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<document id="A779F4400F7B9813C848B0517D081700" ID-DOI="10.5852/ejt.2023.891.2281" ID-GBIF-Dataset="32ba8e03-593e-4dbb-a965-258f61d6e665" ID-ISSN="2118-9773" ID-Zenodo-Dep="8382615" ID-ZooBank="A8753CAC-F181-4E6D-82F3-F4A456025447" IM.bibliography_approvedBy="jonas" IM.illustrations_approvedBy="jonas" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="jonas" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="jonas" checkinTime="1695804784739" checkinUser="plazi" docAuthor="Nethupul, Hasitha, Stöhr, Sabine &amp; Zhang, Haibin" docDate="2023" docId="03D6878CE63EFC1D0F57FD7ECFEF3E32" docLanguage="en" docName="EJT.2023.891.167-185.pdf" docOrigin="European Journal of Taxonomy 891" docSource="https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2281/9835" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Ophiuroglypha fendouzhe Nethupul &amp; Stöhr &amp; Zhang 2023, sp. nov." docType="treatment" docUuid="31A4A22F-7F61-4510-BF10-67851B458559" docUuidSource="ZooBank" docVersion="7" lastPageNumber="178" masterDocId="FFEFFFF4E637FC160D1DFFBCCD4E3858" masterDocTitle="Deepest known novel species of the genus Ophiuroglypha Hertz, 1927 (Echinodermata: Ophiuroidea) from the central rift zone, Philippine Sea" masterLastPageNumber="185" masterPageNumber="167" pageNumber="176" updateTime="1701999656917" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="3EACBA77EB0505DE0674BF751A078886">Deepest known novel species of the genus Ophiuroglypha Hertz, 1927 (Echinodermata: Ophiuroidea) from the central rift zone, Philippine Sea</mods:title>
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<mods:namePart id="F198C382275AF00EC8FE31CB1A6AD080">Nethupul, Hasitha</mods:namePart>
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<mods:affiliation id="928085F1FBB932CED818506B9766646D">Institute of Deep-sea Science and Engineering, Chinese Academy of Sciences, CAS, 57200 Sanya, China. &amp; University of Chinese Academy of Sciences, Beijing 100039, China.</mods:affiliation>
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<mods:namePart id="FF1FEF06F6DA65334FC242FFE95F6C3B">Stöhr, Sabine</mods:namePart>
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<mods:affiliation id="2E93A35E0F120EDD40C8EEDC7D91AE4A">Swedish Museum of Natural History, Dept of Zoology, Box 50007, 10405 Stockholm, Sweden.</mods:affiliation>
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<mods:namePart id="FED798B6DF7B42D309B63B480D1A7C59">Zhang, Haibin</mods:namePart>
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<mods:affiliation id="31AB804CDFE8633BF6A3A9AE337A1E0F">Institute of Deep-sea Science and Engineering, Chinese Academy of Sciences, CAS, 57200 Sanya, China.</mods:affiliation>
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<mods:title id="0FA98AF687DA3F8717B865A89254088B">European Journal of Taxonomy</mods:title>
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<mods:date id="6D9D1FFC3567804492C6FE76A381F502">2023</mods:date>
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<treatment id="03D6878CE63EFC1D0F57FD7ECFEF3E32" ID-DOI="http://doi.org/10.5281/zenodo.8382617" ID-GBIF-Taxon="213502736" ID-Zenodo-Dep="8382617" LSID="urn:lsid:zoobank.org:act:31A4A22F-7F61-4510-BF10-67851B458559" httpUri="http://treatment.plazi.org/id/03D6878CE63EFC1D0F57FD7ECFEF3E32" lastPageId="11" lastPageNumber="178" pageId="9" pageNumber="176">
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<taxonomicName id="4C7F4D19E63EFC1F0F57FD7ECEC93A84" ID-CoL="c4606408-1575-475c-851e-c03d5440d76b" authority="Nethupul &amp; Stöhr &amp; Zhang, 2023" authorityName="Nethupul &amp; Stöhr &amp; Zhang" authorityYear="2023" box="[586,903,706,732]" class="Ophiuroidea" family="Ophiopyrgidae" genus="Ophiuroglypha" kingdom="Animalia" order="Ophiurida" pageId="9" pageNumber="176" phylum="Echinodermata" rank="species" species="fendouzhe" status="sp. nov.">
<emphasis id="B90BEA88E63EFC1F0F57FD7ECEC93A84" bold="true" box="[586,903,706,732]" italics="true" pageId="9" pageNumber="176">Ophiuroglypha fendouzhe</emphasis>
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<paragraph id="8BC0369AE63EFC1F0FFAFCB6CE033B7C" blockId="9.[379,1209,706,804]" box="[743,845,778,804]" pageId="9" pageNumber="176">
<figureCitation id="13442A1FE63EFC1F0FFAFCB6CE033B7C" box="[743,845,778,804]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStart-3="Fig" captionStart-4="Fig" captionStartId-0="1.[189,232,1998,2024]" captionStartId-1="4.[189,232,1931,1957]" captionStartId-2="10.[190,233,1750,1776]" captionStartId-3="12.[189,232,1807,1833]" captionStartId-4="13.[189,232,945,971]" captionTargetBox-0="[190,1394,1166,1962]" captionTargetBox-1="[215,1395,680,1886]" captionTargetBox-2="[189,1399,444,1719]" captionTargetBox-3="[189,1399,265,1760]" captionTargetBox-4="[189,1399,265,895]" captionTargetId-0="figure-325@1.[189,1399,1155,1979]" captionTargetId-1="figure-141@4.[189,1399,675,1894]" captionTargetId-2="figure-68@10.[189,1399,444,1719]" captionTargetId-3="figure-101@12.[189,1399,265,1760]" captionTargetId-4="figure-427@13.[189,1399,265,895]" captionTargetPageId-0="1" captionTargetPageId-1="4" captionTargetPageId-2="10" captionTargetPageId-3="12" captionTargetPageId-4="13" captionText-0="Fig. 1. Map of collecting station of the central rift zone, Philippine Sea, in this study." captionText-1="Fig. 2. Maximum likelihood (ML) tree of the family Ophiopyrgidae, based on partial COI sequences (bootstrap support values were generated with rapid bootstrapping algorithm for 1000 replicates; red = new species). Abbreviation: C = Clade." captionText-2="Fig. 3. Ophiuroglypha fendouzhe sp. nov., holotype (IDSSE-EEB-SW0250). A. Dorsal aspect. B. Ventral aspect. C. Central disc (overlapping disc scales). D. Dorsal arm base. E. Ventral disc, and mouth. F. Ventral arm base. G. Dorsal arm. HJ. Ventral arm (partially broken arm spine highlighted in the Fig. I). KL. Lateral arm. Abbreviations: arc = arm comb; ars = arm spine; as = adoral shield; asp = adoral shield spine; cpp = central primary plate; dap = dorsal arm plate; gs = genital slit; lap = lateral arm plate; m = madreporite; op = oral plate; os = oral shield; otp = oral tentacle pore; rs = radial shield; tp = tentacle pore; ts = tentacle scale; vap = ventral arm plate; vts = ventral tentacle scale. Scale bars: AB = 2 mm; C, EF =1 mm; D, GL = 500 μm." captionText-3="Fig. 4. Ophiuroglypha fendouzhe sp. nov., paratype (IDSSE-EEB-SW0251). AC. Lateral arm plate. DH. Vertebrae. D. Distal view (right side of the structure partially broken). E. Proximal view. F. Ventral view. G. Dorsal view. H. Dorsolateral view. Abbreviations: asa = arm spine articulation; d = dorsal; dist = distal; kn = knob; mo = muscle opening; no = nerve opening; pb = podial basin; prox = proximal; tfp = tube foot pore; v = ventral. Scale bars: A, C, FH = 500 μm; B = 100 μm; DE = 300 μm." captionText-4="Fig. 5. Ophiuroglypha fendouzhe sp. nov., paratype (IDSSE-EEB-SW0251). A. Dorsal aspect. B. Ventral aspect. C. Central disc (overlapping disc scales). D. Ventral arm. E. Dorsal arm. F. Lateral arm. Abbreviations: ars = arm spine; as = adoral shield; asp = adoral shield spine; cpp = central primary plate; dap = dorsal arm plate; gs = genital slit; lap = lateral arm plate; m = madreporite; op = oral plate; os = oral shield; otp = oral tentacle pore; tp = tentacle pore; ts = tentacle scale; vap = ventral arm plate; vts = ventral tentacle scale. Scale bars: AB = 2 mm; C, EF = 500 μm; D = 1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.8382619" figureDoi-1="http://doi.org/10.5281/zenodo.8382621" figureDoi-2="http://doi.org/10.5281/zenodo.8382623" figureDoi-3="http://doi.org/10.5281/zenodo.8382625" figureDoi-4="http://doi.org/10.5281/zenodo.8382627" httpUri-0="https://zenodo.org/record/8382619/files/figure.png" httpUri-1="https://zenodo.org/record/8382621/files/figure.png" httpUri-2="https://zenodo.org/record/8382623/files/figure.png" httpUri-3="https://zenodo.org/record/8382625/files/figure.png" httpUri-4="https://zenodo.org/record/8382627/files/figure.png" pageId="9" pageNumber="176">Figs 15</figureCitation>
</paragraph>
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<subSubSection id="C3656511E63EFC1F0DA0FCF7C85E3C32" pageId="9" pageNumber="176" type="diagnosis">
<paragraph id="8BC0369AE63EFC1F0DA0FCF7CC783B3D" blockId="9.[189,310,843,869]" box="[189,310,843,869]" pageId="9" pageNumber="176">
<heading id="D08881F6E63EFC1F0DA0FCF7CC783B3D" bold="true" box="[189,310,843,869]" fontSize="11" level="3" pageId="9" pageNumber="176" reason="3">
<emphasis id="B90BEA88E63EFC1F0DA0FCF7CC783B3D" bold="true" box="[189,310,843,869]" pageId="9" pageNumber="176">Diagnosis</emphasis>
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<paragraph id="8BC0369AE63EFC1F0DA0FCC7C85E3C32" blockId="9.[189,1399,891,1130]" pageId="9" pageNumber="176">
Disc pentagonal, covered by large irregular overlapping scales. Radial shields broad, slightly wider than long, triangular, barely connected to each other at distal end, and proximal half separated by one elongated disc scale (
<figureCitation id="13442A1FE63EFC1F0CA5FC7FCF743B85" box="[440,570,963,989]" captionStart="Fig" captionStartId="10.[190,233,1750,1776]" captionTargetBox="[189,1399,444,1719]" captionTargetId="figure-68@10.[189,1399,444,1719]" captionTargetPageId="10" captionText="Fig. 3. Ophiuroglypha fendouzhe sp. nov., holotype (IDSSE-EEB-SW0250). A. Dorsal aspect. B. Ventral aspect. C. Central disc (overlapping disc scales). D. Dorsal arm base. E. Ventral disc, and mouth. F. Ventral arm base. G. Dorsal arm. HJ. Ventral arm (partially broken arm spine highlighted in the Fig. I). KL. Lateral arm. Abbreviations: arc = arm comb; ars = arm spine; as = adoral shield; asp = adoral shield spine; cpp = central primary plate; dap = dorsal arm plate; gs = genital slit; lap = lateral arm plate; m = madreporite; op = oral plate; os = oral shield; otp = oral tentacle pore; rs = radial shield; tp = tentacle pore; ts = tentacle scale; vap = ventral arm plate; vts = ventral tentacle scale. Scale bars: AB = 2 mm; C, EF =1 mm; D, GL = 500 μm." figureDoi="http://doi.org/10.5281/zenodo.8382623" httpUri="https://zenodo.org/record/8382623/files/figure.png" pageId="9" pageNumber="176">Fig. 3AC</figureCitation>
). Ventral disc covered by large, overlapping scales, slightly larger than dorsal disc (
<figureCitation id="13442A1FE63EFC1F0C57FC5ACCE93C58" box="[330,423,998,1024]" captionStart="Fig" captionStartId="10.[190,233,1750,1776]" captionTargetBox="[189,1399,444,1719]" captionTargetId="figure-68@10.[189,1399,444,1719]" captionTargetPageId="10" captionText="Fig. 3. Ophiuroglypha fendouzhe sp. nov., holotype (IDSSE-EEB-SW0250). A. Dorsal aspect. B. Ventral aspect. C. Central disc (overlapping disc scales). D. Dorsal arm base. E. Ventral disc, and mouth. F. Ventral arm base. G. Dorsal arm. HJ. Ventral arm (partially broken arm spine highlighted in the Fig. I). KL. Lateral arm. Abbreviations: arc = arm comb; ars = arm spine; as = adoral shield; asp = adoral shield spine; cpp = central primary plate; dap = dorsal arm plate; gs = genital slit; lap = lateral arm plate; m = madreporite; op = oral plate; os = oral shield; otp = oral tentacle pore; rs = radial shield; tp = tentacle pore; ts = tentacle scale; vap = ventral arm plate; vts = ventral tentacle scale. Scale bars: AB = 2 mm; C, EF =1 mm; D, GL = 500 μm." figureDoi="http://doi.org/10.5281/zenodo.8382623" httpUri="https://zenodo.org/record/8382623/files/figure.png" pageId="9" pageNumber="176">Fig. 3D</figureCitation>
). Jaw slender, large, contiguous, pair of pointed ventralmost teeth at apex, and up to five lateral oral papillae (
<figureCitation id="13442A1FE63EFC1F0CFFFBB5CF703C7B" box="[482,574,1033,1059]" captionStart="Fig" captionStartId="10.[190,233,1750,1776]" captionTargetBox="[189,1399,444,1719]" captionTargetId="figure-68@10.[189,1399,444,1719]" captionTargetPageId="10" captionText="Fig. 3. Ophiuroglypha fendouzhe sp. nov., holotype (IDSSE-EEB-SW0250). A. Dorsal aspect. B. Ventral aspect. C. Central disc (overlapping disc scales). D. Dorsal arm base. E. Ventral disc, and mouth. F. Ventral arm base. G. Dorsal arm. HJ. Ventral arm (partially broken arm spine highlighted in the Fig. I). KL. Lateral arm. Abbreviations: arc = arm comb; ars = arm spine; as = adoral shield; asp = adoral shield spine; cpp = central primary plate; dap = dorsal arm plate; gs = genital slit; lap = lateral arm plate; m = madreporite; op = oral plate; os = oral shield; otp = oral tentacle pore; rs = radial shield; tp = tentacle pore; ts = tentacle scale; vap = ventral arm plate; vts = ventral tentacle scale. Scale bars: AB = 2 mm; C, EF =1 mm; D, GL = 500 μm." figureDoi="http://doi.org/10.5281/zenodo.8382623" httpUri="https://zenodo.org/record/8382623/files/figure.png" pageId="9" pageNumber="176">Fig. 3E</figureCitation>
). Second oral tentacle pore large, opening outside mouth slit with four adoral shield spines and five ventral tentacle scales (
<figureCitation id="13442A1FE63EFC1F0E37FB91CEE83C1F" box="[810,934,1069,1095]" captionStart="Fig" captionStartId="10.[190,233,1750,1776]" captionTargetBox="[189,1399,444,1719]" captionTargetId="figure-68@10.[189,1399,444,1719]" captionTargetPageId="10" captionText="Fig. 3. Ophiuroglypha fendouzhe sp. nov., holotype (IDSSE-EEB-SW0250). A. Dorsal aspect. B. Ventral aspect. C. Central disc (overlapping disc scales). D. Dorsal arm base. E. Ventral disc, and mouth. F. Ventral arm base. G. Dorsal arm. HJ. Ventral arm (partially broken arm spine highlighted in the Fig. I). KL. Lateral arm. Abbreviations: arc = arm comb; ars = arm spine; as = adoral shield; asp = adoral shield spine; cpp = central primary plate; dap = dorsal arm plate; gs = genital slit; lap = lateral arm plate; m = madreporite; op = oral plate; os = oral shield; otp = oral tentacle pore; rs = radial shield; tp = tentacle pore; ts = tentacle scale; vap = ventral arm plate; vts = ventral tentacle scale. Scale bars: AB = 2 mm; C, EF =1 mm; D, GL = 500 μm." figureDoi="http://doi.org/10.5281/zenodo.8382623" httpUri="https://zenodo.org/record/8382623/files/figure.png" pageId="9" pageNumber="176">Fig. 3EF</figureCitation>
). Arms slender. Dorsal and ventral arm plates separated (
<figureCitation id="13442A1FE63EFC1F0C95FBECCF483C32" box="[392,518,1104,1130]" captionStart="Fig" captionStartId="10.[190,233,1750,1776]" captionTargetBox="[189,1399,444,1719]" captionTargetId="figure-68@10.[189,1399,444,1719]" captionTargetPageId="10" captionText="Fig. 3. Ophiuroglypha fendouzhe sp. nov., holotype (IDSSE-EEB-SW0250). A. Dorsal aspect. B. Ventral aspect. C. Central disc (overlapping disc scales). D. Dorsal arm base. E. Ventral disc, and mouth. F. Ventral arm base. G. Dorsal arm. HJ. Ventral arm (partially broken arm spine highlighted in the Fig. I). KL. Lateral arm. Abbreviations: arc = arm comb; ars = arm spine; as = adoral shield; asp = adoral shield spine; cpp = central primary plate; dap = dorsal arm plate; gs = genital slit; lap = lateral arm plate; m = madreporite; op = oral plate; os = oral shield; otp = oral tentacle pore; rs = radial shield; tp = tentacle pore; ts = tentacle scale; vap = ventral arm plate; vts = ventral tentacle scale. Scale bars: AB = 2 mm; C, EF =1 mm; D, GL = 500 μm." figureDoi="http://doi.org/10.5281/zenodo.8382623" httpUri="https://zenodo.org/record/8382623/files/figure.png" pageId="9" pageNumber="176">Fig. 3FH</figureCitation>
). Tentacle pores absent after seven or nine arm segments (
<figureCitation id="13442A1FE63EFC1F09ADFBECC84C3C32" box="[1200,1282,1104,1130]" captionStart="Fig" captionStartId="10.[190,233,1750,1776]" captionTargetBox="[189,1399,444,1719]" captionTargetId="figure-68@10.[189,1399,444,1719]" captionTargetPageId="10" captionText="Fig. 3. Ophiuroglypha fendouzhe sp. nov., holotype (IDSSE-EEB-SW0250). A. Dorsal aspect. B. Ventral aspect. C. Central disc (overlapping disc scales). D. Dorsal arm base. E. Ventral disc, and mouth. F. Ventral arm base. G. Dorsal arm. HJ. Ventral arm (partially broken arm spine highlighted in the Fig. I). KL. Lateral arm. Abbreviations: arc = arm comb; ars = arm spine; as = adoral shield; asp = adoral shield spine; cpp = central primary plate; dap = dorsal arm plate; gs = genital slit; lap = lateral arm plate; m = madreporite; op = oral plate; os = oral shield; otp = oral tentacle pore; rs = radial shield; tp = tentacle pore; ts = tentacle scale; vap = ventral arm plate; vts = ventral tentacle scale. Scale bars: AB = 2 mm; C, EF =1 mm; D, GL = 500 μm." figureDoi="http://doi.org/10.5281/zenodo.8382623" httpUri="https://zenodo.org/record/8382623/files/figure.png" pageId="9" pageNumber="176">Fig. 3I</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3656511E63EFC1F0DA0FB2DCC7A3CA6" pageId="9" pageNumber="176" type="etymology">
<paragraph id="8BC0369AE63EFC1F0DA0FB2DCC0A3CF3" blockId="9.[189,324,1169,1195]" box="[189,324,1169,1195]" pageId="9" pageNumber="176">
<heading id="D08881F6E63EFC1F0DA0FB2DCC0A3CF3" bold="true" box="[189,324,1169,1195]" fontSize="11" level="3" pageId="9" pageNumber="176" reason="3">
<emphasis id="B90BEA88E63EFC1F0DA0FB2DCC0A3CF3" bold="true" box="[189,324,1169,1195]" pageId="9" pageNumber="176">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8BC0369AE63EFC1F0DA0FB7CCC7A3CA6" blockId="9.[189,1399,1216,1278]" pageId="9" pageNumber="176">The specific name is dedicated to the manned submersible vessel Fendouzhe, which collected the specimen.</paragraph>
</subSubSection>
<subSubSection id="C3656511E63EFC1F0DA0FA98CE093EDC" pageId="9" pageNumber="176" type="materials_examined">
<paragraph id="8BC0369AE63EFC1F0DA0FA98CCE43D66" blockId="9.[189,426,1316,1342]" box="[189,426,1316,1342]" pageId="9" pageNumber="176">
<heading id="D08881F6E63EFC1F0DA0FA98CCE43D66" bold="true" box="[189,426,1316,1342]" fontSize="11" level="3" pageId="9" pageNumber="176" reason="3">
<emphasis id="B90BEA88E63EFC1F0DA0FA98CCE43D66" bold="true" box="[189,426,1316,1342]" pageId="9" pageNumber="176">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="8BC0369AE63EFC1F0DF7FAEFCC653D80" blockId="9.[189,1399,1363,1496]" pageId="9" pageNumber="176">
<materialsCitation id="3B173CC7E63EFC1F0DF7FAEFCC693D80" ID-GBIF-Occurrence="4426885302" collectingDate="2021-09-20" collectionCode="IDSSE-EEB-SW" collectorName="Fendouzhe" elevation="7729" latitude="16.945667" location="PHILIPPINE SEA" longLatPrecision="13" longitude="129.73883" pageId="9" pageNumber="176" specimenCode="IDSSE-EEB-SW0250" specimenCount="1" typeStatus="holotype">
<emphasis id="B90BEA88E63EFC1F0DF7FAEFCC123D35" bold="true" box="[234,348,1363,1389]" pageId="9" pageNumber="176">
<typeStatus id="54C48838E63EFC1F0DF7FAEFCC123D35" box="[234,348,1363,1389]" pageId="9" pageNumber="176" type="holotype">Holotype</typeStatus>
</emphasis>
<location id="8EA06041E63EFC1F0DA0FACBCCD83DC9" LSID="urn:lsid:plazi:treatment:03D6878CE63EFC1D0F57FD7ECFEF3E32:8EA06041E63EFC1F0DA0FACBCCD83DC9" box="[189,406,1399,1425]" latitude="16.945667" longLatPrecision="13" longitude="129.73883" name="PHILIPPINE SEA" pageId="9" pageNumber="176">PHILIPPINE SEA</location>
• central rift zone;
<geoCoordinate id="EE4B505DE63EFC1F0F6EFACBCE4E3DC9" box="[627,768,1399,1425]" degrees="16" direction="north" minutes="56.74" orientation="latitude" pageId="9" pageNumber="176" precision="9" value="16.945667">16º56.74 N</geoCoordinate>
,
<geoCoordinate id="EE4B505DE63EFC1F0E17FACBCEED3DC9" box="[778,931,1399,1425]" degrees="129" direction="east" minutes="44.33" orientation="longitude" pageId="9" pageNumber="176" precision="9" value="129.73883">129º44.33 E</geoCoordinate>
; depth
<quantity id="4C879B7FE63EFC1F0EEBFACBC91F3DC9" box="[1014,1105,1399,1425]" metricMagnitude="3" metricUnit="m" metricValue="7.728999999999999" pageId="9" pageNumber="176" unit="m" value="7729.0">
<elevation id="0052D1A9E63EFC1F0EEBFACBC91F3DC9" box="[1014,1105,1399,1425]" metricMagnitude="3" metricUnit="m" metricValue="7.728999999999999" pageId="9" pageNumber="176" unit="m" value="7729.0">7729 m</elevation>
</quantity>
;
<date id="FFC1105AE63EFC1F0941FACBC9B63DC9" box="[1116,1272,1399,1425]" pageId="9" pageNumber="176" value="2021-09-20">
<collectingDate id="EF85E9B2E63EFC1F0941FACBC9B63DC9" box="[1116,1272,1399,1425]" pageId="9" pageNumber="176" value="2021-09-20">20 Sep. 2021</collectingDate>
</date>
; collecting event: stn. SC043;
<collectorName id="268A534CE63EFC1F0C87FA27CF553DED" box="[410,539,1435,1461]" pageId="9" pageNumber="176">Fendouzhe</collectorName>
msv leg.; GenBank:
<accessionNumber id="942CAB79E63EFC1F0E0CFA27CED93DED" box="[785,919,1435,1461]" pageId="9" pageNumber="176">ON783074</accessionNumber>
; preserved in 95% ethanol;
<specimenCode id="DBD99EE1E63EFC1F09C7FA27CC693D80" collectionCode="IDSSE-EEB-SW" pageId="9" pageNumber="176">IDSSE-EEB-SW0250</specimenCode>
</materialsCitation>
.
</paragraph>
<paragraph id="8BC0369AE63EFC1F0DF7FA43CC293E41" blockId="9.[189,1398,1535,1668]" box="[234,359,1535,1561]" pageId="9" pageNumber="176">
<emphasis id="B90BEA88E63EFC1F0DF7FA43CC293E41" bold="true" box="[234,359,1535,1561]" pageId="9" pageNumber="176">
<typeStatus id="54C48838E63EFC1F0DF7FA43CC293E41" box="[234,359,1535,1561]" pageId="9" pageNumber="176" type="paratype">Paratypes</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8BC0369AE63EFC1F0DA0F99FCE093EDC" blockId="9.[189,1398,1535,1668]" pageId="9" pageNumber="176">
<materialsCitation id="3B173CC7E63EFC1F0DA0F99FCFCE3E38" ID-GBIF-Occurrence="4426885301" collectingDate="2021-09-20" collectionCode="IDSSE-EEB-SW" collectorName="Fendouzhe" elevation="7729" latitude="16.945667" location="PHILIPPINE SEA" longLatPrecision="13" longitude="129.73883" pageId="9" pageNumber="176" specimenCode="IDSSE-EEB-SW0251" specimenCount="1" typeStatus="paratype">
<location id="8EA06041E63EFC1F0DA0F99FCCD83E65" LSID="urn:lsid:plazi:treatment:03D6878CE63EFC1D0F57FD7ECFEF3E32:8EA06041E63EFC1F0DA0F99FCCD83E65" box="[189,406,1571,1597]" name="PHILIPPINE SEA" pageId="9" pageNumber="176">PHILIPPINE SEA</location>
<specimenCount id="9D79FD13E63EFC1F0CB3F99FCF793E65" box="[430,567,1571,1597]" pageId="9" pageNumber="176" type="generic">1 specimen</specimenCount>
; same collection data as for holotype; GenBank:
<accessionNumber id="942CAB79E63EFC1F096EF99FC9B73E65" box="[1139,1273,1571,1597]" pageId="9" pageNumber="176">ON783074</accessionNumber>
; preserved in 95% ethanol;
<specimenCode id="DBD99EE1E63EFC1F0C61F9FACFCE3E38" box="[380,640,1606,1632]" collectionCode="IDSSE-EEB-SW" pageId="9" pageNumber="176">IDSSE-EEB-SW0251</specimenCode>
</materialsCitation>
<materialsCitation id="3B173CC7E63EFC1F0F8AF9FACE0C3EDC" ID-GBIF-Occurrence="4416297302" collectingDate="2021-09-20" collectionCode="IDSSE-EEB-SW" collectorName="Fendouzhe" elevation="7729" latitude="16.945667" location="PHILIPPINE SEA" longLatPrecision="13" longitude="129.73883" pageId="9" pageNumber="176" specimenCode="IDSSE-EEB-SW0252, IDSSE-EEB-SW0253, IDSSE-EEB-SW0254" specimenCount="3" typeStatus="paratype">
<specimenCount id="9D79FD13E63EFC1F0F8AF9FACE653E38" box="[663,811,1606,1632]" pageId="9" pageNumber="176" type="generic">3 specimens</specimenCount>
; same collection data as for holotype; preserved at -80°C;
<specimenCode id="DBD99EE1E63EFC1F0C0EF9D6CF583EDC" box="[275,534,1642,1668]" collectionCode="IDSSE-EEB-SW" pageId="9" pageNumber="176">IDSSE-EEB-SW0252</specimenCode>
to
<specimenCode id="DBD99EE1E63EFC1F0F26F9D6CE0C3EDC" box="[571,834,1642,1668]" collectionCode="IDSSE-EEB-SW" pageId="9" pageNumber="176">IDSSE-EEB-SW0254</specimenCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3656511E63EFC1D0DA0F916CEA33DAF" lastPageId="11" lastPageNumber="178" pageId="9" pageNumber="176" type="description">
<paragraph id="8BC0369AE63EFC1F0DA0F916CC003E9C" blockId="9.[189,334,1706,1732]" box="[189,334,1706,1732]" pageId="9" pageNumber="176">
<heading id="D08881F6E63EFC1F0DA0F916CC003E9C" bold="true" box="[189,334,1706,1732]" fontSize="11" level="3" pageId="9" pageNumber="176" reason="3">
<emphasis id="B90BEA88E63EFC1F0DA0F916CC003E9C" bold="true" box="[189,334,1706,1732]" pageId="9" pageNumber="176">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8BC0369AE63EFC1F0DA0F966C8553EAC" blockId="9.[189,1307,1754,1780]" box="[189,1307,1754,1780]" pageId="9" pageNumber="176">
<smallCapsWord id="8D26A046E63EFC1F0DA0F966CC223EAA" baselines="1773,1773" box="[189,364,1754,1780]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Measurements" pageId="9" pageNumber="176">MEASUREMENTS</smallCapsWord>
. Disc diameter
<quantity id="4C879B7FE63EFC1F0F3BF966CF323EAC" box="[550,636,1754,1780]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="9" pageNumber="176" unit="mm" value="10.0">10 mm</quantity>
, arm base width
<quantity id="4C879B7FE63EFC1F0E5FF966CED13EAC" box="[834,927,1754,1780]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="9" pageNumber="176" unit="mm" value="0.9">0.9 mm</quantity>
, and arm length
<quantity id="4C879B7FE63EFC1F097CF966C9FA3EAC" box="[1121,1204,1754,1780]" metricMagnitude="-2" metricUnit="m" metricValue="5.6" pageId="9" pageNumber="176" unit="mm" value="56.0">56 mm</quantity>
(
<figureCitation id="13442A1FE63EFC1F09D9F966C8453EAC" box="[1220,1291,1754,1780]" captionStart="Fig" captionStartId="10.[190,233,1750,1776]" captionTargetBox="[189,1399,444,1719]" captionTargetId="figure-68@10.[189,1399,444,1719]" captionTargetPageId="10" captionText="Fig. 3. Ophiuroglypha fendouzhe sp. nov., holotype (IDSSE-EEB-SW0250). A. Dorsal aspect. B. Ventral aspect. C. Central disc (overlapping disc scales). D. Dorsal arm base. E. Ventral disc, and mouth. F. Ventral arm base. G. Dorsal arm. HJ. Ventral arm (partially broken arm spine highlighted in the Fig. I). KL. Lateral arm. Abbreviations: arc = arm comb; ars = arm spine; as = adoral shield; asp = adoral shield spine; cpp = central primary plate; dap = dorsal arm plate; gs = genital slit; lap = lateral arm plate; m = madreporite; op = oral plate; os = oral shield; otp = oral tentacle pore; rs = radial shield; tp = tentacle pore; ts = tentacle scale; vap = ventral arm plate; vts = ventral tentacle scale. Scale bars: AB = 2 mm; C, EF =1 mm; D, GL = 500 μm." figureDoi="http://doi.org/10.5281/zenodo.8382623" httpUri="https://zenodo.org/record/8382623/files/figure.png" pageId="9" pageNumber="176">Fig. 3</figureCitation>
).
</paragraph>
<paragraph id="8BC0369AE63EFC1D0DA0F8A0C96C3983" blockId="9.[189,1399,1820,2025]" lastBlockId="11.[189,1399,266,475]" lastPageId="11" lastPageNumber="178" pageId="9" pageNumber="176">
<smallCapsWord id="8D26A046E63EFC1F0DA0F8A0CDBC3F6B" baselines="1839,1838" box="[189,242,1820,1846]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Disc" pageId="9" pageNumber="176">DISC</smallCapsWord>
. Disc pentagonal, covered by large irregular, and polygonal overlapping scales (
<figureCitation id="13442A1FE63EFC1F09FEF8A0C8243F6E" box="[1251,1386,1820,1846]" captionStart="Fig" captionStartId="10.[190,233,1750,1776]" captionTargetBox="[189,1399,444,1719]" captionTargetId="figure-68@10.[189,1399,444,1719]" captionTargetPageId="10" captionText="Fig. 3. Ophiuroglypha fendouzhe sp. nov., holotype (IDSSE-EEB-SW0250). A. Dorsal aspect. B. Ventral aspect. C. Central disc (overlapping disc scales). D. Dorsal arm base. E. Ventral disc, and mouth. F. Ventral arm base. G. Dorsal arm. HJ. Ventral arm (partially broken arm spine highlighted in the Fig. I). KL. Lateral arm. Abbreviations: arc = arm comb; ars = arm spine; as = adoral shield; asp = adoral shield spine; cpp = central primary plate; dap = dorsal arm plate; gs = genital slit; lap = lateral arm plate; m = madreporite; op = oral plate; os = oral shield; otp = oral tentacle pore; rs = radial shield; tp = tentacle pore; ts = tentacle scale; vap = ventral arm plate; vts = ventral tentacle scale. Scale bars: AB = 2 mm; C, EF =1 mm; D, GL = 500 μm." figureDoi="http://doi.org/10.5281/zenodo.8382623" httpUri="https://zenodo.org/record/8382623/files/figure.png" pageId="9" pageNumber="176">Fig. 3AC</figureCitation>
). Centrodorsal primary plate rounded in disc center (
<figureCitation id="13442A1FE63EFC1F0E3DF883CE303F01" box="[800,894,1855,1881]" captionStart="Fig" captionStartId="10.[190,233,1750,1776]" captionTargetBox="[189,1399,444,1719]" captionTargetId="figure-68@10.[189,1399,444,1719]" captionTargetPageId="10" captionText="Fig. 3. Ophiuroglypha fendouzhe sp. nov., holotype (IDSSE-EEB-SW0250). A. Dorsal aspect. B. Ventral aspect. C. Central disc (overlapping disc scales). D. Dorsal arm base. E. Ventral disc, and mouth. F. Ventral arm base. G. Dorsal arm. HJ. Ventral arm (partially broken arm spine highlighted in the Fig. I). KL. Lateral arm. Abbreviations: arc = arm comb; ars = arm spine; as = adoral shield; asp = adoral shield spine; cpp = central primary plate; dap = dorsal arm plate; gs = genital slit; lap = lateral arm plate; m = madreporite; op = oral plate; os = oral shield; otp = oral tentacle pore; rs = radial shield; tp = tentacle pore; ts = tentacle scale; vap = ventral arm plate; vts = ventral tentacle scale. Scale bars: AB = 2 mm; C, EF =1 mm; D, GL = 500 μm." figureDoi="http://doi.org/10.5281/zenodo.8382623" httpUri="https://zenodo.org/record/8382623/files/figure.png" pageId="9" pageNumber="176">Fig. 3C</figureCitation>
). Radial shields broad, slightly wider than long, triangular, barely connected to each other at distal end, sharp pointed proximal edge, straight distal edge, and proximal half separated by one elongated disc scale (k-plate) (
<figureCitation id="13442A1FE63EFC1F0904F83BC98F3FF9" box="[1049,1217,1927,1953]" captionStart="Fig" captionStartId="10.[190,233,1750,1776]" captionTargetBox="[189,1399,444,1719]" captionTargetId="figure-68@10.[189,1399,444,1719]" captionTargetPageId="10" captionText="Fig. 3. Ophiuroglypha fendouzhe sp. nov., holotype (IDSSE-EEB-SW0250). A. Dorsal aspect. B. Ventral aspect. C. Central disc (overlapping disc scales). D. Dorsal arm base. E. Ventral disc, and mouth. F. Ventral arm base. G. Dorsal arm. HJ. Ventral arm (partially broken arm spine highlighted in the Fig. I). KL. Lateral arm. Abbreviations: arc = arm comb; ars = arm spine; as = adoral shield; asp = adoral shield spine; cpp = central primary plate; dap = dorsal arm plate; gs = genital slit; lap = lateral arm plate; m = madreporite; op = oral plate; os = oral shield; otp = oral tentacle pore; rs = radial shield; tp = tentacle pore; ts = tentacle scale; vap = ventral arm plate; vts = ventral tentacle scale. Scale bars: AB = 2 mm; C, EF =1 mm; D, GL = 500 μm." figureDoi="http://doi.org/10.5281/zenodo.8382623" httpUri="https://zenodo.org/record/8382623/files/figure.png" pageId="9" pageNumber="176">Fig. 3A, CD</figureCitation>
). Median distal interradial area of disc formed by four to five large scales (⅓ larger than other dorsal disc scales), most of them rounded rectangular (
<figureCitation id="13442A1FE63EFC1F0CE9F873CF003FB1" box="[500,590,1999,2025]" captionStart="Fig" captionStartId="10.[190,233,1750,1776]" captionTargetBox="[189,1399,444,1719]" captionTargetId="figure-68@10.[189,1399,444,1719]" captionTargetPageId="10" captionText="Fig. 3. Ophiuroglypha fendouzhe sp. nov., holotype (IDSSE-EEB-SW0250). A. Dorsal aspect. B. Ventral aspect. C. Central disc (overlapping disc scales). D. Dorsal arm base. E. Ventral disc, and mouth. F. Ventral arm base. G. Dorsal arm. HJ. Ventral arm (partially broken arm spine highlighted in the Fig. I). KL. Lateral arm. Abbreviations: arc = arm comb; ars = arm spine; as = adoral shield; asp = adoral shield spine; cpp = central primary plate; dap = dorsal arm plate; gs = genital slit; lap = lateral arm plate; m = madreporite; op = oral plate; os = oral shield; otp = oral tentacle pore; rs = radial shield; tp = tentacle pore; ts = tentacle scale; vap = ventral arm plate; vts = ventral tentacle scale. Scale bars: AB = 2 mm; C, EF =1 mm; D, GL = 500 μm." figureDoi="http://doi.org/10.5281/zenodo.8382623" httpUri="https://zenodo.org/record/8382623/files/figure.png" pageId="9" pageNumber="176">Fig. 3C</figureCitation>
). Ventral disc covered by large, overlapping scales, slightly larger than on central dorsal disc (
<figureCitation id="13442A1FE63DFC1C0CDEFEB6CF11397C" box="[451,607,266,292]" captionStart="Fig" captionStartId="10.[190,233,1750,1776]" captionTargetBox="[189,1399,444,1719]" captionTargetId="figure-68@10.[189,1399,444,1719]" captionTargetPageId="10" captionText="Fig. 3. Ophiuroglypha fendouzhe sp. nov., holotype (IDSSE-EEB-SW0250). A. Dorsal aspect. B. Ventral aspect. C. Central disc (overlapping disc scales). D. Dorsal arm base. E. Ventral disc, and mouth. F. Ventral arm base. G. Dorsal arm. HJ. Ventral arm (partially broken arm spine highlighted in the Fig. I). KL. Lateral arm. Abbreviations: arc = arm comb; ars = arm spine; as = adoral shield; asp = adoral shield spine; cpp = central primary plate; dap = dorsal arm plate; gs = genital slit; lap = lateral arm plate; m = madreporite; op = oral plate; os = oral shield; otp = oral tentacle pore; rs = radial shield; tp = tentacle pore; ts = tentacle scale; vap = ventral arm plate; vts = ventral tentacle scale. Scale bars: AB = 2 mm; C, EF =1 mm; D, GL = 500 μm." figureDoi="http://doi.org/10.5281/zenodo.8382623" httpUri="https://zenodo.org/record/8382623/files/figure.png" pageId="10" pageNumber="177">Fig. 3B, EF</figureCitation>
). Distal end of genital slit bordered by small pointed genital papillae forming arm comb (
<figureCitation id="13442A1FE63DFC1C0CBBFE91CF6E391F" box="[422,544,301,327]" captionStart="Fig" captionStartId="10.[190,233,1750,1776]" captionTargetBox="[189,1399,444,1719]" captionTargetId="figure-68@10.[189,1399,444,1719]" captionTargetPageId="10" captionText="Fig. 3. Ophiuroglypha fendouzhe sp. nov., holotype (IDSSE-EEB-SW0250). A. Dorsal aspect. B. Ventral aspect. C. Central disc (overlapping disc scales). D. Dorsal arm base. E. Ventral disc, and mouth. F. Ventral arm base. G. Dorsal arm. HJ. Ventral arm (partially broken arm spine highlighted in the Fig. I). KL. Lateral arm. Abbreviations: arc = arm comb; ars = arm spine; as = adoral shield; asp = adoral shield spine; cpp = central primary plate; dap = dorsal arm plate; gs = genital slit; lap = lateral arm plate; m = madreporite; op = oral plate; os = oral shield; otp = oral tentacle pore; rs = radial shield; tp = tentacle pore; ts = tentacle scale; vap = ventral arm plate; vts = ventral tentacle scale. Scale bars: AB = 2 mm; C, EF =1 mm; D, GL = 500 μm." figureDoi="http://doi.org/10.5281/zenodo.8382623" httpUri="https://zenodo.org/record/8382623/files/figure.png" pageId="10" pageNumber="177">Fig. 3EF</figureCitation>
). Oral shield as wide as long, triangular, slightly convex at distal end, and connected to genital slit (
<figureCitation id="13442A1FE63DFC1C0CF7FEECCF093932" box="[490,583,336,362]" captionStart="Fig" captionStartId="10.[190,233,1750,1776]" captionTargetBox="[189,1399,444,1719]" captionTargetId="figure-68@10.[189,1399,444,1719]" captionTargetPageId="10" captionText="Fig. 3. Ophiuroglypha fendouzhe sp. nov., holotype (IDSSE-EEB-SW0250). A. Dorsal aspect. B. Ventral aspect. C. Central disc (overlapping disc scales). D. Dorsal arm base. E. Ventral disc, and mouth. F. Ventral arm base. G. Dorsal arm. HJ. Ventral arm (partially broken arm spine highlighted in the Fig. I). KL. Lateral arm. Abbreviations: arc = arm comb; ars = arm spine; as = adoral shield; asp = adoral shield spine; cpp = central primary plate; dap = dorsal arm plate; gs = genital slit; lap = lateral arm plate; m = madreporite; op = oral plate; os = oral shield; otp = oral tentacle pore; rs = radial shield; tp = tentacle pore; ts = tentacle scale; vap = ventral arm plate; vts = ventral tentacle scale. Scale bars: AB = 2 mm; C, EF =1 mm; D, GL = 500 μm." figureDoi="http://doi.org/10.5281/zenodo.8382623" httpUri="https://zenodo.org/record/8382623/files/figure.png" pageId="10" pageNumber="177">Fig. 3E</figureCitation>
). Adoral shields small, narrow, rectangular, slightly curving outward from oral shield, contiguous proximally, and distally connected to first lateral arm plate (
<figureCitation id="13442A1FE63DFC1C09CAFECFC87D39D5" box="[1239,1331,371,397]" captionStart="Fig" captionStartId="10.[190,233,1750,1776]" captionTargetBox="[189,1399,444,1719]" captionTargetId="figure-68@10.[189,1399,444,1719]" captionTargetPageId="10" captionText="Fig. 3. Ophiuroglypha fendouzhe sp. nov., holotype (IDSSE-EEB-SW0250). A. Dorsal aspect. B. Ventral aspect. C. Central disc (overlapping disc scales). D. Dorsal arm base. E. Ventral disc, and mouth. F. Ventral arm base. G. Dorsal arm. HJ. Ventral arm (partially broken arm spine highlighted in the Fig. I). KL. Lateral arm. Abbreviations: arc = arm comb; ars = arm spine; as = adoral shield; asp = adoral shield spine; cpp = central primary plate; dap = dorsal arm plate; gs = genital slit; lap = lateral arm plate; m = madreporite; op = oral plate; os = oral shield; otp = oral tentacle pore; rs = radial shield; tp = tentacle pore; ts = tentacle scale; vap = ventral arm plate; vts = ventral tentacle scale. Scale bars: AB = 2 mm; C, EF =1 mm; D, GL = 500 μm." figureDoi="http://doi.org/10.5281/zenodo.8382623" httpUri="https://zenodo.org/record/8382623/files/figure.png" pageId="10" pageNumber="177">Fig. 3E</figureCitation>
). Jaw slender, large, contiguous, with pair of pointed infradental papillae at apex, and spearhead-shaped teeth (
<figureCitation id="13442A1FE63CFC1D0DDBFE92CC6F3910" box="[198,289,302,328]" captionStart="Fig" captionStartId="10.[190,233,1750,1776]" captionTargetBox="[189,1399,444,1719]" captionTargetId="figure-68@10.[189,1399,444,1719]" captionTargetPageId="10" captionText="Fig. 3. Ophiuroglypha fendouzhe sp. nov., holotype (IDSSE-EEB-SW0250). A. Dorsal aspect. B. Ventral aspect. C. Central disc (overlapping disc scales). D. Dorsal arm base. E. Ventral disc, and mouth. F. Ventral arm base. G. Dorsal arm. HJ. Ventral arm (partially broken arm spine highlighted in the Fig. I). KL. Lateral arm. Abbreviations: arc = arm comb; ars = arm spine; as = adoral shield; asp = adoral shield spine; cpp = central primary plate; dap = dorsal arm plate; gs = genital slit; lap = lateral arm plate; m = madreporite; op = oral plate; os = oral shield; otp = oral tentacle pore; rs = radial shield; tp = tentacle pore; ts = tentacle scale; vap = ventral arm plate; vts = ventral tentacle scale. Scale bars: AB = 2 mm; C, EF =1 mm; D, GL = 500 μm." figureDoi="http://doi.org/10.5281/zenodo.8382623" httpUri="https://zenodo.org/record/8382623/files/figure.png" pageId="11" pageNumber="178">Fig. 3E</figureCitation>
). Up to five lateral oral papillae. Distalmost papilla (ventral compartment plate) much broader than others, and proximal two to three papillae spearhead-shaped to oval, and distal lateral oral papilla (buccal scale) lesspointed and broader (
<figureCitation id="13442A1FE63CFC1D0F81FEC4CFB739CA" box="[668,761,376,402]" captionStart="Fig" captionStartId="10.[190,233,1750,1776]" captionTargetBox="[189,1399,444,1719]" captionTargetId="figure-68@10.[189,1399,444,1719]" captionTargetPageId="10" captionText="Fig. 3. Ophiuroglypha fendouzhe sp. nov., holotype (IDSSE-EEB-SW0250). A. Dorsal aspect. B. Ventral aspect. C. Central disc (overlapping disc scales). D. Dorsal arm base. E. Ventral disc, and mouth. F. Ventral arm base. G. Dorsal arm. HJ. Ventral arm (partially broken arm spine highlighted in the Fig. I). KL. Lateral arm. Abbreviations: arc = arm comb; ars = arm spine; as = adoral shield; asp = adoral shield spine; cpp = central primary plate; dap = dorsal arm plate; gs = genital slit; lap = lateral arm plate; m = madreporite; op = oral plate; os = oral shield; otp = oral tentacle pore; rs = radial shield; tp = tentacle pore; ts = tentacle scale; vap = ventral arm plate; vts = ventral tentacle scale. Scale bars: AB = 2 mm; C, EF =1 mm; D, GL = 500 μm." figureDoi="http://doi.org/10.5281/zenodo.8382623" httpUri="https://zenodo.org/record/8382623/files/figure.png" pageId="11" pageNumber="178">Fig. 3E</figureCitation>
). Second oral tentacle pore large and opening outside mouth slit, with five ventral tentacle scales and four adoral shield spines at each side. Genital slits conspicuous and extending from oral shield to disc periphery (
<figureCitation id="13442A1FE63CFC1D0E87FE7DC95B3983" box="[922,1045,449,475]" captionStart="Fig" captionStartId="10.[190,233,1750,1776]" captionTargetBox="[189,1399,444,1719]" captionTargetId="figure-68@10.[189,1399,444,1719]" captionTargetPageId="10" captionText="Fig. 3. Ophiuroglypha fendouzhe sp. nov., holotype (IDSSE-EEB-SW0250). A. Dorsal aspect. B. Ventral aspect. C. Central disc (overlapping disc scales). D. Dorsal arm base. E. Ventral disc, and mouth. F. Ventral arm base. G. Dorsal arm. HJ. Ventral arm (partially broken arm spine highlighted in the Fig. I). KL. Lateral arm. Abbreviations: arc = arm comb; ars = arm spine; as = adoral shield; asp = adoral shield spine; cpp = central primary plate; dap = dorsal arm plate; gs = genital slit; lap = lateral arm plate; m = madreporite; op = oral plate; os = oral shield; otp = oral tentacle pore; rs = radial shield; tp = tentacle pore; ts = tentacle scale; vap = ventral arm plate; vts = ventral tentacle scale. Scale bars: AB = 2 mm; C, EF =1 mm; D, GL = 500 μm." figureDoi="http://doi.org/10.5281/zenodo.8382623" httpUri="https://zenodo.org/record/8382623/files/figure.png" pageId="11" pageNumber="178">Fig. 3EF</figureCitation>
).
</paragraph>
<caption id="DF006612E63DFC1C0DA3F96ACFE63FBF" ID-DOI="http://doi.org/10.5281/zenodo.8382623" ID-Zenodo-Dep="8382623" httpUri="https://zenodo.org/record/8382623/files/figure.png" pageId="10" pageNumber="177" startId="10.[190,233,1750,1776]" targetBox="[189,1399,444,1719]" targetPageId="10">
<paragraph id="8BC0369AE63DFC1C0DA3F96ACFE63FBF" blockId="10.[189,1399,1750,2023]" pageId="10" pageNumber="177">
<emphasis id="B90BEA88E63DFC1C0DA3F96ACC5B3EA8" bold="true" box="[190,277,1750,1776]" pageId="10" pageNumber="177">Fig. 3.</emphasis>
<taxonomicName id="4C7F4D19E63DFC1C0C3BF96ACF2F3EA8" authority="Nethupul &amp; Stöhr &amp; Zhang, 2023" authorityName="Nethupul &amp; Stöhr &amp; Zhang" authorityYear="2023" box="[294,609,1750,1776]" class="Ophiuroidea" family="Ophiopyrgidae" genus="Ophiuroglypha" kingdom="Animalia" order="Ophiurida" pageId="10" pageNumber="177" phylum="Echinodermata" rank="species" species="fendouzhe" status="sp. nov.">
<emphasis id="B90BEA88E63DFC1C0C3BF96ACF2F3EA8" box="[294,609,1750,1776]" italics="true" pageId="10" pageNumber="177">Ophiuroglypha fendouzhe</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23857F3E63DFC1C0F6FF96BCF943EA9" box="[626,730,1751,1777]" pageId="10" pageNumber="177" rank="species">sp. nov.</taxonomicNameLabel>
, holotype (IDSSE-EEB-SW0250).
<emphasis id="B90BEA88E63DFC1C0985F96AC9E33EA8" bold="true" box="[1176,1197,1750,1776]" pageId="10" pageNumber="177">A</emphasis>
. Dorsal aspect.
<emphasis id="B90BEA88E63DFC1C0DA0F946CD9F3F4C" bold="true" box="[189,209,1786,1812]" pageId="10" pageNumber="177">B</emphasis>
. Ventral aspect.
<emphasis id="B90BEA88E63DFC1C0C84F945CCE03F4B" bold="true" box="[409,430,1785,1811]" pageId="10" pageNumber="177">C</emphasis>
. Central disc (overlapping disc scales).
<emphasis id="B90BEA88E63DFC1C0E8DF946CEEB3F4C" bold="true" box="[912,933,1786,1812]" pageId="10" pageNumber="177">D</emphasis>
. Dorsal arm base.
<emphasis id="B90BEA88E63DFC1C0997F946C9D03F4C" bold="true" box="[1162,1182,1786,1812]" pageId="10" pageNumber="177">E</emphasis>
. Ventral disc, and mouth.
<emphasis id="B90BEA88E63DFC1C0C0BF8A1CC663F6F" bold="true" box="[278,296,1821,1847]" pageId="10" pageNumber="177">F</emphasis>
. Ventral arm base.
<emphasis id="B90BEA88E63DFC1C0F15F8A1CF513F6F" bold="true" box="[520,543,1821,1847]" pageId="10" pageNumber="177">G</emphasis>
. Dorsal arm.
<emphasis id="B90BEA88E63DFC1C0FA0F8A1CFBF3F6F" bold="true" box="[701,753,1821,1847]" pageId="10" pageNumber="177">HJ</emphasis>
. Ventral arm (partially broken arm spine highlighted in the Fig. I).
<emphasis id="B90BEA88E63DFC1C0C22F8FCCC363F02" bold="true" box="[319,376,1856,1882]" pageId="10" pageNumber="177">KL</emphasis>
. Lateral arm. Abbreviations: arc = arm comb; ars = arm spine; as = adoral shield; asp = adoral shield spine; cpp = central primary plate; dap = dorsal arm plate; gs = genital slit; lap = lateral arm plate; m = madreporite; op = oral plate; os = oral shield; otp = oral tentacle pore; rs = radial shield; tp = tentacle pore; ts = tentacle scale; vap = ventral arm plate; vts = ventral tentacle scale. Scale bars: AB = 2 mm; C, EF =1 mm; D, GL = 500 μm.
</paragraph>
</caption>
<paragraph id="8BC0369AE63CFC1D0DA0FDB8CEE43BD6" blockId="11.[189,1399,516,910]" pageId="11" pageNumber="178">
<smallCapsWord id="8D26A046E63CFC1D0DA0FDB8CDB33A44" baselines="535,535" box="[189,253,516,542]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Arms" pageId="11" pageNumber="178">ARMS</smallCapsWord>
. Five slender, and non-moniliform arms. Dorsal arm plates small, diamond-shaped, straight to slightly fan-shaped with convex distal edge, pointed proximal edge, as wide as long, and widely separated, except on first and second arm segments (
<figureCitation id="13442A1FE63CFC1D0E2CFDF2CEF93A30" box="[817,951,590,616]" captionStart="Fig" captionStartId="10.[190,233,1750,1776]" captionTargetBox="[189,1399,444,1719]" captionTargetId="figure-68@10.[189,1399,444,1719]" captionTargetPageId="10" captionText="Fig. 3. Ophiuroglypha fendouzhe sp. nov., holotype (IDSSE-EEB-SW0250). A. Dorsal aspect. B. Ventral aspect. C. Central disc (overlapping disc scales). D. Dorsal arm base. E. Ventral disc, and mouth. F. Ventral arm base. G. Dorsal arm. HJ. Ventral arm (partially broken arm spine highlighted in the Fig. I). KL. Lateral arm. Abbreviations: arc = arm comb; ars = arm spine; as = adoral shield; asp = adoral shield spine; cpp = central primary plate; dap = dorsal arm plate; gs = genital slit; lap = lateral arm plate; m = madreporite; op = oral plate; os = oral shield; otp = oral tentacle pore; rs = radial shield; tp = tentacle pore; ts = tentacle scale; vap = ventral arm plate; vts = ventral tentacle scale. Scale bars: AB = 2 mm; C, EF =1 mm; D, GL = 500 μm." figureDoi="http://doi.org/10.5281/zenodo.8382623" httpUri="https://zenodo.org/record/8382623/files/figure.png" pageId="11" pageNumber="178">Fig. 3D, G</figureCitation>
). First two ventral arm plates slightly wider than long, pentagonal, and contiguous. Following ventral arm plates wider than long, pentagonal to triangular, distally straight to slightly curved, proximally triangular, and increasingly separated along arm (
<figureCitation id="13442A1FE63CFC1D0DE7FD00CCDD3A8E" box="[250,403,700,726]" captionStart="Fig" captionStartId="10.[190,233,1750,1776]" captionTargetBox="[189,1399,444,1719]" captionTargetId="figure-68@10.[189,1399,444,1719]" captionTargetPageId="10" captionText="Fig. 3. Ophiuroglypha fendouzhe sp. nov., holotype (IDSSE-EEB-SW0250). A. Dorsal aspect. B. Ventral aspect. C. Central disc (overlapping disc scales). D. Dorsal arm base. E. Ventral disc, and mouth. F. Ventral arm base. G. Dorsal arm. HJ. Ventral arm (partially broken arm spine highlighted in the Fig. I). KL. Lateral arm. Abbreviations: arc = arm comb; ars = arm spine; as = adoral shield; asp = adoral shield spine; cpp = central primary plate; dap = dorsal arm plate; gs = genital slit; lap = lateral arm plate; m = madreporite; op = oral plate; os = oral shield; otp = oral tentacle pore; rs = radial shield; tp = tentacle pore; ts = tentacle scale; vap = ventral arm plate; vts = ventral tentacle scale. Scale bars: AB = 2 mm; C, EF =1 mm; D, GL = 500 μm." figureDoi="http://doi.org/10.5281/zenodo.8382623" httpUri="https://zenodo.org/record/8382623/files/figure.png" pageId="11" pageNumber="178">Fig. 3EF, H</figureCitation>
). Lateral arm plate with mostly three and rarely four arm spines. All arm spines small, smooth, pointed tip, similar in length, and approximately ½ arm segment long (
<figureCitation id="13442A1FE63CFC1D0961FD5DC9B03AA3" box="[1148,1278,737,763]" captionStart="Fig" captionStartId="10.[190,233,1750,1776]" captionTargetBox="[189,1399,444,1719]" captionTargetId="figure-68@10.[189,1399,444,1719]" captionTargetPageId="10" captionText="Fig. 3. Ophiuroglypha fendouzhe sp. nov., holotype (IDSSE-EEB-SW0250). A. Dorsal aspect. B. Ventral aspect. C. Central disc (overlapping disc scales). D. Dorsal arm base. E. Ventral disc, and mouth. F. Ventral arm base. G. Dorsal arm. HJ. Ventral arm (partially broken arm spine highlighted in the Fig. I). KL. Lateral arm. Abbreviations: arc = arm comb; ars = arm spine; as = adoral shield; asp = adoral shield spine; cpp = central primary plate; dap = dorsal arm plate; gs = genital slit; lap = lateral arm plate; m = madreporite; op = oral plate; os = oral shield; otp = oral tentacle pore; rs = radial shield; tp = tentacle pore; ts = tentacle scale; vap = ventral arm plate; vts = ventral tentacle scale. Scale bars: AB = 2 mm; C, EF =1 mm; D, GL = 500 μm." figureDoi="http://doi.org/10.5281/zenodo.8382623" httpUri="https://zenodo.org/record/8382623/files/figure.png" pageId="11" pageNumber="178">Fig. 3GL</figureCitation>
). Distally, middle arm spine slightly curved inwards (
<figureCitation id="13442A1FE63CFC1D0FDBFCBACE733B78" box="[710,829,774,800]" captionStart="Fig" captionStartId="10.[190,233,1750,1776]" captionTargetBox="[189,1399,444,1719]" captionTargetId="figure-68@10.[189,1399,444,1719]" captionTargetPageId="10" captionText="Fig. 3. Ophiuroglypha fendouzhe sp. nov., holotype (IDSSE-EEB-SW0250). A. Dorsal aspect. B. Ventral aspect. C. Central disc (overlapping disc scales). D. Dorsal arm base. E. Ventral disc, and mouth. F. Ventral arm base. G. Dorsal arm. HJ. Ventral arm (partially broken arm spine highlighted in the Fig. I). KL. Lateral arm. Abbreviations: arc = arm comb; ars = arm spine; as = adoral shield; asp = adoral shield spine; cpp = central primary plate; dap = dorsal arm plate; gs = genital slit; lap = lateral arm plate; m = madreporite; op = oral plate; os = oral shield; otp = oral tentacle pore; rs = radial shield; tp = tentacle pore; ts = tentacle scale; vap = ventral arm plate; vts = ventral tentacle scale. Scale bars: AB = 2 mm; C, EF =1 mm; D, GL = 500 μm." figureDoi="http://doi.org/10.5281/zenodo.8382623" httpUri="https://zenodo.org/record/8382623/files/figure.png" pageId="11" pageNumber="178">Fig. 3IL</figureCitation>
). First tentacle pore with three or four tentacle scales (
<figureCitation id="13442A1FE63CFC1D0C0EFC97CCC13B1D" box="[275,399,811,837]" captionStart="Fig" captionStartId="10.[190,233,1750,1776]" captionTargetBox="[189,1399,444,1719]" captionTargetId="figure-68@10.[189,1399,444,1719]" captionTargetPageId="10" captionText="Fig. 3. Ophiuroglypha fendouzhe sp. nov., holotype (IDSSE-EEB-SW0250). A. Dorsal aspect. B. Ventral aspect. C. Central disc (overlapping disc scales). D. Dorsal arm base. E. Ventral disc, and mouth. F. Ventral arm base. G. Dorsal arm. HJ. Ventral arm (partially broken arm spine highlighted in the Fig. I). KL. Lateral arm. Abbreviations: arc = arm comb; ars = arm spine; as = adoral shield; asp = adoral shield spine; cpp = central primary plate; dap = dorsal arm plate; gs = genital slit; lap = lateral arm plate; m = madreporite; op = oral plate; os = oral shield; otp = oral tentacle pore; rs = radial shield; tp = tentacle pore; ts = tentacle scale; vap = ventral arm plate; vts = ventral tentacle scale. Scale bars: AB = 2 mm; C, EF =1 mm; D, GL = 500 μm." figureDoi="http://doi.org/10.5281/zenodo.8382623" httpUri="https://zenodo.org/record/8382623/files/figure.png" pageId="11" pageNumber="178">Fig. 3FL</figureCitation>
). Second and third tentacle pores with four to two tentacle scales (
<figureCitation id="13442A1FE63CFC1D0986FC97C9BB3B1D" box="[1179,1269,811,837]" captionStart="Fig" captionStartId="10.[190,233,1750,1776]" captionTargetBox="[189,1399,444,1719]" captionTargetId="figure-68@10.[189,1399,444,1719]" captionTargetPageId="10" captionText="Fig. 3. Ophiuroglypha fendouzhe sp. nov., holotype (IDSSE-EEB-SW0250). A. Dorsal aspect. B. Ventral aspect. C. Central disc (overlapping disc scales). D. Dorsal arm base. E. Ventral disc, and mouth. F. Ventral arm base. G. Dorsal arm. HJ. Ventral arm (partially broken arm spine highlighted in the Fig. I). KL. Lateral arm. Abbreviations: arc = arm comb; ars = arm spine; as = adoral shield; asp = adoral shield spine; cpp = central primary plate; dap = dorsal arm plate; gs = genital slit; lap = lateral arm plate; m = madreporite; op = oral plate; os = oral shield; otp = oral tentacle pore; rs = radial shield; tp = tentacle pore; ts = tentacle scale; vap = ventral arm plate; vts = ventral tentacle scale. Scale bars: AB = 2 mm; C, EF =1 mm; D, GL = 500 μm." figureDoi="http://doi.org/10.5281/zenodo.8382623" httpUri="https://zenodo.org/record/8382623/files/figure.png" pageId="11" pageNumber="178">Fig. 3F</figureCitation>
). Fourth to fifth tentacle pores with three to one tentacle scales (
<figureCitation id="13442A1FE63CFC1D0E07FCF3CE3C3B31" box="[794,882,847,873]" captionStart="Fig" captionStartId="10.[190,233,1750,1776]" captionTargetBox="[189,1399,444,1719]" captionTargetId="figure-68@10.[189,1399,444,1719]" captionTargetPageId="10" captionText="Fig. 3. Ophiuroglypha fendouzhe sp. nov., holotype (IDSSE-EEB-SW0250). A. Dorsal aspect. B. Ventral aspect. C. Central disc (overlapping disc scales). D. Dorsal arm base. E. Ventral disc, and mouth. F. Ventral arm base. G. Dorsal arm. HJ. Ventral arm (partially broken arm spine highlighted in the Fig. I). KL. Lateral arm. Abbreviations: arc = arm comb; ars = arm spine; as = adoral shield; asp = adoral shield spine; cpp = central primary plate; dap = dorsal arm plate; gs = genital slit; lap = lateral arm plate; m = madreporite; op = oral plate; os = oral shield; otp = oral tentacle pore; rs = radial shield; tp = tentacle pore; ts = tentacle scale; vap = ventral arm plate; vts = ventral tentacle scale. Scale bars: AB = 2 mm; C, EF =1 mm; D, GL = 500 μm." figureDoi="http://doi.org/10.5281/zenodo.8382623" httpUri="https://zenodo.org/record/8382623/files/figure.png" pageId="11" pageNumber="178">Fig. 3F</figureCitation>
). Number of tentacle scales decreases to one scale, and absent after seven or nine arm segments (
<figureCitation id="13442A1FE63CFC1D0E00FCC8CED33BD6" box="[797,925,884,910]" captionStart="Fig" captionStartId="10.[190,233,1750,1776]" captionTargetBox="[189,1399,444,1719]" captionTargetId="figure-68@10.[189,1399,444,1719]" captionTargetPageId="10" captionText="Fig. 3. Ophiuroglypha fendouzhe sp. nov., holotype (IDSSE-EEB-SW0250). A. Dorsal aspect. B. Ventral aspect. C. Central disc (overlapping disc scales). D. Dorsal arm base. E. Ventral disc, and mouth. F. Ventral arm base. G. Dorsal arm. HJ. Ventral arm (partially broken arm spine highlighted in the Fig. I). KL. Lateral arm. Abbreviations: arc = arm comb; ars = arm spine; as = adoral shield; asp = adoral shield spine; cpp = central primary plate; dap = dorsal arm plate; gs = genital slit; lap = lateral arm plate; m = madreporite; op = oral plate; os = oral shield; otp = oral tentacle pore; rs = radial shield; tp = tentacle pore; ts = tentacle scale; vap = ventral arm plate; vts = ventral tentacle scale. Scale bars: AB = 2 mm; C, EF =1 mm; D, GL = 500 μm." figureDoi="http://doi.org/10.5281/zenodo.8382623" httpUri="https://zenodo.org/record/8382623/files/figure.png" pageId="11" pageNumber="178">Fig. 3H, L</figureCitation>
).
</paragraph>
<paragraph id="8BC0369AE63CFC1D0DA0FC0BCE793B89" blockId="11.[189,823,951,977]" box="[189,823,951,977]" pageId="11" pageNumber="178">
<smallCapsWord id="8D26A046E63CFC1D0DA0FC0BCC473B97" baselines="970,970" box="[189,265,951,977]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Color" pageId="11" pageNumber="178">COLOR</smallCapsWord>
. Creamy white in alcohol specimen (
<figureCitation id="13442A1FE63CFC1D0FA3FC0BCE473B89" box="[702,777,951,977]" captionStart="Fig" captionStartId="10.[190,233,1750,1776]" captionTargetBox="[189,1399,444,1719]" captionTargetId="figure-68@10.[189,1399,444,1719]" captionTargetPageId="10" captionText="Fig. 3. Ophiuroglypha fendouzhe sp. nov., holotype (IDSSE-EEB-SW0250). A. Dorsal aspect. B. Ventral aspect. C. Central disc (overlapping disc scales). D. Dorsal arm base. E. Ventral disc, and mouth. F. Ventral arm base. G. Dorsal arm. HJ. Ventral arm (partially broken arm spine highlighted in the Fig. I). KL. Lateral arm. Abbreviations: arc = arm comb; ars = arm spine; as = adoral shield; asp = adoral shield spine; cpp = central primary plate; dap = dorsal arm plate; gs = genital slit; lap = lateral arm plate; m = madreporite; op = oral plate; os = oral shield; otp = oral tentacle pore; rs = radial shield; tp = tentacle pore; ts = tentacle scale; vap = ventral arm plate; vts = ventral tentacle scale. Scale bars: AB = 2 mm; C, EF =1 mm; D, GL = 500 μm." figureDoi="http://doi.org/10.5281/zenodo.8382623" httpUri="https://zenodo.org/record/8382623/files/figure.png" pageId="11" pageNumber="178">Figs 3</figureCitation>
,
<figureCitation id="13442A1FE63CFC1D0E0AFC0BCE663B89" box="[791,808,951,977]" captionStart="Fig" captionStartId="13.[189,232,945,971]" captionTargetBox="[189,1399,265,895]" captionTargetId="figure-427@13.[189,1399,265,895]" captionTargetPageId="13" captionText="Fig. 5. Ophiuroglypha fendouzhe sp. nov., paratype (IDSSE-EEB-SW0251). A. Dorsal aspect. B. Ventral aspect. C. Central disc (overlapping disc scales). D. Ventral arm. E. Dorsal arm. F. Lateral arm. Abbreviations: ars = arm spine; as = adoral shield; asp = adoral shield spine; cpp = central primary plate; dap = dorsal arm plate; gs = genital slit; lap = lateral arm plate; m = madreporite; op = oral plate; os = oral shield; otp = oral tentacle pore; tp = tentacle pore; ts = tentacle scale; vap = ventral arm plate; vts = ventral tentacle scale. Scale bars: AB = 2 mm; C, EF = 500 μm; D = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.8382627" httpUri="https://zenodo.org/record/8382627/files/figure.png" pageId="11" pageNumber="178">5</figureCitation>
).
</paragraph>
<paragraph id="8BC0369AE63CFC1D0DA0FC46C8183D4E" blockId="11.[189,1399,1018,1302]" pageId="11" pageNumber="178">
<smallCapsWord id="8D26A046E63CFC1D0DA0FC46CC583C4A" baselines="1037,1037" box="[189,278,1018,1044]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Ossicle" pageId="11" pageNumber="178">OSSICLE</smallCapsWord>
<smallCapsWord id="8D26A046E63CFC1D0C3FFBBCCCFD3C4A" baselines="1037" box="[290,435,1024,1042]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="morphology" pageId="11" pageNumber="178">MORPHOLOGY</smallCapsWord>
. (IDSSE-EEB-SW0251) Three arm spine articulations placed at distal edge of lateral arm plate, with clear muscle opening and small nerve opening (
<figureCitation id="13442A1FE63CFC1D0939FBA3C9E73C61" box="[1060,1193,1055,1081]" captionStart="Fig" captionStartId="12.[189,232,1807,1833]" captionTargetBox="[189,1399,265,1760]" captionTargetId="figure-101@12.[189,1399,265,1760]" captionTargetPageId="12" captionText="Fig. 4. Ophiuroglypha fendouzhe sp. nov., paratype (IDSSE-EEB-SW0251). AC. Lateral arm plate. DH. Vertebrae. D. Distal view (right side of the structure partially broken). E. Proximal view. F. Ventral view. G. Dorsal view. H. Dorsolateral view. Abbreviations: asa = arm spine articulation; d = dorsal; dist = distal; kn = knob; mo = muscle opening; no = nerve opening; pb = podial basin; prox = proximal; tfp = tube foot pore; v = ventral. Scale bars: A, C, FH = 500 μm; B = 100 μm; DE = 300 μm." figureDoi="http://doi.org/10.5281/zenodo.8382625" httpUri="https://zenodo.org/record/8382625/files/figure.png" pageId="11" pageNumber="178">Fig. 4AB</figureCitation>
). Tube foot pore developed as large within-plate perforation on lateral arm plates from base to middle part of arm, where tentacle scales begin to disappear from arm segments (
<figureCitation id="13442A1FE63CFC1D0E58FBD5CE863CDB" box="[837,968,1129,1155]" captionStart="Fig" captionStartId="12.[189,232,1807,1833]" captionTargetBox="[189,1399,265,1760]" captionTargetId="figure-101@12.[189,1399,265,1760]" captionTargetPageId="12" captionText="Fig. 4. Ophiuroglypha fendouzhe sp. nov., paratype (IDSSE-EEB-SW0251). AC. Lateral arm plate. DH. Vertebrae. D. Distal view (right side of the structure partially broken). E. Proximal view. F. Ventral view. G. Dorsal view. H. Dorsolateral view. Abbreviations: asa = arm spine articulation; d = dorsal; dist = distal; kn = knob; mo = muscle opening; no = nerve opening; pb = podial basin; prox = proximal; tfp = tube foot pore; v = ventral. Scale bars: A, C, FH = 500 μm; B = 100 μm; DE = 300 μm." figureDoi="http://doi.org/10.5281/zenodo.8382625" httpUri="https://zenodo.org/record/8382625/files/figure.png" pageId="11" pageNumber="178">Fig. 4AB</figureCitation>
). Middle half of inner side of lateral arm plate with depression and two round knobs (
<figureCitation id="13442A1FE63CFC1D0E0CFB31CE3F3CFF" box="[785,881,1165,1191]" captionStart="Fig" captionStartId="12.[189,232,1807,1833]" captionTargetBox="[189,1399,265,1760]" captionTargetId="figure-101@12.[189,1399,265,1760]" captionTargetPageId="12" captionText="Fig. 4. Ophiuroglypha fendouzhe sp. nov., paratype (IDSSE-EEB-SW0251). AC. Lateral arm plate. DH. Vertebrae. D. Distal view (right side of the structure partially broken). E. Proximal view. F. Ventral view. G. Dorsal view. H. Dorsolateral view. Abbreviations: asa = arm spine articulation; d = dorsal; dist = distal; kn = knob; mo = muscle opening; no = nerve opening; pb = podial basin; prox = proximal; tfp = tube foot pore; v = ventral. Scale bars: A, C, FH = 500 μm; B = 100 μm; DE = 300 μm." figureDoi="http://doi.org/10.5281/zenodo.8382625" httpUri="https://zenodo.org/record/8382625/files/figure.png" pageId="11" pageNumber="178">Fig. 4C</figureCitation>
). Vertebrae with moderate zygospondylous articulation, proximal end with podial basins (
<figureCitation id="13442A1FE63CFC1D0FF9FB0ECE273C94" box="[740,873,1202,1228]" captionStart="Fig" captionStartId="12.[189,232,1807,1833]" captionTargetBox="[189,1399,265,1760]" captionTargetId="figure-101@12.[189,1399,265,1760]" captionTargetPageId="12" captionText="Fig. 4. Ophiuroglypha fendouzhe sp. nov., paratype (IDSSE-EEB-SW0251). AC. Lateral arm plate. DH. Vertebrae. D. Distal view (right side of the structure partially broken). E. Proximal view. F. Ventral view. G. Dorsal view. H. Dorsolateral view. Abbreviations: asa = arm spine articulation; d = dorsal; dist = distal; kn = knob; mo = muscle opening; no = nerve opening; pb = podial basin; prox = proximal; tfp = tube foot pore; v = ventral. Scale bars: A, C, FH = 500 μm; B = 100 μm; DE = 300 μm." figureDoi="http://doi.org/10.5281/zenodo.8382625" httpUri="https://zenodo.org/record/8382625/files/figure.png" pageId="11" pageNumber="178">Fig. 4DH</figureCitation>
). Dorsal side of vertebrae distally triangular and proximally slightly curved, shallow longitudinal groove along midline (
<figureCitation id="13442A1FE63CFC1D0924FB6BC9F63CA9" box="[1081,1208,1239,1265]" captionStart="Fig" captionStartId="12.[189,232,1807,1833]" captionTargetBox="[189,1399,265,1760]" captionTargetId="figure-101@12.[189,1399,265,1760]" captionTargetPageId="12" captionText="Fig. 4. Ophiuroglypha fendouzhe sp. nov., paratype (IDSSE-EEB-SW0251). AC. Lateral arm plate. DH. Vertebrae. D. Distal view (right side of the structure partially broken). E. Proximal view. F. Ventral view. G. Dorsal view. H. Dorsolateral view. Abbreviations: asa = arm spine articulation; d = dorsal; dist = distal; kn = knob; mo = muscle opening; no = nerve opening; pb = podial basin; prox = proximal; tfp = tube foot pore; v = ventral. Scale bars: A, C, FH = 500 μm; B = 100 μm; DE = 300 μm." figureDoi="http://doi.org/10.5281/zenodo.8382625" httpUri="https://zenodo.org/record/8382625/files/figure.png" pageId="11" pageNumber="178">Fig. 4FH</figureCitation>
). Ventral side of vertebrae with long and broad ambulacral groove with pair of lateral ambulacral canals (
<figureCitation id="13442A1FE63CFC1D09D6FB40C8043D4E" box="[1227,1354,1276,1302]" captionStart="Fig" captionStartId="12.[189,232,1807,1833]" captionTargetBox="[189,1399,265,1760]" captionTargetId="figure-101@12.[189,1399,265,1760]" captionTargetPageId="12" captionText="Fig. 4. Ophiuroglypha fendouzhe sp. nov., paratype (IDSSE-EEB-SW0251). AC. Lateral arm plate. DH. Vertebrae. D. Distal view (right side of the structure partially broken). E. Proximal view. F. Ventral view. G. Dorsal view. H. Dorsolateral view. Abbreviations: asa = arm spine articulation; d = dorsal; dist = distal; kn = knob; mo = muscle opening; no = nerve opening; pb = podial basin; prox = proximal; tfp = tube foot pore; v = ventral. Scale bars: A, C, FH = 500 μm; B = 100 μm; DE = 300 μm." figureDoi="http://doi.org/10.5281/zenodo.8382625" httpUri="https://zenodo.org/record/8382625/files/figure.png" pageId="11" pageNumber="178">Fig. 4FH</figureCitation>
).
</paragraph>
<paragraph id="8BC0369AE63CFC1D0DA0FA82CCFD3D00" blockId="11.[189,435,1342,1368]" box="[189,435,1342,1368]" pageId="11" pageNumber="178">
<heading id="D08881F6E63CFC1D0DA0FA82CCFD3D00" bold="true" box="[189,435,1342,1368]" fontSize="11" level="3" pageId="11" pageNumber="178" reason="3">
<emphasis id="B90BEA88E63CFC1D0DA0FA82CCFD3D00" bold="true" box="[189,435,1342,1368]" pageId="11" pageNumber="178">
<typeStatus id="54C48838E63CFC1D0DA0FA82CC613D00" box="[189,303,1342,1368]" pageId="11" pageNumber="178" type="paratype">Paratype</typeStatus>
variations
</emphasis>
</heading>
</paragraph>
<paragraph id="8BC0369AE63CFC1D0DA0FAD3CEA33DAF" blockId="11.[189,1399,1391,1527]" pageId="11" pageNumber="178">
Four
<typeStatus id="54C48838E63CFC1D0DE1FAD3CC2F3DD1" box="[252,353,1391,1417]" pageId="11" pageNumber="178" type="paratype">paratype</typeStatus>
specimens were collected from the same location. They are similar to the
<typeStatus id="54C48838E63CFC1D09D2FAD3C8783DD1" box="[1231,1334,1391,1417]" pageId="11" pageNumber="178" type="holotype">holotype</typeStatus>
(disc diameters 8.010.0 mm), but
<specimenCount id="9D79FD13E63CFC1D0F08FA28CFE53DF6" box="[533,683,1428,1454]" pageId="11" pageNumber="178" type="generic" typeStatus="paratype">one paratype</specimenCount>
has a sub-pentagonal disc and an obvious round centrodorsal primary plate (
<figureCitation id="13442A1FE63CFC1D0C78FA04CC8C3D8A" box="[357,450,1464,1490]" captionStart="Fig" captionStartId="13.[189,232,945,971]" captionTargetBox="[189,1399,265,895]" captionTargetId="figure-427@13.[189,1399,265,895]" captionTargetPageId="13" captionText="Fig. 5. Ophiuroglypha fendouzhe sp. nov., paratype (IDSSE-EEB-SW0251). A. Dorsal aspect. B. Ventral aspect. C. Central disc (overlapping disc scales). D. Ventral arm. E. Dorsal arm. F. Lateral arm. Abbreviations: ars = arm spine; as = adoral shield; asp = adoral shield spine; cpp = central primary plate; dap = dorsal arm plate; gs = genital slit; lap = lateral arm plate; m = madreporite; op = oral plate; os = oral shield; otp = oral tentacle pore; tp = tentacle pore; ts = tentacle scale; vap = ventral arm plate; vts = ventral tentacle scale. Scale bars: AB = 2 mm; C, EF = 500 μm; D = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.8382627" httpUri="https://zenodo.org/record/8382627/files/figure.png" pageId="11" pageNumber="178">Fig. 5A</figureCitation>
).All
<typeStatus id="54C48838E63CFC1D0CE0FA04CF233D8A" box="[509,621,1464,1490]" pageId="11" pageNumber="178" type="paratype">paratypes</typeStatus>
are identical to the
<typeStatus id="54C48838E63CFC1D0E54FA04CEFE3D8A" box="[841,944,1464,1490]" pageId="11" pageNumber="178" type="holotype">holotype</typeStatus>
with respect to the other morphological characters and this small variation is considered intraspecific (
<figureCitation id="13442A1FE63CFC1D0E8BFA61CE903DAF" box="[918,990,1501,1527]" captionStart="Fig" captionStartId="13.[189,232,945,971]" captionTargetBox="[189,1399,265,895]" captionTargetId="figure-427@13.[189,1399,265,895]" captionTargetPageId="13" captionText="Fig. 5. Ophiuroglypha fendouzhe sp. nov., paratype (IDSSE-EEB-SW0251). A. Dorsal aspect. B. Ventral aspect. C. Central disc (overlapping disc scales). D. Ventral arm. E. Dorsal arm. F. Lateral arm. Abbreviations: ars = arm spine; as = adoral shield; asp = adoral shield spine; cpp = central primary plate; dap = dorsal arm plate; gs = genital slit; lap = lateral arm plate; m = madreporite; op = oral plate; os = oral shield; otp = oral tentacle pore; tp = tentacle pore; ts = tentacle scale; vap = ventral arm plate; vts = ventral tentacle scale. Scale bars: AB = 2 mm; C, EF = 500 μm; D = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.8382627" httpUri="https://zenodo.org/record/8382627/files/figure.png" pageId="11" pageNumber="178">Fig. 5</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3656511E63CFC1D0DA0F9A3CFEF3E32" pageId="11" pageNumber="178" type="distribution">
<paragraph id="8BC0369AE63CFC1D0DA0F9A3CCA03E61" blockId="11.[189,494,1567,1594]" box="[189,494,1567,1594]" pageId="11" pageNumber="178">
<heading id="D08881F6E63CFC1D0DA0F9A3CCA03E61" bold="true" box="[189,494,1567,1594]" fontSize="11" level="3" pageId="11" pageNumber="178" reason="3">
<emphasis id="B90BEA88E63CFC1D0DA0F9A3CCA03E61" bold="true" box="[189,494,1567,1594]" pageId="11" pageNumber="178">Distribution and habitat</emphasis>
</heading>
</paragraph>
<paragraph id="8BC0369AE63CFC1D0DA0F9ECCFEF3E32" blockId="11.[189,673,1616,1642]" box="[189,673,1616,1642]" pageId="11" pageNumber="178">
So far only known from the
<typeStatus id="54C48838E63CFC1D0F14F9ECCF723E32" box="[521,572,1616,1642]" pageId="11" pageNumber="178">type</typeStatus>
locality.
</paragraph>
</subSubSection>
</treatment>
</document>