906 lines
119 KiB
XML
906 lines
119 KiB
XML
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<mods:title id="F9C28F3315EC2EC9A09EF1AAD7ABBB36">An integrative taxonomic review of the South Asian microhylid genus Uperodon</mods:title>
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<mods:namePart id="C4BD49D17A11DC8635C962A3161E7B62">Garg, Sonali</mods:namePart>
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<mods:namePart id="730CF9372AB8E8F9A91F81FB90DEA354">Senevirathne, Gayani</mods:namePart>
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<mods:namePart id="69DA258D523F8B5E04D75BCC86C62E31">Wijayathilaka, Nayana</mods:namePart>
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<mods:namePart id="CD79B021A315AE3A21D1738D47586A4E">Deuti, Kaushik</mods:namePart>
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<treatment id="03D4A4166C149460FF5ACDB6FB48F858" ID-DOI="http://doi.org/10.5281/zenodo.5587723" ID-GBIF-Taxon="141119107" ID-Zenodo-Dep="5587723" LSID="urn:lsid:zoobank.org:act:D612A25A-E09E-4193-8B89-3DC62746FB55" httpUri="http://treatment.plazi.org/id/03D4A4166C149460FF5ACDB6FB48F858" lastPageId="43" lastPageNumber="44" pageId="41" pageNumber="42">
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<heading id="D08AA26C6C149462FF5ACDB6FE6DFE49" bold="true" box="[151,461,476,502]" fontSize="11" level="1" pageId="41" pageNumber="42" reason="1">
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<taxonomicName id="4C7D6E836C149462FF5ACDB6FEC8FE49" ID-CoL="7DLVH" authority="Garg & Senevirathne & Wijayathilaka & Phuge & Deuti & Manamendra-Arachchi & Meegaskumbura & Biju, 2018" authorityName="Garg & Senevirathne & Wijayathilaka & Phuge & Deuti & Manamendra-Arachchi & Meegaskumbura & Biju" authorityYear="2018" box="[151,360,476,502]" class="Amphibia" family="Microhylidae" genus="Uperodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="41" pageNumber="42" phylum="Chordata" rank="species" species="rohani" status="sp. nov.">
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<emphasis id="B909C9126C149462FF5ACDB6FEC8FE49" bold="true" box="[151,360,476,502]" italics="true" pageId="41" pageNumber="42">Uperodon rohani</emphasis>
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<taxonomicNameLabel id="A23A74696C149462FEA2CDB6FE6DFE49" box="[367,461,476,502]" pageId="41" pageNumber="42" rank="species">sp. nov.</taxonomicNameLabel>
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<paragraph id="8BC215006C149462FF5ACD94FE38FDA8" blockId="41.[151,1176,476,607]" box="[151,408,510,535]" pageId="41" pageNumber="42">Rohan’s Globular Frog</paragraph>
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<paragraph id="8BC215006C149462FF5ACE48FB38FDE0" blockId="41.[151,1176,476,607]" pageId="41" pageNumber="42">
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(
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<figureCitation id="134609856C149462FF52CE48FF4CFD84" box="[159,236,546,571]" captionStart="FIGURE 1" captionStartId="13.[151,250,1632,1654]" captionTargetBox="[154,1433,199,1603]" captionTargetId="figure@13.[151,1436,195,1608]" captionTargetPageId="13" captionText="FIGURE 1. Maximum Likelihood phylogram based on 511 bp of mitochondrial 16S rRNA gene sequences from 73 individuals representing all known members of the genus Uperodon. Three samples representing taxon under synonymy are shown inquotationmarks.Outgroup not shown for graphical purposes.BayesianPosterior Probabilities and bootstrap values of>50 are shown above and below the branches, respectively. Values towards the species labels represent mean uncorrected genetic distances between the species with range values in parentheses." httpUri="https://zenodo.org/record/1183457/files/figure.png" pageId="41" pageNumber="42">Figs. 1</figureCitation>
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,
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<figureCitation id="134609856C149462FF37CE49FEEFFD84" box="[250,335,547,571]" captionStart="FIGURE 2" captionStartId="14.[151,250,1617,1639]" captionTargetBox="[152,1434,196,1594]" captionTargetId="figure@14.[151,1436,193,1595]" captionTargetPageId="14" captionText="FIGURE 2. A. Geographic distribution of the genus Uperodon in Bangladesh, Bhutan, India, Nepal, Pakistan and Sri Lanka; B. Distributionof Uperodon globulosus, U. systomaand U.taprobanicus; C. Distribution of four endemic Uperodon species of the Western Ghats (excluding the widely distributed Indian member U. variegatus, shown in Fig.12A); D. Distribution of four endemic Uperdodon species of Sri Lanka. Details of localities reported in the present study are provided in Table 1. Records from literature are listed in supplementary Table S1 and indicated here based onapproximate coordinates." httpUri="https://zenodo.org/record/1183459/files/figure.png" pageId="41" pageNumber="42">2A, 2D</figureCitation>
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,
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<figureCitation id="134609856C149462FE96CE49FEDCFD84" box="[347,380,547,571]" captionStart="FIGURE 3" captionStartId="15.[151,250,1749,1771]" captionTargetBox="[151,1436,644,1728]" captionTargetId="figure@15.[151,1436,644,1728]" captionTargetPageId="15" captionText="FIGURE 3. Preserved type specimens of Uperodon species in dorsal and ventral view. A. Uperodon systoma, Syntype, ZMB 3551 (female); B. Uperodon globulosus, Syntype, NHM 62.12.29.4 (ex. BMNH 1947.2.10.97) (female); C. Kaloula pulchra taprobanica, Holotype, NHM 1905.3.25.138 (ex. BMNH 1947.2.10.62) (female); D. Ramanella anamalaiensis, Neotype, ZSI/ WGRC/V/A/956 (male); E. Hylaedactylus montanus, Holotype, NHM 72.4.17.212 (ex. BMNH 1947.2.11.19) (female); F. Ramanella mormorata, Neotype, ZSI/WGRC/V/A/958, (female); G. Ramanella nagaoi, Holotype, WHT 2826 (male); H. Callula obscura, Holotype, NHM 58.11.28.42 (ex. BMNH 1947.2.11.20) (female); I. Ramanella palmata, Holotype, NHM 90.11.8.62 (ex.BMNH 1947.2.11.21); J. Callula triangularis, Lectotype, NHM 74.4.29.897 (ex.BMNH 1947.2.11.15) (male); K.Hylaedactylus carnaticus, Lectotype, ZSI 2761(female); L. Uperodonrohani sp. nov., Holotype, DZ1449 (male).(Images: SD Biju and Sonali Garg)." httpUri="https://zenodo.org/record/1183461/files/figure.png" pageId="41" pageNumber="42">3L</figureCitation>
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,
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<figureCitation id="134609856C149462FE45CE48FE0DFD84" box="[392,429,546,571]" captionStart="FIGURE 5" captionStartId="17.[151,250,1589,1611]" captionTargetBox="[155,1432,1093,1561]" captionTargetId="figure@17.[151,1436,1087,1567]" captionTargetPageId="17" captionText="FIGURE 5. Ventral surface of the palate in Uperodon species showing vomerine odontophores separated or fused with dermal projectionsand/or neopalatinalridgeson posterior sides ofthe choanae (indicated witharrows).A. Uperodon systoma, SDBDU 2005.2787 (female) and SDBDU 2005.2806 (male); B. Uperodon globulosus, SDBDU 2014.2673 (female) and SDBDU 2005.4762 (female); C. Uperodon taprobanicus, SDBDU 2007.6364 (female) and SDBDU 2005.4732A (male); D. Uperodon anamalaiensis, SDBDU 2013.2302 (female) and SDBDU 2002.3050 (male); E. Uperodon montanus, SDBDU 2002.24B (female) and SDBDU 2003.40115 (male); F. Uperodon mormorata, SDBDU 2011.507 (male) and SDBDU 2012.43 (male); G. Uperodon obscurus, DZ 1366 (male); H. Uperodon palmatus, DZ 1331 (male); I. Uperodon triangularis, SDBDU 2007.6400 (female) and ZSI/WGRC/V/A/959 (male); J. Uperodon variegatus, SDBDU 2005.4726 (female) and SDBDU 2005.4727 (male); K.Uperodon rohani sp. nov., DZ 1521 (male).(Images:SD Biju)." httpUri="https://zenodo.org/record/1183465/files/figure.png" pageId="41" pageNumber="42">5K</figureCitation>
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,
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<figureCitation id="134609856C149462FE75CE49FE76FD84" box="[440,470,547,571]" captionStart="FIGURE12" captionStartId="39.[151,250,1561,1583]" captionTargetBox="[151,1436,988,1540]" captionTargetId="figure@39.[150,1436,988,1540]" captionTargetPageId="39" captionText="FIGURE12. A.Distributionof Uperodon variegatus andUperodon rohani sp. nov. Detailsof localities reported inthe present study are provided in Table 1. Records from literature are listed in supplementary Table S1 and indicated here based on approximatecoordinates; B–E. Head, hand, footand schematic illustration of foot webbing inthe holotype of Uperodonrohani sp. nov. (DZ 1449, male): B. lateral view of head; C. ventral view of hand; D. ventral view of foot; E. foot webbing; F–J. Uperodonrohani sp. nov. (DZ 1449,male) inlife:F. dorsolateral view; G. dorsal view; H.ventral view; I. Uperodon rohanisp. nov. (Paratype,DZ 1485, female),dorsal view; J. Uperodon rohanisp. nov. (DZ 1130, female), dorsolateral view.(Images:SD Biju)." httpUri="https://zenodo.org/record/1183479/files/figure.png" pageId="41" pageNumber="42">12</figureCitation>
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,
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<figureCitation id="134609856C149462FE2FCE49FDA0FD84" box="[482,512,547,571]" captionStart="FIGURE 13" captionStartId="40.[151,250,716,738]" captionTargetBox="[166,1415,201,683]" captionTargetId="figure@40.[151,1435,193,694]" captionTargetPageId="40" captionText="FIGURE 13. Principal Component Analyses showing morphometric differentiation between Uperodon rohani sp. nov. and U. variegatus based on 17 morphological characters. Left: Plot for the first two factor planes explaining 54.4% of the variations among 28 adult male specimens. Right: Plot for the first two factor planes explaining 73% of the variations among 15 adult female specimens.Factor loadings for each component are provided in Table S2." httpUri="https://zenodo.org/record/1183481/files/figure.png" pageId="41" pageNumber="42">13</figureCitation>
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,
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<figureCitation id="134609856C149462FDC1CE49FD8BFD84" box="[524,555,547,571]" captionStart="FIGURE14" captionStartId="40.[151,250,1533,1555]" captionTargetBox="[163,1428,885,1494]" captionTargetId="figure@40.[151,1436,879,1511]" captionTargetPageId="40" captionText="FIGURE14.Cleared and double stainedspecimens of Uperodon rohani sp. nov. (above) and U.variegatus (below).From left to right: Dorsal view of the crania composed of nasals (NS), frontoparietals (FP), prootics (PR), exoccipitals (EX) and squamosals (SQ); anteroventral view of the crania showing components of the upper jaw, premaxillae (PM) and maxillae (MA); ventral view of the lower jaw comprising of dentaries (DN), angulosplenials (AG) and mentomeckelians (MM). Bones are indicated in red, cartilage in blue. (Images: Gayani Senevirathne)." httpUri="https://zenodo.org/record/1183483/files/figure.png" pageId="41" pageNumber="42">14</figureCitation>
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;
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<tableCitation id="C6FF20BB6C149462FDFACE48FD11FD84" box="[567,689,546,571]" captionStart-0="TABLE 1" captionStart-1="TABLE 2" captionStart-2="TABLE 4" captionStartId-0="4.[151,239,1593,1616]" captionStartId-1="9.[151,239,1084,1107]" captionStartId-2="48.[119,193,547,566]" captionTargetBox-2="[135,1884,616,1625]" captionText-0="TABLE 1. Collection localities for the genetically and/or morphologically studied Uperodon samples in the present study." captionText-1="TABLE 2. Samples used for the molecular study." captionText-2="TABLE 4. Morphometric measurements of Uperodon species.All measurements are in millimetres (mm). The status of each specimen is given after the museum number:Holotype (HT), Paratype (PT), Neotype (NT), Lectotype (LT), Syntype (ST)." httpUri-2="http://table.plazi.org/id/DF0245886C0D947BFFBACE49F8ECFD89" pageId="41" pageNumber="42" tableUuid-2="DF0245886C0D947BFFBACE49F8ECFD89">Tables 1–4</tableCitation>
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) http://zoobank.org/urn:lsid:zoobank.org:act:D612A25A-E09E-4193-8B89-3DC62746FB55
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</paragraph>
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<paragraph id="8BC215006C149462FF5ACEE4FBD5FAB4" blockId="41.[151,1437,654,2011]" pageId="41" pageNumber="42">
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<materialsCitation id="3B151F5D6C149462FF5ACEE4FD18FD74" ID-GBIF-Occurrence="1821690768" collectingDate="2014-10-02" collectorName="Madhava Meegaskumbura" country="Sri Lanka" location="Anuradhapura district" municipality="Mihintale" pageId="41" pageNumber="42" specimenCount="1" stateProvince="Sri" typeStatus="holotype">
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<emphasis id="B909C9126C149462FF5ACEE4FEABFD18" bold="true" box="[151,267,654,679]" pageId="41" pageNumber="42">
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<typeStatus id="54C6ABA26C149462FF5ACEE4FEA6FD18" box="[151,262,654,679]" pageId="41" pageNumber="42" type="holotype">Holotype</typeStatus>
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.
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</emphasis>
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DZ 1449, an adult male, from
|
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<collectingMunicipality id="6BA68F7A6C149462FD44CEE4FD59FD18" box="[649,761,654,679]" pageId="41" pageNumber="42">Mihintale</collectingMunicipality>
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,
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<location id="8EA243DB6C149462FCC4CEE4FBA6FD18" LSID="urn:lsid:plazi:treatment:03D4A4166C149460FF5ACDB6FB48F858:8EA243DB6C149462FCC4CEE4FBA6FD18" box="[777,1030,654,679]" country="Sri Lanka" municipality="Mihintale" name="Anuradhapura district" pageId="41" pageNumber="42" stateProvince="Sri">Anuradhapura district</location>
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,
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<collectingCountry id="F36A55906C149462FBDBCEE4FB2BFD18" box="[1046,1163,654,679]" name="Sri Lanka" pageId="41" pageNumber="42">
|
||
<collectingRegion id="49B9DBE26C149462FBDBCEE4FB97FD18" box="[1046,1079,654,679]" country="United Kingdom" name="Surrey" pageId="41" pageNumber="42">Sri</collectingRegion>
|
||
Lanka
|
||
</collectingCountry>
|
||
, collected by
|
||
<collectorName id="268870D66C149462FAF9CEE4FEF2FD74" pageId="41" pageNumber="42">Madhava Meegaskumbura</collectorName>
|
||
and team, on 0
|
||
<date id="FFC333C06C149462FDCECED9FD18FD74" box="[515,696,690,715]" pageId="41" pageNumber="42" value="2014-10-02">
|
||
<collectingDate id="EF87CA286C149462FDCECED9FD18FD74" box="[515,696,690,715]" pageId="41" pageNumber="42" value="2014-10-02">2 October 2014</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="3B151F5D6C149462FD0ECED8FD6EFCAC" ID-GBIF-Occurrence="1821690777" collectingDate="2013-11-04" country="Sri Lanka" location="Hambantota district" municipality="Bundala" pageId="41" pageNumber="42" specimenCount="2" stateProvince="Sri" typeStatus="paratype">
|
||
<emphasis id="B909C9126C149462FD0ECED8FCE2FD74" bold="true" box="[707,834,690,715]" pageId="41" pageNumber="42">
|
||
<typeStatus id="54C6ABA26C149462FD0ECED8FC9DFD74" box="[707,829,690,715]" pageId="41" pageNumber="42" type="paratype">Paratypes</typeStatus>
|
||
.
