treatments-xml/data/03/DB/87/03DB87DFE32B7A05FD3CF9E9FE916F36.xml
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<document id="C2F8F084BD5B2C126216EBEA7EAC9BF5" ID-DOI="10.5252/z2011n2a3" ID-ISSN="1638-9387" ID-Zenodo-Dep="4546855" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1613601229310" checkinUser="felipe" docAuthor="Bonato, Lucio, Iorio, Étienne &amp; Minelli, Alessandro" docDate="2011" docId="03DB87DFE32B7A05FD3CF9E9FE916F36" docLanguage="en" docName="Zoosystema.33.2.175-205.pdf" docOrigin="Zoosystema 33 (2)" docSource="http://dx.doi.org/10.5252/z2011n2a3" docStyle="DocumentStyle:EB3C2C73672D2E883259A050BD27978B.6:Zoosystema.2011-2012.journal_article" docStyleId="EB3C2C73672D2E883259A050BD27978B" docStyleName="Zoosystema.2011-2012.journal_article" docStyleVersion="6" docTitle="Clinopodes skopljensis" docType="treatment" docVersion="3" lastPageNumber="192" masterDocId="FFE2FFA7E33B7A14FFE7FFC3FFBD6A3E" masterDocTitle="The centipede genus Clinopodes C. L. Koch, 1847 (Chilopoda, Geophilomorpha, Geophilidae): reassessment of species diversity and distribution, with a new species from the Maritime Alps (France)" masterLastPageNumber="205" masterPageNumber="175" pageNumber="191" updateTime="1698873028875" updateUser="plazi" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="CC40CBDB3BEF696EA7E3E17E8CDB3955">The centipede genus Clinopodes C. L. Koch, 1847 (Chilopoda, Geophilomorpha, Geophilidae): reassessment of species diversity and distribution, with a new species from the Maritime Alps (France)</mods:title>
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<mods:namePart id="B0F987AD6DEABC23CA0D86F2ADCF1D3B">Bonato, Lucio</mods:namePart>
<mods:affiliation id="E607D51F457CF97D440F65555A2E246E">Università degli Studi di Padova, Dipartimento di Biologia, via U. Bassi 58 b, I- 35131 Padova (Italy) lucio. bonato @ unipd. it</mods:affiliation>
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<mods:namePart id="69C506815890B96AD7E70D31C832FD4C">Iorio, Étienne</mods:namePart>
<mods:affiliation id="AA7AF9FBA0C155D12FEBA8B5244FAAF8">ECO-MED (Écologie &amp; Médiation), Pôle Entomologie, Tour Méditerranée, 65 av. Jules Cantini, F- 13298 Marseille cedex 20 (France) e. iorio @ ecomed. fr</mods:affiliation>
<mods:nameIdentifier id="71EB7EC71F617253DF3E53C673700199" type="email">iorio@ecomed.fr</mods:nameIdentifier>
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<mods:namePart id="C648FE864EDBBD5317DC3C1970FA28AF">Minelli, Alessandro</mods:namePart>
<mods:affiliation id="1D60CE68830BC1B98E7A4646E030205A">Università degli Studi di Padova, Dipartimento di Biologia, via U. Bassi 58 b, I- 35131 Padova (Italy) alessandro. minelli @ unipd. it</mods:affiliation>
<mods:nameIdentifier id="925955FB2FDDA56894EDAEAD7EE192AF" type="email">minelli@unipd.it</mods:nameIdentifier>
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<mods:title id="8EE5ABE3BFFDD46903A97327BC9DC741">Zoosystema</mods:title>
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<mods:date id="0D4A91E66061683D02198FEDF3A5D723">2011</mods:date>
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<mods:number id="CA62D223E4C82CB5081147867CEF540C">2011-06-30</mods:number>
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<mods:number id="3B8C2021E088459BBFAD9263D21053E8">33</mods:number>
