667 lines
80 KiB
XML
667 lines
80 KiB
XML
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<mods:title id="547257D4275272136F80EF310C4CA339">On the identity of Cyphonoxia maljuzhenkoi Zaitzev, 1928 (Coleoptera Scarabaeidae: Melolonthinae)</mods:title>
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<taxonomicName id="4C7F4D0CA272FFFCFF4505DA77BFFD53" authority="(Zaitzev, 1928)" authorityName="Bezděk & Kalashian & Ghrejyan" authorityYear="2019" baseAuthorityName="Zaitzev" baseAuthorityYear="1928" box="[151,765,723,750]" class="Insecta" family="Melolonthidae" genus="Anoxia" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="440" phylum="Arthropoda" rank="species" species="maljuzhenkoi" status="comb. nov." subGenus="Protanoxia">
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<emphasis id="B90BEA9DA272FFFCFF4505DA75ADFD50" bold="true" box="[151,239,724,750]" italics="true" pageId="1" pageNumber="440">Anoxia</emphasis>
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(
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<emphasis id="B90BEA9DA272FFFCFED205DA74C5FD50" bold="true" box="[256,391,724,750]" italics="true" pageId="1" pageNumber="440">Protanoxia</emphasis>
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)
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<emphasis id="B90BEA9DA272FFFCFE4B05DD777CFD50" bold="true" box="[409,574,723,750]" italics="true" pageId="1" pageNumber="440">maljuzhenkoi</emphasis>
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(
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<bibRefCitation id="EFEE4B7EA272FFFCFD9F05DD77B6FD53" author="Zaitzev, F. A." box="[589,756,723,749]" pageId="1" pageNumber="440" pagination="373 - 397" refId="ref4775" refString="Zaitzev, F. A. (1928) Obzor khrushchey Kavkaza v svyazi s ikh rasprostraneniem v kraye. Zur Verbreitung der Melolonthinen in den Kaukasuslandern. Izvestiya Tifliskogo Gosudarstvennogo Politekhnicheskogo Instituta, 3, 373 - 397. [In Russian, German summary]" type="journal article" year="1928">Zaitzev, 1928</bibRefCitation>
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<taxonomicNameLabel id="A23857E6A272FFFCFCD605DA7698FD53" box="[772,986,723,750]" pageId="1" pageNumber="440" rank="species">new combination</taxonomicNameLabel>
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<subSubSection id="C3656504A272FFFCFF4505F97436FCAC" box="[151,372,759,786]" pageId="1" pageNumber="440" type="description">
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<paragraph id="8BC0368FA272FFFCFF4505F97436FCAC" blockId="1.[151,986,723,786]" box="[151,372,759,786]" pageId="1" pageNumber="440">
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(
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<figureCitation id="13442A0AA272FFFCFF4D05F9742EFCAC" box="[159,364,759,786]" captionStart-0="FIGURES 1–7" captionStart-1="FIGURES 8–16" captionStartId-0="2.[151,264,1920,1945]" captionStartId-1="3.[151,264,1867,1892]" captionTargetBox-0="[205,1358,203,1839]" captionTargetBox-1="[208,1380,206,1822]" captionTargetId-0="figure@2.[164,1423,181,1894]" captionTargetId-1="figure@3.[183,1404,181,1842]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionText-0="FIGURES 1–7. Anoxia (Protanoxia) maljuzhenkoi (Zaitzev, 1928) from “Vordan Karmir” State Sanctuary, Armenia. 1, male (length 22.5 mm), dorsal view; 2, male, lateral view; 3, male aedeagus, lateral view; 4, male aedeagus, dorsal view; 5, female (length 21.4 mm), dorsal view; 6, female, lateral view; 7, female protibia, ventral view. Not to scale." captionText-1="FIGURES 8–16. Comparison of Anoxia (Protanoxia) species. 8, Lectotype of Anoxia maljuzhenkoi (Zaitzev, 1928), male, dorsal view; 9, A. maljuzhenkoi lectotype labels; 10, A. maljuzhenkoi base of pronotum and elytra; 11, A. ciliciensis Baraud, 1989 base of pronotum and elytra; 12, A. maljuzhenkoi pygidium; 13, A. ciliciensis pygidium; 14, A. smyrnensis Petrovitz, 1965 terminal maxillary palpomere; 15, A. maljuzhenkoi terminal maxillary palpomere; 16, A. ciliciensis terminal maxillary palpomere. Not to scale." figureDoi-0="http://doi.org/10.5281/zenodo.3442500" figureDoi-1="http://doi.org/10.5281/zenodo.3442502" httpUri-0="https://zenodo.org/record/3442500/files/figure.png" httpUri-1="https://zenodo.org/record/3442502/files/figure.png" pageId="1" pageNumber="440">Figs. 1–10, 12, 15</figureCitation>
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)
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<subSubSection id="C3656504A272FFFCFF450432776EFC2F" pageId="1" pageNumber="440" type="reference_group">
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<paragraph id="8BC0368FA272FFFCFF45043271D3FCEA" blockId="1.[151,1437,827,914]" box="[151,1169,827,852]" pageId="1" pageNumber="440">
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<emphasis id="B90BEA9DA272FFFCFF45043274DDFCEA" box="[151,415,828,852]" italics="true" pageId="1" pageNumber="440">Cyphonoxia maljuzhenkoi</emphasis>
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<treatmentCitation id="0ADE109EA272FFFCFE7704357720FCEA" author="Zaitzev, F. A." box="[421,610,827,852]" page="397" pageId="1" pageNumber="440" year="1928">
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<bibRefCitation id="EFEE4B7EA272FFFCFE7704357720FCEA" author="Zaitzev, F. A." box="[421,610,827,852]" pageId="1" pageNumber="440" pagination="373 - 397" refId="ref4775" refString="Zaitzev, F. A. (1928) Obzor khrushchey Kavkaza v svyazi s ikh rasprostraneniem v kraye. Zur Verbreitung der Melolonthinen in den Kaukasuslandern. Izvestiya Tifliskogo Gosudarstvennogo Politekhnicheskogo Instituta, 3, 373 - 397. [In Russian, German summary]" type="journal article" year="1928">Zaitzev, 1928: 397</bibRefCitation>
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(original description);
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<bibRefCitation id="EFEE4B7EA272FFFCFC9F043571CFFCEA" author="Iablokoff-Khnzorian, S. M." box="[845,1165,827,852]" pageId="1" pageNumber="440" pagination="2" refId="ref4438" refString="Iablokoff-Khnzorian, S. M. (1967) Nasekomye zhestkokrylye. Tom VI. Plastinchatousye (Scarabaeoidea). Fauna Armyanskoy SSR. [Insects, beetles. Vol. 6. Scarabaeoidea. Fauna of the Armenian SSR]. Izdatel'stvo Akademii Nauk Armyanskoy SSR, Yerevan, Armenia, 223 + [2] pp. [In Russian, Armenian title]" type="journal article" year="1967">Iablokoff-Khnzorian 1967: 176</bibRefCitation>
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<emphasis id="B90BEA9DA272FFFCFF45045574EFFCCC" box="[151,429,858,882]" italics="true" pageId="1" pageNumber="440">Cryptotrogus maljuzhenkoi</emphasis>
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:
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<treatmentCitation id="0ADE109EA272FFFCFE6E045477D8FCCD" author="Medvedev, S. I." box="[444,666,858,883]" page="174" pageId="1" pageNumber="440" year="1951">
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;
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<treatmentCitation id="0ADE109EA272FFFCFD7404547621FCCD" author="Baraud, J." box="[678,867,858,883]" page="441" pageId="1" pageNumber="440" year="1992">
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<bibRefCitation id="EFEE4B7EA272FFFCFD7404547621FCCD" author="Baraud, J." box="[678,867,858,883]" pageId="1" pageNumber="440" refId="ref4186" refString="Baraud, J. (1992) Coleopteres Scarabaeoidea d'Europe. Faune de France et regions limitrophies, 78. Federation Francaise des Societes de Sciences Naturelles & Societe Linneenne de Lyon, Paris, 856 pp." type="book" year="1992">Baraud 1992: 441</bibRefCitation>
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;
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<bibRefCitation id="EFEE4B7EA272FFFCFCBD0454716CFCCD" author="Bezdek, A." box="[879,1070,858,883]" pageId="1" pageNumber="440" pagination="191 - 198" refId="ref4224" refString="Bezdek, A. (2006) Tribus Melolonthini. In: Lobl, I. & Smetana, A. (Eds.), Catalogue of Palearctic Coleoptera. Volume 3. Scara- baeoidea - Scirtoidea - Dascilloidea - Buprestoidea - Byrrhoidea. Apollo Books, Senstrup, pp. 191 - 198." type="book chapter" year="2006">Bezděk 2006: 193</bibRefCitation>
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<bibRefCitation id="EFEE4B7EA272FFFCFBE804547040FCCD" author="Shokhin, I. V." box="[1082,1282,858,883]" pageId="1" pageNumber="440" pagination="105 - 185" refId="ref4689" refString="Shokhin, I. V. (2007) Materialy k faune plastinchatousykh zhukov (Coleoptera, Scarabaeoidea) Yuzhnoy Rossii. Contribution to the fauna of lamellicorn beetles (Coleoptera, Scarabaeoidea) of southern Russia, with some nomenclatural changes in the family Scarabaeidae. Caucasian Entomological Bulletin, 3, 105 - 185. [In Russian, English abstract] https: // doi. org / 10.23885 / 1814 - 3326 - 2007 - 3 - 2 - 105 - 185" type="journal article" year="2007">Shokhin 2007: 154</bibRefCitation>
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;
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<treatmentCitation id="0ADE109EA272FFFCFADC045475B2FC2F" author="Bezdek, A." page="229" pageId="1" pageNumber="440" year="2016">
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<bibRefCitation id="EFEE4B7EA272FFFCFADC045475B2FC2F" author="Bezdek, A." pageId="1" pageNumber="440" pagination="226 - 236" refId="ref4280" refString="Bezdek, A. (2016) Tribus Melolonthini. In: Lobl, I & Lobl, D. (Eds.), Catalogue of Palaearctic Coleoptera. Volume 3. Scara- baeoidea - Scirtoidea - Dascilloidea - Buprestoidea - Byrrhoidea. Revised and Updated Edition. E. J. Brill, Leiden, pp. 226 - 236." type="book chapter" year="2016">Bezděk 2016: 229</bibRefCitation>
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<treatmentCitation id="0ADE109EA272FFFCFF290477776AFC2F" author="Montreuil, O. & Keith, D." box="[251,552,889,914]" page="408" pageId="1" pageNumber="440" year="2017">
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<bibRefCitation id="EFEE4B7EA272FFFCFF290477776AFC2F" author="Montreuil, O. & Keith, D." box="[251,552,889,914]" pageId="1" pageNumber="440" pagination="387 - 412" refId="ref4582" refString="Montreuil, O. & Keith, D. (2017) Les Cryptotrogus Kraatz de la faune d'Iran et des regions limitrophes (Coleoptera: Scarabae- oidea: Melolonthidae). Annales de la Societe Entomologique de France (New Series), 53, 387 - 412. https: // doi. org / 10.1080 / 00379271.2017.1385421" type="journal article" year="2017">Montreuil & Keith 2017: 408</bibRefCitation>
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locality.
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“Nachkraj: Alishar, distr. Sharur” [=
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,
|
||
<location id="8EA06054A272FFFCFA9604B170DAFC67" LSID="urn:lsid:plazi:treatment:03D68799A272FFF8FF4505DA71F3FBE4:8EA06054A272FFFCFA9604B170DAFC67" box="[1348,1432,959,985]" county="Autonomous Republic" latitude="39.533333" longLatPrecision="1168" longitude="44.916668" municipality="Sharur district" name="Alishar" pageId="1" pageNumber="440" stateProvince="Naxcivan">Alishar</location>
|
||
,
|
||
<geoCoordinate id="EE4B5048A272FFFCFF4504EA75BAFC40" box="[151,248,996,1022]" degrees="39" direction="north" minutes="32" orientation="latitude" pageId="1" pageNumber="440" precision="925" value="39.533333">39°32’N</geoCoordinate>
|
||
<geoCoordinate id="EE4B5048A272FFFCFF2D04ED7420FC40" box="[255,354,995,1022]" degrees="44" direction="east" minutes="55" orientation="longitude" pageId="1" pageNumber="440" precision="925" value="44.916668">44°55’E</geoCoordinate>
|
||
]
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BC0368FA272FFFCFF1503067739FAA0" blockId="1.[151,1437,959,2030]" pageId="1" pageNumber="440">
|
||
<emphasis id="B90BEA9DA272FFFCFF15030674B1FB9C" bold="true" box="[199,499,1031,1058]" pageId="1" pageNumber="440">Type material examined.</emphasis>
|
||
<materialsCitation id="3B173CD2A272FFFCFE2903067739FAA0" ID-GBIF-Occurrence="2413161128" collectionCode="IZAY" collectorName="A. Bezdek & M. Kalashian & T. Ghrejyan" determinerName="Zaitzev & Khnzorian" pageId="1" pageNumber="440" specimenCount="5" specimenCount-male="5" typeStatus="lectotype">
|
||
<typeStatus id="54C4882DA272FFFCFE290306772CFB9C" box="[507,622,1032,1058]" pageId="1" pageNumber="440" type="lectotype">Lectotype</typeStatus>
|
||
,
|
||
<specimenCount id="9D79FD06A272FFFCFDA9030677D0FB9C" box="[635,658,1032,1058]" pageId="1" pageNumber="440" type="male">♂</specimenCount>
|
||
, present designation (
|
||
<collectionCode id="ED6EAE4AA272FFFCFC5903067693FB9C" box="[907,977,1032,1058]" pageId="1" pageNumber="440">IZAY</collectionCode>
|
||
), labelled: “Erivansk. g. [printed] | Alis- hari [handwritten, in Cyrillic script] | Maljushenco [printed] || Meganoxia
|
||
<specimenCount id="9D79FD06A272FFFCFC0A032276AFFBF8" box="[984,1005,1068,1094]" pageId="1" pageNumber="440" type="male">♂</specimenCount>
|
||
|
|
||
<taxonomicName id="4C7F4D0CA272FFFCFBD303257138FBFB" authority="Reitt." authorityName="Reitt." box="[1025,1146,1067,1093]" class="Insecta" family="Melolonthidae" genus="Cryptotrogus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="440" phylum="Arthropoda" rank="species" species="orita">orita Reitt.</taxonomicName>
|
||
!! [handwritten] |
|
||
<determinerName id="7E1CDA4BA272FFFCFA9A032570DEFBFB" box="[1352,1436,1067,1093]" pageId="1" pageNumber="440">Zaitzev</determinerName>
|
||
det. [printed] || Alishary | Maljuzhenko [Iablokoff-Khnzorian’s handwriting, in Cyrillic script] ||
|
||
<taxonomicName id="4C7F4D0CA272FFFCFB2A035E70CEFBD4" authorityName="Kraatz" authorityYear="1888" box="[1272,1420,1104,1130]" class="Insecta" family="Melolonthidae" genus="Cryptotrogus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="440" phylum="Arthropoda" rank="genus">Cryptotrogus</taxonomicName>
|
||
| maluzhenkoi [Iablokoff-Khnzorian’s handwriting] ||
|
||
<taxonomicName id="4C7F4D0CA272FFFCFD32037D762BFB33" authorityName=", Iablokoff-Khnzorian" authorityYear="1967" box="[736,873,1139,1165]" class="Insecta" family="Melolonthidae" genus="Cyphonoxia" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="440" phylum="Arthropoda" rank="genus">Cyphonoxia</taxonomicName>
|
||
| maluzhenkoi Zaitz. [handwritten]
|
||
<determinerName id="7E1CDA4BA272FFFCFB25037D702FFB33" box="[1271,1389,1139,1165]" pageId="1" pageNumber="440">Khnzorian</determinerName>
|
||
det. [printed] ||
|
||
<taxonomicName id="4C7F4D0CA272FFFCFEC803997707FB0F" authorityName="Zaitzev" authorityYear="1928" box="[282,581,1175,1201]" class="Insecta" family="Melolonthidae" genus="Cyphonoxia" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="440" phylum="Arthropoda" rank="species" species="maljuzhenkoi">Cyphonoxia maljuzhenkoi</taxonomicName>
|
||
|
|
||
<typeStatus id="54C4882DA272FFFCFDB203967657FB0C" box="[608,789,1176,1202]" pageId="1" pageNumber="440" type="lectotype">LECTOTYPUS</typeStatus>
|
||
,
|
||
<specimenCount id="9D79FD06A272FFFCFCF60396767BFB0C" box="[804,825,1176,1202]" pageId="1" pageNumber="440" type="male">♂</specimenCount>
|
||
|
|
||
<collectorName id="268A5359A272FFFCFC810396768DFB0F" box="[851,975,1175,1202]" pageId="1" pageNumber="440">A. Bezděk</collectorName>
|
||
,
|
||
<collectorName id="268A5359A272FFFCFC0C0396713EFB0F" box="[990,1148,1175,1202]" pageId="1" pageNumber="440">M. Kalashian</collectorName>
|
||
, |
|
||
<collectorName id="268A5359A272FFFCFB4803967060FB0F" box="[1178,1314,1175,1202]" pageId="1" pageNumber="440">T. Ghrejyan</collectorName>
|
||
des. 2019 [printed, red label]” (
|
||
<figureCitation id="13442A0AA272FFFCFE5003B574AEFB68" box="[386,492,1211,1238]" captionStart="FIGURES 8–16" captionStartId="3.[151,264,1867,1892]" captionTargetBox="[208,1380,206,1822]" captionTargetId="figure@3.[183,1404,181,1842]" captionTargetPageId="3" captionText="FIGURES 8–16. Comparison of Anoxia (Protanoxia) species. 8, Lectotype of Anoxia maljuzhenkoi (Zaitzev, 1928), male, dorsal view; 9, A. maljuzhenkoi lectotype labels; 10, A. maljuzhenkoi base of pronotum and elytra; 11, A. ciliciensis Baraud, 1989 base of pronotum and elytra; 12, A. maljuzhenkoi pygidium; 13, A. ciliciensis pygidium; 14, A. smyrnensis Petrovitz, 1965 terminal maxillary palpomere; 15, A. maljuzhenkoi terminal maxillary palpomere; 16, A. ciliciensis terminal maxillary palpomere. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.3442502" httpUri="https://zenodo.org/record/3442502/files/figure.png" pageId="1" pageNumber="440">Figs. 8, 9</figureCitation>
|
||
);
|
||
<typeStatus id="54C4882DA272FFFCFDD003B577DEFB6B" box="[514,668,1211,1237]" pageId="1" pageNumber="440" type="paralectotype">paralectotype</typeStatus>
|
||
,
|
||
<specimenCount id="9D79FD06A272FFFCFD7403B277F9FB68" box="[678,699,1212,1238]" pageId="1" pageNumber="440" type="male">♂</specimenCount>
|
||
(
|
||
<collectionCode id="ED6EAE4AA272FFFCFD1B03B27653FB68" box="[713,785,1212,1238]" pageId="1" pageNumber="440">IZAY</collectionCode>
|
||
): “Erivansk. g. [printed] | Alishari [handwritten, in Cyrillic script] | Maljushenco [printed] ||
|
||
<taxonomicName id="4C7F4D0CA272FFFCFDD603D1766EFB47" authorityName="Zaitzev" authorityYear="1928" box="[516,812,1247,1273]" class="Insecta" family="Melolonthidae" genus="Cyphonoxia" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="440" phylum="Arthropoda" rank="species" species="maljuzhenkoi">Cyphonoxia maljuzhenkoi</taxonomicName>
|
||
|
|
||
<typeStatus id="54C4882DA272FFFCFCED03EE717FFB44" box="[831,1085,1248,1274]" pageId="1" pageNumber="440" type="paralectotype">PARALECTOTYPUS</typeStatus>
|
||
,
|
||
<specimenCount id="9D79FD06A272FFFCFB9A03EE711FFB44" box="[1096,1117,1248,1274]" pageId="1" pageNumber="440" type="male">♂</specimenCount>
|
||
|
|
||
<collectorName id="268A5359A272FFFCFBBC03EE71A5FB47" box="[1134,1255,1247,1274]" pageId="1" pageNumber="440">A. Bezděk</collectorName>
|
||
,
|
||
<collectorName id="268A5359A272FFFCFB2003EE70CEFB47" box="[1266,1420,1247,1274]" pageId="1" pageNumber="440">M. Kalashian</collectorName>
|
||
, |
|
||
<collectorName id="268A5359A272FFFCFF45020A745EFAA3" box="[151,284,1283,1310]" pageId="1" pageNumber="440">T. Ghrejyan</collectorName>
|
||
des. 2019 [printed, red label]”.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="8BC0368FA272FFFCFF1502297174FA10" blockId="1.[151,1437,959,2030]" pageId="1" pageNumber="440">
|
||
<emphasis id="B90BEA9DA272FFFCFF150229776EFAFC" bold="true" box="[199,556,1319,1346]" pageId="1" pageNumber="440">Additional material examined</emphasis>
|
||
(
|
||
<specimenCount id="9D79FD06A272FFFCFDEE02267785FAFC" box="[572,711,1319,1346]" pageId="1" pageNumber="440" type="generic">3 specimens</specimenCount>
|
||
).
|
||
<materialsCitation id="3B173CD2A272FFFCFD090229749AFA34" ID-GBIF-Occurrence="2416712877" collectingDate="2017-06-13" collectionCode="NMPC" country="Armenia" county="Arazap" elevation="853" latitude="40.06333" location="Jan Sumpich" longLatPrecision="19" longitude="44.129444" municipality="Araratskaya Koshenil Reserve" pageId="1" pageNumber="440" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Armavir">
|
||
<emphasis id="B90BEA9DA272FFFCFD090229760CFAFC" bold="true" box="[731,846,1319,1346]" pageId="1" pageNumber="440">
|
||
<collectingCountry id="F368761FA272FFFCFD090229760BFAFC" box="[731,841,1319,1346]" name="Armenia" pageId="1" pageNumber="440">Armenia</collectingCountry>
|
||
:
|
||
</emphasis>
|
||
<collectingCounty id="62A14E03A272FFFCFC87022676E4FAFC" box="[853,934,1320,1346]" pageId="1" pageNumber="440">Arazap</collectingCounty>
|
||
environs,
|
||
<quantity id="4C879B6AA272FFFCFBC802267123FAFF" box="[1050,1121,1319,1346]" metricMagnitude="4" metricUnit="m" metricValue="1.2" pageId="1" pageNumber="440" unit="km" value="12.0">
|
||
<locationDeviation id="9A722FADA272FFFCFBC802267123FAFF" box="[1050,1121,1319,1346]" metricMagnitude="4" metricUnit="m" metricValue="1.2" pageId="1" pageNumber="440" unit="km" value="12.0">12 km</locationDeviation>
|
||
</quantity>
|
||
SE of
|
||
<collectingRegion id="49BBF86DA272FFFCFB790229704EFAFF" box="[1195,1292,1319,1345]" country="Armenia" name="Armavir" pageId="1" pageNumber="440">Armavir</collectingRegion>
|
||
,
|
||
<collectingMunicipality id="6BA4ACF5A272FFFCFAC70229741FFAD8" pageId="1" pageNumber="440">Araratskaya Koshenil Reserve</collectingMunicipality>
|
||
[= “Vordan Karmir” State Sanctuary], salt marsh,
|
||
<geoCoordinate id="EE4B5048A272FFFCFC460242714DFAD8" box="[916,1039,1356,1382]" degrees="40" direction="north" minutes="3" orientation="latitude" pageId="1" pageNumber="440" precision="15" seconds="48" value="40.06333">40°3’48”N</geoCoordinate>
|
||
<geoCoordinate id="EE4B5048A272FFFCFBC4024571D2FADB" box="[1046,1168,1355,1381]" degrees="44" direction="east" minutes="7" orientation="longitude" pageId="1" pageNumber="440" precision="15" seconds="46" value="44.129444">44°7’46”E</geoCoordinate>
|
||
,
|
||
<quantity id="4C879B6AA272FFFCFB4E024571A7FAD8" box="[1180,1253,1355,1382]" metricMagnitude="2" metricUnit="m" metricValue="8.53" pageId="1" pageNumber="440" unit="m" value="853.0">
|
||
<elevation id="0052D1BCA272FFFCFB4E024571A7FAD8" box="[1180,1253,1355,1382]" metricMagnitude="2" metricUnit="m" metricValue="8.53" pageId="1" pageNumber="440" unit="m" value="853.0">853 m</elevation>
|
||
</quantity>
|
||
,
|
||
<date id="FFC1104FA272FFFCFB2302457029FADB" box="[1265,1387,1355,1381]" pageId="1" pageNumber="440" value="2017-06-13">
|
||
<collectingDate id="EF85E9A7A272FFFCFB2302457029FADB" box="[1265,1387,1355,1381]" pageId="1" pageNumber="440" value="2017-06-13">13.vi.2017</collectingDate>
|
||
</date>
|
||
,
|
||
<location id="8EA06054A272FFFCFAA5024275BFFA34" LSID="urn:lsid:plazi:treatment:03D68799A272FFF8FF4505DA71F3FBE4:8EA06054A272FFFCFAA5024275BFFA34" country="Armenia" county="Arazap" latitude="40.06333" longLatPrecision="19" longitude="44.129444" municipality="Araratskaya Koshenil Reserve" name="Jan Sumpich" pageId="1" pageNumber="440" stateProvince="Armavir">Jan Šumpich</location>
|
||
,
|
||
<specimenCount id="9D79FD06A272FFFCFED8027E7474FA34" box="[266,310,1392,1418]" pageId="1" pageNumber="440" type="male">1 ♂</specimenCount>
|
||
<specimenCount id="9D79FD06A272FFFCFEEC027E7428FA34" box="[318,362,1392,1418]" pageId="1" pageNumber="440" type="female">1 ♀</specimenCount>
|
||
(
|
||
<collectionCode id="ED6EAE4AA272FFFCFEA8027E7492FA34" box="[378,464,1392,1418]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="1" pageNumber="440">NMPC</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B173CD2A272FFFCFE31027E7174FA10" ID-GBIF-Occurrence="2416712865" collectingDate="2017-06-13" collectionCode="IECA" country="Armenia" county="Arazap" elevation="853" latitude="40.06333" location="Araratskaya Koshenil Reserve" longLatPrecision="19" longitude="44.129444" municipality="Araratskaya Koshenil Reserve" pageId="1" pageNumber="440" specimenCount="1" specimenCount-female="1" specimenCount-male="1" stateProvince="Armavir">
|
||
Arazap environs,
|
||
<collectingRegion id="49BBF86DA272FFFCFD790261764EFA37" box="[683,780,1391,1417]" country="Armenia" name="Armavir" pageId="1" pageNumber="440">Armavir</collectingRegion>
|
||
,
|
||
<location id="8EA06054A272FFFCFCC50261712DFA34" LSID="urn:lsid:plazi:treatment:03D68799A272FFF8FF4505DA71F3FBE4:8EA06054A272FFFCFCC50261712DFA34" box="[791,1135,1391,1418]" country="Armenia" county="Arazap" latitude="40.06333" longLatPrecision="19" longitude="44.129444" municipality="Araratskaya Koshenil Reserve" name="Araratskaya Koshenil Reserve" pageId="1" pageNumber="440" stateProvince="Armavir">Araratskaya Koshenil Reserve</location>
|
||
[= “Vordan Karmir” State Sanctuary], salt marsh,
|
||
<geoCoordinate id="EE4B5048A272FFFCFE4D029A7758FA10" box="[415,538,1428,1454]" degrees="40" direction="north" minutes="3" orientation="latitude" pageId="1" pageNumber="440" precision="15" seconds="48" value="40.06333">40°3’48”N</geoCoordinate>
|
||
<geoCoordinate id="EE4B5048A272FFFCFDF3029D77D9FA13" box="[545,667,1427,1453]" degrees="44" direction="east" minutes="7" orientation="longitude" pageId="1" pageNumber="440" precision="15" seconds="46" value="44.129444">44°7’46”E</geoCoordinate>
|
||
,
|
||
<quantity id="4C879B6AA272FFFCFD75029D77B2FA10" box="[679,752,1427,1454]" metricMagnitude="2" metricUnit="m" metricValue="8.53" pageId="1" pageNumber="440" unit="m" value="853.0">
|
||
<elevation id="0052D1BCA272FFFCFD75029D77B2FA10" box="[679,752,1427,1454]" metricMagnitude="2" metricUnit="m" metricValue="8.53" pageId="1" pageNumber="440" unit="m" value="853.0">853 m</elevation>
|
||
</quantity>
|
||
, Alois Pavlíčko,
|
||
<specimenCount id="9D79FD06A272FFFCFC7C029A769AFA10" box="[942,984,1428,1454]" pageId="1" pageNumber="440" type="female">1 ♀</specimenCount>
|
||
(
|
||
<collectionCode id="ED6EAE4AA272FFFCFC35029A716DFA10" box="[999,1071,1428,1454]" country="Czech Republic" httpUri="http://grbio.org/cool/06zb-203s" name="Biology Centre of the Academy of Sciences of the Czech Republic, Institute of Entomology" pageId="1" pageNumber="440" type="Other institutional type">IECA</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="8BC0368FA272FFFCFBE4029A7178FA10" blockId="1.[151,1437,959,2030]" box="[1078,1082,1428,1454]" pageId="1" pageNumber="440">.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3656504A272FFF8FF1502B97028FF0D" lastPageId="5" lastPageNumber="444" pageId="1" pageNumber="440" type="description">
|
||
<paragraph id="8BC0368FA272FFFCFF1502B977A7F938" blockId="1.[151,1437,959,2030]" pageId="1" pageNumber="440">
|
||
<emphasis id="B90BEA9DA272FFFCFF1502B977D3FA6C" bold="true" box="[199,657,1463,1490]" pageId="1" pageNumber="440">
|
||
Redescription of the
|
||
<typeStatus id="54C4882DA272FFFCFE1A02B67771FA6C" box="[456,563,1464,1490]" pageId="1" pageNumber="440" type="lectotype">lectotype</typeStatus>
|
||
(male).
|
||
</emphasis>
|
||
Body elongate, almost parallel, convex. Dorsal and ventral surface moderately shiny, brown; macrosetation pale (
|
||
<figureCitation id="13442A0AA272FFFCFD4902D5779DFA48" box="[667,735,1499,1526]" captionStart="FIGURES 8–16" captionStartId="3.[151,264,1867,1892]" captionTargetBox="[208,1380,206,1822]" captionTargetId="figure@3.[183,1404,181,1842]" captionTargetPageId="3" captionText="FIGURES 8–16. Comparison of Anoxia (Protanoxia) species. 8, Lectotype of Anoxia maljuzhenkoi (Zaitzev, 1928), male, dorsal view; 9, A. maljuzhenkoi lectotype labels; 10, A. maljuzhenkoi base of pronotum and elytra; 11, A. ciliciensis Baraud, 1989 base of pronotum and elytra; 12, A. maljuzhenkoi pygidium; 13, A. ciliciensis pygidium; 14, A. smyrnensis Petrovitz, 1965 terminal maxillary palpomere; 15, A. maljuzhenkoi terminal maxillary palpomere; 16, A. ciliciensis terminal maxillary palpomere. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.3442502" httpUri="https://zenodo.org/record/3442502/files/figure.png" pageId="1" pageNumber="440">Fig. 8</figureCitation>
|
||
). Dorsal surface of head, pronotum, scutellar shield, and elytra covered with white and pale yellow, short, recumbent scale-like macrosetae; ventral surface of thorax densely covered with pale yellow, long, erect macrosetae; ventral surface of abdomen with densely spaced, white, recumbent, scale-like macrosetae. Head appendages and legs covered with pale yellow, moderately long macrosetae sparsely mixed with white, recumbent, scale-like macrosetae.
|
||
</paragraph>
|
||
<paragraph id="8BC0368FA272FFFCFF1501817164F853" blockId="1.[151,1437,959,2030]" pageId="1" pageNumber="440">Head. Labrum transverse, deeply bilobed, lobes rounded, irregularly and coarsely punctate, covered with long, erect macrosetae and basally with few scale-like macrosetae. Clypeus transverse, rectangular, anterior margin upturned, anterior angles rounded apically. Frontoclypeal suture weakly visible, forming continuous bisinuate flat line. Clypeus and frons densely, irregularly, coarsely punctate; covered with short, erect, scale-like macrosetae; macrosetae mixed on frons with long, erect macrosetae. Eye canthus narrow, with row of short, scale-like macrosetae and few long, erect macrosetae. Eye moderately large, only minutely extending beyond eye canthus. Antenna with 10 antennomeres; antennomere III somewhat longer than antennomeres IV and V combined; club with five antennomeres, shorter than antennal shaft. Antennomere I with long, erect macrosetae apically and laterally; antennomeres II–IV with few isolated, long macrosetae; club with sparse, short macrosetae. Terminal maxillary palpomere elongate, apically rounded, approximately as long as palpomeres II and III combined.</paragraph>
|
||
<caption id="DF006607A271FFFFFF45008E71D2F85F" ID-DOI="http://doi.org/10.5281/zenodo.3442500" ID-Zenodo-Dep="3442500" httpUri="https://zenodo.org/record/3442500/files/figure.png" pageId="2" pageNumber="441" startId="2.[151,264,1920,1945]" targetBox="[205,1358,203,1839]" targetPageId="2">
|
||
<paragraph id="8BC0368FA271FFFFFF45008E71D2F85F" blockId="2.[151,1436,1920,2017]" pageId="2" pageNumber="441">
|
||
<emphasis id="B90BEA9DA271FFFFFF45008E747EF827" bold="true" box="[151,316,1920,1945]" pageId="2" pageNumber="441">FIGURES 1–7.</emphasis>
|
||
<taxonomicName id="4C7F4D0CA271FFFFFE91008F7605F826" authority="(Zaitzev, 1928)" authorityName="Bezděk & Kalashian & Ghrejyan" authorityYear="2019" baseAuthorityName="Zaitzev" baseAuthorityYear="1928" box="[323,839,1920,1945]" class="Insecta" family="Melolonthidae" genus="Anoxia" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="441" phylum="Arthropoda" rank="species" species="maljuzhenkoi" subGenus="Protanoxia">
|
||
<emphasis id="B90BEA9DA271FFFFFE91008F74C9F826" box="[323,395,1921,1944]" italics="true" pageId="2" pageNumber="441">Anoxia</emphasis>
|
||
(
|
||
<emphasis id="B90BEA9DA271FFFFFE49008F774EF826" box="[411,524,1921,1944]" italics="true" pageId="2" pageNumber="441">Protanoxia</emphasis>
|
||
)
|
||
<emphasis id="B90BEA9DA271FFFFFDCE008E77E1F826" box="[540,675,1920,1944]" italics="true" pageId="2" pageNumber="441">maljuzhenkoi</emphasis>
|
||
(
|
||
<bibRefCitation id="EFEE4B7EA271FFFFFD60008E7602F826" author="Zaitzev, F. A." box="[690,832,1920,1945]" pageId="2" pageNumber="441" pagination="373 - 397" refId="ref4775" refString="Zaitzev, F. A. (1928) Obzor khrushchey Kavkaza v svyazi s ikh rasprostraneniem v kraye. Zur Verbreitung der Melolonthinen in den Kaukasuslandern. Izvestiya Tifliskogo Gosudarstvennogo Politekhnicheskogo Instituta, 3, 373 - 397. [In Russian, German summary]" type="journal article" year="1928">Zaitzev, 1928</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
from “Vordan Karmir” State Sanctuary, Armenia.
|
||
<emphasis id="B90BEA9DA271FFFFFA82008E701FF827" bold="true" box="[1360,1373,1920,1945]" pageId="2" pageNumber="441">1</emphasis>
|
||
, male (length 22.5 mm), dorsal view;
|
||
<emphasis id="B90BEA9DA271FFFFFE0900AA74AAF803" bold="true" box="[475,488,1956,1981]" pageId="2" pageNumber="441">2</emphasis>
|
||
, male, lateral view;
|
||
<emphasis id="B90BEA9DA271FFFFFD6800AA7785F803" bold="true" box="[698,711,1956,1981]" pageId="2" pageNumber="441">3</emphasis>
|
||
, male aedeagus, lateral view;
|
||
<emphasis id="B90BEA9DA271FFFFFC2E00AA714BF803" bold="true" box="[1020,1033,1956,1981]" pageId="2" pageNumber="441">4</emphasis>
|
||
, male aedeagus, dorsal view;
|
||
<emphasis id="B90BEA9DA271FFFFFAEF00AA7008F802" bold="true" box="[1341,1354,1956,1980]" pageId="2" pageNumber="441">5</emphasis>
|
||
, female (length 21.4 mm), dorsal view;
|
||
<emphasis id="B90BEA9DA271FFFFFE0500C674A6F85F" bold="true" box="[471,484,1992,2017]" pageId="2" pageNumber="441">6</emphasis>
|
||
, female, lateral view;
|
||
<emphasis id="B90BEA9DA271FFFFFD1400C67791F85E" bold="true" box="[710,723,1992,2016]" pageId="2" pageNumber="441">7</emphasis>
|
||
, female protibia, ventral view. Not to scale.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF006607A270FFFEFF45004574CDF84A" ID-DOI="http://doi.org/10.5281/zenodo.3442502" ID-Zenodo-Dep="3442502" httpUri="https://zenodo.org/record/3442502/files/figure.png" pageId="3" pageNumber="442" startId="3.[151,264,1867,1892]" targetBox="[208,1380,206,1822]" targetPageId="3">
|
||
<paragraph id="8BC0368FA270FFFEFF45004574CDF84A" blockId="3.[151,1437,1867,2036]" pageId="3" pageNumber="442">
|
||
<emphasis id="B90BEA9DA270FFFEFF4500457408F8DA" bold="true" box="[151,330,1867,1892]" pageId="3" pageNumber="442">FIGURES 8–16.</emphasis>
|
||
Comparison of
|
||
<taxonomicName id="4C7F4D0CA270FFFEFE2500427788F8DA" authority="(Protanoxia)" authorityName="Medvedev" authorityYear="1951" baseAuthorityName="Protanoxia" box="[503,714,1868,1892]" class="Insecta" family="Melolonthidae" genus="Anoxia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="442" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B90BEA9DA270FFFEFE250042777DF8DD" box="[503,575,1868,1891]" italics="true" pageId="3" pageNumber="442">Anoxia</emphasis>
|
||
(
|
||
<emphasis id="B90BEA9DA270FFFEFD8300437780F8DA" box="[593,706,1869,1892]" italics="true" pageId="3" pageNumber="442">Protanoxia</emphasis>
|
||
)
|
||
</taxonomicName>
|
||
species.
|
||
<emphasis id="B90BEA9DA270FFFEFCFE0045767BF8DA" bold="true" box="[812,825,1867,1892]" pageId="3" pageNumber="442">8</emphasis>
|
||
, Lectotype of
|
||
<taxonomicName id="4C7F4D0CA270FFFEFC040042701AF8DA" authority="(Zaitzev, 1928)" authorityName="Bezděk & Kalashian & Ghrejyan" authorityYear="2019" baseAuthorityName="Zaitzev" baseAuthorityYear="1928" box="[982,1368,1867,1892]" class="Insecta" family="Melolonthidae" genus="Anoxia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="442" phylum="Arthropoda" rank="species" species="maljuzhenkoi">
|
||
<emphasis id="B90BEA9DA270FFFEFC04004271ECF8DA" box="[982,1198,1868,1892]" italics="true" pageId="3" pageNumber="442">Anoxia maljuzhenkoi</emphasis>
|
||
(
|
||
<bibRefCitation id="EFEE4B7EA270FFFEFB6C00457012F8DA" author="Zaitzev, F. A." box="[1214,1360,1867,1892]" pageId="3" pageNumber="442" pagination="373 - 397" refId="ref4775" refString="Zaitzev, F. A. (1928) Obzor khrushchey Kavkaza v svyazi s ikh rasprostraneniem v kraye. Zur Verbreitung der Melolonthinen in den Kaukasuslandern. Izvestiya Tifliskogo Gosudarstvennogo Politekhnicheskogo Instituta, 3, 373 - 397. [In Russian, German summary]" type="journal article" year="1928">Zaitzev, 1928</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, male, dorsal view;
|
||
<emphasis id="B90BEA9DA270FFFEFECF00617468F836" bold="true" box="[285,298,1903,1928]" pageId="3" pageNumber="442">9</emphasis>
|
||
,
|
||
<taxonomicName id="4C7F4D0CA270FFFEFEEA007E749FF836" authorityName="Bezděk & Kalashian & Ghrejyan" authorityYear="2019" baseAuthorityName="Zaitzev" baseAuthorityYear="1928" box="[312,477,1904,1928]" class="Insecta" family="Melolonthidae" genus="Anoxia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="442" phylum="Arthropoda" rank="species" species="maljuzhenkoi">
|
||
<emphasis id="B90BEA9DA270FFFEFEEA007E749FF836" box="[312,477,1904,1928]" italics="true" pageId="3" pageNumber="442">A. maljuzhenkoi</emphasis>
|
||
</taxonomicName>
|
||
lectotype labels;
|
||
<emphasis id="B90BEA9DA270FFFEFD46006177EFF836" bold="true" box="[660,685,1903,1928]" pageId="3" pageNumber="442">10</emphasis>
|
||
,
|
||
<taxonomicName id="4C7F4D0CA270FFFEFD6E007E7623F836" authorityName="Bezděk & Kalashian & Ghrejyan" authorityYear="2019" baseAuthorityName="Zaitzev" baseAuthorityYear="1928" box="[700,865,1904,1928]" class="Insecta" family="Melolonthidae" genus="Anoxia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="442" phylum="Arthropoda" rank="species" species="maljuzhenkoi">
|
||
<emphasis id="B90BEA9DA270FFFEFD6E007E7623F836" box="[700,865,1904,1928]" italics="true" pageId="3" pageNumber="442">A. maljuzhenkoi</emphasis>
|
||
</taxonomicName>
|
||
base of pronotum and elytra;
|
||
<emphasis id="B90BEA9DA270FFFEFB49006171F1F836" bold="true" box="[1179,1203,1903,1928]" pageId="3" pageNumber="442">11</emphasis>
|
||
,
|
||
<taxonomicName id="4C7F4D0CA270FFFEFB10007E7588F812" authority="Baraud, 1989" authorityName="Baraud" authorityYear="1989" class="Insecta" family="Melolonthidae" genus="Anoxia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="442" phylum="Arthropoda" rank="species" species="ciliciensis">
|
||
<emphasis id="B90BEA9DA270FFFEFB10007E7006F836" box="[1218,1348,1904,1928]" italics="true" pageId="3" pageNumber="442">A. ciliciensis</emphasis>
|
||
<bibRefCitation id="EFEE4B7EA270FFFEFA9F00617588F812" author="Baraud, J." pageId="3" pageNumber="442" pagination="273 - 284" refId="ref4143" refString="Baraud, J. (1989) Revision des Anoxia Castelnau d'Europe et d'Asie. 1 re note: le sous-genre Protanoxia Medvedev (Col. Melolonthidae). Bulletin de la Societe Entomologique de France, 93, 273 - 284." type="journal article" year="1989">Baraud, 1989</bibRefCitation>
|
||
</taxonomicName>
|
||
base of pronotum and elytra;
|
||
<emphasis id="B90BEA9DA270FFFEFDD8009D7761F812" bold="true" box="[522,547,1939,1964]" pageId="3" pageNumber="442">12</emphasis>
|
||
,
|
||
<taxonomicName id="4C7F4D0CA270FFFEFDE0009A779AF812" authorityName="Bezděk & Kalashian & Ghrejyan" authorityYear="2019" baseAuthorityName="Zaitzev" baseAuthorityYear="1928" box="[562,728,1940,1964]" class="Insecta" family="Melolonthidae" genus="Anoxia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="442" phylum="Arthropoda" rank="species" species="maljuzhenkoi">
|
||
<emphasis id="B90BEA9DA270FFFEFDE0009A779AF812" box="[562,728,1940,1964]" italics="true" pageId="3" pageNumber="442">A. maljuzhenkoi</emphasis>
|
||
</taxonomicName>
|
||
pygidium;
|
||
<emphasis id="B90BEA9DA270FFFEFC80009D7629F812" bold="true" box="[850,875,1939,1964]" pageId="3" pageNumber="442">13</emphasis>
|
||
,
|
||
<taxonomicName id="4C7F4D0CA270FFFEFCA9009A76BDF812" authorityName="Baraud" authorityYear="1989" box="[891,1023,1940,1964]" class="Insecta" family="Melolonthidae" genus="Anoxia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="442" phylum="Arthropoda" rank="species" species="ciliciensis">
|
||
<emphasis id="B90BEA9DA270FFFEFCA9009A76BDF812" box="[891,1023,1940,1964]" italics="true" pageId="3" pageNumber="442">A. ciliciensis</emphasis>
|
||
</taxonomicName>
|
||
pygidium;
|
||
<emphasis id="B90BEA9DA270FFFEFBAB009D71D0F812" bold="true" box="[1145,1170,1939,1964]" pageId="3" pageNumber="442">14</emphasis>
|
||
,
|
||
<taxonomicName id="4C7F4D0CA270FFFEFB70009A7588F86E" authority="Petrovitz, 1965" authorityName="Petrovitz" authorityYear="1965" class="Insecta" family="Melolonthidae" genus="Anoxia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="442" phylum="Arthropoda" rank="species" species="smyrnensis">
|
||
<emphasis id="B90BEA9DA270FFFEFB70009A7070F815" box="[1186,1330,1940,1963]" italics="true" pageId="3" pageNumber="442">A. smyrnensis</emphasis>
|
||
Petrovitz, 1965
|
||
</taxonomicName>
|
||
terminal maxillary palpomere;
|
||
<emphasis id="B90BEA9DA270FFFEFDC100B9776EF86E" bold="true" box="[531,556,1975,2000]" pageId="3" pageNumber="442">15</emphasis>
|
||
,
|
||
<taxonomicName id="4C7F4D0CA270FFFEFDE900B677A3F86E" authorityName="Bezděk & Kalashian & Ghrejyan" authorityYear="2019" baseAuthorityName="Zaitzev" baseAuthorityYear="1928" box="[571,737,1976,2000]" class="Insecta" family="Melolonthidae" genus="Anoxia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="442" phylum="Arthropoda" rank="species" species="maljuzhenkoi">
|
||
<emphasis id="B90BEA9DA270FFFEFDE900B677A3F86E" box="[571,737,1976,2000]" italics="true" pageId="3" pageNumber="442">A. maljuzhenkoi</emphasis>
|
||
</taxonomicName>
|
||
terminal maxillary palpomere;
|
||
<emphasis id="B90BEA9DA270FFFEFBF800B97101F86E" bold="true" box="[1066,1091,1975,2000]" pageId="3" pageNumber="442">16</emphasis>
|
||
,
|
||
<taxonomicName id="4C7F4D0CA270FFFEFB8000B67197F86E" authorityName="Baraud" authorityYear="1989" box="[1106,1237,1976,2000]" class="Insecta" family="Melolonthidae" genus="Anoxia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="442" phylum="Arthropoda" rank="species" species="ciliciensis">
|
||
<emphasis id="B90BEA9DA270FFFEFB8000B67197F86E" box="[1106,1237,1976,2000]" italics="true" pageId="3" pageNumber="442">A. ciliciensis</emphasis>
|
||
</taxonomicName>
|
||
terminal maxillary palpomere. Not to scale.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF006607A277FFF9FF45048D7772FC7E" ID-DOI="http://doi.org/10.5281/zenodo.3442504" ID-Zenodo-Dep="3442504" httpUri="https://zenodo.org/record/3442504/files/figure.png" pageId="4" pageNumber="443" startId="4.[151,250,899,924]" targetBox="[177,1411,286,874]" targetPageId="4">
|
||
<paragraph id="8BC0368FA277FFF9FF45048D7772FC7E" blockId="4.[151,1437,899,960]" pageId="4" pageNumber="443">
|
||
<emphasis id="B90BEA9DA277FFF9FF45048D7463FC22" bold="true" box="[151,289,899,924]" pageId="4" pageNumber="443">FIGURE 17.</emphasis>
|
||
“Vordan Karmir” State Sanctuary, Armenia – the biotope of
|
||
<taxonomicName id="4C7F4D0CA277FFF9FC41048A70DBFC22" authority="(Zaitzev, 1928)" authorityName="Bezděk & Kalashian & Ghrejyan" authorityYear="2019" baseAuthorityName="Zaitzev" baseAuthorityYear="1928" box="[915,1433,899,924]" class="Insecta" family="Melolonthidae" genus="Anoxia" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="443" phylum="Arthropoda" rank="species" species="maljuzhenkoi" subGenus="Protanoxia">
|
||
<emphasis id="B90BEA9DA277FFF9FC41048A7699FC25" box="[915,987,900,923]" italics="true" pageId="4" pageNumber="443">Anoxia</emphasis>
|
||
(
|
||
<emphasis id="B90BEA9DA277FFF9FC39048B711EFC22" box="[1003,1116,901,924]" italics="true" pageId="4" pageNumber="443">Protanoxia</emphasis>
|
||
)
|
||
<emphasis id="B90BEA9DA277FFF9FBBE048A71B1FC22" box="[1132,1267,900,924]" italics="true" pageId="4" pageNumber="443">maljuzhenkoi</emphasis>
|
||
(
|
||
<bibRefCitation id="EFEE4B7EA277FFF9FAD3048D70D0FC22" author="Zaitzev, F. A." box="[1281,1426,899,924]" pageId="4" pageNumber="443" pagination="373 - 397" refId="ref4775" refString="Zaitzev, F. A. (1928) Obzor khrushchey Kavkaza v svyazi s ikh rasprostraneniem v kraye. Zur Verbreitung der Melolonthinen in den Kaukasuslandern. Izvestiya Tifliskogo Gosudarstvennogo Politekhnicheskogo Instituta, 3, 373 - 397. [In Russian, German summary]" type="journal article" year="1928">Zaitzev, 1928</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
. Photograph by Jan Šumpich, June 2017.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BC0368FA277FFF9FF1504E17620FB03" blockId="4.[151,1437,1006,2041]" pageId="4" pageNumber="443">Pronotum transverse, moderately convex with distinct longitudinal groove in the middle, widest approximately in the middle. Basal border present, broadly interrupted in the middle, lateral borders complete, anterior border missing. Lateral margins weakly crenate. Anterior margin regularly, broadly sinuate. Anterior angles obtuse-angular; posterior angles broadly rounded. Punctation consisting of coarse, irregularly-spaced punctures becoming denser in the longitudinal groove, each puncture bearing white, scale-like, recumbent macroseta; two rounded, impunctate, bare lateral areas on each side of pronotum present.</paragraph>
|
||
<paragraph id="8BC0368FA277FFF9FF1503C87107FB5F" blockId="4.[151,1437,1006,2041]" box="[199,1093,1222,1249]" pageId="4" pageNumber="443">Scutellar shield large, transverse, sides and apex broadly rounded; disc punctate</paragraph>
|
||
<paragraph id="8BC0368FA277FFF9FF1503E476AEFABB" blockId="4.[151,1437,1006,2041]" box="[199,1004,1258,1285]" pageId="4" pageNumber="443">laterally, densely covered with white, scale-like, recumbent macrosetae.</paragraph>
|
||
<paragraph id="8BC0368FA277FFF9FF150200702CFACE" blockId="4.[151,1437,1006,2041]" pageId="4" pageNumber="443">Elytra convex, parallel sided, rounded apically; apical angle obtuse-angular. Striae missing; humeral umbones present, vaguely swollen. Surface moderately shiny; punctation coarse, irregularly spaced. Each puncture bearing short, white, scale-like, almost recumbent macroseta. Epipleura narrow, with several macrosetae in basal third.</paragraph>
|
||
<paragraph id="8BC0368FA277FFF9FF1502757428FA2B" blockId="4.[151,1437,1006,2041]" box="[199,362,1403,1429]" pageId="4" pageNumber="443">Macropterous.</paragraph>
|
||
<paragraph id="8BC0368FA277FFF9FF1502917183F92E" blockId="4.[151,1437,1006,2041]" pageId="4" pageNumber="443">Legs. Femora narrow, shiny, irregularly punctate; densely covered with long, semierect, pale yellow macrosetae mixed with short, white, recumbent, scale-like macrosetae. Protibia narrow, tridentate, terminal calcar missing. Mesotibia and metatibia slightly expanded apicad, with two macrosetiferous longitudinal and two incomplete oblique transversal carinae, sparsely covered with short scale-like macrosetae, inner part densely covered by long erect macrosetae. Upper terminal calcar of metatibia flattened, blunt apically, about 1.5 times as long as lower calcar. Tarsal claws bifid, with well-developed, unequal ventrobasal teeth. Protarsus with distinctly longer ventrobasal tooth of inner claw, whereas mesotarsi and metatarsi with more robust ventrobasal teeth on outer claws.</paragraph>
|
||
<paragraph id="8BC0368FA277FFF9FF1501947613F967" blockId="4.[151,1437,1006,2041]" pageId="4" pageNumber="443">Ventral surface of thorax densely covered with pale yellow, long, erect macrosetae; ventral surface of abdomen with densely-spaced, white, recumbent, scale-like macrosetae.</paragraph>
|
||
<paragraph id="8BC0368FA277FFF9FF1501EC77B0F89F" blockId="4.[151,1437,1006,2041]" pageId="4" pageNumber="443">Pygidium almost flat, completely bordered; apically broadly rounded; irregularly covered with coarse punctures bearing short, recumbent, scale-like macrosetae.</paragraph>
|
||
<paragraph id="8BC0368FA277FFF9FF1500247139F8FB" blockId="4.[151,1437,1006,2041]" box="[199,1147,1834,1861]" pageId="4" pageNumber="443">
|
||
Male genitalia. Parameres symmetrical, two times as long as phallobasis (
|
||
<figureCitation id="13442A0AA277FFF9FBD00024712DF8FB" box="[1026,1135,1834,1861]" captionStart="FIGURES 1–7" captionStartId="2.[151,264,1920,1945]" captionTargetBox="[205,1358,203,1839]" captionTargetId="figure@2.[164,1423,181,1894]" captionTargetPageId="2" captionText="FIGURES 1–7. Anoxia (Protanoxia) maljuzhenkoi (Zaitzev, 1928) from “Vordan Karmir” State Sanctuary, Armenia. 1, male (length 22.5 mm), dorsal view; 2, male, lateral view; 3, male aedeagus, lateral view; 4, male aedeagus, dorsal view; 5, female (length 21.4 mm), dorsal view; 6, female, lateral view; 7, female protibia, ventral view. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.3442500" httpUri="https://zenodo.org/record/3442500/files/figure.png" pageId="4" pageNumber="443">Figs. 3–4</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BC0368FA277FFF9FF150040767EF80E" blockId="4.[151,1437,1006,2041]" pageId="4" pageNumber="443">
|
||
<emphasis id="B90BEA9DA277FFF9FF15004074F4F8D7" bold="true" box="[199,438,1870,1897]" pageId="4" pageNumber="443">Sexual dimorphism.</emphasis>
|
||
Female (
|
||
<figureCitation id="13442A0AA277FFF9FDCE004077C7F8D7" box="[540,645,1870,1897]" captionStart="FIGURES 1–7" captionStartId="2.[151,264,1920,1945]" captionTargetBox="[205,1358,203,1839]" captionTargetId="figure@2.[164,1423,181,1894]" captionTargetPageId="2" captionText="FIGURES 1–7. Anoxia (Protanoxia) maljuzhenkoi (Zaitzev, 1928) from “Vordan Karmir” State Sanctuary, Armenia. 1, male (length 22.5 mm), dorsal view; 2, male, lateral view; 3, male aedeagus, lateral view; 4, male aedeagus, dorsal view; 5, female (length 21.4 mm), dorsal view; 6, female, lateral view; 7, female protibia, ventral view. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.3442500" httpUri="https://zenodo.org/record/3442500/files/figure.png" pageId="4" pageNumber="443">Figs. 5–6</figureCitation>
|
||
) differs from male in the following characters: pronotum dark brown to black; clypeus broadly rounded; antennal club with four antennomeres, shorter; terminal calcar of protibia present (
|
||
<figureCitation id="13442A0AA277FFF9FF72009875A6F80F" box="[160,228,1942,1969]" captionStart="FIGURES 1–7" captionStartId="2.[151,264,1920,1945]" captionTargetBox="[205,1358,203,1839]" captionTargetId="figure@2.[164,1423,181,1894]" captionTargetPageId="2" captionText="FIGURES 1–7. Anoxia (Protanoxia) maljuzhenkoi (Zaitzev, 1928) from “Vordan Karmir” State Sanctuary, Armenia. 1, male (length 22.5 mm), dorsal view; 2, male, lateral view; 3, male aedeagus, lateral view; 4, male aedeagus, dorsal view; 5, female (length 21.4 mm), dorsal view; 6, female, lateral view; 7, female protibia, ventral view. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.3442500" httpUri="https://zenodo.org/record/3442500/files/figure.png" pageId="4" pageNumber="443">Fig. 7</figureCitation>
|
||
); tarsal claws of all legs with ventrobasal teeth equal.
|
||
</paragraph>
|
||
<paragraph id="8BC0368FA277FFF9FF1500B47050F846" blockId="4.[151,1437,1006,2041]" pageId="4" pageNumber="443">
|
||
<emphasis id="B90BEA9DA277FFF9FF1500B4740FF86B" bold="true" box="[199,333,1978,2005]" pageId="4" pageNumber="443">Variability.</emphasis>
|
||
In comparison with
|
||
<typeStatus id="54C4882DA277FFF9FDE500B477E3F86A" box="[567,673,1978,2004]" pageId="4" pageNumber="443" type="lectotype">lectotype</typeStatus>
|
||
, recently collected male from “Vordan Karmir” State Sanctuary has partially abraded scale-like macrosetae on pronotum and protibia, as well as external teeth on protibia.
|
||
</paragraph>
|
||
<paragraph id="8BC0368FA276FFF8FF1507977028FF0D" blockId="5.[151,1437,152,1115]" box="[199,1386,152,179]" pageId="5" pageNumber="444">
|
||
<emphasis id="B90BEA9DA276FFF8FF150797743CFF0D" bold="true" box="[199,382,153,179]" pageId="5" pageNumber="444">Measurements.</emphasis>
|
||
Total body length of males:
|
||
<quantity id="4C879B6AA276FFF8FD6C07967623FF0D" box="[702,865,152,179]" metricMagnitude="-2" metricUnit="m" metricValue="2.23" metricValueMax="2.25" metricValueMin="2.21" pageId="5" pageNumber="444" unit="mm" value="22.3" valueMax="22.5" valueMin="22.1">22.1–22.5 mm</quantity>
|
||
(
|
||
<typeStatus id="54C4882DA276FFF8FCA20796769BFF0C" box="[880,985,152,178]" pageId="5" pageNumber="444" type="lectotype">lectotype</typeStatus>
|
||
<quantity id="4C879B6AA276FFF8FC0D07977105FF0D" box="[991,1095,153,179]" metricMagnitude="-2" metricUnit="m" metricValue="2.21" pageId="5" pageNumber="444" unit="mm" value="22.1">22.1 mm</quantity>
|
||
), females:
|
||
<quantity id="4C879B6AA276FFF8FB1207977024FF0D" box="[1216,1382,153,179]" metricMagnitude="-2" metricUnit="m" metricValue="2.16" metricValueMax="2.18" metricValueMin="2.1399999999999997" pageId="5" pageNumber="444" unit="mm" value="21.6" valueMax="21.8" valueMin="21.4">21.4–21.8 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3656504A276FFF8FF1507B2758DFB89" pageId="5" pageNumber="444" type="diagnosis">
|
||
<paragraph id="8BC0368FA276FFF8FF1507B27429FED9" blockId="5.[151,1437,152,1115]" pageId="5" pageNumber="444">
|
||
<emphasis id="B90BEA9DA276FFF8FF1507B27400FF69" bold="true" box="[199,322,188,215]" pageId="5" pageNumber="444">Diagnosis.</emphasis>
|
||
An examination of both sexes helps us to clarify the generic placement of the species. The presence of terminal calcar of protibia in the female, its absence in the male, and the length of antennomere III clearly place this species in the genus
|
||
<taxonomicName id="4C7F4D0CA276FFF8FE27060B7706FEA1" authorityName="Laporte" authorityYear="1832" box="[501,580,261,287]" class="Insecta" family="Melolonthidae" genus="Anoxia" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="444" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B90BEA9DA276FFF8FE27060B7706FEA1" box="[501,580,261,287]" italics="true" pageId="5" pageNumber="444">Anoxia</emphasis>
|
||
</taxonomicName>
|
||
. The rounded apex of the pygidium more precisely place it in the subgenus
|
||
<taxonomicName id="4C7F4D0CA276FFF8FF4506277456FEFC" authorityName="Medvedev" authorityYear="1951" box="[151,276,297,322]" class="Insecta" family="Melolonthidae" genus="Anoxia" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="444" phylum="Arthropoda" rank="subGenus" subGenus="Protanoxia">
|
||
<emphasis id="B90BEA9DA276FFF8FF4506277456FEFC" box="[151,276,297,322]" italics="true" pageId="5" pageNumber="444">Protanoxia</emphasis>
|
||
</taxonomicName>
|
||
. Therefore, the following combination is proposed:
|
||
<taxonomicName id="4C7F4D0CA276FFF8FCB2062770D5FEFD" authority="(Zaitzev, 1928)" authorityName="Bezděk & Kalashian & Ghrejyan" authorityYear="2019" baseAuthorityName="Zaitzev" baseAuthorityYear="1928" box="[864,1431,296,323]" class="Insecta" family="Melolonthidae" genus="Anoxia" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="444" phylum="Arthropoda" rank="species" species="maljuzhenkoi" status="comb. nov." subGenus="Protanoxia">
|
||
<emphasis id="B90BEA9DA276FFF8FCB2062776EDFEFD" box="[864,943,297,323]" italics="true" pageId="5" pageNumber="444">Anoxia</emphasis>
|
||
(
|
||
<emphasis id="B90BEA9DA276FFF8FC120627717FFEFC" box="[960,1085,297,322]" italics="true" pageId="5" pageNumber="444">Protanoxia</emphasis>
|
||
)
|
||
<emphasis id="B90BEA9DA276FFF8FB9F062671A0FEFC" box="[1101,1250,296,322]" italics="true" pageId="5" pageNumber="444">maljuzhenkoi</emphasis>
|
||
(
|
||
<bibRefCitation id="EFEE4B7EA276FFF8FB20062670CDFEFD" author="Zaitzev, F. A." box="[1266,1423,296,323]" pageId="5" pageNumber="444" pagination="373 - 397" refId="ref4775" refString="Zaitzev, F. A. (1928) Obzor khrushchey Kavkaza v svyazi s ikh rasprostraneniem v kraye. Zur Verbreitung der Melolonthinen in den Kaukasuslandern. Izvestiya Tifliskogo Gosudarstvennogo Politekhnicheskogo Instituta, 3, 373 - 397. [In Russian, German summary]" type="journal article" year="1928">Zaitzev, 1928</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="B90BEA9DA276FFF8FF4506437421FED9" bold="true" box="[151,355,332,359]" pageId="5" pageNumber="444">
|
||
<taxonomicNameLabel id="A23857E6A276FFF8FF4506437421FED9" box="[151,355,332,359]" pageId="5" pageNumber="444" rank="species">new combination</taxonomicNameLabel>
|
||
</emphasis>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BC0368FA276FFF8FF15067E76D0FD4C" blockId="5.[151,1437,152,1115]" pageId="5" pageNumber="444">
|
||
Previously, eight members of
|
||
<taxonomicName id="4C7F4D0CA276FFF8FDC1067F77D2FE34" authorityName="Medvedev" authorityYear="1951" box="[531,656,369,394]" class="Insecta" family="Melolonthidae" genus="Anoxia" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="444" phylum="Arthropoda" rank="subGenus" subGenus="Protanoxia">
|
||
<emphasis id="B90BEA9DA276FFF8FDC1067F77D2FE34" box="[531,656,369,394]" italics="true" pageId="5" pageNumber="444">Protanoxia</emphasis>
|
||
</taxonomicName>
|
||
were known (
|
||
<bibRefCitation id="EFEE4B7EA276FFF8FCFF067E7683FE35" author="Baraud, J." box="[813,961,368,395]" pageId="5" pageNumber="444" pagination="253 - 261" refId="ref4106" refString="Baraud, J. (1980) Coleopteres Scarabaeoidea de l'Afrique du Nord. 5 e note: Melolonthinae nouveaux ou meconnus. Bulletin de la Societe Entomologique de France, 85, 253 - 261." type="journal article" year="1980">Baraud 1980</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFEE4B7EA276FFF8FC19067F7144FE35" author="Baraud, J." box="[971,1030,369,395]" pageId="5" pageNumber="444" refId="ref4186" refString="Baraud, J. (1992) Coleopteres Scarabaeoidea d'Europe. Faune de France et regions limitrophies, 78. Federation Francaise des Societes de Sciences Naturelles & Societe Linneenne de Lyon, Paris, 856 pp." type="book" year="1992">1992</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFEE4B7EA276FFF8FBC2067E71EAFE35" author="Bezdek, A." box="[1040,1192,368,395]" pageId="5" pageNumber="444" pagination="226 - 236" refId="ref4280" refString="Bezdek, A. (2016) Tribus Melolonthini. In: Lobl, I & Lobl, D. (Eds.), Catalogue of Palaearctic Coleoptera. Volume 3. Scara- baeoidea - Scirtoidea - Dascilloidea - Buprestoidea - Byrrhoidea. Revised and Updated Edition. E. J. Brill, Leiden, pp. 226 - 236." type="book chapter" year="2016">Bezděk 2016</bibRefCitation>
|
||
). Two of them,
|
||
<emphasis id="B90BEA9DA276FFF8FA81067F7026FE35" box="[1363,1380,369,395]" italics="true" pageId="5" pageNumber="444">A</emphasis>
|
||
.
|
||
<taxonomicName id="4C7F4D0CA276FFF8FAA3067F74E3FE11" authority="Dewailly, 1957" authorityName="Dewailly" authorityYear="1957" class="Insecta" family="Melolonthidae" genus="Anoxia" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="444" phylum="Arthropoda" rank="species" species="kocheri" subGenus="Protanoxia">
|
||
(
|
||
<emphasis id="B90BEA9DA276FFF8FAA9067F70CEFE34" box="[1403,1420,369,394]" italics="true" pageId="5" pageNumber="444">P</emphasis>
|
||
.)
|
||
<emphasis id="B90BEA9DA276FFF8FF45069A75A9FE10" box="[151,235,404,430]" italics="true" pageId="5" pageNumber="444">kocheri</emphasis>
|
||
Dewailly, 1957
|
||
</taxonomicName>
|
||
and
|
||
<emphasis id="B90BEA9DA276FFF8FE09069B74AEFE11" box="[475,492,405,431]" italics="true" pageId="5" pageNumber="444">A</emphasis>
|
||
.
|
||
<taxonomicName id="4C7F4D0CA276FFF8FE2E069B7671FE11" authority="Dewailly, 1957" authorityName="Dewailly" authorityYear="1957" box="[508,819,404,431]" class="Insecta" family="Melolonthidae" genus="Anoxia" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="444" phylum="Arthropoda" rank="species" species="rotroui" subGenus="Protanoxia">
|
||
(
|
||
<emphasis id="B90BEA9DA276FFF8FDD7069B7754FE10" box="[517,534,405,430]" italics="true" pageId="5" pageNumber="444">P</emphasis>
|
||
.)
|
||
<emphasis id="B90BEA9DA276FFF8FDE2069B773FFE10" box="[560,637,405,430]" italics="true" pageId="5" pageNumber="444">rotroui</emphasis>
|
||
Dewailly, 1957
|
||
</taxonomicName>
|
||
are endemitic to
|
||
<collectingCountry id="F368761FA276FFF8FC2F069B7124FE11" box="[1021,1126,405,431]" name="Morocco" pageId="5" pageNumber="444">Morocco</collectingCountry>
|
||
. The remaining species are distributed in southeastern Europe and the Near East.
|
||
<taxonomicName id="4C7F4D0CA276FFF8FD3506B77674FE6D" authorityName="Laporte" authorityYear="1832" box="[743,822,441,467]" class="Insecta" family="Melolonthidae" genus="Anoxia" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="444" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B90BEA9DA276FFF8FD3506B77674FE6D" box="[743,822,441,467]" italics="true" pageId="5" pageNumber="444">Anoxia</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicName id="4C7F4D0CA276FFF8FCEE06B77143FE6C" authorityName="Bezděk & Kalashian & Ghrejyan" authorityYear="2019" baseAuthorityName="Zaitzev" baseAuthorityYear="1928" box="[828,1025,440,467]" class="Insecta" family="Melolonthidae" genus="Anoxia" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="444" phylum="Arthropoda" rank="species" species="maljuzhenkoi" subGenus="Protanoxia">
|
||
(
|
||
<emphasis id="B90BEA9DA276FFF8FC9706B77614FE6C" box="[837,854,441,466]" italics="true" pageId="5" pageNumber="444">P</emphasis>
|
||
.)
|
||
<emphasis id="B90BEA9DA276FFF8FCBE06B67143FE6C" box="[876,1025,440,466]" italics="true" pageId="5" pageNumber="444">maljuzhenkoi</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="B90BEA9DA276FFF8FBDC06B7715DFE6D" box="[1038,1055,441,467]" italics="true" pageId="5" pageNumber="444">A</emphasis>
|
||
.
|
||
<taxonomicName id="4C7F4D0CA276FFF8FBFE06B77028FE6D" authority="Baraud, 1989" authorityName="Baraud" authorityYear="1989" box="[1068,1386,440,467]" class="Insecta" family="Melolonthidae" genus="Anoxia" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="444" phylum="Arthropoda" rank="species" species="ciliciensis" subGenus="Protanoxia">
|
||
(
|
||
<emphasis id="B90BEA9DA276FFF8FBE706B77104FE6C" box="[1077,1094,441,466]" italics="true" pageId="5" pageNumber="444">P</emphasis>
|
||
.)
|
||
<emphasis id="B90BEA9DA276FFF8FB8E06B6718EFE6C" box="[1116,1228,440,466]" italics="true" pageId="5" pageNumber="444">ciliciensis</emphasis>
|
||
<bibRefCitation id="EFEE4B7EA276FFF8FB0006B67028FE6D" author="Baraud, J." box="[1234,1386,440,467]" pageId="5" pageNumber="444" pagination="273 - 284" refId="ref4143" refString="Baraud, J. (1989) Revision des Anoxia Castelnau d'Europe et d'Asie. 1 re note: le sous-genre Protanoxia Medvedev (Col. Melolonthidae). Bulletin de la Societe Entomologique de France, 93, 273 - 284." type="journal article" year="1989">Baraud, 1989</bibRefCitation>
|
||
</taxonomicName>
|
||
, and
|
||
<emphasis id="B90BEA9DA276FFF8FF4506D375EAFE49" box="[151,168,477,503]" italics="true" pageId="5" pageNumber="444">A</emphasis>
|
||
.
|
||
<taxonomicName id="4C7F4D0CA276FFF8FF6406D37754FE49" authority="Petrovitz, 1965" authorityName="Petrovitz" authorityYear="1965" box="[182,534,476,503]" class="Insecta" family="Melolonthidae" genus="Anoxia" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="444" phylum="Arthropoda" rank="species" species="smyrnensis" subGenus="Protanoxia">
|
||
(
|
||
<emphasis id="B90BEA9DA276FFF8FF1206D37593FE48" box="[192,209,477,502]" italics="true" pageId="5" pageNumber="444">P</emphasis>
|
||
.)
|
||
<emphasis id="B90BEA9DA276FFF8FF3A06D37426FE48" box="[232,356,477,502]" italics="true" pageId="5" pageNumber="444">smyrnensis</emphasis>
|
||
Petrovitz, 1965
|
||
</taxonomicName>
|
||
form a trio of habitually similar and rarely collected species, the males of which are characterized by an antennal club as long as the antennal shaft and by the presence of three external teeth on the protibia. Males of other
|
||
<taxonomicName id="4C7F4D0CA276FFF8FE74052B7761FD80" authorityName="Medvedev" authorityYear="1951" box="[422,547,549,574]" class="Insecta" family="Melolonthidae" genus="Anoxia" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="444" phylum="Arthropoda" rank="subGenus" subGenus="Protanoxia">
|
||
<emphasis id="B90BEA9DA276FFF8FE74052B7761FD80" box="[422,547,549,574]" italics="true" pageId="5" pageNumber="444">Protanoxia</emphasis>
|
||
</taxonomicName>
|
||
species differ from this trio in displaying an antennal club that is distinctly longer than the antennal shaft (
|
||
<emphasis id="B90BEA9DA276FFF8FE1D054774A2FDDD" box="[463,480,585,611]" italics="true" pageId="5" pageNumber="444">A</emphasis>
|
||
.
|
||
<taxonomicName id="4C7F4D0CA276FFF8FE3C0547761EFDDD" authority="Petrovitz, 1973" authorityName="Petrovitz" authorityYear="1973" box="[494,860,584,611]" class="Insecta" family="Melolonthidae" genus="Anoxia" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="444" phylum="Arthropoda" rank="species" species="laevimacula" subGenus="Protanoxia">
|
||
(
|
||
<emphasis id="B90BEA9DA276FFF8FE2A0547774BFDDC" box="[504,521,585,610]" italics="true" pageId="5" pageNumber="444">P</emphasis>
|
||
.)
|
||
<emphasis id="B90BEA9DA276FFF8FDF2054677EBFDDC" box="[544,681,584,610]" italics="true" pageId="5" pageNumber="444">laevimacula</emphasis>
|
||
Petrovitz, 1973
|
||
</taxonomicName>
|
||
) or by the presence of only an apical external tooth on the protibia (
|
||
<emphasis id="B90BEA9DA276FFF8FE9A0563741BFD39" box="[328,345,621,647]" italics="true" pageId="5" pageNumber="444">A</emphasis>
|
||
. (
|
||
<emphasis id="B90BEA9DA276FFF8FEA2056374C3FD38" box="[368,385,621,646]" italics="true" pageId="5" pageNumber="444">P</emphasis>
|
||
.)
|
||
<emphasis id="B90BEA9DA276FFF8FE45056274B8FD38" box="[407,506,620,646]" italics="true" pageId="5" pageNumber="444">orentalis</emphasis>
|
||
(Krynicki, 1832),
|
||
<emphasis id="B90BEA9DA276FFF8FD1A0563779BFD39" box="[712,729,621,647]" italics="true" pageId="5" pageNumber="444">A</emphasis>
|
||
.
|
||
<taxonomicName id="4C7F4D0CA276FFF8FD3505637169FD38" authority="Marseul, 1868" authorityName="Marseul" authorityYear="1868" box="[743,1067,620,647]" class="Insecta" family="Melolonthidae" genus="Anoxia" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="444" phylum="Arthropoda" rank="species" species="cingulata" subGenus="Protanoxia">
|
||
(
|
||
<emphasis id="B90BEA9DA276FFF8FD2205637643FD38" box="[752,769,621,646]" italics="true" pageId="5" pageNumber="444">P</emphasis>
|
||
.)
|
||
<emphasis id="B90BEA9DA276FFF8FCCA056276C0FD38" box="[792,898,620,646]" italics="true" pageId="5" pageNumber="444">cingulata</emphasis>
|
||
Marseul, 1868
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="B90BEA9DA276FFF8FBE705637104FD39" box="[1077,1094,621,647]" italics="true" pageId="5" pageNumber="444">A</emphasis>
|
||
.
|
||
<taxonomicName id="4C7F4D0CA276FFF8FB8605637028FD39" authority="Keith, 2003" authorityName="Keith" authorityYear="2003" box="[1108,1386,620,647]" class="Insecta" family="Melolonthidae" genus="Anoxia" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="444" phylum="Arthropoda" rank="species" species="baraudi" subGenus="Protanoxia">
|
||
(
|
||
<emphasis id="B90BEA9DA276FFF8FB8F0563712CFD38" box="[1117,1134,621,646]" italics="true" pageId="5" pageNumber="444">P</emphasis>
|
||
.)
|
||
<emphasis id="B90BEA9DA276FFF8FB570562719CFD38" box="[1157,1246,620,646]" italics="true" pageId="5" pageNumber="444">baraudi</emphasis>
|
||
Keith, 2003
|
||
</taxonomicName>
|
||
, and
|
||
<emphasis id="B90BEA9DA276FFF8FF45059F75EAFD15" box="[151,168,657,683]" italics="true" pageId="5" pageNumber="444">A</emphasis>
|
||
.
|
||
<taxonomicName id="4C7F4D0CA276FFF8FF64059F7476FD14" authorityName="Dewailly" authorityYear="1957" box="[182,308,657,683]" class="Insecta" family="Melolonthidae" genus="Anoxia" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="444" phylum="Arthropoda" rank="species" species="rotroui" subGenus="Protanoxia">
|
||
(
|
||
<emphasis id="B90BEA9DA276FFF8FF6D059F7592FD14" box="[191,208,657,682]" italics="true" pageId="5" pageNumber="444">P</emphasis>
|
||
.)
|
||
<emphasis id="B90BEA9DA276FFF8FF35059F7476FD14" box="[231,308,657,682]" italics="true" pageId="5" pageNumber="444">rotroui</emphasis>
|
||
</taxonomicName>
|
||
), see also
|
||
<bibRefCitation id="EFEE4B7EA276FFF8FE7B059E7709FD15" author="Baraud, J." box="[425,587,656,683]" pageId="5" pageNumber="444" pagination="273 - 284" refId="ref4143" refString="Baraud, J. (1989) Revision des Anoxia Castelnau d'Europe et d'Asie. 1 re note: le sous-genre Protanoxia Medvedev (Col. Melolonthidae). Bulletin de la Societe Entomologique de France, 93, 273 - 284." type="journal article" year="1989">Baraud (1989)</bibRefCitation>
|
||
and
|
||
<bibRefCitation id="EFEE4B7EA276FFF8FD53059E7661FD15" author="Rittner, O." box="[641,803,656,683]" pageId="5" pageNumber="444" pagination="99 - 108" refId="ref4642" refString="Rittner, O. (2016) Synopsis of the Melolonthini (Scarabaeidae: Melolonthinae) of Israel, with a first description of the female of Anoxia (Protanoxia) laevimacula Petrovitz, 1973. Israel Journal of Entomology, 46, 99 - 108." type="journal article" year="2016">Rittner (2016)</bibRefCitation>
|
||
. Males of the Moroccan species
|
||
<emphasis id="B90BEA9DA276FFF8FB5C059F71DDFD15" box="[1166,1183,657,683]" italics="true" pageId="5" pageNumber="444">A</emphasis>
|
||
.
|
||
<taxonomicName id="4C7F4D0CA276FFF8FB7F059F7070FD14" authorityName="Dewailly" authorityYear="1957" box="[1197,1330,656,683]" class="Insecta" family="Melolonthidae" genus="Anoxia" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="444" phylum="Arthropoda" rank="species" species="kocheri" subGenus="Protanoxia">
|
||
(
|
||
<emphasis id="B90BEA9DA276FFF8FB64059F7185FD14" box="[1206,1223,657,682]" italics="true" pageId="5" pageNumber="444">P</emphasis>
|
||
.)
|
||
<emphasis id="B90BEA9DA276FFF8FB0C059E7070FD14" box="[1246,1330,656,682]" italics="true" pageId="5" pageNumber="444">kocheri</emphasis>
|
||
</taxonomicName>
|
||
also bear a barely visible basal external tooth on the protibia (
|
||
<bibRefCitation id="EFEE4B7EA276FFF8FD3A05BA763DFD71" author="Baraud, J." box="[744,895,692,719]" pageId="5" pageNumber="444" pagination="253 - 261" refId="ref4106" refString="Baraud, J. (1980) Coleopteres Scarabaeoidea de l'Afrique du Nord. 5 e note: Melolonthinae nouveaux ou meconnus. Bulletin de la Societe Entomologique de France, 85, 253 - 261." type="journal article" year="1980">Baraud 1980</bibRefCitation>
|
||
). For discrimination of
|
||
<emphasis id="B90BEA9DA276FFF8FB5E05BB71DFFD71" box="[1164,1181,693,719]" italics="true" pageId="5" pageNumber="444">A</emphasis>
|
||
.
|
||
<taxonomicName id="4C7F4D0CA276FFF8FB7F05BB7037FD70" authorityName="Bezděk & Kalashian & Ghrejyan" authorityYear="2019" baseAuthorityName="Zaitzev" baseAuthorityYear="1928" box="[1197,1397,692,719]" class="Insecta" family="Melolonthidae" genus="Anoxia" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="444" phylum="Arthropoda" rank="species" species="maljuzhenkoi" subGenus="Protanoxia">
|
||
(
|
||
<emphasis id="B90BEA9DA276FFF8FB6405BB7185FD70" box="[1206,1223,693,718]" italics="true" pageId="5" pageNumber="444">P</emphasis>
|
||
.)
|
||
<emphasis id="B90BEA9DA276FFF8FB3205BA7037FD70" box="[1248,1397,692,718]" italics="true" pageId="5" pageNumber="444">maljuzhenkoi</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="B90BEA9DA276FFF8FA5605BB70D7FD71" box="[1412,1429,693,719]" italics="true" pageId="5" pageNumber="444">A</emphasis>
|
||
.
|
||
<taxonomicName id="4C7F4D0CA276FFF8FF4505D7747BFD4C" authorityName="Baraud" authorityYear="1989" box="[151,313,728,755]" class="Insecta" family="Melolonthidae" genus="Anoxia" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="444" phylum="Arthropoda" rank="species" species="ciliciensis" subGenus="Protanoxia">
|
||
(
|
||
<emphasis id="B90BEA9DA276FFF8FF7305D775F0FD4C" box="[161,178,729,754]" italics="true" pageId="5" pageNumber="444">P</emphasis>
|
||
.)
|
||
<emphasis id="B90BEA9DA276FFF8FF1B05D6747BFD4C" box="[201,313,728,754]" italics="true" pageId="5" pageNumber="444">ciliciensis</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<emphasis id="B90BEA9DA276FFF8FEA405D774C5FD4D" box="[374,391,729,755]" italics="true" pageId="5" pageNumber="444">A</emphasis>
|
||
.
|
||
<taxonomicName id="4C7F4D0CA276FFF8FE4705D77701FD4C" authorityName="Petrovitz" authorityYear="1965" box="[405,579,729,755]" class="Insecta" family="Melolonthidae" genus="Anoxia" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="444" phylum="Arthropoda" rank="species" species="smyrnensis" subGenus="Protanoxia">
|
||
(
|
||
<emphasis id="B90BEA9DA276FFF8FE4D05D774F2FD4C" box="[415,432,729,754]" italics="true" pageId="5" pageNumber="444">P</emphasis>
|
||
.)
|
||
<emphasis id="B90BEA9DA276FFF8FE1505D77701FD4C" box="[455,579,729,754]" italics="true" pageId="5" pageNumber="444">smyrnensis</emphasis>
|
||
</taxonomicName>
|
||
, see identification key below.
|
||
</paragraph>
|
||
<paragraph id="8BC0368FA276FFF8FF1505F2758DFB89" blockId="5.[151,1437,152,1115]" pageId="5" pageNumber="444">
|
||
<emphasis id="B90BEA9DA276FFF8FF1505F274D8FCA9" bold="true" box="[199,410,764,791]" pageId="5" pageNumber="444">Collecting events.</emphasis>
|
||
All recently recorded specimens from Arazap were collected by light trap (J. Šumpich and A. Pavlíčko, personal communication). The biotope, where the specimens were collected, is very specific (
|
||
<figureCitation id="13442A0AA276FFF8FABD042E75F4FCE1" captionStart="FIGURE 17" captionStartId="4.[151,250,899,924]" captionTargetBox="[177,1411,286,874]" captionTargetId="figure@4.[177,1411,181,874]" captionTargetPageId="4" captionText="FIGURE 17. “Vordan Karmir” State Sanctuary, Armenia – the biotope of Anoxia (Protanoxia) maljuzhenkoi (Zaitzev, 1928). Photograph by Jan Šumpich, June 2017." figureDoi="http://doi.org/10.5281/zenodo.3442504" httpUri="https://zenodo.org/record/3442504/files/figure.png" pageId="5" pageNumber="444">Fig. 17</figureCitation>
|
||
). According to the European Nature Information System habitat classification (
|
||
<bibRefCitation id="EFEE4B7EA276FFF8FBFF044A71BDFCE1" author="Davies, C. E. & Moss, D. & Hill, M. O." box="[1069,1279,836,863]" pageId="5" pageNumber="444" refId="ref4343" refString="Davies, C. E., Moss, D. & Hill, M. O. (2004) EUNIS habitat classification revised 2004. European Environment Agency, European Topic Centre on Nature Protection and Biodiversity, Paris, 310 pp." type="book" year="2004">
|
||
Davies
|
||
<emphasis id="B90BEA9DA276FFF8FB56044B71FEFCE0" box="[1156,1212,836,862]" italics="true" pageId="5" pageNumber="444">et al.</emphasis>
|
||
2004
|
||
</bibRefCitation>
|
||
), it belongs to the category
|
||
<geoCoordinate id="EE4B5048A276FFF8FEFB04667422FC3C" box="[297,352,872,898]" degrees="6.2" direction="east" orientation="longitude" pageId="5" pageNumber="444" precision="5555" value="6.2">E6.2</geoCoordinate>
|
||
. (continental inland salt steppes), habitats completely included in Resolution 4 of the Bern Convention as specifically protected. The particular biotope belongs to the subcategories
|
||
<geoCoordinate id="EE4B5048A276FFF8FBB2048271E0FC19" box="[1120,1186,908,935]" degrees="6.25" direction="east" orientation="longitude" pageId="5" pageNumber="444" precision="555" value="6.25">E6.25</geoCoordinate>
|
||
-AM (salt steppes and solonchaks grasslands) and
|
||
<geoCoordinate id="EE4B5048A276FFF8FE0204BE7763FC75" box="[464,545,944,971]" degrees="6.251" direction="east" orientation="longitude" pageId="5" pageNumber="444" precision="55" value="6.251">E6.251</geoCoordinate>
|
||
-AM (
|
||
<emphasis id="B90BEA9DA276FFF8FDBA04BE7798FC74" box="[616,730,944,970]" italics="true" pageId="5" pageNumber="444">Aleuropus</emphasis>
|
||
(
|
||
<taxonomicName id="4C7F4D0CA276FFF8FD3B04BF760AFC75" box="[745,840,945,971]" class="Liliopsida" family="Poaceae" kingdom="Plantae" order="Poales" pageId="5" pageNumber="444" phylum="Tracheophyta" rank="family">Poaceae</taxonomicName>
|
||
) dominated habitats) (
|
||
<bibRefCitation id="EFEE4B7EA276FFF8FB9604BE70D3FC75" author="Fayvush, G. M. & Aleksanyan, A. S." box="[1092,1425,944,971]" pageId="5" pageNumber="444" refId="ref4388" refString="Fayvush, G. M. & Aleksanyan, A. S. (2016) Habitats of Armenia. Mestoobitaniya Armenii. National Academy of Sciences of the Republic of Armenia, Institute of Botany, Yerevan, Armenia, 360 pp. [In Russian and English]" type="book" year="2016">Fayvush & Aleksanyan 2016</bibRefCitation>
|
||
). The latter being one of the few remained spots of this
|
||
<typeStatus id="54C4882DA276FFF8FD2304DB7663FC51" box="[753,801,981,1007]" pageId="5" pageNumber="444">type</typeStatus>
|
||
of habitat and it is protected as a biotope of the critically endangered Araratian cochineal or Vordan karmir –
|
||
<taxonomicName id="4C7F4D0CA276FFF8FD0504F671CBFBAD" authority="(Brandt, 1833)" authorityName="Brandt" authorityYear="1833" baseAuthorityName="Brandt" baseAuthorityYear="1833" box="[727,1161,1016,1043]" class="Insecta" family="Margarodidae" genus="Porphyrophora" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="444" phylum="Arthropoda" rank="species" species="hamelii">
|
||
<emphasis id="B90BEA9DA276FFF8FD0504F6769DFBAC" box="[727,991,1016,1042]" italics="true" pageId="5" pageNumber="444">Porphyrophora hamelii</emphasis>
|
||
(Brandt, 1833)
|
||
</taxonomicName>
|
||
(
|
||
<taxonomicName id="4C7F4D0CA276FFF8FB4504F67051FBAC" box="[1175,1299,1016,1042]" class="Insecta" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="444" phylum="Arthropoda" rank="order">Hemiptera</taxonomicName>
|
||
: Margarodi- dae).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3656504A276FFF8FF15034E71F3FBE4" box="[199,1201,1088,1115]" pageId="5" pageNumber="444" type="distribution">
|
||
<paragraph id="8BC0368FA276FFF8FF15034E71F3FBE4" blockId="5.[151,1437,152,1115]" box="[199,1201,1088,1115]" pageId="5" pageNumber="444">
|
||
<emphasis id="B90BEA9DA276FFF8FF15034E7423FBE5" bold="true" box="[199,353,1088,1115]" pageId="5" pageNumber="444">Distribution.</emphasis>
|
||
<collectingCountry id="F368761FA276FFF8FEB4034E74A3FBE4" box="[358,481,1088,1114]" name="Azerbaijan" pageId="5" pageNumber="444">Azerbaijan</collectingCountry>
|
||
(
|
||
<collectingRegion id="49BBF86DA276FFF8FE3D034E77C7FBE4" box="[495,645,1088,1114]" country="Azerbaijan" name="Naxcivan" pageId="5" pageNumber="444">Nakhichevan</collectingRegion>
|
||
Autonomous Republic), first record for
|
||
<collectingCountry id="F368761FA276FFF8FB95034E71EFFBE4" box="[1095,1197,1088,1114]" name="Armenia" pageId="5" pageNumber="444">Armenia</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |