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<document ID-DOI="10.11865/zs.2022303" ID-ISSN="2095-6827" ID-Zenodo-Dep="7175859" ID-ZooBank="6DED5057-996E-4DD5-BDE3-A8C366E59BB9" approvalRequired="7" approvalRequired_for_textStreams="7" checkinTime="1665295167977" checkinUser="felipe" docAuthor="Lin, Yejie, Zhao, Huifeng, Koh, Joseph K H &amp; Li, Shuqiang" docDate="2022" docId="03D687D8AA45FFF5FF37FE70062DFA5F" docLanguage="en" docName="ZoolSyst.47.3.198270.pdf" docOrigin="Zoological Systematics 47 (3)" docSource="http://zoobank.org/6ded5057-996e-4dd5-bde3-a8c366e59bb9" docStyle="DocumentStyle:07E7829F32901B127B59E960199959BD.1:ZoolSyst.2014-.journal_article" docStyleId="07E7829F32901B127B59E960199959BD" docStyleName="ZoolSyst.2014-.journal_article" docStyleVersion="1" docTitle="Pandava banna Lin &amp; Li 2022, sp. nov." docType="treatment" docVersion="3" lastPageNumber="269" masterDocId="FFEFFFA0AA03FFB2FFBFFFCC026CFFAD" masterDocTitle="Taxonomy notes on twenty-eight spider species (Arachnida: Araneae) from Asia" masterLastPageNumber="270" masterPageNumber="198" pageNumber="268" updateTime="1665313746843" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Taxonomy notes on twenty-eight spider species (Arachnida: Araneae) from Asia</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Lin, Yejie</mods:namePart>
<mods:affiliation>Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Zhao, Huifeng</mods:namePart>
<mods:affiliation>Hebei Key Laboratory of Animal Diversity, College of Life Science, Langfang Normal University, Langfang 065000, China</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Koh, Joseph K H</mods:namePart>
<mods:affiliation>National Biodiversity Centre, National Parks Board, 259569, Singapore Corresponding author, E-mail: lisq @ ioz. ac. cn</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Li, Shuqiang</mods:namePart>
<mods:affiliation>Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
<mods:title>Zoological Systematics</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2022</mods:date>
<mods:detail type="volume">
<mods:number>47</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>3</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>198</mods:start>
<mods:end>270</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://zoobank.org/6ded5057-996e-4dd5-bde3-a8c366e59bb9</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.11865/zs.2022303</mods:identifier>
<mods:identifier type="ISSN">2095-6827</mods:identifier>
<mods:identifier type="Zenodo-Dep">7175859</mods:identifier>
<mods:identifier type="ZooBank">6DED5057-996E-4DD5-BDE3-A8C366E59BB9</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.7172312" ID-Zenodo-Dep="7172312" LSID="urn:lsid:plazi:treatment:03D687D8AA45FFF5FF37FE70062DFA5F" httpUri="http://treatment.plazi.org/id/03D687D8AA45FFF5FF37FE70062DFA5F" lastPageId="71" lastPageNumber="269" pageId="70" pageNumber="268">
<subSubSection box="[136,674,444,469]" pageId="70" pageNumber="268" type="nomenclature">
<paragraph blockId="70.[136,674,444,469]" box="[136,674,444,469]" pageId="70" pageNumber="268">
<heading box="[136,674,444,469]" centered="true" fontSize="10" level="2" pageId="70" pageNumber="268" reason="2">
<emphasis bold="true" box="[136,515,444,469]" pageId="70" pageNumber="268">
<taxonomicName authority="Lin &amp; Li" authorityName="Lin &amp; Li" authorityYear="2022" box="[136,418,444,468]" class="Arachnida" family="Titanoecidae" genus="Pandava" kingdom="Animalia" order="Araneae" pageId="70" pageNumber="268" phylum="Arthropoda" rank="species" species="banna" status="sp. nov.">
<emphasis bold="true" box="[136,309,444,468]" italics="true" pageId="70" pageNumber="268">Pandava banna</emphasis>
Lin &amp; Li
</taxonomicName>
,
