336 lines
43 KiB
XML
336 lines
43 KiB
XML
<document id="AD2C6F7115A68AC87D3C2BF530A12682" ID-CLB-Dataset="27465" ID-DOI="10.11646/zootaxa.4550.4.6" ID-GBIF-Dataset="aed90136-53f9-4de1-944a-0fd6c8433dc8" ID-ISSN="1175-5326" ID-Zenodo-Dep="2625675" ID-ZooBank="188DEE45-B408-4766-98B0-109C7E7B3B7E" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1554277448317" checkinUser="plazi" docAuthor="Bukejs, Andris, Alekseev, Vitalii I., Cooper, David M. L., King, Gavin A. & Mckellar, Ryan C." docDate="2019" docId="03D48789FFB10E77FF4E90BDFA69FCC2" docLanguage="en" docName="zootaxa.4550.4.6.pdf" docOrigin="Zootaxa 4550 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pycnomerus agtsteinicus Bukejs & Alekseev & Cooper & King & Mckellar 2019, sp. nov." docType="treatment" docUuid="64C8B794-3D69-49AC-B3B1-A9EA13C31314" docUuidSource="ZooBank" docVersion="9" lastPageNumber="571" masterDocId="FFEDFFF1FFB00E71FFD9972AFFF5FFD7" masterDocTitle="A new fossil species of Pycnomerus Erichson (Coleoptera: Zopheridae) from Baltic amber, and a replacement name for a Recent North American congener" masterLastPageNumber="572" masterPageNumber="565" pageNumber="566" updateTime="1698717167131" updateUser="ExternalLinkService">
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<mods:title id="3BEB6C6D9F433412162EDC8CAD1264C1">A new fossil species of Pycnomerus Erichson (Coleoptera: Zopheridae) from Baltic amber, and a replacement name for a Recent North American congener</mods:title>
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<mods:name id="FB7078F102A00174DE9DC4DE555A3983" type="personal">
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<mods:namePart id="9043F8FF5A072E27290605DAD0053702">Bukejs, Andris</mods:namePart>
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</mods:name>
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<mods:roleTerm id="C06D33EA6E7294D77EA42D778651EC7C">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="859CFF374BB5B45904AF754DB4399C85">Alekseev, Vitalii I.</mods:namePart>
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</mods:name>
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<mods:name id="A1FC873571E20374E0EC7060A886EDD3" type="personal">
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<mods:roleTerm id="3D40C7E9400328E1F0C391769DFD76D9">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="3F850B41A1D25A41B9A44D70ED58CC76">Cooper, David M. L.</mods:namePart>
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</mods:role>
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<mods:namePart id="69396DF6792BA719C0B56033AB6B54E3">King, Gavin A.</mods:namePart>
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<mods:name id="9A27AC3CFF4867C747F0DA82073169C5" type="personal">
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<mods:roleTerm id="729196BC7DCA5E9D8D2657D7EE38E4B0">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="428AF87F4DCD5F2EAC92C2ACF078494C">Mckellar, Ryan C.</mods:namePart>
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<mods:typeOfResource id="ED70F9AF5A0842B5563B22E364C2CD0F">text</mods:typeOfResource>
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<mods:title id="69CE30F7F0036E4E685369DA9B162234">Zootaxa</mods:title>
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<mods:part id="587A97A94142B13957ED6FEA07B3618C">
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<mods:date id="5C61155EC86DCB15B1500728CE145064">2019</mods:date>
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<mods:number id="E2C6800B7E48F34083045062C60BCDB8">2019-01-29</mods:number>
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<mods:detail id="2D8A87FC1F146C2F185224CD242DEBC2" type="volume">
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<mods:number id="A1937C70958FE3897293210C577DB2B6">4550</mods:number>
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<mods:number id="DA87D79EE449EC34082BA55AFCE7711B">4</mods:number>
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<mods:identifier id="12897515EB7B0D1C84B92F812D0F8934" type="DOI">10.11646/zootaxa.4550.4.6</mods:identifier>
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<mods:identifier id="F206C17A8BB3BCC30FBEA4E5A9F8391B" type="ZooBank">188DEE45-B408-4766-98B0-109C7E7B3B7E</mods:identifier>
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<treatment id="03D48789FFB10E77FF4E90BDFA69FCC2" ID-DOI="http://doi.org/10.5281/zenodo.5921816" ID-GBIF-Taxon="154533193" ID-Zenodo-Dep="5921816" LSID="urn:lsid:zoobank.org:act:64C8B794-3D69-49AC-B3B1-A9EA13C31314" httpUri="http://treatment.plazi.org/id/03D48789FFB10E77FF4E90BDFA69FCC2" lastPageId="6" lastPageNumber="571" pageId="1" pageNumber="566">
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<subSubSection id="C3676514FFB10E70FF4E90BDFC8AF878" box="[151,895,1942,1967]" pageId="1" pageNumber="566" type="nomenclature">
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<paragraph id="8BC2369FFFB10E70FF4E90BDFC8AF878" blockId="1.[151,911,1942,2036]" box="[151,895,1942,1967]" pageId="1" pageNumber="566">
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<heading id="D08A81F3FFB10E70FF4E90BDFC8AF878" bold="true" box="[151,895,1942,1967]" fontSize="10" level="2" pageId="1" pageNumber="566" reason="2">
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<emphasis id="B909EA8DFFB10E70FF4E90BDFC8AF878" bold="true" box="[151,895,1942,1967]" pageId="1" pageNumber="566">
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<taxonomicName id="4C7D4D1CFFB10E70FF4E90BDFCEAF878" authority="Bukejs, Alekseev & McKellar" authorityName="Bukejs & Alekseev & Cooper & King & Mckellar" authorityYear="2019" box="[151,799,1942,1967]" class="Insecta" family="Zopheridae" genus="Pycnomerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="566" phylum="Arthropoda" rank="species" species="agtsteinicus" status="sp. nov.">
