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<document id="1AB4B9A807B730F9F1CD45032C677E52" ID-DOI="10.5281/zenodo.276769" ID-GBIF-Dataset="f0d9b732-7675-4d3c-9902-2619c3576c4d" ID-ISSN="1175-5326" ID-Zenodo-Dep="276769" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460235349831" checkinUser="plazi" docAuthor="David, Patrick, Pham, Cuong The, Nguyen, Truong Quang &amp; Ziegler, Thomas" docDate="2011" docId="03D487A2FA72FFBCFF29FBF4FDA1FC87" docLanguage="en" docName="zt02758p056.pdf" docOrigin="Zootaxa 2758" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Opisthotropis" docType="key" docVersion="7" lastPageNumber="53" masterDocId="FFEDFFDAFA7BFFB6FFBEFFEFFFA8FF92" masterDocTitle="A new species of the genus Opisthotropis Günther, 1872 (Squamata: Natricidae) from the highlands of Kon Tum Province, Vietnam" masterLastPageNumber="56" masterPageNumber="43" pageNumber="52" updateTime="1698239489785" updateUser="plazi">
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<mods:title id="5A8A3EE39314C3A35C62A20EADDFE161">A new species of the genus Opisthotropis Günther, 1872 (Squamata: Natricidae) from the highlands of Kon Tum Province, Vietnam</mods:title>
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<mods:namePart id="573CF972417254934FBC980F966DE88C">David, Patrick</mods:namePart>
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<mods:namePart id="75DA15EF726C0ECCECDD0A1C01B1BFE2">Pham, Cuong The</mods:namePart>
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<mods:namePart id="09A12BAB6DA2A7410654AD572AF73D9B">Nguyen, Truong Quang</mods:namePart>
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<mods:namePart id="A1F1FA902D830DD75DB54F31019ACE4C">Ziegler, Thomas</mods:namePart>
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<mods:date id="6A577D7B6B729E0EA09F71F247761705">2011</mods:date>
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<mods:number id="BE16EFF6E60E58F208F3CA422F6000DE">2758</mods:number>
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<treatment id="03D487A2FA72FFBCFF29FBF4FDA1FC87" ID-DOI="http://doi.org/10.5281/zenodo.6182680" ID-GBIF-Taxon="119388036" ID-Zenodo-Dep="6182680" LSID="urn:lsid:plazi:treatment:03D487A2FA72FFBCFF29FBF4FDA1FC87" httpUri="http://treatment.plazi.org/id/03D487A2FA72FFBCFF29FBF4FDA1FC87" lastPageId="10" lastPageNumber="53" pageId="9" pageNumber="52">
<subSubSection id="C367653FFA72FFBFFF29FBF4FBFAFB50" pageId="9" pageNumber="52" type="nomenclature">
<paragraph id="8BC236B4FA72FFBFFF29FBF4FC98FBA7" blockId="9.[151,816,1051,1078]" box="[151,816,1051,1078]" pageId="9" pageNumber="52">
<heading id="D08A81D8FA72FFBFFF29FBF4FC98FBA7" bold="true" box="[151,816,1051,1078]" fontSize="11" level="1" pageId="9" pageNumber="52" reason="1">
<emphasis id="B909EAA6FA72FFBFFF29FBF4FC98FBA7" bold="true" box="[151,816,1051,1078]" pageId="9" pageNumber="52">
Key to the species of
<taxonomicName id="4C7D4D37FA72FFBFFE22FBF3FD97FBA4" box="[412,575,1052,1078]" class="Reptilia" family="Colubridae" genus="Opisthotropis" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="52" phylum="Chordata" rank="genus">
<emphasis id="B909EAA6FA72FFBFFE22FBF3FD97FBA4" bold="true" box="[412,575,1052,1078]" italics="true" pageId="9" pageNumber="52">Opisthotropis</emphasis>
</taxonomicName>
and related genera
</emphasis>
</heading>
</paragraph>
<paragraph id="8BC236B4FA72FFBFFF29FB8DFBFAFB50" blockId="9.[151,1436,1122,1218]" pageId="9" pageNumber="52">
On the basis of recent taxonomic modifications and of new material available subsequent to
<bibRefCitation id="EFEC4B45FA72FFBFFB0FFB8DFA3FFBE8" box="[1201,1431,1122,1147]" pageId="9" pageNumber="55" refString="Ziegler, T., David, P. &amp; Vu, N. T. (2008) A new natricine snake of the genus Opisthotropis from Tam Dao, Vinh Phuc Province, northern Vietnam (Squamata, Colubridae). Zoosystematics and Evolution, Berlin, 84 (2), 197 - 203." type="journal article">
Ziegler
<emphasis id="B909EAA6FA72FFBFFAB7FB8BFA93FBE9" box="[1289,1339,1123,1147]" italics="true" pageId="9" pageNumber="52">et al</emphasis>
. (2008)
</bibRefCitation>
, we provide a new key to the genus
<taxonomicName id="4C7D4D37FA72FFBFFD9BFB69FD16FB0C" box="[549,702,1158,1182]" class="Reptilia" family="Colubridae" genus="Opisthotropis" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="52" phylum="Chordata" rank="genus">
<emphasis id="B909EAA6FA72FFBFFD9BFB69FD16FB0C" box="[549,702,1158,1182]" italics="true" pageId="9" pageNumber="52">Opisthotropis</emphasis>
</taxonomicName>
and related genera. This key, as well as Table 1 above, are based on specimens examined by us and the references listed in the Materials and Methods.
