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<document id="FE1959A92D42A811AFA1E801C406D120" ID-DOI="10.5281/zenodo.213946" ID-GBIF-Dataset="2a54e127-4d40-404e-ab7f-5b48f6e37e83" ID-ISSN="1175-5326" ID-Zenodo-Dep="213946" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460564534622" checkinUser="plazi" docAuthor="Grisales, Diana, Wolff, Marta &amp; De, Claudio J. B." docDate="2012" docId="03BE87F8522DD434C09420A4FC35F9E6" docLanguage="en" docName="zt03591p046.pdf" docOrigin="Zootaxa 3591" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Fannia katios Grisales, Wolff &amp; De, 2012, sp. nov." docType="treatment" docVersion="8" lastPageNumber="30" masterDocId="FF87FF805231D429C0032033FFBBFFA9" masterDocTitle="Neotropical Fanniidae (Insecta, Diptera): new species of Fannia from Colombia" masterLastPageNumber="46" masterPageNumber="1" pageNumber="29" updateTime="1698320797981" updateUser="plazi">
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<mods:title id="CCA3B43E26A1EDA0601314A3778D90B4">Neotropical Fanniidae (Insecta, Diptera): new species of Fannia from Colombia</mods:title>
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<mods:namePart id="37B3E634EB20AF91F060C6C7F1A64C54">Grisales, Diana</mods:namePart>
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<mods:namePart id="1C9CBCAAAADD60D62BD2918D40D75411">Wolff, Marta</mods:namePart>
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<mods:namePart id="9900E95F70F7B1AAC929B1F9ECB1DCE0">De, Claudio J. B.</mods:namePart>
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<mods:date id="2085074AD560449134217B1B248FF7E6">2012</mods:date>
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<treatment id="03BE87F8522DD434C09420A4FC35F9E6" ID-DOI="http://doi.org/10.5281/zenodo.6175061" ID-GBIF-Taxon="119557553" ID-Zenodo-Dep="6175061" LSID="urn:lsid:plazi:treatment:03BE87F8522DD434C09420A4FC35F9E6" httpUri="http://treatment.plazi.org/id/03BE87F8522DD434C09420A4FC35F9E6" lastPageId="29" lastPageNumber="30" pageId="28" pageNumber="29">
<subSubSection id="C30D6565522DD435C09420A4FD2CFF7B" pageId="28" pageNumber="29" type="nomenclature">
<paragraph id="8BA836EE522DD435C09420A4FE1FFF18" blockId="28.[151,663,151,210]" box="[151,420,151,177]" pageId="28" pageNumber="29">
<heading id="D0E08182522DD435C09420A4FE1FFF18" bold="true" box="[151,420,151,177]" fontSize="11" level="1" pageId="28" pageNumber="29" reason="1">
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<taxonomicName id="4C174D6D522DD435C09420A4FEFBFF18" ID-CoL="9J2KD" box="[151,320,151,177]" class="Insecta" family="Fanniidae" genus="Fannia" kingdom="Animalia" order="Diptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="katios" status="sp. nov.">
<emphasis id="B963EAFC522DD435C09420A4FEFBFF18" bold="true" box="[151,320,151,177]" italics="true" pageId="28" pageNumber="29">Fannia katios</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2505787522DD435C14420A4FE1FFF18" box="[327,420,151,177]" pageId="28" pageNumber="29" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BA836EE522DD435C0942089FD2CFF7B" blockId="28.[151,663,151,210]" box="[151,663,186,210]" pageId="28" pageNumber="29">
(
<figureCitation id="132C2A6B522DD435C09C2089FEFAFF7B" box="[159,321,186,210]" captionStart="FIGURES 1 12" captionStartId="8.[151,264,1845,1868]" captionTargetBox="[171,1399,339,1823]" captionTargetId="figure@8.[171,1417,339,1823]" captionTargetPageId="8" captionText="FIGURES 1 12. Fannia: (1) F. canicularis, male, head, anterior view; (2) F. chibcha sp. nov., male holotype, hind leg, anterior view; (3) F. dodgei, male, abdomen, posterior view; (4) F. dorsomaculata sp. nov., male holotype, lateral view; (5) F. dorsomaculata sp. nov., male holotype, dorsal view; (6) F. grandis sp. nov., male paratype, lateral view; (7) F. grandis, male, dorsal view; (8) F. iguaque sp. nov., male holotype, head, anterior view; (9) F. katios sp. nov., female paratype, head, anterior view; (10) F. katios sp. nov., male holotype, posterior view; (11) F. katios sp. nov., female paratype, posterior view; (12) F. lamosca sp. nov., male holotype, dorsal view. Scale: 1 mm." httpUri="https://zenodo.org/record/213947/files/figure.png" pageId="28" pageNumber="29">Figs. 9, 10, 11</figureCitation>
,
<figureCitation id="132C2A6B522DD435C14E2089FED0FF7B" box="[333,363,186,210]" captionStart="FIGURES 20 37" captionStartId="13.[151,264,1716,1739]" captionTargetBox="[151,1435,193,1694]" captionTargetId="figure@13.[151,1436,193,1695]" captionTargetPageId="13" captionText="FIGURES 20 37. Fannia: male, hind leg, anterior view: (20) F. canicularis; (21) F. chibcha sp. nov., holotype; (22) F. chingaza sp. nov., holotype; (23) F. dodgei; (24) F. dorsomaculata sp. nov., holotype; (25) F. e m b e r a sp. nov., paratype; (26) F. grandis; (27) F. iguaque sp. nov., holotype; (28) F. katios sp. nov., holotype; (29) F. l a m o s c a sp. nov., holotype; (30) F. magdalena sp. nov., holotype; (31) F. pijao sp. nov., holotype; (32) F. p o rc e sp. nov., holotype; (33) F. pu s i o; (34) F. quimbaya sp. nov., holotype; (35) F. sumapaz sp. nov., holotype; (36) F. sutagao sp. nov., holotype; (37) F. trimaculata. Scale: 1 mm." httpUri="https://zenodo.org/record/213949/files/figure.png" pageId="28" pageNumber="29">28</figureCitation>
,
<figureCitation id="132C2A6B522DD435C1742089FE2EFF7B" box="[375,405,186,210]" captionStart="FIGURES 38 55" captionStartId="14.[151,264,1744,1767]" captionTargetBox="[151,1435,193,1723]" captionTargetId="figure@14.[151,1436,193,1723]" captionTargetPageId="14" captionText="FIGURES 38 55. Fannia: male, hind leg, posterior view: (38) F. canicularis; (39) F. chibcha sp. nov., holotype; (40) F. chingaza sp. nov., holotype; (41) F. dodgei; (42) F. dorsomaculata sp. nov., holotype; (43) F. e m b e r a sp. nov., holotype; (44) F. grandis; (45) F. iguaque sp. nov., holotype; (46) F. katios sp. nov., holotype; (47) F. l a m o s c a sp. nov., holotype; (48) F. magdalena sp. nov., holotype; (49) F. pijao sp. nov., holotype; (50) F. p o rc e sp. nov., holotype; (51) F. pu s i o; (52) F. quimbaya sp. nov., holotype; (53) F. sumapaz sp. nov., holotype; (54) F. sutagao sp. nov., holotype; (55) F. trimaculata. Scale: 1 mm." httpUri="https://zenodo.org/record/213950/files/figure.png" pageId="28" pageNumber="29">46</figureCitation>
,
