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<document id="4D80D20CF8070C830D575251D1F46846" ID-CLB-Dataset="51163" ID-DOI="10.11646/zootaxa.4066.5.7" ID-GBIF-Dataset="10a50e95-023c-4d71-bf69-70071162f79e" ID-ISSN="1175-5326" ID-Zenodo-Dep="266479" ID-ZooBank="4D6292DE-EE09-45FE-971C-B3408DC751CD" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459692109250" checkinUser="plazi" docAuthor="Smetana, Aleš" docDate="2016" docId="03E74E0E480CEB6EFF3524E5FB6165AE" docLanguage="en" docName="zootaxa.4066.5.7.pdf" docOrigin="Zootaxa 4066 (5)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Geodromicus taiwanensis Shavrin &amp; Smetana, sp. n." docType="treatment" docVersion="9" lastPageNumber="597" masterDocId="FFDE3676480FEB68FFA22228FFA56706" masterDocTitle="New species and records of the genus Geodromicus Redtenbacher, 1857 (Coleoptera: Staphylinidae: Omaliinae) from Taiwan" masterLastPageNumber="599" masterPageNumber="591" pageNumber="594" updateTime="1698209388784" updateUser="ExternalLinkService">
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<mods:title id="34C4F9B24B76A34477F8D7289B5C927F">New species and records of the genus Geodromicus Redtenbacher, 1857 (Coleoptera: Staphylinidae: Omaliinae) from Taiwan</mods:title>
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<mods:namePart id="ED42AE59859A8985BCA99CA26B978EEF">Smetana, Aleš</mods:namePart>
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<mods:title id="8D1DA50DA3A99F13B8519FB32E2DEBBC">Zootaxa</mods:title>
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<mods:date id="C5D795C8F0345A8C352A56FC18BC2B5B">2016</mods:date>
<mods:detail id="FB299D8D7E13A5C5822AAB83665B0C2E" type="volume">
<mods:number id="C8C0683A018B5AC4A0CFCB7E67487755">4066</mods:number>
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<mods:classification id="027A6B0F90D48D34AC83E2726AD42841">journal article</mods:classification>
<mods:identifier id="CC6CBFAE6798821340A7FD4FD68C1ACD" type="CLB-Dataset">51163</mods:identifier>
<mods:identifier id="1D426E0BD3D124C435695F0215AC435B" type="DOI">10.11646/zootaxa.4066.5.7</mods:identifier>
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<treatment id="03E74E0E480CEB6EFF3524E5FB6165AE" ID-DOI="http://doi.org/10.5281/zenodo.6091757" ID-GBIF-Taxon="119333484" ID-Zenodo-Dep="6091757" LSID="urn:lsid:plazi:treatment:03E74E0E480CEB6EFF3524E5FB6165AE" httpUri="http://treatment.plazi.org/id/03E74E0E480CEB6EFF3524E5FB6165AE" lastPageId="6" lastPageNumber="597" pageId="3" pageNumber="594">
<subSubSection id="C354AC93480CEB6BFF3524E5FE1C600F" pageId="3" pageNumber="594" type="nomenclature">
<paragraph id="8BF1FF18480CEB6BFF3524E5FCB961EE" blockId="3.[151,796,1741,1801]" box="[151,796,1741,1768]" pageId="3" pageNumber="594">
<heading id="D0B94874480CEB6BFF3524E5FCB961EE" bold="true" box="[151,796,1741,1768]" fontSize="11" level="1" pageId="3" pageNumber="594" reason="1">
<emphasis id="B93A230A480CEB6BFF3524E5FCB961EE" bold="true" box="[151,796,1741,1768]" pageId="3" pageNumber="594">
<taxonomicName id="4C4E849B480CEB6BFF3524E5FD6861EE" ID-CoL="6KBXM" authority="Shavrin &amp; Smetana" authorityName="Shavrin &amp; Smetana" box="[151,717,1741,1768]" class="Insecta" family="Staphylinidae" genus="Geodromicus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="594" phylum="Arthropoda" rank="species" species="taiwanensis" status="sp. nov.">
<emphasis id="B93A230A480CEB6BFF3524E5FE6961E1" bold="true" box="[151,460,1741,1767]" italics="true" pageId="3" pageNumber="594">Geodromicus taiwanensis</emphasis>
