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<document id="9DEF0A248981C30DE37AAAD5385C4881" ID-DOI="10.5281/zenodo.179321" ID-GBIF-Dataset="0d9c9123-7c7e-4def-b770-0fd81f6f1cba" ID-ISSN="1175-5326" ID-Zenodo-Dep="179321" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459964011400" checkinUser="plazi" docAuthor="Judson, Mark L. I." docDate="2007" docId="03D487C4250FFF8A34A8FB44070E12C1" docLanguage="en" docName="zt01627p068.pdf" docOrigin="Zootaxa 1627" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Lagynochthonius fragilis Judson, 2007, n. sp." docType="treatment" docVersion="8" lastPageNumber="63" masterDocId="FFEDFFBC2509FF80343FFF810316174B" masterDocTitle="A new and endangered species of the pseudoscorpion genus Lagynochthonius from a cave in Vietnam, with notes on chelal morphology and the composition of the Tyrannochthoniini (Arachnida, Chelonethi, Chthoniidae)" masterLastPageNumber="68" masterPageNumber="53" pageNumber="59" updateTime="1698231047183" updateUser="plazi">
<mods:mods id="9954D2D414AB307CE3985547933170C1" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="BCABB65213A7287F96F5CCFD4DDA05E2">
<mods:title id="B68FE70EC05F674858C0219A03BA0357">A new and endangered species of the pseudoscorpion genus Lagynochthonius from a cave in Vietnam, with notes on chelal morphology and the composition of the Tyrannochthoniini (Arachnida, Chelonethi, Chthoniidae)</mods:title>
</mods:titleInfo>
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</mods:role>
<mods:namePart id="6FAB194FC6005B162040F8C1A4859597">Judson, Mark L. I.</mods:namePart>
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<mods:typeOfResource id="090FE8EA99B718F032281861909206A9">text</mods:typeOfResource>
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<mods:title id="D69C78FE645A80CAF824D79A3DD470E1">Zootaxa</mods:title>
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<mods:part id="89AB5A1E3733343B314D29A8FF816268">
<mods:date id="D047C73A0943F0C27D1930FBFAA78FE5">2007</mods:date>
<mods:detail id="3DBF8B430768D2E48F6A21DCE8C4AC53" type="volume">
<mods:number id="886B0A1D2C6B0A5161646B2EAB181858">1627</mods:number>
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<mods:classification id="25441A92D75804B186B9E2E06AEA9C69">journal article</mods:classification>
<mods:identifier id="1965C81C278E1C846AFB9C4D51617512" type="DOI">10.5281/zenodo.179321</mods:identifier>
<mods:identifier id="713279206C43D81B6491D522E5B16EFB" type="GBIF-Dataset">0d9c9123-7c7e-4def-b770-0fd81f6f1cba</mods:identifier>
<mods:identifier id="D8DEDFE7EA27DFA39EA3D28B4B970B47" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="238E803FE0208173E755BDA1E957B84D" type="Zenodo-Dep">179321</mods:identifier>
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<treatment id="03D487C4250FFF8A34A8FB44070E12C1" ID-DOI="http://doi.org/10.5281/zenodo.6246948" ID-GBIF-Taxon="119377212" ID-Zenodo-Dep="6246948" LSID="urn:lsid:plazi:treatment:03D487C4250FFF8A34A8FB44070E12C1" httpUri="http://treatment.plazi.org/id/03D487C4250FFF8A34A8FB44070E12C1" lastPageId="10" lastPageNumber="63" pageId="6" pageNumber="59">
<subSubSection id="C3676559250FFF8634A8FB440208124C" pageId="6" pageNumber="59" type="nomenclature">
<paragraph id="8BC236D2250FFF8634A8FB4401041394" blockId="6.[151,530,1221,1287]" box="[151,530,1221,1247]" pageId="6" pageNumber="59">
<heading id="D08A81BE250FFF8634A8FB4401041394" bold="true" box="[151,530,1221,1247]" fontSize="11" level="1" pageId="6" pageNumber="59" reason="1">
<emphasis id="B909EAC0250FFF8634A8FB4401041394" bold="true" box="[151,530,1221,1247]" pageId="6" pageNumber="59">
<taxonomicName id="4C7D4D51250FFF8634A8FB4402DE1394" ID-CoL="3RY28" box="[151,456,1221,1247]" class="Arachnida" family="Chthoniidae" genus="Lagynochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="6" pageNumber="59" phylum="Arthropoda" rank="species" species="fragilis" status="sp. nov.">
<emphasis id="B909EAC0250FFF8634A8FB4402DE1394" bold="true" box="[151,456,1221,1247]" italics="true" pageId="6" pageNumber="59">Lagynochthonius fragilis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23A57BB250FFF8635F0FB4401041394" box="[463,530,1221,1247]" pageId="6" pageNumber="59" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BC236D2250FFF8634A8FB6C0208124C" blockId="6.[151,530,1221,1287]" box="[151,286,1261,1287]" pageId="6" pageNumber="59">
(
<figureCitation id="13462A57250FFF8634A0FB6C03F2124C" box="[159,228,1261,1287]" captionStart="FIGURE 1" captionStartId="7.[151,244,775,796]" captionText="FIGURE 1. Lagynochthonius fragilis n. sp., dorsal view of female holotype." pageId="6" pageNumber="59">Figs 1</figureCitation>
–21)
</paragraph>
</subSubSection>
<subSubSection id="C3676559250FFF8634A8FABC068A1124" pageId="6" pageNumber="59" type="materials_examined">
<paragraph id="8BC236D2250FFF8634A8FABC068A1124" blockId="6.[151,1437,1341,2007]" pageId="6" pageNumber="59">
<emphasis id="B909EAC0250FFF8634A8FABC025B121C" bold="true" box="[151,333,1341,1367]" pageId="6" pageNumber="59">
<typeStatus id="54C68870250FFF8634A8FABC03C2121C" box="[151,212,1341,1367]" pageId="6" pageNumber="59">Type</typeStatus>
material.
</emphasis>
<typeStatus id="54C68870250FFF86356BFABC02D7121C" box="[340,449,1341,1367]" pageId="6" pageNumber="59" type="holotype">Holotype</typeStatus>
Ψ,
<collectingCountry id="F36A7642250FFF8635D6FABC0165121C" box="[489,627,1341,1367]" name="Vietnam" pageId="6" pageNumber="59">VIETNAM</collectingCountry>
, Kien Giang Prov., Hon Chong karst, Nui Bai Voi, “grotte-hôpital de Mo So” (
<geoCoordinate id="EE495015250FFF86353AFAE402871234" box="[261,401,1381,1407]" direction="north" orientation="latitude" pageId="6" pageNumber="59" precision="15" value="10.231388">10°13'53''N</geoCoordinate>
,
<geoCoordinate id="EE495015250FFF8635A1FAE401231234" box="[414,565,1381,1407]" direction="east" orientation="longitude" pageId="6" pageNumber="59" precision="15" value="104.61639">104°36'59''E</geoCoordinate>
),
<date id="FFC31012250FFF863674FAE400071234" box="[587,785,1381,1407]" pageId="6" pageNumber="59" value="2003-01-29">29 January 2003</date>
, hand-collected by A. Bedos &amp; L. Deharveng (
<collectionCode id="ED6CAE17250FFF86317FFAE4068A1234" LSID="urn:lsid:biocol.org:col:34988" box="[1344,1436,1381,1407]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="6" pageNumber="59">MNHN</collectionCode>
Ps-704-41, in alcohol). The
<typeStatus id="54C68870250FFF8635DCFA0C015F12EC" box="[483,585,1421,1447]" pageId="6" pageNumber="59" type="holotype">holotype</typeStatus>
is gravid: the distended opisthosoma contains about 15 small eggs. It is in fairly good condition, except that many hairs are lost and there is a transverse dorsal tear in the opisthosoma.