<taxonomicNameLabel box="[430,515,445,469]" pageId="70" pageNumber="268" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation box="[544,667,444,468]" captionStart-0="Figure 63" captionStart-1="Figure 64" captionStart-2="Figure 65" captionStartId-0="68.[155,218,1995,2017]" captionStartId-1="69.[155,218,1994,2016]" captionStartId-2="70.[155,218,2024,2046]" captionTargetBox-0="[189,1395,801,1958]" captionTargetBox-1="[199,1392,726,1955]" captionTargetBox-2="[189,1398,550,1982]" captionTargetId-0="figure-244@68.[189,1399,801,1965]" captionTargetId-1="figure-248@69.[189,1398,720,1960]" captionTargetId-2="figure-72@70.[189,1398,550,1982]" captionTargetPageId-0="68" captionTargetPageId-1="69" captionTargetPageId-2="70" captionText-0="Figure 63. Pandava banna Lin &amp; Li, sp. nov., holotype male, left palp. A. Ventral view; B. Dorsal view. Abbreviations: MA—median apophysis; MLT—median lobe of tibial apophysis; PLT—prolateral lobe of tibial apophysis; RLT—retrolateral lobe of tibial apophysis; SD—sperm duct; ST—subtegulum; TP—Tegular process. Scale bars=0.1mm." captionText-1="Figure 64. Pandava banna Lin &amp; Li, sp. nov., holotype male, left palp. A. Prolateral view; B. Retrolateral view. Abbreviations: E— embolus; MA—median apophysis; MLT—median lobe of tibial apophysis; PLT—prolateral lobe of tibial apophysis; RLT— retrolateral lobe of tibial apophysis; SD—sperm duct; ST—subtegulum; TP—Tegular process. Scale bars=0.1mm." captionText-2="Figure 65. Pandava banna Lin &amp; Li, sp. nov., paratype female.A. Epigyne, ventral view; B. Vulva, dorsal view.Abbreviations: CD— copulatory duct; CO—copulatory opening; ER—epigynal rim; FD—fertilization duct; S—spermatheca. Scale bars=0.1mm." figureDoi-0="http://doi.org/10.5281/zenodo.7176006" figureDoi-1="http://doi.org/10.5281/zenodo.7176008" figureDoi-2="http://doi.org/10.5281/zenodo.7176010" httpUri-0="https://zenodo.org/record/7176006/files/figure.png" httpUri-1="https://zenodo.org/record/7176008/files/figure.png" httpUri-2="https://zenodo.org/record/7176010/files/figure.png" pageId="70" pageNumber="268">Figs 6365</figureCitation>
)
</heading>
</paragraph>
</subSubSection>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7176010" ID-Zenodo-Dep="7176010" httpUri="https://zenodo.org/record/7176010/files/figure.png" pageId="70" pageNumber="268" startId="70.[155,218,2024,2046]" targetBox="[189,1398,550,1982]" targetPageId="70">
<paragraph blockId="70.[155,1433,2024,2078]" pageId="70" pageNumber="268">
Figure 65.
<taxonomicName authority="Lin &amp; Li" authorityName="Lin &amp; Li" authorityYear="2022" box="[259,501,2024,2046]" class="Arachnida" family="Titanoecidae" genus="Pandava" kingdom="Animalia" order="Araneae" pageId="70" pageNumber="268" phylum="Arthropoda" rank="species" species="banna" status="sp. nov.">
<emphasis box="[259,410,2024,2045]" italics="true" pageId="70" pageNumber="268">Pandava banna</emphasis>
Lin &amp; Li
</taxonomicName>
,
<emphasis bold="true" box="[510,586,2025,2046]" pageId="70" pageNumber="268">
<taxonomicNameLabel box="[510,586,2025,2046]" pageId="70" pageNumber="268" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, paratype female. A. Epigyne, ventral view; B. Vulva, dorsal view. Abbreviations: CD— copulatory duct; CO—copulatory opening; ER—epigynal rim; FD—fertilization duct; S—spermatheca. Scale bars = 0.1 mm.
</paragraph>
</caption>
<subSubSection pageId="71" pageNumber="269" type="diagnosis">
<paragraph blockId="71.[136,1452,236,1058]" pageId="71" pageNumber="269">
Diagnosis. Males of the new species resemble all other
<taxonomicName box="[787,966,236,260]" pageId="71" pageNumber="269">
<emphasis box="[787,882,236,260]" italics="true" pageId="71" pageNumber="269">Pandava</emphasis>
species
</taxonomicName>
by the divided median apophysis and having the median lobe of the tibial apophysis longer than the retrolateral lobe. Females differ from other species by the long, coiled copulatory duct.