Shavrin &amp; Smetana
</taxonomicName>
,
<taxonomicNameLabel id="A2099E71480CEB6BFD7824E6FCB961EE" box="[730,796,1742,1768]" pageId="3" pageNumber="594" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BF1FF18480CEB6BFF3524D8FE1C600F" blockId="3.[151,796,1741,1801]" box="[151,441,1776,1801]" pageId="3" pageNumber="594">
(
<figureCitation id="1375E39D480CEB6BFF3D24D8FF4B600F" box="[159,238,1776,1801]" captionStart="FIGURES 1 – 2" captionStartId="2.[151,264,1170,1193]" captionTargetBox="[151,1426,197,1138]" captionTargetId="figure@2.[151,1436,193,1149]" captionTargetPageId="2" captionText="FIGURES 1 – 2. Habitus of Geodromicus: 1 — G. philydroides sp. n., holotype, 2 — G. taiwanensis, sp. n., male, paratype. Scale bar: 1.0 mm." httpUri="https://zenodo.org/record/266480/files/figure.png" pageId="3" pageNumber="594">Figs. 2</figureCitation>
,
<figureCitation id="1375E39D480CEB6BFF5824D8FE91600F" box="[250,308,1776,1801]" captionStart="FIGURES 3 – 11" captionStartId="3.[151,264,1577,1600]" captionTargetBox="[199,1397,452,1555]" captionTargetId="figure@3.[163,1423,447,1556]" captionTargetPageId="3" captionText="FIGURES 3 – 11. Abdominal segments (dorsal view) of Geodromicus philydroides sp. n. (3 − 5) and G. taiwanensis, sp. n. (6 − 11): 3, 6 — male abdominal tergite VIII, 4, 7 — male abdominal sternite VIII, 5, 8 — male genital segment, 9 — female abdominal tergite VIII, 10 — female abdominal sternite VIII, 11 — female genital segment. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/266481/files/figure.png" pageId="3" pageNumber="594">6–11</figureCitation>
,
<figureCitation id="1375E39D480CEB6BFE9D24D8FE2D600F" box="[319,392,1776,1801]" captionStart="FIGURES 14 – 16" captionStartId="5.[151,264,1239,1262]" captionTargetBox="[154,1428,308,1208]" captionTargetId="figure@5.[151,1436,301,1218]" captionTargetPageId="5" captionText="FIGURES 14 – 16. Aedeagus of Geodromicus taiwanensis sp. n.: 14, 16 — parameral view, 15 — lateral view. Scale bar: 0.1 mm." httpUri="https://zenodo.org/record/266483/files/figure.png" pageId="3" pageNumber="594">14–16</figureCitation>
,
<figureCitation id="1375E39D480CEB6BFE3124D9FE15600F" box="[403,432,1777,1801]" captionStart="FIGURES 17 – 18" captionStartId="6.[151,264,1785,1808]" captionTargetBox="[171,1428,745,1758]" captionTargetId="figure@6.[151,1436,735,1764]" captionTargetPageId="6" captionText="FIGURES 17 – 18. Distribution of Geodromicus philydroides sp. n. (17) and G. taiwanensis sp. n. (18) in Taiwan." httpUri="https://zenodo.org/record/266484/files/figure.png" pageId="3" pageNumber="594">18</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C354AC93480CEB6CFF352510FF5B6143" lastPageId="4" lastPageNumber="595" pageId="3" pageNumber="594" type="materials_examined">
<paragraph id="8BF1FF18480CEB6BFF352510FEEA60BB" blockId="3.[151,1436,1848,1981]" pageId="3" pageNumber="594">
<emphasis id="B93A230A480CEB6BFF352510FE186057" bold="true" box="[151,445,1848,1873]" pageId="3" pageNumber="594">
<typeStatus id="54F541BA480CEB6BFF352510FF776057" box="[151,210,1848,1873]" pageId="3" pageNumber="594">Type</typeStatus>
material examined.
</emphasis>
<typeStatus id="54F541BA480CEB6BFE662510FD896057" box="[452,556,1848,1873]" pageId="3" pageNumber="594" type="holotype">Holotype</typeStatus>
♂: ‘
<collectingCountry id="F359BF88480CEB6BFDFB2511FD686057" box="[601,717,1849,1873]" name="Taiwan" pageId="3" pageNumber="594">TAIWAN</collectingCountry>
, Kaohsiung | Hsien, Crk.