<typeStatus id="54C68870250FFF8634DAFA5C025D12BC" box="[229,331,1501,1527]" pageId="6" pageNumber="59" type="paratype">Paratype</typeStatus>
Ψ, right palp only, same details as
<typeStatus id="54C68870250FFF8636CDFA5C004D12BC" box="[754,859,1501,1527]" pageId="6" pageNumber="59" type="holotype">holotype</typeStatus>
, except collected by Le Cong
<collectingCountry id="F36A7642250FFF8630F6FA5C07E912BC" box="[1225,1279,1501,1527]" name="Isle of Man" pageId="6" pageNumber="59">Man</collectingCountry>
&amp; L. Deharveng (
<collectionCode id="ED6CAE17250FFF8634DFF984022A1154" LSID="urn:lsid:biocol.org:col:34988" box="[224,316,1541,1567]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="6" pageNumber="59">MNHN</collectionCode>
Ps-704-42, in alcohol). The rest of this specimen was not present in the sample from which it was sorted. Both
<typeStatus id="54C68870250FFF86354AF9AC02A5110C" box="[373,435,1581,1607]" pageId="6" pageNumber="59">types</typeStatus>
are deposited in the Muséum national d’Histoire naturelle, Paris (
<collectionCode id="ED6CAE17250FFF8630CBF9AC0643110C" LSID="urn:lsid:biocol.org:col:34988" box="[1268,1365,1581,1607]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="6" pageNumber="59">MNHN</collectionCode>
). The “grotte-hôpital de Mo So” is a horizontal calcareous cave that extends for over
<quantity id="4C859B37250FFF863010F9D4076E1124" box="[1071,1144,1621,1647]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="6" pageNumber="59" unit="m" value="500.0">500 m</quantity>
(
<bibRefCitation id="EFEC4B23250FFF8630B8F9D406991124" author="Deharveng" box="[1159,1423,1621,1647]" pageId="6" pageNumber="66" refString="Deharveng, L., Truong Quang Tam &amp; Duong Tien Dung (1995) Explorations au centre et au sud du Vietnam. Spelunca, 59, 8 - 10." type="journal article" year="1995">
Deharveng
<emphasis id="B909EAC0250FFF86312FF9D4065C1125" box="[1296,1354,1621,1646]" italics="true" pageId="6" pageNumber="59">et al.</emphasis>
1995
</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3676559250FFF8634F9F9FC000F11F4" pageId="6" pageNumber="59" type="diagnosis">
<paragraph id="8BC236D2250FFF8634F9F9FC000F11F4" blockId="6.[151,1437,1341,2007]" pageId="6" pageNumber="59">
<emphasis id="B909EAC0250FFF8634F9F9FC025011DC" bold="true" box="[198,326,1661,1687]" pageId="6" pageNumber="59">Diagnosis.</emphasis>
Troglomorphic species with reduced eyes and elongate appendages; tergites I–VII with 4 setae; chela of female 6.2–6.3 times longer than broad.
</paragraph>
</subSubSection>
<subSubSection id="C3676559250FFF8A34F9F94C071516E9" lastPageId="10" lastPageNumber="63" pageId="6" pageNumber="59" type="description">
<paragraph id="8BC236D2250FFF8634F9F94C03F3109C" blockId="6.[151,1437,1341,2007]" pageId="6" pageNumber="59">
<emphasis id="B909EAC0250FFF8634F9F94C024911AC" bold="true" box="[198,351,1741,1767]" pageId="6" pageNumber="59">Description.</emphasis>
Moderately large species with troglomorphic facies: pallid, eyes reduced and appendages elongate (
<figureCitation id="13462A57250FFF863532F97402431044" box="[269,341,1781,1807]" captionStart="FIGURE 1" captionStartId="7.[151,244,775,796]" captionText="FIGURE 1. Lagynochthonius fragilis n. sp., dorsal view of female holotype." pageId="6" pageNumber="59">Fig. 1</figureCitation>
). Carapace distinctly constricted posteriorly (
<figureCitation id="13462A57250FFF863741F97400DF1044" box="[894,969,1781,1807]" captionStart="FIGURES 2 – 12" captionStartId="8.[151,257,1822,1843]" captionTargetBox="[171,1429,534,1792]" captionTargetId="figure@8.[151,1436,514,1798]" captionTargetPageId="8" captionText="FIGURES 2 – 12. Lagynochthonius fragilis n. sp., female holotype. 2, carapace; 3, process of coxa I, lateral view; 4, process of coxa I, ventral view; 5, anterior margin of carapace; 6, right leg IV, lateral view; 7, tip of movable finger of chelicera, showing spinneret, distoventral view. 8, right chelicera. 9, apotele of right leg IV, lateral view; 10, arolium and basal piece of apotele of right leg IV, ventral view; 11, right leg I, lateral view; 12, flagellum of right chelicera. Abbreviations: Δ condyle of palpcoxa; λ base of cheliceral tectum; Μ muscles; po preocular seta; tap tapetum of anterior eye; tec tectum of chelicera. Divisions of scales lines 0.1 mm (figs 2 - 5, 8, 11), 0.05 mm (fig. 6) or 0.01 mm (figs 7, 9 - 10, 12)." httpUri="https://zenodo.org/record/179322/files/figure.png" pageId="6" pageNumber="59">Fig. 2</figureCitation>
); anterior eyes with very weak lenses, posterior eyes not visible externally; both pairs of eyes with reduced tapeta, though these still reflect incident light; anteromedian process absent, space between median setae slightly rounded, weakly depressed and wellsclerotized; hispid anterolaterally and reticulate posterolaterally; anterior margin (apart from median region) extremely thin and difficult to see without dissection of chelicerae, denticulate (
<figureCitation id="13462A57250FFF863001F814079010E4" box="[1086,1158,1941,1967]" captionStart="FIGURES 2 – 12" captionStartId="8.[151,257,1822,1843]" captionTargetBox="[171,1429,534,1792]" captionTargetId="figure@8.[151,1436,514,1798]" captionTargetPageId="8" captionText="FIGURES 2 – 12. Lagynochthonius fragilis n. sp., female holotype. 2, carapace; 3, process of coxa I, lateral view; 4, process of coxa I, ventral view; 5, anterior margin of carapace; 6, right leg IV, lateral view; 7, tip of movable finger of chelicera, showing spinneret, distoventral view. 8, right chelicera. 9, apotele of right leg IV, lateral view; 10, arolium and basal piece of apotele of right leg IV, ventral view; 11, right leg I, lateral view; 12, flagellum of right chelicera. Abbreviations: Δ condyle of palpcoxa; λ base of cheliceral tectum; Μ muscles; po preocular seta; tap tapetum of anterior eye; tec tectum of chelicera. Divisions of scales lines 0.1 mm (figs 2 - 5, 8, 11), 0.05 mm (fig. 6) or 0.01 mm (figs 7, 9 - 10, 12)." httpUri="https://zenodo.org/record/179322/files/figure.png" pageId="6" pageNumber="59">Fig. 5</figureCitation>
); chaetotaxy s4s-4-4-2- 2 (18).
