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<document id="7AC5C81CFB47C3BEE450E2315108103E" ID-CLB-Dataset="36776" ID-DOI="10.11646/zootaxa.3876.1.1" ID-GBIF-Dataset="c40a76a9-ec38-46e1-b9ff-869544e5f692" ID-ISSN="1175-5326" ID-Zenodo-Dep="287281" ID-ZooBank="071AFC59-C2EA-429A-8511-BAC5C4539C04" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460940313376" checkinUser="plazi" docAuthor="Smit, Harry &amp; Pešić, Vladimir" docDate="2014" docId="03D48784A927FFB8FF35120DFE45FE54" docLanguage="en" docName="zt03876p071.pdf" docOrigin="Zootaxa 3876 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Sperchonopsis orientalis Smit &amp; Pešić, 2014, n. sp." docType="treatment" docVersion="9" lastPageNumber="15" masterDocId="FFEDFFFCA92BFFB6FFA2166CFF84FF9B" masterDocTitle="Water mites from Mount Kinabalu and the Crocker Range, Borneo, Malaysia (Acari: Hydrachnidia), with the description of 34 new species" masterLastPageNumber="71" masterPageNumber="1" pageNumber="13" updateTime="1698584280452" updateUser="ExternalLinkService">
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<mods:title id="D8A9E4CE344C9A35D6E7CD8507617379">Water mites from Mount Kinabalu and the Crocker Range, Borneo, Malaysia (Acari: Hydrachnidia), with the description of 34 new species</mods:title>
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<mods:namePart id="20BE233CD1E8527A3C7F5769D613EFF5">Smit, Harry</mods:namePart>
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<mods:namePart id="2B2DB90F82928295BB985842680D22EE">Pešić, Vladimir</mods:namePart>
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<mods:date id="591D766424756F7D7808FC7BE65FBC9E">2014</mods:date>
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<treatment id="03D48784A927FFB8FF35120DFE45FE54" ID-DOI="http://doi.org/10.5281/zenodo.5695940" ID-GBIF-Taxon="127670514" ID-Zenodo-Dep="5695940" LSID="urn:lsid:plazi:treatment:03D48784A927FFB8FF35120DFE45FE54" httpUri="http://treatment.plazi.org/id/03D48784A927FFB8FF35120DFE45FE54" lastPageId="14" lastPageNumber="15" pageId="12" pageNumber="13">
<subSubSection id="C3676519A927FFBAFF35120DFE9AFB06" pageId="12" pageNumber="13" type="nomenclature">
<paragraph id="8BC23692A927FFBAFF35120DFD8DFBE0" blockId="12.[151,521,1121,1181]" box="[151,521,1121,1147]" pageId="12" pageNumber="13">
<heading id="D08A81FEA927FFBAFF35120DFD8DFBE0" bold="true" box="[151,521,1121,1147]" fontSize="11" level="1" pageId="12" pageNumber="13" reason="1">
<emphasis id="B909EA80A927FFBAFF35120DFD8DFBE0" bold="true" box="[151,521,1121,1147]" pageId="12" pageNumber="13">
<taxonomicName id="4C7D4D11A927FFBAFF35120DFE3BFBE0" box="[151,447,1121,1147]" class="Arachnida" family="Sperchonidae" genus="Sperchonopsis" kingdom="Animalia" order="Trombidiformes" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="orientalis" status="sp. nov.">
<emphasis id="B909EA80A927FFBAFF35120DFE3BFBE0" bold="true" box="[151,447,1121,1147]" italics="true" pageId="12" pageNumber="13">Sperchonopsis orientalis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23A57FBA927FFBAFE65120DFD8DFBE0" box="[455,521,1121,1147]" pageId="12" pageNumber="13" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BC23692A927FFBAFF3512E8FE9AFB06" blockId="12.[151,521,1121,1181]" box="[151,286,1156,1181]" pageId="12" pageNumber="13">
(
<figureCitation id="13462A17A927FFBAFF3D12E8FE93FB06" box="[159,279,1156,1181]" captionStart="FIGURE 6 A E" captionStartId="13.[151,250,1953,1975]" captionTargetBox="[176,1410,195,1904]" captionTargetId="figure@13.[176,1410,193,1932]" captionTargetPageId="13" captionText="FIGURE 6 A E. Sperchonopsis orientalis n. sp., female holotype, Mahua stream: A = idiosoma, dorsal view; C D = palp (P 1 missing); D = capitulum and chelicera. Scale bars = 100 Μm." httpUri="https://zenodo.org/record/287287/files/figure.png" pageId="12" pageNumber="13">Fig. 6AE</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3676519A927FFBAFF3512A0FCF6FACB" pageId="12" pageNumber="13" type="materials_examined">
<paragraph id="8BC23692A927FFBAFF3512A0FCF6FACB" blockId="12.[151,1437,1228,2009]" pageId="12" pageNumber="13">
<emphasis id="B909EA80A927FFBAFF3512A0FE9AFB7E" bold="true" box="[151,286,1228,1253]" pageId="12" pageNumber="13">
<typeStatus id="54C68830A927FFBAFF3512A0FF56FB7E" box="[151,210,1228,1253]" pageId="12" pageNumber="13">Type</typeStatus>
series
</emphasis>
.
<typeStatus id="54C68830A927FFBAFE8C12A0FE12FB7E" box="[302,406,1228,1253]" pageId="12" pageNumber="13" type="holotype">Holotype</typeStatus>
female, dissected and slide mounted, Mahua stream at crossing with main road,
<taxonomicName id="4C7D4D11A927FFBAFAE012A0FA19FB7E" box="[1346,1437,1228,1253]" class="Arachnida" family="Axonopsidae" genus="Albia" kingdom="Animalia" order="Trombidiformes" pageId="12" pageNumber="63" phylum="Arthropoda" rank="species" species="crocker" status="sp. nov.">Crocker</taxonomicName>
Range, Borneo,
<collectingCountry id="F36A7602A927FFBAFEED1283FE3DFA93" box="[335,441,1263,1288]" name="Malaysia" pageId="12" pageNumber="13">Malaysia</collectingCountry>
, 5º
<geoCoordinate id="EE495055A927FFBAFE781283FDCCFA9C" box="[474,584,1263,1288]" direction="north" orientation="latitude" pageId="12" pageNumber="13" precision="55" value="45.225">45.225 N</geoCoordinate>
, 116º
<geoCoordinate id="EE495055A927FFBAFD271283FD6AFA9C" box="[645,750,1263,1288]" direction="east" orientation="longitude" pageId="12" pageNumber="13" precision="55" value="26.085">26.085 E</geoCoordinate>
, alt.
<quantity id="4C859B77A927FFBAFC851283FCEBFA9C" box="[807,879,1263,1288]" metricMagnitude="2" metricUnit="m" metricValue="7.52" pageId="12" pageNumber="13" unit="m" value="752.0">752 m</quantity>
asl.,
<date id="FFC31052A927FFBAFC0E1283FBACFA93" box="[940,1064,1263,1288]" pageId="12" pageNumber="13" value="2012-09-22">22-ix-2012</date>
(
<collectionCode id="ED6CAE57A927FFBAFB9A129CFB17FA93" LSID="urn:lsid:biocol.org:col:34992" box="[1080,1171,1264,1288]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="12" pageNumber="13">RMNH</collectionCode>
).
<typeStatus id="54C68830A927FFBAFB04129CFA89FA93" box="[1190,1293,1264,1288]" pageId="12" pageNumber="13" type="paratype">Paratype</typeStatus>
: one female, same data as
<typeStatus id="54C68830A927FFBAFE8F1378FE0BFAB6" box="[301,399,1300,1325]" pageId="12" pageNumber="13" type="holotype">holotype</typeStatus>
(
<collectionCode id="ED6CAE57A927FFBAFE3C1378FE7CFAB7" LSID="urn:lsid:biocol.org:col:34992" box="[414,504,1300,1324]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="12" pageNumber="13">RMNH</collectionCode>
); one female, stream Kemantis, Sayap, Mt. Kinabalu, Borneo,
<collectingCountry id="F36A7602A927FFBAFB1E1378FAA1FAB6" box="[1212,1317,1300,1325]" name="Malaysia" pageId="12" pageNumber="13">Malaysia</collectingCountry>
, 6º
<geoCoordinate id="EE495055A927FFBAFAED1378FF2FFAD4" direction="north" orientation="latitude" pageId="12" pageNumber="13" precision="1" value="0.16401666">0 9.841 N</geoCoordinate>
116º
<geoCoordinate id="EE495055A927FFBAFF49135BFED6FAD4" box="[235,338,1335,1360]" direction="east" orientation="longitude" pageId="12" pageNumber="13" precision="55" value="33.936">33.936 E</geoCoordinate>
, alt.
<quantity id="4C859B77A927FFBAFE2A1354FE4BFAD4" box="[392,463,1336,1360]" metricMagnitude="2" metricUnit="m" metricValue="9.28" pageId="12" pageNumber="13" unit="m" value="928.0">928 m</quantity>
asl,
<date id="FFC31052A927FFBAFDA1135BFDFBFACB" box="[515,639,1335,1360]" pageId="12" pageNumber="13" value="2012-09-16">16-ix-2012</date>
(bar-coded,
<collectionCode id="ED6CAE57A927FFBAFCAF1354FCE2FACB" LSID="urn:lsid:biocol.org:col:34992" box="[781,870,1336,1360]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="12" pageNumber="13">RMNH</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="C3676519A927FFBAFF651330FE04FA03" pageId="12" pageNumber="13" type="diagnosis">
<paragraph id="8BC23692A927FFBAFF651330FE04FA03" blockId="12.[151,1437,1228,2009]" pageId="12" pageNumber="13">
<emphasis id="B909EA80A927FFBAFF651330FEBEFAEE" bold="true" box="[199,314,1372,1397]" pageId="12" pageNumber="13">Diagnosis</emphasis>
(Male unknown). Colour yellow; idiosoma slender (length/width ratio 1.5); mid dorsum with a pair of elongate platelets.
</paragraph>
</subSubSection>
<subSubSection id="C3676519A927FFBAFF6513C8FEFBF8D3" pageId="12" pageNumber="13" type="description">
<paragraph id="8BC23692A927FFBAFF6513C8FD26F9D6" blockId="12.[151,1437,1228,2009]" pageId="12" pageNumber="13">
<emphasis id="B909EA80A927FFBAFF6513C8FEDDFA26" bold="true" box="[199,345,1444,1469]" pageId="12" pageNumber="13">Description.</emphasis>
Female: Colour yellow. Glandularia enlarged, mid dorsum with a pair of elongate plates (
<figureCitation id="13462A17A927FFBAFACD13C8FF38FA7B" captionStart="FIGURE 6 A E" captionStartId="13.[151,250,1953,1975]" captionTargetBox="[176,1410,195,1904]" captionTargetId="figure@13.[176,1410,193,1932]" captionTargetPageId="13" captionText="FIGURE 6 A E. Sperchonopsis orientalis n. sp., female holotype, Mahua stream: A = idiosoma, dorsal view; C D = palp (P 1 missing); D = capitulum and chelicera. Scale bars = 100 Μm." httpUri="https://zenodo.org/record/287287/files/figure.png" pageId="12" pageNumber="13">Fig. 6A</figureCitation>
); integument papillate. Coxal field: tips of Cx-I projecting and covered by numerous setae. Genital field: Ac-1- 2 longish, Ac-3 roundish. Excretory pore surrounded by a sclerotized ring. Capitulum with rostrum slightly shorter than base (
<figureCitation id="13462A17A927FFBAFEB21063FEE3F9B3" box="[272,359,1551,1576]" captionStart="FIGURE 6 A E" captionStartId="13.[151,250,1953,1975]" captionTargetBox="[176,1410,195,1904]" captionTargetId="figure@13.[176,1410,193,1932]" captionTargetPageId="13" captionText="FIGURE 6 A E. Sperchonopsis orientalis n. sp., female holotype, Mahua stream: A = idiosoma, dorsal view; C D = palp (P 1 missing); D = capitulum and chelicera. Scale bars = 100 Μm." httpUri="https://zenodo.org/record/287287/files/figure.png" pageId="12" pageNumber="13">Fig. 6E</figureCitation>
). Palp: P2 with a long ventrodistal projection bearing 2 setae, P4 with a ventral projection close to segment base, bearing 2 setae (
<figureCitation id="13462A17A927FFBAFDAD1058FD12F9D6" box="[527,662,1588,1613]" captionStart="FIGURE 6 A E" captionStartId="13.[151,250,1953,1975]" captionTargetBox="[176,1410,195,1904]" captionTargetId="figure@13.[176,1410,193,1932]" captionTargetPageId="13" captionText="FIGURE 6 A E. Sperchonopsis orientalis n. sp., female holotype, Mahua stream: A = idiosoma, dorsal view; C D = palp (P 1 missing); D = capitulum and chelicera. Scale bars = 100 Μm." httpUri="https://zenodo.org/record/287287/files/figure.png" pageId="12" pageNumber="13">Figs. 6CD</figureCitation>
).
</paragraph>
<paragraph id="8BC23692A927FFBAFF651035FF4CF8BF" blockId="12.[151,1437,1228,2009]" pageId="12" pageNumber="13">
<emphasis id="B909EA80A927FFBAFF651035FE06F9EB" box="[199,386,1625,1648]" italics="true" pageId="12" pageNumber="13">Measurements—</emphasis>
Idiosoma (ventral view:
<figureCitation id="13462A17A927FFBAFD31103BFD63F9EB" box="[659,743,1623,1648]" captionStart="FIGURE 6 A E" captionStartId="13.[151,250,1953,1975]" captionTargetBox="[176,1410,195,1904]" captionTargetId="figure@13.[176,1410,193,1932]" captionTargetPageId="13" captionText="FIGURE 6 A E. Sperchonopsis orientalis n. sp., female holotype, Mahua stream: A = idiosoma, dorsal view; C D = palp (P 1 missing); D = capitulum and chelicera. Scale bars = 100 Μm." httpUri="https://zenodo.org/record/287287/files/figure.png" pageId="12" pageNumber="13">Fig. 6B</figureCitation>
) length 640 (664), width 420 (478); distance between anterior tip of Cx-I and posterior margin of Cx-IV 347, Cx-III width 423; genital flap length 102, width 37, length Ac-13: 2731, 2831, 2627. Capitulum ventral length 200; chelicera total length 195; palp: total length 363369, dorsal length/height, dorsal length/height ratio: P1, 33/43, 0.77; P2, 105/54, 1.94; P3, 103105/4548, 2.22.3; P4, 8891/2729, 3.23.3; P5, 34/15, 2.3; length P2/P4 ratio 1.19. Legs: dorsal length of I-leg-16: 47, 63, 69, 98, 109, 109.
</paragraph>
<paragraph id="8BC23692A927FFBAFF651143FEFBF8D3" blockId="12.[151,1437,1228,2009]" box="[199,383,1839,1864]" pageId="12" pageNumber="13">Male: unknown.</paragraph>
</subSubSection>
<subSubSection id="C3676519A927FFBAFF651138FC82F8F6" box="[199,774,1876,1901]" pageId="12" pageNumber="13" type="etymology">
<paragraph id="8BC23692A927FFBAFF651138FC82F8F6" blockId="12.[151,1437,1228,2009]" box="[199,774,1876,1901]" pageId="12" pageNumber="13">
<emphasis id="B909EA80A927FFBAFF651138FECAF8F6" bold="true" box="[199,334,1876,1901]" pageId="12" pageNumber="13">Etymology.</emphasis>
Named after the Oriental fauna region.