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<emphasis id="B909EA8DFFB10E70FF4E90BDFE43F879" bold="true" box="[151,438,1942,1966]" italics="true" pageId="1" pageNumber="566">Pycnomerus agtsteinicus</emphasis>
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Bukejs, Alekseev & McKellar
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</taxonomicName>
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<taxonomicNameLabel id="A23A57F6FFB10E70FCFF90BCFC8AF878" box="[806,895,1942,1967]" pageId="1" pageNumber="566" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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||
<subSubSection id="C3676514FFB10E70FF4E9090FC7AF823" pageId="1" pageNumber="566" type="description">
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<paragraph id="8BC2369FFFB10E70FF4E9090FEFFF804" blockId="1.[151,911,1942,2036]" box="[151,266,1978,2003]" pageId="1" pageNumber="566">
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(
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<figureCitation id="13462A1AFFB10E70FF469090FEF4F804" box="[159,257,1978,2003]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStartId-0="3.[151,250,1944,1966]" captionStartId-1="4.[151,250,1946,1968]" captionStartId-2="5.[151,250,1561,1583]" captionTargetBox-0="[209,1387,193,1924]" captionTargetBox-1="[205,1382,193,1924]" captionTargetBox-2="[171,1415,193,1545]" captionTargetId-0="figure@3.[209,1387,193,1924]" captionTargetId-1="figure@4.[205,1382,193,1924]" captionTargetId-2="figure@5.[171,1415,193,1545]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionText-0="FIGURE 1. Pycnomerus agtsteinicus sp. nov., holotype 273-2 [CCHH]: A, B—dorsal habitus photomicrograph, and corresponding SR X-ray micro-CT rendering; C, D—ventral habitus photomicrograph, and corresponding SR X-ray micro-CT rendering. Scale bars represent 0.5 mm." captionText-1="FIGURE 2. Pycnomerus agtsteinicus sp. nov., holotype 273-2 [CCHH]: A, B—lateral left habitus photomicrograph, and corresponding SR X-ray micro-CT rendering; C, D—lateral right habitus photomicrograph, and corresponding SR X-ray micro-CT rendering. Scale bars represent 1 mm." captionText-2="FIGURE 3. Pycnomerus agtsteinicus sp. nov., holotype 273-2 [CCHH]: A, B—frontal habitus photomicrograph, and corresponding SR X-ray micro-CT rendering; C—caudal habitus SR X-ray micro-CT rendering; D—left antenna, ventral view; E—left antenna, dorsal view; F—left protarsus, ventral view; G—left middle leg, ventral view. Scale bars represent 0.5 mm (A–C), 0.25 mm (D–G)." httpUri-0="https://zenodo.org/record/2625677/files/figure.png" httpUri-1="https://zenodo.org/record/2625679/files/figure.png" httpUri-2="https://zenodo.org/record/2625681/files/figure.png" pageId="1" pageNumber="566">Figs 1–3</figureCitation>
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)
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</paragraph>
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<paragraph id="8BC2369FFFB10E70FF4E90F4FC7AF823" blockId="1.[151,911,1942,2036]" box="[151,911,2014,2036]" pageId="1" pageNumber="566">
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<uri id="FFEC3A9DFFB10E70FF4E90F4FC7AF823" box="[151,911,2014,2036]" pageId="1" pageNumber="566">
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urn:lsid:zoobank.org:act:
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<uuid id="FFDB0C4AFFB10E70FE5790F4FC7AF823" box="[398,911,2014,2036]" pageId="1" pageNumber="566">5BDD33E3-D9FB-492E-B87E-525ADA5D7CFF</uuid>
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</uri>
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||
</paragraph>
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||
</subSubSection>
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<subSubSection id="C3676514FFB20E73FF4E97BDFAA3FEB2" pageId="2" pageNumber="567" type="materials_examined">
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<paragraph id="8BC2369FFFB20E73FF4E97BDFD14FECA" blockId="2.[151,1437,151,2013]" pageId="2" pageNumber="567">
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<emphasis id="B909EA8DFFB20E73FF4E97BDFEBEFF67" bold="true" box="[151,331,151,176]" pageId="2" pageNumber="567">Type material.</emphasis>
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<materialsCitation id="3B153CC2FFB20E73FE8097BDFD14FECA" ID-GBIF-Occurrence="2234218678" collectionCode="CCHH" pageId="2" pageNumber="567" specimenCount="1" typeStatus="holotype">
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<typeStatus id="54C6883DFFB20E73FE8097BDFE33FF67" box="[345,454,151,176]" pageId="2" pageNumber="567" type="holotype">Holotype</typeStatus>
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: collection number 273-2 [
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<collectionCode id="ED6CAE5AFFB20E73FCD297B2FC94FF67" box="[779,865,152,176]" pageId="2" pageNumber="567">CCHH</collectionCode>
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], adult, female (submentum without setose pit). Complete beetle with partially exposed hind wings, included in small, transparent yellow amber piece embedded in block of GTS-polyester resin with dimensions 13×10×
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<quantity id="4C859B7AFFB20E73FCCB97F5FC84FF20" box="[786,881,223,248]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="2" pageNumber="567" unit="mm" value="4.5">4.5 mm</quantity>
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. Syninclusions consist of few stellate
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<taxonomicName id="4C7D4D1CFFB20E73FAED97CAFA69FF2F" authorityName="" authorityYear="1829" box="[1332,1436,224,248]" class="Magnoliopsida" family="Fagaceae" higherTaxonomySource="GBIF" kingdom="Plantae" order="Fagales" pageId="2" pageNumber="567" phylum="Tracheophyta" rank="family">Fagaceae</taxonomicName>
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trichomes, and few small pieces of organic material.
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</materialsCitation>
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</paragraph>
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<paragraph id="8BC2369FFFB20E73FF1E960DFCBEFEE8" blockId="2.[151,1437,151,2013]" box="[199,843,295,320]" pageId="2" pageNumber="567">
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<materialsCitation id="3B153CC2FFB20E73FF1E960DFCB2FEE8" ID-GBIF-Occurrence="2234218689" box="[199,839,295,320]" location="Baltic" municipality="Upper Eocene" pageId="2" pageNumber="567" specimenCount="1" typeStatus="holotype">
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<emphasis id="B909EA8DFFB20E73FF1E960DFEA0FE97" bold="true" box="[199,341,295,320]" pageId="2" pageNumber="567">
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<typeStatus id="54C6883DFFB20E73FF1E960DFEF7FE97" box="[199,258,295,320]" pageId="2" pageNumber="567">Type</typeStatus>
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strata.
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</emphasis>
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<location id="8EA26044FFB20E73FE85960DFE6AFE97" LSID="urn:lsid:plazi:treatment:03D48789FFB10E77FF4E90BDFA69FCC2:8EA26044FFB20E73FE85960DFE6AFE97" box="[348,415,295,320]" municipality="Upper Eocene" name="Baltic" pageId="2" pageNumber="567">Baltic</location>
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amber, mid-Eocene to
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<collectingMunicipality id="6BA6ACE5FFB20E73FD7D9602FCB2FEE8" box="[676,839,296,319]" pageId="2" pageNumber="567">Upper Eocene</collectingMunicipality>
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</materialsCitation>
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.
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</paragraph>
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<paragraph id="8BC2369FFFB20E73FF1E9666FAA3FEB2" blockId="2.[151,1437,151,2013]" box="[199,1366,332,357]" pageId="2" pageNumber="567">
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<materialsCitation id="3B153CC2FFB20E73FF1E9666FAA4FEB2" ID-GBIF-Occurrence="2234218655" box="[199,1361,332,357]" country="Russia" location="Baltic Sea" municipality="Yantarny" pageId="2" pageNumber="567" specimenCount="1" stateProvince="Kaliningrad" typeStatus="holotype">
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<emphasis id="B909EA8DFFB20E73FF1E9666FE90FEB2" bold="true" box="[199,357,332,357]" pageId="2" pageNumber="567">
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<typeStatus id="54C6883DFFB20E73FF1E9666FEF7FEB2" box="[199,258,332,357]" pageId="2" pageNumber="567">Type</typeStatus>
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locality.
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</emphasis>
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<location id="8EA26044FFB20E73FEB59666FE2AFEB3" LSID="urn:lsid:plazi:treatment:03D48789FFB10E77FF4E90BDFA69FCC2:8EA26044FFB20E73FEB59666FE2AFEB3" box="[364,479,332,357]" country="Russia" municipality="Yantarny" name="Baltic Sea" pageId="2" pageNumber="567" stateProvince="Kaliningrad">Baltic Sea</location>
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coast,
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<collectingMunicipality id="6BA6ACE5FFB20E73FDF49667FD67FEB2" box="[557,658,333,357]" pageId="2" pageNumber="567">Yantarny</collectingMunicipality>
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settlement (formerly Palmnicken),
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<collectingRegion id="49B9F87DFFB20E73FBF89666FB52FEB2" box="[1057,1191,332,357]" country="Russia" name="Kaliningrad" pageId="2" pageNumber="567">Kaliningrad</collectingRegion>
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region,
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<collectingCountry id="F36A760FFFB20E73FADA9666FAA4FEB2" box="[1283,1361,332,357]" name="Russia" pageId="2" pageNumber="567">Russia</collectingCountry>
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</materialsCitation>
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.
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C3676514FFB20E73FF1E9645FE61FE7A" pageId="2" pageNumber="567" type="etymology">
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<paragraph id="8BC2369FFFB20E73FF1E9645FE61FE7A" blockId="2.[151,1437,151,2013]" pageId="2" pageNumber="567">
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<emphasis id="B909EA8DFFB20E73FF1E9645FEBBFE5F" bold="true" box="[199,334,367,392]" pageId="2" pageNumber="567">Etymology.</emphasis>
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The specific epithet refers to the amber source and is derived from the Upper German word for amber, “der Agtstein”.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3676514FFB20E73FF1E969DFC32FBC7" pageId="2" pageNumber="567" type="diagnosis">
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||
<paragraph id="8BC2369FFFB20E73FF1E969DFAE7FD1B" blockId="2.[151,1437,151,2013]" pageId="2" pageNumber="567">
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||
<emphasis id="B909EA8DFFB20E73FF1E969DFE25FE07" bold="true" box="[199,464,439,464]" pageId="2" pageNumber="567">Differential diagnosis.</emphasis>
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<taxonomicName id="4C7D4D1CFFB20E73FE019693FD07FE18" authority="Bukejs & Alekseev & Cooper & King & Mckellar, 2019" authorityName="Bukejs & Alekseev & Cooper & King & Mckellar" authorityYear="2019" box="[472,754,440,464]" class="Insecta" family="Zopheridae" genus="Pycnomerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="567" phylum="Arthropoda" rank="species" species="agtsteinicus" status="sp. nov.">
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<emphasis id="B909EA8DFFB20E73FE019693FD07FE18" box="[472,754,440,464]" italics="true" pageId="2" pageNumber="567">Pycnomerus agtsteinicus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A23A57F6FFB20E73FD22969DFCA3FE18" box="[763,854,439,464]" pageId="2" pageNumber="567" rank="species">
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||
<emphasis id="B909EA8DFFB20E73FD22969DFCE0FE07" bold="true" box="[763,789,439,464]" pageId="2" pageNumber="567">sp</emphasis>
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||
.
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<emphasis id="B909EA8DFFB20E73FCFC969DFCBAFE07" bold="true" box="[805,847,439,464]" pageId="2" pageNumber="567">nov</emphasis>
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.
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</taxonomicNameLabel>
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differs from
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<taxonomicName id="4C7D4D1CFFB20E73FC299693FB16FE07" authority="Alekseev" authorityName="Alekseev" authorityYear="2015" box="[1008,1251,439,464]" class="Insecta" family="Zopheridae" genus="Pycnomerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="567" phylum="Arthropoda" rank="species" species="simukovi">
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||
<emphasis id="B909EA8DFFB20E73FC299693FB84FE07" box="[1008,1137,440,464]" italics="true" pageId="2" pageNumber="567">P. simukovi</emphasis>
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Alekseev
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</taxonomicName>
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, described from Baltic amber, in the following morphological characters: (1) anterior portion of head with oblong pits (rounded pits in
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<taxonomicName id="4C7D4D1CFFB20E73FF6E952BFECDFDCF" authorityName="Alekseev" authorityYear="2015" box="[183,312,512,536]" class="Insecta" family="Zopheridae" genus="Pycnomerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="567" phylum="Arthropoda" rank="species" species="simukovi">
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<emphasis id="B909EA8DFFB20E73FF6E952BFECDFDCF" box="[183,312,512,536]" italics="true" pageId="2" pageNumber="567">P. simukovi</emphasis>
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</taxonomicName>
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); (2) pronotum more transverse, 1.2× as wide as long (0.9× as wide as long in
|
||
<taxonomicName id="4C7D4D1CFFB20E73FB07952BFA95FDCF" authorityName="Alekseev" authorityYear="2015" box="[1246,1376,512,536]" class="Insecta" family="Zopheridae" genus="Pycnomerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="567" phylum="Arthropoda" rank="species" species="simukovi">
|
||
<emphasis id="B909EA8DFFB20E73FB07952BFA95FDCF" box="[1246,1376,512,536]" italics="true" pageId="2" pageNumber="567">P. simukovi</emphasis>
|
||
</taxonomicName>
|
||
); (3) antennomere 11 transverse, rounded apically, and almost as wide as antennomere 10 (oblong, tapered, and distinctly narrower than antennomere
|
||
<quantity id="4C859B7AFFB20E73FD9A9562FD89FDB7" box="[579,636,583,608]" metricMagnitude="-1" metricUnit="m" metricValue="2.54" pageId="2" pageNumber="567" unit="in" value="10.0">10 in</quantity>
|
||
<taxonomicName id="4C7D4D1CFFB20E73FD5A9563FCF6FDB7" authorityName="Alekseev" authorityYear="2015" box="[643,771,584,608]" class="Insecta" family="Zopheridae" genus="Pycnomerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="567" phylum="Arthropoda" rank="species" species="simukovi">
|
||
<emphasis id="B909EA8DFFB20E73FD5A9563FCF6FDB7" box="[643,771,584,608]" italics="true" pageId="2" pageNumber="567">P. simukovi</emphasis>
|
||
</taxonomicName>
|
||
); (4) scutellar shield more transverse, 2.6× as wide as long (1.5× as wide as long in
|
||
<taxonomicName id="4C7D4D1CFFB20E73FE689547FDD8FD53" authorityName="Alekseev" authorityYear="2015" box="[433,557,620,644]" class="Insecta" family="Zopheridae" genus="Pycnomerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="567" phylum="Arthropoda" rank="species" species="simukovi">
|
||
<emphasis id="B909EA8DFFB20E73FE689547FDD8FD53" box="[433,557,620,644]" italics="true" pageId="2" pageNumber="567">P. simukovi</emphasis>
|
||
</taxonomicName>
|
||
); (5) sparser punctation of head and pronotum; (6) pronotum behind anterior angles more widely rounded (pronotal sides more parallel and almost straight within anterior one-third of length in
|
||
<taxonomicName id="4C7D4D1CFFB20E73FF4E959FFEE3FD1B" authorityName="Alekseev" authorityYear="2015" box="[151,278,692,716]" class="Insecta" family="Zopheridae" genus="Pycnomerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="567" phylum="Arthropoda" rank="species" species="simukovi">
|
||
<emphasis id="B909EA8DFFB20E73FF4E959FFEE3FD1B" box="[151,278,692,716]" italics="true" pageId="2" pageNumber="567">P. simukovi</emphasis>
|
||
</taxonomicName>
|
||
); and (7) ventrite 5 longer than ventrite 4 (ventrite 5 shorter than ventrite
|
||
<quantity id="4C859B7AFFB20E73FB88959EFB89FD1A" box="[1105,1148,692,717]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="2" pageNumber="567" unit="in" value="4.0">4 in</quantity>
|
||
<taxonomicName id="4C7D4D1CFFB20E73FB5A959FFAF7FD1B" authorityName="Alekseev" authorityYear="2015" box="[1155,1282,692,716]" class="Insecta" family="Zopheridae" genus="Pycnomerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="567" phylum="Arthropoda" rank="species" species="simukovi">
|
||
<emphasis id="B909EA8DFFB20E73FB5A959FFAF7FD1B" box="[1155,1282,692,716]" italics="true" pageId="2" pageNumber="567">P. simukovi</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BC2369FFFB20E73FF1E95FDFABAFCEF" blockId="2.[151,1437,151,2013]" pageId="2" pageNumber="567">
|
||
Differences between
|
||
<taxonomicName id="4C7D4D1CFFB20E73FE6E95F3FDACFD38" authority="Bukejs & Alekseev & Cooper & King & Mckellar, 2019" authorityName="Bukejs & Alekseev & Cooper & King & Mckellar" authorityYear="2019" box="[439,601,728,752]" class="Insecta" family="Zopheridae" genus="Pycnomerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="567" phylum="Arthropoda" rank="species" species="agtsteinicus" status="sp. nov.">
|
||
<emphasis id="B909EA8DFFB20E73FE6E95F3FDACFD38" box="[439,601,728,752]" italics="true" pageId="2" pageNumber="567">P. agtsteinicus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23A57F6FFB20E73FDBA95FDFD48FD38" box="[611,701,727,752]" pageId="2" pageNumber="567" rank="species">
|
||
<emphasis id="B909EA8DFFB20E73FDBA95FDFD88FD27" bold="true" box="[611,637,727,752]" pageId="2" pageNumber="567">sp</emphasis>
|
||
.
|
||
<emphasis id="B909EA8DFFB20E73FD5595FDFD43FD27" bold="true" box="[652,694,727,752]" pageId="2" pageNumber="567">nov</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
and subfossils of extinct
|
||
<taxonomicName id="4C7D4D1CFFB20E73FC3A95F3FB91FD38" authorityName="Porch & Smith" authorityYear="2017" box="[995,1124,728,752]" class="Insecta" family="Zopheridae" genus="Pycnomerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="567" phylum="Arthropoda" rank="species" species="rimatara">
|
||
<emphasis id="B909EA8DFFB20E73FC3A95F3FB91FD38" box="[995,1124,728,752]" italics="true" pageId="2" pageNumber="567">P. rimatara</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C7D4D1CFFB20E73FB4495F3FAE3FD27" authorityName="Porch & Smith" authorityYear="2017" box="[1181,1302,728,752]" class="Insecta" family="Zopheridae" genus="Pycnomerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="567" phylum="Arthropoda" rank="species" species="prebblei">
|
||
<emphasis id="B909EA8DFFB20E73FB4495F3FAE3FD27" box="[1181,1302,728,752]" italics="true" pageId="2" pageNumber="567">P. prebblei</emphasis>
|
||
</taxonomicName>
|
||
include the following combination of characters: (1) anterior portion of head with oblong pits; (2) pronotum more transverse; (3) flattened pronotal disc; (4) pronotal lateral margins without crenulation; and (5) habitus distinctly wider.
|
||
</paragraph>
|
||
<paragraph id="8BC2369FFFB20E73FF1E946EFC32FBC7" blockId="2.[151,1437,151,2013]" pageId="2" pageNumber="567">
|
||
The newly described fossil species can be easily distinguished from the modern European natives
|
||
<taxonomicName id="4C7D4D1CFFB20E73FAC8946FFF1FFC57" baseAuthorityName="Ganglbauer" baseAuthorityYear="1899" class="Insecta" family="Zopheridae" genus="Pycnomerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="567" phylum="Arthropoda" rank="species" species="italicus">
|
||
<emphasis id="B909EA8DFFB20E73FAC8946FFF1FFC57" italics="true" pageId="2" pageNumber="567">Pycnomerus italicus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C7D4D1CFFB20E73FEF89443FE5AFC57" baseAuthorityName="Germar" baseAuthorityYear="1824" box="[289,431,872,896]" class="Insecta" family="Zopheridae" genus="Pycnomerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="567" phylum="Arthropoda" rank="species" species="sulcicollis">
|
||
<emphasis id="B909EA8DFFB20E73FEF89443FE5AFC57" box="[289,431,872,896]" italics="true" pageId="2" pageNumber="567">P. sulcicollis</emphasis>
|
||
</taxonomicName>
|
||
based on its non-reduced, 11-segmented antennae; and lack of deep, paired, longitudinal impressions on pronotal disc. It can be distinguished from
|
||
<taxonomicName id="4C7D4D1CFFB20E73FC9594A7FC23FC73" baseAuthorityName="Olivier" baseAuthorityYear="1790" box="[844,982,908,932]" class="Insecta" family="Zopheridae" genus="Pycnomerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="567" phylum="Arthropoda" rank="species" species="terebrans">
|
||
<emphasis id="B909EA8DFFB20E73FC9594A7FC23FC73" box="[844,982,908,932]" italics="true" pageId="2" pageNumber="567">P. terebrans</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C7D4D1CFFB20E73FBCD94A7FB5FFC73" authorityName="Jaquelin du Val" authorityYear="1859" box="[1044,1194,909,932]" class="Insecta" family="Zopheridae" genus="Pycnomerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="567" phylum="Arthropoda" rank="species" species="inexpectus">
|
||
<emphasis id="B909EA8DFFB20E73FBCD94A7FB5FFC73" box="[1044,1194,909,932]" italics="true" pageId="2" pageNumber="567">P. inexpectus</emphasis>
|
||
</taxonomicName>
|
||
based upon reduced anterior pronotal angles, distinctly 2-segmented antennal club, and elytral apices that are not flattened. Additionally, the new species differs from
|
||
<taxonomicName id="4C7D4D1CFFB20E73FDA594FFFCE5FC3B" authorityName="Jaquelin du Val" authorityYear="1859" box="[636,784,981,1004]" class="Insecta" family="Zopheridae" genus="Pycnomerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="567" phylum="Arthropoda" rank="species" species="inexpectus">
|
||
<emphasis id="B909EA8DFFB20E73FDA594FFFCE5FC3B" box="[636,784,981,1004]" italics="true" pageId="2" pageNumber="567">P. inexpectus</emphasis>
|
||
</taxonomicName>
|
||
in possessing pronotal disc without shallow impressions, and from
|
||
<taxonomicName id="4C7D4D1CFFB20E73FEDD94D3FE7CFBC7" baseAuthorityName="Olivier" baseAuthorityYear="1790" box="[260,393,1016,1040]" class="Insecta" family="Zopheridae" genus="Pycnomerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="567" phylum="Arthropoda" rank="species" species="terebrans">
|
||
<emphasis id="B909EA8DFFB20E73FEDD94D3FE7CFBC7" box="[260,393,1016,1040]" italics="true" pageId="2" pageNumber="567">P. terebrans</emphasis>
|
||
</taxonomicName>
|
||
in possessing pronotal sides that are not explanate.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3676514FFB20E77FF1E9336FD8DFF02" lastPageId="6" lastPageNumber="571" pageId="2" pageNumber="567" type="description">
|
||
<paragraph id="8BC2369FFFB20E73FF1E9336FBF5FB8F" blockId="2.[151,1437,151,2013]" pageId="2" pageNumber="567">
|
||
<emphasis id="B909EA8DFFB20E73FF1E9336FEACFBE2" bold="true" box="[199,345,1052,1077]" pageId="2" pageNumber="567">Description.</emphasis>
|
||
Body length
|
||
<quantity id="4C859B7AFFB20E73FE259336FDADFBE3" box="[508,600,1052,1076]" metricMagnitude="-3" metricUnit="m" metricValue="3.1" pageId="2" pageNumber="567" unit="mm" value="3.1">3.1 mm</quantity>
|
||
, maximum width 1.0 mm; body shape elongated, parallel-sided, dorsoventrally flattened; body shiny, unicolor, ferruginous (as preserved), glabrous.