<quantity id="4CB652FD480CEB6BFC4B2510FBB96057" box="[1001,1052,1848,1873]" metricMagnitude="3" metricUnit="m" metricValue="4.0" pageId="3" pageNumber="594" unit="km" value="4.0">4km</quantity>
E | Yakou,
<quantity id="4CB652FD480CEB6BFB382510FB4D6057" box="[1178,1256,1848,1873]" metricMagnitude="3" metricUnit="m" metricValue="2.6" pageId="3" pageNumber="594" unit="m" value="2600.0">2600m</quantity>
<date id="FFF0D9D8480CEB6BFB4C2511FAE56057" box="[1262,1344,1849,1873]" pageId="3" pageNumber="594">23.VII.</date>
| [19]93 A.Smetana [T162]’, ‘
<taxonomicName id="4C4E849B480CEB6BFE392575FD956073" box="[411,560,1885,1909]" class="Insecta" family="Staphylinidae" genus="Geodromicus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="594" phylum="Arthropoda" rank="genus">
<emphasis id="B93A230A480CEB6BFE392575FD956073" box="[411,560,1885,1909]" italics="true" pageId="3" pageNumber="594">Geodromicus</emphasis>
</taxonomicName>
2 [handwritten] | A. Smetana det. 2015’ &lt;rectangular label, printed&gt;, ‘
<typeStatus id="54F541BA480CEB6BFF3925A9FE91609F" box="[155,308,1921,1945]" pageId="3" pageNumber="594" type="holotype">HOLOTYPE</typeStatus>
|
<taxonomicName id="4C4E849B480CEB6BFEEE25A8FE47609E" box="[332,482,1920,1944]" class="Insecta" family="Staphylinidae" genus="Geodromicus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="594" phylum="Arthropoda" rank="genus">
<emphasis id="B93A230A480CEB6BFEEE25A8FE47609E" box="[332,482,1920,1944]" italics="true" pageId="3" pageNumber="594">Geodromicus</emphasis>
</taxonomicName>
|
<taxonomicName id="4C4E849B480CEB6BFE5B25A9FDD8609E" box="[505,637,1921,1944]" class="Insecta" family="Staphylinidae" genus="Geodromicus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="594" phylum="Arthropoda" rank="species" species="taiwanensis" status="sp. nov.">
<emphasis id="B93A230A480CEB6BFE5B25A9FDD8609E" box="[505,637,1921,1944]" italics="true" pageId="3" pageNumber="594">taiwanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2099E71480CEB6BFD2425A9FD1E609E" box="[646,699,1921,1944]" pageId="3" pageNumber="594" rank="species">sp.n.</taxonomicNameLabel>
| Shavrin A.V. &amp; Smetana A. des. 2015’ &lt;red rectangular label, printed&gt; (CSm).
</paragraph>
<caption id="DF31AF90480BEB6CFF3526DBFA20620F" box="[151,1413,1267,1290]" httpUri="https://zenodo.org/record/266482/files/figure.png" pageId="4" pageNumber="595" targetBox="[217,1369,199,1233]" targetPageId="4">
<paragraph id="8BF1FF18480BEB6CFF3526DBFA20620F" blockId="4.[151,1413,1267,1290]" box="[151,1413,1267,1290]" pageId="4" pageNumber="595">
<emphasis id="B93A230A480BEB6CFF3526DBFEF6620E" bold="true" box="[151,339,1267,1290]" pageId="4" pageNumber="595">FIGURES 12–13.</emphasis>
Aedeagus of
<taxonomicName id="4C4E849B480BEB6CFE7D26DBFD4E620F" box="[479,747,1267,1289]" class="Insecta" family="Staphylinidae" genus="Geodromicus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="595" phylum="Arthropoda" rank="species" species="philydroides" status="sp. nov.">
<emphasis id="B93A230A480BEB6CFE7D26DBFD4E620F" box="[479,747,1267,1289]" italics="true" pageId="4" pageNumber="595">Geodromicus philydroides</emphasis>
</taxonomicName>
<emphasis id="B93A230A480BEB6CFD5326DCFC8F620F" bold="true" box="[753,810,1268,1289]" pageId="4" pageNumber="595">
<taxonomicNameLabel id="A2099E71480BEB6CFD5326DCFC8F620F" box="[753,810,1268,1289]" pageId="4" pageNumber="595" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
: 12—parameral view, 13—lateral view. Scale bar: 0.1 mm.
</paragraph>
</caption>
<paragraph id="8BF1FF18480BEB6CFF652719FF5B6143" blockId="4.[151,1437,1328,2038]" pageId="4" pageNumber="595">
<typeStatus id="54F541BA480BEB6CFF652719FE91624F" box="[199,308,1329,1353]" pageId="4" pageNumber="595" type="paratype">Paratypes</typeStatus>
[Three males (two from Yakou and one from Nanhushi) were dissected: last abdominal segments of the specimens were glued under the beetles; plastic plate with the aedeagus in Canadian balsam was pinned under beetles]:
<specimenCount id="9D483491480BEB6CFF5E2750FE836296" box="[252,294,1400,1425]" pageId="4" pageNumber="595" type="generic">5 ♂</specimenCount>
, 14 ♀: same data as the
<typeStatus id="54F541BA480BEB6CFD962750FD336297" box="[564,662,1400,1425]" pageId="4" pageNumber="595" type="holotype">holotype</typeStatus>
(CSm, CSh); 1 ♀: ‘