</paragraph>
</subSubSection>
<subSubSection id="C3676519A927FFB8FF65111BFCDFFE36" lastPageId="14" lastPageNumber="15" pageId="12" pageNumber="13" type="discussion">
<paragraph id="8BC23692A927FFB8FF65111BFC84FE86" blockId="12.[151,1437,1228,2009]" lastBlockId="14.[151,1437,151,464]" lastPageId="14" lastPageNumber="15" pageId="12" pageNumber="13">
<emphasis id="B909EA80A927FFBAFF65111BFEB0F80B" bold="true" box="[199,308,1911,1936]" pageId="12" pageNumber="13">Remarks</emphasis>
. The specimens examined from Borneo agree in general morphology of idiosoma and palps with
<taxonomicName id="4C7D4D11A927FFBAFF3511F0FDDCF82E" authority="Protz, 1896" authorityName="Protz" authorityYear="1896" box="[151,600,1948,1973]" class="Arachnida" family="Sperchonidae" genus="Sperchonopsis" kingdom="Animalia" order="Trombidiformes" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="verrucosa">
<emphasis id="B909EA80A927FFBAFF3511F0FE30F82F" box="[151,436,1948,1973]" italics="true" pageId="12" pageNumber="13">Sperchonopsis verrucosa</emphasis>
(Protz, 1896)
</taxonomicName>
, a species widely distributed in the Holarctic and Oriental regions. The latter species differs in the brownish colour (sometimes with bluish sclerotized parts) and in more stout idiosoma (length/width ratio &lt;1.5, data taken from Di Sabatino
<emphasis id="B909EA80A925FFB8FD5416F5FCA1FF2B" box="[758,805,151,176]" italics="true" pageId="14" pageNumber="15">et al</emphasis>
. 2010). These small differences obviously do not reflect the genetic distance as the molecular distance between a
<taxonomicName id="4C7D4D11A925FFB8FCAC16D0FBA6FF4E" box="[782,1058,188,213]" class="Arachnida" family="Sperchonidae" genus="Sperchonopsis" kingdom="Animalia" order="Trombidiformes" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="orientalis" status="sp. nov.">
<emphasis id="B909EA80A925FFB8FCAC16D0FBA6FF4E" box="[782,1058,188,213]" italics="true" pageId="14" pageNumber="15">Sperchonopsis orientalis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23A57FBA925FFB8FB8B16D0FBE3FF4F" box="[1065,1127,188,213]" pageId="14" pageNumber="15" rank="species">
<emphasis id="B909EA80A925FFB8FB8B16D0FBE4FF4E" bold="true" box="[1065,1120,188,213]" pageId="14" pageNumber="15">n. sp</emphasis>
.
</taxonomicNameLabel>
specimen from Borneo and
<taxonomicName id="4C7D4D11A925FFB8FF35168CFEA7FF63" box="[151,291,224,248]" class="Arachnida" family="Sperchonidae" genus="Sperchonopsis" kingdom="Animalia" order="Trombidiformes" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="verrucosa">
<emphasis id="B909EA80A925FFB8FF35168CFEA7FF63" box="[151,291,224,248]" italics="true" pageId="14" pageNumber="15">S. verrucosa</emphasis>
</taxonomicName>
from
<collectingCountry id="F36A7602A925FFB8FEC516B3FDA5FF63" box="[359,545,223,248]" name="Netherlands" pageId="14" pageNumber="15">The Netherlands</collectingCountry>
were found to be high (18.8% in COI). These high divergence values suggest a long independent evolutionary history of these species.
</paragraph>
<caption id="DF02661AA926FFBBFF3511CDFD7AF84D" httpUri="https://zenodo.org/record/287287/files/figure.png" pageId="13" pageNumber="14" targetBox="[176,1410,195,1904]" targetPageId="13">
<paragraph id="8BC23692A926FFBBFF3511CDFD7AF84D" blockId="13.[151,1435,1953,2006]" pageId="13" pageNumber="14">
<emphasis id="B909EA80A926FFBBFF3511CDFEC7F82D" bold="true" box="[151,323,1953,1975]" pageId="13" pageNumber="14">FIGURE 6AE.</emphasis>
<taxonomicName id="4C7D4D11A926FFBBFEEB11CDFDC0F82C" box="[329,580,1953,1975]" class="Arachnida" family="Sperchonidae" genus="Sperchonopsis" kingdom="Animalia" order="Trombidiformes" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="orientalis" status="sp. nov.">
<emphasis id="B909EA80A926FFBBFEEB11CDFDC0F82C" box="[329,580,1953,1975]" italics="true" pageId="13" pageNumber="14">Sperchonopsis orientalis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23A57FBA926FFBBFDE811CDFD07F82C" box="[586,643,1953,1975]" pageId="13" pageNumber="14" rank="species">
<emphasis id="B909EA80A926FFBBFDE811CDFDF9F82D" bold="true" box="[586,637,1953,1974]" pageId="13" pageNumber="14">n. sp</emphasis>
.