|
||
</paragraph>
|
||
<paragraph id="8BC2369FFFB20E73FF1E934EFEA4F9FB" blockId="2.[151,1437,151,2013]" pageId="2" pageNumber="567">Head prognathous, retracted into prothorax up to posterior margins of eyes, nearly as long as wide, with indistinct subantennal grooves; covered with small and dense punctation; distance between punctures about 1.0– 3.0× diameter of one puncture, punctures slightly smaller and denser in anterior portion of head, and interspaces glabrous; anterior portion of head with pair of oblong pits dorsally. Clypeus with widely and shallowly concave anterior margin. Compound eyes large, oval, slightly convex, entire, apparently without interfacetal setae, with distinct facets; each facet about as large as one head puncture (in posterior portion of head). Distance between compound eyes dorsally nearly equal to 5× transverse diameter of one eye. Mandibles bidentate apically. Submentum without setose pit. Antennae with 11 antennomeres, clavate, with distinct 2-segmented club, stout, short, reaching middle of pronotum, covered with fine, sparse, erect to semierect setation; antennomeres transverse; scape about 1.1× as wide and 0.8× as long as pedicel; antennomeres 3–9 subequal in size and shape; antennomere 9 about 2.7× as wide as long; antennomere 10 about 2× as wide as long, and 1.6× as wide as antennomere 9; antennomere 11 about 1.2× as wide as long, with widely rounded apex, and slightly narrower than antennomere 10.</paragraph>
|
||
<paragraph id="8BC2369FFFB20E73FF1E9113FBAAF937" blockId="2.[151,1437,151,2013]" pageId="2" pageNumber="567">Pronotum transverse, 1.2× as wide as long, widest in anterior one-fourth of pronotal length; pronotal disc flat; covered with small, sparse punctures. Lateral margins of pronotum apparently smooth, almost straight, slightly converging posterior to widest point, becoming rounded posterolaterally, with narrow bordering; anterior margin or pronotum slightly concave, apparently without bordering, unmodified; posterior margin convex, with narrow bordering. Anterior angles acute, slightly protruding; posterior angles widely rounded.</paragraph>
|
||
<paragraph id="8BC2369FFFB20E73FF1E91C6FBDFF8D2" blockId="2.[151,1437,151,2013]" box="[199,1066,1772,1797]" pageId="2" pageNumber="567">Scutellar shield small, elongated oval, transverse, about 2.6× as wide as long.</paragraph>
|
||
<paragraph id="8BC2369FFFB20E73FF1E9025FBC9F8A7" blockId="2.[151,1437,151,2013]" pageId="2" pageNumber="567">Elytra elongated oval, parallel-sided, flattened, about 2.1× as long as wide, with distinct humeri. Each elytron with 10 deep striae; striae distinct throughout entire length of elytron; interstrial intervals distinctly wider than stria, and convex, with sparse, minute punctures; scutellary striole absent. Macropterous.</paragraph>
|
||
<paragraph id="8BC2369FFFB20E73FF1E9057FD1AF80A" blockId="2.[151,1437,151,2013]" pageId="2" pageNumber="567">Pro- and metaventrite with large, sparse, rounded punctures (similar to punctures of ventrite 1, and larger than pronotal punctation); hypomeron without antennal cavities; prosternal intercoxal process wide, distinctly wider than diameter of procoxa, expanded in apical portion.</paragraph>
|
||
<caption id="DF026617FFB30E72FF4E90B2FDDFF83C" httpUri="https://zenodo.org/record/2625677/files/figure.png" pageId="3" pageNumber="568" startId="3.[151,250,1944,1966]" targetBox="[209,1387,193,1924]" targetPageId="3">
|
||
<paragraph id="8BC2369FFFB30E72FF4E90B2FDDFF83C" blockId="3.[151,1436,1944,2027]" pageId="3" pageNumber="568">
|
||
<emphasis id="B909EA8DFFB30E72FF4E90B2FEEFF87A" bold="true" box="[151,282,1944,1966]" pageId="3" pageNumber="568">FIGURE 1.</emphasis>
|
||
<taxonomicName id="4C7D4D1CFFB30E72FEF190B3FDD9F879" authority="Bukejs & Alekseev & Cooper & King & Mckellar, 2019" authorityName="Bukejs & Alekseev & Cooper & King & Mckellar" authorityYear="2019" box="[296,556,1945,1966]" class="Insecta" family="Zopheridae" genus="Pycnomerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="568" phylum="Arthropoda" rank="species" species="agtsteinicus" status="sp. nov.">
|
||
<emphasis id="B909EA8DFFB30E72FEF190B3FDD9F879" box="[296,556,1945,1966]" italics="true" pageId="3" pageNumber="568">Pycnomerus agtsteinicus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B909EA8DFFB30E72FDE390B2FD64F87A" bold="true" box="[570,657,1944,1965]" pageId="3" pageNumber="568">
|
||
<taxonomicNameLabel id="A23A57F6FFB30E72FDE390B2FD64F87A" box="[570,657,1944,1965]" pageId="3" pageNumber="568" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, holotype 273-2 [CCHH]: A, B—dorsal habitus photomicrograph, and corresponding SR X-ray micro-CT rendering; C, D—ventral habitus photomicrograph, and corresponding SR X-ray micro-CT rendering. Scale bars represent 0.5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF026617FFB40E75FF4E90B0FD74F83A" httpUri="https://zenodo.org/record/2625679/files/figure.png" pageId="4" pageNumber="569" startId="4.[151,250,1946,1968]" targetBox="[205,1382,193,1924]" targetPageId="4">
|
||
<paragraph id="8BC2369FFFB40E75FF4E90B0FD74F83A" blockId="4.[151,1436,1946,2029]" pageId="4" pageNumber="569">
|
||
<emphasis id="B909EA8DFFB40E75FF4E90B0FEE2F878" bold="true" box="[151,279,1946,1968]" pageId="4" pageNumber="569">FIGURE 2.</emphasis>
|
||
<taxonomicName id="4C7D4D1CFFB40E75FEF890B1FDD4F867" authority="Bukejs & Alekseev & Cooper & King & Mckellar, 2019" authorityName="Bukejs & Alekseev & Cooper & King & Mckellar" authorityYear="2019" box="[289,545,1947,1968]" class="Insecta" family="Zopheridae" genus="Pycnomerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="569" phylum="Arthropoda" rank="species" species="agtsteinicus" status="sp. nov.">
|
||
<emphasis id="B909EA8DFFB40E75FEF890B1FDD4F867" box="[289,545,1947,1968]" italics="true" pageId="4" pageNumber="569">Pycnomerus agtsteinicus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B909EA8DFFB40E75FDF590B0FD75F878" bold="true" box="[556,640,1946,1967]" pageId="4" pageNumber="569">
|
||
<taxonomicNameLabel id="A23A57F6FFB40E75FDF590B0FD75F878" box="[556,640,1946,1967]" pageId="4" pageNumber="569" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, holotype 273-2 [CCHH]: A, B—lateral left habitus photomicrograph, and corresponding SR X-ray micro-CT rendering; C, D—lateral right habitus photomicrograph, and corresponding SR X-ray micro-CT rendering. Scale bars represent 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF026617FFB50E74FF4E9133FE7BF95C" httpUri="https://zenodo.org/record/2625681/files/figure.png" pageId="5" pageNumber="570" startId="5.[151,250,1561,1583]" targetBox="[171,1415,193,1545]" targetPageId="5">
|
||
<paragraph id="8BC2369FFFB50E74FF4E9133FE7BF95C" blockId="5.[151,1436,1561,1675]" pageId="5" pageNumber="570">
|
||
<emphasis id="B909EA8DFFB50E74FF4E9133FEECF9F9" bold="true" box="[151,281,1561,1583]" pageId="5" pageNumber="570">FIGURE 3.</emphasis>
|
||
<taxonomicName id="4C7D4D1CFFB50E74FEFE9130FDDEF9F8" authority="Bukejs & Alekseev & Cooper & King & Mckellar, 2019" authorityName="Bukejs & Alekseev & Cooper & King & Mckellar" authorityYear="2019" box="[295,555,1562,1583]" class="Insecta" family="Zopheridae" genus="Pycnomerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="570" phylum="Arthropoda" rank="species" species="agtsteinicus" status="sp. nov.">
|
||
<emphasis id="B909EA8DFFB50E74FEFE9130FDDEF9F8" box="[295,555,1562,1583]" italics="true" pageId="5" pageNumber="570">Pycnomerus agtsteinicus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B909EA8DFFB50E74FDE19133FD7AF9F9" bold="true" box="[568,655,1561,1582]" pageId="5" pageNumber="570">
|
||
<taxonomicNameLabel id="A23A57F6FFB50E74FDE19133FD7AF9F9" box="[568,655,1561,1582]" pageId="5" pageNumber="570" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, holotype 273-2 [CCHH]: A, B—frontal habitus photomicrograph, and corresponding SR X-ray micro-CT rendering; C—caudal habitus SR X-ray micro-CT rendering; D—left antenna, ventral view; E—left antenna, dorsal view; F—left protarsus, ventral view; G—left middle leg, ventral view. Scale bars represent 0.5 mm (A–C), 0.25 mm (D–G).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BC2369FFFB50E74FF1E9193FB91F8B6" blockId="5.[151,1437,1720,2033]" pageId="5" pageNumber="570">Legs short, robust, with sparse and fine punctation. Procoxal cavities externally closed. Procoxae globose; meso- and metacoxae oval, transverse; all coxae widely separated by more than one transverse diameter of coxa. Femora clavate; tibiae dilated apically, almost straight, slightly longer than femora, with two apical spines at inner margin. Tarsi nearly as long as tibia, without dense setae ventrally, with few semierect setae; tarsal formula 4-4-4. Relative length ratios of metatarsomeres 1–4 equal to 5:3:3:7. Claws thin, free, simple.</paragraph>
|
||
<paragraph id="8BC2369FFFB50E77FF1E9046FD8DFF02" blockId="5.[151,1437,1720,2033]" lastBlockId="6.[151,1437,151,789]" lastPageId="6" lastPageNumber="571" pageId="5" pageNumber="570">Abdomen with five visible ventrites; abdominal sutures distinct throughout length; ventrites 1–3 more solidly fused together than ventrites 4–5; ventrites 1–4 with large, sparse punctures, and distance between punctures about 0.5–3.0× diameter of one puncture; ventrite 5 without preapical groove, covered with large and distinctly denser punctation (distance between punctures less than diameter of one puncture), and with small punctures at posterior margin. Intermetacoxal apophysis (intercoxal process of ventrite 1) widely rounded. Relative length ratios of ventrites 1–5 equal to 9:7:7:5:6 (medially).</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3676514FFB60E77FF1E97F5FA69FCC2" pageId="6" pageNumber="571" type="discussion">
|
||
<paragraph id="8BC2369FFFB60E77FF1E97F5FA69FCC2" blockId="6.[151,1437,151,789]" pageId="6" pageNumber="571">
|
||
<emphasis id="B909EA8DFFB60E77FF1E97F5FEC1FF2F" bold="true" box="[199,308,223,248]" pageId="6" pageNumber="571">Remarks</emphasis>
|
||
. Several morphological characters mentioned in the comparison of the newly described species with
|
||
<taxonomicName id="4C7D4D1CFFB60E77FF4E962FFEE2FECB" authorityName="Alekseev" authorityYear="2015" box="[151,279,260,284]" class="Insecta" family="Zopheridae" genus="Pycnomerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="571" phylum="Arthropoda" rank="species" species="simukovi">
|
||
<emphasis id="B909EA8DFFB60E77FF4E962FFEE2FECB" box="[151,279,260,284]" italics="true" pageId="6" pageNumber="571">P. simukovi</emphasis>
|
||
</taxonomicName>
|
||
, can vary sexually or individually in some well-studied extant species of
|
||
<taxonomicName id="4C7D4D1CFFB60E77FB8E962FFB14FECB" box="[1111,1249,261,284]" class="Insecta" family="Zopheridae" genus="Pycnomerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="571" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B909EA8DFFB60E77FB8E962FFB14FECB" box="[1111,1249,261,284]" italics="true" pageId="6" pageNumber="571">Pycnomerus</emphasis>
|
||
</taxonomicName>
|
||
. In addition, the original description of
|
||
<taxonomicName id="4C7D4D1CFFB60E77FE4F9603FDE0FE97" authorityName="Alekseev" authorityYear="2015" box="[406,533,296,320]" class="Insecta" family="Zopheridae" genus="Pycnomerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="571" phylum="Arthropoda" rank="species" species="simukovi">
|
||
<emphasis id="B909EA8DFFB60E77FE4F9603FDE0FE97" box="[406,533,296,320]" italics="true" pageId="6" pageNumber="571">P. simukovi</emphasis>
|
||
</taxonomicName>
|
||
from Baltic amber was based on a specimen that is possibly a little distorted and of unknown sex (the area of the submentum was not discernible, and the presence or absence of a setose pit there is unclear). Consequently, the description of
|
||
<taxonomicName id="4C7D4D1CFFB60E77FDA2965BFD09FE5F" authorityName="Alekseev" authorityYear="2015" box="[635,764,368,392]" class="Insecta" family="Zopheridae" genus="Pycnomerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="571" phylum="Arthropoda" rank="species" species="simukovi">
|
||
<emphasis id="B909EA8DFFB60E77FDA2965BFD09FE5F" box="[635,764,368,392]" italics="true" pageId="6" pageNumber="571">P. simukovi</emphasis>
|
||
</taxonomicName>
|
||
was provided with heavily schematized drawings. Both of the specimens under consideration (i.e., the specimen described here as
|
||
<taxonomicName id="4C7D4D1CFFB60E77FC1D96BFFB92FE7B" authority="Bukejs & Alekseev & Cooper & King & Mckellar, 2019" authorityName="Bukejs & Alekseev & Cooper & King & Mckellar" authorityYear="2019" box="[964,1127,405,428]" class="Insecta" family="Zopheridae" genus="Pycnomerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="571" phylum="Arthropoda" rank="species" species="agtsteinicus" status="sp. nov.">
|
||
<emphasis id="B909EA8DFFB60E77FC1D96BFFB92FE7B" box="[964,1127,405,428]" italics="true" pageId="6" pageNumber="571">P. agtsteinicus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23A57F6FFB60E77FBB696BEFB3DFE7B" box="[1135,1224,404,429]" pageId="6" pageNumber="571" rank="species">
|
||
<emphasis id="B909EA8DFFB60E77FBB696BEFB7CFE7A" bold="true" box="[1135,1161,404,429]" pageId="6" pageNumber="571">sp</emphasis>
|
||
.
|
||
<emphasis id="B909EA8DFFB60E77FB4196BEFB37FE7A" bold="true" box="[1176,1218,404,429]" pageId="6" pageNumber="571">nov</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
, and the specimen previously described as
|
||
<taxonomicName id="4C7D4D1CFFB60E77FE739693FDD2FE07" authorityName="Alekseev" authorityYear="2015" box="[426,551,440,464]" class="Insecta" family="Zopheridae" genus="Pycnomerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="571" phylum="Arthropoda" rank="species" species="simukovi">
|
||
<emphasis id="B909EA8DFFB60E77FE739693FDD2FE07" box="[426,551,440,464]" italics="true" pageId="6" pageNumber="571">P. simukovi</emphasis>
|
||
</taxonomicName>
|
||
) possess a similar set of general morphological characters, and are without a doubt, morphologically close species that clearly differ from Recent European congeners. It is possible that the new species could be the female of the previously described Eocene species, and that the differences described herein could be interpreted as sexual characters or intraspecific variation. Despite the general morphological similarity, between the two fossil species we suggest that the new species is distinct, and that it can be discerned based upon the full combination of aforementioned characters. However, it is worth noting that assessing reliable specific differences in Eocene
|
||
<taxonomicName id="4C7D4D1CFFB60E77FE2C95BBFD75FD7F" box="[501,640,657,680]" class="Insecta" family="Zopheridae" genus="Pycnomerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="571" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B909EA8DFFB60E77FE2C95BBFD75FD7F" box="[501,640,657,680]" italics="true" pageId="6" pageNumber="571">Pycnomerus</emphasis>
|
||
</taxonomicName>
|
||
species would require further study of additional fossil material from this deposit. The detailed study of the Recent
|
||
<taxonomicName id="4C7D4D1CFFB60E77FD65959FFCB3FD1B" box="[700,838,693,716]" class="Insecta" family="Zopheridae" genus="Pycnomerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="571" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B909EA8DFFB60E77FD65959FFCB3FD1B" box="[700,838,693,716]" italics="true" pageId="6" pageNumber="571">Pycnomerus</emphasis>
|
||
</taxonomicName>
|
||
fauna from the tropics and temperate areas of the Southern Hemisphere, as well as the world-wide revision of the genus, will be necessary elements of future studies required for a more precise systematic placement of the extinct Eocene
|
||
<taxonomicName id="4C7D4D1CFFB60E77FC6895D7FBCEFCC3" box="[945,1083,765,788]" class="Insecta" family="Zopheridae" genus="Pycnomerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="571" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B909EA8DFFB60E77FC6895D7FBCEFCC3" box="[945,1083,765,788]" italics="true" pageId="6" pageNumber="571">Pycnomerus</emphasis>
|
||
</taxonomicName>
|
||
species within this large genus.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |