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<mods:title id="55E9D585ED57430A3485298B1903714E">An illustrated key to the species of subgenus Gyrostoma Kirby, 1828 (Hymenoptera, Vespidae, Polistinae) from China, with discovery of Polistes (Gyrostoma) tenuispunctia Kim, 2001</mods:title>
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<treatment id="03D687963E7CFFC477DAFA86928D9EA7" ID-DOI="http://doi.org/10.5281/zenodo.5696268" ID-GBIF-Taxon="119422061" ID-Zenodo-Dep="5696268" LSID="urn:lsid:plazi:treatment:03D687963E7CFFC477DAFA86928D9EA7" httpUri="http://treatment.plazi.org/id/03D687963E7CFFC477DAFA86928D9EA7" lastPageId="13" lastPageNumber="389" pageId="10" pageNumber="387">
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<paragraph id="8BC036803E7CFFC377DAFA8693839A79" blockId="10.[151,708,1371,1431]" box="[151,708,1371,1398]" pageId="10" pageNumber="387">
<heading id="D08881EC3E7CFFC377DAFA8693839A79" bold="true" box="[151,708,1371,1398]" fontSize="11" level="1" pageId="10" pageNumber="387" reason="1">
<taxonomicName id="4C7F4D033E7CFFC377DAFA8693839A79" ID-CoL="4KYH3" authority="Cameron, 1900" authorityName="Cameron" authorityYear="1900" box="[151,708,1371,1398]" class="Insecta" family="Eumenidae" genus="Polistes" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="387" phylum="Arthropoda" rank="species" species="rothneyi" subGenus="Gyrostoma">
<emphasis id="B90BEA923E7CFFC377DAFA8693839A79" bold="true" box="[151,708,1371,1398]" pageId="10" pageNumber="387">
<emphasis id="B90BEA923E7CFFC377DAFA8691B39A7A" bold="true" box="[151,244,1371,1397]" italics="true" pageId="10" pageNumber="387">Polistes</emphasis>
(
<emphasis id="B90BEA923E7CFFC37649FA8690CF9A7A" bold="true" box="[260,392,1371,1397]" italics="true" pageId="10" pageNumber="387">Gyrostoma</emphasis>
)
<emphasis id="B90BEA923E7CFFC376D7FA8690B89A7A" bold="true" box="[410,511,1371,1397]" italics="true" pageId="10" pageNumber="387">rothneyi</emphasis>
<bibRefCitation id="EFEE4B713E7CFFC3754BFA8193839A79" author="Cameron" box="[518,708,1372,1398]" pageId="10" pageNumber="387" refString="Cameron, P. (1900) Descriptions of new genera and species of Hymenoptera. Annals and Magazine of Natural History, 6, 410 - 419, 495 - 506, 530 - 539." type="journal article" year="1900">
<collectingCountry id="F36876103E7CFFC3754BFA81933A9A79" box="[518,637,1372,1398]" name="Cameroon" pageId="10" pageNumber="387">Cameron</collectingCountry>
, 1900
</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BC036803E7CFFC377DAFAA390609A98" blockId="10.[151,708,1371,1431]" box="[151,295,1406,1431]" pageId="10" pageNumber="387">
(
<figureCitation id="13442A053E7CFFC377D2FAA390589A98" box="[159,287,1406,1431]" captionStart-0="FIGURE 20" captionStart-1="FIGURE 21" captionStart-2="FIGURE 22" captionStartId-0="11.[151,250,1485,1507]" captionStartId-1="13.[151,250,1776,1798]" captionStartId-2="14.[151,250,1341,1363]" captionTargetBox-0="[302,1284,409,1462]" captionTargetBox-1="[302,1284,468,1753]" captionTargetBox-2="[302,1284,193,1319]" captionTargetId-0="figure@11.[302,1284,409,1463]" captionTargetId-1="figure@13.[302,1284,468,1754]" captionTargetId-2="figure@14.[302,1284,193,1319]" captionTargetPageId-0="11" captionTargetPageId-1="13" captionTargetPageId-2="14" captionText-0="FIGURE 20. Polistes rothneyi Cameron, 1900, Southwest plateau group (female). A, C, E, G, dorsal view; B, D, F, H, lateral view: A B, f. rothneyi (det. van der Vecht) from Shillong, India; C D, f. tibetanus van der Vecht, 1968 from Tibet, paratype; E F, specimens from Yunnan, showing the color pattern between f. rothneyi and f. tibetanus; G H, specimens from Yunnan, showing the color pattern as? subs. A (van der Vecht, 1968: 103, fig. 7)." captionText-1="FIGURE 21. Polistes rothneyi Cameron, 1900, Common yellow-black group (female). A, C, E, G, I, dorsal view; B, D, F, H, J, lateral view: A B, f. grahami van der Vecht, 1968 from Sichuan, paratype (showing mesonotum with two yellow stripes); C D, f. grahami van der Vecht, 1968 from Fujian, paratype (showing mesonotum with four yellow stripes); E F, f. iwatai van der Vecht, 1968 from Japan, paratype (showing color pattern very similar as A-B f. grahami, but pronotum mostly yellow); G H, specimens from Hainan with the color pattern same as f. hainanensis van der Vecht, 1968, showing the apical band of the first metasomal tergite entirely yellow; I J, f. gressitti van der Vecht, 1968 from Taiwan, paratype (showing pronotum entirely yellow); K L, f. yayeyamae Matsumura, 1908 from Iriomote Loochoo Is., paratype (showing body mainly yellow)." captionText-2="FIGURE 22. Polistes rothneyi Cameron, 1900, Northeast group (female) (showing the colour pattern variation). A, C, E, G, I, J, dorsal view; B, D, F, H, lateral view: A B, specimen from Shenyang, Liaoning; C D, specimen from Shenyang, Kuandian; E F, specimen from Qianshan, Liaoning; G H, f. koreanus van der Vecht, 1968 from Seoul area, paratype (with pronotum black except narrow yellow margin, first metasomal tergite black with a narrow apical band); I, specimen from Siping, Jilin; J, specimen from Xinbin, Liaoning." httpUri-0="https://zenodo.org/record/253078/files/figure.png" httpUri-1="https://zenodo.org/record/253079/files/figure.png" httpUri-2="https://zenodo.org/record/253080/files/figure.png" pageId="10" pageNumber="387">Figs 2022</figureCitation>
)
</paragraph>
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<paragraph id="8BC036803E7CFFC377DAFA1B92B29AD3" blockId="10.[151,1436,1478,1930]" box="[151,1013,1478,1500]" pageId="10" pageNumber="387">
<treatmentCitationGroup id="AB6F11AE3E7CFFC377DAFA1B92B29AD3" box="[151,1013,1478,1500]" pageId="10" pageNumber="387">
<taxonomicName id="4C7F4D033E7CFFC377DAFA1B93579AD3" ID-CoL="4KYH3" authority="Cameron 1900: 410" authorityName="Cameron" authorityPageNumber="410" authorityYear="1900" box="[151,528,1478,1500]" class="Insecta" family="Eumenidae" genus="Polistes" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="387" phylum="Arthropoda" rank="species" species="rothneyi">
<emphasis id="B90BEA923E7CFFC377DAFA1B90079AD3" box="[151,320,1478,1500]" italics="true" pageId="10" pageNumber="387">Polistes rothneyi</emphasis>
<treatmentCitation id="0ADE10913E7CFFC3760AFA1B93579AD3" author="Cameron" box="[327,528,1478,1500]" page="410" pageId="10" pageNumber="387" year="1900">
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<collectingCountry id="F36876103E7CFFC3760AFA1B90E39AD3" box="[327,420,1478,1500]" name="Cameroon" pageId="10" pageNumber="387">Cameron</collectingCountry>
1900
</bibRefCitation>
: 410
</treatmentCitation>
</taxonomicName>
(♂, Bengal,
<collectingCountry id="F36876103E7CFFC375DEFA1B938E9AD3" box="[659,713,1478,1500]" name="India" pageId="10" pageNumber="387">India</collectingCountry>
, deposited in OUM, Oxford).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BC036803E7CFFC377DAFA3895B19915" blockId="10.[151,1436,1478,1930]" pageId="10" pageNumber="387">
<treatmentCitationGroup id="AB6F11AE3E7CFFC377DAFA3895B19915" pageId="10" pageNumber="387">
<taxonomicName id="4C7F4D033E7CFFC377DAFA38930E9AF4" ID-CoL="4KYHB" authority="Cameron 1900: 411" authorityName="Cameron" authorityPageNumber="411" authorityYear="1900" box="[151,585,1509,1531]" class="Insecta" family="Eumenidae" genus="Polistes" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="387" phylum="Arthropoda" rank="species" species="rufolineatus">
<emphasis id="B90BEA923E7CFFC377DAFA38902B9AF4" box="[151,364,1509,1531]" italics="true" pageId="10" pageNumber="387">Polistes rufolineatus</emphasis>
<treatmentCitation id="0ADE10913E7CFFC37635FA38930E9AF4" author="Cameron" box="[376,585,1509,1531]" page="411" pageId="10" pageNumber="387" year="1900">
<bibRefCitation id="EFEE4B713E7CFFC37635FA3893519AF4" author="Cameron" box="[376,534,1509,1531]" pageId="10" pageNumber="387" refString="Cameron, P. (1900) Descriptions of new genera and species of Hymenoptera. Annals and Magazine of Natural History, 6, 410 - 419, 495 - 506, 530 - 539." type="journal article" year="1900">
<collectingCountry id="F36876103E7CFFC37635FA3890929AF4" box="[376,469,1509,1531]" name="Cameroon" pageId="10" pageNumber="387">Cameron</collectingCountry>
1900
</bibRefCitation>
: 411
</treatmentCitation>
</taxonomicName>
(
<typeStatus id="54C488223E7CFFC37511FA3893FA9AF4" box="[604,701,1509,1531]" pageId="10" pageNumber="387" type="lectotype">lectotype</typeStatus>
, ♂, designated by van der Vecht, 1964, Khasia Hills, Assam,
<collectingCountry id="F36876103E7CFFC3722CFA3894DF9AF4" box="[1377,1432,1509,1531]" name="India" pageId="10" pageNumber="387">India</collectingCountry>
, deposited in BMNH, London). Synonymized under
<taxonomicName id="4C7F4D033E7CFFC3759BF9D9928B9915" ID-CoL="4KYH3" authority="Cameron, 1900" authorityName="Cameron" authorityYear="1900" box="[726,972,1540,1562]" class="Insecta" family="Eumenidae" genus="Polistes" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="387" phylum="Arthropoda" rank="species" species="rothneyi">
<emphasis id="B90BEA923E7CFFC3759BF9D9926D9915" box="[726,810,1540,1562]" italics="true" pageId="10" pageNumber="387">rothneyi</emphasis>
<bibRefCitation id="EFEE4B713E7CFFC3747DF9D9928B9915" author="Cameron" box="[816,972,1540,1562]" pageId="10" pageNumber="387" refString="Cameron, P. (1900) Descriptions of new genera and species of Hymenoptera. Annals and Magazine of Natural History, 6, 410 - 419, 495 - 506, 530 - 539." type="journal article" year="1900">
<collectingCountry id="F36876103E7CFFC3747DF9D992D79915" box="[816,912,1540,1562]" name="Cameroon" pageId="10" pageNumber="387">Cameron</collectingCountry>
, 1900
</bibRefCitation>
</taxonomicName>
by van der
<treatmentCitation id="0ADE10913E7CFFC3730EF9D995B59915" author="Vecht" box="[1091,1266,1540,1562]" page="102" pageId="10" pageNumber="387" year="1968">
<bibRefCitation id="EFEE4B713E7CFFC3730EF9D995F89915" author="Vecht" box="[1091,1215,1540,1562]" pageId="10" pageNumber="387" refString="van der Vecht, J. (1968) The geographic variation of Polistes (Megapolistes subg. n.) rothneyi Cameron. Bijdragen Tot de Dierkunde, 38, 97 - 109." type="journal article" year="1968">Vecht, 1968</bibRefCitation>
: 102
</treatmentCitation>
.
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</paragraph>
<paragraph id="8BC036803E7CFFC377DAF9FE958B9937" blockId="10.[151,1436,1478,1930]" box="[151,1228,1571,1593]" pageId="10" pageNumber="387">
<treatmentCitationGroup id="AB6F11AE3E7CFFC377DAF9FE958B9937" box="[151,1228,1571,1593]" pageId="10" pageNumber="387">
<taxonomicName id="4C7F4D033E7CFFC377DAF9FE93C29936" ID-CoL="5KKCY" authority="Matsumura 1908: 6" authorityName="Matsumura" authorityPageNumber="6" authorityYear="1908" box="[151,645,1571,1593]" class="Insecta" family="Eumenidae" genus="Polistes" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="387" phylum="Arthropoda" rank="subSpecies" species="rothneyi" subSpecies="yayeyamae">
<emphasis id="B90BEA923E7CFFC377DAF9FE90F09936" box="[151,439,1571,1593]" italics="true" pageId="10" pageNumber="387">Polistes rothneyi yayeyamae</emphasis>
<treatmentCitation id="0ADE10913E7CFFC376F3F9F993C29936" author="Matsumura" box="[446,645,1571,1593]" page="6" pageId="10" pageNumber="387" year="1908">
<bibRefCitation id="EFEE4B713E7CFFC376F3F9F9932A9936" author="Matsumura" box="[446,621,1571,1593]" pageId="10" pageNumber="387" refString="Matsumura, S. (1908) Nihon Ekichu Mokuroku [List of Japanese Beneficial Insects]. Rokumeikan, Tokyo, 174 pp. [in Japanese]" type="book" year="1908">Matsumura 1908</bibRefCitation>
: 6
</treatmentCitation>
</taxonomicName>
, nom nudum (
<typeStatus id="54C488223E7CFFC3745AF9FE92339936" box="[791,884,1571,1593]" pageId="10" pageNumber="387" type="holotype">holotype</typeStatus>
, ♀, Ryukyu Isl.,
<collectingCountry id="F36876103E7CFFC37352F9F9951B9936" box="[1055,1116,1572,1593]" name="Japan" pageId="10" pageNumber="387">Japan</collectingCountry>
),
<emphasis id="B90BEA923E7CFFC37323F9FE958B9937" bold="true" box="[1134,1228,1571,1592]" pageId="10" pageNumber="387">syn. nov.</emphasis>
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BC036803E7CFFC377DAF99C93E3997A" blockId="10.[151,1436,1478,1930]" pageId="10" pageNumber="387">
<treatmentCitationGroup id="AB6F11AE3E7CFFC377DAF99C93E3997A" pageId="10" pageNumber="387">
<taxonomicName id="4C7F4D033E7CFFC377DAF99C90E29958" ID-CoL="5KKCK" box="[151,421,1601,1623]" class="Insecta" family="Eumenidae" genus="Polistes" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="387" phylum="Arthropoda" rank="subSpecies" species="rothneyi" subSpecies="grahami">
<emphasis id="B90BEA923E7CFFC377DAF99C90E29958" box="[151,421,1601,1623]" italics="true" pageId="10" pageNumber="387">Polistes rothneyi grahami</emphasis>
</taxonomicName>
van der
<treatmentCitation id="0ADE10913E7CFFC37544F99C93F09958" author="Vecht" box="[521,695,1601,1623]" page="104" pageId="10" pageNumber="387" year="1968">
<bibRefCitation id="EFEE4B713E7CFFC37544F99C93C59958" author="Vecht" box="[521,642,1601,1623]" pageId="10" pageNumber="387" refString="van der Vecht, J. (1968) The geographic variation of Polistes (Megapolistes subg. n.) rothneyi Cameron. Bijdragen Tot de Dierkunde, 38, 97 - 109." type="journal article" year="1968">Vecht 1968</bibRefCitation>
: 104
</treatmentCitation>
(
<typeStatus id="54C488223E7CFFC37585F99C92629958" box="[712,805,1601,1623]" pageId="10" pageNumber="387" type="holotype">holotype</typeStatus>
, ♀, Hangzhou, Zhejiang,
<collectingCountry id="F36876103E7CFFC37374F99C953F9958" box="[1081,1144,1601,1623]" name="China" pageId="10" pageNumber="387">China</collectingCountry>
, deposited in
<collectingCountry id="F36876103E7CFFC3725DF99C94DC9958" box="[1296,1435,1601,1623]" name="United States of America" pageId="10" pageNumber="387">United States</collectingCountry>
National Museum, Washington, DC),
<emphasis id="B90BEA923E7CFFC3750AF9BD93E3997A" bold="true" box="[583,676,1632,1653]" pageId="10" pageNumber="387">syn. nov.</emphasis>
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</paragraph>
<paragraph id="8BC036803E7CFFC377DAF9A392E199BC" blockId="10.[151,1436,1478,1930]" pageId="10" pageNumber="387">
<treatmentCitationGroup id="AB6F11AE3E7CFFC377DAF9A392E199BC" pageId="10" pageNumber="387">
<taxonomicName id="4C7F4D033E7CFFC377DAF9A39089999B" ID-CoL="5KKCM" box="[151,462,1662,1684]" class="Insecta" family="Eumenidae" genus="Polistes" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="387" phylum="Arthropoda" rank="subSpecies" species="rothneyi" subSpecies="hainanensis">
<emphasis id="B90BEA923E7CFFC377DAF9A39089999B" box="[151,462,1662,1684]" italics="true" pageId="10" pageNumber="387">Polistes rothneyi hainanensis</emphasis>
</taxonomicName>
van der
<treatmentCitation id="0ADE10913E7CFFC37571F9A393B7999B" author="Vecht" box="[572,752,1662,1684]" page="106" pageId="10" pageNumber="387" year="1968">
<bibRefCitation id="EFEE4B713E7CFFC37571F9A393FD999B" author="Vecht" box="[572,698,1662,1684]" pageId="10" pageNumber="387" refString="van der Vecht, J. (1968) The geographic variation of Polistes (Megapolistes subg. n.) rothneyi Cameron. Bijdragen Tot de Dierkunde, 38, 97 - 109." type="journal article" year="1968">Vecht 1968</bibRefCitation>
: 106
</treatmentCitation>
(
<typeStatus id="54C488223E7CFFC37448F9A39225999B" box="[773,866,1662,1684]" pageId="10" pageNumber="387" type="holotype">holotype</typeStatus>
, ♀, Nada, Hainan,
<collectingCountry id="F36876103E7CFFC3730DF9A39538999B" box="[1088,1151,1662,1684]" name="China" pageId="10" pageNumber="387">China</collectingCountry>
, deposited in Museum of Comparative Zoology, Harvard University, Cambridge, MCZ),
<emphasis id="B90BEA923E7CFFC37404F94392E199BC" bold="true" box="[841,934,1694,1715]" pageId="10" pageNumber="387">syn. nov.</emphasis>
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BC036803E7CFFC377DAF961943D99DE" blockId="10.[151,1436,1478,1930]" box="[151,1402,1724,1746]" pageId="10" pageNumber="387">
<treatmentCitationGroup id="AB6F11AE3E7CFFC377DAF961943D99DE" box="[151,1402,1724,1746]" pageId="10" pageNumber="387">
<taxonomicName id="4C7F4D033E7CFFC377DAF96190C199DD" ID-CoL="5KKCQ" box="[151,390,1724,1746]" class="Insecta" family="Eumenidae" genus="Polistes" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="387" phylum="Arthropoda" rank="subSpecies" species="rothneyi" subSpecies="iwatai">
<emphasis id="B90BEA923E7CFFC377DAF96190C199DD" box="[151,390,1724,1746]" italics="true" pageId="10" pageNumber="387">Polistes rothneyi iwatai</emphasis>
</taxonomicName>
van der
<treatmentCitation id="0ADE10913E7CFFC37693F96193C399DD" author="Vecht" box="[478,644,1724,1746]" page="104" pageId="10" pageNumber="387" year="1968">
<bibRefCitation id="EFEE4B713E7CFFC37693F961931399DD" author="Vecht" box="[478,596,1724,1746]" pageId="10" pageNumber="387" refString="van der Vecht, J. (1968) The geographic variation of Polistes (Megapolistes subg. n.) rothneyi Cameron. Bijdragen Tot de Dierkunde, 38, 97 - 109." type="journal article" year="1968">Vecht 1968</bibRefCitation>
: 104
</treatmentCitation>
(
<typeStatus id="54C488223E7CFFC375DCF96193AA99DD" box="[657,749,1724,1746]" pageId="10" pageNumber="387" type="holotype">holotype</typeStatus>
, ♀, Takayama,
<collectingCountry id="F36876103E7CFFC374C1F960928F99DD" box="[908,968,1725,1746]" name="Japan" pageId="10" pageNumber="387">Japan</collectingCountry>
, deposited in MCZ, Cambridge),
<emphasis id="B90BEA923E7CFFC37250F961943D99DE" bold="true" box="[1309,1402,1724,1745]" pageId="10" pageNumber="387">syn. nov.</emphasis>
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BC036803E7CFFC377DAF90690639800" blockId="10.[151,1436,1478,1930]" pageId="10" pageNumber="387">
<treatmentCitationGroup id="AB6F11AE3E7CFFC377DAF90690639800" pageId="10" pageNumber="387">
<taxonomicName id="4C7F4D033E7CFFC377DAF90690DB99FE" ID-CoL="5KKCL" box="[151,412,1755,1777]" class="Insecta" family="Eumenidae" genus="Polistes" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="387" phylum="Arthropoda" rank="subSpecies" species="rothneyi" subSpecies="gressitti">
<emphasis id="B90BEA923E7CFFC377DAF90690DB99FE" box="[151,412,1755,1777]" italics="true" pageId="10" pageNumber="387">Polistes rothneyi gressitti</emphasis>
</taxonomicName>
van der
<treatmentCitation id="0ADE10913E7CFFC376B6F90693E299FE" author="Vecht" box="[507,677,1755,1777]" page="106" pageId="10" pageNumber="387" year="1968">
<bibRefCitation id="EFEE4B713E7CFFC376B6F906933499FE" author="Vecht" box="[507,627,1755,1777]" pageId="10" pageNumber="387" refString="van der Vecht, J. (1968) The geographic variation of Polistes (Megapolistes subg. n.) rothneyi Cameron. Bijdragen Tot de Dierkunde, 38, 97 - 109." type="journal article" year="1968">Vecht 1968</bibRefCitation>
: 106
</treatmentCitation>
(
<typeStatus id="54C488223E7CFFC375F8F906925699FE" box="[693,785,1755,1777]" pageId="10" pageNumber="387" type="holotype">holotype</typeStatus>
, ♀, Taihanroku,
<collectingCountry id="F36876103E7CFFC3748CF906954999FE" box="[961,1038,1755,1777]" name="Taiwan" pageId="10" pageNumber="387">Taiwan</collectingCountry>
,
<collectingCountry id="F36876103E7CFFC37357F906951E99FE" box="[1050,1113,1755,1777]" name="China" pageId="10" pageNumber="387">China</collectingCountry>
, deposited in RMNH, Leiden),
<emphasis id="B90BEA923E7CFFC3778AF92790639800" bold="true" box="[199,292,1786,1807]" pageId="10" pageNumber="387">syn. nov.</emphasis>
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BC036803E7CFFC377DAF8C594159822" blockId="10.[151,1436,1478,1930]" box="[151,1362,1816,1838]" pageId="10" pageNumber="387">
<treatmentCitationGroup id="AB6F11AE3E7CFFC377DAF8C594159822" box="[151,1362,1816,1838]" pageId="10" pageNumber="387">
<taxonomicName id="4C7F4D033E7CFFC377DAF8C590E39821" ID-CoL="5KKCW" box="[151,420,1816,1838]" class="Insecta" family="Eumenidae" genus="Polistes" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="387" phylum="Arthropoda" rank="subSpecies" species="rothneyi" subSpecies="tibetanus">
<emphasis id="B90BEA923E7CFFC377DAF8C590E39821" box="[151,420,1816,1838]" italics="true" pageId="10" pageNumber="387">Polistes rothneyi tibetanus</emphasis>
</taxonomicName>
van der
<treatmentCitation id="0ADE10913E7CFFC376B1F8C593E69821" author="Vecht" box="[508,673,1816,1838]" page="102" pageId="10" pageNumber="387" year="1968">
<bibRefCitation id="EFEE4B713E7CFFC376B1F8C593359821" author="Vecht" box="[508,626,1816,1838]" pageId="10" pageNumber="387" refString="van der Vecht, J. (1968) The geographic variation of Polistes (Megapolistes subg. n.) rothneyi Cameron. Bijdragen Tot de Dierkunde, 38, 97 - 109." type="journal article" year="1968">Vecht 1968</bibRefCitation>
: 102
</treatmentCitation>
(
<typeStatus id="54C488223E7CFFC375E2F8C5924B9821" box="[687,780,1816,1838]" pageId="10" pageNumber="387" type="holotype">holotype</typeStatus>
, ♀, Tibet,
<collectingCountry id="F36876103E7CFFC3743BF8C592F29821" box="[886,949,1816,1838]" name="China" pageId="10" pageNumber="387">China</collectingCountry>
, deposited in RMNH, Leiden),
<emphasis id="B90BEA923E7CFFC373B8F8C594159822" bold="true" box="[1269,1362,1816,1837]" pageId="10" pageNumber="387">syn. nov.</emphasis>
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BC036803E7CFFC377DAF8EB93639864" blockId="10.[151,1436,1478,1930]" pageId="10" pageNumber="387">
<treatmentCitationGroup id="AB6F11AE3E7CFFC377DAF8EB93639864" pageId="10" pageNumber="387">
<taxonomicName id="4C7F4D033E7CFFC377DAF8EB90E49843" ID-CoL="5KKCR" box="[151,419,1846,1868]" class="Insecta" family="Eumenidae" genus="Polistes" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="387" phylum="Arthropoda" rank="subSpecies" species="rothneyi" subSpecies="koreanus">
<emphasis id="B90BEA923E7CFFC377DAF8EB90E49843" box="[151,419,1846,1868]" italics="true" pageId="10" pageNumber="387">Polistes rothneyi koreanus</emphasis>
</taxonomicName>
van der
<treatmentCitation id="0ADE10913E7CFFC376B6F8EB93E59843" author="Vecht" box="[507,674,1846,1868]" page="104" pageId="10" pageNumber="387" year="1968">
<bibRefCitation id="EFEE4B713E7CFFC376B6F8EB93369843" author="Vecht" box="[507,625,1846,1868]" pageId="10" pageNumber="387" refString="van der Vecht, J. (1968) The geographic variation of Polistes (Megapolistes subg. n.) rothneyi Cameron. Bijdragen Tot de Dierkunde, 38, 97 - 109." type="journal article" year="1968">Vecht 1968</bibRefCitation>
: 104
</treatmentCitation>
(
<typeStatus id="54C488223E7CFFC375E2F8EB924B9843" box="[687,780,1846,1868]" pageId="10" pageNumber="387" type="holotype">holotype</typeStatus>
, ♀, Seoul,
<collectingCountry id="F36876103E7CFFC37431F8EA92FB9843" box="[892,956,1847,1868]" name="South Korea" pageId="10" pageNumber="387">Korea</collectingCountry>
, deposited in University of Michigan, Museum of Zoology, Ann Arbor),
<emphasis id="B90BEA923E7CFFC3768BF88B93639864" bold="true" box="[454,548,1878,1899]" pageId="10" pageNumber="387">syn. nov.</emphasis>
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BC036803E7CFFC377DAF8A9943F9886" blockId="10.[151,1436,1478,1930]" box="[151,1400,1908,1930]" pageId="10" pageNumber="387">
<treatmentCitationGroup id="AB6F11AE3E7CFFC377DAF8A9943F9886" box="[151,1400,1908,1930]" pageId="10" pageNumber="387">
<taxonomicName id="4C7F4D033E7CFFC377DAF8A990FD9885" ID-CoL="5KKCV" box="[151,442,1908,1930]" class="Insecta" family="Eumenidae" genus="Polistes" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="387" phylum="Arthropoda" rank="subSpecies" species="rothneyi" subSpecies="sikkimensis">
<emphasis id="B90BEA923E7CFFC377DAF8A990FD9885" box="[151,442,1908,1930]" italics="true" pageId="10" pageNumber="387">Polistes rothneyi sikkimensis</emphasis>
</taxonomicName>
van der
<treatmentCitation id="0ADE10913E7CFFC3755DF8A993F49885" author="Vecht" box="[528,691,1908,1930]" page="102" pageId="10" pageNumber="387" year="1968">
<bibRefCitation id="EFEE4B713E7CFFC3755DF8A993C49885" author="Vecht" box="[528,643,1908,1930]" pageId="10" pageNumber="387" refString="van der Vecht, J. (1968) The geographic variation of Polistes (Megapolistes subg. n.) rothneyi Cameron. Bijdragen Tot de Dierkunde, 38, 97 - 109." type="journal article" year="1968">Vecht 1968</bibRefCitation>
: 102
</treatmentCitation>
(
<typeStatus id="54C488223E7CFFC3758DF8A9925A9885" box="[704,797,1908,1930]" pageId="10" pageNumber="387" type="holotype">holotype</typeStatus>
, ♀, Sikkim,
<collectingCountry id="F36876103E7CFFC374D1F8A992949885" box="[924,979,1908,1930]" name="India" pageId="10" pageNumber="387">India</collectingCountry>
, deposited in BMNH, London),
<emphasis id="B90BEA923E7CFFC37256F8A9943F9886" bold="true" box="[1307,1400,1908,1929]" pageId="10" pageNumber="387">syn. nov.</emphasis>
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C365650B3E7CFFC277DAF86B93B69E13" lastPageId="11" lastPageNumber="388" pageId="10" pageNumber="387" type="materials_examined">
<paragraph id="8BC036803E7CFFC377DAF86B94DB98FB" blockId="10.[151,1436,1974,2036]" pageId="10" pageNumber="387">
<emphasis id="B90BEA923E7CFFC377DAF86B938E98C0" bold="true" box="[151,713,1974,1999]" pageId="10" pageNumber="387">Material examined. Southwest plateau group</emphasis>
(
<figureCitation id="13442A053E7CFFC37592F86B927C98C0" box="[735,827,1974,1999]" captionStart="FIGURE 20" captionStartId="11.[151,250,1485,1507]" captionTargetBox="[302,1284,409,1462]" captionTargetId="figure@11.[302,1284,409,1463]" captionTargetPageId="11" captionText="FIGURE 20. Polistes rothneyi Cameron, 1900, Southwest plateau group (female). A, C, E, G, dorsal view; B, D, F, H, lateral view: A B, f. rothneyi (det. van der Vecht) from Shillong, India; C D, f. tibetanus van der Vecht, 1968 from Tibet, paratype; E F, specimens from Yunnan, showing the color pattern between f. rothneyi and f. tibetanus; G H, specimens from Yunnan, showing the color pattern as? subs. A (van der Vecht, 1968: 103, fig. 7)." httpUri="https://zenodo.org/record/253078/files/figure.png" pageId="10" pageNumber="387">Fig. 20</figureCitation>
): f.
<taxonomicName id="4C7F4D033E7CFFC3743FF86A928998C0" box="[882,974,1975,1999]" class="Insecta" family="Eumenidae" genus="Polistes" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="387" phylum="Arthropoda" rank="species" species="rothneyi">
<emphasis id="B90BEA923E7CFFC3743FF86A928998C0" box="[882,974,1975,1999]" italics="true" pageId="10" pageNumber="387">rothneyi</emphasis>
</taxonomicName>
:
<emphasis id="B90BEA923E7CFFC374A9F86B956A98C0" bold="true" box="[996,1069,1974,1999]" pageId="10" pageNumber="387">
<collectingCountry id="F36876103E7CFFC374A9F86B956F98C0" box="[996,1064,1974,1999]" name="India" pageId="10" pageNumber="387">India</collectingCountry>
:
</emphasis>
♀,
<typeStatus id="54C488223E7CFFC37329F86B958998C0" box="[1124,1230,1974,1999]" pageId="10" pageNumber="387" type="lectotype">lectotype</typeStatus>
, Khasia Hills (
<taxonomicName id="4C7F4D033E7CFFC372CAF86590D798FC" authority="Cameron" authorityName="Cameron" class="Insecta" family="Eumenidae" genus="Polistes" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="387" phylum="Arthropoda" rank="species" species="rufolineatus">
<emphasis id="B90BEA923E7CFFC372CAF865905998FC" italics="true" pageId="10" pageNumber="387">P. rufolineatus</emphasis>
<collectingCountry id="F36876103E7CFFC3766AF80690D798FC" box="[295,400,2011,2035]" name="Cameroon" pageId="10" pageNumber="387">Cameron</collectingCountry>
</taxonomicName>
, BM,
<typeStatus id="54C488223E7CFFC37697F801934A98FB" box="[474,525,2012,2036]" pageId="10" pageNumber="387">type</typeStatus>
, no. 18.895a) Cam. coll. 1901-137; 1♀, Shillong,
<date id="FFC110403E7CFFC3731DF806958B98FB" box="[1104,1228,2011,2036]" pageId="10" pageNumber="387" value="1903-08-06">6.viii.1903</date>
, 5.02 (
<collectionCode id="ED6EAE453E7CFFC37251F806943198FC" LSID="urn:lsid:biocol.org:col:34992" box="[1308,1398,2011,2035]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="10" pageNumber="387">RMNH</collectionCode>
); f.
</paragraph>
<paragraph id="8BC036803E7DFFC277DAFF4593B69E13" blockId="11.[151,1436,151,357]" pageId="11" pageNumber="388">
<emphasis id="B90BEA923E7DFFC277DAFF4591B99FBF" box="[151,254,152,176]" italics="true" pageId="11" pageNumber="388">tibetanus</emphasis>
:
<emphasis id="B90BEA923E7DFFC2765DFF4A90259FBF" bold="true" box="[272,354,151,176]" pageId="11" pageNumber="388">
<collectingCountry id="F36876103E7DFFC2765DFF4A901A9FBF" box="[272,349,151,176]" name="China" pageId="11" pageNumber="388">China</collectingCountry>
:
</emphasis>
1♀,
<typeStatus id="54C488223E7DFFC276EBFF45934E9FBF" box="[422,521,152,176]" pageId="11" pageNumber="388" type="paratype">paratype</typeStatus>
, Tibet, 1954 (
<collectionCode id="ED6EAE453E7DFFC275F4FF4592549FBF" LSID="urn:lsid:biocol.org:col:34992" box="[697,787,152,176]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="11" pageNumber="388">RMNH</collectionCode>
).? f. A: 1♀, Deqing, Yunnan,
<quantity id="4C879B653E7DFFC273D9FF4595A49FBF" box="[1172,1251,151,176]" metricMagnitude="4" metricUnit="m" metricValue="2.8" pageId="11" pageNumber="388" unit="km" value="28.0">28 km</quantity>
SE (28.315˚N, 99.1252˚E),
<date id="FFC110403E7DFFC2766AFF6190E69FDA" box="[295,417,188,213]" pageId="11" pageNumber="388" value="2009-06-21">21.vi.2009</date>
, Blank, Liston &amp; Taeger (
<collectionCode id="ED6EAE453E7DFFC2759BFF61925D9FDB" box="[726,794,188,212]" pageId="11" pageNumber="388">SDEI</collectionCode>
). 2♀, Fugong, Yunnan,
<date id="FFC110403E7DFFC2737FFF6195FC9FDA" box="[1074,1211,188,213]" pageId="11" pageNumber="388" value="2003-08-24">24.viii.2003</date>
, 20046637-8, Q Li (
<collectionCode id="ED6EAE453E7DFFC277D2FF3D91AE9FF7" box="[159,233,224,248]" pageId="11" pageNumber="388">ZJUH</collectionCode>
), 4♀, Ninglangningli, Lijiang, Yunnan, no. 20046553, -6554, -6556, -6557,
<date id="FFC110403E7DFFC27377FF0295109FF7" box="[1082,1111,223,248]" pageId="11" pageNumber="388" value="2003-08-23">23</date>
.viii.2003, Q Li (
<collectionCode id="ED6EAE453E7DFFC27259FF3D94189FF7" box="[1300,1375,224,248]" pageId="11" pageNumber="388">ZJUH</collectionCode>
); 1♀, id., but
<date id="FFC110403E7DFFC277A6FED990329E12" box="[235,373,260,285]" pageId="11" pageNumber="388" value="2004-08-22">22.viii.2004</date>
, no. 20046541, ZS Geng (
<collectionCode id="ED6EAE453E7DFFC275D7FED993A29E13" box="[666,741,260,284]" pageId="11" pageNumber="388">ZJUH</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="C365650B3E7DFFC5778AFEFA928B9BD7" lastPageId="12" lastPageNumber="389" pageId="11" pageNumber="388" type="distribution">
<paragraph id="8BC036803E7DFFC2778AFEFA93029E6A" blockId="11.[151,1436,151,357]" pageId="11" pageNumber="388">
<emphasis id="B90BEA923E7DFFC2778AFEFA90279E4F" bold="true" box="[199,352,295,320]" pageId="11" pageNumber="388">Distribution.</emphasis>
<collectingCountry id="F36876103E7DFFC27622FEFA90F59E4F" box="[367,434,295,320]" name="China" pageId="11" pageNumber="388">China</collectingCountry>
(Yunnan, Tibet);
<collectingCountry id="F36876103E7DFFC275C6FEFA93839E4F" box="[651,708,295,320]" name="India" pageId="11" pageNumber="388">India</collectingCountry>
(Delhi, Uttar Pradesh, West Bengal, Sikkim, Bihar, Assam, Meghalaya);
<collectingCountry id="F36876103E7DFFC27665FE91902A9E6A" box="[296,365,332,357]" name="Nepal" pageId="11" pageNumber="388">Nepal</collectingCountry>
. (
<bibRefCitation id="EFEE4B713E7DFFC27632FE91937A9E6A" author="Carpenter" box="[383,573,332,357]" pageId="11" pageNumber="388" refString="Carpenter, J. M. (1996 b) Distributional Checklist of Species of the Genus Polistes (Hymenoptera: Vespidae; Polistinae, Polistini). American Museum Novitates, 3188, 1 - 39. Available from: http: // iunh 2. sci. ibaraki. ac. jp / wasp / list. html (accessed 21 November 2013)" type="journal article" year="1996" yearSuffix="b">Carpenter 1996b</bibRefCitation>
)
</paragraph>
<caption id="DF0066083E7DFFC277DAFA1092289931" httpUri="https://zenodo.org/record/253078/files/figure.png" pageId="11" pageNumber="388" targetBox="[302,1284,409,1462]" targetPageId="11">
<paragraph id="8BC036803E7DFFC277DAFA1092289931" blockId="11.[151,1436,1484,1598]" pageId="11" pageNumber="388">
<emphasis id="B90BEA923E7DFFC277DAFA1090679AED" bold="true" box="[151,288,1485,1507]" pageId="11" pageNumber="388">FIGURE 20.</emphasis>
<taxonomicName id="4C7F4D033E7DFFC2766AFA11933F9AED" authority="Cameron, 1900" authorityName="Cameron" authorityYear="1900" box="[295,632,1484,1506]" class="Insecta" family="Eumenidae" genus="Polistes" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="388" phylum="Arthropoda" rank="species" species="rothneyi">
<emphasis id="B90BEA923E7DFFC2766AFA1190969AED" box="[295,465,1484,1506]" italics="true" pageId="11" pageNumber="388">Polistes rothneyi</emphasis>
Cameron, 1900
</taxonomicName>
, Southwest plateau group (female). A, C, E, G, dorsal view; B, D, F, H, lateral view:
<emphasis id="B90BEA923E7DFFC2779BFA36904B990F" bold="true" box="[214,268,1515,1536]" pageId="11" pageNumber="388">AB,</emphasis>
f.
<taxonomicName id="4C7F4D033E7DFFC27667FA36903A990E" box="[298,381,1515,1537]" class="Insecta" family="Eumenidae" genus="Polistes" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="388" phylum="Arthropoda" rank="species" species="rothneyi">
<emphasis id="B90BEA923E7DFFC27667FA36903A990E" box="[298,381,1515,1537]" italics="true" pageId="11" pageNumber="388">rothneyi</emphasis>
</taxonomicName>
(det. van der Vecht) from Shillong, India;
<emphasis id="B90BEA923E7DFFC27478FA36922A990E" bold="true" box="[821,877,1515,1537]" pageId="11" pageNumber="388">CD,</emphasis>
f.
<emphasis id="B90BEA923E7DFFC274C7FA3692A0990E" box="[906,999,1515,1537]" italics="true" pageId="11" pageNumber="388">tibetanus</emphasis>
van der Vecht, 1968 from Tibet, paratype;
<emphasis id="B90BEA923E7DFFC277DAF9D7918F9910" bold="true" box="[151,200,1546,1567]" pageId="11" pageNumber="388">EF,</emphasis>
specimens from Yunnan, showing the color pattern between f.
<taxonomicName id="4C7F4D033E7DFFC27419F9D792E0992F" box="[852,935,1546,1568]" class="Insecta" family="Eumenidae" genus="Polistes" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="388" phylum="Arthropoda" rank="species" species="rothneyi">
<emphasis id="B90BEA923E7DFFC27419F9D792E0992F" box="[852,935,1546,1568]" italics="true" pageId="11" pageNumber="388">rothneyi</emphasis>
</taxonomicName>
and f.
<emphasis id="B90BEA923E7DFFC274BCF9D79509992F" box="[1009,1102,1546,1568]" italics="true" pageId="11" pageNumber="388">tibetanus</emphasis>
;
<emphasis id="B90BEA923E7DFFC27311F9D795D0992F" bold="true" box="[1116,1175,1546,1568]" pageId="11" pageNumber="388">GH,</emphasis>
specimens from Yunnan, showing the color pattern as?subs. A (van der Vecht, 1968: 103, fig. 7).
</paragraph>
</caption>
<paragraph id="8BC036803E7DFFC5778AF9B6953F9CB2" blockId="11.[151,1436,1643,2022]" lastBlockId="12.[151,1437,151,2021]" lastPageId="12" lastPageNumber="389" pageId="11" pageNumber="388">
<emphasis id="B90BEA923E7DFFC2778AF9B69360998B" bold="true" box="[199,551,1643,1668]" pageId="11" pageNumber="388">Common yellow-black group</emphasis>
(
<figureCitation id="13442A053E7DFFC27576F9B693C8998B" box="[571,655,1643,1668]" captionStart="FIGURE 21" captionStartId="13.[151,250,1776,1798]" captionTargetBox="[302,1284,468,1753]" captionTargetId="figure@13.[302,1284,468,1754]" captionTargetPageId="13" captionText="FIGURE 21. Polistes rothneyi Cameron, 1900, Common yellow-black group (female). A, C, E, G, I, dorsal view; B, D, F, H, J, lateral view: A B, f. grahami van der Vecht, 1968 from Sichuan, paratype (showing mesonotum with two yellow stripes); C D, f. grahami van der Vecht, 1968 from Fujian, paratype (showing mesonotum with four yellow stripes); E F, f. iwatai van der Vecht, 1968 from Japan, paratype (showing color pattern very similar as A-B f. grahami, but pronotum mostly yellow); G H, specimens from Hainan with the color pattern same as f. hainanensis van der Vecht, 1968, showing the apical band of the first metasomal tergite entirely yellow; I J, f. gressitti van der Vecht, 1968 from Taiwan, paratype (showing pronotum entirely yellow); K L, f. yayeyamae Matsumura, 1908 from Iriomote Loochoo Is., paratype (showing body mainly yellow)." httpUri="https://zenodo.org/record/253079/files/figure.png" pageId="11" pageNumber="388">Fig. 21</figureCitation>
): f.
<emphasis id="B90BEA923E7DFFC27589F9B6920E998C" box="[708,841,1643,1667]" italics="true" pageId="11" pageNumber="388">hainanensis</emphasis>
(Hainan):
<emphasis id="B90BEA923E7DFFC27485F9B6955C998B" bold="true" box="[968,1051,1643,1668]" pageId="11" pageNumber="388">
<collectingCountry id="F36876103E7DFFC27485F9B69551998B" box="[968,1046,1643,1668]" name="China" pageId="11" pageNumber="388">China</collectingCountry>
:
</emphasis>
♀,
<typeStatus id="54C488223E7DFFC27306F9B195E9998B" box="[1099,1198,1644,1668]" pageId="11" pageNumber="388" type="paratype">paratype</typeStatus>
, Nodoa, Hainan,
<date id="FFC110403E7DFFC27234F9B691BA99A8" pageId="11" pageNumber="388" value="1935-06" valueMax="1935-08-31" valueMin="1935-06-00">vi viii.1935</date>
, L. Gressitt (RMNH). 1♀, Baisha (19˚13'N, 109˚26'E), Hainan,
<quantity id="4C879B653E7DFFC27480F952955199A8" box="[973,1046,1679,1703]" metricMagnitude="2" metricUnit="m" metricValue="2.02" pageId="11" pageNumber="388" unit="m" value="202.0">202 m</quantity>
,
<date id="FFC110403E7DFFC2736FF95395E599A8" box="[1058,1186,1678,1703]" pageId="11" pageNumber="388" value="2011-07-21">21.vii.2011</date>
, no. hzs0700439, YH Zhong (CATAS); 1♀, Nanluo Village (19˚47'N, 109˚63'E), Nanyang, Hainan,
<quantity id="4C879B653E7DFFC27352F96E952A99C5" box="[1055,1133,1715,1739]" metricMagnitude="2" metricUnit="m" metricValue="2.02" pageId="11" pageNumber="388" unit="m" value="202.0">202 m</quantity>
,
<date id="FFC110403E7DFFC27335F96F95AE99C4" box="[1144,1257,1714,1739]" pageId="11" pageNumber="388" value="2011-07-06">6.vii.2011</date>
, no. 0 7008, WJ Zhu (CATAS); 2♀, Jianfenling Nature Reserve (18˚44'38”N 108˚50'36”E), Hainan,
<quantity id="4C879B653E7DFFC27377F908958F99E1" box="[1082,1224,1749,1774]" metricMagnitude="3" metricUnit="m" metricValue="1.065" metricValueMax="1.15" metricValueMin="0.98" pageId="11" pageNumber="388" unit="m" value="1065.0" valueMax="1150.0" valueMin="980.0">9801150 m</quantity>
,
<date id="FFC110403E7DFFC27398F908943999E1" box="[1237,1406,1749,1774]" pageId="11" pageNumber="388" value="2013-07-13" valueMax="2013-07-14" valueMin="2013-07-13">1314.vii.2013</date>
, C van Achterberg (RMNH). f.
<emphasis id="B90BEA923E7DFFC2769BF9249373981E" box="[470,564,1785,1809]" italics="true" pageId="11" pageNumber="388">grahami</emphasis>
:
<emphasis id="B90BEA923E7DFFC27508F92493DF981D" bold="true" box="[581,664,1785,1810]" pageId="11" pageNumber="388">
<collectingCountry id="F36876103E7DFFC27508F92493D5981D" box="[581,658,1785,1810]" name="China" pageId="11" pageNumber="388">China</collectingCountry>
:
</emphasis>
♂,
<typeStatus id="54C488223E7DFFC27588F927926F981D" box="[709,808,1786,1810]" pageId="11" pageNumber="388" type="paratype">paratype</typeStatus>
, no.663, Peiping, Hepei,
<collectingCountry id="F36876103E7DFFC2730BF92495CC981D" box="[1094,1163,1785,1810]" name="China" pageId="11" pageNumber="388">China</collectingCountry>
, 193, Liu (RMNH); 9♀,
<typeStatus id="54C488223E7DFFC277DAF8C09043983A" box="[151,260,1821,1845]" pageId="11" pageNumber="388" type="paratype">paratypes</typeStatus>
, Kuatun, Fukien,
<date id="FFC110403E7DFFC27680F8C19362983A" box="[461,549,1820,1845]" pageId="11" pageNumber="388" value="1946-03">iii.1946</date>
, Tschung, Sen (RMNH); 1♀,
<specimenCount id="9D79FD093E7DFFC27439F8C092DE983A" box="[884,921,1821,1845]" pageId="11" pageNumber="388" type="generic">1♂</specimenCount>
, Museum Paris, Kouy-chéou, Rég. de Pin-fa, Père Cavalerie, 1909 (MNHN); 1♀, Museum Paris, Kouyyang, Kouy-Tchéou, PP Cavalerie et Fortunat, 1906 (MNHN); 2♀, Tsé-Kou, Yun-Nan (Mgr Soulie), R Oberthür, 1898 (MNHN); 1♀, Rainforestry Path, Mt. Wuzhi, Hainan,
<collectingCountry id="F36876103E7DFFC277B7F85A907898AF" box="[250,319,1927,1952]" name="China" pageId="11" pageNumber="388">China</collectingCountry>
,
<date id="FFC110403E7DFFC27603F85A90E898AF" box="[334,431,1927,1952]" pageId="11" pageNumber="388" value="2010-05-06">6.v.2010</date>
, JL Tan (NWUM); 2♀,
<specimenCount id="9D79FD093E7DFFC27584F85A93A898AF" box="[713,751,1927,1952]" pageId="11" pageNumber="388" type="generic">5♂</specimenCount>
, Quannan, Jiangxi,
<date id="FFC110403E7DFFC2749DF85A954498AF" box="[976,1027,1927,1952]" pageId="11" pageNumber="388">1.xii</date>
or
<date id="FFC110403E7DFFC27360F85A95E098AF" box="[1069,1191,1927,1952]" pageId="11" pageNumber="388" value="2010-11-27">27.xi.2010</date>
, SC Li (GSFA);
<specimenCount id="9D79FD093E7DFFC2723FF85A94D19890" box="[1394,1430,1927,1951]" pageId="11" pageNumber="388" type="generic">1♂</specimenCount>
, Yushan, Jiangxi,
<quantity id="4C879B653E7DFFC27616F87690F798CD" box="[347,432,1963,1987]" metricMagnitude="3" metricUnit="m" metricValue="1.1199999999999999" pageId="11" pageNumber="388" unit="m" value="1120.0">1120 m</quantity>
,
<date id="FFC110403E7DFFC276F2F877930598CC" box="[447,578,1962,1987]" pageId="11" pageNumber="388" value="1985-07-20">20.vii.1985</date>
; 1♀, Mt. Tianzhu, Qianshan, Anhui,
<date id="FFC110403E7DFFC274BCF87795EB98CC" box="[1009,1196,1962,1987]" pageId="11" pageNumber="388" value="2011-03-10" valueMax="2011-04-11" valueMin="2011-03-10">10.iii11.iv.2011</date>
, ML Sheng (GSFA); 1♀, Mt. Gutian, Kaihua, Zhejiang,
<date id="FFC110403E7DFFC27506F810939C98E9" box="[587,731,1997,2022]" pageId="11" pageNumber="388" value="2005-07-01" valueMax="2005-07-03" valueMin="2005-07-01">13.vii.2005</date>
, 200604253, XX Chen (ZJUH); 1♀, Nanyuechansi Temple, Yixing, Jiangsu,
<collectingCountry id="F36876103E7AFFC57613FF4A90E49FBF" box="[350,419,151,176]" name="China" pageId="12" pageNumber="389">China</collectingCountry>
,
<date id="FFC110403E7AFFC576FBFF4A93759FBF" box="[438,562,151,176]" pageId="12" pageNumber="389" value="2010-08-02">2.viii.2010</date>
, JL Tan (ZJUH); 1♀, West Mt. Tianmu, Lin'an, Zhejiang,
<date id="FFC110403E7AFFC57269FF4A94D19FBF" box="[1316,1430,151,176]" pageId="12" pageNumber="389" value="2000-07-07">7.vii.2000</date>
, 200103816, XX Chen (ZJUH);
<specimenCount id="9D79FD093E7AFFC57563FF6193159FDB" box="[558,594,188,212]" pageId="12" pageNumber="389" type="generic">1♂</specimenCount>
, Suichang, Zhejiang,
<date id="FFC110403E7AFFC57429FF6195479FDB" box="[868,1024,188,212]" pageId="12" pageNumber="389" value="1973-10-18" valueMax="1973-10-19" valueMin="1973-10-18">1819.x.1973</date>
, 73057.4, JH He (ZJUH); 1♀, S.
<collectingCountry id="F36876103E7AFFC577DAFF0291B49FF8" box="[151,243,223,247]" name="China" pageId="12" pageNumber="389">CHINA</collectingCountry>
: NE of Guangzhou, Nankun Mt. Tree Res. (23˚40'N, 113˚56'E) Guangdong,
<date id="FFC110403E7AFFC573CFFF0295B79FF8" box="[1154,1264,223,247]" pageId="12" pageNumber="389" value="2012-05-24">24.v.2012</date>
, c.
<quantity id="4C879B653E7AFFC57257FF0294229FF8" box="[1306,1381,223,247]" metricMagnitude="2" metricUnit="m" metricValue="3.4" pageId="12" pageNumber="389" unit="m" value="340.0">340 m</quantity>
, C v Achterberg (RMNH); 1♀, Dongxianfeng, Dongyang, Zhejiang,
<date id="FFC110403E7AFFC5742FFEDF92A39E14" box="[866,996,258,283]" pageId="12" pageNumber="389" value="2010-07-28">28.vii.2010</date>
, JL Tan &amp; XX Chen (ZJUH); 1♀, Mt. Wuyun, Hangzhou, Zhejiang,
<date id="FFC110403E7AFFC576B5FEF8933D9E31" box="[504,634,293,318]" pageId="12" pageNumber="389" value="2010-07-25">25.vii.2010</date>
, JL Tan (ZJUH); 1♀, Xiangshuwan, Kuankuoshui, Suiyang, Guizhou,
<date id="FFC110403E7AFFC577DAFE9490449E6D" box="[151,259,329,354]" pageId="12" pageNumber="389" value="2010-06-05">5.vi.2010</date>
, JL Tan (ZJUH); 1♀, Mt. Yuhuang, Hangzhou, Zhejiang,
<date id="FFC110403E7AFFC574DAFE94955E9E6D" box="[919,1049,329,354]" pageId="12" pageNumber="389" value="2010-07-24">24.vii.2010</date>
, JL Tan (ZJUH); 1♀, Mt. Tianmu, Linan, Zhejiang,
<date id="FFC110403E7AFFC57629FEB1909F9E8A" box="[356,472,364,389]" pageId="12" pageNumber="389" value="2010-07-03">3.vii.2010</date>
, JL. Tan (ZJUH); 2♀, Shunxi, Qingliangfeng, Linan, Zhejiang,
<date id="FFC110403E7AFFC573F4FEB194049E8A" box="[1209,1347,364,389]" pageId="12" pageNumber="389" value="2013-08-23">23.viii.2013</date>
, JL Tan (NWUM); 4♀, Mt. Nankun, Guangdong,
<date id="FFC110403E7AFFC57534FE4D93A29EA6" box="[633,741,400,425]" pageId="12" pageNumber="389" value="2002-06-08">8.vi.2002</date>
, no. 20028901, -8904, -8906, -8787, ZF Xu (ZJUH); 6 ♀, Mt. Tai, Taian, Shandong,
<date id="FFC110403E7AFFC57690FE6E93109EC3" box="[477,599,435,460]" pageId="12" pageNumber="389" value="1997-06-23">23.vi.1997</date>
, no. 200011329, -325, -336, -320, -311, -245, Q. Li (ZJUH); 1♀, Taiyi Palace, Changan, Xian, Shaanxi,
<date id="FFC110403E7AFFC57558FE0A93CD9EFF" box="[533,650,471,496]" pageId="12" pageNumber="389" value="2011-04-25">25.iv.2011</date>
, GH Wang (NWUM); 1♀, Fengyukou, Xian, Shaanxi (NWUM); 2♀, Guanghuojie, Ningshan, Shaanxi,
<date id="FFC110403E7AFFC57560FE2793E59D1C" box="[557,674,506,531]" pageId="12" pageNumber="389" value="2013-07-02">2.vii.2013</date>
, JL Tan (NWUM); 14♀, Foping (33˚30'N, 107˚58'E), Shaanxi,
<date id="FFC110403E7AFFC577DAFDC0905E9D39" box="[151,281,541,566]" pageId="12" pageNumber="389" value="2013-07-31">31.vii.2013</date>
, JL Tan (NWUM);
<specimenCount id="9D79FD093E7AFFC57541FDC393779D39" box="[524,560,542,566]" pageId="12" pageNumber="389" type="generic">1♂</specimenCount>
, Zaoyuan, Yanan, Shaanxi,
<date id="FFC110403E7AFFC574C4FDC392BC9D39" box="[905,1019,542,566]" pageId="12" pageNumber="389" value="2010-10-17">17.x.2010</date>
, JL Tan (NWUM); 1♀,
<specimenCount id="9D79FD093E7AFFC5726EFDC3940F9D39" box="[1315,1352,542,566]" pageId="12" pageNumber="389" type="generic">1♂</specimenCount>
, NWU Taoyuan Campus, Xian, Shaanxi,
<date id="FFC110403E7AFFC5757CFD9C93D99D55" box="[561,670,577,602]" pageId="12" pageNumber="389" value="2010-09-04">4.ix.2010</date>
, T Zhou (NWUM); 1♀, Yichuan, Shaanxi,
<date id="FFC110403E7AFFC573E9FD9C945F9D55" box="[1188,1304,577,602]" pageId="12" pageNumber="389" value="1980-07-09">9.vii.1980</date>
, YZ Zhang (NWAM);
<specimenCount id="9D79FD093E7AFFC5765FFDB890719D72" box="[274,310,613,637]" pageId="12" pageNumber="389" type="generic">1♂</specimenCount>
, Liping National Forestry Park, Hanzhong, Shaanxi,
<date id="FFC110403E7AFFC574EAFDB895509D71" box="[935,1047,613,638]" pageId="12" pageNumber="389" value="2011-10-15">15.x.2011</date>
, JL Tan (NWUM); 1♀, Mt. Hua, Shaanxi,
<date id="FFC110403E7AFFC5764CFD54900A9DAE" box="[257,333,649,673]" pageId="12" pageNumber="389" value="1984-05">v.1984</date>
, GQ Han (NWAM); 5♀,
<specimenCount id="9D79FD093E7AFFC5753EFD5493D09DAE" box="[627,663,649,673]" pageId="12" pageNumber="389" type="generic">1♂</specimenCount>
, Liyuanping, Mt. Qinling, Shaanxi,
<date id="FFC110403E7AFFC57376FD5595859DAE" box="[1083,1218,648,673]" pageId="12" pageNumber="389" value="2011-08-18">18.viii.2011</date>
, JL Tan (NWUM); 17♀,
<specimenCount id="9D79FD093E7AFFC57795FD7191BA9DCA" box="[216,253,684,709]" pageId="12" pageNumber="389" type="generic">5♂</specimenCount>
, Jinsixia (33˚22'N, 110˚33'E), Shangnan, Shaanxi,
<quantity id="4C879B653E7AFFC57405FD7192D39DCB" box="[840,916,684,708]" metricMagnitude="2" metricUnit="m" metricValue="7.08" pageId="12" pageNumber="389" unit="m" value="708.0">708 m</quantity>
,
<date id="FFC110403E7AFFC574EEFD7195629DCA" box="[931,1061,684,709]" pageId="12" pageNumber="389" value="2013-07-24">24.vii.2013</date>
, JL Tan (NWUM); 2♀, Huishui, Guizhou,
<date id="FFC110403E7AFFC57648FD1290399DE7" box="[261,382,719,744]" pageId="12" pageNumber="389" value="2009-06-25">25.vi.2009</date>
,
<date id="FFC110403E7AFFC576C7FD1290B99DE7" box="[394,510,719,744]" pageId="12" pageNumber="389" value="2009-07-09">9.vii.2009</date>
, ZD Li (NWUM); 1♀, Qianyang, Guizhou,
<date id="FFC110403E7AFFC574A7FD1295249DE7" box="[1002,1123,719,744]" pageId="12" pageNumber="389" value="2009-06-18">18.vi.2009</date>
, PQ Shi (NWUM); 2♀, Mt. Dinghu, Guangdong,
<date id="FFC110403E7AFFC576C1FD2E934A9C03" box="[396,525,755,780]" pageId="12" pageNumber="389" value="1985-07-17">17.vii.1985</date>
, YL Zhang (NWAM); 2♀, Binzhou, Hunan, YL Zhang, YH Chai (NWAM); 1♀, Yuelu Mt. Park (28˚12'N 112˚56'E), Changsha, Hunan, c.
<quantity id="4C879B653E7AFFC5745FFCCA921D9C21" box="[786,858,791,815]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="12" pageNumber="389" unit="m" value="300.0">300 m</quantity>
,
<date id="FFC110403E7AFFC57428FCCB929A9C20" box="[869,989,790,815]" pageId="12" pageNumber="389" value="2012-06-14">14.vi.2012</date>
, C van Achterberg (RMNH). f.
<emphasis id="B90BEA923E7AFFC57271FCCA94D39C21" box="[1340,1428,791,814]" italics="true" pageId="12" pageNumber="389">gressitti</emphasis>
:
<emphasis id="B90BEA923E7AFFC577DAFCE491AD9C5D" bold="true" box="[151,234,825,850]" pageId="12" pageNumber="389">
<collectingCountry id="F36876103E7AFFC577DAFCE491A29C5D" box="[151,229,825,850]" name="China" pageId="12" pageNumber="389">China</collectingCountry>
:
</emphasis>
1♀,
<typeStatus id="54C488223E7AFFC5766FFCE490C19C5D" box="[290,390,825,850]" pageId="12" pageNumber="389" type="holotype">holotype</typeStatus>
, Taihanroku,
<collectingCountry id="F36876103E7AFFC57550FCE793C59C5E" box="[541,642,826,849]" name="Taiwan" pageId="12" pageNumber="389">Formosa</collectingCountry>
,
<date id="FFC110403E7AFFC575C0FCE492449C5D" box="[653,771,825,850]" pageId="12" pageNumber="389" value="1908-04-26">26.iv.1908</date>
, H. Sauter (RMNH); f.
<taxonomicName id="4C7F4D033E7AFFC57341FCE795CF9C5E" box="[1036,1160,826,849]" class="Insecta" family="Eumenidae" genus="Polistes" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="389" phylum="Arthropoda" rank="species" species="yayeyamae">
<emphasis id="B90BEA923E7AFFC57341FCE795CF9C5E" box="[1036,1160,826,849]" italics="true" pageId="12" pageNumber="389">yayeyamae</emphasis>
</taxonomicName>
(possibly distributed in Ta i w a n)
<emphasis id="B90BEA923E7AFFC577B5FC8090079C79" bold="true" box="[248,320,861,886]" pageId="12" pageNumber="389">
<collectingCountry id="F36876103E7AFFC577B5FC8090079C79" box="[248,320,861,886]" name="Japan" pageId="12" pageNumber="389">Japan</collectingCountry>
</emphasis>
: ♀,
<typeStatus id="54C488223E7AFFC57636FC8390999C79" box="[379,478,862,886]" pageId="12" pageNumber="389" type="paratype">paratype</typeStatus>
, Iriomote Loochoo Island, Southern Ryukyu Is,
<date id="FFC110403E7AFFC57352FC8095ED9C79" box="[1055,1194,861,886]" pageId="12" pageNumber="389" value="1934-08-22">22.viii.1934</date>
, L Gressitt (RMNH).
<emphasis id="B90BEA923E7AFFC577DAFC5D90779C96" bold="true" box="[151,304,896,921]" pageId="12" pageNumber="389">Distribution.</emphasis>
<collectingCountry id="F36876103E7AFFC57676FC5D90399C96" box="[315,382,896,921]" name="China" pageId="12" pageNumber="389">China</collectingCountry>
(Heilongjiang, Beijing,
<collectingCountry id="F36876103E7AFFC575D4FC5D93AC9C96" box="[665,747,896,921]" name="China" pageId="12" pageNumber="389">Tianjin</collectingCountry>
, Shandong, Sichuan, Chongqing, Anhui, Jiangsu, Zhejiang, Guangdong, Jiangxi, Fujian, Guizhou, Shaanxi, Yunnan, Hainan, Hebei, Hunan,
<collectingCountry id="F36876103E7AFFC5735AFC79952B9CB2" box="[1047,1132,932,957]" name="Taiwan" pageId="12" pageNumber="389">Taiwan</collectingCountry>
).
</paragraph>
<paragraph id="8BC036803E7AFFC5778AFC1A928B9BD7" blockId="12.[151,1437,151,2021]" pageId="12" pageNumber="389">
<emphasis id="B90BEA923E7AFFC5778AFC1A90CD9CEF" bold="true" box="[199,394,967,992]" pageId="12" pageNumber="389">Northeast group</emphasis>
(
<figureCitation id="13442A053E7AFFC576D4FC1A90AE9CEF" box="[409,489,967,992]" captionStart="FIGURE 22" captionStartId="14.[151,250,1341,1363]" captionTargetBox="[302,1284,193,1319]" captionTargetId="figure@14.[302,1284,193,1319]" captionTargetPageId="14" captionText="FIGURE 22. Polistes rothneyi Cameron, 1900, Northeast group (female) (showing the colour pattern variation). A, C, E, G, I, J, dorsal view; B, D, F, H, lateral view: A B, specimen from Shenyang, Liaoning; C D, specimen from Shenyang, Kuandian; E F, specimen from Qianshan, Liaoning; G H, f. koreanus van der Vecht, 1968 from Seoul area, paratype (with pronotum black except narrow yellow margin, first metasomal tergite black with a narrow apical band); I, specimen from Siping, Jilin; J, specimen from Xinbin, Liaoning." httpUri="https://zenodo.org/record/253080/files/figure.png" pageId="12" pageNumber="389">Fig. 22</figureCitation>
):
<emphasis id="B90BEA923E7AFFC5754FFC1A93139CEF" bold="true" box="[514,596,967,992]" pageId="12" pageNumber="389">
<collectingCountry id="F36876103E7AFFC5754FFC1A93089CEF" box="[514,591,967,992]" name="China" pageId="12" pageNumber="389">China</collectingCountry>
:
</emphasis>
1♀, Lingxi (west tomb), Shenyang,
<date id="FFC110403E7AFFC574A1FC1A95329CEF" box="[1004,1141,967,992]" pageId="12" pageNumber="389" value="2009-08-27">27.viii.2009</date>
, S. Li (GSFA);
<specimenCount id="9D79FD093E7AFFC5726CFC1594029CEF" box="[1313,1349,968,992]" pageId="12" pageNumber="389" type="generic">1♂</specimenCount>
, Lingxi (west tomb), Shenyang,
<date id="FFC110403E7AFFC576E6FC3693739B0B" box="[427,564,1003,1028]" pageId="12" pageNumber="389" value="2008-08-15">15.viii.2008</date>
, S. Li (GSFA); 2♀, Qianshan, Shenyang, Liaoning,
<date id="FFC110403E7AFFC573C8FC3694499B0B" box="[1157,1294,1003,1028]" pageId="12" pageNumber="389" value="2006-08-27">27.viii.2006</date>
, YM Li, YT Zhang; 1♀,
<specimenCount id="9D79FD093E7AFFC5766FFBD290009B28" box="[290,327,1039,1063]" pageId="12" pageNumber="389" type="generic">2♂</specimenCount>
, Xinbin, Liaoning,
<date id="FFC110403E7AFFC5756AFBD393D49B28" box="[551,659,1038,1063]" pageId="12" pageNumber="389" value="2009-06-01">1.vi.2009</date>
, ML Sheng (GSFA); 1♀, Shenyang, Liaoning, 1991; 1♀, Kuandian, Liaoning,
<date id="FFC110403E7AFFC57641FBEF90309B44" box="[268,375,1074,1099]" pageId="12" pageNumber="389" value="2007-06-07">7.vi.2007</date>
, SP Sun (GSFA); 1♀, Shenyang, Liaoning,
<date id="FFC110403E7AFFC5743CFBEF92BC9B44" box="[881,1019,1074,1099]" pageId="12" pageNumber="389" value="2010-08-17">17.viii.2010</date>
, L Jiang (NWUM); 2♀,
<specimenCount id="9D79FD093E7AFFC57254FBEF94799B45" box="[1305,1342,1074,1098]" pageId="12" pageNumber="389" type="generic">3♂</specimenCount>
, Siping, Jilin,
<date id="FFC110403E7AFFC57796FB8890759B61" box="[219,306,1109,1134]" pageId="12" pageNumber="389" value="1999-06">vi.1999</date>
, BZ Ren; 1♀, North Tomb, Shenyang, Liaoning,
<date id="FFC110403E7AFFC574C8FB8B92B09B61" box="[901,1015,1110,1134]" pageId="12" pageNumber="389" value="2010-10-18">18.x.2010</date>
, XB Nie; 1♀, Shenyang, Liaoning,
<date id="FFC110403E7AFFC577DAFBA491BD9B9E" box="[151,250,1145,1169]" pageId="12" pageNumber="389" value="2010-10-08">8.x.2010</date>
, SP Sun. f.
<emphasis id="B90BEA923E7AFFC576DDFBA490B39B9E" box="[400,500,1145,1169]" italics="true" pageId="12" pageNumber="389">koreanus</emphasis>
:
<emphasis id="B90BEA923E7AFFC57544FBA4931A9B9D" bold="true" box="[521,605,1145,1170]" pageId="12" pageNumber="389">
<collectingCountry id="F36876103E7AFFC57544FBA4931F9B9D" box="[521,600,1145,1170]" name="South Korea" pageId="12" pageNumber="389">Korea</collectingCountry>
:
</emphasis>
♀,
<typeStatus id="54C488223E7AFFC575DFFBA793B29B9D" box="[658,757,1146,1170]" pageId="12" pageNumber="389" type="paratype">paratype</typeStatus>
, Seoul area,
<date id="FFC110403E7AFFC574DEFBA4954E9B9D" box="[915,1033,1145,1170]" pageId="12" pageNumber="389" value="1954-04-25">25.iv.1954</date>
, GW Byers. f.
<emphasis id="B90BEA923E7AFFC5738BFBA7944C9B9E" box="[1222,1291,1146,1169]" italics="true" pageId="12" pageNumber="389">iwatai</emphasis>
:
<emphasis id="B90BEA923E7AFFC5726DFBA494349B9D" bold="true" box="[1312,1395,1145,1170]" pageId="12" pageNumber="389">
<collectingCountry id="F36876103E7AFFC5726DFBA494299B9D" box="[1312,1390,1145,1170]" name="Japan" pageId="12" pageNumber="389">Japan</collectingCountry>
:
</emphasis>
♀,
<typeStatus id="54C488223E7AFFC577DAFB4091BD9BBA" box="[151,250,1181,1205]" pageId="12" pageNumber="389" type="paratype">paratype</typeStatus>
, Sasayama, Honshu,
<date id="FFC110403E7AFFC576BDFB4193199BBA" box="[496,606,1180,1205]" pageId="12" pageNumber="389" value="1953-05-16">16.v.1953</date>
, M Miki from Prof. Iwata;
<specimenCount id="9D79FD093E7AFFC574EDFB4092829BBA" box="[928,965,1181,1205]" pageId="12" pageNumber="389" type="generic">2♂</specimenCount>
,
<collectingCountry id="F36876103E7AFFC57499FB4095519BBB" box="[980,1046,1181,1204]" name="Japan" pageId="12" pageNumber="389">Japan</collectingCountry>
, 1982, 1983.
<emphasis id="B90BEA923E7AFFC573FBFB4194089BBA" bold="true" box="[1206,1359,1180,1205]" pageId="12" pageNumber="389">Distribution.</emphasis>
<collectingCountry id="F36876103E7AFFC57214FB4194DC9BBA" box="[1369,1435,1180,1205]" name="China" pageId="12" pageNumber="389">China</collectingCountry>
(Shandong (Weihai), Liaoning, Jilin);
<collectingCountry id="F36876103E7AFFC57573FB1C93C69BD7" box="[574,641,1217,1240]" name="Japan" pageId="12" pageNumber="389">Japan</collectingCountry>
;
<collectingCountry id="F36876103E7AFFC575C1FB1C93939BD7" box="[652,724,1217,1240]" name="South Korea" pageId="12" pageNumber="389">Korea</collectingCountry>
. (van der
<bibRefCitation id="EFEE4B713E7AFFC5740CFB6292F89BD7" author="Vecht" box="[833,959,1215,1240]" pageId="12" pageNumber="389" refString="van der Vecht, J. (1968) The geographic variation of Polistes (Megapolistes subg. n.) rothneyi Cameron. Bijdragen Tot de Dierkunde, 38, 97 - 109." type="journal article" year="1968">Vecht 1968</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C365650B3E7AFFC4778AFB3E928D9EA7" lastPageId="13" lastPageNumber="390" pageId="12" pageNumber="389" type="discussion">
<paragraph id="8BC036803E7AFFC4778AFB3E928D9EA7" blockId="12.[151,1437,151,2021]" lastBlockId="13.[151,1436,151,425]" lastPageId="13" lastPageNumber="390" pageId="12" pageNumber="389">
<emphasis id="B90BEA923E7AFFC5778AFB3E907D9BF3" bold="true" box="[199,314,1251,1276]" pageId="12" pageNumber="389">Remarks.</emphasis>
<taxonomicName id="4C7F4D033E7AFFC5760BFB3990F99BF4" box="[326,446,1251,1275]" class="Insecta" family="Eumenidae" genus="Polistes" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="389" phylum="Arthropoda" rank="species" species="rothneyi">
<emphasis id="B90BEA923E7AFFC5760BFB3990F99BF4" box="[326,446,1251,1275]" italics="true" pageId="12" pageNumber="389">P. rothneyi</emphasis>
</taxonomicName>
is easily confused with the closely allied species
<taxonomicName id="4C7F4D033E7AFFC5735CFB3995DD9BF4" box="[1041,1178,1251,1275]" class="Insecta" family="Eumenidae" genus="Polistes" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="389" phylum="Arthropoda" rank="species" species="olivaceus">
<emphasis id="B90BEA923E7AFFC5735CFB3995DD9BF4" box="[1041,1178,1251,1275]" italics="true" pageId="12" pageNumber="389">P. olivaceus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C7F4D033E7AFFC573E1FB39940F9BF4" box="[1196,1352,1251,1275]" class="Insecta" family="Eumenidae" genus="Polistes" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="389" phylum="Arthropoda" rank="species" species="jokahamae">
<emphasis id="B90BEA923E7AFFC573E1FB39940F9BF4" box="[1196,1352,1251,1275]" italics="true" pageId="12" pageNumber="389">P. jokahamae</emphasis>
</taxonomicName>
and
<taxonomicName id="4C7F4D033E7AFFC572CAFB39906A9A10" class="Insecta" family="Eumenidae" genus="Polistes" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="389" phylum="Arthropoda" rank="species" species="tenuispunctia">
<emphasis id="B90BEA923E7AFFC572CAFB39906A9A10" italics="true" pageId="12" pageNumber="389">P. tenuispunctia</emphasis>
</taxonomicName>
. It is a polytypic species with a very interesting variable colour pattern. Van der
<bibRefCitation id="EFEE4B713E7AFFC573F7FADB940F9A10" author="Vecht" box="[1210,1352,1286,1311]" pageId="12" pageNumber="389" refString="van der Vecht, J. (1968) The geographic variation of Polistes (Megapolistes subg. n.) rothneyi Cameron. Bijdragen Tot de Dierkunde, 38, 97 - 109." type="journal article" year="1968">Vecht (1968)</bibRefCitation>
treated them as 16 subspecies and two uncertain? subspecies A and B. Das &amp; Gupta (1989) added one. Even though, these more than 17 colour forms still could not cover all the variation. Among them, seven “subspecies”, i.e.
<emphasis id="B90BEA923E7AFFC5727CFA9394D29A69" box="[1329,1429,1358,1382]" italics="true" pageId="12" pageNumber="389">koreanus</emphasis>
,
<emphasis id="B90BEA923E7AFFC577DAFAAC91BA9A86" box="[151,253,1393,1417]" italics="true" pageId="12" pageNumber="389">tibetanus</emphasis>
,
<emphasis id="B90BEA923E7AFFC57646FAAF90249A86" box="[267,355,1394,1417]" italics="true" pageId="12" pageNumber="389">gressitti</emphasis>
,
<emphasis id="B90BEA923E7AFFC5763CFAAC90B19A86" box="[369,502,1393,1417]" italics="true" pageId="12" pageNumber="389">hainanensis</emphasis>
,
<emphasis id="B90BEA923E7AFFC57549FAAC93259A86" box="[516,610,1393,1417]" italics="true" pageId="12" pageNumber="389">grahami</emphasis>
,? subspecies A and B, are recorded from
<collectingCountry id="F36876103E7AFFC57374FAAC95389A85" box="[1081,1151,1393,1418]" name="China" pageId="12" pageNumber="389">China</collectingCountry>
.
<taxonomicName id="4C7F4D033E7AFFC573C7FAAF94599A86" box="[1162,1310,1394,1417]" class="Insecta" family="Eumenidae" genus="Polistes" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="389" phylum="Arthropoda" rank="species" species="yayeyamae">
<emphasis id="B90BEA923E7AFFC573C7FAAF94599A86" box="[1162,1310,1394,1417]" italics="true" pageId="12" pageNumber="389">P. yayeyamae</emphasis>
</taxonomicName>
is possibly also distributed in
<collectingCountry id="F36876103E7AFFC57627FA4990F99AA2" box="[362,446,1428,1453]" name="Taiwan" pageId="12" pageNumber="389">Taiwan</collectingCountry>
. The colour differences between
<emphasis id="B90BEA923E7AFFC5747AFA4892D29AA2" box="[823,917,1429,1453]" italics="true" pageId="12" pageNumber="389">grahami</emphasis>
,
<emphasis id="B90BEA923E7AFFC574E8FA48956D9AA2" box="[933,1066,1429,1453]" italics="true" pageId="12" pageNumber="389">hainanensis</emphasis>
and
<emphasis id="B90BEA923E7AFFC5732EFA4895FC9AA3" box="[1123,1211,1429,1452]" italics="true" pageId="12" pageNumber="389">gressitti</emphasis>
, are very small and mainly related to the colour of the pronotum and the first metasomal tergite. The colour forms
<emphasis id="B90BEA923E7AFFC57244FA6594209ADF" box="[1289,1383,1464,1488]" italics="true" pageId="12" pageNumber="389">grahami</emphasis>
and
<emphasis id="B90BEA923E7AFFC577DAFA01905B9AFB" box="[151,284,1500,1524]" italics="true" pageId="12" pageNumber="389">hainanensis</emphasis>
overlap in Hainan and it is better to unite these colour forms. Even within f.
<emphasis id="B90BEA923E7AFFC573D0FA0195BC9AFB" box="[1181,1275,1500,1524]" italics="true" pageId="12" pageNumber="389">grahami</emphasis>
there are two colour patterns (mesoscutum with four stripes or two stripes) and are sympatric in
<collectingCountry id="F36876103E7AFFC57319FA2295DC9917" box="[1108,1179,1535,1560]" name="China" pageId="12" pageNumber="389">China</collectingCountry>
. The colour pattern of some specimens collected in Yunnan intergrades f.
<taxonomicName id="4C7F4D033E7AFFC57585F9FE92649934" box="[712,803,1571,1595]" class="Insecta" family="Eumenidae" genus="Polistes" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="389" phylum="Arthropoda" rank="species" species="rothneyi">
<emphasis id="B90BEA923E7AFFC57585F9FE92649934" box="[712,803,1571,1595]" italics="true" pageId="12" pageNumber="389">rothneyi</emphasis>
</taxonomicName>
and f.
<emphasis id="B90BEA923E7AFFC57423F9FE92939934" box="[878,980,1571,1595]" italics="true" pageId="12" pageNumber="389">tibetanus</emphasis>
having four reddish brown stripes on the mesoscutum but the first tergite mostly black (
<figureCitation id="13442A053E7AFFC575D7F99B93AA9950" box="[666,749,1606,1631]" captionStart="FIGURE 19" captionStartId="10.[151,250,1213,1235]" captionTargetBox="[302,1284,301,1191]" captionTargetId="figure@10.[302,1284,301,1192]" captionTargetPageId="10" captionText="FIGURE 19. Polistes olivaceus (De Geer, 1773), habitus. A, C, E, dorsal view; B, D, F, lateral view: A B, female, specimens from Hainan, China; C D, male, specimens from Somoya; E F, specimens from Assam, India (as Polistes hebraeus Fabricius, 1787)." httpUri="https://zenodo.org/record/253077/files/figure.png" pageId="12" pageNumber="389">Fig. 19</figureCitation>
). However, one specimen collected in Yunnan (no. 20046541) has the head reddish yellow and the bands and stripes yellow with reddish brown tinge, what fits f.
<emphasis id="B90BEA923E7AFFC573ADF9B49418998E" box="[1248,1375,1641,1665]" italics="true" pageId="12" pageNumber="389">sikkimensis</emphasis>
well. In addition, the two uncertain A and B forms show a more or less continuous variation series. Similar situation occurs between the specimens from northeastern
<collectingCountry id="F36876103E7AFFC575F0F96D924599C6" box="[701,770,1712,1737]" name="China" pageId="12" pageNumber="389">China</collectingCountry>
, f.
<emphasis id="B90BEA923E7AFFC57469F96C922D99C7" box="[804,874,1713,1736]" italics="true" pageId="12" pageNumber="389">iwatai</emphasis>
(area on basal band and waved line on the second metasomal tergite often somewhat reddish) and f.
<emphasis id="B90BEA923E7AFFC57584F909926A99E3" box="[713,813,1748,1772]" italics="true" pageId="12" pageNumber="389">koreanus</emphasis>
(the first metasomal tergite black with a narrow apical band). It is interesting that the colour pattern of f.
<emphasis id="B90BEA923E7AFFC5758CF92592409800" box="[705,775,1784,1807]" italics="true" pageId="12" pageNumber="389">iwatai</emphasis>
is very similar to that of f.
<emphasis id="B90BEA923E7AFFC57379F92A95D59800" box="[1076,1170,1783,1807]" italics="true" pageId="12" pageNumber="389">grahami</emphasis>
so that the latter can be considered transitional. Van der
<bibRefCitation id="EFEE4B713E7AFFC57541F8C693D9983B" author="Vecht" box="[524,670,1819,1844]" pageId="12" pageNumber="389" refString="van der Vecht, J. (1968) The geographic variation of Polistes (Megapolistes subg. n.) rothneyi Cameron. Bijdragen Tot de Dierkunde, 38, 97 - 109." type="journal article" year="1968">Vecht (1968)</bibRefCitation>
mentioned that the colour of the body ranges from paler or dark yellow through various shades of brownish red to black. He roughly divided the specimens into three groups i.e. yellow and black, brownish-red and more or less transitional forms. Our observations showed that the Chinese colour forms of
<taxonomicName id="4C7F4D033E7AFFC57600F85B90869892" box="[333,449,1925,1949]" class="Insecta" family="Eumenidae" genus="Polistes" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="389" phylum="Arthropoda" rank="species" species="rothneyi">
<emphasis id="B90BEA923E7AFFC57600F85B90869892" box="[333,449,1925,1949]" italics="true" pageId="12" pageNumber="389">P. rothneyi</emphasis>
</taxonomicName>
can be defined only by colour patterns that intergrade in some degree. This leads us to synonymize all (as far seen from
<collectingCountry id="F36876103E7AFFC5755EF875931F98CE" box="[531,600,1960,1985]" name="China" pageId="12" pageNumber="389">China</collectingCountry>
) as colour forms of
<taxonomicName id="4C7F4D033E7AFFC57473F87492F698CE" box="[830,945,1961,1985]" class="Insecta" family="Eumenidae" genus="Polistes" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="389" phylum="Arthropoda" rank="species" species="rothneyi">
<emphasis id="B90BEA923E7AFFC57473F87492F698CE" box="[830,945,1961,1985]" italics="true" pageId="12" pageNumber="389">P. rothneyi</emphasis>
</taxonomicName>
. Since it is difficult and tedious to describe the colour patterns in words, we prefer to show its variation in a series photographs (
<figureCitation id="13442A053E7AFFC57300F811959698EA" box="[1101,1233,1996,2021]" captionStart-0="FIGURE 19" captionStart-1="FIGURE 20" captionStart-2="FIGURE 21" captionStartId-0="10.[151,250,1213,1235]" captionStartId-1="11.[151,250,1485,1507]" captionStartId-2="13.[151,250,1776,1798]" captionTargetBox-0="[302,1284,301,1191]" captionTargetBox-1="[302,1284,409,1462]" captionTargetBox-2="[302,1284,468,1753]" captionTargetId-0="figure@10.[302,1284,301,1192]" captionTargetId-1="figure@11.[302,1284,409,1463]" captionTargetId-2="figure@13.[302,1284,468,1754]" captionTargetPageId-0="10" captionTargetPageId-1="11" captionTargetPageId-2="13" captionText-0="FIGURE 19. Polistes olivaceus (De Geer, 1773), habitus. A, C, E, dorsal view; B, D, F, lateral view: A B, female, specimens from Hainan, China; C D, male, specimens from Somoya; E F, specimens from Assam, India (as Polistes hebraeus Fabricius, 1787)." captionText-1="FIGURE 20. Polistes rothneyi Cameron, 1900, Southwest plateau group (female). A, C, E, G, dorsal view; B, D, F, H, lateral view: A B, f. rothneyi (det. van der Vecht) from Shillong, India; C D, f. tibetanus van der Vecht, 1968 from Tibet, paratype; E F, specimens from Yunnan, showing the color pattern between f. rothneyi and f. tibetanus; G H, specimens from Yunnan, showing the color pattern as? subs. A (van der Vecht, 1968: 103, fig. 7)." captionText-2="FIGURE 21. Polistes rothneyi Cameron, 1900, Common yellow-black group (female). A, C, E, G, I, dorsal view; B, D, F, H, J, lateral view: A B, f. grahami van der Vecht, 1968 from Sichuan, paratype (showing mesonotum with two yellow stripes); C D, f. grahami van der Vecht, 1968 from Fujian, paratype (showing mesonotum with four yellow stripes); E F, f. iwatai van der Vecht, 1968 from Japan, paratype (showing color pattern very similar as A-B f. grahami, but pronotum mostly yellow); G H, specimens from Hainan with the color pattern same as f. hainanensis van der Vecht, 1968, showing the apical band of the first metasomal tergite entirely yellow; I J, f. gressitti van der Vecht, 1968 from Taiwan, paratype (showing pronotum entirely yellow); K L, f. yayeyamae Matsumura, 1908 from Iriomote Loochoo Is., paratype (showing body mainly yellow)." httpUri-0="https://zenodo.org/record/253077/files/figure.png" httpUri-1="https://zenodo.org/record/253078/files/figure.png" httpUri-2="https://zenodo.org/record/253079/files/figure.png" pageId="12" pageNumber="389">Figs 1921</figureCitation>
). Referring to van der
<bibRefCitation id="EFEE4B713E7BFFC4778CFF4A90169FBF" author="Vecht" box="[193,337,151,176]" pageId="13" pageNumber="390" refString="van der Vecht, J. (1968) The geographic variation of Polistes (Megapolistes subg. n.) rothneyi Cameron. Bijdragen Tot de Dierkunde, 38, 97 - 109." type="journal article" year="1968">Vecht (1968)</bibRefCitation>
, we divide the examined Chinese specimens into three colour groups i.e. Southwest Plateau group (brownish-red to black, distributed in southeastern
<collectingCountry id="F36876103E7BFFC47582FF6692539FDB" box="[719,788,187,212]" name="China" pageId="13" pageNumber="390">China</collectingCountry>
, Yunnan and Tibet), Common yellow-black group (widely distributed in the mainland of
<collectingCountry id="F36876103E7BFFC4755CFF0393139FF8" box="[529,596,222,247]" name="China" pageId="13" pageNumber="390">China</collectingCountry>
spreading southward to Hainan and
<collectingCountry id="F36876103E7BFFC47356FF0395299FF8" box="[1051,1134,222,247]" name="Taiwan" pageId="13" pageNumber="390">Taiwan</collectingCountry>
, including f.
<emphasis id="B90BEA923E7BFFC4725AFF0294329FF8" box="[1303,1397,223,247]" italics="true" pageId="13" pageNumber="390">grahami</emphasis>
, f.
<emphasis id="B90BEA923E7BFFC477DAFEDF905B9E15" box="[151,284,258,282]" italics="true" pageId="13" pageNumber="390">hainanensis</emphasis>
, f.
<emphasis id="B90BEA923E7BFFC47609FEDE90DB9E15" box="[324,412,259,282]" italics="true" pageId="13" pageNumber="390">gressitti</emphasis>
and f.
<taxonomicName id="4C7F4D033E7BFFC476A3FEDE932E9E15" box="[494,617,259,282]" class="Insecta" family="Eumenidae" genus="Polistes" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="390" phylum="Arthropoda" rank="species" species="yayeyamae">
<emphasis id="B90BEA923E7BFFC476A3FEDE932E9E15" box="[494,617,259,282]" italics="true" pageId="13" pageNumber="390">yayeyamae</emphasis>
</taxonomicName>
) and Northeast group (colour pattern between f.
<emphasis id="B90BEA923E7BFFC473D4FEDE95989E15" box="[1177,1247,259,282]" italics="true" pageId="13" pageNumber="390">iwatai</emphasis>
and f.
<emphasis id="B90BEA923E7BFFC4727CFEDF94D29E15" box="[1329,1429,258,282]" italics="true" pageId="13" pageNumber="390">koreanus</emphasis>
, generally with reddish brown blotches, the first metasomal tergite black, at most with a narrow apical band; distributed in Northeast
<collectingCountry id="F36876103E7BFFC476EAFE9490AA9E6D" box="[423,493,329,354]" name="China" pageId="13" pageNumber="390">China</collectingCountry>
, Shandong (Weihai), Liaoning, Jilin). For convenience we use the available names as colour forms for arranging the specimens into different colour groups. We did not find a colour gap between the Common black yellow group (
<figureCitation id="13442A053E7BFFC476A1FE4D937A9EA7" box="[492,573,400,425]" captionStart="FIGURE 21" captionStartId="13.[151,250,1776,1798]" captionTargetBox="[302,1284,468,1753]" captionTargetId="figure@13.[302,1284,468,1754]" captionTargetPageId="13" captionText="FIGURE 21. Polistes rothneyi Cameron, 1900, Common yellow-black group (female). A, C, E, G, I, dorsal view; B, D, F, H, J, lateral view: A B, f. grahami van der Vecht, 1968 from Sichuan, paratype (showing mesonotum with two yellow stripes); C D, f. grahami van der Vecht, 1968 from Fujian, paratype (showing mesonotum with four yellow stripes); E F, f. iwatai van der Vecht, 1968 from Japan, paratype (showing color pattern very similar as A-B f. grahami, but pronotum mostly yellow); G H, specimens from Hainan with the color pattern same as f. hainanensis van der Vecht, 1968, showing the apical band of the first metasomal tergite entirely yellow; I J, f. gressitti van der Vecht, 1968 from Taiwan, paratype (showing pronotum entirely yellow); K L, f. yayeyamae Matsumura, 1908 from Iriomote Loochoo Is., paratype (showing body mainly yellow)." httpUri="https://zenodo.org/record/253079/files/figure.png" pageId="13" pageNumber="390">Fig. 21</figureCitation>
) and the Northeast group (
<figureCitation id="13442A053E7BFFC47424FE4D92FA9EA7" box="[873,957,400,425]" captionStart="FIGURE 22" captionStartId="14.[151,250,1341,1363]" captionTargetBox="[302,1284,193,1319]" captionTargetId="figure@14.[302,1284,193,1319]" captionTargetPageId="14" captionText="FIGURE 22. Polistes rothneyi Cameron, 1900, Northeast group (female) (showing the colour pattern variation). A, C, E, G, I, J, dorsal view; B, D, F, H, lateral view: A B, specimen from Shenyang, Liaoning; C D, specimen from Shenyang, Kuandian; E F, specimen from Qianshan, Liaoning; G H, f. koreanus van der Vecht, 1968 from Seoul area, paratype (with pronotum black except narrow yellow margin, first metasomal tergite black with a narrow apical band); I, specimen from Siping, Jilin; J, specimen from Xinbin, Liaoning." httpUri="https://zenodo.org/record/253080/files/figure.png" pageId="13" pageNumber="390">Fig. 22</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>