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<document id="7B64F3ED30EAF162A7FBAF492AE5F6E1" ID-CLB-Dataset="4026" ID-DOI="10.11646/zootaxa.5052.1.9" ID-GBIF-Dataset="07baf883-204c-4608-a8b3-924711319ef3" ID-ISSN="1175-5326" ID-Zenodo-Dep="5566148" ID-ZooBank="3E6DA6AB-32EB-473A-BD67-C8C59D4357BF" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1634107457561" checkinUser="plazi" docAuthor="Viidalepp, Jaan &amp; Kostjuk, Igor" docDate="2021" docId="03D48798FFA0347554FB1D3BEC03F838" docLanguage="en" docName="zootaxa.5052.1.9.pdf" docOrigin="Zootaxa 5052 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Hierochthonia Prout 1912" docType="treatment" docVersion="9" lastPageNumber="140" masterDocId="FFEDFFE0FFA33476546C1864E845FF81" masterDocTitle="Taxonomic notes on two endemic geometrine genera from Middle Asia (Lepidoptera: Geometridae, Geometrinae)" masterLastPageNumber="144" masterPageNumber="137" pageNumber="140" updateTime="1699459903341" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="5E458E42F41EB963399A24DFFE50FF5D">Taxonomic notes on two endemic geometrine genera from Middle Asia (Lepidoptera: Geometridae, Geometrinae)</mods:title>
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<mods:namePart id="6C4331C4F94207FA500ABA3951E934B0">Viidalepp, Jaan</mods:namePart>
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<mods:namePart id="41141B0D2A8AD753FE450FD3966AB481">Kostjuk, Igor</mods:namePart>
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<mods:affiliation id="30A5182E9A7DB604056B63C7188AA1DB">Zoological museum, Kyiv National Taras Shevchenko University, Volodymyrska str. 60, UA- 01601 Kyiv, UKRAINE. ikostjuk @ univ. kiev. ua; https: // orcid. org / 0000 - 0002 - 8656 - 5330</mods:affiliation>
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<treatment id="03D48798FFA0347554FB1D3BEC03F838" ID-DOI="http://doi.org/10.5281/zenodo.5699320" ID-GBIF-Taxon="188647881" ID-Zenodo-Dep="5699320" LSID="urn:lsid:plazi:treatment:03D48798FFA0347554FB1D3BEC03F838" httpUri="http://treatment.plazi.org/id/03D48798FFA0347554FB1D3BEC03F838" lastPageNumber="140" pageId="3" pageNumber="140">
<subSubSection id="C3676505FFA0347554FB1D3BEBD3FAF8" box="[151,918,1375,1402]" pageId="3" pageNumber="140" type="nomenclature">
<paragraph id="8BC2368EFFA0347554FB1D3BEBD3FAF8" blockId="3.[151,918,1375,1402]" box="[151,918,1375,1402]" pageId="3" pageNumber="140">
<heading id="D08A81E2FFA0347554FB1D3BEBD3FAF8" bold="true" box="[151,918,1375,1402]" fontSize="11" level="1" pageId="3" pageNumber="140" reason="1">
<emphasis id="B909EA9CFFA0347554FB1D3BEBD3FAF8" bold="true" box="[151,918,1375,1402]" pageId="3" pageNumber="140">
<taxonomicName id="4C7D4D0DFFA0347554FB1D3BE9A4FAF8" ID-CoL="92BLV" authority="Prout, 1912" authorityName="Prout" authorityYear="1912" box="[151,481,1375,1402]" class="Insecta" family="Geometridae" genus="Hierochthonia" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="140" phylum="Arthropoda" rank="genus">
<emphasis id="B909EA9CFFA0347554FB1D3BE90CFAFB" bold="true" box="[151,329,1375,1402]" italics="true" pageId="3" pageNumber="140">Hierochthonia</emphasis>
Prout, 1912
</taxonomicName>
and “
<emphasis id="B909EA9CFFA0347556411D3BEA01FAF8" bold="true" box="[557,580,1375,1401]" italics="true" pageId="3" pageNumber="140">H</emphasis>
.”
<emphasis id="B909EA9CFFA03475560D1D3BEAB8FAFB" bold="true" box="[609,765,1375,1402]" italics="true" pageId="3" pageNumber="140">alexandraria</emphasis>
Prout, 1912
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3676505FFA0347554FB1DCCEC03F838" pageId="3" pageNumber="140" type="discussion">
<paragraph id="8BC2368EFFA0347554FB1DCCE9FDF9F4" blockId="3.[151,1437,1447,1978]" pageId="3" pageNumber="140">
Prout diagnosed the genus
<taxonomicName id="4C7D4D0DFFA0347555B21DC3EAC5FA40" authorityName="Prout" authorityYear="1912" box="[478,640,1447,1473]" class="Insecta" family="Geometridae" genus="Hierochthonia" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="140" phylum="Arthropoda" rank="genus">
<emphasis id="B909EA9CFFA0347555B21DC3EAC5FA40" box="[478,640,1447,1473]" italics="true" pageId="3" pageNumber="140">Hierochthonia</emphasis>
</taxonomicName>
(
<typeStatus id="54C6882CFFA0347556F91DC3EA8BFA40" box="[661,718,1447,1473]" pageId="3" pageNumber="140">Type</typeStatus>
species:
<taxonomicName id="4C7D4D0DFFA03475572E1DC3ECA7FA40" authority="Warren, 1901" authorityName="Warren" authorityYear="1901" box="[834,1250,1447,1474]" class="Insecta" family="Geometridae" genus="Microloxia" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="140" phylum="Arthropoda" rank="species" species="pulverata">
<emphasis id="B909EA9CFFA03475572E1DC3EC73FA40" box="[834,1078,1447,1473]" italics="true" pageId="3" pageNumber="140">Microloxia pulverata</emphasis>
Warren, 1901
</taxonomicName>
) using external characters as antennae bipectinate up to the last apical segments; hindwing with the subcostal vein fused with the fore margin of the discal cell near to its end; hindtibia provided with one pair of spurs (Prout 1912: 204).
<bibRefCitation id="EFEC4B7FFFA0347551481D94E8A7F9AF" author="Hausmann, A." pageId="3" pageNumber="140" pagination="3 - 90" refId="ref4374" refString="Hausmann, A. (1996) The morphology of the geometrid moths of the Levant and neighbouring countries. Part I: Orthostixinae, Geometrinae. Nota lepidopterologica, 19 (1 / 2), 3 - 90." type="journal article" year="1996">Hausmann (1996)</bibRefCitation>
redescribed the genus
<taxonomicName id="4C7D4D0DFFA0347555881E77EAC3F9AC" authorityName="Prout" authorityYear="1912" box="[484,646,1555,1581]" class="Insecta" family="Geometridae" genus="Hierochthonia" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="140" phylum="Arthropoda" rank="genus">
<emphasis id="B909EA9CFFA0347555881E77EAC3F9AC" box="[484,646,1555,1581]" italics="true" pageId="3" pageNumber="140">Hierochthonia</emphasis>
</taxonomicName>
using
<taxonomicName id="4C7D4D0DFFA0347556BE1E70EBFCF9AF" authority="Warren" authorityName="Warren" authorityYear="1901" box="[722,953,1555,1582]" class="Insecta" family="Geometridae" genus="Microloxia" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="140" phylum="Arthropoda" rank="species" species="pulverata">
<emphasis id="B909EA9CFFA0347556BE1E70EB25F9AC" box="[722,864,1555,1581]" italics="true" pageId="3" pageNumber="140">H. pulverata</emphasis>
Warren
</taxonomicName>
and
<taxonomicName id="4C7D4D0DFFA03475579C1E70ED3BF9AC" authority="Pungeler. Comparison" authorityName="Pungeler. Comparison" baseAuthorityName="Pungeler" baseAuthorityYear="1902" box="[1008,1406,1555,1581]" class="Insecta" family="Geometridae" genus="Hierochthonia" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="140" phylum="Arthropoda" rank="species" species="semitaria">
<emphasis id="B909EA9CFFA03475579C1E70EC38F9AC" box="[1008,1149,1556,1581]" italics="true" pageId="3" pageNumber="140">H. semitaria</emphasis>
Püngeler. Comparison
</taxonomicName>
of the characteristics of
<taxonomicName id="4C7D4D0DFFA0347555E51E5CEA7AF9D0" authorityName="Prout" authorityYear="1912" box="[393,575,1591,1617]" class="Insecta" family="Geometridae" genus="Hierochthonia" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="140" phylum="Arthropoda" rank="species" species="alexandraria">
<emphasis id="B909EA9CFFA0347555E51E5CEA7AF9D0" box="[393,575,1591,1617]" italics="true" pageId="3" pageNumber="140">H. alexandraria</emphasis>
</taxonomicName>
(also
<figureCitation id="13462A0BFFA0347556E91E53EAB2F9D0" box="[645,759,1591,1617]" captionStart="FIGURES 113" captionStartId="1.[151,264,1398,1422]" captionTargetBox="[202,1386,800,1372]" captionTargetId="figure-205@1.[202,1386,800,1372]" captionTargetPageId="1" captionText="FIGURES 113. Adult moths of the studied taxa. 12: Hissarica postalbida: 1—male, Tajikistan, Kondara 2—female, Kondara; 34: Xenochlorodes albicostaria: 3—male, Iran, Comèe, 4female, Comèe; 56: Xenochlorodes olympiaria: 5—male, Dalmatia, Gravosa, 6female, Griechenland, Delphi; 78: Hierochthonia pulverata, 7—male, Syria, 8—female, Syria; 911: Ratsa alexandraria: 9—Holotype, female, “Alexandergebirge”, 10—male, Kazakhstan, Kentau, 11—female, Kentau; 1213: Ratsa? petitaria: 12—Lectotype, female, “Askhabad” 13—female, Turkmenistan, Imam-Baba." figureDoi="http://doi.org/10.5281/zenodo.5566152" httpUri="https://zenodo.org/record/5566152/files/figure.png" pageId="3" pageNumber="140">Figs 911</figureCitation>
,
<figureCitation id="13462A0BFFA0347557681E53EB64F9D0" box="[772,801,1591,1617]" captionStart="FIGURES 2226" captionStartId="5.[151,264,1897,1921]" captionTargetBox="[253,1325,784,1858]" captionTargetId="figure-271@5.[248,1339,777,1872]" captionTargetPageId="5" captionText="FIGURES 2226. Genitalia structures of Hierochthonia, Ratsa, gen. nov. and Xenochlorodes. 22: Hierochthonia pulverata: 22a—male armature, 22b—aedeagus, 22c—VIII sternite, 22d—lateral view; 2324: Ratsa alexandraria: 23a—male armature, 23b—aedeagus, 23c—VIII sternite, 23d—lateral view, 24a—female genitalia, lateral view, 24b—female genitalia, ventral view; 25—female genitalia of Ratsa? petitaria; 26—female genitalia of Xenochlorodes albicostaria. Scale bar, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5566160" httpUri="https://zenodo.org/record/5566160/files/figure.png" pageId="3" pageNumber="140">23</figureCitation>
) with those of southern species (
<figureCitation id="13462A0BFFA0347550FB1E53ECB8F9D0" box="[1175,1277,1591,1618]" captionStart="FIGURES 113" captionStartId="1.[151,264,1398,1422]" captionTargetBox="[202,1386,800,1372]" captionTargetId="figure-205@1.[202,1386,800,1372]" captionTargetPageId="1" captionText="FIGURES 113. Adult moths of the studied taxa. 12: Hissarica postalbida: 1—male, Tajikistan, Kondara 2—female, Kondara; 34: Xenochlorodes albicostaria: 3—male, Iran, Comèe, 4female, Comèe; 56: Xenochlorodes olympiaria: 5—male, Dalmatia, Gravosa, 6female, Griechenland, Delphi; 78: Hierochthonia pulverata, 7—male, Syria, 8—female, Syria; 911: Ratsa alexandraria: 9—Holotype, female, “Alexandergebirge”, 10—male, Kazakhstan, Kentau, 11—female, Kentau; 1213: Ratsa? petitaria: 12—Lectotype, female, “Askhabad” 13—female, Turkmenistan, Imam-Baba." figureDoi="http://doi.org/10.5281/zenodo.5566152" httpUri="https://zenodo.org/record/5566152/files/figure.png" pageId="3" pageNumber="140">Figs 7, 8</figureCitation>