</paragraph>
<paragraph blockId="71.[136,1452,236,1058]" pageId="71" pageNumber="269">
Description. Male (
<typeStatus box="[405,505,340,364]" pageId="71" pageNumber="269">holotype</typeStatus>
). Total length 3.13. Carapace 1.53 long; 1.19 wide. Abdomen 1.52 long; 1.03 wide. Carapace brown with cephalic area dark brown. Chelicerae brown, with 3 promarginal and 2 retromarginal teeth. Sternum, labium and endites yellowish brown. Eye diameters and interdistances: AME 0.05; ALE 0.11; PME 0.11; PLE 0.10; AME AME 0.04; AMEALE 0.06; PMEPME 0.09; PMEPLE 0.12; AMEPME 0.10; ALEPLE 0.08. Legs brown. Leg measurements: I 5.32 (1.56, 1.94, 1.20, 0.62), II 4.38 (1.34, 1.53, 0.99, 0.52), III 3.61 (1.17, 1.21, 0.82, 0.41), IV 3.52 (1.10, 1.18, 0.80, 0.44). Abdomen beige, with brown pattern.
</paragraph>
<paragraph blockId="71.[136,1452,236,1058]" pageId="71" pageNumber="269">
Palp (
<figureCitation box="[252,377,548,572]" captionStart-0="Figure 63" captionStart-1="Figure 64" captionStartId-0="68.[155,218,1995,2017]" captionStartId-1="69.[155,218,1994,2016]" captionTargetBox-0="[189,1395,801,1958]" captionTargetBox-1="[199,1392,726,1955]" captionTargetId-0="figure-244@68.[189,1399,801,1965]" captionTargetId-1="figure-248@69.[189,1398,720,1960]" captionTargetPageId-0="68" captionTargetPageId-1="69" captionText-0="Figure 63. Pandava banna Lin &amp; Li, sp. nov., holotype male, left palp. A. Ventral view; B. Dorsal view. Abbreviations: MA—median apophysis; MLT—median lobe of tibial apophysis; PLT—prolateral lobe of tibial apophysis; RLT—retrolateral lobe of tibial apophysis; SD—sperm duct; ST—subtegulum; TP—Tegular process. Scale bars=0.1mm." captionText-1="Figure 64. Pandava banna Lin &amp; Li, sp. nov., holotype male, left palp. A. Prolateral view; B. Retrolateral view. Abbreviations: E— embolus; MA—median apophysis; MLT—median lobe of tibial apophysis; PLT—prolateral lobe of tibial apophysis; RLT— retrolateral lobe of tibial apophysis; SD—sperm duct; ST—subtegulum; TP—Tegular process. Scale bars=0.1mm." figureDoi-0="http://doi.org/10.5281/zenodo.7176006" figureDoi-1="http://doi.org/10.5281/zenodo.7176008" httpUri-0="https://zenodo.org/record/7176006/files/figure.png" httpUri-1="https://zenodo.org/record/7176008/files/figure.png" pageId="71" pageNumber="269">Figs 6364</figureCitation>
). Median apophysis conical, bifurcated. Sperm duct visible in middle of tegulum. Prolateral lobe of tibial apophysis bifurcated, median lobe of tibial apophysis longest, terminus slightly curved, retrolateral lobe of tibial apophysis semi-circular with notch in the middle.
</paragraph>
<paragraph blockId="71.[136,1452,236,1058]" pageId="71" pageNumber="269">Female (IZCAS-Ar43336). Total length 4.13. Carapace 1.73 long; 1.25 wide. Abdomen 2.31 long; 1.25 wide. Chelicerae with 2 promarginal and 2 retromarginal teeth. Eye diameters and interdistances: AME 0.07; ALE 0.12; PME 0.11; PLE 0.12; AMEAME 0.05; AMEALE 0.07; PMEPME 0.10; PMEPLE 0.13; AMEPME 0.10; ALEPLE 0.04. Leg measurements: I 4.25 (1.31, 1.60, 0.75, 0.59), II 3.69 (1.26, 1.31, 0.66, 0.46), III 3.05 (0.93, 1.20, 0.53, 0.39), IV 4.00 (1.00, 1.01, 0.91, 0.48). Habitus similar to that of male.</paragraph>
<paragraph blockId="71.[136,1452,236,1058]" pageId="71" pageNumber="269">
Female genitalia (
<figureCitation box="[392,477,825,850]" captionStart="Figure 65" captionStartId="70.[155,218,2024,2046]" captionTargetBox="[189,1398,550,1982]" captionTargetId="figure-72@70.[189,1398,550,1982]" captionTargetPageId="70" captionText="Figure 65. Pandava banna Lin &amp; Li, sp. nov., paratype female.A. Epigyne, ventral view; B. Vulva, dorsal view.Abbreviations: CD— copulatory duct; CO—copulatory opening; ER—epigynal rim; FD—fertilization duct; S—spermatheca. Scale bars=0.1mm." figureDoi="http://doi.org/10.5281/zenodo.7176010" httpUri="https://zenodo.org/record/7176010/files/figure.png" pageId="71" pageNumber="269">Fig. 65</figureCitation>
). Epigyne with wide plate, long, coiled copulatory duct. Single elongate spermatheca. Fertilization duct transverse.
</paragraph>
<paragraph blockId="71.[136,1452,236,1058]" pageId="71" pageNumber="269">
<materialsCitation collectingDate="2009-11-22" collectorName="Menglun Township, G &amp; G. Tang" country="China" county="Mengla County" elevation="627" latitude="21.9064" location="Xishuangbanna Dai Autonomous Prefecture" longLatPrecision="7" longitude="101.28" municipality="Material" pageId="71" pageNumber="269" specimenCount="6" specimenCount-female="2" specimenCount-male="4" stateProvince="Yunnan" typeStatus="holotype">
<collectingMunicipality box="[189,279,895,919]" pageId="71" pageNumber="269">Material</collectingMunicipality>
examined.
<typeStatus box="[415,514,895,919]" pageId="71" pageNumber="269">Holotype</typeStatus>
<specimenCount box="[526,546,895,918]" pageId="71" pageNumber="269" type="male"></specimenCount>
(IZCAS-Ar43331),
<collectingCountry box="[776,844,895,919]" name="China" pageId="71" pageNumber="269">China</collectingCountry>
:
<collectingRegion box="[858,942,895,918]" country="China" name="Yunnan" pageId="71" pageNumber="269">Yunnan</collectingRegion>
,
<location LSID="urn:lsid:plazi:treatment:03D687D8AA45FFF5FF37FE70062DFA5F:8EA06015AA44FFF5FC01FCB307CBFC3A" box="[958,1447,895,919]" country="China" county="Mengla County" latitude="21.9064" longLatPrecision="7" longitude="101.28" municipality="Material" name="Xishuangbanna Dai Autonomous Prefecture" pageId="71" pageNumber="269" stateProvince="Yunnan">Xishuangbanna Dai Autonomous Prefecture</location>
,
<collectingCounty box="[136,305,930,954]" pageId="71" pageNumber="269">Mengla County</collectingCounty>
,
<collectorName box="[316,558,930,954]" pageId="71" pageNumber="269">
<location LSID="urn:lsid:plazi:treatment:03D687D8AA45FFF5FF37FE70062DFA5F:8EA06015AA44FFF5FE83FC6E007CFC17" box="[316,528,930,954]" country="China" county="Mengla County" latitude="21.9064" longLatPrecision="7" longitude="101.28" municipality="Material" name="Menglun Township" pageId="71" pageNumber="269" stateProvince="Yunnan">Menglun Township</location>
, G
</collectorName>
213 road,
<location LSID="urn:lsid:plazi:treatment:03D687D8AA45FFF5FF37FE70062DFA5F:8EA06015AA44FFF5FD24FC6E0099FC17" box="[667,757,930,954]" country="China" county="Mengla County" latitude="21.9064" longLatPrecision="7" longitude="101.28" municipality="Material" name="Bamboo" pageId="71" pageNumber="269" stateProvince="Yunnan">Bamboo</location>
plantation (
<geoCoordinate box="[888,1008,930,954]" degrees="21.9064" direction="north" orientation="latitude" pageId="71" pageNumber="269" precision="5" value="21.9064">21.9064°N</geoCoordinate>
,
<geoCoordinate box="[1019,1149,930,954]" degrees="101.2800" direction="east" orientation="longitude" pageId="71" pageNumber="269" precision="5" value="101.28">101.2800°E</geoCoordinate>
, elev.
<quantity box="[1217,1285,930,954]" metricMagnitude="2" metricUnit="m" metricValue="6.27" pageId="71" pageNumber="269" unit="m" value="627.0">
<elevation box="[1217,1285,930,954]" metricMagnitude="2" metricUnit="m" metricValue="6.27" pageId="71" pageNumber="269" unit="m" value="627.0">627 m</elevation>
</quantity>
),
<date pageId="71" pageNumber="269" value="2009-11-22">
<collectingDate pageId="71" pageNumber="269" value="2009-11-22">22 November 2009</collectingDate>
</date>
,
<collectorName box="[203,288,965,989]" pageId="71" pageNumber="269">G. Tang</collectorName>
leg.
<typeStatus box="[341,448,965,989]" pageId="71" pageNumber="269">Paratypes</typeStatus>
.
<specimenCount box="[457,493,965,989]" pageId="71" pageNumber="269" type="male">3♂</specimenCount>
<specimenCount box="[493,524,965,989]" pageId="71" pageNumber="269" type="female">2♀</specimenCount>
(IZCAS-Ar43332Ar43336), same data as holotype
</materialsCitation>
.
</paragraph>
<paragraph blockId="71.[136,1452,236,1058]" box="[189,707,999,1024]" pageId="71" pageNumber="269">
Distribution. Known only from the
<typeStatus box="[568,614,1000,1024]" pageId="71" pageNumber="269">type</typeStatus>
locality.
</paragraph>
<paragraph blockId="71.[136,1452,236,1058]" box="[189,1070,1034,1058]" pageId="71" pageNumber="269">
Etymology. The specific name is a noun in apposition taken from the
<typeStatus box="[931,977,1034,1058]" pageId="71" pageNumber="269">type</typeStatus>
locality.
</paragraph>
<paragraph blockId="71.[136,1275,1110,1134]" box="[136,1275,1110,1134]" pageId="71" pageNumber="269">
<emphasis bold="true" box="[136,232,1110,1134]" inLineHeading="true" pageId="71" pageNumber="269" reason="1">Funding</emphasis>
This research was supported by the National Natural Science Foundation of
<collectingCountry box="[1073,1137,1110,1134]" name="China" pageId="71" pageNumber="269">China</collectingCountry>
(31972869).
</paragraph>
</subSubSection>
<subSubSection pageId="71" pageNumber="269" type="description">
<paragraph blockId="71.[136,1459,1186,1522]" pageId="71" pageNumber="269">
<emphasis bold="true" box="[136,357,1186,1210]" inLineHeading="true" pageId="71" pageNumber="269" reason="1">Acknowledgements</emphasis>
The manuscript benefited greatly from comments by Yuri Marusik, Anna Sestakova, and Yucheng Lin. Theo Blick checked the etymology. Sarah Crews checked the English of the final version. Yi Li (Yunnan,
<collectingCountry box="[1246,1314,1221,1245]" name="China" pageId="71" pageNumber="269">China</collectingCountry>
), Rixin Jiang (
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,
<collectingCountry box="[265,333,1255,1279]" name="China" pageId="71" pageNumber="269">China</collectingCountry>
), Yujie Zhao (
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,
<collectingCountry box="[575,643,1255,1279]" name="China" pageId="71" pageNumber="269">China</collectingCountry>
), Zhigang Chen (
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,
<collectingCountry box="[923,991,1255,1279]" name="China" pageId="71" pageNumber="269">China</collectingCountry>
), Jincheng Liu (
<collectingRegion box="[1162,1245,1255,1279]" country="China" name="Beijing" pageId="71" pageNumber="269">Beijing</collectingRegion>
,
<collectingCountry box="[1256,1324,1255,1279]" name="China" pageId="71" pageNumber="269">China</collectingCountry>
), Wei Wang (
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,
<collectingCountry box="[283,351,1290,1314]" name="China" pageId="71" pageNumber="269">China</collectingCountry>
), Fan Gao (
<collectingRegion box="[476,560,1290,1314]" country="China" name="Jiangsu" pageId="71" pageNumber="269">Jiangsu</collectingRegion>
,
<collectingCountry box="[572,640,1290,1314]" name="China" pageId="71" pageNumber="269">China</collectingCountry>
), Ruiyang Wang (
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,
<collectingCountry box="[927,995,1290,1314]" name="China" pageId="71" pageNumber="269">China</collectingCountry>
), Meiying Lin (
<collectingRegion box="[1163,1246,1290,1314]" country="China" name="Beijing" pageId="71" pageNumber="269">Beijing</collectingRegion>
,
<collectingCountry box="[1257,1325,1290,1314]" name="China" pageId="71" pageNumber="269">China</collectingCountry>
), Qianle Lu (
<collectingRegion box="[143,272,1325,1349]" country="China" name="Guangdong" pageId="71" pageNumber="269">Guangdong</collectingRegion>
,