<collectingCountry id="F359BF88480BEB6CFCCD2751FC466297" box="[879,995,1401,1425]" name="Taiwan" pageId="4" pageNumber="595">TAIWAN</collectingCountry>
, Kaohsiung | Hsien, Peinantashan | trail
<quantity id="4CB652FD480BEB6CFF3527B4FF4D62B3" box="[151,232,1436,1461]" metricMagnitude="3" metricUnit="m" metricValue="2.45" pageId="4" pageNumber="595" unit="m" value="2450.0">2450m</quantity>
,
<date id="FFF0D9D8480BEB6CFF5127B4FEFD62B3" box="[243,344,1436,1461]" pageId="4" pageNumber="595" value="1995-05-02">2.V.1995</date>
| A. Smetana [T170]’ (cSm);
<specimenCount id="9D483491480BEB6CFD0827B5FD7262B2" box="[682,727,1437,1461]" pageId="4" pageNumber="595" type="generic">1 ♂</specimenCount>
, 7 ♀: ‘
<collectingCountry id="F359BF88480BEB6CFC8327B5FC3662B3" box="[801,915,1437,1461]" name="Taiwan" pageId="4" pageNumber="595">TAIWAN</collectingCountry>
Hualien | Hsien, Taroko N.P. | Nanhushi Hut |
<quantity id="4CB652FD480BEB6CFF3527E9FF4062DF" box="[151,229,1473,1497]" metricMagnitude="3" metricUnit="m" metricValue="2.2199999999999998" pageId="4" pageNumber="595" unit="m" value="2220.0">2220m</quantity>
<date id="FFF0D9D8480BEB6CFF4E27E9FEF562DF" box="[236,336,1473,1497]" pageId="4" pageNumber="595" value="1990-05-08">8.V.1990</date>
| A. Smetana [T48]’ (cSm, cSh); 1 ♀: ‘
<collectingCountry id="F359BF88480BEB6CFCAB27E9FCDE62DF" box="[777,891,1473,1497]" name="Taiwan" pageId="4" pageNumber="595">TAIWAN</collectingCountry>
Hualien | Hsien, Taroko N. P. | Chungyantienshi | (Riv.) Waterfall |
<quantity id="4CB652FD480BEB6CFE2627CDFE7462FB" box="[388,465,1509,1533]" metricMagnitude="3" metricUnit="m" metricValue="2.3" pageId="4" pageNumber="595" unit="m" value="2300.0">2300m</quantity>
<date id="FFF0D9D8480BEB6CFE4227CDFDF662FB" box="[480,595,1509,1533]" pageId="4" pageNumber="595" value="1990-05-10">10.V.1990</date>
| A. Smetana [T50]’ (cSm). All
<typeStatus id="54F541BA480BEB6CFC5327CDFBF862FB" box="[1009,1117,1509,1533]" pageId="4" pageNumber="595" type="paratype">paratypes</typeStatus>
with our additional label: ‘
<typeStatus id="54F541BA480BEB6CFF392421FE896127" box="[155,300,1545,1569]" pageId="4" pageNumber="595" type="paratype">PARATYPE</typeStatus>
|
<taxonomicName id="4C4E849B480BEB6CFEE62421FE7F6127" box="[324,474,1545,1569]" class="Insecta" family="Staphylinidae" genus="Geodromicus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="595" phylum="Arthropoda" rank="genus">
<emphasis id="B93A230A480BEB6CFEE62421FE7F6127" box="[324,474,1545,1569]" italics="true" pageId="4" pageNumber="595">Geodromicus</emphasis>
</taxonomicName>
|
<taxonomicName id="4C4E849B480BEB6CFE502421FDD26126" box="[498,631,1545,1568]" class="Insecta" family="Staphylinidae" genus="Geodromicus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="595" phylum="Arthropoda" rank="species" species="taiwanensis" status="sp. nov.">
<emphasis id="B93A230A480BEB6CFE502421FDD26126" box="[498,631,1545,1568]" italics="true" pageId="4" pageNumber="595">taiwanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2099E71480BEB6CFD222421FD106126" box="[640,693,1545,1568]" pageId="4" pageNumber="595" rank="species">sp.n.</taxonomicNameLabel>
| Shavrin A.V. &amp; Smetana A. des. 2015’ &lt;red rectangular label, printed&gt;.
</paragraph>
</subSubSection>
<subSubSection id="C354AC93480BEB6EFF652478FC52668E" lastPageId="6" lastPageNumber="597" pageId="4" pageNumber="595" type="description">
<paragraph id="8BF1FF18480BEB6CFF652478FA9161B7" blockId="4.[151,1437,1328,2038]" pageId="4" pageNumber="595">
<emphasis id="B93A230A480BEB6CFF652478FEFC616F" bold="true" box="[199,345,1616,1641]" pageId="4" pageNumber="595">Description.</emphasis>
Measurements (n=30): WH: 0.70–0.87; LH: 0.50–0.70; LA (
<typeStatus id="54F541BA480BEB6CFBAA2478FBD4616F" box="[1032,1137,1616,1641]" pageId="4" pageNumber="595" type="holotype">holotype</typeStatus>
): 2.13; LE: 0.18−0.25; LT: 0.12−0.22; LPM × WPM (
<typeStatus id="54F541BA480BEB6CFE1C245CFD82618B" box="[446,551,1652,1677]" pageId="4" pageNumber="595" type="holotype">holotype</typeStatus>
): III: 0.16 × 0.07, IV: 0.15 × 0.05; LP: 0.60–0.75; WPMax: 0.75–0.87; WPMin: 0.51–0.60; LEl: 1.14–1.37; WE: 1.24–1.47; WA: 1.21−1.47; LAed: 0.65; TL: 3.30–4.60 (
<typeStatus id="54F541BA480BEB6CFBDD24B0FB4261B7" box="[1151,1255,1688,1713]" pageId="4" pageNumber="595" type="holotype">holotype</typeStatus>
: 4.25).