</paragraph>
<caption id="DF02665A250EFF8734A8FC86006F1457" box="[151,889,775,797]" pageId="7" pageNumber="60">
<paragraph id="8BC236D2250EFF8734A8FC86006F1457" blockId="7.[151,889,775,797]" box="[151,889,775,797]" pageId="7" pageNumber="60">
<emphasis id="B909EAC0250EFF8734A8FC86021B1457" bold="true" box="[151,269,775,796]" pageId="7" pageNumber="60">FIGURE 1.</emphasis>
<taxonomicName id="4C7D4D51250EFF87352DFC8901141456" box="[274,514,776,797]" class="Arachnida" family="Chthoniidae" genus="Lagynochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="7" pageNumber="60" phylum="Arthropoda" rank="species" species="fragilis" status="sp. nov.">
<emphasis id="B909EAC0250EFF87352DFC8901141456" box="[274,514,776,797]" italics="true" pageId="7" pageNumber="60">Lagynochthonius fragilis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23A57BB250EFF873637FC8601281457" box="[520,574,775,796]" pageId="7" pageNumber="60" rank="species">n. sp.</taxonomicNameLabel>
, dorsal view of female holotype.
</paragraph>
</caption>
<paragraph id="8BC236D2250EFF8734F9FC0C024C130C" blockId="7.[151,1437,909,1984]" pageId="7" pageNumber="60">
Tergal chaetotaxy 4:4:4:4:4:4:4:5:5:4:TT:0; tergites VIII and IX each with an unpaired median seta. Sternal setae 10:(
<quantity id="4C859B37250EFF873507FC3402721484" box="[312,356,949,975]" metricMagnitude="0" metricUnit="m" metricValue="3.0" pageId="7" pageNumber="60" unit="m" value="3.0">3m</quantity>
)s8s(
<quantity id="4C859B37250EFF8735A3FC3402D11484" box="[412,455,949,975]" metricMagnitude="0" metricUnit="m" metricValue="3.0" pageId="7" pageNumber="60" unit="m" value="3.0">3m</quantity>
):(
<quantity id="4C859B37250EFF8735E1FC34011C1484" box="[478,522,949,975]" metricMagnitude="0" metricUnit="m" metricValue="3.0" pageId="7" pageNumber="60" unit="m" value="3.0">3m</quantity>
)s6s(
<quantity id="4C859B37250EFF87367BFC3401661484" box="[580,624,949,975]" metricMagnitude="0" metricUnit="m" metricValue="3.0" pageId="7" pageNumber="60" unit="m" value="3.0">3m</quantity>
):10?:9:9:9:9:9:0:2; sternites VI–X with unpaired median seta. Coxae typical, anterior process of coxa I long (
<figureCitation id="13462A57250EFF873656FC5C01C514BC" box="[617,723,989,1015]" captionStart="FIGURES 2 – 12" captionStartId="8.[151,257,1822,1843]" captionTargetBox="[171,1429,534,1792]" captionTargetId="figure@8.[151,1436,514,1798]" captionTargetPageId="8" captionText="FIGURES 2 – 12. Lagynochthonius fragilis n. sp., female holotype. 2, carapace; 3, process of coxa I, lateral view; 4, process of coxa I, ventral view; 5, anterior margin of carapace; 6, right leg IV, lateral view; 7, tip of movable finger of chelicera, showing spinneret, distoventral view. 8, right chelicera. 9, apotele of right leg IV, lateral view; 10, arolium and basal piece of apotele of right leg IV, ventral view; 11, right leg I, lateral view; 12, flagellum of right chelicera. Abbreviations: Δ condyle of palpcoxa; λ base of cheliceral tectum; Μ muscles; po preocular seta; tap tapetum of anterior eye; tec tectum of chelicera. Divisions of scales lines 0.1 mm (figs 2 - 5, 8, 11), 0.05 mm (fig. 6) or 0.01 mm (figs 7, 9 - 10, 12)." httpUri="https://zenodo.org/record/179322/files/figure.png" pageId="7" pageNumber="60">Figs 3–4</figureCitation>
); setae P 5,
<date id="FFC31012250EFF873760FC5C009614BC" box="[863,896,989,1015]" pageId="7" pageNumber="60">I 3</date>
,
<date id="FFC31012250EFF8737B2FC5C00C014BC" box="[909,982,989,1015]" pageId="7" pageNumber="60">II 3–4</date>
,
<date id="FFC31012250EFF8737DDFC5C070114BC" box="[994,1047,989,1015]" pageId="7" pageNumber="60">III 4</date>
,
<date id="FFC31012250EFF87301BFC5C074D14BC" box="[1060,1115,989,1015]" pageId="7" pageNumber="60">IV 5</date>
; intercoxal tubercle absent. Coxae II each with 10–11 spines, arranged in an arc, distal (middle) spines longest, spines incised for about half their length.