<collectingCountry box="[285,353,1325,1349]" name="China" pageId="71" pageNumber="269">China</collectingCountry>
), Tongyao Jiang (IZCAS), Dongdong Wang (
<collectingRegion box="[848,929,1325,1349]" country="China" name="Hainan" pageId="71" pageNumber="269">Hainan</collectingRegion>
,
<collectingCountry box="[942,1010,1325,1349]" name="China" pageId="71" pageNumber="269">China</collectingCountry>
), Yu Wen (
<collectingRegion box="[1131,1212,1325,1349]" country="China" name="Jiangxi" pageId="71" pageNumber="269">Jiangxi</collectingRegion>
,
<collectingCountry box="[1225,1293,1325,1349]" name="China" pageId="71" pageNumber="269">China</collectingCountry>
), Zhiyuan Yao (Shenyang,
<collectingCountry box="[262,330,1359,1383]" name="China" pageId="71" pageNumber="269">China</collectingCountry>
), Zelun Li (
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,
<collectingCountry box="[543,611,1359,1383]" name="China" pageId="71" pageNumber="269">China</collectingCountry>
), Yunhu Mo (
<collectingRegion box="[751,832,1359,1383]" country="China" name="Hainan" pageId="71" pageNumber="269">Hainan</collectingRegion>
,
<collectingCountry box="[842,910,1359,1383]" name="China" pageId="71" pageNumber="269">China</collectingCountry>
), Yunmeng Wang (
<collectingRegion box="[1107,1190,1359,1383]" country="China" name="Beijing" pageId="71" pageNumber="269">Beijing</collectingRegion>
,
<collectingCountry box="[1199,1264,1359,1383]" name="China" pageId="71" pageNumber="269">China</collectingCountry>
) and Yiduo Wang (
<collectingRegion box="[143,272,1394,1418]" country="China" name="Guangdong" pageId="71" pageNumber="269">Guangdong</collectingRegion>
,
<collectingCountry box="[282,347,1394,1418]" name="China" pageId="71" pageNumber="269">China</collectingCountry>
) helped with field work. Jishen Wang (
<collectingRegion box="[755,841,1394,1418]" country="China" name="Yunnan" pageId="71" pageNumber="269">Yunnan</collectingRegion>
,
<collectingCountry box="[850,918,1394,1418]" name="China" pageId="71" pageNumber="269">China</collectingCountry>
), Jing Yuan (
<collectingRegion box="[1050,1121,1394,1418]" country="China" name="Anhui" pageId="71" pageNumber="269">Anhui</collectingRegion>
,
<collectingCountry box="[1130,1198,1394,1418]" name="China" pageId="71" pageNumber="269">China</collectingCountry>
), and Saiyu Hou (
<collectingRegion box="[1383,1454,1394,1418]" country="China" name="Anhui" pageId="71" pageNumber="269">Anhui</collectingRegion>
,
<collectingCountry box="[136,201,1429,1453]" name="China" pageId="71" pageNumber="269">China</collectingCountry>
) provided helpful suggestions. Fan Gao (
<collectingRegion box="[632,716,1429,1453]" country="China" name="Jiangsu" pageId="71" pageNumber="269">Jiangsu</collectingRegion>
,
<collectingCountry box="[726,794,1429,1453]" name="China" pageId="71" pageNumber="269">China</collectingCountry>
), Dongdong Wang (
<collectingRegion box="[1001,1082,1429,1453]" country="China" name="Hainan" pageId="71" pageNumber="269">Hainan</collectingRegion>
,
<collectingCountry box="[1091,1159,1429,1453]" name="China" pageId="71" pageNumber="269">China</collectingCountry>
), Yiduo Wang (
<collectingRegion box="[1319,1448,1429,1453]" country="China" name="Guangdong" pageId="71" pageNumber="269">Guangdong</collectingRegion>
,
<collectingCountry box="[136,201,1463,1487]" name="China" pageId="71" pageNumber="269">China</collectingCountry>
) and Qianle Lu (
<collectingRegion box="[394,523,1463,1487]" country="China" name="Guangdong" pageId="71" pageNumber="269">Guangdong</collectingRegion>
,
<collectingCountry box="[537,602,1463,1487]" name="China" pageId="71" pageNumber="269">China</collectingCountry>
) took the photographs. We also thank Changteng Yang, Tao Zhang and their colleagues, Nanling National Forest Park Administration, for their help in our field work.
</paragraph>
</subSubSection>
</treatment>
</document>