</paragraph>
</subSubSection>
<subSubSection id="C367653FFA72FFBCFF29FB1DFDA1FC87" lastPageId="10" lastPageNumber="53" pageId="9" pageNumber="52" type="key">
<keyStep id="30892E11FA72FFBFFF29FB1DFA34FACD" pageId="9" pageNumber="52">
<paragraph id="8BC236B4FA72FFBFFF29FB1DFA33FAB7" blockId="9.[151,1437,1266,2020]" pageId="9" pageNumber="52">
<keyLead id="308C9581FA72FFBFFF29FB1DFA33FAB7" pageId="9" pageNumber="52">
1 Posterior upper maxillary teeth distinctly enlarged; nostrils rounded, dorsolateral, laterally directed; first supralabials distinctly longer than high...................................................................
<taxonomicName id="4C7D4D37FA72FFBFFB1BFAFFFA33FAB7" box="[1189,1435,1296,1317]" class="Reptilia" family="Colubridae" genus="Parahelicops" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="52" phylum="Chordata" rank="species" species="annamensis">
<emphasis id="B909EAA6FA72FFBFFB1BFAFFFA33FAB7" box="[1189,1435,1296,1317]" italics="true" pageId="9" pageNumber="52">Parahelicops annamensis</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BC236B4FA72FFBFFF29FAC2FA34FACD" blockId="9.[151,1437,1266,2020]" pageId="9" pageNumber="52">
<keyLead id="308C9581FA72FFBFFF29FAC2FA34FACD" pageId="9" pageNumber="52">- Upper maxillary teeth subequal; nostrils crescentic, in dorsal position, directed upwards; first supralabials distinctly higher than long............................................................................................ 2</keyLead>
</paragraph>
</keyStep>
<keyStep id="30892E11FA72FFBFFF29FA88FA33FA08" pageId="9" pageNumber="52">
<paragraph id="8BC236B4FA72FFBFFF29FA88FA34FAEE" blockId="9.[151,1437,1266,2020]" box="[151,1436,1383,1404]" pageId="9" pageNumber="52">
<keyLead id="308C9581FA72FFBFFF29FA88FA34FAEE" box="[151,1436,1383,1404]" pageId="9" pageNumber="52">2 15 or 17 scale rows at midbody and on neck............................................................... 3</keyLead>
</paragraph>
<paragraph id="8BC236B4FA72FFBFFF29FA6AFA33FA08" blockId="9.[151,1437,1266,2020]" box="[151,1435,1413,1434]" pageId="9" pageNumber="52">
<keyLead id="308C9581FA72FFBFFF29FA6AFA33FA08" box="[151,1435,1413,1434]" pageId="9" pageNumber="52">- 19 rows at midbody, 1923 rows on neck.................................................................. 16</keyLead>
</paragraph>
</keyStep>
<keyStep id="30892E11FA72FFBFFF29FA4DFA34FA46" pageId="9" pageNumber="52">
<paragraph id="8BC236B4FA72FFBFFF29FA4DFA34FA25" blockId="9.[151,1437,1266,2020]" box="[151,1436,1442,1463]" pageId="9" pageNumber="52">
<keyLead id="308C9581FA72FFBFFF29FA4DFA34FA25" box="[151,1436,1442,1463]" pageId="9" pageNumber="52">3 15 scale rows at midbody, 1517 scale rows on neck.......................................................... 4</keyLead>
</paragraph>
<paragraph id="8BC236B4FA72FFBFFF29FA50FA34FA46" blockId="9.[151,1437,1266,2020]" box="[151,1436,1471,1492]" pageId="9" pageNumber="52">
<keyLead id="308C9581FA72FFBFFF29FA50FA34FA46" box="[151,1436,1471,1492]" pageId="9" pageNumber="52">- 17 rows at midbody.................................................................................... 7</keyLead>
</paragraph>
</keyStep>
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<paragraph id="8BC236B4FA72FFBFFF29FA32FA34FA60" blockId="9.[151,1437,1266,2020]" box="[151,1436,1501,1522]" pageId="9" pageNumber="52">
<keyLead id="308C9581FA72FFBFFF29FA32FA34FA60" box="[151,1436,1501,1522]" pageId="9" pageNumber="52">
4 Internasal in contact with loreal...................................................................
<taxonomicName id="4C7D4D37FA72FFBFFAFDFA32FA34FA60" box="[1347,1436,1501,1522]" class="Reptilia" family="Colubridae" genus="Opisthotropis" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="52" phylum="Chordata" rank="species" species="jacobi">
<emphasis id="B909EAA6FA72FFBFFAFDFA32FA34FA60" box="[1347,1436,1501,1522]" italics="true" pageId="9" pageNumber="52">
<collectingCountry id="F36A7624FA72FFBFFAFDFA32FAFFFA60" box="[1347,1367,1501,1522]" name="Iceland" pageId="9" pageNumber="52">O</collectingCountry>
. jacobi
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BC236B4FA72FFBFFF29FA15FA34F99D" blockId="9.[151,1437,1266,2020]" box="[151,1436,1530,1551]" pageId="9" pageNumber="52">
<keyLead id="308C9581FA72FFBFFF29FA15FA34F99D" box="[151,1436,1530,1551]" pageId="9" pageNumber="52">- Internasal not in contact with loreal....................................................................... 5</keyLead>
</paragraph>
</keyStep>
<keyStep id="30892E11FA72FFBFFF29F9F8FA34F9D8" pageId="9" pageNumber="52">
<paragraph id="8BC236B4FA72FFBFFF29F9F8FA34F9BE" blockId="9.[151,1437,1266,2020]" box="[151,1436,1559,1580]" pageId="9" pageNumber="52">
<keyLead id="308C9581FA72FFBFFF29F9F8FA34F9BE" box="[151,1436,1559,1580]" pageId="9" pageNumber="52">5 1 preocular; 1 anterior temporal; dorsal scales entirely smooth.................................................. 6</keyLead>
</paragraph>
<paragraph id="8BC236B4FA72FFBFFF29F9DAFA34F9D8" blockId="9.[151,1437,1266,2020]" box="[151,1436,1589,1610]" pageId="9" pageNumber="52">
<keyLead id="308C9581FA72FFBFFF29F9DAFA34F9D8" box="[151,1436,1589,1610]" pageId="9" pageNumber="52">
- 2 preoculars; 2 anterior temporals; dorsal scales keeled posteriorly......................................
<taxonomicName id="4C7D4D37FA72FFBFFA96F9DAFA34F9D8" box="[1320,1436,1589,1610]" class="Reptilia" family="Colubridae" genus="Opisthotropis" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="52" phylum="Chordata" rank="species" species="kikuzatoi">
<emphasis id="B909EAA6FA72FFBFFA96F9DAFA34F9D8" box="[1320,1436,1589,1610]" italics="true" pageId="9" pageNumber="52">
<collectingCountry id="F36A7624FA72FFBFFA96F9DAFA94F9D8" box="[1320,1340,1589,1610]" name="Iceland" pageId="9" pageNumber="52">O</collectingCountry>
. kikuzatoi
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30892E11FA72FFBFFF29F9BDFA34F917" pageId="9" pageNumber="52">
<paragraph id="8BC236B4FA72FFBFFF29F9BDFA34F9F5" blockId="9.[151,1437,1266,2020]" box="[151,1436,1618,1639]" pageId="9" pageNumber="52">
<keyLead id="308C9581FA72FFBFFF29F9BDFA34F9F5" box="[151,1436,1618,1639]" pageId="9" pageNumber="52">
6 9 SL; body dark with pale crossbars..........................................................
<taxonomicName id="4C7D4D37FA72FFBFFABFF9BDFA34F9F5" box="[1281,1436,1618,1639]" class="Reptilia" family="Colubridae" genus="Opisthotropis" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="52" phylum="Chordata" rank="species" species="guangxiensis">
<emphasis id="B909EAA6FA72FFBFFABFF9BDFA34F9F5" box="[1281,1436,1618,1639]" italics="true" pageId="9" pageNumber="52">
<collectingCountry id="F36A7624FA72FFBFFABFF9BDFABDF9F5" box="[1281,1301,1618,1639]" name="Iceland" pageId="9" pageNumber="52">O</collectingCountry>
. guangxiensis
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BC236B4FA72FFBFFF29F980FA34F917" blockId="9.[151,1437,1266,2020]" box="[151,1436,1647,1669]" pageId="9" pageNumber="52">
<keyLead id="308C9581FA72FFBFFF29F980FA34F917" box="[151,1436,1647,1669]" pageId="9" pageNumber="52">
- 8 SL; body black with a tallow spot on each dorsal scale.............................................
<taxonomicName id="4C7D4D37FA72FFBFFA9DF99FFA34F917" box="[1315,1436,1648,1669]" class="Reptilia" family="Colubridae" genus="Opisthotropis" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="52" phylum="Chordata" rank="species" species="maculosa">
<emphasis id="B909EAA6FA72FFBFFA9DF99FFA34F917" box="[1315,1436,1648,1669]" italics="true" pageId="9" pageNumber="52">
<collectingCountry id="F36A7624FA72FFBFFA9DF99FFA9FF917" box="[1315,1335,1648,1669]" name="Iceland" pageId="9" pageNumber="52">O</collectingCountry>
. maculosa
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30892E11FA72FFBFFF29F962FA34F92D" pageId="9" pageNumber="52">
<paragraph id="8BC236B4FA72FFBFFF29F962FA34F930" blockId="9.[151,1437,1266,2020]" box="[151,1436,1677,1698]" pageId="9" pageNumber="52">
<keyLead id="308C9581FA72FFBFFF29F962FA34F930" box="[151,1436,1677,1698]" pageId="9" pageNumber="52">
7 Dorsal scales strongly keeled (endemic to Sumatra)..................................................
<taxonomicName id="4C7D4D37FA72FFBFFA82F962FA34F930" box="[1340,1436,1677,1698]" class="Reptilia" family="Colubridae" genus="Opisthotropis" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="52" phylum="Chordata" rank="species" species="rugosa">
<emphasis id="B909EAA6FA72FFBFFA82F962FA34F930" box="[1340,1436,1677,1698]" italics="true" pageId="9" pageNumber="52">
<collectingCountry id="F36A7624FA72FFBFFA82F962FAF8F930" box="[1340,1360,1677,1698]" name="Iceland" pageId="9" pageNumber="52">O</collectingCountry>
. rugosa
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BC236B4FA72FFBFFF29F945FA34F92D" blockId="9.[151,1437,1266,2020]" box="[151,1436,1706,1727]" pageId="9" pageNumber="52">
<keyLead id="308C9581FA72FFBFFF29F945FA34F92D" box="[151,1436,1706,1727]" pageId="9" pageNumber="52">- Dorsal scales smooth or weakly keeled on posterior part of body................................................ 8</keyLead>
</paragraph>
</keyStep>
<keyStep id="30892E11FA72FFBFFF29F928FA34F968" pageId="9" pageNumber="52">
<paragraph id="8BC236B4FA72FFBFFF29F928FA33F94F" blockId="9.[151,1437,1266,2020]" box="[151,1435,1735,1757]" pageId="9" pageNumber="52">
<keyLead id="308C9581FA72FFBFFF29F928FA33F94F" box="[151,1435,1735,1757]" pageId="9" pageNumber="52">
8 No preocular; elongate loreal entering orbit.......................................................
<taxonomicName id="4C7D4D37FA72FFBFFA97F927FA33F94F" box="[1321,1435,1736,1757]" class="Reptilia" family="Colubridae" genus="Opisthotropis" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="52" phylum="Chordata" rank="species" species="latouchii">
<emphasis id="B909EAA6FA72FFBFFA97F927FA33F94F" box="[1321,1435,1736,1757]" italics="true" pageId="9" pageNumber="52">
<collectingCountry id="F36A7624FA72FFBFFA97F927FA95F94F" box="[1321,1341,1736,1757]" name="Iceland" pageId="9" pageNumber="52">O</collectingCountry>
. latouchii
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BC236B4FA72FFBFFF29F90AFA34F968" blockId="9.[151,1437,1266,2020]" box="[151,1436,1765,1786]" pageId="9" pageNumber="52">
<keyLead id="308C9581FA72FFBFFF29F90AFA34F968" box="[151,1436,1765,1786]" pageId="9" pageNumber="52">- At least 1 preocular................................................................................... 9</keyLead>
</paragraph>
</keyStep>
<keyStep id="30892E11FA72FFBFFF29F8EDFA33F8A6" pageId="9" pageNumber="52">
<paragraph id="8BC236B4FA72FFBFFF29F8EDFA33F885" blockId="9.[151,1437,1266,2020]" box="[151,1435,1794,1815]" pageId="9" pageNumber="52">
<keyLead id="308C9581FA72FFBFFF29F8EDFA33F885" box="[151,1435,1794,1815]" pageId="9" pageNumber="52">9 Internasal in contact with loreal......................................................................... 10</keyLead>
</paragraph>
<paragraph id="8BC236B4FA72FFBFFF29F8F0FA33F8A6" blockId="9.[151,1437,1266,2020]" box="[151,1435,1823,1844]" pageId="9" pageNumber="52">
<keyLead id="308C9581FA72FFBFFF29F8F0FA33F8A6" box="[151,1435,1823,1844]" pageId="9" pageNumber="52">- Internasal not in contact with loreal...................................................................... 12</keyLead>
</paragraph>
</keyStep>
<keyStep id="30892E11FA72FFBFFF29F8D2FA33F8FD" pageId="9" pageNumber="52">
<paragraph id="8BC236B4FA72FFBFFF29F8D2FA34F8C0" blockId="9.[151,1437,1266,2020]" box="[151,1436,1853,1874]" pageId="9" pageNumber="52">
<keyLead id="308C9581FA72FFBFFF29F8D2FA34F8C0" box="[151,1436,1853,1874]" pageId="9" pageNumber="52">
10 Dorsal pattern uniformly blackish-brown; dorsal scales keeled posteriorly....................................
<taxonomicName id="4C7D4D37FA72FFBFFAE8F8D2FA34F8C0" box="[1366,1436,1853,1874]" class="Reptilia" family="Colubridae" genus="Opisthotropis" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="52" phylum="Chordata" rank="species" species="atra">
<emphasis id="B909EAA6FA72FFBFFAE8F8D2FA34F8C0" box="[1366,1436,1853,1874]" italics="true" pageId="9" pageNumber="52">
<collectingCountry id="F36A7624FA72FFBFFAE8F8D2FAC2F8C0" box="[1366,1386,1853,1874]" name="Iceland" pageId="9" pageNumber="52">O</collectingCountry>
. atra
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BC236B4FA72FFBFFF29F8B5FA33F8FD" blockId="9.[151,1437,1266,2020]" box="[151,1435,1882,1903]" pageId="9" pageNumber="52">
<keyLead id="308C9581FA72FFBFFF29F8B5FA33F8FD" box="[151,1435,1882,1903]" pageId="9" pageNumber="52">- Dorsal pattern uniformly greyish-brown; dorsal scales smooth................................................. 11</keyLead>
</paragraph>
</keyStep>
<keyStep id="30892E11FA72FFBFFF29F898FA34F838" pageId="9" pageNumber="52">
<paragraph id="8BC236B4FA72FFBFFF29F898FA34F81F" blockId="9.[151,1437,1266,2020]" box="[151,1436,1911,1933]" pageId="9" pageNumber="52">
<keyLead id="308C9581FA72FFBFFF29F898FA34F81F" box="[151,1436,1911,1933]" pageId="9" pageNumber="52">
11 Greyish-olive above; 189194 VEN; endemic to southern
<collectingCountry id="F36A7624FA72FFBFFCAFF898FCCAF81E" box="[785,866,1911,1932]" name="Vietnam" pageId="9" pageNumber="52">Vietnam</collectingCountry>
...................................
<taxonomicName id="4C7D4D37FA72FFBFFAB3F897FA34F81F" box="[1293,1436,1912,1933]" class="Reptilia" family="Colubridae" genus="Opisthotropis" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="52" phylum="Chordata" rank="species" species="daovantieni">
<emphasis id="B909EAA6FA72FFBFFAB3F897FA34F81F" box="[1293,1436,1912,1933]" italics="true" pageId="9" pageNumber="52">
<collectingCountry id="F36A7624FA72FFBFFAB3F897FA89F81F" box="[1293,1313,1912,1933]" name="Iceland" pageId="9" pageNumber="52">O</collectingCountry>
. daovantieni
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BC236B4FA72FFBFFF29F87AFA34F838" blockId="9.[151,1437,1266,2020]" box="[151,1436,1941,1962]" pageId="9" pageNumber="52">
<keyLead id="308C9581FA72FFBFFF29F87AFA34F838" box="[151,1436,1941,1962]" pageId="9" pageNumber="52">
- Greyish-brown above; 183 VEN; endemic to northern
<collectingCountry id="F36A7624FA72FFBFFD53F87AFCEAF838" box="[749,834,1941,1962]" name="Thailand" pageId="9" pageNumber="52">Thailand</collectingCountry>
.......................................
<taxonomicName id="4C7D4D37FA72FFBFFA93F87AFA34F838" box="[1325,1436,1941,1962]" class="Reptilia" family="Colubridae" genus="Opisthotropis" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="52" phylum="Chordata" rank="species" species="spenceri">
<emphasis id="B909EAA6FA72FFBFFA93F87AFA34F838" box="[1325,1436,1941,1962]" italics="true" pageId="9" pageNumber="52">
<collectingCountry id="F36A7624FA72FFBFFA93F87AFAE9F838" box="[1325,1345,1941,1962]" name="Iceland" pageId="9" pageNumber="52">O</collectingCountry>
. spenceri
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30892E11FA72FFBFFF29F85DFA34F876" pageId="9" pageNumber="52">
<paragraph id="8BC236B4FA72FFBFFF29F85DFA34F85A" blockId="9.[151,1437,1266,2020]" box="[151,1436,1970,1992]" pageId="9" pageNumber="52">
<keyLead id="308C9581FA72FFBFFF29F85DFA34F85A" box="[151,1436,1970,1992]" pageId="9" pageNumber="52">
12 Dorsal scales entirely smooth; dorsum with pale crossbars...............................................
<taxonomicName id="4C7D4D37FA72FFBFFAF5F85CFA34F85A" box="[1355,1436,1971,1992]" class="Reptilia" family="Colubridae" genus="Opisthotropis" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="52" phylum="Chordata" rank="species" species="cheni">
<emphasis id="B909EAA6FA72FFBFFAF5F85CFA34F85A" box="[1355,1436,1971,1992]" italics="true" pageId="9" pageNumber="52">
<collectingCountry id="F36A7624FA72FFBFFAF5F85CFAF7F85A" box="[1355,1375,1971,1992]" name="Iceland" pageId="9" pageNumber="52">O</collectingCountry>
. cheni
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BC236B4FA72FFBFFF29F820FA34F876" blockId="9.[151,1437,1266,2020]" box="[151,1436,1999,2020]" pageId="9" pageNumber="52">
<keyLead id="308C9581FA72FFBFFF29F820FA34F876" box="[151,1436,1999,2020]" pageId="9" pageNumber="52">- Dorsal scales at least keeled on posterior part of body; dorsal pattern not mainly made of crossbars.................... 13</keyLead>
</paragraph>
</keyStep>
<keyStep id="30892E11FA71FFBCFF29FF78FA33FF58" pageId="10" pageNumber="53">
<paragraph id="8BC236B4FA71FFBCFF29FF78FA34FF3F" blockId="10.[151,1436,151,789]" box="[151,1436,151,173]" pageId="10" pageNumber="53">
<keyLead id="308C9581FA71FFBCFF29FF78FA34FF3F" box="[151,1436,151,173]" pageId="10" pageNumber="53">
13 2 anterior temporals........................................................................
<taxonomicName id="4C7D4D37FA71FFBCFAB8FF77FA34FF3F" box="[1286,1436,152,173]" class="Reptilia" family="Colubridae" genus="Opisthotropis" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="53" phylum="Chordata" rank="species" species="tamdaoensis">
<emphasis id="B909EAA6FA71FFBCFAB8FF77FA34FF3F" box="[1286,1436,152,173]" italics="true" pageId="10" pageNumber="53">
<collectingCountry id="F36A7624FA71FFBCFAB8FF77FAB2FF3F" box="[1286,1306,152,173]" name="Iceland" pageId="10" pageNumber="53">O</collectingCountry>
. tamdaoensis
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BC236B4FA71FFBCFF29FF5AFA33FF58" blockId="10.[151,1436,151,789]" box="[151,1435,181,202]" pageId="10" pageNumber="53">
<keyLead id="308C9581FA71FFBCFF29FF5AFA33FF58" box="[151,1435,181,202]" pageId="10" pageNumber="53">- 1 anterior temporal................................................................................... 14</keyLead>
</paragraph>
</keyStep>
<keyStep id="30892E11FA71FFBCFF29FF3DFA34FE96" pageId="10" pageNumber="53">
<paragraph id="8BC236B4FA71FFBCFF29FF3DFA34FF75" blockId="10.[151,1436,151,789]" box="[151,1436,210,231]" pageId="10" pageNumber="53">
<keyLead id="308C9581FA71FFBCFF29FF3DFA34FF75" box="[151,1436,210,231]" pageId="10" pageNumber="53">
14 Body uniformly dark with pale-edged scales; 2 suboculars...........................................
<taxonomicName id="4C7D4D37FA71FFBCFAA6FF3DFA34FF75" box="[1304,1436,210,231]" class="Reptilia" family="Colubridae" genus="Opisthotropis" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="53" phylum="Chordata" rank="species" species="andersonii">
<emphasis id="B909EAA6FA71FFBCFAA6FF3DFA34FF75" box="[1304,1436,210,231]" italics="true" pageId="10" pageNumber="53">
<collectingCountry id="F36A7624FA71FFBCFAA6FF3DFA84FF75" box="[1304,1324,210,231]" name="Iceland" pageId="10" pageNumber="53">O</collectingCountry>
. andersonii
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BC236B4FA71FFBCFF29FF00FA34FE96" blockId="10.[151,1436,151,789]" box="[151,1436,239,260]" pageId="10" pageNumber="53">
<keyLead id="308C9581FA71FFBCFF29FF00FA34FE96" box="[151,1436,239,260]" pageId="10" pageNumber="53">- Body pattern not as such; no suboculars................................................................. 15</keyLead>
</paragraph>
</keyStep>
<keyStep id="30892E11FA71FFBCFF29FEE2FA34FEE8" pageId="10" pageNumber="53">
<paragraph id="8BC236B4FA71FFBCFF29FEE2FA34FEAD" blockId="10.[151,1436,151,789]" pageId="10" pageNumber="53">
<keyLead id="308C9581FA71FFBCFF29FEE2FA34FEAD" pageId="10" pageNumber="53">
15 A black lateral stripe separating abruptly pale colour of venter from dark dorsum colour; 159173 VEN; 911 SL; 1 preocular; dorsal scales keeled throughout body.............................................................
<taxonomicName id="4C7D4D37FA71FFBCFA90FEC5FA34FEAD" box="[1326,1436,298,319]" class="Reptilia" family="Colubridae" genus="Opisthotropis" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="53" phylum="Chordata" rank="species" species="lateralis">
<emphasis id="B909EAA6FA71FFBCFA90FEC5FA34FEAD" box="[1326,1436,298,319]" italics="true" pageId="10" pageNumber="53">
<collectingCountry id="F36A7624FA71FFBCFA90FEC5FAEAFEAD" box="[1326,1346,298,319]" name="Iceland" pageId="10" pageNumber="53">O</collectingCountry>
. lateralis
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BC236B4FA71FFBCFF29FEA8FA34FEE8" blockId="10.[151,1436,151,789]" pageId="10" pageNumber="53">
<keyLead id="308C9581FA71FFBCFF29FEA8FA34FEE8" pageId="10" pageNumber="53">
- No black lateral stripe separating pale colour of venter from dark dorsum colour; 147155 VEN; 68 SL; 2 preoculars; dorsal scales smooth on anterior part of body............................................................
<taxonomicName id="4C7D4D37FA71FFBCFA96FE8AFA34FEE8" box="[1320,1436,357,378]" class="Reptilia" family="Colubridae" genus="Opisthotropis" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="53" phylum="Chordata" rank="species" species="maxwelli">
<emphasis id="B909EAA6FA71FFBCFA96FE8AFA34FEE8" box="[1320,1436,357,378]" italics="true" pageId="10" pageNumber="53">
<collectingCountry id="F36A7624FA71FFBCFA96FE8AFA94FEE8" box="[1320,1340,357,378]" name="Iceland" pageId="10" pageNumber="53">O</collectingCountry>
. maxwelli
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30892E11FA71FFBCFF29FE6DFA34FE26" pageId="10" pageNumber="53">
<paragraph id="8BC236B4FA71FFBCFF29FE6DFA33FE05" blockId="10.[151,1436,151,789]" box="[151,1435,386,407]" pageId="10" pageNumber="53">
<keyLead id="308C9581FA71FFBCFF29FE6DFA33FE05" box="[151,1435,386,407]" pageId="10" pageNumber="53">16 Internasal in contact with loreal......................................................................... 17</keyLead>
</paragraph>
<paragraph id="8BC236B4FA71FFBCFF29FE70FA34FE26" blockId="10.[151,1436,151,789]" box="[151,1436,415,436]" pageId="10" pageNumber="53">
<keyLead id="308C9581FA71FFBCFF29FE70FA34FE26" box="[151,1436,415,436]" pageId="10" pageNumber="53">- Internasal not in contact with loreal..................................................................... 19</keyLead>
</paragraph>
</keyStep>
<keyStep id="30892E11FA71FFBCFF29FE52FA33FE7D" pageId="10" pageNumber="53">
<paragraph id="8BC236B4FA71FFBCFF29FE52FA34FE40" blockId="10.[151,1436,151,789]" box="[151,1436,445,466]" pageId="10" pageNumber="53">
<keyLead id="308C9581FA71FFBCFF29FE52FA34FE40" box="[151,1436,445,466]" pageId="10" pageNumber="53">
17 1 prefrontal...................................................................
<emphasis id="B909EAA6FA71FFBCFB3AFE52FA34FE40" bold="true" box="[1156,1436,445,466]" pageId="10" pageNumber="53">
<taxonomicName id="4C7D4D37FA71FFBCFB3AFE52FAE0FE40" box="[1156,1352,445,466]" class="Reptilia" family="Colubridae" genus="Opisthotropis" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="53" phylum="Chordata" rank="species" species="cucae" status="sp. nov.">
<emphasis id="B909EAA6FA71FFBCFB3AFE52FAE0FE40" bold="true" box="[1156,1352,445,466]" italics="true" pageId="10" pageNumber="53">Opisthotropis cucae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23A57DDFA71FFBCFAF1FE52FA34FE40" box="[1359,1436,445,466]" pageId="10" pageNumber="53" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8BC236B4FA71FFBCFF29FE35FA33FE7D" blockId="10.[151,1436,151,789]" box="[151,1435,474,495]" pageId="10" pageNumber="53">
<keyLead id="308C9581FA71FFBCFF29FE35FA33FE7D" box="[151,1435,474,495]" pageId="10" pageNumber="53">- 2 prefrontals......................................................................................... 18</keyLead>
</paragraph>
</keyStep>
<keyStep id="30892E11FA71FFBCFF29FE18FA34FDB8" pageId="10" pageNumber="53">
<paragraph id="8BC236B4FA71FFBCFF29FE18FA34FD9F" blockId="10.[151,1436,151,789]" box="[151,1436,503,525]" pageId="10" pageNumber="53">
<keyLead id="308C9581FA71FFBCFF29FE18FA34FD9F" box="[151,1436,503,525]" pageId="10" pageNumber="53">
18 Dorsal scales smooth; 195 VEN; endemic to the
<collectingCountry id="F36A7624FA71FFBCFD0AFE18FC88FD9E" box="[692,800,503,524]" name="Philippines" pageId="10" pageNumber="53">Philippines</collectingCountry>
............................................
<taxonomicName id="4C7D4D37FA71FFBCFA82FE17FA34FD9F" box="[1340,1436,504,525]" class="Reptilia" family="Colubridae" genus="Opisthotropis" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="53" phylum="Chordata" rank="species" species="alcalai">
<emphasis id="B909EAA6FA71FFBCFA82FE17FA34FD9F" box="[1340,1436,504,525]" italics="true" pageId="10" pageNumber="53">
<collectingCountry id="F36A7624FA71FFBCFA82FE17FAF8FD9F" box="[1340,1360,504,525]" name="Iceland" pageId="10" pageNumber="53">O</collectingCountry>
. alcalai
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BC236B4FA71FFBCFF29FDFAFA34FDB8" blockId="10.[151,1436,151,789]" box="[151,1436,533,554]" pageId="10" pageNumber="53">
<keyLead id="308C9581FA71FFBCFF29FDFAFA34FDB8" box="[151,1436,533,554]" pageId="10" pageNumber="53">
- Dorsal scales keeled; 160176 VEN; endemic to Borneo...............................................
<taxonomicName id="4C7D4D37FA71FFBCFAFAFDFAFA34FDB8" box="[1348,1436,533,554]" class="Reptilia" family="Colubridae" genus="Opisthotropis" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="53" phylum="Chordata" rank="species" species="typica">
<emphasis id="B909EAA6FA71FFBCFAFAFDFAFA34FDB8" box="[1348,1436,533,554]" italics="true" pageId="10" pageNumber="53">
<collectingCountry id="F36A7624FA71FFBCFAFAFDFAFAF0FDB8" box="[1348,1368,533,554]" name="Iceland" pageId="10" pageNumber="53">O</collectingCountry>
. typica
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30892E11FA71FFBCFF29FDDDFA33FDF6" pageId="10" pageNumber="53">
<paragraph id="8BC236B4FA71FFBCFF29FDDDFA33FDD5" blockId="10.[151,1436,151,789]" box="[151,1435,562,583]" pageId="10" pageNumber="53">
<keyLead id="308C9581FA71FFBCFF29FDDDFA33FDD5" box="[151,1435,562,583]" pageId="10" pageNumber="53">19 Dorsal scales strongly keeled........................................................................... 20</keyLead>
</paragraph>
<paragraph id="8BC236B4FA71FFBCFF29FDA0FA33FDF6" blockId="10.[151,1436,151,789]" box="[151,1435,591,612]" pageId="10" pageNumber="53">
<keyLead id="308C9581FA71FFBCFF29FDA0FA33FDF6" box="[151,1435,591,612]" pageId="10" pageNumber="53">- Dorsal scales smooth or only weakly keeled posteriorly...................................................... 21</keyLead>
</paragraph>
</keyStep>
<keyStep id="30892E11FA71FFBCFF29FD82FA34FD0D" pageId="10" pageNumber="53">
<paragraph id="8BC236B4FA71FFBCFF29FD82FA34FD10" blockId="10.[151,1436,151,789]" box="[151,1436,621,642]" pageId="10" pageNumber="53">
<keyLead id="308C9581FA71FFBCFF29FD82FA34FD10" box="[151,1436,621,642]" pageId="10" pageNumber="53">
20 Dorsal scales with strong, thick keels; 136140 VEN; 89 SL; dorsal pattern uniform.....................
<taxonomicName id="4C7D4D37FA71FFBCFAA5FD82FA34FD10" box="[1307,1436,621,642]" class="Reptilia" family="Colubridae" genus="Opisthotropis" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="53" phylum="Chordata" rank="species" species="boonsongi">
<emphasis id="B909EAA6FA71FFBCFAA5FD82FA34FD10" box="[1307,1436,621,642]" italics="true" pageId="10" pageNumber="53">
<collectingCountry id="F36A7624FA71FFBCFAA5FD82FA87FD10" box="[1307,1327,621,642]" name="Iceland" pageId="10" pageNumber="53">O</collectingCountry>
. boonsongi
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BC236B4FA71FFBCFF29FD65FA34FD0D" blockId="10.[151,1436,151,789]" box="[151,1436,650,671]" pageId="10" pageNumber="53">
<keyLead id="308C9581FA71FFBCFF29FD65FA34FD0D" box="[151,1436,650,671]" pageId="10" pageNumber="53">
- Dorsal scales with strong but narrow keels; 146154 VEN; 1316 SL; body with black stripes.............
<taxonomicName id="4C7D4D37FA71FFBCFAB3FD65FA34FD0D" box="[1293,1436,650,671]" class="Reptilia" family="Colubridae" genus="Opisthotropis" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="53" phylum="Chordata" rank="species" species="kuatunensis">
<emphasis id="B909EAA6FA71FFBCFAB3FD65FA34FD0D" box="[1293,1436,650,671]" italics="true" pageId="10" pageNumber="53">
<collectingCountry id="F36A7624FA71FFBCFAB3FD65FA89FD0D" box="[1293,1313,650,671]" name="Iceland" pageId="10" pageNumber="53">O</collectingCountry>
. kuatunensis
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30892E11FA71FFBCFF29FD48FDA1FC87" pageId="10" pageNumber="53">
<paragraph id="8BC236B4FA71FFBCFF29FD48FA34FD48" blockId="10.[151,1436,151,789]" pageId="10" pageNumber="53">
<keyLead id="308C9581FA71FFBCFF29FD48FA34FD48" pageId="10" pageNumber="53">
21 Dorsal scales keeled on posterior part of body; 190205 VEN; strong dorsal pattern made of broad, black, pale-centred cross- bars........................................................................................
<taxonomicName id="4C7D4D37FA71FFBCFA8EFD2AFA34FD48" box="[1328,1436,709,730]" class="Reptilia" family="Colubridae" genus="Opisthotropis" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="53" phylum="Chordata" rank="species" species="balteata">
<emphasis id="B909EAA6FA71FFBCFA8EFD2AFA34FD48" box="[1328,1436,709,730]" italics="true" pageId="10" pageNumber="53">
<collectingCountry id="F36A7624FA71FFBCFA8EFD2AFAECFD48" box="[1328,1348,709,730]" name="Iceland" pageId="10" pageNumber="53">O</collectingCountry>
. balteata
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BC236B4FA71FFBCFF29FD0DFDA1FC87" blockId="10.[151,1436,151,789]" pageId="10" pageNumber="53">
<keyLead id="308C9581FA71FFBCFF29FD0DFDA1FC87" pageId="10" pageNumber="53">
- Dorsal scales entirely smooth; 145155 VEN; subdued pattern with faint dark crossbands or dark, large blotches..........
<taxonomicName id="4C7D4D37FA71FFBCFF5FFCEFFDA1FC87" box="[225,521,768,789]" class="Reptilia" family="Colubridae" genus="Paratapinophis" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="53" phylum="Chordata" rank="species" species="praemaxillaris">
<emphasis id="B909EAA6FA71FFBCFF5FFCEFFDA1FC87" box="[225,521,768,789]" italics="true" pageId="10" pageNumber="53">Paratapinophis praemaxillaris</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
</document>