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<mods:identifier id="D2E19F807CC4B064DAD6D7D195A6223D" type="ISSN">1638-9387</mods:identifier>
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<treatment id="03DB87DFE32B7A05FD3CF9E9FE916F36" LSID="urn:lsid:plazi:treatment:03DB87DFE32B7A05FD3CF9E9FE916F36" httpUri="http://treatment.plazi.org/id/03DB87DFE32B7A05FD3CF9E9FE916F36" lastPageId="17" lastPageNumber="192" pageId="16" pageNumber="191">
<subSubSection id="C3686542E32B7A04FD3CF9E9FB3E6C7B" box="[731,1155,1578,1605]" pageId="16" pageNumber="191" type="nomenclature">
<paragraph id="8BCD36C9E32B7A04FD3CF9E9FB3E6C7B" blockId="16.[731,1155,1578,1605]" box="[731,1155,1578,1605]" pageId="16" pageNumber="191">
<heading id="D08581A5E32B7A04FD3CF9E9FB3E6C7B" box="[731,1155,1578,1605]" centered="true" fontSize="11" level="2" pageId="16" pageNumber="191" reason="2">
<taxonomicName id="4C724D4AE32B7A04FD3CF9E9FB3E6C7B" ID-CoL="W6NB" authority="(Verhoeff, 1938)" baseAuthorityName="Verhoeff" baseAuthorityYear="1938" box="[731,1155,1578,1605]" class="Chilopoda" family="Geophilidae" genus="Clinopodes" kingdom="Animalia" order="Geophilomorpha" pageId="16" pageNumber="191" phylum="Arthropoda" rank="species" species="skopljensis">
<emphasis id="B906EADBE32B7A04FD3CF9E9FC756C7B" bold="true" box="[731,968,1578,1605]" italics="true" pageId="16" pageNumber="191">Clinopodes skopljensis</emphasis>
(
<bibRefCitation id="EFE34B38E32B7A04FC31F9E9FBC66C7B" author="VERHOEFF K. W." box="[982,1147,1578,1605]" pageId="16" pageNumber="191" pagination="339 - 388" refId="ref24424" refString="VERHOEFF K. W. 1938. - Chilopoden-Studien zur Kenntnis der Epimorphen. Zoologische Jahrbucher, Abteilung fur Systematik, Okologie und Geographie der Tiere 71: 339 - 388." type="journal article" year="1938">Verhoeff, 1938</bibRefCitation>
)
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3686542E32B7A05FD42F9B2FEF86AD1" lastPageId="17" lastPageNumber="192" pageId="16" pageNumber="191" type="materials_examined">
<paragraph id="8BCD36C9E32B7A04FD42F9B2FB076C9D" blockId="16.[674,1211,1649,1699]" pageId="16" pageNumber="191">
<materialsCitation id="3B1A3C94E32B7A04FD42F9B2FB0A6CB7" box="[677,1207,1649,1673]" collectionCode="TYPE, MATERIAL, AND, LOCALITY" pageId="16" pageNumber="191" specimenCount="1" specimenCount-juvenile="1" typeStatus="holotype">
<collectionCode id="ED63AE0CE32B7A04FD42F9B2FC636CB9" box="[677,990,1649,1673]" pageId="16" pageNumber="191">TYPE MATERIAL AND TYPE LOCALITY</collectionCode>
. —
<specimenCount id="9D74FD40E32B7A04FBE3F9B2FBE06CB7" box="[1028,1117,1649,1673]" pageId="16" pageNumber="191" type="juvenile">1 juvenile</specimenCount>
,
<typeStatus id="54C9886BE32B7A04FB83F9B2FB0A6CB7" box="[1124,1207,1649,1673]" pageId="16" pageNumber="191" type="holotype">holotype</typeStatus>
</materialsCitation>
; from “Skoplje” (= Skopje [
<collectingCountry id="F3657659E32B7A04FC7EF948FBB56C9D" box="[921,1032,1675,1699]" name="Macedonia" pageId="16" pageNumber="191">Macedonia</collectingCountry>
]) (
<bibRefCitation id="EFE34B38E32B7A04FBC7F948FB126C9D" author="VERHOEFF K. W." box="[1056,1199,1675,1699]" pageId="16" pageNumber="191" pagination="339 - 388" refId="ref24424" refString="VERHOEFF K. W. 1938. - Chilopoden-Studien zur Kenntnis der Epimorphen. Zoologische Jahrbucher, Abteilung fur Systematik, Okologie und Geographie der Tiere 71: 339 - 388." type="journal article" year="1938">Verhoeff 1938</bibRefCitation>
).