,
<figureCitation id="13462A0BFFA0347551671E53ED6DF9D0" box="[1291,1320,1591,1617]" captionStart="FIGURES 2226" captionStartId="5.[151,264,1897,1921]" captionTargetBox="[253,1325,784,1858]" captionTargetId="figure-271@5.[248,1339,777,1872]" captionTargetPageId="5" captionText="FIGURES 2226. Genitalia structures of Hierochthonia, Ratsa, gen. nov. and Xenochlorodes. 22: Hierochthonia pulverata: 22a—male armature, 22b—aedeagus, 22c—VIII sternite, 22d—lateral view; 2324: Ratsa alexandraria: 23a—male armature, 23b—aedeagus, 23c—VIII sternite, 23d—lateral view, 24a—female genitalia, lateral view, 24b—female genitalia, ventral view; 25—female genitalia of Ratsa? petitaria; 26—female genitalia of Xenochlorodes albicostaria. Scale bar, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5566160" httpUri="https://zenodo.org/record/5566160/files/figure.png" pageId="3" pageNumber="140">22</figureCitation>
) indicates the following differences.
</paragraph>
<paragraph id="8BC2368EFFA0347554AB1E1BEBF2F963" blockId="3.[151,1437,1447,1978]" pageId="3" pageNumber="140">
Both
<emphasis id="B909EA9CFFA0347555641E1BEA1DF918" box="[264,600,1663,1689]" italics="true" pageId="3" pageNumber="140">
Hierochthonia--
<taxonomicName id="4C7D4D0DFFA0347555AB1EE4EA1DF918" authorityName="Warren" authorityYear="1901" box="[455,600,1663,1689]" class="Insecta" family="Geometridae" genus="Microloxia" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="140" phylum="Arthropoda" rank="species" species="pulverata">H. pulverata</taxonomicName>
</emphasis>
and
<emphasis id="B909EA9CFFA0347556F91EE4EAECF918" box="[661,681,1664,1689]" italics="true" pageId="3" pageNumber="140">H</emphasis>
.
<emphasis id="B909EA9CFFA0347556D71EE4EB72F918" box="[699,823,1664,1689]" italics="true" pageId="3" pageNumber="140">semitaria--</emphasis>
possess long, flat postvaginal lamellae and roundish lateral sclerites attached to the ostium in female genitalia (
<bibRefCitation id="EFEC4B7FFFA03475575B1EC7EBB1F93C" author="Hausmann, A." box="[823,1012,1699,1725]" pageId="3" pageNumber="140" pagination="3 - 90" refId="ref4374" refString="Hausmann, A. (1996) The morphology of the geometrid moths of the Levant and neighbouring countries. Part I: Orthostixinae, Geometrinae. Nota lepidopterologica, 19 (1 / 2), 3 - 90." type="journal article" year="1996">Hausmann 1996</bibRefCitation>
, Figs 144, 145). These structures are missing in “
<emphasis id="B909EA9CFFA0347555731EACE9A7F960" box="[287,482,1735,1762]" italics="true" pageId="3" pageNumber="140">
H.”
<taxonomicName id="4C7D4D0DFFA03475553C1EA3E9A7F960" authorityName="Prout" authorityYear="1912" box="[336,482,1735,1761]" class="Insecta" family="Geometridae" genus="Hierochthonia" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="140" phylum="Arthropoda" rank="species" species="alexandraria">alexandraria</taxonomicName>
</emphasis>
and
<emphasis id="B909EA9CFFA0347556751EACEAFFF960" box="[537,698,1736,1762]" italics="true" pageId="3" pageNumber="140">
“H.”
<taxonomicName id="4C7D4D0DFFA0347556361EACEAFFF960" authorityName="Christoph" authorityYear="1887" box="[602,698,1736,1761]" class="Insecta" family="Geometridae" genus="Eucrostes" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="140" phylum="Arthropoda" rank="species" species="petitaria">petitaria</taxonomicName>
</emphasis>
females (
<figureCitation id="13462A0BFFA0347557441EA3EBEFF963" box="[808,938,1735,1762]" captionStart="FIGURES 2226" captionStartId="5.[151,264,1897,1921]" captionTargetBox="[253,1325,784,1858]" captionTargetId="figure-271@5.[248,1339,777,1872]" captionTargetPageId="5" captionText="FIGURES 2226. Genitalia structures of Hierochthonia, Ratsa, gen. nov. and Xenochlorodes. 22: Hierochthonia pulverata: 22a—male armature, 22b—aedeagus, 22c—VIII sternite, 22d—lateral view; 2324: Ratsa alexandraria: 23a—male armature, 23b—aedeagus, 23c—VIII sternite, 23d—lateral view, 24a—female genitalia, lateral view, 24b—female genitalia, ventral view; 25—female genitalia of Ratsa? petitaria; 26—female genitalia of Xenochlorodes albicostaria. Scale bar, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5566160" httpUri="https://zenodo.org/record/5566160/files/figure.png" pageId="3" pageNumber="140">Figs 24, 25</figureCitation>
).
</paragraph>
<paragraph id="8BC2368EFFA0347554AB1E8FEC03F838" blockId="3.[151,1437,1447,1978]" pageId="3" pageNumber="140">
The valva in male genitalia of typical
<taxonomicName id="4C7D4D0DFFA03475560B1E8FEB4CF884" authorityName="Prout" authorityYear="1912" box="[615,777,1771,1797]" class="Insecta" family="Geometridae" genus="Hierochthonia" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="140" phylum="Arthropoda" rank="genus">
<emphasis id="B909EA9CFFA03475560B1E8FEB4CF884" box="[615,777,1771,1797]" italics="true" pageId="3" pageNumber="140">Hierochthonia</emphasis>
</taxonomicName>
has a medial harpe (
<figureCitation id="13462A0BFFA0347557861E8FEC05F884" box="[1002,1088,1771,1797]" captionStart="FIGURES 2226" captionStartId="5.[151,264,1897,1921]" captionTargetBox="[253,1325,784,1858]" captionTargetId="figure-271@5.[248,1339,777,1872]" captionTargetPageId="5" captionText="FIGURES 2226. Genitalia structures of Hierochthonia, Ratsa, gen. nov. and Xenochlorodes. 22: Hierochthonia pulverata: 22a—male armature, 22b—aedeagus, 22c—VIII sternite, 22d—lateral view; 2324: Ratsa alexandraria: 23a—male armature, 23b—aedeagus, 23c—VIII sternite, 23d—lateral view, 24a—female genitalia, lateral view, 24b—female genitalia, ventral view; 25—female genitalia of Ratsa? petitaria; 26—female genitalia of Xenochlorodes albicostaria. Scale bar, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5566160" httpUri="https://zenodo.org/record/5566160/files/figure.png" pageId="3" pageNumber="140">Figs 22</figureCitation>
), and aedeagus is spoon shaped (distally dilated) with a complex apical sclerotization. However, “
<emphasis id="B909EA9CFFA0347557E11F74EC16F8A8" box="[909,1107,1807,1834]" italics="true" pageId="3" pageNumber="140">
H.”
<taxonomicName id="4C7D4D0DFFA0347557AD1F6BEC16F8A8" authorityName="Prout" authorityYear="1912" box="[961,1107,1807,1833]" class="Insecta" family="Geometridae" genus="Hierochthonia" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="140" phylum="Arthropoda" rank="species" species="alexandraria">alexandraria</taxonomicName>
</emphasis>
has a tubular aedeagus with two thorn shaped cornuti on the vesica, and a simple valva (
<figureCitation id="13462A0BFFA03475575A1F57EBCEF8CC" box="[822,907,1843,1869]" captionStart="FIGURES 2226" captionStartId="5.[151,264,1897,1921]" captionTargetBox="[253,1325,784,1858]" captionTargetId="figure-271@5.[248,1339,777,1872]" captionTargetPageId="5" captionText="FIGURES 2226. Genitalia structures of Hierochthonia, Ratsa, gen. nov. and Xenochlorodes. 22: Hierochthonia pulverata: 22a—male armature, 22b—aedeagus, 22c—VIII sternite, 22d—lateral view; 2324: Ratsa alexandraria: 23a—male armature, 23b—aedeagus, 23c—VIII sternite, 23d—lateral view, 24a—female genitalia, lateral view, 24b—female genitalia, ventral view; 25—female genitalia of Ratsa? petitaria; 26—female genitalia of Xenochlorodes albicostaria. Scale bar, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5566160" httpUri="https://zenodo.org/record/5566160/files/figure.png" pageId="3" pageNumber="140">Fig. 23</figureCitation>
). Prout (1912) stressed the peculiar venation of hindwing with Sc free when characterizing “
<emphasis id="B909EA9CFFA0347556FB1F33EBA1F8F0" box="[663,996,1879,1905]" italics="true" pageId="3" pageNumber="140">
<taxonomicName id="4C7D4D0DFFA0347556FB1F33EB7AF8F0" authorityName="Prout" authorityYear="1912" box="[663,831,1879,1905]" class="Insecta" family="Geometridae" genus="Hierochthonia" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="140" phylum="Arthropoda" rank="genus">Hierochthonia</taxonomicName>
<taxonomicName id="4C7D4D0DFFA03475573E1F33EBA1F8F0" authorityName="Prout" authorityYear="1912" box="[850,996,1879,1905]" class="Insecta" family="Geometridae" genus="Hierochthonia" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="140" phylum="Arthropoda" rank="species" species="alexandraria">alexandraria</taxonomicName>
</emphasis>
. We use this synergy of differences in wing venation and in male and female genitalia characteristics to separate “
<emphasis id="B909EA9CFFA0347557861F18EBBBF814" box="[1002,1022,1916,1941]" italics="true" pageId="3" pageNumber="140">H</emphasis>
.”
<taxonomicName id="4C7D4D0DFFA0347550751F1FECEEF814" authorityName="Prout" authorityYear="1912" box="[1049,1195,1915,1941]" class="Insecta" family="Geometridae" genus="Hierochthonia" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="140" phylum="Arthropoda" rank="species" species="alexandraria">
<emphasis id="B909EA9CFFA0347550751F1FECEEF814" box="[1049,1195,1915,1941]" italics="true" pageId="3" pageNumber="140">alexandraria</emphasis>
</taxonomicName>
in a new genus of its own,
<taxonomicName id="4C7D4D0DFFA0347554B91FC4EA4DF838" authority="Viidalepp &amp; Kostjuk" authorityName="Viidalepp &amp; Kostjuk" authorityYear="2021" box="[213,520,1951,1977]" class="Insecta" family="Geometridae" genus="Ratsa" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="140" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B909EA9CFFA0347554B91FC4E950F838" box="[213,277,1952,1977]" italics="true" pageId="3" pageNumber="140">Ratsa</emphasis>
Viidalepp &amp; Kostjuk
</taxonomicName>
,
<taxonomicNameLabel id="A23A57E7FFA03475567F1FFBEA38F83B" box="[531,637,1951,1978]" pageId="3" pageNumber="140" rank="genus">
<emphasis id="B909EA9CFFA03475567F1FFBEA32F838" bold="true" box="[531,631,1951,1977]" pageId="3" pageNumber="140">gen. nov</emphasis>
.
</taxonomicNameLabel>
will be described below for this species.
</paragraph>
</subSubSection>
</treatment>
</document>