</taxonomicNameLabel>
, female holotype, Mahua stream: A = idiosoma, dorsal view; CD = palp (P1 missing); D = capitulum and chelicera. Scale bars = 100 Μm.
</paragraph>
</caption>
<paragraph id="8BC23692A925FFB8FF65174BFCDFFE36" blockId="14.[151,1437,151,464]" pageId="14" pageNumber="15">
<bibRefCitation id="EFEC4B63A925FFB8FF65174BFEE4FEDB" author="Wiles" box="[199,352,295,320]" pageId="14" pageNumber="15" refString="Wiles, P. R. (1999) The water mites (Acari: Hydrachnidia) of Borneo and additional new species from Thailand and Sulawesi. The Raffles Bulletin of Zoology, 47, 409 - 439." type="journal article" year="1999">Wiles (1999)</bibRefCitation>
reported
<taxonomicName id="4C7D4D11A925FFB8FE78174BFD7EFEDB" box="[474,762,295,320]" class="Arachnida" family="Sperchonidae" genus="Sperchonopsis" kingdom="Animalia" order="Trombidiformes" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="verrucosa">
<emphasis id="B909EA80A925FFB8FE78174BFD7EFEDB" box="[474,762,295,320]" italics="true" pageId="14" pageNumber="15">Sperchonopsis verrucosa</emphasis>
</taxonomicName>
from several sites in the catchment area of the Rivers Temburong and Belalong in
<collectingCountry id="F36A7602A925FFB8FE401720FDAAFEFE" box="[482,558,332,357]" name="Brunei" pageId="14" pageNumber="15">Brunei</collectingCountry>
Darussalem, without discussion of differences with the original description. Most probably his specimens refer to
<taxonomicName id="4C7D4D11A925FFB8FDE7171CFD4AFE13" box="[581,718,367,392]" class="Arachnida" family="Sperchonidae" genus="Sperchonopsis" kingdom="Animalia" order="Trombidiformes" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="orientalis" status="sp. nov.">
<emphasis id="B909EA80A925FFB8FDE7171CFD4AFE13" box="[581,718,367,392]" italics="true" pageId="14" pageNumber="15">S. orientalis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23A57FBA925FFB8FD741703FC92FE1C" box="[726,790,367,392]" pageId="14" pageNumber="15" rank="species">
<emphasis id="B909EA80A925FFB8FD741703FC8BFE13" bold="true" box="[726,783,367,392]" pageId="14" pageNumber="15">n. sp</emphasis>
.
</taxonomicNameLabel>
Further, all Oriental records of
<taxonomicName id="4C7D4D11A925FFB8FB231703FA18FE13" box="[1153,1436,367,392]" class="Arachnida" family="Sperchonidae" genus="Sperchonopsis" kingdom="Animalia" order="Trombidiformes" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="verrucosa">
<emphasis id="B909EA80A925FFB8FB231703FA18FE13" box="[1153,1436,367,392]" italics="true" pageId="14" pageNumber="15">Sperchonopsis verrucosa</emphasis>
</taxonomicName>
need revision, at best with application of molecular techniques.
</paragraph>
</subSubSection>
<subSubSection id="C3676519A925FFB8FF6517DBFE45FE54" box="[199,449,439,464]" pageId="14" pageNumber="15" type="distribution">
<paragraph id="8BC23692A925FFB8FF6517DBFE45FE54" blockId="14.[151,1437,151,464]" box="[199,449,439,464]" pageId="14" pageNumber="15">
<emphasis id="B909EA80A925FFB8FF6517DBFEE4FE4B" bold="true" box="[199,352,439,464]" pageId="14" pageNumber="15">Distribution.</emphasis>
Borneo.
</paragraph>
</subSubSection>
</treatment>
</document>