</paragraph>
<paragraph id="8BCD36C9E32A7A05FF96FF14FEF86AD1" blockId="17.[113,325,215,239]" box="[113,325,215,239]" pageId="17" pageNumber="192">SYNONYMS. — None.</paragraph>
</subSubSection>
<subSubSection id="C3686542E32A7A05FF96FEC4FD3A6B57" pageId="17" pageNumber="192" type="description">
<paragraph id="8BCD36C9E32A7A05FF96FEC4FE166B21" blockId="17.[113,427,263,287]" box="[113,427,263,287]" pageId="17" pageNumber="192">NOMINAL SUBSPECIES. — None.</paragraph>
<paragraph id="8BCD36C9E32A7A05FF96FEF5FD3A6B57" blockId="17.[113,647,310,361]" pageId="17" pageNumber="192">
MAIN REFERENCES. —
<bibRefCitation id="EFE34B38E32A7A05FEDAFEF5FE566B70" author="VERHOEFF K. W." box="[317,491,310,334]" pageId="17" pageNumber="192" pagination="339 - 388" refId="ref24424" refString="VERHOEFF K. W. 1938. - Chilopoden-Studien zur Kenntnis der Epimorphen. Zoologische Jahrbucher, Abteilung fur Systematik, Okologie und Geographie der Tiere 71: 339 - 388." type="journal article" year="1938">Verhoeff 1938: 341</bibRefCitation>
(original description, as
<taxonomicName id="4C724D4AE32A7A05FF5CFE92FECE6B57" authorityName="Verhoeff" authorityYear="1938" box="[187,371,337,361]" class="Chilopoda" family="Geophilidae" genus="Geophilus" kingdom="Animalia" order="Geophilomorpha" pageId="17" pageNumber="192" phylum="Arthropoda" rank="species" species="skopljensis">
<emphasis id="B906EADBE32A7A05FF5CFE92FECE6B57" box="[187,371,337,361]" italics="true" pageId="17" pageNumber="192">Geophilus skopljensis</emphasis>
</taxonomicName>
);
<bibRefCitation id="EFE34B38E32A7A05FE60FE92FD8C6B57" author="ATTEMS C." box="[391,561,337,361]" pageId="17" pageNumber="192" pagination="50 - 149" refId="ref20136" refString="ATTEMS C. 1947. - Neue Geophilomorpha des Wiener Museums. Annalen des Naturhistorischen Museums in Wien 55: 50 - 149." type="journal article" year="1947">Attems 1947: 120</bibRefCitation>
(in key).
</paragraph>
</subSubSection>
<subSubSection id="C3686542E32A7A05FF96FE43FE156BA6" box="[113,424,384,408]" pageId="17" pageNumber="192" type="materials_examined">
<paragraph id="8BCD36C9E32A7A05FF96FE43FE156BA6" blockId="17.[113,424,384,408]" box="[113,424,384,408]" pageId="17" pageNumber="192">
<collectionCode id="ED63AE0CE32A7A05FF96FE43FE836BA9" box="[113,318,384,408]" pageId="17" pageNumber="192">MATERIAL EXAMINED</collectionCode>
. — None.
</paragraph>
</subSubSection>
<subSubSection id="C3686542E32A7A05FF96FE73FEEC693E" pageId="17" pageNumber="192" type="diagnosis">
<paragraph id="8BCD36C9E32A7A05FF96FE73FDF26888" blockId="17.[112,646,432,694]" pageId="17" pageNumber="192">
DIAGNOSIS. — A
<taxonomicName id="4C724D4AE32A7A05FEF1FE73FEC46BF6" authorityName="C. L. Koch" authorityYear="1847" box="[278,377,432,456]" class="Chilopoda" family="Geophilidae" genus="Clinopodes" kingdom="Animalia" order="Geophilomorpha" pageId="17" pageNumber="192" phylum="Arthropoda" rank="genus">
<emphasis id="B906EADBE32A7A05FEF1FE73FEC46BF6" box="[278,377,432,456]" italics="true" pageId="17" pageNumber="192">Clinopodes</emphasis>
</taxonomicName>
species with around 61 pairs of legs; denticles of the forcipular coxosternite moderately elongate, slightly wider than long; chitin-lines vanishing before reaching the condyles; even the largest sternal porefields on the posterior leg-bearing segments well behind the mid-length of the metasternite; all canals of the coxal organs opening through independent pores, more dense close to the lateral margins of the metasternites but most of them not covered by the latter, without a distinctly isolated pore. See also
<tableCitation id="C6F00372E32A7A05FEACFD5DFE2E6888" box="[331,403,670,694]" captionStart="TABLE" captionStartId="10.[110,121,525,542]" captionText="TABLE 3.— Differential characters between the species of Clinopodes C. L. Koch,1847.Data are from original descriptions,subsequent redescriptions based on specimens confidently identified correctly (see “Main references” in the text, under each species), and direct observations of representative specimens (see “Material examined” in the text, under each species). An asterisk (*) marks those species for which the range of variation is estimated based on 1-3 specimens only." pageId="17" pageNumber="192">Table 3</tableCitation>
and key to species.