<figureCitation id="132C2A6B522DD435C1A22089FE04FF7B" box="[417,447,186,210]" captionStart="FIGURES 56 73" captionStartId="15.[151,264,1875,1898]" captionTargetBox="[200,1393,193,1844]" captionTargetId="figure@15.[200,1394,193,1845]" captionTargetPageId="15" captionText="FIGURES 56 73. Fannia: male, sternite 5, ventral view: (56) F. c a ni c ul a ri s; (57) F. c h i b c h a sp. nov., paratype; (58) F. chingaza sp. nov., holotype; (59) F. dodgei; (60) F. dorsomaculata sp. nov., paratype; (61) F. e m b e r a sp. nov., paratype; (62) F. grandis; (63) F. iguaque sp. nov., paratype; (64) F. katios sp. nov., paratype; (65) F. l a m o s c a sp. nov., paratype; (66) F. magdalena sp. nov., paratype; (67) F. pijao sp. nov., paratype; (68) F. porce sp. nov., holotype; (69) F. p u s i o; (70) F. quimbaya sp. nov., paratype; (71) F. sumapaz sp. nov., paratype; (72) F. sutagao sp. nov., paratype; (73) F. trimaculata. Scale: 0.1 mm." httpUri="https://zenodo.org/record/213951/files/figure.png" pageId="28" pageNumber="29">64</figureCitation>
,
<figureCitation id="132C2A6B522DD435C1C82089FE51FF7B" box="[459,490,186,210]" captionStart="FIGURES 74 82" captionStartId="16.[151,264,1809,1832]" captionTargetBox="[152,1435,193,1787]" captionTargetId="figure@16.[151,1436,193,1789]" captionTargetPageId="16" captionText="FIGURES 74 82. Fannia: male, epandrium, cercal plate and surstyli, dorsal view: (74) F. c an i c ul a r i s; (75) F. c hi b c ha sp. nov., paratype; (76) F. c h i n g a z a sp. nov., holotype; (77) F. dodgei; (78) F. dorsomaculata sp. nov., paratype; (79) F. e m b e r a sp. nov., paratype; (80) F. grandis; (81) F. iguaque sp. nov., paratype; (82) F. katios sp. nov., paratype. Scale: 0.1 mm." httpUri="https://zenodo.org/record/213952/files/figure.png" pageId="28" pageNumber="29">82</figureCitation>
,
<figureCitation id="132C2A6B522DD435C1F62089FD99FF7B" box="[501,546,186,210]" captionStart="FIGURES 92 100" captionStartId="18.[151,264,1950,1973]" captionTargetBox="[246,1354,193,1930]" captionTargetId="figure@18.[237,1355,193,1931]" captionTargetPageId="18" captionText="FIGURES 92 100. Fannia: male, epandrium, cercal plate and surstyli, lateral view: (92) F. canicularis; (93) F. c h i b c h a sp. nov., paratype; (94) F. c h i n g a z a sp. nov., holotype; (95) F. dodgei; (96) F. dorsomaculata sp. nov., paratype; (97) F. e m b e r a sp. nov., paratype; (98) F. grandis; (99) F. iguaque sp. nov., paratype; (100) F. katios sp. nov., paratype. Scale: 0.1 mm." httpUri="https://zenodo.org/record/213954/files/figure.png" pageId="28" pageNumber="29">100</figureCitation>
,
<figureCitation id="132C2A6B522DD435C22E2089FDE3FF7B" box="[557,600,186,210]" captionStart="FIGURES 110 118" captionStartId="20.[151,264,1635,1658]" captionTargetBox="[165,1427,193,1608]" captionTargetId="figure@20.[163,1428,193,1609]" captionTargetPageId="20" captionText="FIGURES 110 118. Fannia: male, hypandrium and associated structures, dorsal view: (110) F. canicularis; (111) F. chibcha sp. nov., paratype; (112) F. c h i n g a z a sp. nov., holotype; (113) F. dodgei; (114) F. dorsomaculata sp. nov., paratype; (115) F. embera sp. nov., paratype; (116) F. grandis; (117) F. iguaque sp. nov., paratype. Scale: 0.1 mm." httpUri="https://zenodo.org/record/213956/files/figure.png" pageId="28" pageNumber="29">118</figureCitation>
, 133)
</paragraph>
</subSubSection>
<subSubSection id="C30D6565522DD435C0942131FD03FE5B" pageId="28" pageNumber="29" type="diagnosis">
<paragraph id="8BA836EE522DD435C0942131FD03FE5B" blockId="28.[151,1437,258,2014]" pageId="28" pageNumber="29">
<emphasis id="B963EAFC522DD435C0942131FEAAFEB2" bold="true" box="[151,273,258,283]" pageId="28" pageNumber="29">Diagnosis.</emphasis>
these characters apply only to the male sex. Arista brown with base yellowish; scape brown with a apical margin yellowish; postpedicel yellow with apex and outer surface brownish-grey; palpus yellow with base brownish; scutum with weak and narrow median vitta, and two brown marks on intra-alar area; hind femur on ventral surface without pre-apical protuberance and with weak apical constriction, anterior and posterior surfaces swollen on basal third, 2 pre-apical
<emphasis id="B963EAFC522DD435C22F21A7FDFEFE02" box="[556,581,404,427]" italics="true" pageId="28" pageNumber="29">av</emphasis>
, apical seta longer, posterior surface on apical half bearing weak setae with hooked apices, setae increasing in length up to swollen area, not forming a tuft; abdomen with tergites 14 laterally translucent-yellow; abdomen brown trimaculate.
</paragraph>
</subSubSection>
<subSubSection id="C30D6565522DD434C0C421CDFDEAFCF5" lastPageId="29" lastPageNumber="30" pageId="28" pageNumber="29" type="description">
<paragraph id="8BA836EE522DD435C0C421CDFC70FDBF" blockId="28.[151,1437,258,2014]" box="[199,971,510,535]" pageId="28" pageNumber="29">
<emphasis id="B963EAFC522DD435C0C421CDFEEAFDBE" bold="true" box="[199,337,510,535]" pageId="28" pageNumber="29">Description</emphasis>
.
<emphasis id="B963EAFC522DD435C15C21CCFDB7FDBE" bold="true" box="[351,524,511,535]" pageId="28" pageNumber="29">
<typeStatus id="54AC884C522DD435C15C21CCFE77FDBE" box="[351,460,511,535]" pageId="28" pageNumber="29" type="holotype">Holotype</typeStatus>
male
</emphasis>
: body length
<quantity id="4CEF9B0B522DD435C2A421CDFD50FDBF" box="[679,747,510,534]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="28" pageNumber="29" unit="mm" value="3.0">3 mm</quantity>
; wing length
<quantity id="4CEF9B0B522DD435C38121CDFC7DFDBF" box="[898,966,510,534]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="28" pageNumber="29" unit="mm" value="3.0">3 mm</quantity>
.
</paragraph>
<paragraph id="8BA836EE522DD435C0C42210FC18FCBB" blockId="28.[151,1437,258,2014]" pageId="28" pageNumber="29">
<emphasis id="B963EAFC522DD435C0C42210FEBFFD92" box="[199,260,547,571]" italics="true" pageId="28" pageNumber="29">Head</emphasis>
: eye bare. Frontal vitta velvety brown, posterior area velvety black and greyish pilose centrally. Interocular space 0.12 mm.
<emphasis id="B963EAFC522DD435C1D1227BFE55FDF6" box="[466,494,584,607]" italics="true" pageId="28" pageNumber="29">Fr</emphasis>
810. Ocellar triangle black;
<emphasis id="B963EAFC522DD435C343227BFCE0FDF6" box="[832,859,584,607]" italics="true" pageId="28" pageNumber="29">oc</emphasis>
proclinate and developed, 3 less developed setulae on ocellar area.