|
||
</emphasis>
|
||
DZ 1448, DZ 1450 and DZ 1374, three adult males, collected along with the
|
||
<typeStatus id="54C6ABA26C149462FE75CEBCFDBEFD50" box="[440,542,726,751]" pageId="41" pageNumber="42" type="holotype">holotype</typeStatus>
|
||
; DZ 1336, an adult male, from
|
||
<collectingMunicipality id="6BA68F7A6C149462FC5ACEBCFC54FD50" box="[919,1012,726,751]" pageId="41" pageNumber="42">Bundala</collectingMunicipality>
|
||
NP,
|
||
<location id="8EA243DB6C149462FBFCCEBCFAB5FD50" LSID="urn:lsid:plazi:treatment:03D4A4166C149460FF5ACDB6FB48F858:8EA243DB6C149462FBFCCEBCFAB5FD50" box="[1073,1301,726,751]" country="Sri Lanka" municipality="Bundala" name="Hambantota district" pageId="41" pageNumber="42" stateProvince="Sri">Hambantota district</location>
|
||
,
|
||
<collectingCountry id="F36A55906C149462FAE9CEBCFA38FD50" box="[1316,1432,726,751]" name="Sri Lanka" pageId="41" pageNumber="42">
|
||
<collectingRegion id="49B9DBE26C149462FAE9CEBCFAE5FD50" box="[1316,1349,726,751]" country="United Kingdom" name="Surrey" pageId="41" pageNumber="42">Sri</collectingRegion>
|
||
Lanka
|
||
</collectingCountry>
|
||
, collected by MM and team, on 0
|
||
<date id="FFC333C06C149462FDCCCE91FD6EFCAC" box="[513,718,762,787]" pageId="41" pageNumber="42" value="2013-11-04">
|
||
<collectingDate id="EF87CA286C149462FDCCCE91FD6EFCAC" box="[513,718,762,787]" pageId="41" pageNumber="42" value="2013-11-04">4 November 2013</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B151F5D6C149462FD14CE91FACAFC88" ID-GBIF-Occurrence="1821690769" collectingDate="2014-09-11" country="Sri Lanka" location="Kurunegala district" municipality="Makandura" pageId="41" pageNumber="42" specimenCount="1" stateProvince="Sri" typeStatus="paratype">
|
||
DZ 1520 and DZ 1521, two adult males, and DZ 1425, an adult female, from
|
||
<collectingMunicipality id="6BA68F7A6C149462FEFCCF74FE13FC88" box="[305,435,798,823]" pageId="41" pageNumber="42">Makandura</collectingMunicipality>
|
||
,
|
||
<location id="8EA243DB6C149462FE0DCF74FD3BFC88" LSID="urn:lsid:plazi:treatment:03D4A4166C149460FF5ACDB6FB48F858:8EA243DB6C149462FE0DCF74FD3BFC88" box="[448,667,798,823]" country="Sri Lanka" municipality="Makandura" name="Kurunegala district" pageId="41" pageNumber="42" stateProvince="Sri">Kurunegala district</location>
|
||
,
|
||
<collectingCountry id="F36A55906C149462FD6ACF74FCB9FC88" box="[679,793,798,823]" name="Sri Lanka" pageId="41" pageNumber="42">
|
||
<collectingRegion id="49B9DBE26C149462FD6ACF74FD68FC88" box="[679,712,798,823]" country="United Kingdom" name="Surrey" pageId="41" pageNumber="42">Sri</collectingRegion>
|
||
Lanka
|
||
</collectingCountry>
|
||
, collected by MM and team, on
|
||
<date id="FFC333C06C149462FB41CF75FACAFC88" box="[1164,1386,798,823]" pageId="41" pageNumber="42" value="2014-09-11">
|
||
<collectingDate id="EF87CA286C149462FB41CF75FACAFC88" box="[1164,1386,798,823]" pageId="41" pageNumber="42" value="2014-09-11">11 September 2014</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B151F5D6C149462FABACF75FE4FFCC0" ID-GBIF-Occurrence="1821690767" collectingDate="2012-10-10" country="Sri Lanka" location="Ampara district" municipality="Ampara" pageId="41" pageNumber="42" specimenCount="1" stateProvince="Sri" typeStatus="paratype">
|
||
DZ 1486, an adult male, and DZ 1485, an adult female, from
|
||
<collectingMunicipality id="6BA68F7A6C149462FCE2CF29FC2CFCE4" box="[815,908,835,859]" pageId="41" pageNumber="42">Ampara</collectingMunicipality>
|
||
,
|
||
<location id="8EA243DB6C149462FC55CF29FBEBFCE4" LSID="urn:lsid:plazi:treatment:03D4A4166C149460FF5ACDB6FB48F858:8EA243DB6C149462FC55CF29FBEBFCE4" box="[920,1099,834,859]" country="Sri Lanka" municipality="Ampara" name="Ampara district" pageId="41" pageNumber="42" stateProvince="Sri">Ampara district</location>
|
||
,
|
||
<collectingCountry id="F36A55906C149462FB94CF28FB6BFCE4" box="[1113,1227,834,859]" name="Sri Lanka" pageId="41" pageNumber="42">
|
||
<collectingRegion id="49B9DBE26C149462FB94CF28FBDAFCE4" box="[1113,1146,834,859]" country="United Kingdom" name="Surrey" pageId="41" pageNumber="42">Sri</collectingRegion>
|
||
Lanka
|
||
</collectingCountry>
|
||
, collected by MM and team, on
|
||
<date id="FFC333C06C149462FEE2CF0DFE4FFCC0" box="[303,495,870,895]" pageId="41" pageNumber="42" value="2012-10-10">
|
||
<collectingDate id="EF87CA286C149462FEE2CF0DFE4FFCC0" box="[303,495,870,895]" pageId="41" pageNumber="42" value="2012-10-10">10 October 2012</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B151F5D6C149462FE31CF0DFD6BFC1C" ID-GBIF-Occurrence="1821690771" collectingDate="2013-01-04" country="Sri Lanka" location="Matale district" municipality="Dambulla" pageId="41" pageNumber="42" specimenCount="1" stateProvince="Sri" typeStatus="paratype">
|
||
DZ 1129–DZ 1132, four adult females, from
|
||
<collectingMunicipality id="6BA68F7A6C149462FC34CF0CFBCAFCC0" box="[1017,1130,870,895]" pageId="41" pageNumber="42">Dambulla</collectingMunicipality>
|
||
,
|
||
<location id="8EA243DB6C149462FBBBCF0CFABCFCC0" LSID="urn:lsid:plazi:treatment:03D4A4166C149460FF5ACDB6FB48F858:8EA243DB6C149462FBBBCF0CFABCFCC0" box="[1142,1308,870,895]" country="Sri Lanka" municipality="Dambulla" name="Matale district" pageId="41" pageNumber="42" stateProvince="Sri">Matale district</location>
|
||
,
|
||
<collectingCountry id="F36A55906C149462FAEACF0CFA38FCC0" box="[1319,1432,870,895]" name="Sri Lanka" pageId="41" pageNumber="42">
|
||
<collectingRegion id="49B9DBE26C149462FAEACF0CFAE8FCC0" box="[1319,1352,870,895]" country="United Kingdom" name="Surrey" pageId="41" pageNumber="42">Sri</collectingRegion>
|
||
Lanka
|
||
</collectingCountry>
|
||
, collected by MM and team, on 0
|
||
<date id="FFC333C06C149462FDDBCFE1FD6BFC1C" box="[534,715,907,931]" pageId="41" pageNumber="42" value="2013-01-04">
|
||
<collectingDate id="EF87CA286C149462FDDBCFE1FD6BFC1C" box="[534,715,907,931]" pageId="41" pageNumber="42" value="2013-01-04">4 January 2013</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="3B151F5D6C149462FD1ACFE0FD04FC78" ID-GBIF-Occurrence="1821690778" collectionCode="NHM" collectorName="GH Herry" country="Sri Lanka" location="Ceylon" municipality="Referred" pageId="41" pageNumber="42" specimenCode="NHM 1931.2" specimenCount="1" stateProvince="Sri" typeStatus="paratype">
|
||
<emphasis id="B909C9126C149462FD1ACFE0FC6DFC1C" bold="true" box="[727,973,906,931]" pageId="41" pageNumber="42">
|
||
<collectingMunicipality id="6BA68F7A6C149462FD1ACFE0FCE2FC1C" box="[727,834,906,931]" pageId="41" pageNumber="42">Referred</collectingMunicipality>
|
||
specimens.
|
||
</emphasis>
|
||
<specimenCode id="DBDBBD7B6C149462FC1BCFE1FBD5FC1C" box="[982,1141,907,931]" collectionCode="NHM" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14908" name="University of Nottingham" pageId="41" pageNumber="42">NHM 1931.2</specimenCode>
|
||
.1.21, an adult male, from “
|
||
<location id="8EA243DB6C149462FF6CCFC4FF58FC78" LSID="urn:lsid:plazi:treatment:03D4A4166C149460FF5ACDB6FB48F858:8EA243DB6C149462FF6CCFC4FF58FC78" box="[161,248,942,967]" country="Sri Lanka" municipality="Referred" name="Ceylon" pageId="41" pageNumber="42" stateProvince="Sri">Ceylon</location>
|
||
”,
|
||
<collectingCountry id="F36A55906C149462FEDFCFC4FE27FC78" box="[274,391,942,967]" name="Sri Lanka" pageId="41" pageNumber="42">
|
||
<collectingRegion id="49B9DBE26C149462FEDFCFC4FE93FC78" box="[274,307,942,967]" country="United Kingdom" name="Surrey" pageId="41" pageNumber="42">Sri</collectingRegion>
|
||
Lanka
|
||
</collectingCountry>
|
||
, collected by
|
||
<collectorName id="268870D66C149462FDE1CFC5FD04FC78" box="[556,676,943,967]" pageId="41" pageNumber="42">GH Herry</collectorName>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B151F5D6C149462FD7ECFC5FCC1FC54" ID-GBIF-Occurrence="1821690773" collectionCode="NHM" collectorName="R Sananayake" country="Sri Lanka" location="Kurunegala dist." municipality="Kurunegala" pageId="41" pageNumber="42" specimenCode="NHM 1973.3015" specimenCount="1" stateProvince="Sri" typeStatus="paratype">
|
||
<specimenCode id="DBDBBD7B6C149462FD7ECFC5FCD9FC78" box="[691,889,942,967]" collectionCode="NHM" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14908" name="University of Nottingham" pageId="41" pageNumber="42">NHM 1973.3015</specimenCode>
|
||
, an adult male, from
|
||
<collectingMunicipality id="6BA68F7A6C149462FBB4CFC4FB5FFC78" box="[1145,1279,942,967]" pageId="41" pageNumber="42">Kurunegala</collectingMunicipality>
|
||
, “
|
||
<location id="8EA243DB6C149462FADACFC4FF6AFC54" LSID="urn:lsid:plazi:treatment:03D4A4166C149460FF5ACDB6FB48F858:8EA243DB6C149462FADACFC4FF6AFC54" country="Sri Lanka" municipality="Kurunegala" name="Kurunegala dist." pageId="41" pageNumber="42" stateProvince="Sri">Kurunegala dist.</location>
|
||
,”
|
||
<collectingCountry id="F36A55906C149462FF29CFB8FEF7FC54" box="[228,343,978,1003]" name="Sri Lanka" pageId="41" pageNumber="42">
|
||
<collectingRegion id="49B9DBE26C149462FF29CFB8FEA5FC54" box="[228,261,978,1003]" country="United Kingdom" name="Surrey" pageId="41" pageNumber="42">Sri</collectingRegion>
|
||
Lanka
|
||
</collectingCountry>
|
||
, collected by
|
||
<collectorName id="268870D66C149462FE37CFB9FD00FC54" box="[506,672,978,1003]" pageId="41" pageNumber="42">R Sananayake</collectorName>
|
||
; NMH 1976.264
|
||
</materialsCitation>
|
||
–
|
||
<materialsCitation id="3B151F5D6C149462FCBDCFB8FAF1FB8C" ID-GBIF-Occurrence="1821690774" collectingDate="2000-11-18" collectionCode="NHM" collectorName="JC Naggs & MM Bahir, C & Gans & C Yahalmulla" country="Sri Lanka" county="Badulla" location="Nochchiyagama" municipality="Hambantota dist." pageId="41" pageNumber="42" specimenCode="NHM 1976.265" specimenCount="1" stateProvince="Sri" typeStatus="paratype">
|
||
<specimenCode id="DBDBBD7B6C149462FCBDCFB8FB88FC54" box="[880,1064,978,1003]" collectionCode="NHM" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14908" name="University of Nottingham" pageId="41" pageNumber="42">NHM 1976.265</specimenCode>
|
||
, two adult males, from
|
||
<collectingCounty id="62A36D8C6C149462FAF3CFB8FA38FC54" box="[1342,1432,978,1003]" pageId="41" pageNumber="42">Badulla</collectingCounty>
|
||
, “
|
||
<collectingMunicipality id="6BA68F7A6C149462FF6CCF9CFECEFBB0" box="[161,366,1014,1039]" pageId="41" pageNumber="42">Hambantota dist.</collectingMunicipality>
|
||
,”
|
||
<collectingCountry id="F36A55906C149462FE40CF9CFDA5FBB0" box="[397,517,1014,1039]" name="Sri Lanka" pageId="41" pageNumber="42">
|
||
<collectingRegion id="49B9DBE26C149462FE40CF9CFE0EFBB0" box="[397,430,1014,1039]" country="United Kingdom" name="Surrey" pageId="41" pageNumber="42">Sri</collectingRegion>
|
||
Lanka
|
||
</collectingCountry>
|
||
, collected by
|
||
<collectorName id="268870D66C149462FD7ACF9DFC8EFBB0" box="[695,814,1015,1039]" pageId="41" pageNumber="42">JC Naggs</collectorName>
|
||
; WHT 5320, an adult male, from
|
||
<location id="8EA243DB6C149462FB12CF9CFA38FBB0" LSID="urn:lsid:plazi:treatment:03D4A4166C149460FF5ACDB6FB48F858:8EA243DB6C149462FB12CF9CFA38FBB0" box="[1247,1432,1014,1039]" country="Sri Lanka" county="Badulla" municipality="Hambantota dist." name="Nochchiyagama" pageId="41" pageNumber="42" stateProvince="Sri">Nochchiyagama</location>
|
||
,
|
||
<location id="8EA243DB6C149462FF5AC870FE2EFB8C" LSID="urn:lsid:plazi:treatment:03D4A4166C149460FF5ACDB6FB48F858:8EA243DB6C149462FF5AC870FE2EFB8C" box="[151,398,1050,1075]" country="Sri Lanka" county="Badulla" municipality="Hambantota dist." name="Anuradhapura district" pageId="41" pageNumber="42" stateProvince="Sri">Anuradhapura district</location>
|
||
,
|
||
<collectingCountry id="F36A55906C149462FE54C870FDA9FB8C" box="[409,521,1050,1075]" name="Sri Lanka" pageId="41" pageNumber="42">
|
||
<collectingRegion id="49B9DBE26C149462FE54C870FE1AFB8C" box="[409,442,1050,1075]" country="United Kingdom" name="Surrey" pageId="41" pageNumber="42">Sri</collectingRegion>
|
||
Lanka
|
||
</collectingCountry>
|
||
, collected by
|
||
<collectorName id="268870D66C149462FD6FC871FCD8FB8C" box="[674,888,1050,1075]" pageId="41" pageNumber="42">MM Bahir, C Gans</collectorName>
|
||
and
|
||
<collectorName id="268870D66C149462FC60C871FBEAFB8C" box="[941,1098,1050,1075]" pageId="41" pageNumber="42">C Yahalmulla</collectorName>
|
||
, on
|
||
<date id="FFC333C06C149462FBBAC871FAF1FB8C" box="[1143,1361,1050,1075]" pageId="41" pageNumber="42" value="2000-11-18">
|
||
<collectingDate id="EF87CA286C149462FBBAC871FAF1FB8C" box="[1143,1361,1050,1075]" pageId="41" pageNumber="42" value="2000-11-18">18 November 2000</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B151F5D6C149462FA90C871FEE1FBC4" ID-GBIF-Occurrence="1821690776" collectingDate="1996-02-08" collectorName="D Gabadge" country="Sri Lanka" location="Matale district" pageId="41" pageNumber="42" specimenCount="1" stateProvince="Sri" typeStatus="paratype">
|
||
WHT 0 1188, an adult male, from Sigiriya, Near Rest house,
|
||
<location id="8EA243DB6C149462FCCAC854FC0EFBE8" LSID="urn:lsid:plazi:treatment:03D4A4166C149460FF5ACDB6FB48F858:8EA243DB6C149462FCCAC854FC0EFBE8" box="[775,942,1086,1111]" country="Sri Lanka" name="Matale district" pageId="41" pageNumber="42" stateProvince="Sri">Matale district</location>
|
||
,
|
||
<collectingCountry id="F36A55906C149462FC77C854FB8DFBE8" box="[954,1069,1086,1111]" name="Sri Lanka" pageId="41" pageNumber="42">
|
||
<collectingRegion id="49B9DBE26C149462FC77C854FC7BFBE8" box="[954,987,1086,1111]" country="United Kingdom" name="Surrey" pageId="41" pageNumber="42">Sri</collectingRegion>
|
||
Lanka
|
||
</collectingCountry>
|
||
, collected by
|
||
<collectorName id="268870D66C149462FB01C855FAEFFBE8" box="[1228,1359,1086,1111]" pageId="41" pageNumber="42">D Gabadge</collectorName>
|
||
, on 0
|
||
<date id="FFC333C06C149462FA42C855FEE1FBC4" pageId="41" pageNumber="42" value="1996-02-08">
|
||
<collectingDate id="EF87CA286C149462FA42C855FEE1FBC4" pageId="41" pageNumber="42" value="1996-02-08">8 February 1996</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B151F5D6C149462FE82C809FE1DFB20" ID-GBIF-Occurrence="1821690770" collectingDate="1997-07-28" collectorName="A Nollert" country="Sri Lanka" location="Polonnaruwa district" municipality="Polonnaruwa" pageId="41" pageNumber="42" specimenCount="1" stateProvince="Sri" typeStatus="paratype">
|
||
WHT 1749, an adult male, from
|
||
<collectingMunicipality id="6BA68F7A6C149462FD1DC808FCC7FBC4" box="[720,871,1122,1147]" pageId="41" pageNumber="42">Polonnaruwa</collectingMunicipality>
|
||
,
|
||
<location id="8EA243DB6C149462FCB8C808FBC4FBC4" LSID="urn:lsid:plazi:treatment:03D4A4166C149460FF5ACDB6FB48F858:8EA243DB6C149462FCB8C808FBC4FBC4" box="[885,1124,1122,1147]" country="Sri Lanka" municipality="Polonnaruwa" name="Polonnaruwa district" pageId="41" pageNumber="42" stateProvince="Sri">Polonnaruwa district</location>
|
||
,
|
||
<collectingCountry id="F36A55906C149462FBBCC808FB45FBC4" box="[1137,1253,1122,1147]" name="Sri Lanka" pageId="41" pageNumber="42">
|
||
<collectingRegion id="49B9DBE26C149462FBBCC808FB32FBC4" box="[1137,1170,1122,1147]" country="United Kingdom" name="Surrey" pageId="41" pageNumber="42">Sri</collectingRegion>
|
||
Lanka
|
||
</collectingCountry>
|
||
, collected by
|
||
<collectorName id="268870D66C149462FA45C809FF49FB20" pageId="41" pageNumber="42">A Nollert</collectorName>
|
||
, on
|
||
<date id="FFC333C06C149462FEEDC8EDFE1DFB20" box="[288,445,1158,1183]" pageId="41" pageNumber="42" value="1997-07-28">
|
||
<collectingDate id="EF87CA286C149462FEEDC8EDFE1DFB20" box="[288,445,1158,1183]" pageId="41" pageNumber="42" value="1997-07-28">28 July 1997</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B151F5D6C149462FE00C8EDFCE2FB7C" ID-GBIF-Occurrence="1821690772" collectingDate="1995-08-11" collectorName="D Gabadage & MM Bahir" country="Sri Lanka" location="Anuradhapura district" municipality="Anuradhapura" pageId="41" pageNumber="42" specimenCount="1" stateProvince="Sri" typeStatus="paratype">
|
||
WHT 0 1189, an adult male, from
|
||
<collectingMunicipality id="6BA68F7A6C149462FCA9C8ECFBA7FB20" box="[868,1031,1158,1183]" pageId="41" pageNumber="42">Anuradhapura</collectingMunicipality>
|
||
,
|
||
<location id="8EA243DB6C149462FBDAC8ECFAB4FB20" LSID="urn:lsid:plazi:treatment:03D4A4166C149460FF5ACDB6FB48F858:8EA243DB6C149462FBDAC8ECFAB4FB20" box="[1047,1300,1158,1183]" country="Sri Lanka" municipality="Anuradhapura" name="Anuradhapura district" pageId="41" pageNumber="42" stateProvince="Sri">Anuradhapura district</location>
|
||
,
|
||
<collectingCountry id="F36A55906C149462FAEEC8ECFA38FB20" box="[1315,1432,1158,1183]" name="Sri Lanka" pageId="41" pageNumber="42">
|
||
<collectingRegion id="49B9DBE26C149462FAEEC8ECFAE4FB20" box="[1315,1348,1158,1183]" country="United Kingdom" name="Surrey" pageId="41" pageNumber="42">Sri</collectingRegion>
|
||
Lanka
|
||
</collectingCountry>
|
||
, collected by
|
||
<collectorName id="268870D66C149462FEEBC8C1FE11FB7C" box="[294,433,1194,1219]" pageId="41" pageNumber="42">D Gabadage</collectorName>
|
||
and
|
||
<collectorName id="268870D66C149462FE2AC8C1FDC0FB7C" box="[487,608,1194,1219]" pageId="41" pageNumber="42">MM Bahir</collectorName>
|
||
, on
|
||
<date id="FFC333C06C149462FD40C8C1FCE2FB7C" box="[653,834,1194,1219]" pageId="41" pageNumber="42" value="1995-08-11">
|
||
<collectingDate id="EF87CA286C149462FD40C8C1FCE2FB7C" box="[653,834,1194,1219]" pageId="41" pageNumber="42" value="1995-08-11">11 August 1995</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B151F5D6C149462FC80C8C1FBD1FAB4" ID-GBIF-Occurrence="1821690775" collectingDate="1995-08" collectionCode="NHM" collectorName="R Sananayake & U. Premachandra" country="Sri Lanka" county="Kurunegala" location="Kurunegala dist." municipality="Ceylon" pageId="41" pageNumber="42" specimenCode="NHM 1973.3014" specimenCount="1" stateProvince="Sri" typeStatus="paratype">
|
||
<specimenCode id="DBDBBD7B6C149462FC80C8C1FBB0FB7C" box="[845,1040,1195,1219]" collectionCode="NHM" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14908" name="University of Nottingham" pageId="41" pageNumber="42">NHM 1973.3014</specimenCode>
|
||
, an adult female, from
|
||
<collectingCounty id="62A36D8C6C149462FADFC8C0FA39FB7C" box="[1298,1433,1194,1219]" pageId="41" pageNumber="42">Kurunegala</collectingCounty>
|
||
:
|
||
<collectingMunicipality id="6BA68F7A6C149462FF5AC8A4FF4BFB58" box="[151,235,1230,1255]" pageId="41" pageNumber="42">Ceylon</collectingMunicipality>
|
||
, “
|
||
<location id="8EA243DB6C149462FF32C8A4FE1EFB58" LSID="urn:lsid:plazi:treatment:03D4A4166C149460FF5ACDB6FB48F858:8EA243DB6C149462FF32C8A4FE1EFB58" box="[255,446,1230,1255]" country="Sri Lanka" county="Kurunegala" municipality="Ceylon" name="Kurunegala dist." pageId="41" pageNumber="42" stateProvince="Sri">Kurunegala dist.</location>
|
||
,”
|
||
<collectingCountry id="F36A55906C149462FE18C8A4FDE5FB58" box="[469,581,1230,1255]" name="Sri Lanka" pageId="41" pageNumber="42">
|
||
<collectingRegion id="49B9DBE26C149462FE18C8A4FE56FB58" box="[469,502,1230,1255]" country="United Kingdom" name="Surrey" pageId="41" pageNumber="42">Sri</collectingRegion>
|
||
Lanka
|
||
</collectingCountry>
|
||
, collected by
|
||
<collectorName id="268870D66C149462FD13C8A5FC20FB58" box="[734,896,1230,1255]" pageId="41" pageNumber="42">R Sananayake</collectorName>
|
||
; WHT 0 1191, an adult female, from Palatupana, Kirinda, Hambantota district,
|
||
<collectingCountry id="F36A55906C149462FE2BC898FDF6FAB4" box="[486,598,1266,1291]" name="Sri Lanka" pageId="41" pageNumber="42">
|
||
<collectingRegion id="49B9DBE26C149462FE2BC898FDA7FAB4" box="[486,519,1266,1291]" country="United Kingdom" name="Surrey" pageId="41" pageNumber="42">Sri</collectingRegion>
|
||
Lanka
|
||
</collectingCountry>
|
||
, collected by
|
||
<collectorName id="268870D66C149462FD3CC899FC16FAB4" box="[753,950,1266,1291]" pageId="41" pageNumber="42">U. Premachandra</collectorName>
|
||
, in
|
||
<date id="FFC333C06C149462FC13C899FBD1FAB4" box="[990,1137,1266,1291]" pageId="41" pageNumber="42" value="1995-08">
|
||
<collectingDate id="EF87CA286C149462FC13C899FBD1FAB4" box="[990,1137,1266,1291]" pageId="41" pageNumber="42" value="1995-08">August 1995</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C367468B6C149462FF0AC97CFA3CF864" pageId="41" pageNumber="42" type="etymology">
|
||
<paragraph id="8BC215006C149462FF0AC97CFE90FAC9" blockId="41.[151,1437,654,2011]" pageId="41" pageNumber="42">
|
||
<emphasis id="B909C9126C149462FF0AC97CFEEEFA90" bold="true" box="[199,334,1302,1327]" pageId="41" pageNumber="42">Etymology.</emphasis>
|
||
This species is named after Rohan Pethiyagoda, in recognition of his contribution to biodiversity studies in
|
||
<collectingCountry id="F36A55906C149462FEDDC950FEEAFAEC" box="[272,330,1338,1363]" name="India" pageId="41" pageNumber="42">India</collectingCountry>
|
||
and
|
||
<collectingCountry id="F36A55906C149462FE45C950FE5DFAEC" box="[392,509,1338,1363]" name="Sri Lanka" pageId="41" pageNumber="42">
|
||
<collectingRegion id="49B9DBE26C149462FE45C950FE09FAEC" box="[392,425,1338,1363]" country="United Kingdom" name="Surrey" pageId="41" pageNumber="42">Sri</collectingRegion>
|
||
Lanka
|
||
</collectingCountry>
|
||
, especially on frogs and fishes. The species epithet is formed as a noun in the genitive case.
|
||
</paragraph>
|
||
<paragraph id="8BC215006C149462FF0AC9E8FA3CF864" blockId="41.[151,1437,654,2011]" pageId="41" pageNumber="42">
|
||
<emphasis id="B909C9126C149462FF0AC9E8FEC0FA24" bold="true" box="[199,352,1410,1435]" pageId="41" pageNumber="42">Comparison.</emphasis>
|
||
<taxonomicName id="4C7D6E836C149462FEBCC9E8FD99FA24" authority="Garg & Senevirathne & Wijayathilaka & Phuge & Deuti & Manamendra-Arachchi & Meegaskumbura & Biju, 2018" authorityName="Garg & Senevirathne & Wijayathilaka & Phuge & Deuti & Manamendra-Arachchi & Meegaskumbura & Biju" authorityYear="2018" box="[369,569,1410,1435]" class="Amphibia" family="Microhylidae" genus="Uperodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="41" pageNumber="42" phylum="Chordata" rank="species" species="rohani" status="sp. nov.">
|
||
<emphasis id="B909C9126C149462FEBCC9E8FD99FA24" box="[369,569,1410,1435]" italics="true" pageId="41" pageNumber="42">Uperodon rohani</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B909C9126C149462FD87C9E8FD0CFA24" bold="true" box="[586,684,1410,1435]" pageId="41" pageNumber="42">
|
||
<taxonomicNameLabel id="A23A74696C149462FD87C9E8FD0CFA24" box="[586,684,1410,1435]" pageId="41" pageNumber="42" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
cannot be confused with
|
||
<taxonomicName id="4C7D6E836C149462FC35C9EEFB3CFA24" box="[1016,1180,1410,1435]" class="Amphibia" family="Microhylidae" genus="Uperodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="41" pageNumber="42" phylum="Chordata" rank="species" species="globulosus">
|
||
<emphasis id="B909C9126C149462FC35C9EEFB3CFA24" box="[1016,1180,1410,1435]" italics="true" pageId="41" pageNumber="42">U. globulosus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C7D6E836C149462FB7EC9EEFA98FA24" baseAuthorityName="Schneider" baseAuthorityYear="1799" box="[1203,1336,1412,1435]" class="Amphibia" family="Microhylidae" genus="Uperodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="41" pageNumber="42" phylum="Chordata" rank="species" species="systoma">
|
||
<emphasis id="B909C9126C149462FB7EC9EEFA98FA24" box="[1203,1336,1412,1435]" italics="true" pageId="41" pageNumber="42">U. systoma</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C7D6E836C149462FA4CC9EEFE88FA00" baseAuthorityName="Parker" baseAuthorityYear="1934" class="Amphibia" family="Microhylidae" genus="Uperodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="41" pageNumber="42" phylum="Chordata" rank="species" species="taprobanicus">
|
||
<emphasis id="B909C9126C149462FA4CC9EEFE88FA00" italics="true" pageId="41" pageNumber="42">U. taprobanicus</emphasis>
|
||
</taxonomicName>
|
||
due to its smaller snout-vent size, male SVL
|
||
<quantity id="4C85B8E56C149462FCE0C9CCFC08FA01" box="[813,936,1446,1471]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" metricValueMax="3.4" metricValueMin="2.6" pageId="41" pageNumber="42" unit="mm" value="30.0" valueMax="34.0" valueMin="26.0">26–34 mm</quantity>
|
||
,
|
||
<emphasis id="B909C9126C149462FC79C9CDFC68FA01" box="[948,968,1447,1470]" italics="true" pageId="41" pageNumber="42">N</emphasis>
|
||
= 16, female SVL
|
||
<quantity id="4C85B8E56C149462FB6FC9CCFABEFA01" box="[1186,1310,1446,1471]" metricMagnitude="-2" metricUnit="m" metricValue="3.15" metricValueMax="3.5" metricValueMin="2.8" pageId="41" pageNumber="42" unit="mm" value="31.5" valueMax="35.0" valueMin="28.0">28–35 mm</quantity>
|
||
,
|
||
<emphasis id="B909C9126C149462FAE7C9CDFA9DFA01" box="[1322,1341,1447,1470]" italics="true" pageId="41" pageNumber="42">N</emphasis>
|
||
= 8 (vs. larger,
|
||
<taxonomicName id="4C7D6E836C149462FF2EC9A6FEDCFA5C" box="[227,380,1482,1507]" class="Amphibia" family="Microhylidae" genus="Uperodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="41" pageNumber="42" phylum="Chordata" rank="species" species="globulosus">
|
||
<emphasis id="B909C9126C149462FF2EC9A6FEDCFA5C" box="[227,380,1482,1507]" italics="true" pageId="41" pageNumber="42">U. globulosus</emphasis>
|
||
</taxonomicName>
|
||
male SVL
|
||
<quantity id="4C85B8E56C149462FE37C9A0FDD4FA5D" box="[506,628,1482,1507]" metricMagnitude="-2" metricUnit="m" metricValue="5.35" metricValueMax="5.6" metricValueMin="5.1" pageId="41" pageNumber="42" unit="mm" value="53.5" valueMax="56.0" valueMin="51.0">51–56 mm</quantity>
|
||
,
|
||
<emphasis id="B909C9126C149462FDB3C9A6FD31FA5C" box="[638,657,1484,1507]" italics="true" pageId="41" pageNumber="42">N</emphasis>
|
||
= 2, female SVL
|
||
<quantity id="4C85B8E56C149462FC98C9A0FC6FFA5D" box="[853,975,1482,1507]" metricMagnitude="-2" metricUnit="m" metricValue="6.9" metricValueMax="7.2" metricValueMin="6.6" pageId="41" pageNumber="42" unit="mm" value="69.0" valueMax="72.0" valueMin="66.0">66–72 mm</quantity>
|
||
,
|
||
<emphasis id="B909C9126C149462FC17C9A6FC4DFA5C" box="[986,1005,1484,1507]" italics="true" pageId="41" pageNumber="42">N</emphasis>
|
||
= 3;
|
||
<taxonomicName id="4C7D6E836C149462FBE8C9A6FB00FA5C" baseAuthorityName="Schneider" baseAuthorityYear="1799" box="[1061,1184,1484,1507]" class="Amphibia" family="Microhylidae" genus="Uperodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="41" pageNumber="42" phylum="Chordata" rank="species" species="systoma">
|
||
<emphasis id="B909C9126C149462FBE8C9A6FB00FA5C" box="[1061,1184,1484,1507]" italics="true" pageId="41" pageNumber="42">U. systoma</emphasis>
|
||
</taxonomicName>
|
||
male SVL
|
||
<quantity id="4C85B8E56C149462FAD3C9A0FA38FA5D" box="[1310,1432,1482,1507]" metricMagnitude="-2" metricUnit="m" metricValue="5.35" metricValueMax="5.6" metricValueMin="5.1" pageId="41" pageNumber="42" unit="mm" value="53.5" valueMax="56.0" valueMin="51.0">51–56 mm</quantity>
|
||
,
|
||
<emphasis id="B909C9126C149462FF5AC985FF0AF9B9" box="[151,170,1519,1542]" italics="true" pageId="41" pageNumber="42">N</emphasis>
|
||
= 2, female SVL
|
||
<quantity id="4C85B8E56C149462FEBCC984FE4BF9B9" box="[369,491,1518,1543]" metricMagnitude="-2" metricUnit="m" metricValue="5.55" metricValueMax="6.4" metricValueMin="4.7" pageId="41" pageNumber="42" unit="mm" value="55.5" valueMax="64.0" valueMin="47.0">47–64 mm</quantity>
|
||
,
|
||
<emphasis id="B909C9126C149462FE3BC985FDA9F9B9" box="[502,521,1519,1542]" italics="true" pageId="41" pageNumber="42">N</emphasis>
|
||
= 4; and
|
||
<taxonomicName id="4C7D6E836C149462FDBFC985FC85F9B8" baseAuthorityName="Parker" baseAuthorityYear="1934" box="[626,805,1518,1543]" class="Amphibia" family="Microhylidae" genus="Uperodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="41" pageNumber="42" phylum="Chordata" rank="species" species="taprobanicus">
|
||
<emphasis id="B909C9126C149462FDBFC985FC85F9B8" box="[626,805,1518,1543]" italics="true" pageId="41" pageNumber="42">U. taprobanicus</emphasis>
|
||
</taxonomicName>
|
||
male SVL
|
||
<quantity id="4C85B8E56C149462FC68C984FBBFF9B9" box="[933,1055,1518,1543]" metricMagnitude="-2" metricUnit="m" metricValue="4.7" metricValueMax="5.2" metricValueMin="4.2" pageId="41" pageNumber="42" unit="mm" value="47.0" valueMax="52.0" valueMin="42.0">42–52 mm</quantity>
|
||
,
|
||
<emphasis id="B909C9126C149462FBE7C985FB9DF9B9" box="[1066,1085,1519,1542]" italics="true" pageId="41" pageNumber="42">N</emphasis>
|
||
= 3, female SVL
|
||
<quantity id="4C85B8E56C149462FAC9C984FADEF9B9" box="[1284,1406,1518,1543]" metricMagnitude="-2" metricUnit="m" metricValue="5.4" metricValueMax="5.5" metricValueMin="5.3" pageId="41" pageNumber="42" unit="mm" value="54.0" valueMax="55.0" valueMin="53.0">53–55 mm</quantity>
|
||
,
|
||
<emphasis id="B909C9126C149462FA44C985FA3CF9B9" box="[1417,1436,1519,1542]" italics="true" pageId="41" pageNumber="42">N</emphasis>
|
||
= 2).
|
||
<taxonomicName id="4C7D6E836C149462FF19CA78FE33F994" authority="Garg & Senevirathne & Wijayathilaka & Phuge & Deuti & Manamendra-Arachchi & Meegaskumbura & Biju, 2018" authorityName="Garg & Senevirathne & Wijayathilaka & Phuge & Deuti & Manamendra-Arachchi & Meegaskumbura & Biju" authorityYear="2018" box="[212,403,1554,1579]" class="Amphibia" family="Microhylidae" genus="Uperodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="41" pageNumber="42" phylum="Chordata" rank="species" species="rohani" status="sp. nov.">
|
||
<emphasis id="B909C9126C149462FF19CA78FE33F994" box="[212,403,1554,1579]" italics="true" pageId="41" pageNumber="42">Uperodon rohani</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B909C9126C149462FE56CA78FE55F994" bold="true" box="[411,501,1554,1579]" pageId="41" pageNumber="42">
|
||
<taxonomicNameLabel id="A23A74696C149462FE56CA78FE55F994" box="[411,501,1554,1579]" pageId="41" pageNumber="42" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
specifically also differs from the former two by its finger tips having truncate discs (vs. rounded without discs), and presence of two weakly-developed neopalatinal ridges on posterior side of each choana, fused with the vomerine odontophores (vs. two prominent dermal projections on the neopalatinal ridges on posterior side of each choana, separated from the vomerine odontophores); and from
|
||
<taxonomicName id="4C7D6E836C149462FB8FCA15FB55F928" baseAuthorityName="Parker" baseAuthorityYear="1934" box="[1090,1269,1662,1687]" class="Amphibia" family="Microhylidae" genus="Uperodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="41" pageNumber="42" phylum="Chordata" rank="species" species="taprobanicus">
|
||
<emphasis id="B909C9126C149462FB8FCA15FBF6F929" box="[1090,1110,1663,1686]" italics="true" pageId="41" pageNumber="42">U</emphasis>
|
||
.
|
||
<emphasis id="B909C9126C149462FBA9CA14FB55F928" box="[1124,1269,1662,1687]" italics="true" pageId="41" pageNumber="42">taprobanicus</emphasis>
|
||
</taxonomicName>
|
||
by presence of two weakly-developed neopalatinal ridges on posterior side of each choana, fused with the vomerine odontophores (vs. well-developed neopalatinal ridges). Further,
|
||
<taxonomicName id="4C7D6E836C149462FD70CAADFC87F960" authority="Garg & Senevirathne & Wijayathilaka & Phuge & Deuti & Manamendra-Arachchi & Meegaskumbura & Biju, 2018" authorityName="Garg & Senevirathne & Wijayathilaka & Phuge & Deuti & Manamendra-Arachchi & Meegaskumbura & Biju" authorityYear="2018" box="[701,807,1734,1759]" class="Amphibia" family="Microhylidae" genus="Uperodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="41" pageNumber="42" phylum="Chordata" rank="species" species="rohani" status="sp. nov.">
|
||
<emphasis id="B909C9126C149462FD70CAADFC87F960" box="[701,807,1734,1759]" italics="true" pageId="41" pageNumber="42">U. rohani</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B909C9126C149462FCE3CAACFC26F960" bold="true" box="[814,902,1734,1759]" pageId="41" pageNumber="42">
|
||
<taxonomicNameLabel id="A23A74696C149462FCE3CAACFC26F960" box="[814,902,1734,1759]" pageId="41" pageNumber="42" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
differs from
|
||
<taxonomicName id="4C7D6E836C149462FBDACAADFB79F960" authorityName="M Parambikulam" baseAuthorityName="Rao" baseAuthorityYear="1937" box="[1047,1241,1734,1759]" class="Amphibia" family="Microhylidae" genus="Uperodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="41" pageNumber="42" phylum="Chordata" rank="species" species="anamalaiensis">
|
||
<emphasis id="B909C9126C149462FBDACAADFB79F960" box="[1047,1241,1734,1759]" italics="true" pageId="41" pageNumber="42">U. anamalaiensis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C7D6E836C149462FB2BCAADFAD4F961" box="[1254,1396,1735,1758]" class="Amphibia" family="Microhylidae" genus="Uperodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="41" pageNumber="42" phylum="Chordata" rank="species" species="montanus">
|
||
<emphasis id="B909C9126C149462FB2BCAADFAD4F961" box="[1254,1396,1735,1758]" italics="true" pageId="41" pageNumber="42">U. montanus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C7D6E836C149462FA4CCAADFF5BF8BC" class="Amphibia" family="Microhylidae" genus="Uperodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="41" pageNumber="42" phylum="Chordata" rank="species" species="obscurus">
|
||
<emphasis id="B909C9126C149462FA4CCAADFF5BF8BC" italics="true" pageId="41" pageNumber="42">U. obscurus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C7D6E836C149462FE8CCA86FE70F8BC" box="[321,464,1770,1795]" class="Amphibia" family="Microhylidae" genus="Uperodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="41" pageNumber="42" phylum="Chordata" rank="species" species="palmatus">
|
||
<emphasis id="B909C9126C149462FE8CCA86FE70F8BC" box="[321,464,1770,1795]" italics="true" pageId="41" pageNumber="42">U. palmatus</emphasis>
|
||
</taxonomicName>
|
||
by its ventral skin being white to off-white in life (that turns greyish-white in preservation) without any spots or blotches (vs. light or dark brown with prominent spots or blotches), and absence of webbing between toes in both the sexes (vs. present).
|
||
<taxonomicName id="4C7D6E836C149462FCEFCB58FC43F8F4" authority="Garg & Senevirathne & Wijayathilaka & Phuge & Deuti & Manamendra-Arachchi & Meegaskumbura & Biju, 2018" authorityName="Garg & Senevirathne & Wijayathilaka & Phuge & Deuti & Manamendra-Arachchi & Meegaskumbura & Biju" authorityYear="2018" box="[802,995,1842,1867]" class="Amphibia" family="Microhylidae" genus="Uperodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="41" pageNumber="42" phylum="Chordata" rank="species" species="rohani" status="sp. nov.">
|
||
<emphasis id="B909C9126C149462FCEFCB58FC43F8F4" box="[802,995,1842,1867]" italics="true" pageId="41" pageNumber="42">Uperodon rohani</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B909C9126C149462FC20CB58FBE8F8F4" bold="true" box="[1005,1096,1842,1867]" pageId="41" pageNumber="42">
|
||
<taxonomicNameLabel id="A23A74696C149462FC20CB58FBE8F8F4" box="[1005,1096,1842,1867]" pageId="41" pageNumber="42" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
differs from
|
||
<taxonomicName id="4C7D6E836C149462FB2FCB5EFAF3F8F4" baseAuthorityName="Manamendra-Arachchi and Pethiyagoda" baseAuthorityYear="2001" box="[1250,1363,1844,1867]" class="Amphibia" family="Microhylidae" genus="Uperodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="41" pageNumber="42" phylum="Chordata" rank="species" species="nagaoi">
|
||
<emphasis id="B909C9126C149462FB2FCB5EFAF3F8F4" box="[1250,1363,1844,1867]" italics="true" pageId="41" pageNumber="42">U. nagaoi</emphasis>
|
||
</taxonomicName>
|
||
by its ventral skin being white to off-white in life (that turns greyish-white in preservation) without any spots or speckles (vs. brown to dark brown in life that turns brown or dark greyish-brown in preservation, with or without inconspicuous spots or speckles). For detailed comparison of
|
||
<taxonomicName id="4C7D6E836C149462FC89CBF5FC0EF808" authority="Garg & Senevirathne & Wijayathilaka & Phuge & Deuti & Manamendra-Arachchi & Meegaskumbura & Biju, 2018" authorityName="Garg & Senevirathne & Wijayathilaka & Phuge & Deuti & Manamendra-Arachchi & Meegaskumbura & Biju" authorityYear="2018" box="[836,942,1950,1975]" class="Amphibia" family="Microhylidae" genus="Uperodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="41" pageNumber="42" phylum="Chordata" rank="species" species="rohani" status="sp. nov.">
|
||
<emphasis id="B909C9126C149462FC89CBF5FC0EF808" box="[836,942,1950,1975]" italics="true" pageId="41" pageNumber="42">U. rohani</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B909C9126C149462FC7BCBF4FBAFF808" bold="true" box="[950,1039,1950,1975]" pageId="41" pageNumber="42">
|
||
<taxonomicNameLabel id="A23A74696C149462FC7BCBF4FBAFF808" box="[950,1039,1950,1975]" pageId="41" pageNumber="42" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
with
|
||
<taxonomicName id="4C7D6E836C149462FB82CBF5FB49F808" box="[1103,1257,1950,1975]" class="Amphibia" family="Microhylidae" genus="Uperodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="41" pageNumber="42" phylum="Chordata" rank="species" species="globulosus">
|
||
<emphasis id="B909C9126C149462FB82CBF5FB49F808" box="[1103,1257,1950,1975]" italics="true" pageId="41" pageNumber="42">U. globulosus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C7D6E836C149462FB3ACBF5FAD3F809" baseAuthorityName="Schneider" baseAuthorityYear="1799" box="[1271,1395,1951,1974]" class="Amphibia" family="Microhylidae" genus="Uperodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="41" pageNumber="42" phylum="Chordata" rank="species" species="systoma">
|
||
<emphasis id="B909C9126C149462FB3ACBF5FAD3F809" box="[1271,1395,1951,1974]" italics="true" pageId="41" pageNumber="42">U. systoma</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C7D6E836C149462FA4CCBF5FE88F864" baseAuthorityName="Parker" baseAuthorityYear="1934" class="Amphibia" family="Microhylidae" genus="Uperodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="41" pageNumber="42" phylum="Chordata" rank="species" species="taprobanicus">
|
||
<emphasis id="B909C9126C149462FA4CCBF5FE88F864" italics="true" pageId="41" pageNumber="42">U. taprobanicus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C7D6E836C149462FEF0CBAEFDA6F864" authorityName="M Parambikulam" baseAuthorityName="Rao" baseAuthorityYear="1937" box="[317,518,1986,2011]" class="Amphibia" family="Microhylidae" genus="Uperodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="41" pageNumber="42" phylum="Chordata" rank="species" species="anamalaiensis">
|
||
<emphasis id="B909C9126C149462FEF0CBAEFDA6F864" box="[317,518,1986,2011]" italics="true" pageId="41" pageNumber="42">U. anamalaiensis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C7D6E836C149462FDD6CBAEFD10F864" box="[539,688,1988,2011]" class="Amphibia" family="Microhylidae" genus="Uperodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="41" pageNumber="42" phylum="Chordata" rank="species" species="montanus">
|
||
<emphasis id="B909C9126C149462FDD6CBAEFD10F864" box="[539,688,1988,2011]" italics="true" pageId="41" pageNumber="42">U. montanus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C7D6E836C149462FD08CBAEFCCBF864" baseAuthorityName="Rao" baseAuthorityYear="1937" box="[709,875,1988,2011]" class="Amphibia" family="Microhylidae" genus="Uperodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="41" pageNumber="42" phylum="Chordata" rank="species" species="mormorata">
|
||
<emphasis id="B909C9126C149462FD08CBAEFCCBF864" box="[709,875,1988,2011]" italics="true" pageId="41" pageNumber="42">U. mormorata</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C7D6E836C149462FC4DCBAEFC56F864" baseAuthorityName="Manamendra-Arachchi and Pethiyagoda" baseAuthorityYear="2001" box="[896,1014,1988,2011]" class="Amphibia" family="Microhylidae" genus="Uperodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="41" pageNumber="42" phylum="Chordata" rank="species" species="nagaoi">
|
||
<emphasis id="B909C9126C149462FC4DCBAEFC56F864" box="[896,1014,1988,2011]" italics="true" pageId="41" pageNumber="42">U. nagaoi</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C7D6E836C149462FBC1CBAEFB38F864" box="[1036,1176,1986,2011]" class="Amphibia" family="Microhylidae" genus="Uperodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="41" pageNumber="42" phylum="Chordata" rank="species" species="obscurus">
|
||
<emphasis id="B909C9126C149462FBC1CBAEFB38F864" box="[1036,1176,1986,2011]" italics="true" pageId="41" pageNumber="42">U. obscurus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C7D6E836C149462FB63CBAEFA9CF864" box="[1198,1340,1986,2011]" class="Amphibia" family="Microhylidae" genus="Uperodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="41" pageNumber="42" phylum="Chordata" rank="species" species="palmatus">
|
||
<emphasis id="B909C9126C149462FB63CBAEFA9CF864" box="[1198,1340,1986,2011]" italics="true" pageId="41" pageNumber="42">U. palmatus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<emphasis id="B909C9126C149462FA4CCBAEFA3CF864" box="[1409,1436,1988,2011]" italics="true" pageId="41" pageNumber="42">U.</emphasis>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C367468B6C179460FF5ACCFDFD14F81F" lastPageId="43" lastPageNumber="44" pageId="42" pageNumber="43" type="description">
|
||
<paragraph id="8BC215006C179461FF5ACCFDFC5AFE37" blockId="42.[151,1437,151,2013]" pageId="42" pageNumber="43">
|
||
<taxonomicName id="4C7D6E836C179461FF5ACCFDFEBBFF0F" box="[151,283,151,176]" class="Amphibia" family="Microhylidae" genus="Uperodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="42" pageNumber="43" phylum="Chordata" rank="species" species="triangularis">
|
||
<emphasis id="B909C9126C179461FF5ACCFDFEBBFF0F" box="[151,283,151,176]" italics="true" pageId="42" pageNumber="43">triangularis</emphasis>
|
||
</taxonomicName>
|
||
see ‘comparison’ section of those species.