</paragraph>
<paragraph id="8BF1FF18480BEB6DFF652494FCE967FE" blockId="4.[151,1437,1328,2038]" lastBlockId="5.[151,1437,151,248]" lastPageId="5" lastPageNumber="596" pageId="4" pageNumber="595">
Body and antennomeres reddish brown to brown; apical margins, basal third of each elytron and paratergites yellow brown to reddish brown; legs yellow brown, apical part of femora and/or entire length of tibiae brown; ocelli, basis of antennomeres 1–3 and mouthparts yellow to reddish yellow. Punctation of head dense, deep and coarse, interspaces between punctures on infraorbital ridge as broad as diameters of 1–2 nearest punctures, punctation somewhat coarser on interocellar impression; punctation of pronotum same as that on head, with impunctate oval area between longitudinal and basal depressions; scutellum without punctures; punctation of elytra deep, sparse and regular; abdomen with very small and regular punctures. Body glossy; head with distinct coarse cellular microsculpture, clypeus with smoother microsculpture; microsculpture of pronotum variable: same as that on head, especially on apical third, or more gentle, indistict or entirely reduced, impunctate area of basal third of pronotum without microsculpture, longitudinal wide depression with very dense and coarse microsculpture; scutellum with distinct isodiametric microsculpture or without it; elytra without microsculpture; abdomen with distinct fine isodiametric microsculpture. Habitus as in
<figureCitation id="1375E39D480AEB6DFCA122F7FCED67FE" box="[771,840,223,248]" captionStart="FIGURES 1 – 2" captionStartId="2.[151,264,1170,1193]" captionTargetBox="[151,1426,197,1138]" captionTargetId="figure@2.[151,1436,193,1149]" captionTargetPageId="2" captionText="FIGURES 1 – 2. Habitus of Geodromicus: 1 — G. philydroides sp. n., holotype, 2 — G. taiwanensis, sp. n., male, paratype. Scale bar: 1.0 mm." httpUri="https://zenodo.org/record/266480/files/figure.png" pageId="5" pageNumber="596">Fig. 2</figureCitation>
.
</paragraph>
<caption id="DF31AF90480AEB6DFF3526FFFA3963E8" box="[151,1436,1239,1262]" httpUri="https://zenodo.org/record/266483/files/figure.png" pageId="5" pageNumber="596" targetBox="[154,1428,308,1208]" targetPageId="5">
<paragraph id="8BF1FF18480AEB6DFF3526FFFA3963E8" blockId="5.[151,1436,1239,1262]" box="[151,1436,1239,1262]" pageId="5" pageNumber="596">
<emphasis id="B93A230A480AEB6DFF3526FFFEF663EB" bold="true" box="[151,339,1239,1262]" pageId="5" pageNumber="596">FIGURES 14–16.</emphasis>
Aedeagus of
<taxonomicName id="4C4E849B480AEB6DFE7F26F0FD4763E8" box="[477,738,1240,1262]" class="Insecta" family="Staphylinidae" genus="Geodromicus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="596" phylum="Arthropoda" rank="species" species="taiwanensis" status="sp. nov.">
<emphasis id="B93A230A480AEB6DFE7F26F0FD4763E8" box="[477,738,1240,1262]" italics="true" pageId="5" pageNumber="596">Geodromicus taiwanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2099E71480AEB6DFD4526F0FCBA63EB" box="[743,799,1240,1261]" pageId="5" pageNumber="596" rank="species">
<emphasis id="B93A230A480AEB6DFD4526F0FCBA63EB" bold="true" box="[743,799,1240,1261]" pageId="5" pageNumber="596">sp. n.</emphasis>
</taxonomicNameLabel>
: 14, 16—parameral view, 15—lateral view. Scale bar: 0.1 mm.
</paragraph>
</caption>
<paragraph id="8BF1FF18480AEB6DFF65273DFF43610F" blockId="5.[151,1437,1301,2013]" pageId="5" pageNumber="596">Head 1.2–1.4 times as broad as long, with moderately deep triangular to rectangular interocellar impression and with convex posterior parts of infraorbital ridges, with moderately narrow postocular parts. Eyes mediumsized, markedly convex, 1.1–1.5 times as long as temples. Ocelli large, distance between ocelli slightly longer than distance between ocellus and posterior margin of eye. Maxillary palp segment 3 slightly longer than apical (4th) segment. Antennae reaching half length or posterior third of elytra when reclined; antennomeres with lengths × widths: 1: 0.27 × 0.10; 2: 0.17 × 0.07; 3: 0.18 × 0.07; 4: 0.17 × 0.07; 5−7: 0.18 × 0.07; 8−10: 0.18 × 0.08; 11: 0.26 × 0.08.</paragraph>
<paragraph id="8BF1FF18480AEB6DFF65243DFB1F6173" blockId="5.[151,1437,1301,2013]" pageId="5" pageNumber="596">Pronotum small and convex, 1.1−1.2 times as broad as long, as broad as head or slightly wider; anterior margin of pronotum slightly rounded; posterior angles nearly rectangular; longitudinal depression moderately wide and deep, widest in apical part; basal part of pronotum with moderately deep transverse depression</paragraph>
<paragraph id="8BF1FF18480AEB6DFF6524A8FD3661BA" blockId="5.[151,1437,1301,2013]" pageId="5" pageNumber="596">Elytra somewhat wider than long, slightly depressed in basal third, 1.8−1.9 times as long as pronotum; hind margins of elytra straight or slightly truncate.</paragraph>
<paragraph id="8BF1FF18480AEB6DFF6524E0FAB56002" blockId="5.[151,1437,1301,2013]" pageId="5" pageNumber="596">Abdomen as broad as elytra or slightly narrower, with two moderately wide wing-folding patches (tomentose spots) on abdominal tergite IV, with distinct palisade fringe on apical margin of abdominal tergite VII.</paragraph>
<paragraph id="8BF1FF18480AEB6DFF652538FBAC60DB" blockId="5.[151,1437,1301,2013]" pageId="5" pageNumber="596">
Male. Pronotum and front femora wider. Apical margin of abdominal tergite VIII straight (
<figureCitation id="1375E39D480AEB6DFB662538FAAD602F" box="[1220,1288,1808,1833]" captionStart="FIGURES 3 – 11" captionStartId="3.[151,264,1577,1600]" captionTargetBox="[199,1397,452,1555]" captionTargetId="figure@3.[163,1423,447,1556]" captionTargetPageId="3" captionText="FIGURES 3 – 11. Abdominal segments (dorsal view) of Geodromicus philydroides sp. n. (3 − 5) and G. taiwanensis, sp. n. (6 − 11): 3, 6 — male abdominal tergite VIII, 4, 7 — male abdominal sternite VIII, 5, 8 — male genital segment, 9 — female abdominal tergite VIII, 10 — female abdominal sternite VIII, 11 — female genital segment. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/266481/files/figure.png" pageId="5" pageNumber="596">Fig. 6</figureCitation>
). Abdominal sternite VIII (
<figureCitation id="1375E39D480AEB6DFE9B251CFE24604B" box="[313,385,1844,1869]" captionStart="FIGURES 3 – 11" captionStartId="3.[151,264,1577,1600]" captionTargetBox="[199,1397,452,1555]" captionTargetId="figure@3.[163,1423,447,1556]" captionTargetPageId="3" captionText="FIGURES 3 – 11. Abdominal segments (dorsal view) of Geodromicus philydroides sp. n. (3 − 5) and G. taiwanensis, sp. n. (6 − 11): 3, 6 — male abdominal tergite VIII, 4, 7 — male abdominal sternite VIII, 5, 8 — male genital segment, 9 — female abdominal tergite VIII, 10 — female abdominal sternite VIII, 11 — female genital segment. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/266481/files/figure.png" pageId="5" pageNumber="596">Fig. 7</figureCitation>
) with wide and moderately deep apical emargination. Male genital segment as in
<figureCitation id="1375E39D480AEB6DFAED251CFA32604B" box="[1359,1431,1844,1869]" captionStart="FIGURES 3 – 11" captionStartId="3.[151,264,1577,1600]" captionTargetBox="[199,1397,452,1555]" captionTargetId="figure@3.[163,1423,447,1556]" captionTargetPageId="3" captionText="FIGURES 3 – 11. Abdominal segments (dorsal view) of Geodromicus philydroides sp. n. (3 − 5) and G. taiwanensis, sp. n. (6 − 11): 3, 6 — male abdominal tergite VIII, 4, 7 — male abdominal sternite VIII, 5, 8 — male genital segment, 9 — female abdominal tergite VIII, 10 — female abdominal sternite VIII, 11 — female genital segment. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/266481/files/figure.png" pageId="5" pageNumber="596">Fig. 8</figureCitation>
. Aedeagus (
<figureCitation id="1375E39D480AEB6DFEBA2570FE006077" box="[280,421,1880,1905]" captionStart="FIGURES 14 – 16" captionStartId="5.[151,264,1239,1262]" captionTargetBox="[154,1428,308,1208]" captionTargetId="figure@5.[151,1436,301,1218]" captionTargetPageId="5" captionText="FIGURES 14 – 16. Aedeagus of Geodromicus taiwanensis sp. n.: 14, 16 — parameral view, 15 — lateral view. Scale bar: 0.1 mm." httpUri="https://zenodo.org/record/266483/files/figure.png" pageId="5" pageNumber="596">Figs. 14, 16</figureCitation>
) with long and moderately narrow median lobe, gradually narrowing from middle part towards small rounded apex; parameres moderately wide, long, almost reaching apex of aedeagus, with four setae on apical part, placed in pairs laterally and apically on setiferous surface. Structure of endophallus in general similar to that of
<taxonomicName id="4C4E849B480AEB6DFEFA25EDFE5B60DA" box="[344,510,1988,2012]" class="Insecta" family="Staphylinidae" genus="Geodromicus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="596" phylum="Arthropoda" rank="species" species="philydroides" status="sp. nov.">
<emphasis id="B93A230A480AEB6DFEFA25EDFE5B60DA" box="[344,510,1988,2012]" italics="true" pageId="5" pageNumber="596">P. philydroides</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2099E71480AEB6DFDA725ECFDE060DB" box="[517,581,1988,2013]" pageId="5" pageNumber="596" rank="species">
<emphasis id="B93A230A480AEB6DFDA725ECFDE060DB" bold="true" box="[517,581,1988,2013]" pageId="5" pageNumber="596">sp. n.</emphasis>
</taxonomicNameLabel>
Aedeagus in lateral aspect as in
<figureCitation id="1375E39D480AEB6DFC1025ECFBA060DB" box="[946,1029,1988,2013]" captionStart="FIGURES 14 – 16" captionStartId="5.[151,264,1239,1262]" captionTargetBox="[154,1428,308,1208]" captionTargetId="figure@5.[151,1436,301,1218]" captionTargetPageId="5" captionText="FIGURES 14 – 16. Aedeagus of Geodromicus taiwanensis sp. n.: 14, 16 — parameral view, 15 — lateral view. Scale bar: 0.1 mm." httpUri="https://zenodo.org/record/266483/files/figure.png" pageId="5" pageNumber="596">Fig. 15</figureCitation>
.
</paragraph>
<paragraph id="8BF1FF184809EB6EFF6522BFFC3967D2" blockId="6.[151,1437,151,680]" pageId="6" pageNumber="597">
Female. Pronotum and front femora narrower. Apical margins of abdominal tergite VIII (
<figureCitation id="1375E39D4809EB6EFB0422BFFB4267B6" box="[1190,1255,151,176]" captionStart="FIGURES 3 – 11" captionStartId="3.[151,264,1577,1600]" captionTargetBox="[199,1397,452,1555]" captionTargetId="figure@3.[163,1423,447,1556]" captionTargetPageId="3" captionText="FIGURES 3 – 11. Abdominal segments (dorsal view) of Geodromicus philydroides sp. n. (3 − 5) and G. taiwanensis, sp. n. (6 − 11): 3, 6 — male abdominal tergite VIII, 4, 7 — male abdominal sternite VIII, 5, 8 — male genital segment, 9 — female abdominal tergite VIII, 10 — female abdominal sternite VIII, 11 — female genital segment. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/266481/files/figure.png" pageId="6" pageNumber="597">Fig. 9</figureCitation>
) and abdominal sternite VIII (
<figureCitation id="1375E39D4809EB6EFE922294FE2467D2" box="[304,385,188,213]" captionStart="FIGURES 3 – 11" captionStartId="3.[151,264,1577,1600]" captionTargetBox="[199,1397,452,1555]" captionTargetId="figure@3.[163,1423,447,1556]" captionTargetPageId="3" captionText="FIGURES 3 – 11. Abdominal segments (dorsal view) of Geodromicus philydroides sp. n. (3 − 5) and G. taiwanensis, sp. n. (6 − 11): 3, 6 — male abdominal tergite VIII, 4, 7 — male abdominal sternite VIII, 5, 8 — male genital segment, 9 — female abdominal tergite VIII, 10 — female abdominal sternite VIII, 11 — female genital segment. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/266481/files/figure.png" pageId="6" pageNumber="597">Fig. 10</figureCitation>
) rounded. Female genital segment as in
<figureCitation id="1375E39D4809EB6EFCE42294FC3D67D2" box="[838,920,188,213]" captionStart="FIGURES 3 – 11" captionStartId="3.[151,264,1577,1600]" captionTargetBox="[199,1397,452,1555]" captionTargetId="figure@3.[163,1423,447,1556]" captionTargetPageId="3" captionText="FIGURES 3 – 11. Abdominal segments (dorsal view) of Geodromicus philydroides sp. n. (3 − 5) and G. taiwanensis, sp. n. (6 − 11): 3, 6 — male abdominal tergite VIII, 4, 7 — male abdominal sternite VIII, 5, 8 — male genital segment, 9 — female abdominal tergite VIII, 10 — female abdominal sternite VIII, 11 — female genital segment. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/266481/files/figure.png" pageId="6" pageNumber="597">Fig. 11</figureCitation>
.
</paragraph>
<paragraph id="8BF1FF184809EB6EFF6522F7FC52668E" blockId="6.[151,1437,151,680]" pageId="6" pageNumber="597">
<emphasis id="B93A230A4809EB6EFF6522F7FE1767FE" bold="true" box="[199,434,223,248]" pageId="6" pageNumber="597">Comparative notes.</emphasis>
Based on the narrow body, the character of punctation and microsculpture of the body, the wide longitudinal depression on the pronotum, and the very similar shape of parameres and internal structure of the aedeagus,
<taxonomicName id="4C4E849B4809EB6EFE962300FE726639" box="[308,471,296,319]" class="Insecta" family="Staphylinidae" genus="Geodromicus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="597" phylum="Arthropoda" rank="species" species="taiwanensis" status="sp. nov.">
<emphasis id="B93A230A4809EB6EFE962300FEE06639" box="[308,325,296,319]" italics="true" pageId="6" pageNumber="597">G</emphasis>
.
<emphasis id="B93A230A4809EB6EFEF12300FE726639" box="[339,471,296,319]" italics="true" pageId="6" pageNumber="597">taiwanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2099E714809EB6EFE7C230FFDB86646" box="[478,541,295,320]" pageId="6" pageNumber="597" rank="species">
<emphasis id="B93A230A4809EB6EFE7C230FFDB86646" bold="true" box="[478,541,295,320]" pageId="6" pageNumber="597">sp. n.</emphasis>
</taxonomicNameLabel>
is similar to
<taxonomicName id="4C4E849B4809EB6EFD0C2300FCF26646" box="[686,855,296,320]" class="Insecta" family="Staphylinidae" genus="Geodromicus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="597" phylum="Arthropoda" rank="species" species="philydroides" status="sp. nov.">
<emphasis id="B93A230A4809EB6EFD0C2300FCF26646" box="[686,855,296,320]" italics="true" pageId="6" pageNumber="597">G. philydroides</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2099E714809EB6EFCFC230FFC386646" box="[862,925,295,320]" pageId="6" pageNumber="597" rank="species">
<emphasis id="B93A230A4809EB6EFCFC230FFC386646" bold="true" box="[862,925,295,320]" pageId="6" pageNumber="597">sp. n.</emphasis>
</taxonomicNameLabel>
, from which it differs by the paler coloration, narrower postocular parts of head and pronotum, shorter apical antennomere, the length of elytra, and by the shape of the median lobe of the aedeagus (for details see above and the key below).
</paragraph>
</subSubSection>
<subSubSection id="C354AC934809EB6EFF6523BCFB7366AB" box="[199,1238,404,429]" pageId="6" pageNumber="597" type="etymology">
<paragraph id="8BF1FF184809EB6EFF6523BCFB7366AB" blockId="6.[151,1437,151,680]" box="[199,1238,404,429]" pageId="6" pageNumber="597">
<emphasis id="B93A230A4809EB6EFF6523BCFEEB66AB" bold="true" box="[199,334,404,429]" pageId="6" pageNumber="597">Etymology.</emphasis>
The specific epithet is an adjective derived from the geographical name
<collectingCountry id="F359BF884809EB6EFBDE23BCFB7766AB" box="[1148,1234,404,429]" name="Taiwan" pageId="6" pageNumber="597">Taiwan</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C354AC934809EB6EFF65239FFB6165AE" pageId="6" pageNumber="597" type="distribution">
<paragraph id="8BF1FF184809EB6EFF65239FFEFC66F2" blockId="6.[151,1437,151,680]" pageId="6" pageNumber="597">
<emphasis id="B93A230A4809EB6EFF65239FFEC566D6" bold="true" box="[199,352,439,464]" pageId="6" pageNumber="597">Distribution.</emphasis>
The species is at present known from central (Hualien Hsien) and southern (Kaohsiung Hsien)
<collectingCountry id="F359BF884809EB6EFF3523F4FF4C66F3" box="[151,233,476,501]" name="Taiwan" pageId="6" pageNumber="597">Taiwan</collectingCountry>
(
<figureCitation id="1375E39D4809EB6EFF5A23F4FEE966F2" box="[248,332,476,501]" captionStart="FIGURES 17 – 18" captionStartId="6.[151,264,1785,1808]" captionTargetBox="[171,1428,745,1758]" captionTargetId="figure@6.[151,1436,735,1764]" captionTargetPageId="6" captionText="FIGURES 17 – 18. Distribution of Geodromicus philydroides sp. n. (17) and G. taiwanensis sp. n. (18) in Taiwan." httpUri="https://zenodo.org/record/266484/files/figure.png" pageId="6" pageNumber="597">Fig. 18</figureCitation>
).
</paragraph>
<paragraph id="8BF1FF184809EB6EFF6523D7FB6165AE" blockId="6.[151,1437,151,680]" pageId="6" pageNumber="597">
<emphasis id="B93A230A4809EB6EFF6523D7FEEF651E" bold="true" box="[199,330,511,536]" pageId="6" pageNumber="597">Bionomics.</emphasis>
Specimens were taken in original mixed forest (T162, T170) and in old coniferous forest with lush undergrowth (T48) by sifting wet moss from large rocks directly in a creek (T162), by sifting wet leaf litter and various debris on two small seepages (T170) and at a large creek by sifting wet layers of fallen leaves, accumulated at bases of large rocks close to the water (T48). One specimen (T50) was collected near a large waterfall in an old mixed forest by sifting various debris, fallen leaves and moss in the spray zone of the waterfall.
</paragraph>
</subSubSection>
</treatment>
</document>
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