</paragraph>
<paragraph id="8BC236D2250EFF8734F9FBD4025D1214" blockId="7.[151,1437,909,1984]" pageId="7" pageNumber="60">
Cheliceral palm (
<figureCitation id="13462A57250EFF8735A8FBD402F61324" box="[407,480,1109,1135]" captionStart="FIGURES 2 – 12" captionStartId="8.[151,257,1822,1843]" captionTargetBox="[171,1429,534,1792]" captionTargetId="figure@8.[151,1436,514,1798]" captionTargetPageId="8" captionText="FIGURES 2 – 12. Lagynochthonius fragilis n. sp., female holotype. 2, carapace; 3, process of coxa I, lateral view; 4, process of coxa I, ventral view; 5, anterior margin of carapace; 6, right leg IV, lateral view; 7, tip of movable finger of chelicera, showing spinneret, distoventral view. 8, right chelicera. 9, apotele of right leg IV, lateral view; 10, arolium and basal piece of apotele of right leg IV, ventral view; 11, right leg I, lateral view; 12, flagellum of right chelicera. Abbreviations: Δ condyle of palpcoxa; λ base of cheliceral tectum; Μ muscles; po preocular seta; tap tapetum of anterior eye; tec tectum of chelicera. Divisions of scales lines 0.1 mm (figs 2 - 5, 8, 11), 0.05 mm (fig. 6) or 0.01 mm (figs 7, 9 - 10, 12)." httpUri="https://zenodo.org/record/179322/files/figure.png" pageId="7" pageNumber="60">Fig. 8</figureCitation>
) with 5 setae, ventrobasal seta shorter than others. Palm with moderate hispid granulation dorsally. Movable finger with a few hispid granules in middle of ventral face, but otherwise smooth. Fixed cheliceral finger with 10–11 teeth, distal tooth larger than others; movable finger with 9–10 teeth; spinneret weakly raised, keel-like, with 2 silk-duct openings (
<figureCitation id="13462A57250EFF873784FB4C071513AC" box="[955,1027,1229,1255]" captionStart="FIGURES 2 – 12" captionStartId="8.[151,257,1822,1843]" captionTargetBox="[171,1429,534,1792]" captionTargetId="figure@8.[151,1436,514,1798]" captionTargetPageId="8" captionText="FIGURES 2 – 12. Lagynochthonius fragilis n. sp., female holotype. 2, carapace; 3, process of coxa I, lateral view; 4, process of coxa I, ventral view; 5, anterior margin of carapace; 6, right leg IV, lateral view; 7, tip of movable finger of chelicera, showing spinneret, distoventral view. 8, right chelicera. 9, apotele of right leg IV, lateral view; 10, arolium and basal piece of apotele of right leg IV, ventral view; 11, right leg I, lateral view; 12, flagellum of right chelicera. Abbreviations: Δ condyle of palpcoxa; λ base of cheliceral tectum; Μ muscles; po preocular seta; tap tapetum of anterior eye; tec tectum of chelicera. Divisions of scales lines 0.1 mm (figs 2 - 5, 8, 11), 0.05 mm (fig. 6) or 0.01 mm (figs 7, 9 - 10, 12)." httpUri="https://zenodo.org/record/179322/files/figure.png" pageId="7" pageNumber="60">Fig. 7</figureCitation>
). Serrula exterior with 20 and serrula interior with 14 blades. Rallum (‘flagellum’) (
<figureCitation id="13462A57250EFF8736D7FB74002A1244" box="[744,828,1269,1295]" captionStart="FIGURES 2 – 12" captionStartId="8.[151,257,1822,1843]" captionTargetBox="[171,1429,534,1792]" captionTargetId="figure@8.[151,1436,514,1798]" captionTargetPageId="8" captionText="FIGURES 2 – 12. Lagynochthonius fragilis n. sp., female holotype. 2, carapace; 3, process of coxa I, lateral view; 4, process of coxa I, ventral view; 5, anterior margin of carapace; 6, right leg IV, lateral view; 7, tip of movable finger of chelicera, showing spinneret, distoventral view. 8, right chelicera. 9, apotele of right leg IV, lateral view; 10, arolium and basal piece of apotele of right leg IV, ventral view; 11, right leg I, lateral view; 12, flagellum of right chelicera. Abbreviations: Δ condyle of palpcoxa; λ base of cheliceral tectum; Μ muscles; po preocular seta; tap tapetum of anterior eye; tec tectum of chelicera. Divisions of scales lines 0.1 mm (figs 2 - 5, 8, 11), 0.05 mm (fig. 6) or 0.01 mm (figs 7, 9 - 10, 12)." httpUri="https://zenodo.org/record/179322/files/figure.png" pageId="7" pageNumber="60">Fig. 12</figureCitation>
) with 8 blades, distal blade recumbent basally, with fine barbules and set apart from the other blades, latter tightly grouped and with long pinnae, some of which are subdivided.
</paragraph>
<paragraph id="8BC236D2250EFF8734F9FAEC029A108B" blockId="7.[151,1437,909,1984]" pageId="7" pageNumber="60">
Palp (Figs 13–21) smooth, apart from a patch of minute denticles on ventral surface of patella (Fig. 21). Femur 1.7 times longer than carapace; setal formula 4-5-1-3-5-0. Chelal palm (Figs 13, 18) gradually constricted towards fingers; dorsal surface with a single row of 5 chemosensory setae between trichobothria
<emphasis id="B909EAC0250EFF87314AFA3C068A129D" box="[1397,1436,1469,1494]" italics="true" pageId="7" pageNumber="60">esb</emphasis>
and
<emphasis id="B909EAC0250EFF8734F4FA64021A12B5" box="[203,268,1509,1534]" italics="true" pageId="7" pageNumber="60">ib/isb</emphasis>
; distal paraxial seta of palm only moderately enlarged (length
<quantity id="4C859B37250EFF87303CFA64079612B4" box="[1027,1152,1509,1535]" metricMagnitude="-5" metricUnit="m" metricValue="8.7" pageId="7" pageNumber="60" unit="mm" value="0.087">0.087 mm</quantity>
); antiaxial lyrifissure at base of fixed finger unusually small. Fingers weakly curved in dorsal view (Fig. 18). Fixed finger with 16 macrodenticles and 7 (
<quantity id="4C859B37250EFF87359FF9B402DB1104" box="[416,461,1589,1615]" metricMagnitude="-1" metricUnit="m" metricValue="1.524" pageId="7" pageNumber="60" unit="in" value="6.0">6 in</quantity>
<typeStatus id="54C68870250EFF8735EBF9B4012D1104" box="[468,571,1589,1615]" pageId="7" pageNumber="60" type="paratype">paratype</typeStatus>
) intercalary microdenticles, latter without canals; first 5 and last 4 macrodenticles smaller than others. Fixed finger also with a modified accessory tooth on dorso-antiaxial face (Figs 14 and 19:
<emphasis id="B909EAC0250EFF873524F904022711D5" box="[283,305,1669,1694]" italics="true" pageId="7" pageNumber="60">td</emphasis>
), between the sensilla
<emphasis id="B909EAC0250EFF873600F904014311D5" box="[575,597,1669,1694]" italics="true" pageId="7" pageNumber="60">af</emphasis>
and trichobothria
<emphasis id="B909EAC0250EFF873710F904005D11D5" box="[815,843,1669,1694]" italics="true" pageId="7" pageNumber="60">dx</emphasis>
. Movable finger with 7 (
<quantity id="4C859B37250EFF87304CF90407B711D4" box="[1139,1185,1669,1695]" metricMagnitude="-1" metricUnit="m" metricValue="1.524" pageId="7" pageNumber="60" unit="in" value="6.0">6 in</quantity>
<typeStatus id="54C68870250EFF873097F904061911D4" box="[1192,1295,1669,1695]" pageId="7" pageNumber="60" type="paratype">paratype</typeStatus>
) macrodenticles and 5 (
<quantity id="4C859B37250EFF873512F92C024C118C" box="[301,346,1709,1735]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="7" pageNumber="60" unit="in" value="3.0">3 in</quantity>
<typeStatus id="54C68870250EFF87355FF92C02D1118C" box="[352,455,1709,1735]" pageId="7" pageNumber="60" type="paratype">paratype</typeStatus>
) intercalary microdenticles, latter without dental canals; base of finger with about 6 (
<quantity id="4C859B37250EFF87349FF95403CD11A4" box="[160,219,1749,1775]" metricMagnitude="-1" metricUnit="m" metricValue="3.048" pageId="7" pageNumber="60" unit="in" value="12.0">12 in</quantity>
<typeStatus id="54C68870250EFF8734DDF954025F11A4" box="[226,329,1749,1775]" pageId="7" pageNumber="60" type="paratype">paratype</typeStatus>
) very low, vestigial teeth (without canals and therefore difficult to count accurately). Fixed finger with sensilla
<emphasis id="B909EAC0250EFF873562F97C0262105D" box="[349,372,1789,1814]" italics="true" pageId="7" pageNumber="60">af</emphasis>
1–2 close together, near tip (Fig. 14). Movable finger with 4 sensilla:
<emphasis id="B909EAC0250EFF8730B4F97C07B9105D" box="[1163,1199,1789,1814]" italics="true" pageId="7" pageNumber="60">am</emphasis>
1–2 near tip (Fig. 17);
<emphasis id="B909EAC0250EFF8734A8F8AA03B0100F" box="[151,166,1835,1860]" italics="true" pageId="7" pageNumber="60">p</emphasis>
1 just distad of last (7th) macrodenticle (Fig. 16);
<emphasis id="B909EAC0250EFF8736DBF8AA01E5100F" box="[740,755,1835,1860]" italics="true" pageId="7" pageNumber="60">p</emphasis>
2 near dental margin, just behind 7th macrodenticle, with a long internal tubule (Fig. 16). Apodeme complex of movable finger strongly sclerotized. Tip of fixed finger (Fig. 14) with 1 enlarged, slightly lanceolate paraxial seta near tip, movable finger with 2 enlarged setae (not lanceolate) (Fig. 17).