<emphasis id="B963EAFC522DD435C15D225FFE31FD2A" box="[350,394,620,643]" italics="true" pageId="28" pageNumber="29">Poc</emphasis>
converging. Fronto-orbital plate greyish pollinose, posterior area to eye margin black. Parafacial greyish pollinose and without short setae. Gena grey. Lunule brown. Scape brown with apical margin yellowish and 2 short setae. Pedicel yellow, golden pollinose, dorsal setae short. Postpedicel yellow with apex and outer surface brownish-grey, yellow pilosity and measuring 2 times the length of pedicel. Arista brown with base yellow and pubescent. Palpus yellow with base brownish and filiform
</paragraph>
<paragraph id="8BA836EE522DD435C0C4232DFBFDFBA7" blockId="28.[151,1437,258,2014]" pageId="28" pageNumber="29">
<emphasis id="B963EAFC522DD435C0C4232DFEA3FC9F" box="[199,280,798,822]" italics="true" pageId="28" pageNumber="29">Thorax</emphasis>
: grey pollinose, particularly on
<emphasis id="B963EAFC522DD435C27E2313FD19FC9E" box="[637,674,800,823]" italics="true" pageId="28" pageNumber="29">acr</emphasis>
and
<emphasis id="B963EAFC522DD435C2DA232DFD48FC9F" box="[729,755,798,822]" italics="true" pageId="28" pageNumber="29">dc</emphasis>
areas; scutum with median vitta narrow and weak up to base of scutellum, two postsutural marks between intraalar area and the second pair of
<emphasis id="B963EAFC522DD435C47F2370FB2CFCF2" box="[1148,1175,835,859]" italics="true" pageId="28" pageNumber="29">dc</emphasis>
, pre-sutural area with inconspicuous brown marks. Scutellum with basal, brownish mark, apex yellowish.
<emphasis id="B963EAFC522DD435C44C2355FBCCFCD7" box="[1103,1143,870,894]" italics="true" pageId="28" pageNumber="29">Acr</emphasis>
3:4, not arranged in rows;
<emphasis id="B963EAFC522DD435C09423B8FF09FC0A" box="[151,178,907,931]" italics="true" pageId="28" pageNumber="29">dc</emphasis>
2:3, with postsutural
<emphasis id="B963EAFC522DD435C1B923B8FE6FFC0A" box="[442,468,907,931]" italics="true" pageId="28" pageNumber="29">dc</emphasis>
setae forming an inverted cone. Postpronotum with 2 developed setae, 1 shorter. Posterior
<emphasis id="B963EAFC522DD435C100239CFEA2FC6F" box="[259,281,943,966]" italics="true" pageId="28" pageNumber="29">ia</emphasis>
longer than weakly differentiated anterior seta.
<emphasis id="B963EAFC522DD435C3332383FCE1FC6E" box="[816,858,944,967]" italics="true" pageId="28" pageNumber="29">Pra</emphasis>
1, weakly differentiated and near
<emphasis id="B963EAFC522DD435C4D4239DFABDFC6F" box="[1239,1286,942,966]" italics="true" pageId="28" pageNumber="29">spal</emphasis>
. Proepimeral area with ground setulae. Pre-basal pre-scutellar seta absent, area with short setae. Subapical scutellar setae 1 pair. Discal scutellar setae 1 pair. Discal area with short setae. Lateral scutellar setae 24.
</paragraph>
<paragraph id="8BA836EE522DD435C0C4242FFA27FB9B" blockId="28.[151,1437,258,2014]" box="[199,1436,1050,1080]" pageId="28" pageNumber="29">
<emphasis id="B963EAFC522DD435C0C4242FFF44FB9A" box="[199,255,1052,1075]" italics="true" pageId="28" pageNumber="29">Wing</emphasis>
: yellowish; R1 with weak dilatation on basal third. Calypters whitish. Haltere yellowish with base yellow.
</paragraph>
<paragraph id="8BA836EE522DD435C0C42471FA92F9DC" blockId="28.[151,1437,258,2014]" pageId="28" pageNumber="29">
<emphasis id="B963EAFC522DD435C0C42471FF47FBF0" box="[199,252,1090,1113]" italics="true" pageId="28" pageNumber="29">Legs</emphasis>
: brown; fore coxa brown-yellowish; fore and mid trochanter yellow; apex of fore femur and basal third of fore tibia yellow; apex of mid and hind femora and base and apex of tibiae yellow; tarsi darkened and pulvilli yellow-brownish. Fore femur on ventral surface bare; 1 row of short and strong
<emphasis id="B963EAFC522DD435C42724BAFB89FB08" box="[1060,1074,1161,1185]" italics="true" pageId="28" pageNumber="29">d</emphasis>
and
<emphasis id="B963EAFC522DD435C46824BAFB3CFB08" box="[1131,1159,1161,1185]" italics="true" pageId="28" pageNumber="29">pd</emphasis>
setae; 1 row of thin and short
<emphasis id="B963EAFC522DD435C0D4249DFF5EFB6C" box="[215,229,1198,1221]" italics="true" pageId="28" pageNumber="29">p</emphasis>
;
<emphasis id="B963EAFC522DD435C0F5249DFEABFB6C" box="[246,272,1198,1221]" italics="true" pageId="28" pageNumber="29">pv</emphasis>
short and sparse on basal half, ending in 7 developed pre-apical
<emphasis id="B963EAFC522DD435C3F9249EFBB3FB6C" box="[1018,1032,1197,1221]" italics="true" pageId="28" pageNumber="29">d</emphasis>
setae. Fore tibia with 1 strong preapical
<emphasis id="B963EAFC522DD435C0E124E2FF4BFB40" box="[226,240,1233,1257]" italics="true" pageId="28" pageNumber="29">d</emphasis>
; 1 short apical
<emphasis id="B963EAFC522DD435C1A724E1FE0BFB40" box="[420,432,1234,1257]" italics="true" pageId="28" pageNumber="29">v</emphasis>
; 1 short apical
<emphasis id="B963EAFC522DD435C26724E1FDC6FB40" box="[612,637,1234,1257]" italics="true" pageId="28" pageNumber="29">pv</emphasis>
. Fore tarsus with strong basal
<emphasis id="B963EAFC522DD435C3DE24E1FC52FB40" box="[989,1001,1234,1257]" italics="true" pageId="28" pageNumber="29">v</emphasis>
on first tarsomere and 12 thin setae; tarsomere 13 with 1 thin and long apical-lateral
<emphasis id="B963EAFC522DD435C2D724C4FD5BFAA7" box="[724,736,1271,1294]" italics="true" pageId="28" pageNumber="29">v</emphasis>
; tarsomere 5 slightly flattened with respect to the remaining tarsomeres and yellow apically. Mid femur weakly constricted on pre-apical ventral surface; 1 row of
<emphasis id="B963EAFC522DD435C54F2529FADCFA98" box="[1356,1383,1306,1329]" italics="true" pageId="28" pageNumber="29">av</emphasis>
that decrease in length and become scarce towards apex; two rows of unordered
<emphasis id="B963EAFC522DD435C3EC250CFC46FAFF" box="[1007,1021,1343,1366]" italics="true" pageId="28" pageNumber="29">a</emphasis>
ending in long and strong pre-apical setae; 1 row of short
<emphasis id="B963EAFC522DD435C1842551FE28FAD0" box="[391,403,1378,1401]" italics="true" pageId="28" pageNumber="29">v</emphasis>
with hooked apices on basal third; 1 row of strong
<emphasis id="B963EAFC522DD435C3DC2551FC42FAD0" box="[991,1017,1378,1401]" italics="true" pageId="28" pageNumber="29">pv</emphasis>
with hooked apices; 1 row of long
<emphasis id="B963EAFC522DD435C58D2551FA27FAD0" box="[1422,1436,1378,1401]" italics="true" pageId="28" pageNumber="29">p</emphasis>
with hooked apices, 4 apical most setae stronger; 1 row of short
<emphasis id="B963EAFC522DD435C37025B5FC34FA37" box="[883,911,1414,1438]" italics="true" pageId="28" pageNumber="29">pd</emphasis>
on basal third. Mid tibia with 1
<emphasis id="B963EAFC522DD435C50325B5FAA7FA37" box="[1280,1308,1414,1438]" italics="true" pageId="28" pageNumber="29">ad</emphasis>
on edge of apical third; 1 pre-apical
<emphasis id="B963EAFC522DD435C1B12599FE77FA68" box="[434,460,1450,1473]" italics="true" pageId="28" pageNumber="29">av</emphasis>
; 1 strong apical
<emphasis id="B963EAFC522DD435C28A2599FD2EFA68" box="[649,661,1450,1473]" italics="true" pageId="28" pageNumber="29">v</emphasis>
; 1pre-apical
<emphasis id="B963EAFC522DD435C329259AFC83FA68" box="[810,824,1449,1473]" italics="true" pageId="28" pageNumber="29">d</emphasis>
; 1 short median
<emphasis id="B963EAFC522DD435C3F62599FBB8FA68" box="[1013,1027,1450,1473]" italics="true" pageId="28" pageNumber="29">p</emphasis>
; 1 short apical
<emphasis id="B963EAFC522DD435C4B12599FB70FA68" box="[1202,1227,1450,1473]" italics="true" pageId="28" pageNumber="29">pv</emphasis>
.
<collectingCountry id="F300767E522DD435C4DA259AFAAAFA68" box="[1241,1297,1449,1473]" name="India" pageId="28" pageNumber="29">Hind</collectingCountry>
coxa with 1 seta on posterior margin.
<collectingCountry id="F300767E522DD435C1D225FEFDB2FA4C" box="[465,521,1485,1509]" name="India" pageId="28" pageNumber="29">Hind</collectingCountry>
femur on ventral surface without pre-apical protuberance, and weak apical constriction (
<figureCitation id="132C2A6B522DD435C12E25C2FE3FF9A0" box="[301,388,1521,1545]" captionStart="FIGURES 20 37" captionStartId="13.[151,264,1716,1739]" captionTargetBox="[151,1435,193,1694]" captionTargetId="figure@13.[151,1436,193,1695]" captionTargetPageId="13" captionText="FIGURES 20 37. Fannia: male, hind leg, anterior view: (20) F. canicularis; (21) F. chibcha sp. nov., holotype; (22) F. chingaza sp. nov., holotype; (23) F. dodgei; (24) F. dorsomaculata sp. nov., holotype; (25) F. e m b e r a sp. nov., paratype; (26) F. grandis; (27) F. iguaque sp. nov., holotype; (28) F. katios sp. nov., holotype; (29) F. l a m o s c a sp. nov., holotype; (30) F. magdalena sp. nov., holotype; (31) F. pijao sp. nov., holotype; (32) F. p o rc e sp. nov., holotype; (33) F. pu s i o; (34) F. quimbaya sp. nov., holotype; (35) F. sumapaz sp. nov., holotype; (36) F. sutagao sp. nov., holotype; (37) F. trimaculata. Scale: 1 mm." httpUri="https://zenodo.org/record/213949/files/figure.png" pageId="28" pageNumber="29">Fig. 28</figureCitation>
); swollen on anterior and posterior surfaces and on median third; 2 pre-apical
<emphasis id="B963EAFC522DD435C51025C1FA97F9A0" box="[1299,1324,1522,1545]" italics="true" pageId="28" pageNumber="29">av</emphasis>
, 1 longer apical seta (
<figureCitation id="132C2A6B522DD435C11E2626FEC9F984" box="[285,370,1557,1581]" captionStart="FIGURES 20 37" captionStartId="13.[151,264,1716,1739]" captionTargetBox="[151,1435,193,1694]" captionTargetId="figure@13.[151,1436,193,1695]" captionTargetPageId="13" captionText="FIGURES 20 37. Fannia: male, hind leg, anterior view: (20) F. canicularis; (21) F. chibcha sp. nov., holotype; (22) F. chingaza sp. nov., holotype; (23) F. dodgei; (24) F. dorsomaculata sp. nov., holotype; (25) F. e m b e r a sp. nov., paratype; (26) F. grandis; (27) F. iguaque sp. nov., holotype; (28) F. katios sp. nov., holotype; (29) F. l a m o s c a sp. nov., holotype; (30) F. magdalena sp. nov., holotype; (31) F. pijao sp. nov., holotype; (32) F. p o rc e sp. nov., holotype; (33) F. pu s i o; (34) F. quimbaya sp. nov., holotype; (35) F. sumapaz sp. nov., holotype; (36) F. sutagao sp. nov., holotype; (37) F. trimaculata. Scale: 1 mm." httpUri="https://zenodo.org/record/213949/files/figure.png" pageId="28" pageNumber="29">Fig. 28</figureCitation>
); ventral surface bare; 1 row of short
<emphasis id="B963EAFC522DD435C31E2625FC82F987" box="[797,825,1558,1582]" italics="true" pageId="28" pageNumber="29">ad</emphasis>
on basal half; 2 developed pre-apical
<emphasis id="B963EAFC522DD435C4EA2625FB4CF987" box="[1257,1271,1558,1582]" italics="true" pageId="28" pageNumber="29">d</emphasis>
;
<emphasis id="B963EAFC522DD435C5052625FA99F987" box="[1286,1314,1558,1582]" italics="true" pageId="28" pageNumber="29">pd</emphasis>
unordered on basal half, thin and with hooked apices;
<emphasis id="B963EAFC522DD435C2872609FD29F9F8" box="[644,658,1594,1617]" italics="true" pageId="28" pageNumber="29">p</emphasis>
weak with hooked apices on apical half that increase in length up to swollen area, not forming a tuft, last 2 setae with straight apices (
<figureCitation id="132C2A6B522DD435C372266EFC7EF9DC" box="[881,965,1629,1653]" captionStart="FIGURES 38 55" captionStartId="14.[151,264,1744,1767]" captionTargetBox="[151,1435,193,1723]" captionTargetId="figure@14.[151,1436,193,1723]" captionTargetPageId="14" captionText="FIGURES 38 55. Fannia: male, hind leg, posterior view: (38) F. canicularis; (39) F. chibcha sp. nov., holotype; (40) F. chingaza sp. nov., holotype; (41) F. dodgei; (42) F. dorsomaculata sp. nov., holotype; (43) F. e m b e r a sp. nov., holotype; (44) F. grandis; (45) F. iguaque sp. nov., holotype; (46) F. katios sp. nov., holotype; (47) F. l a m o s c a sp. nov., holotype; (48) F. magdalena sp. nov., holotype; (49) F. pijao sp. nov., holotype; (50) F. p o rc e sp. nov., holotype; (51) F. pu s i o; (52) F. quimbaya sp. nov., holotype; (53) F. sumapaz sp. nov., holotype; (54) F. sutagao sp. nov., holotype; (55) F. trimaculata. Scale: 1 mm." httpUri="https://zenodo.org/record/213950/files/figure.png" pageId="28" pageNumber="29">Fig. 46</figureCitation>
). Tibia and hind tarsus missing.
</paragraph>
<paragraph id="8BA836EE522DD435C0C426B2FCCBF810" blockId="28.[151,1437,258,2014]" pageId="28" pageNumber="29">
<emphasis id="B963EAFC522DD435C0C426B2FE8AF930" box="[199,305,1665,1689]" italics="true" pageId="28" pageNumber="29">Abdomen</emphasis>
: yellowish with brown marks (
<figureCitation id="132C2A6B522DD435C29026B2FD5CF930" box="[659,743,1665,1689]" captionStart="FIGURES 1 12" captionStartId="8.[151,264,1845,1868]" captionTargetBox="[171,1399,339,1823]" captionTargetId="figure@8.[171,1417,339,1823]" captionTargetPageId="8" captionText="FIGURES 1 12. Fannia: (1) F. canicularis, male, head, anterior view; (2) F. chibcha sp. nov., male holotype, hind leg, anterior view; (3) F. dodgei, male, abdomen, posterior view; (4) F. dorsomaculata sp. nov., male holotype, lateral view; (5) F. dorsomaculata sp. nov., male holotype, dorsal view; (6) F. grandis sp. nov., male paratype, lateral view; (7) F. grandis, male, dorsal view; (8) F. iguaque sp. nov., male holotype, head, anterior view; (9) F. katios sp. nov., female paratype, head, anterior view; (10) F. katios sp. nov., male holotype, posterior view; (11) F. katios sp. nov., female paratype, posterior view; (12) F. lamosca sp. nov., male holotype, dorsal view. Scale: 1 mm." httpUri="https://zenodo.org/record/213947/files/figure.png" pageId="28" pageNumber="29">Fig. 10</figureCitation>
), syntergite 1+2, tergites 3 and 4 with 2 brown lateral marks; tergites 3 and 4 with median vitta brown, expanded on apical margin of tergite, not reaching lateral area; syntergite 1+2, tergites 3 and 4 translucent-yellow, very pale; tergite 5 brown-yellowish pollinose. Syntergite 1+2, tergites 34 with developed marginal setae; tergite 5 with one pair of lateral setae. Sternite 1 bare. Sternite 5 as in
<figureCitation id="132C2A6B522DD435C54626DEFA23F8AC" box="[1349,1432,1773,1797]" captionStart="FIGURES 56 73" captionStartId="15.[151,264,1875,1898]" captionTargetBox="[200,1393,193,1844]" captionTargetId="figure@15.[200,1394,193,1845]" captionTargetPageId="15" captionText="FIGURES 56 73. Fannia: male, sternite 5, ventral view: (56) F. c a ni c ul a ri s; (57) F. c h i b c h a sp. nov., paratype; (58) F. chingaza sp. nov., holotype; (59) F. dodgei; (60) F. dorsomaculata sp. nov., paratype; (61) F. e m b e r a sp. nov., paratype; (62) F. grandis; (63) F. iguaque sp. nov., paratype; (64) F. katios sp. nov., paratype; (65) F. l a m o s c a sp. nov., paratype; (66) F. magdalena sp. nov., paratype; (67) F. pijao sp. nov., paratype; (68) F. porce sp. nov., holotype; (69) F. p u s i o; (70) F. quimbaya sp. nov., paratype; (71) F. sumapaz sp. nov., paratype; (72) F. sutagao sp. nov., paratype; (73) F. trimaculata. Scale: 0.1 mm." httpUri="https://zenodo.org/record/213951/files/figure.png" pageId="28" pageNumber="29">Fig. 64</figureCitation>
. Terminalia (
<figureCitation id="132C2A6B522DD435C1232722FEC5F880" box="[288,382,1809,1833]" captionStart="FIGURES 74 82" captionStartId="16.[151,264,1809,1832]" captionTargetBox="[152,1435,193,1787]" captionTargetId="figure@16.[151,1436,193,1789]" captionTargetPageId="16" captionText="FIGURES 74 82. Fannia: male, epandrium, cercal plate and surstyli, dorsal view: (74) F. c an i c ul a r i s; (75) F. c hi b c ha sp. nov., paratype; (76) F. c h i n g a z a sp. nov., holotype; (77) F. dodgei; (78) F. dorsomaculata sp. nov., paratype; (79) F. e m b e r a sp. nov., paratype; (80) F. grandis; (81) F. iguaque sp. nov., paratype; (82) F. katios sp. nov., paratype. Scale: 0.1 mm." httpUri="https://zenodo.org/record/213952/files/figure.png" pageId="28" pageNumber="29">Figs. 82</figureCitation>
,
<figureCitation id="132C2A6B522DD435C1882722FE01F880" box="[395,442,1809,1833]" captionStart="FIGURES 92 100" captionStartId="18.[151,264,1950,1973]" captionTargetBox="[246,1354,193,1930]" captionTargetId="figure@18.[237,1355,193,1931]" captionTargetPageId="18" captionText="FIGURES 92 100. Fannia: male, epandrium, cercal plate and surstyli, lateral view: (92) F. canicularis; (93) F. c h i b c h a sp. nov., paratype; (94) F. c h i n g a z a sp. nov., holotype; (95) F. dodgei; (96) F. dorsomaculata sp. nov., paratype; (97) F. e m b e r a sp. nov., paratype; (98) F. grandis; (99) F. iguaque sp. nov., paratype; (100) F. katios sp. nov., paratype. Scale: 0.1 mm." httpUri="https://zenodo.org/record/213954/files/figure.png" pageId="28" pageNumber="29">100</figureCitation>
): epandrium slightly wider than long, with weakly developed setae on basal half; cercal plate wide and concave, fused up to basal third and with 2 pointed apical projections, with short setae, most of which are straight, ground setulae scarce; surstylus long and articulated with epandrium, apex expanded ventrally and with many thin and short setae, inner basal margin setulose; bacilliform process spiralled and short, on apex of cercal plate; hypandrium and associated structures as in
<figureCitation id="132C2A6B522DD435C30E2792FCD7F810" box="[781,876,1953,1977]" captionStart="FIGURES 110 118" captionStartId="20.[151,264,1635,1658]" captionTargetBox="[165,1427,193,1608]" captionTargetId="figure@20.[163,1428,193,1609]" captionTargetPageId="20" captionText="FIGURES 110 118. Fannia: male, hypandrium and associated structures, dorsal view: (110) F. canicularis; (111) F. chibcha sp. nov., paratype; (112) F. c h i n g a z a sp. nov., holotype; (113) F. dodgei; (114) F. dorsomaculata sp. nov., paratype; (115) F. embera sp. nov., paratype; (116) F. grandis; (117) F. iguaque sp. nov., paratype. Scale: 0.1 mm." httpUri="https://zenodo.org/record/213956/files/figure.png" pageId="28" pageNumber="29">Fig. 118</figureCitation>
.
</paragraph>
<paragraph id="8BA836EE522DD434C0C427F5FDEAFCF5" blockId="28.[151,1437,258,2014]" lastBlockId="29.[151,1437,151,1616]" lastPageId="29" lastPageNumber="30" pageId="28" pageNumber="29">
<emphasis id="B963EAFC522DD435C0C427F5FEA5F877" bold="true" box="[199,286,1990,2014]" pageId="28" pageNumber="29">Female</emphasis>
: similar to male, except: body length 2.9 3.1 mm; wing 22.4 mm. Dichoptic (
<figureCitation id="132C2A6B522DD435C4B727F6FB41F874" box="[1204,1274,1989,2013]" captionStart="FIGURES 1 12" captionStartId="8.[151,264,1845,1868]" captionTargetBox="[171,1399,339,1823]" captionTargetId="figure@8.[171,1417,339,1823]" captionTargetPageId="8" captionText="FIGURES 1 12. Fannia: (1) F. canicularis, male, head, anterior view; (2) F. chibcha sp. nov., male holotype, hind leg, anterior view; (3) F. dodgei, male, abdomen, posterior view; (4) F. dorsomaculata sp. nov., male holotype, lateral view; (5) F. dorsomaculata sp. nov., male holotype, dorsal view; (6) F. grandis sp. nov., male paratype, lateral view; (7) F. grandis, male, dorsal view; (8) F. iguaque sp. nov., male holotype, head, anterior view; (9) F. katios sp. nov., female paratype, head, anterior view; (10) F. katios sp. nov., male holotype, posterior view; (11) F. katios sp. nov., female paratype, posterior view; (12) F. lamosca sp. nov., male holotype, dorsal view. Scale: 1 mm." httpUri="https://zenodo.org/record/213947/files/figure.png" pageId="28" pageNumber="29">Fig. 9</figureCitation>
). Frontal vitta velvety brown up to ocellar triangle.
<emphasis id="B963EAFC522CD434C24620AAFDDAFF19" box="[581,609,153,176]" italics="true" pageId="29" pageNumber="30">Fr</emphasis>
8, converging. Upper
<emphasis id="B963EAFC522CD434C36420ABFC35FF19" box="[871,910,152,176]" italics="true" pageId="29" pageNumber="30">orb</emphasis>
reclinate and lower
<emphasis id="B963EAFC522CD434C47E20ABFB1FFF19" box="[1149,1188,152,176]" italics="true" pageId="29" pageNumber="30">orb</emphasis>
lateroclinate.
<emphasis id="B963EAFC522CD434C54A20AAFAC9FF19" box="[1353,1394,153,176]" italics="true" pageId="29" pageNumber="30">Fro</emphasis>
12 rows unordered along plate, or unordered starting at first
<emphasis id="B963EAFC522CD434C319208FFC97FF7D" box="[794,812,188,212]" italics="true" pageId="29" pageNumber="30">fr</emphasis>
and up to lower
<emphasis id="B963EAFC522CD434C3E7208FFBB0FF7D" box="[996,1035,188,212]" italics="true" pageId="29" pageNumber="30">orb</emphasis>
. Setulae on ocellar triangle varying in position and number.
<emphasis id="B963EAFC522CD434C1A420D2FE68FF51" box="[423,467,225,248]" italics="true" pageId="29" pageNumber="30">Poc</emphasis>
divergent. Fronto-orbital plate greyish pollinose. Gena grey-brownish. Scutum on
<emphasis id="B963EAFC522CD434C57520D2FA20FF51" box="[1398,1435,225,248]" italics="true" pageId="29" pageNumber="30">acr</emphasis>
and
<emphasis id="B963EAFC522CD434C0C52137FF5BFEB5" box="[198,224,260,284]" italics="true" pageId="29" pageNumber="30">dc</emphasis>
regions with weak pollinose vittae, some specimens with 3 differentiated and complete vittae.
<emphasis id="B963EAFC522CD434C5092137FA89FEB5" box="[1290,1330,260,284]" italics="true" pageId="29" pageNumber="30">Acr</emphasis>
34:67. One weakly differentiated
<emphasis id="B963EAFC522CD434C1C6211AFE57FEE9" box="[453,492,297,320]" italics="true" pageId="29" pageNumber="30">pra</emphasis>
, in some specimens with a seta next to it on one side. One pre-basal scutellar seta. Haltere whitish or yellowish with base yellow. Mid femur with 1 row of short and uniform
<emphasis id="B963EAFC522CD434C4BD217EFB63FECD" box="[1214,1240,333,356]" italics="true" pageId="29" pageNumber="30">av</emphasis>
; anterior surface setulose; 1 row of
<emphasis id="B963EAFC522CD434C1692143FE3DFE21" box="[362,390,368,392]" italics="true" pageId="29" pageNumber="30">ad</emphasis>
; 1 row of short
<emphasis id="B963EAFC522CD434C23C2142FDF6FE21" box="[575,589,369,392]" italics="true" pageId="29" pageNumber="30">p</emphasis>
without hooked apices and ending in 3 strong apical setae; 1 row of
<emphasis id="B963EAFC522CD434C55E2142FACCFE21" box="[1373,1399,369,392]" italics="true" pageId="29" pageNumber="30">pv</emphasis>
on basal half that decrease in length towards apex. Mid tibia on anterior surface without seta on edge of apical third, 2 apical
<emphasis id="B963EAFC522CD434C0E2218AFF54FE79" box="[225,239,441,464]" italics="true" pageId="29" pageNumber="30">a</emphasis>
, one long and one short, strong; 1
<emphasis id="B963EAFC522CD434C274218BFD28FE79" box="[631,659,440,464]" italics="true" pageId="29" pageNumber="30">ad</emphasis>
on border of basal third; 1
<emphasis id="B963EAFC522CD434C3CB218AFC59FE79" box="[968,994,441,464]" italics="true" pageId="29" pageNumber="30">av</emphasis>
on edge of apical third; 1 strong apical and 2 short pre-apical
<emphasis id="B963EAFC522CD434C19121EEFE25FE5D" box="[402,414,477,500]" italics="true" pageId="29" pageNumber="30">v</emphasis>
; 3 short apical
<emphasis id="B963EAFC522CD434C24F21EEFDE1FE5D" box="[588,602,477,500]" italics="true" pageId="29" pageNumber="30">p</emphasis>
.
<collectingCountry id="F300767E522CD434C26B21EFFD1BFE5D" box="[616,672,476,500]" name="India" pageId="29" pageNumber="30">Hind</collectingCountry>
femur with 1 row of short
<emphasis id="B963EAFC522CD434C3D321EEFC52FE5D" box="[976,1001,477,500]" italics="true" pageId="29" pageNumber="30">av</emphasis>
, 2 apical long and sub equal in length; dorsal surface bare; posterior surface without a row of setae; 1 row of short, not uniform, unordered
<emphasis id="B963EAFC522CD434C5072232FAA7FDB1" box="[1284,1308,513,536]" italics="true" pageId="29" pageNumber="30">pv</emphasis>
.
<collectingCountry id="F300767E522CD434C52921CCFAD9FDBE" box="[1322,1378,511,535]" name="India" pageId="29" pageNumber="30">Hind</collectingCountry>
tibia with 2 short apical
<emphasis id="B963EAFC522CD434C1752216FE3FFD95" box="[374,388,549,572]" italics="true" pageId="29" pageNumber="30">a</emphasis>
; 34
<emphasis id="B963EAFC522CD434C1CA2216FE5FFD95" box="[457,484,549,572]" italics="true" pageId="29" pageNumber="30">av</emphasis>
on basal third and 1 strong apical; 3
<emphasis id="B963EAFC522CD434C3952217FC09FD95" box="[918,946,548,572]" italics="true" pageId="29" pageNumber="30">ad</emphasis>
on basal third and 1 pre-apical; 1 median, long,and 1 shorter, pre-apical,
<emphasis id="B963EAFC522CD434C1ED227BFE47FDC9" box="[494,508,584,608]" italics="true" pageId="29" pageNumber="30">d</emphasis>
; 1 row of short
<emphasis id="B963EAFC522CD434C2B2227BFD76FDC9" box="[689,717,584,608]" italics="true" pageId="29" pageNumber="30">pd</emphasis>
that increase in length towards apex; posterior surface setulose, particularly on apical third; 1 apical
<emphasis id="B963EAFC522CD434C235225EFDEAFD2D" box="[566,593,621,644]" italics="true" pageId="29" pageNumber="30">pv</emphasis>
; 1 apical
<emphasis id="B963EAFC522CD434C2C0225EFD75FD2D" box="[707,718,621,644]" italics="true" pageId="29" pageNumber="30">v</emphasis>
. Abdomen trimaculate, with brown marks (
<figureCitation id="132C2A6B522CD434C4C3225FFAA8FD2D" box="[1216,1299,620,644]" captionStart="FIGURES 1 12" captionStartId="8.[151,264,1845,1868]" captionTargetBox="[171,1399,339,1823]" captionTargetId="figure@8.[171,1417,339,1823]" captionTargetPageId="8" captionText="FIGURES 1 12. Fannia: (1) F. canicularis, male, head, anterior view; (2) F. chibcha sp. nov., male holotype, hind leg, anterior view; (3) F. dodgei, male, abdomen, posterior view; (4) F. dorsomaculata sp. nov., male holotype, lateral view; (5) F. dorsomaculata sp. nov., male holotype, dorsal view; (6) F. grandis sp. nov., male paratype, lateral view; (7) F. grandis, male, dorsal view; (8) F. iguaque sp. nov., male holotype, head, anterior view; (9) F. katios sp. nov., female paratype, head, anterior view; (10) F. katios sp. nov., male holotype, posterior view; (11) F. katios sp. nov., female paratype, posterior view; (12) F. lamosca sp. nov., male holotype, dorsal view. Scale: 1 mm." httpUri="https://zenodo.org/record/213947/files/figure.png" pageId="29" pageNumber="30">Fig. 11</figureCitation>
). Syntergite 1+2 brown, translucent-yellow; tergites 34 translucent-yellow, tergite 5 brown. Syntergite 1+2 with conspicuous lateral setae; tergites 34 with marginal lateral setae visible, not as developed as in the male. Terminalia (Fig. 133): cercus shorter and narrower than hypoproct, with long and sinuous setae on apex; hypoproct round and with apex pointed, with long and weak setae; sternite 8 reduced to two membranous plates with short setae and two thin, bare, more sclerotized basal plates; sternites 6 and 7 longer than wide and sternite 7 with apex round. Spermathecae 3, shape irregular, one reduced (Fig. 133).
</paragraph>
</subSubSection>
<subSubSection id="C30D6565522CD434C0C42354FB59FC0D" pageId="29" pageNumber="30" type="biology_ecology">
<paragraph id="8BA836EE522CD434C0C42354FB59FC0D" blockId="29.[151,1437,151,1616]" pageId="29" pageNumber="30">
<emphasis id="B963EAFC522CD434C0C42354FE9CFC29" bold="true" box="[199,295,871,896]" pageId="29" pageNumber="30">Biology.</emphasis>
according to label data,
<taxonomicName id="4C174D6D522CD434C240235AFD64FC29" box="[579,735,872,896]" class="Insecta" family="Fanniidae" genus="Fannia" kingdom="Animalia" order="Diptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="katios" status="sp. nov.">
<emphasis id="B963EAFC522CD434C240235AFD64FC29" box="[579,735,872,896]" italics="true" pageId="29" pageNumber="30">Fannia katios</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2505787522CD434C2EB235BFCF8FC29" box="[744,835,872,896]" pageId="29" pageNumber="30" rank="species">
<emphasis id="B963EAFC522CD434C2EB235BFCB9FC29" bold="true" box="[744,770,872,896]" pageId="29" pageNumber="30">sp</emphasis>
.
<emphasis id="B963EAFC522CD434C311235BFC87FC29" bold="true" box="[786,828,872,896]" pageId="29" pageNumber="30">nov</emphasis>
.
</taxonomicNameLabel>
is associated with preserved areas of humid tropical forest, at Tapón del Darién, biogeographic province of Chocó (
<bibRefCitation id="EF864B1F522CD434C35623BFFC40FC0D" author="Morrone" box="[853,1019,908,932]" pageId="29" pageNumber="30" refString="Morrone, J. J. (2006) Biogeographic areas and transition zones of Latin America and the Caribbean Islands based on panbiogeographic and cladistic analyses of the entomofauna. Annual Review of Entomology, 51, 467 - 94." type="journal article" year="2006">Morrone 2006</bibRefCitation>
) limited by
<collectingCountry id="F300767E522CD434C48123BEFB65FC0D" box="[1154,1246,909,932]" name="Panama" pageId="29" pageNumber="30">Panama</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C30D6565522CD434C0C4239CFB26FB36" pageId="29" pageNumber="30" type="discussion">
<paragraph id="8BA836EE522CD434C0C4239CFB26FB36" blockId="29.[151,1437,151,1616]" pageId="29" pageNumber="30">
<emphasis id="B963EAFC522CD434C0C4239CFEF4FC61" bold="true" box="[199,335,943,968]" pageId="29" pageNumber="30">Comments.</emphasis>
these characters apply only to the male sex.
<taxonomicName id="4C174D6D522CD434C3422382FC67FC61" box="[833,988,944,968]" class="Insecta" family="Fanniidae" genus="Fannia" kingdom="Animalia" order="Diptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="katios" status="sp. nov.">
<emphasis id="B963EAFC522CD434C3422382FC67FC61" box="[833,988,944,968]" italics="true" pageId="29" pageNumber="30">Fannia katios</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2505787522CD434C3E12383FB83FC6E" box="[994,1080,943,968]" pageId="29" pageNumber="30" rank="species">
<emphasis id="B963EAFC522CD434C3E12383FC47FC61" bold="true" box="[994,1020,944,968]" pageId="29" pageNumber="30">sp</emphasis>
.
<emphasis id="B963EAFC522CD434C4092383FB8FFC61" bold="true" box="[1034,1076,944,968]" pageId="29" pageNumber="30">nov</emphasis>
.
</taxonomicNameLabel>
is similar to
<taxonomicName id="4C174D6D522CD434C4C62382FA27FC6E" authority="Chillcott" authorityName="Chillcott" box="[1221,1436,943,968]" class="Insecta" family="Fanniidae" genus="Fannia" kingdom="Animalia" order="Diptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="clavata">
<emphasis id="B963EAFC522CD434C4C62382FA8FFC61" box="[1221,1332,944,968]" italics="true" pageId="29" pageNumber="30">F. clavata</emphasis>
Chillcott
</taxonomicName>
and
<taxonomicName id="4C174D6D522CD434C0C523E6FE8CFC45" box="[198,311,980,1004]" class="Insecta" family="Fanniidae" genus="Fannia" kingdom="Animalia" order="Diptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="grandis">
<emphasis id="B963EAFC522CD434C0C523E6FE8CFC45" box="[198,311,980,1004]" italics="true" pageId="29" pageNumber="30">F. grandis</emphasis>
</taxonomicName>
(group
<taxonomicName id="4C174D6D522CD434C19323E7FE5DFC45" box="[400,486,980,1004]" class="Insecta" family="Fanniidae" genus="Fannia" kingdom="Animalia" order="Diptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="grandis">
<emphasis id="B963EAFC522CD434C19323E7FE5DFC45" box="[400,486,980,1004]" italics="true" pageId="29" pageNumber="30">grandis</emphasis>
</taxonomicName>
) in the shape of the genitalia, particularly the wide and concave cercal plate, the two apical median projections, and the spiral-shaped bacilliform process. It is similar to
<taxonomicName id="4C174D6D522CD434C44423CAFF63FB9D" authority="Albuquerque et al. 1981" authorityName="Albuquerque et al." authorityYear="1981" class="Insecta" family="Fanniidae" genus="Fannia" kingdom="Animalia" order="Diptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="grandis">
<emphasis id="B963EAFC522CD434C44423CAFB02FBB9" box="[1095,1209,1016,1040]" italics="true" pageId="29" pageNumber="30">F. grandis</emphasis>
(
<bibRefCitation id="EF864B1F522CD434C4CB23C4FF6BFB9D" author="Albuquerque" pageId="29" pageNumber="30" refString="Albuquerque, D. de O., Pamplona, D. &amp; de Carvalho, C. J. B. (1981) Contribuicao ao conhecimento dos Fannia R. D., 1830 da regiao Neotropical (Diptera, Fanniidae). Arquivo do Museu Nacional do Rio de Janeiro, 56, 9 - 34." type="journal article" year="1981">
Albuquerque
<emphasis id="B963EAFC522CD434C56723CAFA27FBB9" box="[1380,1436,1016,1040]" italics="true" pageId="29" pageNumber="30">et al.</emphasis>
1981
</bibRefCitation>
)
</taxonomicName>
in the shape of the hypandrium and associated structures.
<taxonomicName id="4C174D6D522CD434C36B242EFBB8FB9D" box="[872,1027,1052,1076]" class="Insecta" family="Fanniidae" genus="Fannia" kingdom="Animalia" order="Diptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="katios">
<emphasis id="B963EAFC522CD434C36B242EFBB8FB9D" box="[872,1027,1052,1076]" italics="true" pageId="29" pageNumber="30">Fannia katios</emphasis>
</taxonomicName>
also has characters of the
<emphasis id="B963EAFC522CD434C52F242FFA27FB9D" box="[1324,1436,1052,1076]" italics="true" pageId="29" pageNumber="30">benjamini</emphasis>
group, such as antenna and palpus yellowish, thorax with vittae, base of scutellum brown, hind coxa with setulae, abdomen yellowish with median vitta triangular and lateral marks on tergites (
<bibRefCitation id="EF864B1F522CD434C4272457FB75FBD5" author="Chillcott" box="[1060,1230,1124,1148]" pageId="29" pageNumber="30" refString="Chillcott, J. (1961) A Revision of the Neartic Species of Fanniinae (Diptera: Muscidae). Canadian Entomologist, 92, 1 - 295." type="journal article" year="1961">Chillcott 1961</bibRefCitation>
). Until a cladistic analysis is performed it is not possible to establish to which species group
<emphasis id="B963EAFC522CD434C3DA24BAFB8FFB09" box="[985,1076,1160,1184]" italics="true" pageId="29" pageNumber="30">F. k a t i o s</emphasis>
belongs.
</paragraph>
</subSubSection>
<subSubSection id="C30D6565522CD434C0C4249FFC09FA86" pageId="29" pageNumber="30" type="etymology">
<paragraph id="8BA836EE522CD434C0C4249FFC09FA86" blockId="29.[151,1437,151,1616]" pageId="29" pageNumber="30">
<emphasis id="B963EAFC522CD434C0C4249FFEF5FB6D" bold="true" box="[199,334,1196,1220]" pageId="29" pageNumber="30">Etymology.</emphasis>
name in apposition. The species epithet refers to the Parque Nacional Natural Los
<taxonomicName id="4C174D6D522CD434C50A249FFAEEFB6D" box="[1289,1365,1196,1220]" class="Insecta" family="Fanniidae" genus="Fannia" kingdom="Animalia" order="Diptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="katios">Katíos</taxonomicName>
, typelocality of the species. The park was designated a Natural World Heritage area in 1994 by UNESCO. This area is important because it is located in the biogeographic region of Chocó, on the border with
<collectingCountry id="F300767E522CD434C4A824C7FAA2FAA5" box="[1195,1305,1268,1292]" name="Colombia" pageId="29" pageNumber="30">Colombia</collectingCountry>
<collectingCountry id="F300767E522CD434C53F24C6FA23FAA5" box="[1340,1432,1269,1292]" name="Panama" pageId="29" pageNumber="30">Panama</collectingCountry>
, which favours faunal interchange between Central and South
<collectingCountry id="F300767E522CD434C34A2524FC16FA86" box="[841,941,1303,1327]" name="United States of America" pageId="29" pageNumber="30">America</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C30D6565522CD434C0C4250FFC4CF985" pageId="29" pageNumber="30" type="materials_examined">
<paragraph id="8BA836EE522CD434C0C4250FFC4CF985" blockId="29.[151,1437,151,1616]" pageId="29" pageNumber="30">
<emphasis id="B963EAFC522CD434C0C4250FFED6FAFD" bold="true" box="[199,365,1339,1364]" pageId="29" pageNumber="30">
<typeStatus id="54AC884C522CD434C0C4250FFEB9FAFD" box="[199,258,1340,1364]" pageId="29" pageNumber="30">Type</typeStatus>
material
</emphasis>
.
<emphasis id="B963EAFC522CD434C178250FFE53FAFD" bold="true" box="[379,488,1340,1364]" pageId="29" pageNumber="30">
<typeStatus id="54AC884C522CD434C178250FFE53FAFD" box="[379,488,1340,1364]" pageId="29" pageNumber="30" type="holotype">Holotype</typeStatus>
</emphasis>
male (IAvH). CO [
<collectingCountry id="F300767E522CD434C2BD250FFC8EFAFD" box="[702,821,1340,1364]" name="Colombia" pageId="29" pageNumber="30">Colombia</collectingCountry>
]. Chocó. P.N.N. [Parque Nacional Natural] Los
<taxonomicName id="4C174D6D522CD434C54F250FFA22FAFD" box="[1356,1433,1340,1364]" class="Insecta" family="Fanniidae" genus="Fannia" kingdom="Animalia" order="Diptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="katios">Katíos</taxonomicName>
. Centro\ administrativo Sautatá. Dentro Del\ Bosque.7˚51N; 77˚8W.
<specimenCount id="9D11FD67522CD434C3A3256CFC65FAD1" box="[928,990,1375,1400]" pageId="29" pageNumber="30" type="generic">
<quantity id="4CEF9B0B522CD434C3A3256CFC61FAD1" box="[928,986,1375,1400]" metricMagnitude="1" metricUnit="m" metricValue="3.0" pageId="29" pageNumber="30" unit="m" value="30.0">30 m</quantity>
.
</specimenCount>
[metros] \ Malaise 2.
<date id="FFA9102E522CD434C4CE256CFA90FADE" box="[1229,1323,1375,1399]" pageId="29" pageNumber="30" value="1913-05-29">29.V. 13</date>
.VI. 2000\ P. López. Leg. M. 3755 IAvH.
<emphasis id="B963EAFC522CD434C1F825B7FDD3FA35" bold="true" box="[507,616,1412,1436]" pageId="29" pageNumber="30">
<typeStatus id="54AC884C522CD434C1F825B7FDD3FA35" box="[507,616,1412,1436]" pageId="29" pageNumber="30" type="paratype">Paratype</typeStatus>
</emphasis>
: CO [
<collectingCountry id="F300767E522CD434C2AC25B7FC9CFA35" box="[687,807,1412,1436]" name="Colombia" pageId="29" pageNumber="30">Colombia</collectingCountry>
]. Chocó. P.N.N. [Parque Nacional Natural] Los
<taxonomicName id="4C174D6D522CD434C54F25B7FA22FA35" box="[1356,1433,1412,1436]" class="Insecta" family="Fanniidae" genus="Fannia" kingdom="Animalia" order="Diptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="katios">Katíos</taxonomicName>
. Centro\ administrativo Sautatá. Dentro del\ Bosque.7˚51N; 77˚8W.
<specimenCount id="9D11FD67522CD434C39F2594FC67FA16" box="[924,988,1447,1471]" pageId="29" pageNumber="30" type="generic">
<quantity id="4CEF9B0B522CD434C39F2594FC6CFA16" box="[924,983,1447,1471]" metricMagnitude="1" metricUnit="m" metricValue="3.0" pageId="29" pageNumber="30" unit="m" value="30.0">30 m</quantity>
.
</specimenCount>
[metros] \ Malaise 2.
<date id="FFA9102E522CD434C4D32594FA90FA16" box="[1232,1323,1447,1472]" pageId="29" pageNumber="30" value="1913-05-29">
29.
<emphasis id="B963EAFC522CD434C4F0259AFABBFA69" box="[1267,1280,1449,1472]" italics="true" pageId="29" pageNumber="30">V</emphasis>
. 13
</date>
.VI. 2000\ P. López. Leg. M. 3755 (
<specimenCount id="9D11FD67522CD434C1AE25FFFDB2FA4D" box="[429,521,1484,1508]" pageId="29" pageNumber="30" type="male">3 Males</specimenCount>
, IAvH); idem\ Fuera de\ bosque\ idem\ Malaise 3.
<date id="FFA9102E522CD434C43C25FFFB5CFA4D" box="[1087,1255,1484,1508]" pageId="29" pageNumber="30" value="2003-06-13" valueMax="2003-06-29" valueMin="2003-06-13">1329.VI.2003</date>
\ idem. M. 3763 (
<specimenCount id="9D11FD67522CD434C09C25DCFF46F9AE" box="[159,253,1519,1543]" pageId="29" pageNumber="30" type="male">4 Males</specimenCount>
, IAvH); idem\ Fuera del\ bosque\ Malaise 4.
<date id="FFA9102E522CD434C2F825DCFCE1F9AE" box="[763,858,1519,1544]" pageId="29" pageNumber="30" value="1913-05-29">
29.
<emphasis id="B963EAFC522CD434C31D25C2FC90F9A1" box="[798,811,1521,1544]" italics="true" pageId="29" pageNumber="30">V</emphasis>
. 13
</date>
. VI. 2003\ P. López. Leg. M. 3757 (
<specimenCount id="9D11FD67522CD434C4F625DCFAF9F9AE" box="[1269,1346,1519,1543]" pageId="29" pageNumber="30" type="male">1 male</specimenCount>
, IAvH). The
<typeStatus id="54AC884C522CD434C0CA2627FE90F985" box="[201,299,1556,1580]" pageId="29" pageNumber="30" type="holotype">holotype</typeStatus>
is missing the hind left leg and tibia and tarsus on its right side.
</paragraph>
</subSubSection>
<subSubSection id="C30D6565522CD434C0C42604FC35F9E6" box="[199,910,1591,1616]" pageId="29" pageNumber="30" type="distribution">
<paragraph id="8BA836EE522CD434C0C42604FC35F9E6" blockId="29.[151,1437,151,1616]" box="[199,910,1591,1616]" pageId="29" pageNumber="30">
<emphasis id="B963EAFC522CD434C0C42604FEDBF9F9" bold="true" box="[199,352,1591,1616]" pageId="29" pageNumber="30">Distribution.</emphasis>
<collectingCountry id="F300767E522CD434C1642604FE61F9E6" box="[359,474,1591,1615]" name="Colombia" pageId="29" pageNumber="30">Colombia</collectingCountry>
: department of Chocó (Pacific region).
</paragraph>
</subSubSection>
</treatment>
</document>