</paragraph>
<paragraph id="8BCD36C9E32A7A05FF96FD0DFEEC693E" blockId="17.[112,646,718,768]" pageId="17" pageNumber="192">
GEOGRAPHIC DISTRIBUTION. —
<collectingCountry id="F3657659E32A7A05FE47FD0DFDB768D8" box="[416,522,718,742]" name="Macedonia" pageId="17" pageNumber="192">Macedonia</collectingCountry>
(only known from the
<typeStatus id="54C9886BE32A7A05FF2CFD2BFF48693E" box="[203,245,744,768]" pageId="17" pageNumber="192">type</typeStatus>
locality).
</paragraph>
</subSubSection>
<subSubSection id="C3686542E32A7A05FF96FCF1FE916F36" pageId="17" pageNumber="192" type="description">
<paragraph id="8BCD36C9E32A7A05FF96FCF1FE916F36" blockId="17.[111,649,818,1288]" pageId="17" pageNumber="192">
TAXONOMIC AND NOMENCLATURAL NOTES
<taxonomicName id="4C724D4AE32A7A05FF96FC92FEE96955" baseAuthorityName="Verhoeff" baseAuthorityYear="1938" box="[113,340,849,875]" class="Chilopoda" family="Geophilidae" genus="Clinopodes" kingdom="Animalia" order="Geophilomorpha" pageId="17" pageNumber="192" phylum="Arthropoda" rank="species" species="skopljensis">
<emphasis id="B906EADBE32A7A05FF96FC92FEE96955" box="[113,340,849,875]" italics="true" pageId="17" pageNumber="192">Clinopodes skopljensis</emphasis>
</taxonomicName>
was originally described by
<bibRefCitation id="EFE34B38E32A7A05FF94FCB2FEA669B2" author="VERHOEFF K. W." box="[115,283,881,908]" pageId="17" pageNumber="192" pagination="339 - 388" refId="ref24424" refString="VERHOEFF K. W. 1938. - Chilopoden-Studien zur Kenntnis der Epimorphen. Zoologische Jahrbucher, Abteilung fur Systematik, Okologie und Geographie der Tiere 71: 339 - 388." type="journal article" year="1938">Verhoeff (1938)</bibRefCitation>
as a species in the subgenus
<taxonomicName id="4C724D4AE32A7A05FDA4FCB2FF1A6995" authorityName="C. L. Koch" authorityYear="1847" class="Chilopoda" family="Geophilidae" genus="Clinopodes" kingdom="Animalia" order="Geophilomorpha" pageId="17" pageNumber="192" phylum="Arthropoda" rank="genus">
<emphasis id="B906EADBE32A7A05FDA4FCB2FF1A6995" italics="true" pageId="17" pageNumber="192">Clinopodes</emphasis>
</taxonomicName>
of the genus
<taxonomicName id="4C724D4AE32A7A05FED7FC52FE286995" authorityName="Leach" authorityYear="1814" box="[304,405,913,939]" class="Chilopoda" family="Geophilidae" genus="Geophilus" kingdom="Animalia" order="Geophilomorpha" pageId="17" pageNumber="192" phylum="Arthropoda" rank="genus">
<emphasis id="B906EADBE32A7A05FED7FC52FE286995" box="[304,405,913,939]" italics="true" pageId="17" pageNumber="192">Geophilus</emphasis>
</taxonomicName>
, and was transferred to
<taxonomicName id="4C724D4AE32A7A05FF97FC73FF5D69F4" authorityName="C. L. Koch" authorityYear="1847" box="[112,224,944,970]" class="Chilopoda" family="Geophilidae" genus="Clinopodes" kingdom="Animalia" order="Geophilomorpha" pageId="17" pageNumber="192" phylum="Arthropoda" rank="genus">
<emphasis id="B906EADBE32A7A05FF97FC73FF5D69F4" box="[112,224,944,970]" italics="true" pageId="17" pageNumber="192">Clinopodes</emphasis>
</taxonomicName>
by
<bibRefCitation id="EFE34B38E32A7A05FEEFFC73FE1A69F5" author="ATTEMS C." box="[264,423,944,971]" pageId="17" pageNumber="192" pagination="50 - 149" refId="ref20136" refString="ATTEMS C. 1947. - Neue Geophilomorpha des Wiener Museums. Annalen des Naturhistorischen Museums in Wien 55: 50 - 149." type="journal article" year="1947">Attems (1947)</bibRefCitation>
. No other specimens were identified, it was almost completely ignored with the exception of mere citations in faunistic catalogues (
<bibRefCitation id="EFE34B38E32A7A05FF91FBD3FF586E15" author="KOS I." box="[118,229,1040,1067]" pageId="17" pageNumber="192" pagination="353 - 360" refId="ref22283" refString="KOS I. 1992. - A review of the taxonomy, geographical distribution and ecology of the centipedes of Yugoslavia (Myriapoda, Chilopoda). Berichte des naturwissenschaftlich-medizinischen Vereins in Innsbruck 10 (suppl.): 353 - 360." type="journal article" year="1992">Kos 1992</bibRefCitation>
;
<bibRefCitation id="EFE34B38E32A7A05FF15FBD3FECC6E15" author="STOEV P." box="[242,369,1040,1067]" pageId="17" pageNumber="192" pagination="87 - 105" refId="ref23841" refString="STOEV P. 1997. - A check-list of the centipedes of the Balkan Peninsula with some taxonomic notes and a complete bibliography (Chilopoda). Entomologica Scandinavica 51 (Suppl.): 87 - 105." type="journal article" year="1997">Stoev 1997</bibRefCitation>
,
<bibRefCitation id="EFE34B38E32A7A05FE99FBD3FE756E15" author="STOEV P." box="[382,456,1040,1067]" pageId="17" pageNumber="192" pagination="93 - 107" refId="ref23905" refString="STOEV P. 2001 b. - On the centipedes (Chilopoda) of the Republic of Macedonia. Historia naturalis bulgarica 13: 93 - 107." type="journal article" year="2001">2001b</bibRefCitation>
), and its identity has been regarded as uncertain (
<bibRefCitation id="EFE34B38E32A7A05FE5DFBECFDF96E74" author="STOEV P." box="[442,580,1071,1098]" pageId="17" pageNumber="192" pagination="93 - 107" refId="ref23905" refString="STOEV P. 2001 b. - On the centipedes (Chilopoda) of the Republic of Macedonia. Historia naturalis bulgarica 13: 93 - 107." type="journal article" year="2001">Stoev 2001b</bibRefCitation>
). Even though the original description is incomplete for some characters and is based on a single juvenile, it suggests that
<taxonomicName id="4C724D4AE32A7A05FF10FB4CFEC56E96" baseAuthorityName="Verhoeff" baseAuthorityYear="1938" box="[247,376,1166,1193]" class="Chilopoda" family="Geophilidae" genus="Clinopodes" kingdom="Animalia" order="Geophilomorpha" pageId="17" pageNumber="192" phylum="Arthropoda" rank="species" species="skopljensis">
<emphasis id="B906EADBE32A7A05FF10FB4CFEC56E96" box="[247,376,1166,1193]" italics="true" pageId="17" pageNumber="192">C. skopljensis</emphasis>
</taxonomicName>
could actually be different from all other species of
<taxonomicName id="4C724D4AE32A7A05FE98FB6DFE526EF6" authorityName="C. L. Koch" authorityYear="1847" box="[383,495,1198,1224]" class="Chilopoda" family="Geophilidae" genus="Clinopodes" kingdom="Animalia" order="Geophilomorpha" pageId="17" pageNumber="192" phylum="Arthropoda" rank="genus">
<emphasis id="B906EADBE32A7A05FE98FB6DFE526EF6" box="[383,495,1198,1224]" italics="true" pageId="17" pageNumber="192">Clinopodes</emphasis>
</taxonomicName>
(see
<tableCitation id="C6F00372E32A7A05FDC1FB6DFDCA6EF7" box="[550,631,1198,1225]" captionStart="TABLE" captionStartId="10.[110,121,525,542]" captionText="TABLE 3.— Differential characters between the species of Clinopodes C. L. Koch,1847.Data are from original descriptions,subsequent redescriptions based on specimens confidently identified correctly (see “Main references” in the text, under each species), and direct observations of representative specimens (see “Material examined” in the text, under each species). An asterisk (*) marks those species for which the range of variation is estimated based on 1-3 specimens only." pageId="17" pageNumber="192">Table 3</tableCitation>
), and therefore it is provisionally maintained here as a distinct species.
</paragraph>
</subSubSection>
</treatment>
</document>