|
||
<taxonomicName id="4C7D6E836C179461FD35CCFDFC17FF0F" authority="Garg & Senevirathne & Wijayathilaka & Phuge & Deuti & Manamendra-Arachchi & Meegaskumbura & Biju, 2018" authorityName="Garg & Senevirathne & Wijayathilaka & Phuge & Deuti & Manamendra-Arachchi & Meegaskumbura & Biju" authorityYear="2018" box="[760,951,151,176]" class="Amphibia" family="Microhylidae" genus="Uperodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="42" pageNumber="43" phylum="Chordata" rank="species" species="rohani" status="sp. nov.">
|
||
<emphasis id="B909C9126C179461FD35CCFDFC17FF0F" box="[760,951,151,176]" italics="true" pageId="42" pageNumber="43">Uperodon rohani</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B909C9126C179461FC73CCFDFBB6FF0F" bold="true" box="[958,1046,151,176]" pageId="42" pageNumber="43">
|
||
<taxonomicNameLabel id="A23A74696C179461FC73CCFDFBB6FF0F" box="[958,1046,151,176]" pageId="42" pageNumber="43" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is morphologically and genetically close to
|
||
<taxonomicName id="4C7D6E836C179461FF3FCCD7FE29FF6B" baseAuthorityName="Stoliczka" baseAuthorityYear="1872" box="[242,393,189,212]" class="Amphibia" family="Microhylidae" genus="Uperodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="42" pageNumber="43" phylum="Chordata" rank="species" species="variegatus">
|
||
<emphasis id="B909C9126C179461FF3FCCD7FE29FF6B" box="[242,393,189,212]" italics="true" pageId="42" pageNumber="43">U. variegatus</emphasis>
|
||
</taxonomicName>
|
||
, from which it differs by its snout nearly equal to eye diameter, male SL/EL ratio 1.0–1.2,
|
||
<emphasis id="B909C9126C179461FABECCD7FA26FF6B" box="[1395,1414,189,212]" italics="true" pageId="42" pageNumber="43">N</emphasis>
|
||
= 16, female SL/EL ratio 0.8–1.0,
|
||
<emphasis id="B909C9126C179461FE36CC8BFDAEFF47" box="[507,526,225,248]" italics="true" pageId="42" pageNumber="43">N</emphasis>
|
||
= 8 (vs. snout relatively longer than eye, male SL/EL ratio 1.2–1.6,
|
||
<emphasis id="B909C9126C179461FB34CC8BFAACFF47" box="[1273,1292,225,248]" italics="true" pageId="42" pageNumber="43">N</emphasis>
|
||
= 12, female SL/EL ratio 1.2–1.4,
|
||
<emphasis id="B909C9126C179461FE4FCD6FFE35FEA3" box="[386,405,261,284]" italics="true" pageId="42" pageNumber="43">N</emphasis>
|
||
= 7); head relatively narrower, male HW/HL 1.0–1.1,
|
||
<emphasis id="B909C9126C179461FC3CCD6FFBA4FEA3" box="[1009,1028,261,284]" italics="true" pageId="42" pageNumber="43">N</emphasis>
|
||
= 16, female HW/HL 1.0–1.2,
|
||
<emphasis id="B909C9126C179461FA93CD6FFAD1FEA3" box="[1374,1393,261,284]" italics="true" pageId="42" pageNumber="43">N</emphasis>
|
||
= 8 (vs. head relatively wider, male HW/HL 1.1–1.4,
|
||
<emphasis id="B909C9126C179461FD19CD43FD47FEFF" box="[724,743,297,320]" italics="true" pageId="42" pageNumber="43">N</emphasis>
|
||
= 12, female HW/HL 1.1–1.4,
|
||
<emphasis id="B909C9126C179461FB9BCD43FBC9FEFF" box="[1110,1129,297,320]" italics="true" pageId="42" pageNumber="43">N</emphasis>
|
||
= 7); and shank relatively longer than thigh, male SHL/TL ratio 1.0–1.1,
|
||
<emphasis id="B909C9126C179461FD65CD27FD1BFEDB" box="[680,699,333,356]" italics="true" pageId="42" pageNumber="43">N</emphasis>
|
||
= 8, female SHL/TL ratio 1.1,
|
||
<emphasis id="B909C9126C179461FBEDCD27FB93FEDB" box="[1056,1075,333,356]" italics="true" pageId="42" pageNumber="43">N</emphasis>
|
||
= 8 (vs. relatively shorter, male SHL/TL ratio 0.9–1.0,
|
||
<emphasis id="B909C9126C179461FE5BCD1BFE09FE37" box="[406,425,369,392]" italics="true" pageId="42" pageNumber="43">N</emphasis>
|
||
= 12, female SHL/TL ratio 0.8–0.9,
|
||
<emphasis id="B909C9126C179461FC8DCD1BFCF3FE37" box="[832,851,369,392]" italics="true" pageId="42" pageNumber="43">N</emphasis>
|
||
= 7) (
|
||
<tableCitation id="C6FF20BB6C179461FC5BCD05FC4DFE37" box="[918,1005,367,392]" captionStart="TABLE 4" captionStartId="48.[119,193,547,566]" captionTargetBox="[135,1884,616,1625]" captionText="TABLE 4. Morphometric measurements of Uperodon species.All measurements are in millimetres (mm). The status of each specimen is given after the museum number:Holotype (HT), Paratype (PT), Neotype (NT), Lectotype (LT), Syntype (ST)." httpUri="http://table.plazi.org/id/DF0245886C0D947BFFBACE49F8ECFD89" pageId="42" pageNumber="43" tableUuid="DF0245886C0D947BFFBACE49F8ECFD89">Table 4</tableCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BC215006C179461FF0ACDFEFDF4FC3F" blockId="42.[151,1437,151,2013]" pageId="42" pageNumber="43">
|
||
The new species cannot be confused with previously available names,
|
||
<taxonomicName id="4C7D6E836C179461FC27CDFFFA3CFE13" authority="Cuvier, 1829" authorityName="Cuvier" authorityYear="1829" box="[1002,1436,404,429]" class="Amphibia" family="Microhylidae" genus="Engystoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="42" pageNumber="43" phylum="Chordata" rank="species" species="marmoratum">
|
||
<emphasis id="B909C9126C179461FC27CDFFFAA1FE13" box="[1002,1281,405,428]" italics="true" pageId="42" pageNumber="43">Engystoma marmoratum</emphasis>
|
||
Cuvier, 1829
|
||
</taxonomicName>
|
||
and
|
||
<emphasis id="B909C9126C179461FF04CDD2FE1FFE6F" box="[201,447,439,464]" italics="true" pageId="42" pageNumber="43">Systoma leschenaultii</emphasis>
|
||
Tschudi, 1838, currently under the synonymy of
|
||
<taxonomicName id="4C7D6E836C179461FC32CDDDFA38FE6F" authority="Schneider, 1799" authorityName="Schneider" authorityYear="1799" box="[1023,1432,439,464]" class="Amphibia" family="Microhylidae" genus="Uperodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="42" pageNumber="43" phylum="Chordata" rank="species" species="systoma">
|
||
<emphasis id="B909C9126C179461FC32CDDDFB73FE6F" box="[1023,1235,439,464]" italics="true" pageId="42" pageNumber="43">Uperodon systoma</emphasis>
|
||
<bibRefCitation id="EFEC68F16C179461FB10CDDDFA38FE6F" author="Schneider, J. G." bookContentInfo="266 pp." box="[1245,1432,439,464]" journalOrPublisher="Friederici Frommanni, Jena" pageId="42" pageNumber="43" refId="ref50384" refString="Schneider, J. G. (1799) Historia amphibiorum naturalis et literarariae. Fasciculus primus. Continens Ranas, Calamitas, Bufones, Salamandras et Hydros in genera et species Descriptos notisque suis distinctos. Friederici Frommanni, Jena, 266 pp." title="Historia amphibiorum naturalis et literarariae. Fasciculus primus. Continens Ranas, Calamitas, Bufones, Salamandras et Hydros in genera et species Descriptos notisque suis distinctos" type="book" year="1799">Schneider, 1799</bibRefCitation>
|
||
</taxonomicName>
|
||
. The former was described from “l’Inde” (=
|
||
<collectingCountry id="F36A55906C179461FD4BCDB6FD61FE4A" box="[646,705,476,501]" name="India" pageId="42" pageNumber="43">India</collectingCountry>
|
||
) and as stated by
|
||
<bibRefCitation id="EFEC68F16C179461FC40CDB7FBBAFE4A" author="Peters, W. C. H." box="[909,1050,476,501]" journalOrPublisher="Monatsberichte der Koniglichen Preussische Akademie des Wissenschaften zu Berlin, 1863" pageId="42" pageNumber="43" pagination="445 - 470" part="1863" refId="ref38630" refString="Peters, W. C. H. (1864 " 1863 ") Fernere mittheilungen uber neue batrachier. Monatsberichte der Koniglichen Preussische Akademie des Wissenschaften zu Berlin, 1863, 445 - 470." title="Fernere mittheilungen uber neue batrachier" type="journal article" year="1864">Peters (1864</bibRefCitation>
|
||
“1863”)—“bereits richtig erkannt batten =
|
||
<taxonomicName id="4C7D6E836C179461FF32CD95FDACFDA7" box="[255,524,511,536]" class="Amphibia" family="Microhylidae" genus="Uperodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="42" pageNumber="43" phylum="Chordata" rank="species" species="marmoratum">
|
||
<emphasis id="B909C9126C179461FF32CD95FDACFDA7" box="[255,524,511,536]" italics="true" pageId="42" pageNumber="43">Uperodon marmoratum</emphasis>
|
||
</taxonomicName>
|
||
D. B. und miifste daher den Species namen “
|
||
<taxonomicName id="4C7D6E836C179461FBEECE6AFB23FDA7" authorityName="Schneider" authorityYear="1799" box="[1059,1155,512,536]" class="Amphibia" family="Microhylidae" genus="Uperodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="42" pageNumber="43" phylum="Chordata" rank="species" species="systoma">systoma</taxonomicName>
|
||
” behalten”, was already correctly recognized as
|
||
<taxonomicName id="4C7D6E836C179461FE68CE4EFD10FD83" box="[421,688,548,573]" class="Amphibia" family="Microhylidae" genus="Uperodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="42" pageNumber="43" phylum="Chordata" rank="species" species="marmoratum">
|
||
<emphasis id="B909C9126C179461FE68CE4EFD10FD83" box="[421,688,548,573]" italics="true" pageId="42" pageNumber="43">Uperodon marmoratum</emphasis>
|
||
</taxonomicName>
|
||
(=
|
||
<taxonomicName id="4C7D6E836C179461FD17CE4EFC0DFD83" authorityName="Schneider" authorityYear="1799" box="[730,941,548,573]" class="Amphibia" family="Microhylidae" genus="Uperodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="42" pageNumber="43" phylum="Chordata" rank="species" species="systoma">
|
||
<emphasis id="B909C9126C179461FD17CE4EFC0DFD83" box="[730,941,548,573]" italics="true" pageId="42" pageNumber="43">Uperodon systoma</emphasis>
|
||
</taxonomicName>
|
||
) by
|
||
<bibRefCitation id="EFEC68F16C179461FC2FCE4EFABDFD83" author="Dumeril, A. M. C. & Bibron, G." bookContentInfo="430 pp." box="[994,1309,548,573]" journalOrPublisher="Roret, Paris" pageId="42" pageNumber="43" refId="ref36844" refString="Dumeril, A. M. C. & Bibron, G. (1841) Erpetologie Generale ou Histoire Naturelle complete des Reptiles. Roret, Paris, 430 pp." title="Erpetologie Generale ou Histoire Naturelle complete des Reptiles" type="book" year="1841">Duméril and Bibron (1841)</bibRefCitation>
|
||
. The latter, also stated by
|
||
<bibRefCitation id="EFEC68F16C179461FE8BCE2DFD23FDDF" author="Dumeril, A. M. C. & Bibron, G." bookContentInfo="430 pp." box="[326,643,583,608]" journalOrPublisher="Roret, Paris" pageId="42" pageNumber="43" refId="ref36844" refString="Dumeril, A. M. C. & Bibron, G. (1841) Erpetologie Generale ou Histoire Naturelle complete des Reptiles. Roret, Paris, 430 pp." title="Erpetologie Generale ou Histoire Naturelle complete des Reptiles" type="book" year="1841">Duméril and Bibron (1841)</bibRefCitation>
|
||
—“peut-être de la même espèce que notre Upérodonte marbré”, is perhaps the same as
|
||
<taxonomicName id="4C7D6E836C179461FE5ACE06FDCFFD3B" authorityName="Schneider" authorityYear="1799" box="[407,623,620,645]" class="Amphibia" family="Microhylidae" genus="Uperodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="42" pageNumber="43" phylum="Chordata" rank="species" species="systoma">
|
||
<emphasis id="B909C9126C179461FE5ACE06FDCFFD3B" box="[407,623,620,645]" italics="true" pageId="42" pageNumber="43">Uperodon systoma</emphasis>
|
||
</taxonomicName>
|
||
, from which the new species cannot be confused based on several morphological characters such as smaller body size (vs. larger), finger tips with truncate discs (vs. rounded without discs) and vomerine odontophores fused with neopalatinal ridges (vs. separated). Furthermore, three other names—
|
||
<emphasis id="B909C9126C179461FF5ACEBDFE14FD4F" box="[151,436,727,752]" italics="true" pageId="42" pageNumber="43">Hylaedactylus carnaticus</emphasis>
|
||
<bibRefCitation id="EFEC68F16C179461FE76CEBDFDECFD4F" author="Jerdon, T. C." box="[443,588,727,752]" journalOrPublisher="Journal of the Asiatic Society of Bengal, 22" pageId="42" pageNumber="43" pagination="522 - 534" part="1853" refId="ref45860" refString="Jerdon, T. C. (1854 " 1853 ") Catalogue of reptiles inhabiting the Peninsula of India. Journal of the Asiatic Society of Bengal, 22, 522 - 534.?" title="Catalogue of reptiles inhabiting the Peninsula of India" type="journal article" year="1854">Jerdon, 1854</bibRefCitation>
|
||
“1853”,
|
||
<taxonomicName id="4C7D6E836C179461FD7FCEBDFBB4FD4F" authority="Gunther, 1876" authorityName="Gunther" authorityYear="1876" box="[690,1044,727,752]" class="Amphibia" family="Microhylidae" genus="Callula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="42" pageNumber="43" phylum="Chordata" rank="species" species="olivacea">
|
||
<emphasis id="B909C9126C179461FD7FCEBDFCCCFD4F" box="[690,876,727,752]" italics="true" pageId="42" pageNumber="43">Callula olivacea</emphasis>
|
||
<bibRefCitation id="EFEC68F16C179461FCBECEBDFBB4FD4F" author="Gunther, A. C. L. G." box="[883,1044,727,752]" journalOrPublisher="Proceedings of the Zoological Society of London, 1875" pageId="42" pageNumber="43" pagination="567 - 577" part="1875" refId="ref44979" refString="Gunther, A. C. L. G. (1876 " 1875 ") Third report on collections of Indian reptiles obtained by the British Museum. Proceedings of the Zoological Society of London, 1875, 567 - 577." title="Third report on collections of Indian reptiles obtained by the British Museum" type="journal article" year="1876">Günther, 1876</bibRefCitation>
|
||
</taxonomicName>
|
||
“1875” and
|
||
<taxonomicName id="4C7D6E836C179461FB6FCEBDFE1FFCAA" authority="Rao and Ramanna, 1925" authorityName="Rao and Ramanna" authorityYear="1925" class="Amphibia" family="Microhylidae" genus="Ramanella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="42" pageNumber="43" phylum="Chordata" rank="species" species="symbioitca">
|
||
<emphasis id="B909C9126C179461FB6FCEBDFA3CFD4F" box="[1186,1436,727,752]" italics="true" pageId="42" pageNumber="43">Ramanella symbioitca</emphasis>
|
||
<bibRefCitation id="EFEC68F16C179461FF5ACE97FE1FFCAA" author="Rao, C. R. & Ramanna, B. S." box="[151,447,764,789]" journalOrPublisher="Journal of Zoology" pageId="42" pageNumber="43" pagination="587 - 597" part="95" publicationUrl="https://doi.org/10.1111/j.1096-3642.1925.tb01529.x" refId="ref38907" refString="Rao, C. R. & Ramanna, B. S. (1925) On a new Genus of the Family Engystomatidae (Batrachia). Journal of Zoology, 95 (2), 587 - 597 https: // doi. org / 10.1111 / j. 1096 - 3642.1925. tb 01529. x" title="On a new Genus of the Family Engystomatidae (Batrachia)" type="journal article" year="1925">Rao and Ramanna, 1925</bibRefCitation>
|
||
</taxonomicName>
|
||
, currently under the synonymy of
|
||
<taxonomicName id="4C7D6E836C179461FCAECE96FABAFCAB" authority="Stoliczka, 1872" authorityName="Stoliczka" authorityYear="1872" box="[867,1306,764,789]" class="Amphibia" family="Microhylidae" genus="Uperodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="42" pageNumber="43" phylum="Chordata" rank="species" species="variegatus">
|
||
<emphasis id="B909C9126C179461FCAECE96FBF6FCAB" box="[867,1110,764,789]" italics="true" pageId="42" pageNumber="43">Uperodon variegatus</emphasis>
|
||
<bibRefCitation id="EFEC68F16C179461FBA9CE96FABAFCAB" author="Stoliczka, F." box="[1124,1306,764,789]" journalOrPublisher="Proceedings of the Asiatic Society of Bengal, 1872" pageId="42" pageNumber="43" pagination="101 - 113" refId="ref51385" refString="Stoliczka, F. (1872) Observations on Indian Batrachia. Proceedings of the Asiatic Society of Bengal, 1872, 101 - 113." title="Observations on Indian Batrachia" type="proceedings paper" year="1872">Stoliczka, 1872</bibRefCitation>
|
||
</taxonomicName>
|
||
, were also described from regions in
|
||
<collectingCountry id="F36A55906C179461FE0CCF75FE5BFC87" box="[449,507,799,824]" name="India" pageId="42" pageNumber="43">India</collectingCountry>
|
||
(“Carnatic”, “Yellagherry hills” and “Ellore”, respectively), while the distribution of
|
||
<taxonomicName id="4C7D6E836C179461FF78CF2FFE81FCE2" authority="Garg & Senevirathne & Wijayathilaka & Phuge & Deuti & Manamendra-Arachchi & Meegaskumbura & Biju, 2018" authorityName="Garg & Senevirathne & Wijayathilaka & Phuge & Deuti & Manamendra-Arachchi & Meegaskumbura & Biju" authorityYear="2018" box="[181,289,836,861]" class="Amphibia" family="Microhylidae" genus="Uperodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="42" pageNumber="43" phylum="Chordata" rank="species" species="rohani" status="sp. nov.">
|
||
<emphasis id="B909C9126C179461FF78CF2FFE81FCE2" box="[181,289,836,861]" italics="true" pageId="42" pageNumber="43">U. rohani</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B909C9126C179461FEE4CF2EFE22FCE2" bold="true" box="[297,386,836,861]" pageId="42" pageNumber="43">
|
||
<taxonomicNameLabel id="A23A74696C179461FEE4CF2EFE22FCE2" box="[297,386,836,861]" pageId="42" pageNumber="43" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is restricted to
|
||
<collectingCountry id="F36A55906C179461FDFDCF2EFD02FCE2" box="[560,674,836,861]" name="Sri Lanka" pageId="42" pageNumber="43">
|
||
<collectingRegion id="49B9DBE26C179461FDFDCF2EFDF1FCE2" box="[560,593,836,861]" country="United Kingdom" name="Surrey" pageId="42" pageNumber="43">Sri</collectingRegion>
|
||
Lanka
|
||
</collectingCountry>
|
||
. For specific differences with
|
||
<taxonomicName id="4C7D6E836C179461FC39CF2FFBD0FCE3" authorityName="Schneider" authorityYear="1799" box="[1012,1136,837,860]" class="Amphibia" family="Microhylidae" genus="Uperodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="42" pageNumber="43" phylum="Chordata" rank="species" species="systoma">
|
||
<emphasis id="B909C9126C179461FC39CF2FFBD0FCE3" box="[1012,1136,837,860]" italics="true" pageId="42" pageNumber="43">U. systoma</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C7D6E836C179461FB65CF2FFAE0FCE3" authorityName="Stoliczka" authorityYear="1872" box="[1192,1344,837,860]" class="Amphibia" family="Microhylidae" genus="Uperodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="42" pageNumber="43" phylum="Chordata" rank="species" species="variegatus">
|
||
<emphasis id="B909C9126C179461FB65CF2FFAE0FCE3" box="[1192,1344,837,860]" italics="true" pageId="42" pageNumber="43">U. variegatus</emphasis>
|
||
</taxonomicName>
|
||
, see the detailed comparison with those species.
|
||
</paragraph>
|
||
<paragraph id="8BC215006C179461FF0ACFE6FB20FAC7" blockId="42.[151,1437,151,2013]" pageId="42" pageNumber="43">
|
||
<emphasis id="B909C9126C179461FF0ACFE6FD97FC1A" bold="true" box="[199,567,908,933]" pageId="42" pageNumber="43">Morphometric differentiation.</emphasis>
|
||
<taxonomicName id="4C7D6E836C179461FD8ECFE6FCA9FC1A" authority="Garg & Senevirathne & Wijayathilaka & Phuge & Deuti & Manamendra-Arachchi & Meegaskumbura & Biju, 2018" authorityName="Garg & Senevirathne & Wijayathilaka & Phuge & Deuti & Manamendra-Arachchi & Meegaskumbura & Biju" authorityYear="2018" box="[579,777,908,933]" class="Amphibia" family="Microhylidae" genus="Uperodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="42" pageNumber="43" phylum="Chordata" rank="species" species="rohani" status="sp. nov.">
|
||
<emphasis id="B909C9126C179461FD8ECFE6FCA9FC1A" box="[579,777,908,933]" italics="true" pageId="42" pageNumber="43">Uperodon rohani</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B909C9126C179461FCD8CFE6FCD5FC1A" bold="true" box="[789,885,908,933]" pageId="42" pageNumber="43">
|
||
<taxonomicNameLabel id="A23A74696C179461FCD8CFE6FCD5FC1A" box="[789,885,908,933]" pageId="42" pageNumber="43" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
showed morphological differentiation from its closely related species,
|
||
<taxonomicName id="4C7D6E836C179461FE61CFDBFDE8FC77" authorityName="Stoliczka" authorityYear="1872" box="[428,584,945,968]" class="Amphibia" family="Microhylidae" genus="Uperodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="42" pageNumber="43" phylum="Chordata" rank="species" species="variegatus">
|
||
<emphasis id="B909C9126C179461FE61CFDBFDE8FC77" box="[428,584,945,968]" italics="true" pageId="42" pageNumber="43">U. variegatus</emphasis>
|
||
</taxonomicName>
|
||
. Principal Component Analysis (PCA) for the adult male specimens (
|
||
<taxonomicName id="4C7D6E836C179461FA4CCFDBFF41FC52" authority="Garg & Senevirathne & Wijayathilaka & Phuge & Deuti & Manamendra-Arachchi & Meegaskumbura & Biju, 2018" authorityName="Garg & Senevirathne & Wijayathilaka & Phuge & Deuti & Manamendra-Arachchi & Meegaskumbura & Biju" authorityYear="2018" class="Amphibia" family="Microhylidae" genus="Uperodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="42" pageNumber="43" phylum="Chordata" rank="species" species="rohani" status="sp. nov.">
|
||
<emphasis id="B909C9126C179461FA4CCFDBFF41FC52" italics="true" pageId="42" pageNumber="43">U. rohani</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B909C9126C179461FF25CFBEFEE3FC52" bold="true" box="[232,323,980,1005]" pageId="42" pageNumber="43">
|
||
<taxonomicNameLabel id="A23A74696C179461FF25CFBEFEE3FC52" box="[232,323,980,1005]" pageId="42" pageNumber="43" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
:
|
||
<emphasis id="B909C9126C179461FE9ECFBFFEC6FC53" box="[339,358,981,1004]" italics="true" pageId="42" pageNumber="43">N</emphasis>
|
||
= 12;
|
||
<taxonomicName id="4C7D6E836C179461FE7CCFBFFDEAFC53" authorityName="Stoliczka" authorityYear="1872" box="[433,586,981,1004]" class="Amphibia" family="Microhylidae" genus="Uperodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="42" pageNumber="43" phylum="Chordata" rank="species" species="variegatus">
|
||
<emphasis id="B909C9126C179461FE7CCFBFFDEAFC53" box="[433,586,981,1004]" italics="true" pageId="42" pageNumber="43">U. variegatus</emphasis>
|
||
</taxonomicName>
|
||
:
|
||
<emphasis id="B909C9126C179461FD97CFBFFDCDFC53" box="[602,621,981,1004]" italics="true" pageId="42" pageNumber="43">N</emphasis>
|
||
= 16) recovered five Principal Components (PC) with eigenvalues>1.0 that accounted for 77.2% of the total variance; PC 1 explained 32.3% of the variations, with highest loadings for SVL, HW, IUE, IFE, MN, HAL and TL; PC 2 explained 22.1% of the variations, with highest loadings for SL, IBE, EN, SHL and FOL; PC 3 explained 10% of the variations, with highest loadings for UEW and NS; PC 4 explained 6.9% of the variations, with highest loadings for EL; PC 5 explained 6% of the variations; and the remaining factors altogether explained 22.8% of the variations (
|
||
<tableCitation id="C6FF20BB6C179461FCA2C8EDFC78FB1F" box="[879,984,1159,1184]" captionStart="TABLE 2" captionStartId="9.[151,239,1084,1107]" captionText="TABLE 2. Samples used for the molecular study." pageId="42" pageNumber="43">Table S2</tableCitation>
|
||
). Principal Component Analysis (PCA) for the adult female specimens (
|
||
<taxonomicName id="4C7D6E836C179461FDD4C8C7FD2BFB7A" authority="Garg & Senevirathne & Wijayathilaka & Phuge & Deuti & Manamendra-Arachchi & Meegaskumbura & Biju, 2018" authorityName="Garg & Senevirathne & Wijayathilaka & Phuge & Deuti & Manamendra-Arachchi & Meegaskumbura & Biju" authorityYear="2018" box="[537,651,1196,1221]" class="Amphibia" family="Microhylidae" genus="Uperodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="42" pageNumber="43" phylum="Chordata" rank="species" species="rohani" status="sp. nov.">
|
||
<emphasis id="B909C9126C179461FDD4C8C7FD2BFB7A" box="[537,651,1196,1221]" italics="true" pageId="42" pageNumber="43">U. rohani</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B909C9126C179461FD5AC8C6FD56FB7A" bold="true" box="[663,758,1196,1221]" pageId="42" pageNumber="43">
|
||
<taxonomicNameLabel id="A23A74696C179461FD5AC8C6FD56FB7A" box="[663,758,1196,1221]" pageId="42" pageNumber="43" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
:
|
||
<emphasis id="B909C9126C179461FCC7C8C7FCBDFB7B" box="[778,797,1197,1220]" italics="true" pageId="42" pageNumber="43">N</emphasis>
|
||
= 7;
|
||
<taxonomicName id="4C7D6E836C179461FCAAC8C7FBA5FB7B" authorityName="Stoliczka" authorityYear="1872" box="[871,1029,1197,1220]" class="Amphibia" family="Microhylidae" genus="Uperodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="42" pageNumber="43" phylum="Chordata" rank="species" species="variegatus">
|
||
<emphasis id="B909C9126C179461FCAAC8C7FBA5FB7B" box="[871,1029,1197,1220]" italics="true" pageId="42" pageNumber="43">U. variegatus</emphasis>
|
||
</taxonomicName>
|
||
:
|
||
<emphasis id="B909C9126C179461FBD4C8C7FB8CFB7B" box="[1049,1068,1197,1220]" italics="true" pageId="42" pageNumber="43">N</emphasis>
|
||
= 8) recovered three Principal Components (PC) with eigenvalues>1.0 that accounted for 82.3% of the total variance; PC 1 explained 46.2% of the variations, with highest loadings for SVL, HW, SL, IUE, UEW, IFE, IBE, MN, EN and TL; PC 2 explained 26.9% of the variations, with highest loadings for HL, FAL, HAL and SHL; PC3 explained 9.2% of the variations, with highest loadings for EN; and the remaining factors altogether explained 17.7% of the variations (
|
||
<tableCitation id="C6FF20BB6C179461FAEBC956FA2FFAEB" box="[1318,1423,1340,1365]" captionStart="TABLE 2" captionStartId="9.[151,239,1084,1107]" captionText="TABLE 2. Samples used for the molecular study." pageId="42" pageNumber="43">Table S2</tableCitation>
|
||
). Projections of factor planes 1 and 2 showed distinct clusters for the two species (
|
||
<figureCitation id="134609856C179461FBD2C935FBD2FAC7" box="[1055,1138,1375,1400]" captionStart="FIGURE 13" captionStartId="40.[151,250,716,738]" captionTargetBox="[166,1415,201,683]" captionTargetId="figure@40.[151,1435,193,694]" captionTargetPageId="40" captionText="FIGURE 13. Principal Component Analyses showing morphometric differentiation between Uperodon rohani sp. nov. and U. variegatus based on 17 morphological characters. Left: Plot for the first two factor planes explaining 54.4% of the variations among 28 adult male specimens. Right: Plot for the first two factor planes explaining 73% of the variations among 15 adult female specimens.Factor loadings for each component are provided in Table S2." httpUri="https://zenodo.org/record/1183481/files/figure.png" pageId="42" pageNumber="43">Fig. 13</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BC215006C179461FF0AC9EEFB21F863" blockId="42.[151,1437,151,2013]" pageId="42" pageNumber="43">
|
||
<emphasis id="B909C9126C179461FF0AC9EEFE55FA22" bold="true" box="[199,501,1412,1437]" pageId="42" pageNumber="43">Osteological comparison.</emphasis>
|
||
<taxonomicName id="4C7D6E836C179461FE33C9EEFD61FA22" authority="Garg & Senevirathne & Wijayathilaka & Phuge & Deuti & Manamendra-Arachchi & Meegaskumbura & Biju, 2018" authorityName="Garg & Senevirathne & Wijayathilaka & Phuge & Deuti & Manamendra-Arachchi & Meegaskumbura & Biju" authorityYear="2018" box="[510,705,1412,1437]" class="Amphibia" family="Microhylidae" genus="Uperodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="42" pageNumber="43" phylum="Chordata" rank="species" species="rohani" status="sp. nov.">
|
||
<emphasis id="B909C9126C179461FE33C9EEFD61FA22" box="[510,705,1412,1437]" italics="true" pageId="42" pageNumber="43">Uperodon rohani</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B909C9126C179461FD07C9EEFC86FA22" bold="true" box="[714,806,1412,1437]" pageId="42" pageNumber="43">
|
||
<taxonomicNameLabel id="A23A74696C179461FD07C9EEFC86FA22" box="[714,806,1412,1437]" pageId="42" pageNumber="43" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
and its closely related species,
|
||
<taxonomicName id="4C7D6E836C179461FB5CC9EFFA8BFA23" authorityName="Stoliczka" authorityYear="1872" box="[1169,1323,1413,1436]" class="Amphibia" family="Microhylidae" genus="Uperodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="42" pageNumber="43" phylum="Chordata" rank="species" species="variegatus">
|
||
<emphasis id="B909C9126C179461FB5CC9EFFA8BFA23" box="[1169,1323,1413,1436]" italics="true" pageId="42" pageNumber="43">U. variegatus</emphasis>
|
||
</taxonomicName>
|
||
, have the following skeletal features: Anteriorly slender, basally wider crania, with widest at anterior margins of tympanic annulus in both species. Cranium length is 93% of its width in
|
||
<taxonomicName id="4C7D6E836C179461FCB5C9A7FC47FA5A" authority="Garg & Senevirathne & Wijayathilaka & Phuge & Deuti & Manamendra-Arachchi & Meegaskumbura & Biju, 2018" authorityName="Garg & Senevirathne & Wijayathilaka & Phuge & Deuti & Manamendra-Arachchi & Meegaskumbura & Biju" authorityYear="2018" box="[888,999,1484,1509]" class="Amphibia" family="Microhylidae" genus="Uperodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="42" pageNumber="43" phylum="Chordata" rank="species" species="rohani" status="sp. nov.">
|
||
<emphasis id="B909C9126C179461FCB5C9A7FC47FA5A" box="[888,999,1484,1509]" italics="true" pageId="42" pageNumber="43">U. rohani</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B909C9126C179461FC3DC9A6FBECFA5A" bold="true" box="[1008,1100,1484,1509]" pageId="42" pageNumber="43">
|
||
<taxonomicNameLabel id="A23A74696C179461FC3DC9A6FBECFA5A" box="[1008,1100,1484,1509]" pageId="42" pageNumber="43" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
and 84.2% in
|
||
<taxonomicName id="4C7D6E836C179461FB37C9A7FA35FA5B" authorityName="Stoliczka" authorityYear="1872" box="[1274,1429,1485,1508]" class="Amphibia" family="Microhylidae" genus="Uperodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="42" pageNumber="43" phylum="Chordata" rank="species" species="variegatus">
|
||
<emphasis id="B909C9126C179461FB37C9A7FA35FA5B" box="[1274,1429,1485,1508]" italics="true" pageId="42" pageNumber="43">U. variegatus</emphasis>
|
||
</taxonomicName>
|
||
. Frontoparietals are largest, covering more than half of crania, slender and quadrangular shaped in both species. Nasals, lying dorsal to nasal capsules, triangular-shaped, medial edge 20.6% of the mid-cranial width in
|
||
<taxonomicName id="4C7D6E836C179461FA4CCA7FFEADF9EF" authorityName="Stoliczka" authorityYear="1872" class="Amphibia" family="Microhylidae" genus="Uperodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="42" pageNumber="43" phylum="Chordata" rank="species" species="variegatus">
|
||
<emphasis id="B909C9126C179461FA4CCA7FFEADF9EF" italics="true" pageId="42" pageNumber="43">U. variegatus</emphasis>
|
||
</taxonomicName>
|
||
and 20.9% in
|
||
<taxonomicName id="4C7D6E836C179461FE7ECA53FD80F9EF" authority="Garg & Senevirathne & Wijayathilaka & Phuge & Deuti & Manamendra-Arachchi & Meegaskumbura & Biju, 2018" authorityName="Garg & Senevirathne & Wijayathilaka & Phuge & Deuti & Manamendra-Arachchi & Meegaskumbura & Biju" authorityYear="2018" box="[435,544,1591,1616]" class="Amphibia" family="Microhylidae" genus="Uperodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="42" pageNumber="43" phylum="Chordata" rank="species" species="rohani" status="sp. nov.">
|
||
<emphasis id="B909C9126C179461FE7ECA53FD80F9EF" box="[435,544,1591,1616]" italics="true" pageId="42" pageNumber="43">U. rohani</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B909C9126C179461FDE5CA5DFD22F9EF" bold="true" box="[552,642,1591,1616]" pageId="42" pageNumber="43">
|
||
<taxonomicNameLabel id="A23A74696C179461FDE5CA5DFD22F9EF" box="[552,642,1591,1616]" pageId="42" pageNumber="43" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
Posterolateral margins of frontoparietals slightly overlay anterolateral margins of prootics. Major differences between the two species concentrate mainly on three bones of the lower and upper jaws—premaxillae, mentomeckelians and dentaries. Premaxillae usually composed of alary, dentary and maxillary processes. In both species, premaxillae are edentate. However, shape of dentary process differs immensely, wherein
|
||
<taxonomicName id="4C7D6E836C179461FE4CCAA3FE4EF95F" authority="Garg & Senevirathne & Wijayathilaka & Phuge & Deuti & Manamendra-Arachchi & Meegaskumbura & Biju, 2018" authorityName="Garg & Senevirathne & Wijayathilaka & Phuge & Deuti & Manamendra-Arachchi & Meegaskumbura & Biju" authorityYear="2018" box="[385,494,1735,1760]" class="Amphibia" family="Microhylidae" genus="Uperodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="42" pageNumber="43" phylum="Chordata" rank="species" species="rohani" status="sp. nov.">
|
||
<emphasis id="B909C9126C179461FE4CCAA3FE4EF95F" box="[385,494,1735,1760]" italics="true" pageId="42" pageNumber="43">U. rohani</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B909C9126C179461FE3BCAADFDF0F95F" bold="true" box="[502,592,1735,1760]" pageId="42" pageNumber="43">
|
||
<taxonomicNameLabel id="A23A74696C179461FE3BCAADFDF0F95F" box="[502,592,1735,1760]" pageId="42" pageNumber="43" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
has well-bifurcated dentary process with 74.9% height compared to width of premaxillae. Comparatively, dentary process of
|
||
<taxonomicName id="4C7D6E836C179461FD25CA87FC25F8BB" authorityName="Stoliczka" authorityYear="1872" box="[744,901,1773,1796]" class="Amphibia" family="Microhylidae" genus="Uperodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="42" pageNumber="43" phylum="Chordata" rank="species" species="variegatus">
|
||
<emphasis id="B909C9126C179461FD25CA87FC25F8BB" box="[744,901,1773,1796]" italics="true" pageId="42" pageNumber="43">U. variegatus</emphasis>
|
||
</taxonomicName>
|
||
is rectangular in shape, with a slight median depression, and 27.4% height compared to premaxillary width. Mentomeckelians are observed as cylindrical endochondral elements, anterior to the anterior ends of Meckel’s cartilages, only in
|
||
<taxonomicName id="4C7D6E836C179461FB23CB5FFA35F8F3" authorityName="Stoliczka" authorityYear="1872" box="[1262,1429,1845,1868]" class="Amphibia" family="Microhylidae" genus="Uperodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="42" pageNumber="43" phylum="Chordata" rank="species" species="variegatus">
|
||
<emphasis id="B909C9126C179461FB23CB5FFA35F8F3" box="[1262,1429,1845,1868]" italics="true" pageId="42" pageNumber="43">U. variegatus</emphasis>
|
||
</taxonomicName>
|
||
. Mentomeckelians are absent in
|
||
<taxonomicName id="4C7D6E836C179461FDDBCB33FD29F8CF" authority="Garg & Senevirathne & Wijayathilaka & Phuge & Deuti & Manamendra-Arachchi & Meegaskumbura & Biju, 2018" authorityName="Garg & Senevirathne & Wijayathilaka & Phuge & Deuti & Manamendra-Arachchi & Meegaskumbura & Biju" authorityYear="2018" box="[534,649,1879,1904]" class="Amphibia" family="Microhylidae" genus="Uperodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="42" pageNumber="43" phylum="Chordata" rank="species" species="rohani" status="sp. nov.">
|
||
<emphasis id="B909C9126C179461FDDBCB33FD29F8CF" box="[534,649,1879,1904]" italics="true" pageId="42" pageNumber="43">U. rohani</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B909C9126C179461FD5ACB3DFD57F8CF" bold="true" box="[663,759,1879,1904]" pageId="42" pageNumber="43">
|
||
<taxonomicNameLabel id="A23A74696C179461FD5ACB3DFD57F8CF" box="[663,759,1879,1904]" pageId="42" pageNumber="43" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
; however, the infrarostrals, which are the precursors of Mentomeckelians are still present and can be seen clearly (
|
||
<figureCitation id="134609856C179461FCA4CB16FC63F82B" box="[873,963,1916,1941]" captionStart="FIGURE14" captionStartId="40.[151,250,1533,1555]" captionTargetBox="[163,1428,885,1494]" captionTargetId="figure@40.[151,1436,879,1511]" captionTargetPageId="40" captionText="FIGURE14.Cleared and double stainedspecimens of Uperodon rohani sp. nov. (above) and U.variegatus (below).From left to right: Dorsal view of the crania composed of nasals (NS), frontoparietals (FP), prootics (PR), exoccipitals (EX) and squamosals (SQ); anteroventral view of the crania showing components of the upper jaw, premaxillae (PM) and maxillae (MA); ventral view of the lower jaw comprising of dentaries (DN), angulosplenials (AG) and mentomeckelians (MM). Bones are indicated in red, cartilage in blue. (Images: Gayani Senevirathne)." httpUri="https://zenodo.org/record/1183483/files/figure.png" pageId="42" pageNumber="43">Fig. 14</figureCitation>
|
||
). Development of dentaries also shows differences between the two species. Dentaries are well-developed, extend for about one-third and two-third of the length of the mandible arc in
|
||
<taxonomicName id="4C7D6E836C179461FE2DCBAFFDEDF862" authority="Garg & Senevirathne & Wijayathilaka & Phuge & Deuti & Manamendra-Arachchi & Meegaskumbura & Biju, 2018" authorityName="Garg & Senevirathne & Wijayathilaka & Phuge & Deuti & Manamendra-Arachchi & Meegaskumbura & Biju" authorityYear="2018" box="[480,589,1988,2013]" class="Amphibia" family="Microhylidae" genus="Uperodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="42" pageNumber="43" phylum="Chordata" rank="species" species="rohani" status="sp. nov.">
|
||
<emphasis id="B909C9126C179461FE2DCBAFFDEDF862" box="[480,589,1988,2013]" italics="true" pageId="42" pageNumber="43">U. rohani</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B909C9126C179461FD9ECBAEFD0DF862" bold="true" box="[595,685,1988,2013]" pageId="42" pageNumber="43">
|
||
<taxonomicNameLabel id="A23A74696C179461FD9ECBAEFD0DF862" box="[595,685,1988,2013]" pageId="42" pageNumber="43" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="4C7D6E836C179461FD2ECBAFFCDBF863" authorityName="Stoliczka" authorityYear="1872" box="[739,891,1989,2012]" class="Amphibia" family="Microhylidae" genus="Uperodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="42" pageNumber="43" phylum="Chordata" rank="species" species="variegatus">
|
||
<emphasis id="B909C9126C179461FD2ECBAFFCDBF863" box="[739,891,1989,2012]" italics="true" pageId="42" pageNumber="43">U. variegatus</emphasis>
|
||
</taxonomicName>
|
||
, respectively (
|
||
<figureCitation id="134609856C179461FBEDCBAEFBD4F863" box="[1056,1140,1988,2013]" captionStart="FIGURE14" captionStartId="40.[151,250,1533,1555]" captionTargetBox="[163,1428,885,1494]" captionTargetId="figure@40.[151,1436,879,1511]" captionTargetPageId="40" captionText="FIGURE14.Cleared and double stainedspecimens of Uperodon rohani sp. nov. (above) and U.variegatus (below).From left to right: Dorsal view of the crania composed of nasals (NS), frontoparietals (FP), prootics (PR), exoccipitals (EX) and squamosals (SQ); anteroventral view of the crania showing components of the upper jaw, premaxillae (PM) and maxillae (MA); ventral view of the lower jaw comprising of dentaries (DN), angulosplenials (AG) and mentomeckelians (MM). Bones are indicated in red, cartilage in blue. (Images: Gayani Senevirathne)." httpUri="https://zenodo.org/record/1183483/files/figure.png" pageId="42" pageNumber="43">Fig. 14</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BC215006C169460FF0ACCFDFBE1FEFF" blockId="43.[151,1437,151,2024]" pageId="43" pageNumber="44">
|
||
<emphasis id="B909C9126C169460FF0ACCFDFE12FF0F" bold="true" box="[199,434,151,176]" pageId="43" pageNumber="44">Genetic divergence.</emphasis>
|
||
For 16S mitochondrial gene sequences, the sampled populations of
|
||
<taxonomicName id="4C7D6E836C169460FB7FCCFDFAD4FF0F" authority="Garg & Senevirathne & Wijayathilaka & Phuge & Deuti & Manamendra-Arachchi & Meegaskumbura & Biju, 2018" authorityName="Garg & Senevirathne & Wijayathilaka & Phuge & Deuti & Manamendra-Arachchi & Meegaskumbura & Biju" authorityYear="2018" box="[1202,1396,151,176]" class="Amphibia" family="Microhylidae" genus="Uperodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="43" pageNumber="44" phylum="Chordata" rank="species" species="rohani" status="sp. nov.">
|
||
<emphasis id="B909C9126C169460FB7FCCFDFAD4FF0F" box="[1202,1396,151,176]" italics="true" pageId="43" pageNumber="44">Uperodon rohani</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B909C9126C169460FAB7CCFDFF68FF6A" bold="true" pageId="43" pageNumber="44">
|
||
<taxonomicNameLabel id="A23A74696C169460FAB7CCFDFF68FF6A" pageId="43" pageNumber="44" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
showed an average intraspecific distance of 0.5% (range 0–1.4%,
|
||
<emphasis id="B909C9126C169460FC7FCCD7FC65FF6B" box="[946,965,189,212]" italics="true" pageId="43" pageNumber="44">N</emphasis>
|
||
= 6). The sample from Bundala showed a divergence of up to 1.4% from other populations. Genetically,
|
||
<taxonomicName id="4C7D6E836C169460FC83CC8BFC1AFF47" authority="Garg & Senevirathne & Wijayathilaka & Phuge & Deuti & Manamendra-Arachchi & Meegaskumbura & Biju, 2018" authorityName="Garg & Senevirathne & Wijayathilaka & Phuge & Deuti & Manamendra-Arachchi & Meegaskumbura & Biju" authorityYear="2018" box="[846,954,223,248]" class="Amphibia" family="Microhylidae" genus="Uperodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="43" pageNumber="44" phylum="Chordata" rank="species" species="rohani" status="sp. nov.">
|
||
<emphasis id="B909C9126C169460FC83CC8BFC1AFF47" box="[846,954,223,248]" italics="true" pageId="43" pageNumber="44">U. rohani</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B909C9126C169460FC0DCCB5FBB9FF47" bold="true" box="[960,1049,223,248]" pageId="43" pageNumber="44">
|
||
<taxonomicNameLabel id="A23A74696C169460FC0DCCB5FBB9FF47" box="[960,1049,223,248]" pageId="43" pageNumber="44" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is closely related to
|
||
<taxonomicName id="4C7D6E836C169460FB30CC8BFA35FF47" authorityName="Stoliczka" authorityYear="1872" box="[1277,1429,225,248]" class="Amphibia" family="Microhylidae" genus="Uperodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="43" pageNumber="44" phylum="Chordata" rank="species" species="variegatus">
|
||
<emphasis id="B909C9126C169460FB30CC8BFA35FF47" box="[1277,1429,225,248]" italics="true" pageId="43" pageNumber="44">U. variegatus</emphasis>
|
||
</taxonomicName>
|
||
, from which it was found to differ by an average uncorrected genetic distance of 2.8% (range 2.4–3.4%,
|
||
<emphasis id="B909C9126C169460FAF1CD6FFAEFFEA3" box="[1340,1359,261,284]" italics="true" pageId="43" pageNumber="44">N</emphasis>
|
||
= 24). For interspecific genetic distances with all other members of the genus, see Table 3.
|
||
</paragraph>
|
||
<paragraph id="8BC215006C169460FF0ACD26FCDCFC0F" blockId="43.[151,1437,151,2024]" pageId="43" pageNumber="44">
|
||
<emphasis id="B909C9126C169460FF0ACD26FE40FEDA" bold="true" box="[199,480,332,357]" pageId="43" pageNumber="44">
|
||
Description of
|
||
<typeStatus id="54C6ABA26C169460FEB4CD26FE40FEDA" box="[377,480,332,357]" pageId="43" pageNumber="44" type="holotype">holotype</typeStatus>
|
||
</emphasis>
|
||
(measurements in mm) (3L, 5K, 12B–J). Small-sized (SVL 26.0), relatively slender adult male; head small (HW 6.7, HL 6.2, IFE 3.5, IBE 5.5), less than one-fourth (23.4%) of body length, wider than long (HW/HL ratio 1.1); snout subovoid in dorsal, subovoid to nearly truncate in ventral view, rounded in lateral view, its length (SL 2.6) slightly longer than horizontal diameter of eye (EL 2.5); loreal region rounded with vertical canthus rostralis; interorbital space wider (IUE 2.9) than upper eyelid (UEW 1.6) and internarial distance (IN 1.9); nostril closer to tip of snout (NS 0.7) than to eye (EN 1.4); supratympanic fold distinct, extending from posterior corner of upper eyelid near to insertion of forelimb at axilla; eye diameter (EL 2.5); vomerine odontophores present on the palate, without teeth; presence of two weakly-developed neopalatinal ridges on posterior side of each choana, fused with the vomerine odontophores; tongue emarginate. Forelimbs moderately long and thin; forearm (FAL 5.2) shorter than hand length (HAL 7.2); finger length formula I<II<IV<III, tips of all fingers with oval discs, discs moderately wide compared to finger width (FD I 0.9, FW I 0.4; FD II 1.3, FW II 0.4; FD III 1.2, FW III 0.4; FD IV 1.2, FW IV 0.5); subarticular tubercles prominent, oval, all present; two well-developed palmar tubercles, oval. Hind limbs relatively long and thin, thigh length (TL 9.2) slightly shorter than shank (SHL 9.4) and shorter than foot (FOL 11.1); relative digit lengths I<II<V<III<IV; tips of all toes slightly enlarged with truncate to rounded discs, discs rather wide compared to toe width (TD I 0.7, TW I 0.6; TD II 0.8, TW II 0.6; TD III 0.7, TW III 0.5; TD IV 0.7, TW IV 0.5; TD V 0.5, TW V 0.5); foot webbing absent; subarticular tubercles prominent, oval; two smooth metatarsal tubercles, oval, outer one slightly longer than the inner.
|
||
</paragraph>
|
||
<paragraph id="8BC215006C169460FF0ACFD1FEF4FBA3" blockId="43.[151,1437,151,2024]" pageId="43" pageNumber="44">Skin of dorsum, snout, and between eyes, shagreened; sides of head, posterior part of back, and upper and lower parts of flank, smooth; dorsal surfaces of forelimb, thigh, tibia and tarsus, smooth to shagreened; ventral surfaces smooth.</paragraph>
|
||
<paragraph id="8BC215006C169460FF0AC84DFE72FA74" blockId="43.[151,1437,151,2024]" pageId="43" pageNumber="44">
|
||
<emphasis id="B909C9126C169460FF0AC84DFEFFFBFF" bold="true" box="[199,351,1063,1088]" pageId="43" pageNumber="44">Colouration.</emphasis>
|
||
<emphasis id="B909C9126C169460FEA5C842FDB5FB80" box="[360,533,1064,1087]" italics="true" pageId="43" pageNumber="44">In preservation</emphasis>
|
||
: Dorsum dark brown with irregular olive grey coloured blotches, pair of faint and irregular olive grey longitudinal streaks extending from upper eyelids up to vent; dorsal surfaces of forelimbs and hind limbs lighter in colour compared to dorsum and with faint yellowish-grey spots or blotches; sides of head brown; ventral surfaces (including limbs) greyish-white, throat with blackish-brown calling patch (
|
||
<figureCitation id="134609856C169460FB2EC8F9FA9AFB14" box="[1251,1338,1171,1196]" captionStart="FIGURE 3" captionStartId="15.[151,250,1749,1771]" captionTargetBox="[151,1436,644,1728]" captionTargetId="figure@15.[151,1436,644,1728]" captionTargetPageId="15" captionText="FIGURE 3. Preserved type specimens of Uperodon species in dorsal and ventral view. A. Uperodon systoma, Syntype, ZMB 3551 (female); B. Uperodon globulosus, Syntype, NHM 62.12.29.4 (ex. BMNH 1947.2.10.97) (female); C. Kaloula pulchra taprobanica, Holotype, NHM 1905.3.25.138 (ex. BMNH 1947.2.10.62) (female); D. Ramanella anamalaiensis, Neotype, ZSI/ WGRC/V/A/956 (male); E. Hylaedactylus montanus, Holotype, NHM 72.4.17.212 (ex. BMNH 1947.2.11.19) (female); F. Ramanella mormorata, Neotype, ZSI/WGRC/V/A/958, (female); G. Ramanella nagaoi, Holotype, WHT 2826 (male); H. Callula obscura, Holotype, NHM 58.11.28.42 (ex. BMNH 1947.2.11.20) (female); I. Ramanella palmata, Holotype, NHM 90.11.8.62 (ex.BMNH 1947.2.11.21); J. Callula triangularis, Lectotype, NHM 74.4.29.897 (ex.BMNH 1947.2.11.15) (male); K.Hylaedactylus carnaticus, Lectotype, ZSI 2761(female); L. Uperodonrohani sp. nov., Holotype, DZ1449 (male).(Images: SD Biju and Sonali Garg)." httpUri="https://zenodo.org/record/1183461/files/figure.png" pageId="43" pageNumber="44">Fig. 3L</figureCitation>
|
||
).
|
||
<emphasis id="B909C9126C169460FA81C8F9FF7BFB6F" italics="true" pageId="43" pageNumber="44">Colour in life</emphasis>
|
||
: Dorsum reddish brown with pair of irregular olive yellow coloured longitudinal streaks extending from upper eyelids up to vent, scattered olive yellow coloured spots and blotches especially towards dorsolateral surfaces and posterior part of back, larger olive yellow blotch in center of dorsum, few scattered spots on head, olive yellow patch between nostrils and upper eyelids; lateral sides of head brown with scattered olive yellow spots near margins of mouth; anterior and posterior parts of groin light brown; dorsal surfaces of limbs brown to light brown with scattered olive yellow spots; throat light brown with bluish-black calling patch and irregular light grey speckles; chest, belly and thighs, greyish-white with light brown margins; limbs light brown with irregular light grey speckles (
|
||
<figureCitation id="134609856C169460FEF3C9D9FE66FA74" box="[318,454,1459,1484]" captionStart="FIGURE12" captionStartId="39.[151,250,1561,1583]" captionTargetBox="[151,1436,988,1540]" captionTargetId="figure@39.[150,1436,988,1540]" captionTargetPageId="39" captionText="FIGURE12. A.Distributionof Uperodon variegatus andUperodon rohani sp. nov. Detailsof localities reported inthe present study are provided in Table 1. Records from literature are listed in supplementary Table S1 and indicated here based on approximatecoordinates; B–E. Head, hand, footand schematic illustration of foot webbing inthe holotype of Uperodonrohani sp. nov. (DZ 1449, male): B. lateral view of head; C. ventral view of hand; D. ventral view of foot; E. foot webbing; F–J. Uperodonrohani sp. nov. (DZ 1449,male) inlife:F. dorsolateral view; G. dorsal view; H.ventral view; I. Uperodon rohanisp. nov. (Paratype,DZ 1485, female),dorsal view; J. Uperodon rohanisp. nov. (DZ 1130, female), dorsolateral view.(Images:SD Biju)." httpUri="https://zenodo.org/record/1183479/files/figure.png" pageId="43" pageNumber="44">Figs. 12F–J</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BC215006C169460FF0AC9BDFD47F91B" blockId="43.[151,1437,151,2024]" pageId="43" pageNumber="44">
|
||
<emphasis id="B909C9126C169460FF0AC9BDFEE7FA4F" bold="true" box="[199,327,1495,1520]" pageId="43" pageNumber="44">Variations.</emphasis>
|
||
Morphometric measurements for
|
||
<specimenCount id="9D7BDE896C169460FD01C9BDFC91FA4F" box="[716,817,1495,1520]" pageId="43" pageNumber="44" type="male">16 males</specimenCount>
|
||
and eight females, including the
|
||
<typeStatus id="54C6ABA26C169460FB61C9B2FB7CFA4F" box="[1196,1244,1496,1520]" pageId="43" pageNumber="44">type</typeStatus>
|
||
series, are given in
|
||
<tableCitation id="C6FF20BB6C169460FF7EC991FEA8F9AC" box="[179,264,1531,1556]" captionStart="TABLE 4" captionStartId="48.[119,193,547,566]" captionTargetBox="[135,1884,616,1625]" captionText="TABLE 4. Morphometric measurements of Uperodon species.All measurements are in millimetres (mm). The status of each specimen is given after the museum number:Holotype (HT), Paratype (PT), Neotype (NT), Lectotype (LT), Syntype (ST)." httpUri="http://table.plazi.org/id/DF0245886C0D947BFFBACE49F8ECFD89" pageId="43" pageNumber="44" tableUuid="DF0245886C0D947BFFBACE49F8ECFD89">Table 4</tableCitation>
|
||
. DZ 1374 and DZ 1336: dorsal skin smooth; DZ 1129: lateral surfaces slightly granular; DZ 1374: lateral surfaces of head slightly granular. DZ 1374: dorsum coffee brown in colour with prominent irregular olive grey spots or patches; DZ 1521: dorsum dark brown with olive grey blotches; DZ 1425: dorsum dark brown with small olive grey spots scattered throughout, scattered spots also seen on surfaces of hand, foot, sides of head and groin; DZ 1485: dorsum darker brown with fewer blotches.
|
||
</paragraph>
|
||
<paragraph id="8BC215006C169460FF0ACAC5FCEFF954" blockId="43.[151,1437,151,2024]" pageId="43" pageNumber="44">
|
||
<emphasis id="B909C9126C169460FF0ACAC5FD88F977" bold="true" box="[199,552,1711,1736]" pageId="43" pageNumber="44">Secondary sexual characters.</emphasis>
|
||
Male (DZ 1449): vocal sac externally visible on the lower jaw; female (DZ 1425): ova white, pigmented on poles (diameter 0.6–1.0 mm).
|
||
</paragraph>
|
||
<paragraph id="8BC215006C169460FF0ACA9DFD14F81F" blockId="43.[151,1437,151,2024]" pageId="43" pageNumber="44">
|
||
<emphasis id="B909C9126C169460FF0ACA9DFD34F8AF" bold="true" box="[199,660,1783,1808]" pageId="43" pageNumber="44">Geographical distribution and habitat.</emphasis>
|
||
<taxonomicName id="4C7D6E836C169460FD57CA9DFCFBF8AF" authority="Garg & Senevirathne & Wijayathilaka & Phuge & Deuti & Manamendra-Arachchi & Meegaskumbura & Biju, 2018" authorityName="Garg & Senevirathne & Wijayathilaka & Phuge & Deuti & Manamendra-Arachchi & Meegaskumbura & Biju" authorityYear="2018" box="[666,859,1783,1808]" class="Amphibia" family="Microhylidae" genus="Uperodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="43" pageNumber="44" phylum="Chordata" rank="species" species="rohani" status="sp. nov.">
|
||
<emphasis id="B909C9126C169460FD57CA9DFCFBF8AF" box="[666,859,1783,1808]" italics="true" pageId="43" pageNumber="44">Uperodon rohani</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B909C9126C169460FCACCA9DFC1AF8AF" bold="true" box="[865,954,1783,1808]" pageId="43" pageNumber="44">
|
||
<taxonomicNameLabel id="A23A74696C169460FCACCA9DFC1AF8AF" box="[865,954,1783,1808]" pageId="43" pageNumber="44" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is endemic to
|
||
<collectingCountry id="F36A55906C169460FB96CA9DFB6BF8AF" box="[1115,1227,1783,1808]" name="Sri Lanka" pageId="43" pageNumber="44">
|
||
<collectingRegion id="49B9DBE26C169460FB96CA9DFBDCF8AF" box="[1115,1148,1783,1808]" country="United Kingdom" name="Surrey" pageId="43" pageNumber="44">Sri</collectingRegion>
|
||
Lanka
|
||
</collectingCountry>
|
||
, where it is widely distributed throughout the island (
|
||
<figureCitation id="134609856C169460FDD8CB71FDD0F88C" box="[533,624,1819,1844]" captionStart="FIGURE 2" captionStartId="14.[151,250,1617,1639]" captionTargetBox="[152,1434,196,1594]" captionTargetId="figure@14.[151,1436,193,1595]" captionTargetPageId="14" captionText="FIGURE 2. A. Geographic distribution of the genus Uperodon in Bangladesh, Bhutan, India, Nepal, Pakistan and Sri Lanka; B. Distributionof Uperodon globulosus, U. systomaand U.taprobanicus; C. Distribution of four endemic Uperodon species of the Western Ghats (excluding the widely distributed Indian member U. variegatus, shown in Fig.12A); D. Distribution of four endemic Uperdodon species of Sri Lanka. Details of localities reported in the present study are provided in Table 1. Records from literature are listed in supplementary Table S1 and indicated here based onapproximate coordinates." httpUri="https://zenodo.org/record/1183459/files/figure.png" pageId="43" pageNumber="44">Fig. 2D</figureCitation>
|
||
). In the present study, we genetically and/or morphologically confirmed the occurrence of this species at several localities in the
|
||
<collectingRegion id="49B9DBE26C169460FCDDCB55FCC5F8E7" box="[784,869,1855,1880]" country="Sri Lanka" name="Central" pageId="43" pageNumber="44">Central</collectingRegion>
|
||
,
|
||
<collectingRegion id="49B9DBE26C169460FCBCCB2AFC68F8E8" box="[881,968,1856,1879]" country="Sri Lanka" name="Eastern" pageId="43" pageNumber="44">Eastern</collectingRegion>
|
||
,
|
||
<collectingRegion id="49B9DBE26C169460FC19CB55FB9CF8E7" box="[980,1084,1855,1880]" country="Sri Lanka" name="Northern" pageId="43" pageNumber="44">Northern</collectingRegion>
|
||
,
|
||
<collectingRegion id="49B9DBE26C169460FB85CB55FB45F8E7" box="[1096,1253,1855,1880]" country="Sri Lanka" name="North Central" pageId="43" pageNumber="44">North Central</collectingRegion>
|
||
,
|
||
<collectingRegion id="49B9DBE26C169460FB3CCB55FA38F8E8" box="[1265,1432,1855,1880]" country="Sri Lanka" name="North Western" pageId="43" pageNumber="44">
|
||
North
|
||
<collectingRegion id="49B9DBE26C169460FAF7CB2AFA38F8E8" box="[1338,1432,1856,1879]" country="American Samoa" name="Western" pageId="43" pageNumber="44">Western</collectingRegion>
|
||
</collectingRegion>
|
||
,
|
||
<collectingRegion id="49B9DBE26C169460FF5ACB09FF5FF8C3" box="[151,255,1891,1916]" country="Sri Lanka" name="Southern" pageId="43" pageNumber="44">Southern</collectingRegion>
|
||
, and
|
||
<collectingRegion id="49B9DBE26C169460FEF7CB0EFEC9F8C4" box="[314,361,1892,1915]" country="Sri Lanka" name="Uva" pageId="43" pageNumber="44">Uva</collectingRegion>
|
||
provinces (
|
||
<tableCitation id="C6FF20BB6C169460FE21CB09FDE0F8C4" box="[492,576,1891,1916]" captionStart="TABLE 1" captionStartId="4.[151,239,1593,1616]" captionText="TABLE 1. Collection localities for the genetically and/or morphologically studied Uperodon samples in the present study." pageId="43" pageNumber="44">Table 1</tableCitation>
|
||
). Previous records of
|
||
<taxonomicName id="4C7D6E836C169460FCF7CB0EFC73F8C4" authorityName="Stoliczka" authorityYear="1872" box="[826,979,1892,1915]" class="Amphibia" family="Microhylidae" genus="Uperodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="43" pageNumber="44" phylum="Chordata" rank="species" species="variegatus">
|
||
<emphasis id="B909C9126C169460FCF7CB0EFC73F8C4" box="[826,979,1892,1915]" italics="true" pageId="43" pageNumber="44">U. variegatus</emphasis>
|
||
</taxonomicName>
|
||
from
|
||
<collectingCountry id="F36A55906C169460FBD5CB09FB26F8C3" box="[1048,1158,1891,1916]" name="Sri Lanka" pageId="43" pageNumber="44">
|
||
<collectingRegion id="49B9DBE26C169460FBD5CB09FB99F8C3" box="[1048,1081,1891,1916]" country="United Kingdom" name="Surrey" pageId="43" pageNumber="44">Sri</collectingRegion>
|
||
Lanka
|
||
</collectingCountry>
|
||
refer to this species (see supplementary
|
||
<tableCitation id="C6FF20BB6C169460FE8ECBEDFE05F820" box="[323,421,1927,1952]" captionStart="TABLE 1" captionStartId="4.[151,239,1593,1616]" captionText="TABLE 1. Collection localities for the genetically and/or morphologically studied Uperodon samples in the present study." pageId="43" pageNumber="44">Table S1</tableCitation>
|
||
and references thereto).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C367468B6C169460FF0ACBC1FB48F858" pageId="43" pageNumber="44" type="discussion">
|
||
<paragraph id="8BC215006C169460FF0ACBC1FB48F858" blockId="43.[151,1437,151,2024]" pageId="43" pageNumber="44">
|
||
<emphasis id="B909C9126C169460FF0ACBC1FEE4F87B" bold="true" box="[199,324,1963,1988]" pageId="43" pageNumber="44">Comment.</emphasis>
|
||
This species was historically considered to be conspecific with the Indian species
|
||
<taxonomicName id="4C7D6E836C169460FAE6CBC1FEADF858" authorityName="Stoliczka" authorityYear="1872" class="Amphibia" family="Microhylidae" genus="Uperodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="43" pageNumber="44" phylum="Chordata" rank="species" species="variegatus">
|
||
<emphasis id="B909C9126C169460FAE6CBC1FEADF858" italics="true" pageId="43" pageNumber="44">Uperodon variegatus</emphasis>
|
||
</taxonomicName>
|
||
and widely reported as
|
||
<taxonomicName id="4C7D6E836C169460FDD4CBA5FCA5F858" baseAuthorityName="Stoliczka" baseAuthorityYear="1872" box="[537,773,1999,2024]" class="Amphibia" family="Microhylidae" genus="Ramanella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="43" pageNumber="44" phylum="Chordata" rank="species" species="variegata">
|
||
<emphasis id="B909C9126C169460FDD4CBA5FCA5F858" box="[537,773,1999,2024]" italics="true" pageId="43" pageNumber="44">Ramanella variegata</emphasis>
|
||
</taxonomicName>
|
||
(see
|
||
<tableCitation id="C6FF20BB6C169460FC8DCBA5FC33F858" box="[832,915,1999,2024]" captionStart="TABLE 1" captionStartId="4.[151,239,1593,1616]" captionText="TABLE 1. Collection localities for the genetically and/or morphologically studied Uperodon samples in the present study." pageId="43" pageNumber="44">Table 1</tableCitation>
|
||
and supplementary
|
||
<tableCitation id="C6FF20BB6C169460FBB8CBA5FB79F858" box="[1141,1241,1999,2024]" captionStart="TABLE 1" captionStartId="4.[151,239,1593,1616]" captionText="TABLE 1. Collection localities for the genetically and/or morphologically studied Uperodon samples in the present study." pageId="43" pageNumber="44">Table S1</tableCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |