307096" httpUri="https://zenodo.org/record/7307096/files/figure.png" pageId="24" pageNumber="367">Fig. 19D</figureCitation>
).
</paragraph>
<caption id="DF2C0CA6F02DFF8656C1F89EF33EF85D" ID-DOI="http://doi.org/10.5281/zenodo.7307096" ID-Zenodo-Dep="7307096" httpUri="https://zenodo.org/record/7307096/files/figure.png" pageId="23" pageNumber="366" startId="23.[152,255,1851,1875]" targetBox="[197,1385,188,1822]" targetPageId="23">
<paragraph id="8BEC5C2EF02DFF8656C1F89EF33EF85D" blockId="23.[151,1437,1851,1984]" pageId="23" pageNumber="366">
<emphasis id="B927803CF02DFF8656C1F89EF143F8B6" bold="true" box="[152,292,1851,1876]" pageId="23" pageNumber="366">FIGURE 19.</emphasis>
<taxonomicName id="4C5327ADF02DFF865770F899F2E0F8B6" authority="(Simon, 1909)" authorityName="Eb.Bonaldo &amp; Ramírez &amp; Om.Labarque &amp; Shimano &amp; Silva-Junior &amp; Haddad" authorityYear="2022" baseAuthorityName="Simon" baseAuthorityYear="1909" box="[297,647,1852,1876]" class="Arachnida" family="Clubionidae" genus="Carteronius" kingdom="Animalia" order="Araneae" pageId="23" pageNumber="366" phylum="Arthropoda" rank="species" species="gentilis" status="comb. nov.">
<emphasis id="B927803CF02DFF865770F899F195F8B6" box="[297,498,1852,1876]" italics="true" pageId="23" pageNumber="366">Carteronius gentilis</emphasis>
(
<bibRefCitation id="EFC221DFF02DFF8657A7F899F2E7F8B6" author="Simon, E." box="[510,640,1852,1876]" pageId="23" pageNumber="366" pagination="335 - 449" refId="ref18798" refString="Simon, E. (1909) Arachnides recueillis par L. Fea sur la cote occidentale d'Afrique. 2 e partie. Annali del Museo Civico di Storia Naturale di Genova, 44, 335 - 449." type="journal article" year="1909">Simon, 1909</bibRefCitation>
)
</taxonomicName>
<emphasis id="B927803CF02DFF8654D5F899F29BF8B6" bold="true" box="[652,764,1852,1876]" pageId="23" pageNumber="366">
<taxonomicNameLabel id="A2143D47F02DFF8654D5F899F29BF8B6" box="[652,764,1852,1876]" pageId="23" pageNumber="366" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
: 
<emphasis id="B927803CF02DFF865551F89EF37EF8B1" bold="true" box="[776,793,1851,1875]" pageId="23" pageNumber="366">A</emphasis>
male palp, ventral view; 
<emphasis id="B927803CF02DFF86524AF899F443F8B6" bold="true" box="[1043,1060,1852,1876]" pageId="23" pageNumber="366">B</emphasis>
same, retrolateral view; 
<emphasis id="B927803CF02DFF865340F89EF54CF8B1" bold="true" box="[1305,1323,1851,1875]" pageId="23" pageNumber="366">C</emphasis>
epigynum, ventral view; 
<emphasis id="B927803CF02DFF86577FF8C5F15FF89A" bold="true" box="[294,312,1888,1912]" pageId="23" pageNumber="366">D</emphasis>
same, dorsal view. Abbreviations: E, embolus; C, conductor; MA, median apophysis; VL, ventral lobe; ML, medial lobe; DL, dorsal lobe; AS, spur; CDv, ventral sector of the copulatory duct; CDd, dorsal sector of the copulatory duct; ST2 secondary spermatheca; ST1 primary spermatheca. Scale: 1 mm.
</paragraph>
</caption>
<paragraph id="8BEC5C2EF022FF89569EFE13F56AFE17" blockId="24.[151,1437,150,537]" pageId="24" pageNumber="367">
<emphasis id="B927803CF022FF89569EFE13F192FE32" bold="true" box="[199,501,438,464]" pageId="24" pageNumber="367">Other material examined</emphasis>
: 
<materialsCitation id="3B3B5673F022FF89545DFE12F46EFE17" ID-GBIF-Occurrence="3962979342" collectingDate="1981-03" collectionCode="II-, V" collectorName="Bosmans &amp; Van Stalle" country="Cameroon" elevation="1400" latitude="4.2" location="Mount Cameroun" longLatPrecision="1307" longitude="9.183333" municipality="Buea" pageId="24" pageNumber="367" specimenCount="1">
<collectingCountry id="F3441CBEF022FF89545DFE12F2CEFE33" box="[516,681,439,465]" name="Cameroon" pageId="24" pageNumber="367">CAMEROON</collectingCountry>
: 
<location id="8E8C0AF5F022FF8954EAFE12F31AFE33" LSID="urn:lsid:plazi:treatment:03FAED38F02FFF8956CEFADBF4ADFDFB:8E8C0AF5F022FF8954EAFE12F31AFE33" box="[691,893,439,465]" country="Cameroon" latitude="4.2" longLatPrecision="1307" longitude="9.183333" municipality="Buea" name="Mount Cameroun" pageId="24" pageNumber="367">Mount Cameroun</location>
, near 
<collectingMunicipality id="6B88C654F022FF895599FE12F39BFE33" box="[960,1020,439,465]" pageId="24" pageNumber="367">Buea</collectingMunicipality>
, 
<geoCoordinate id="EE673AE9F022FF895251FE12F40DFE33" box="[1032,1130,439,465]" degrees="04" direction="north" minutes="12" orientation="latitude" pageId="24" pageNumber="367" precision="925" value="4.2">04°12’N</geoCoordinate>
, 
<geoCoordinate id="EE673AE9F022FF89522EFE12F4B2FE33" box="[1143,1237,439,465]" degrees="09" direction="east" minutes="11" orientation="longitude" pageId="24" pageNumber="367" precision="925" value="9.183333">09°11’E</geoCoordinate>
, 
<quantity id="4CABF1CBF022FF8952BBFE12F559FE33" box="[1250,1342,439,465]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="24" pageNumber="367" unit="m" value="1400.0">
<elevation id="007EBB1DF022FF8952BBFE12F559FE33" box="[1250,1342,439,465]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="24" pageNumber="367" unit="m" value="1400.0">1400 m</elevation>
</quantity>
.a.s.l., 
<date id="FFED7AEEF022FF8953D9FE12F092FE17" pageId="24" pageNumber="367" value="1981-03">
<collectingDate id="EFA98306F022FF8953D9FE12F092FE17" pageId="24" pageNumber="367" value="1981-03">
<collectionCode id="ED42C4EBF022FF8953D9FE12F0D1FE17" pageId="24" pageNumber="367">II-III</collectionCode>
.1981
</collectingDate>
</date>
, 
<collectorName id="26A639F8F022FF895759FE7EF100FE17" box="[256,359,475,501]" pageId="24" pageNumber="367">Bosmans</collectorName>
&amp; 
<collectorName id="26A639F8F022FF8957D3FE7EF19AFE17" box="[394,509,475,501]" pageId="24" pageNumber="367">Van Stalle</collectorName>
leg. (montane forest with arable fields, 
<collectionCode id="ED42C4EBF022FF8955E2FE7EF3B6FE17" box="[955,977,475,501]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="24" pageNumber="367" type="Museum">V</collectionCode>
.H.F)
</materialsCitation>
, 
<materialsCitation id="3B3B5673F022FF89524CFE7EF56EFE17" ID-GBIF-Occurrence="3962979332" box="[1045,1289,475,501]" collectionCode="MRAC" country="Cameroon" pageId="24" pageNumber="367" specimenCode="MRAC 162.128" specimenCount="1" specimenCount-female="1">
<specimenCount id="9D5597A7F022FF89524CFE7EF45FFE17" box="[1045,1080,475,501]" pageId="24" pageNumber="367" type="female">1♀</specimenCount>
(
<specimenCode id="DBF5F455F022FF89521FFE7EF566FE17" box="[1094,1281,475,501]" collectionCode="MRAC" country="Belgium" httpUri="http://grbio.org/cool/z14b-bb31" name="Musée Royal de l’Afrique Centrale" pageId="24" pageNumber="367" type="Museum">MRAC 162.128</specimenCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3490FA5F022FF89569EFE5BF4ADFDFB" box="[199,1226,510,537]" pageId="24" pageNumber="367" type="distribution">
<paragraph id="8BEC5C2EF022FF89569EFE5BF4ADFDFB" blockId="24.[151,1437,150,537]" box="[199,1226,510,537]" pageId="24" pageNumber="367">
<emphasis id="B927803CF022FF89569EFE5BF13AFDFA" bold="true" box="[199,349,510,536]" pageId="24" pageNumber="367">Distribution.</emphasis>
Recorded from the island of Bioko (
<collectingCountry id="F3441CBEF022FF8954B2FE5AF3D1FDFB" box="[747,950,511,537]" name="Equatorial Guinea" pageId="24" pageNumber="367">Equatorial Guinea</collectingCountry>
) and 
<collectingCountry id="F3441CBEF022FF8955A8FE5AF404FDFB" box="[1009,1123,511,537]" name="Cameroon" pageId="24" pageNumber="367">Cameroon</collectingCountry>
(
<figureCitation id="136840ABF022FF89522BFE5AF4DEFDFB" box="[1138,1209,511,537]" captionStart="MAP 1" captionStartId="28.[152,211,1701,1725]" captionTargetBox="[151,1436,777,1678]" captionTargetId="figure-256@28.[151,1436,777,1678]" captionTargetPageId="28" captionText="MAP 1. Distribution of Carteronius species and Bunyoronius femoralis sp. nov." figureDoi="http://doi.org/10.5281/zenodo.7310962" httpUri="https://zenodo.org/record/7310962/files/figure.png" pageId="24" pageNumber="367">Map 1</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>