</paragraph>
<paragraph id="8BC236D22501FF8834F9FF1903CA16E9" blockId="8.[151,1436,152,418]" pageId="8" pageNumber="61">
Legs (
<figureCitation id="13462A572501FF88352EFF1902BE17F9" box="[273,424,152,178]" captionStart="FIGURES 2 – 12" captionStartId="8.[151,257,1822,1843]" captionTargetBox="[171,1429,534,1792]" captionTargetId="figure@8.[151,1436,514,1798]" captionTargetPageId="8" captionText="FIGURES 2 – 12. Lagynochthonius fragilis n. sp., female holotype. 2, carapace; 3, process of coxa I, lateral view; 4, process of coxa I, ventral view; 5, anterior margin of carapace; 6, right leg IV, lateral view; 7, tip of movable finger of chelicera, showing spinneret, distoventral view. 8, right chelicera. 9, apotele of right leg IV, lateral view; 10, arolium and basal piece of apotele of right leg IV, ventral view; 11, right leg I, lateral view; 12, flagellum of right chelicera. Abbreviations: Δ condyle of palpcoxa; λ base of cheliceral tectum; Μ muscles; po preocular seta; tap tapetum of anterior eye; tec tectum of chelicera. Divisions of scales lines 0.1 mm (figs 2 - 5, 8, 11), 0.05 mm (fig. 6) or 0.01 mm (figs 7, 9 - 10, 12)." httpUri="https://zenodo.org/record/179322/files/figure.png" pageId="8" pageNumber="61">Figs 6, 9–11</figureCitation>
) typical, elongate. Fine granulation present on anterodorsal faces of femora I, II, IV and patella IV, and on anterior face of trochanters III and IV. Patella of leg IV 1.8 times longer than femur (
<figureCitation id="13462A572501FF8834A0FF6903F11649" box="[159,231,232,258]" captionStart="FIGURES 2 – 12" captionStartId="8.[151,257,1822,1843]" captionTargetBox="[171,1429,534,1792]" captionTargetId="figure@8.[151,1436,514,1798]" captionTargetPageId="8" captionText="FIGURES 2 – 12. Lagynochthonius fragilis n. sp., female holotype. 2, carapace; 3, process of coxa I, lateral view; 4, process of coxa I, ventral view; 5, anterior margin of carapace; 6, right leg IV, lateral view; 7, tip of movable finger of chelicera, showing spinneret, distoventral view. 8, right chelicera. 9, apotele of right leg IV, lateral view; 10, arolium and basal piece of apotele of right leg IV, ventral view; 11, right leg I, lateral view; 12, flagellum of right chelicera. Abbreviations: Δ condyle of palpcoxa; λ base of cheliceral tectum; Μ muscles; po preocular seta; tap tapetum of anterior eye; tec tectum of chelicera. Divisions of scales lines 0.1 mm (figs 2 - 5, 8, 11), 0.05 mm (fig. 6) or 0.01 mm (figs 7, 9 - 10, 12)." httpUri="https://zenodo.org/record/179322/files/figure.png" pageId="8" pageNumber="61">Fig. 6</figureCitation>
). Tarsi with small, simple, gland pores on dorsal surface (1 on tarsi I–III, 2 on tarsus IV). Arolia normal (figs 9–10). Setae of leg I (trochanter to tibia) 4:12:10:13, setae of leg IV (trochanter to basitarsus) 2:3:7:10:9; patellae of legs III and IV each with 3 setae in dorsal row; femora III and IV without dorsal setae; trochanter II with 4 setae, trochanter III with 2 setae; tactile setae present on basitarsus (TS = 0.31) and telotarsus (TS = 0.32).
</paragraph>
<caption id="DF02665A2501FF8834A8F89F014E1080" httpUri="https://zenodo.org/record/179322/files/figure.png" pageId="8" pageNumber="61" targetBox="[171,1429,534,1792]" targetPageId="8">
<paragraph id="8BC236D22501FF8834A8F89F014E1080" blockId="8.[151,1436,1822,1995]" pageId="8" pageNumber="61">
<emphasis id="B909EAC02501FF8834A8F89F022A1078" bold="true" box="[151,316,1822,1843]" pageId="8" pageNumber="61">FIGURES 2–12.</emphasis>
<taxonomicName id="4C7D4D512501FF88357CF89F01251078" box="[323,563,1822,1843]" class="Arachnida" family="Chthoniidae" genus="Lagynochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="8" pageNumber="61" phylum="Arthropoda" rank="species" species="fragilis" status="sp. nov.">
<emphasis id="B909EAC02501FF88357CF89F01251078" box="[323,563,1822,1843]" italics="true" pageId="8" pageNumber="61">Lagynochthonius fragilis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23A57BB2501FF883606F89F01781078" box="[569,622,1822,1843]" pageId="8" pageNumber="61" rank="species">n. sp.</taxonomicNameLabel>
, female holotype. 2, carapace; 3, process of coxa I, lateral view; 4, process of coxa I, ventral view; 5, anterior margin of carapace; 6, right leg IV, lateral view; 7, tip of movable finger of chelicera, showing spinneret, distoventral view. 8, right chelicera. 9, apotele of right leg IV, lateral view; 10, arolium and basal piece of apotele of right leg IV, ventral view; 11, right leg I, lateral view; 12, flagellum of right chelicera. Abbreviations: Δ condyle of palpcoxa; λ base of cheliceral tectum; Μ muscles;
<emphasis id="B909EAC02501FF883537F819023610E6" box="[264,288,1944,1965]" italics="true" pageId="8" pageNumber="61">po</emphasis>
preocular seta;
<emphasis id="B909EAC02501FF883583F81902CD10E6" box="[444,475,1944,1965]" italics="true" pageId="8" pageNumber="61">tap</emphasis>
tapetum of anterior eye;
<emphasis id="B909EAC02501FF8836EEF81901FB10E6" box="[721,749,1944,1965]" italics="true" pageId="8" pageNumber="61">tec</emphasis>
tectum of chelicera. Divisions of scales lines 0.1 mm (figs 2-5, 8, 11), 0.05 mm (fig. 6) or 0.01 mm (figs 7, 9-10, 12).
</paragraph>
</caption>
<paragraph id="8BC236D22500FF8934A8F95100D11016" blockId="9.[151,1436,1744,2035]" pageId="9" pageNumber="62">
<emphasis id="B909EAC02500FF8934A8F951025C11AE" bold="true" box="[151,330,1744,1765]" pageId="9" pageNumber="62">FIGURES 13–21.</emphasis>
<taxonomicName id="4C7D4D512500FF89356EF950015411AD" box="[337,578,1745,1766]" class="Arachnida" family="Chthoniidae" genus="Lagynochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="9" pageNumber="62" phylum="Arthropoda" rank="species" species="fragilis" status="sp. nov.">
<emphasis id="B909EAC02500FF89356EF950015411AD" box="[337,578,1745,1766]" italics="true" pageId="9" pageNumber="62">Lagynochthonius fragilis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23A57BB2500FF893675F950019611AD" box="[586,640,1745,1766]" pageId="9" pageNumber="62" rank="species">n. sp.</taxonomicNameLabel>
, female
<typeStatus id="54C688702500FF8936EBF950003C11AD" box="[724,810,1745,1766]" pageId="9" pageNumber="62" type="holotype">holotype</typeStatus>
, right palp. 13, chela, lateral view [hairs of chemosensory setae, which are lost in
<typeStatus id="54C688702500FF89357FF96F02801048" box="[320,406,1774,1795]" pageId="9" pageNumber="62" type="holotype">holotype</typeStatus>
, added from
<typeStatus id="54C688702500FF893628F96F017C1048" box="[535,618,1774,1795]" pageId="9" pageNumber="62" type="paratype">paratype</typeStatus>
female]; 14, tip of fixed finger, antiaxial view; 15, detail of paraxial face at level of third tooth, showing pores; 16, detail of movable finger, antiaxial view, showing bothridium
<emphasis id="B909EAC02500FF893036F88D0708106A" box="[1033,1054,1804,1825]" italics="true" pageId="9" pageNumber="62">sb</emphasis>
and sensilla
<emphasis id="B909EAC02500FF8930A6F88D07B3106A" box="[1177,1189,1804,1825]" italics="true" pageId="9" pageNumber="62">p</emphasis>
1–2; 17, tip of movable finger, antiaxial view; 18, chela, dorsal view [chemosensory hairs of palm added from
<typeStatus id="54C688702500FF893780F8AA0701100B" box="[959,1047,1835,1856]" pageId="9" pageNumber="62" type="paratype">paratype</typeStatus>
]; 19, tip of fixed finger, dorsal view; 20, palp (minus chela), dorsal view; 21, patella, ventral view, showing weak granulation.
</paragraph>
<caption id="DF02665A2500FF8934A8F8E7011810D3" httpUri="https://zenodo.org/record/179323/files/figure.png" pageId="9" pageNumber="62" targetBox="[156,1429,165,1744]" targetPageId="9">
<paragraph id="8BC236D22500FF8934A8F8E7011810D3" blockId="9.[151,1436,1744,2035]" pageId="9" pageNumber="62">
<emphasis id="B909EAC02500FF8934A8F8E7025F1030" bold="true" box="[151,329,1894,1915]" pageId="9" pageNumber="62">FIGURES 22–23.</emphasis>
<taxonomicName id="4C7D4D512500FF893570F8E702E21030" box="[335,500,1894,1915]" class="Arachnida" family="Chthoniidae" genus="Lagynochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="9" pageNumber="62" phylum="Arthropoda" rank="genus">
<emphasis id="B909EAC02500FF893570F8E702E21030" box="[335,500,1894,1915]" italics="true" pageId="9" pageNumber="62">Lagynochthonius</emphasis>
</taxonomicName>
undescribed sp., male from Tobago, tip of fixed finger of right chela, showing displaced terminal tooth. 22, dorsal view; 23, lateral view.
</paragraph>
</caption>
<paragraph id="8BC236D22500FF8934A8F821015410B8" blockId="9.[151,1436,1744,2035]" pageId="9" pageNumber="62">
Abbreviations:
<emphasis id="B909EAC02500FF893514F820022B10FD" box="[299,317,1953,1974]" italics="true" pageId="9" pageNumber="62">af</emphasis>
apical sensilla of fixed chelal finger;
<emphasis id="B909EAC02500FF893696F82001D010FD" box="[681,710,1953,1974]" italics="true" pageId="9" pageNumber="62">am</emphasis>
apical sensilla of movable chelal finger;
<emphasis id="B909EAC02500FF89306DF820077F10FD" box="[1106,1129,1953,1974]" italics="true" pageId="9" pageNumber="62">dx</emphasis>
trichobothria
<emphasis id="B909EAC02500FF8930CFF820061110FD" box="[1264,1287,1953,1974]" italics="true" pageId="9" pageNumber="62">dx</emphasis>
;
<emphasis id="B909EAC02500FF89312BF820063310FD" box="[1300,1317,1953,1974]" italics="true" pageId="9" pageNumber="62">et</emphasis>
, bothridium of trichobothrium
<emphasis id="B909EAC02500FF893576F83F024C1098" box="[329,346,1982,2003]" italics="true" pageId="9" pageNumber="62">et</emphasis>
;
<emphasis id="B909EAC02500FF893558F83F02651098" box="[359,371,1982,2003]" italics="true" pageId="9" pageNumber="62">p</emphasis>
1–2, proximal sensilla of movable chelal finger;
<emphasis id="B909EAC02500FF893703F83F00471098" box="[828,849,1982,2003]" italics="true" pageId="9" pageNumber="62">sb</emphasis>
, bothridium of trichobothrium
<emphasis id="B909EAC02500FF8930BDF83F07811098" box="[1154,1175,1982,2003]" italics="true" pageId="9" pageNumber="62">sb</emphasis>
;
<emphasis id="B909EAC02500FF89309AF83F07AF1098" box="[1189,1209,1982,2003]" italics="true" pageId="9" pageNumber="62">sc</emphasis>
chemosensory seta;
<emphasis id="B909EAC02500FF8931BCF83F06801098" box="[1411,1430,1982,2003]" italics="true" pageId="9" pageNumber="62">td</emphasis>
, modified tooth;
<emphasis id="B909EAC02500FF89350EF85F022C10B8" box="[305,314,2014,2035]" italics="true" pageId="9" pageNumber="62">v</emphasis>
, vestibulum of bothridium.
</paragraph>
<paragraph id="8BC236D22503FF8A34F9FF19071516E9" blockId="10.[151,1437,152,1418]" pageId="10" pageNumber="63">
Measurements (in mm, ratios in parentheses; for the palp, the larger measurements and lower ratios are those of the
<typeStatus id="54C688702503FF8A3518FF4102871791" box="[295,401,192,218]" pageId="10" pageNumber="63" type="paratype">paratype</typeStatus>
). Body length 1.6. Carapace 0.48 × 0.48 (1.0). Chelicera 0.43 × 0.20 (2.1), movable finger 0.24. Palp femur 0.81–0.90 × 0.13–0.14 (6.3–6.6), patella 0.32–0.35 × 0.15–0.16 (2.1), chela 1.19–1.32 × 0.19–0.21 (6.2–6.3), palm length 0.55–0.61 (2.9), movable finger length 0.71–0.73 (1.2–1.3 × palm). Leg I femur 0.433 × 0.073 (5.9), patella 0.248 × 0.062 (4.0), tibia 0.225 × 0.050 (4.5), tarsus 0.438 × 0.044 (10.0). Leg IV femur 0.255 × 0.235 (1.1), patella 0.470 × 0.218 (2.2), femoropatella length 0.676 (2.9), tibia 0.487 × 0.084 (5.8), basitarsus 0.216 × 0.071 (3.0), telotarsus 0.527 × 0.040 (13.1).
</paragraph>
</subSubSection>
<subSubSection id="C36765592503FF8A34F9FE2E070E12C1" pageId="10" pageNumber="63" type="discussion">
<paragraph id="8BC236D22503FF8A34F9FE2E07A715F1" blockId="10.[151,1437,152,1418]" pageId="10" pageNumber="63">
<emphasis id="B909EAC02503FF8A34F9FE2E02561682" bold="true" box="[198,320,431,457]" pageId="10" pageNumber="63">Remarks.</emphasis>
<taxonomicName id="4C7D4D512503FF8A3579FE31017A1682" box="[326,620,432,457]" class="Arachnida" family="Chthoniidae" genus="Lagynochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="10" pageNumber="63" phylum="Arthropoda" rank="species" species="fragilis">
<emphasis id="B909EAC02503FF8A3579FE31017A1682" box="[326,620,432,457]" italics="true" pageId="10" pageNumber="63">Lagynochthonius fragilis</emphasis>
</taxonomicName>
is the third species of the genus to be recorded from
<collectingCountry id="F36A76422503FF8A30EEFE3106211681" box="[1233,1335,432,458]" name="Vietnam" pageId="10" pageNumber="63">Vietnam</collectingCountry>
, the others being
<taxonomicName id="4C7D4D512503FF8A3535FE59011916B9" authority="Beier" authorityName="Beier" box="[266,527,472,498]" class="Arachnida" family="Chthoniidae" genus="Lagynochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="10" pageNumber="63" phylum="Arthropoda" rank="species" species="annamensis">
<emphasis id="B909EAC02503FF8A3535FE5902A216BA" box="[266,436,472,497]" italics="true" pageId="10" pageNumber="63">L. annamensis</emphasis>
(Beier)
</taxonomicName>
and
<taxonomicName id="4C7D4D512503FF8A3676FE59005E16B9" authority="Beier" authorityName="Beier" box="[585,840,472,498]" class="Arachnida" family="Chthoniidae" genus="Lagynochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="10" pageNumber="63" phylum="Arthropoda" rank="species" species="tonkinensis">
<emphasis id="B909EAC02503FF8A3676FE5901FB16BA" box="[585,749,472,497]" italics="true" pageId="10" pageNumber="63">L. tonkinensis</emphasis>
(
<bibRefCitation id="EFEC4B232503FF8A36C2FE59005E16B9" author="Beier" box="[765,840,472,498]" pageId="10" pageNumber="66" refString="Beier, M. (1951) Die Pseudoscorpione Indochinas. Memoires du Museum national d'Histoire naturelle, Paris, N. S. (A) 1, 47 - 123." type="journal article" year="1951">Beier)</bibRefCitation>
</taxonomicName>
(Beier 1951; Jędryczkowski, 1998). The new species can easily be separated from these epigean species by its much larger size, more attenuated appendages and reduced eyes. Samples taken by A. Bedos and L. Deharveng from outside the cave at Nui Bai Voi included specimens of an epigean
<taxonomicName id="4C7D4D512503FF8A3611FDD101EE1522" box="[558,760,592,617]" class="Arachnida" family="Chthoniidae" genus="Lagynochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="10" pageNumber="63" phylum="Arthropoda" rank="genus">
<emphasis id="B909EAC02503FF8A3611FDD101EE1522" box="[558,760,592,617]" italics="true" pageId="10" pageNumber="63">Lagynochthonius</emphasis>
</taxonomicName>
(1 ɗ, 2 Ψ, 2 tritonymphs, 1 deutonymph: Berlese extraction of secondary forest litter; MNHN) that agree with Beier’s (1951) description of
<taxonomicName id="4C7D4D512503FF8A3053FDF9061915DA" box="[1132,1295,632,657]" class="Arachnida" family="Chthoniidae" genus="Lagynochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="10" pageNumber="63" phylum="Arthropoda" rank="species" species="tonkinensis">
<emphasis id="B909EAC02503FF8A3053FDF9061915DA" box="[1132,1295,632,657]" italics="true" pageId="10" pageNumber="63">L. tonkinensis</emphasis>
</taxonomicName>
, except that the measurements of the females are slightly smaller than those reported for the
<typeStatus id="54C688702503FF8A307DFD2107BA15F1" box="[1090,1196,672,698]" pageId="10" pageNumber="63" type="holotype">holotype</typeStatus>
.
</paragraph>
<paragraph id="8BC236D22503FF8A34F9FD4900931339" blockId="10.[151,1437,152,1418]" pageId="10" pageNumber="63">
The new species is similar to
<taxonomicName id="4C7D4D512503FF8A3621FD4900F715A9" authority="Harvey, 1989" authorityName="Harvey" authorityYear="1989" box="[542,993,712,738]" class="Arachnida" family="Chthoniidae" genus="Lagynochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="10" pageNumber="63" phylum="Arthropoda" rank="species" species="modor">
<emphasis id="B909EAC02503FF8A3621FD49002D15AA" box="[542,827,712,737]" italics="true" pageId="10" pageNumber="63">Lagynochthonius modor</emphasis>
<bibRefCitation id="EFEC4B232503FF8A377DFD4900F715A9" author="Harvey" box="[834,993,712,738]" pageId="10" pageNumber="67" refString="Harvey, M. S. (1989) Two new cavernicolous chthoniids from Australia, with notes on the generic placement of the southwestern Pacific species attributed to the genera Paraliochthonius Beier and Morikawia Chamberlin (Pseudoscorpionida: Chthoniidae). Bulletin of the British Arachnological Society, 8, 21 - 29." type="journal article" year="1989">Harvey, 1989</bibRefCitation>
</taxonomicName>
, described from a cave in Queensland (
<collectingCountry id="F36A76422503FF8A34A0FD7102071441" box="[159,273,752,778]" name="Australia" pageId="10" pageNumber="63">Australia</collectingCountry>
), but differs from the latter in having slightly more robust palps, fewer chelal teeth, and the coxal spines arranged in an arc, rather than a straight line. The only other cavernicolous species known from Southeast Asia is
<taxonomicName id="4C7D4D512503FF8A351DFCC1000C1411" authority="Mahnert, 1988" authorityName="Mahnert" authorityYear="1988" box="[290,794,832,858]" class="Arachnida" family="Chthoniidae" genus="Lagynochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="10" pageNumber="63" phylum="Arthropoda" rank="species" species="guasirih">
<emphasis id="B909EAC02503FF8A351DFCC101431412" box="[290,597,832,857]" italics="true" pageId="10" pageNumber="63">Lagynochthonius guasirih</emphasis>
(Mahnert, 1988)
</taxonomicName>
from
<collectingCountry id="F36A76422503FF8A375EFCC100DF1411" box="[865,969,832,858]" name="Malaysia" pageId="10" pageNumber="63">Sarawak</collectingCountry>
, which differs from
<taxonomicName id="4C7D4D512503FF8A308AFCC1063E1412" box="[1205,1320,832,857]" class="Arachnida" family="Chthoniidae" genus="Lagynochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="10" pageNumber="63" phylum="Arthropoda" rank="species" species="fragilis">
<emphasis id="B909EAC02503FF8A308AFCC1063E1412" box="[1205,1320,832,857]" italics="true" pageId="10" pageNumber="63">L. fragilis</emphasis>
</taxonomicName>
in having the anterior eyes well developed, in retaining a vestige of the anteromedian process of the carapace and in having only 2 setae on tergites I and II. Although small eyes with tapeta are still present in
<taxonomicName id="4C7D4D512503FF8A3084FC11063914E2" box="[1211,1327,912,937]" class="Arachnida" family="Chthoniidae" genus="Lagynochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="10" pageNumber="63" phylum="Arthropoda" rank="species" species="fragilis">
<emphasis id="B909EAC02503FF8A3084FC11063914E2" box="[1211,1327,912,937]" italics="true" pageId="10" pageNumber="63">L. fragilis</emphasis>
</taxonomicName>
, the general form of this species leaves little doubt that it is adapted to a hypogean existence. It is the first troglomorphic species of Tyrannochthoniini to be named from continental Asia; cavernicolous species of
<taxonomicName id="4C7D4D512503FF8A34A8FB8902E91369" authority="Chamberlin" authorityName="Chamberlin" box="[151,511,1032,1058]" class="Arachnida" family="Chthoniidae" genus="Tyrannochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="10" pageNumber="63" phylum="Arthropoda" rank="genus">
<emphasis id="B909EAC02503FF8A34A8FB89027C136A" box="[151,362,1032,1057]" italics="true" pageId="10" pageNumber="63">Tyrannochthonius</emphasis>
Chamberlin
</taxonomicName>
have been recorded from
<collectingCountry id="F36A76422503FF8A3777FB8900A61369" box="[840,944,1032,1058]" name="Thailand" pageId="10" pageNumber="63">Thailand</collectingCountry>
(
<bibRefCitation id="EFEC4B232503FF8A37FCFB8906031369" author="Deharveng" box="[963,1301,1032,1058]" pageId="10" pageNumber="66" refString="Deharveng, L. &amp; Leclerc, P. (1989) Recherches sur les faunes cavernicoles d'Asie du Sud-est. Memoires de Biospeologie, 16, 91 - 110." type="journal article" year="1989">Deharveng &amp; Leclerc 1989</bibRefCitation>
), but these have yet to be described. Given that
<taxonomicName id="4C7D4D512503FF8A367EFBB1001D1302" box="[577,779,1072,1097]" class="Arachnida" family="Chthoniidae" genus="Lagynochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="10" pageNumber="63" phylum="Arthropoda" rank="genus">
<emphasis id="B909EAC02503FF8A367EFBB1001D1302" box="[577,779,1072,1097]" italics="true" pageId="10" pageNumber="63">Lagynochthonius</emphasis>
</taxonomicName>
is abundant in SE Asia, it is likely that other troglomorphic species of this genus remain to be discovered in the region.
</paragraph>
<paragraph id="8BC236D22503FF8A34F9FB01070E12C1" blockId="10.[151,1437,152,1418]" pageId="10" pageNumber="63">
The very thin cuticle of the anterior part of the carapace seen in
<taxonomicName id="4C7D4D512503FF8A378DFB01073013D2" box="[946,1062,1152,1177]" class="Arachnida" family="Chthoniidae" genus="Lagynochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="10" pageNumber="63" phylum="Arthropoda" rank="species" species="fragilis">
<emphasis id="B909EAC02503FF8A378DFB01073013D2" box="[946,1062,1152,1177]" italics="true" pageId="10" pageNumber="63">L. fragilis</emphasis>
</taxonomicName>
appears to be typical for Tyrannochthoniini, although it has not been emphasized previously. When the chelicerae are left undissected, the anterior margin can be very difficult to observe in transmitted light and it is likely that the line at which the cheliceral membrane joins the carapace (labelled λ in
<figureCitation id="13462A572503FF8A371AFB79007D1259" box="[805,875,1272,1298]" captionStart="FIGURES 2 – 12" captionStartId="8.[151,257,1822,1843]" captionTargetBox="[171,1429,534,1792]" captionTargetId="figure@8.[151,1436,514,1798]" captionTargetPageId="8" captionText="FIGURES 2 – 12. Lagynochthonius fragilis n. sp., female holotype. 2, carapace; 3, process of coxa I, lateral view; 4, process of coxa I, ventral view; 5, anterior margin of carapace; 6, right leg IV, lateral view; 7, tip of movable finger of chelicera, showing spinneret, distoventral view. 8, right chelicera. 9, apotele of right leg IV, lateral view; 10, arolium and basal piece of apotele of right leg IV, ventral view; 11, right leg I, lateral view; 12, flagellum of right chelicera. Abbreviations: Δ condyle of palpcoxa; λ base of cheliceral tectum; Μ muscles; po preocular seta; tap tapetum of anterior eye; tec tectum of chelicera. Divisions of scales lines 0.1 mm (figs 2 - 5, 8, 11), 0.05 mm (fig. 6) or 0.01 mm (figs 7, 9 - 10, 12)." httpUri="https://zenodo.org/record/179322/files/figure.png" pageId="10" pageNumber="63">Fig. 5</figureCitation>
) has sometimes been mistaken for the anterior margin of the carapace (especially when the margin has been drawn as being concave on either side of the epistome). The presence of denticulations is therefore probably more widespread in Tyrannochthoniini than previous descriptions would suggest (it is certainly not unique to
<taxonomicName id="4C7D4D512503FF8A37ABFAF1071E12C2" box="[916,1032,1392,1417]" class="Arachnida" family="Chthoniidae" genus="Lagynochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="10" pageNumber="63" phylum="Arthropoda" rank="species" species="fragilis">
<emphasis id="B909EAC02503FF8A37ABFAF1071E12C2" box="[916,1032,1392,1417]" italics="true" pageId="10" pageNumber="63">L. fragilis</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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