254 lines
37 KiB
XML
254 lines
37 KiB
XML
<document id="76E9EEA5EDCACE7EF928FB5D55BF96E7" ID-CLB-Dataset="27879" ID-DOI="10.11646/zootaxa.4526.4.1" ID-GBIF-Dataset="69cd7636-9ec1-4c3b-98ce-d6ce90bc7042" ID-ISSN="1175-5326" ID-Zenodo-Dep="2611755" ID-ZooBank="3CFFF3AC-C447-4FCE-B6F8-D2B7BAE8B678" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1553697705242" checkinUser="plazi" docAuthor="Valdés, Ángel, Lundsten, Lonny & Wilson, Nerida G." docDate="2018" docId="03BE87CCFFAFDE079B8E9FE67C175930" docLanguage="en" docName="zootaxa.4526.4.1.pdf" docOrigin="Zootaxa 4526 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Tritonia Cuvier 1797" docType="treatment" docVersion="9" lastPageNumber="410" masterDocId="FF87FFB4FFA6DE0E9B199F7E7D365F43" masterDocTitle="Five new deep-sea species of nudibranchs (Gastropoda: Heterobranchia: Cladobranchia) from the Northeast Pacific" masterLastPageNumber="433" masterPageNumber="401" pageNumber="410" updateTime="1698714326354" updateUser="ExternalLinkService">
|
||
<mods:mods id="D67ED3AD7DA5BBA6E096CE135BA2578B" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="8FB8CF6E00CC43C052C343E24357B7DC">
|
||
<mods:title id="567E0A463E85A0A9C8C4D48E4F704AE1">Five new deep-sea species of nudibranchs (Gastropoda: Heterobranchia: Cladobranchia) from the Northeast Pacific</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="FF5896A8D3BA32404255019458DD9E82" type="personal">
|
||
<mods:role id="99A4AC93C552134B2B924812093BD5D7">
|
||
<mods:roleTerm id="F371002220251021C9446CA641F58CD0">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="82B216768337163CF80F98B95E861D6D">Valdés, Ángel</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="BAAFC6B68E4E28208A146F05B1855731" type="personal">
|
||
<mods:role id="07E266F9B4031D66B5C2734F3541798A">
|
||
<mods:roleTerm id="07A57F3CDBD9FAB3D7ECDF93EA1DCA55">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="145E3ED4960E79F2426C473AC5F471C3">Lundsten, Lonny</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="EA185AE4601C6F30EFD236FA692C5D3A" type="personal">
|
||
<mods:role id="FAA96ED99E2F12CEBBFB56870746CAF6">
|
||
<mods:roleTerm id="03D071F27AFDDDFF30906F311210C88B">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="9D055745D4C592C00BAF21193FCE5E98">Wilson, Nerida G.</mods:namePart>
|
||
</mods:name>
|
||
<mods:typeOfResource id="BDCC77E1BB663CC3665CFE9975F35C2E">text</mods:typeOfResource>
|
||
<mods:relatedItem id="B0C8F8EEC8D374F3C7723BFEFB45B4D6" type="host">
|
||
<mods:titleInfo id="82F4F9E7CA4EF5907F271FC515872C29">
|
||
<mods:title id="C1A3F95FC636990D5E6AE5E6E0019A70">Zootaxa</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="453E88DE675332243CA8D6E6B9866DBD">
|
||
<mods:date id="D9E01AD03BE747CEC5BBE9BF21C62B60">2018</mods:date>
|
||
<mods:detail id="D2416F9103EEA41FB91865D43F149DCD" type="pubDate">
|
||
<mods:number id="F494E9087F58D1241A7BA0778EEF0B39">2018-12-03</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="BC45D66ECA2F1346D3A11FC674F04987" type="volume">
|
||
<mods:number id="9EC59C1D59DE2CD7EDDC22DD74ECC459">4526</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="DCC075991C1D3AC6D983CDA2E6779250" type="issue">
|
||
<mods:number id="6C80721D0D3746A6308FFCACD90A86D4">4</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="B5493668D89EBE7A00EE13FFB17CC5F2" unit="page">
|
||
<mods:start id="26ED0097BC1BDEF6F251F1B5C29AC025">401</mods:start>
|
||
<mods:end id="33FF653DF2F641358A8A8BBAC3F9A416">433</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:classification id="D8463F0D63EC28FD6AB4A631936844AA">journal article</mods:classification>
|
||
<mods:identifier id="A06654C929DCE5199EBD7E3E1E8A29F6" type="CLB-Dataset">27879</mods:identifier>
|
||
<mods:identifier id="1AD84B45D46D8F7300183B33AB8403D9" type="DOI">10.11646/zootaxa.4526.4.1</mods:identifier>
|
||
<mods:identifier id="B5B593B1EB23BEE03B9A7669F8C8FD61" type="GBIF-Dataset">69cd7636-9ec1-4c3b-98ce-d6ce90bc7042</mods:identifier>
|
||
<mods:identifier id="4FE32880529A48CB41DB3042D5648041" type="ISSN">1175-5326</mods:identifier>
|
||
<mods:identifier id="2F9C1596B0F2053DC88E1715E46BCFF4" type="Zenodo-Dep">2611755</mods:identifier>
|
||
<mods:identifier id="A37CFA97D38AA3E90ABA1B113751844D" type="ZooBank">3CFFF3AC-C447-4FCE-B6F8-D2B7BAE8B678</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="03BE87CCFFAFDE079B8E9FE67C175930" ID-DOI="http://doi.org/10.5281/zenodo.5971398" ID-GBIF-Taxon="154507774" ID-Zenodo-Dep="5971398" LSID="urn:lsid:plazi:treatment:03BE87CCFFAFDE079B8E9FE67C175930" httpUri="http://treatment.plazi.org/id/03BE87CCFFAFDE079B8E9FE67C175930" lastPageNumber="410" pageId="9" pageNumber="410">
|
||
<subSubSection id="C30D6551FFAFDE079B8E9FE67C125FF1" box="[151,292,152,178]" pageId="9" pageNumber="410" type="nomenclature">
|
||
<paragraph id="8BA836DAFFAFDE079B8E9FE67C125FF1" blockId="9.[151,401,152,211]" box="[151,292,152,178]" pageId="9" pageNumber="410">
|
||
<heading id="D0E081B6FFAFDE079B8E9FE67C125FF1" bold="true" box="[151,292,152,178]" fontSize="11" level="1" pageId="9" pageNumber="410" reason="1">
|
||
<emphasis id="B963EAC8FFAFDE079B8E9FE67C125FF1" bold="true" box="[151,292,152,178]" pageId="9" pageNumber="410">
|
||
<taxonomicName id="4C174D59FFAFDE079B8E9FE67DCE5FF1" ID-CoL="7PJZ6" authorityName="Cuvier" authorityYear="1797" box="[151,248,152,178]" class="Gastropoda" family="Tritoniidae" genus="Tritonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="9" pageNumber="410" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B963EAC8FFAFDE079B8E9FE67DCE5FF1" bold="true" box="[151,248,152,178]" italics="true" pageId="9" pageNumber="410">Tritonia</emphasis>
|
||
</taxonomicName>
|
||
sp.
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30D6551FFAFDE079B8E9FC47CA75F90" box="[151,401,186,211]" pageId="9" pageNumber="410" type="description">
|
||
<paragraph id="8BA836DAFFAFDE079B8E9FC47CA75F90" blockId="9.[151,401,152,211]" box="[151,401,186,211]" pageId="9" pageNumber="410">
|
||
(
|
||
<figureCitation id="132C2A5FFFAFDE079B869FC47C345F90" box="[159,258,186,211]" captionStart="FIGURE 2" captionStartId="4.[151,250,1838,1860]" captionTargetBox="[151,1436,193,1817]" captionTargetId="figure@4.[151,1436,193,1817]" captionTargetPageId="4" captionText="FIGURE 2. Digital images of live animals showing color in life. A. Tritonia nigritigris sp. nov., holotype (LACM 3553), in situ, right side invaginations indicated with arrows. B. Tritonia sp. (SIO-BIC M12395), lab photo. C. Dendronotus claguei sp. nov., holotype (LACM 3555), in situ. D. Ziminella vrijenhoeki sp. nov., holotype (SIO-BIC M12137), lab photo. E. Ziminella vrijenhoeki sp. nov., paratype (SIO-BIC M12135), lab photo. F. Cuthona methana sp. nov., holotype (SIO-BIC M12413), lab photo. G. Cuthona methana sp. nov., paratype (SIO-BIC M12412), lab photo. H. Aeolidia libitinaria sp. nov., holotype (SIO- BIC M12138), lab photo." httpUri="https://zenodo.org/record/2611759/files/figure.png" pageId="9" pageNumber="410">Figs. 2B</figureCitation>
|
||
,
|
||
<figureCitation id="132C2A5FFFAFDE079A149FC57C2F5F90" box="[269,281,187,211]" captionStart="FIGURE 3" captionStartId="5.[151,250,1211,1233]" captionTargetBox="[151,1437,193,1189]" captionTargetId="figure@5.[151,1437,193,1189]" captionTargetPageId="5" captionText="FIGURE 3. Photographs of the external morphology of preserved specimens of Tritonioidea and Dendronotoidea. A. Holotype of Tritonia nigritigris sp. nov. (LACM 3553), lateral view. B. Paratype of Dendronotus claguei sp. nov. (LACM 3555), lateral view. C–D. Tritonia sp. (SIO-BIC M12395), C. dorsal view of the head, D. lateral view. Abbreviations: a, anus; pe, penis. Arrow indicates damage on the right side of the body." httpUri="https://zenodo.org/record/2611761/files/figure.png" pageId="9" pageNumber="410">3</figureCitation>
|
||
C–D, 7–8)
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30D6551FFAFDE079B8E9E7C7E405E20" pageId="9" pageNumber="410" type="materials_examined">
|
||
<paragraph id="8BA836DAFFAFDE079B8E9E7C7E405E20" blockId="9.[151,1437,258,1651]" pageId="9" pageNumber="410">
|
||
<emphasis id="B963EAC8FFAFDE079B8E9E7C7CB65E58" bold="true" box="[151,384,258,283]" pageId="9" pageNumber="410">Material examined.</emphasis>
|
||
Hydrate Ridge,
|
||
<collectingRegion id="49D3F838FFAFDE07992E9E7D7FB85E58" box="[567,654,259,283]" country="United States of America" name="Oregon" pageId="9" pageNumber="410">Oregon</collectingRegion>
|
||
, Northeast Pacific Ocean (
|
||
<geoCoordinate id="EE23501DFFAFDE0798AF9E7C79295E59" box="[950,1055,258,283]" direction="north" orientation="latitude" pageId="9" pageNumber="410" precision="555" value="44.67">44.67° N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE23501DFFAFDE079F339E7C79835E58" box="[1066,1205,258,283]" direction="west" orientation="longitude" pageId="9" pageNumber="410" precision="55" value="-125.098">125.098° W</geoCoordinate>
|
||
),
|
||
<quantity id="4CEF9B3FFFAFDE079FD09E7C78395E59" box="[1225,1295,258,283]" metricMagnitude="2" metricUnit="m" metricValue="5.87" pageId="9" pageNumber="410" unit="m" value="587.0">587 m</quantity>
|
||
depth, ROV
|
||
<taxonomicName id="4C174D59FFAFDE079B8E9E597DE35E7D" authorityName="M. C. Miller" authorityYear="1974" box="[151,213,295,318]" class="Gastropoda" family="Facelinidae" genus="Jason" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="9" pageNumber="429" phylum="Mollusca" rank="genus">Jason</taxonomicName>
|
||
II (dive 593),
|
||
<date id="FFA9101AFFAFDE079A9C9E587F3A5E7C" box="[389,524,294,319]" pageId="9" pageNumber="410" value="2011-09-06">6 Sep 2011</date>
|
||
,
|
||
<specimenCount id="9D11FD53FFAFDE0799039E597FAA5E7C" box="[538,668,294,319]" pageId="9" pageNumber="410" type="generic">1 specimen</specimenCount>
|
||
<quantity id="4CEF9B3FFFAFDE0799BE9E597FCF5E7D" box="[679,761,295,319]" metricMagnitude="-2" metricUnit="m" metricValue="1.7" pageId="9" pageNumber="410" unit="mm" value="17.0">17 mm</quantity>
|
||
preserved length (SIO-BIC M12395), GenBank accession numbers:
|
||
<accessionNumber id="9444AB39FFAFDE079A1F9E347CB15E20" box="[262,391,330,355]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MH756139" pageId="9" pageNumber="410">MH756139</accessionNumber>
|
||
(
|
||
<emphasis id="B963EAC8FFAFDE079A8E9E357CF15E21" box="[407,455,331,354]" italics="true" pageId="9" pageNumber="410">COI</emphasis>
|
||
),
|
||
<accessionNumber id="9444AB39FFAFDE079AC69E347F565E20" box="[479,608,330,355]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MH756134" pageId="9" pageNumber="410">MH756134</accessionNumber>
|
||
(
|
||
<emphasis id="B963EAC8FFAFDE0799699E347FAC5E20" box="[624,666,330,355]" italics="true" pageId="9" pageNumber="410">16S</emphasis>
|
||
),
|
||
<materialsCitation id="3B7F3C87FFAFDE0799AB9E347E055E20" ID-GBIF-Occurrence="2029165978" box="[690,819,330,355]" collectionCode="MH" pageId="9" pageNumber="410" specimenCount="1">
|
||
<accessionNumber id="9444AB39FFAFDE0799AB9E347E055E20" box="[690,819,330,355]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MH756145" pageId="9" pageNumber="410">
|
||
<collectionCode id="ED06AE1FFFAFDE0799AB9E347FE95E20" box="[690,735,330,355]" country="Switzerland" name="Naturhistorisches Museum, Basel" pageId="9" pageNumber="410">MH</collectionCode>
|
||
756145
|
||
</accessionNumber>
|
||
</materialsCitation>
|
||
(
|
||
<emphasis id="B963EAC8FFAFDE07985A9E357E505E20" box="[835,870,331,355]" italics="true" pageId="9" pageNumber="410">H3</emphasis>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30D6551FFAFDE079BDE9E107EA65B70" pageId="9" pageNumber="410" type="description">
|
||
<paragraph id="8BA836DAFFAFDE079BDE9E1079255DAC" blockId="9.[151,1437,258,1651]" pageId="9" pageNumber="410">
|
||
<emphasis id="B963EAC8FFAFDE079BDE9E107C6F5EC4" bold="true" box="[199,345,366,391]" pageId="9" pageNumber="410">Description.</emphasis>
|
||
Body elongate, narrow, tapering towards the posterior end (
|
||
<figureCitation id="132C2A5FFFAFDE079F049E10794F5EC4" box="[1053,1145,366,391]" captionStart="FIGURE 2" captionStartId="4.[151,250,1838,1860]" captionTargetBox="[151,1436,193,1817]" captionTargetId="figure@4.[151,1436,193,1817]" captionTargetPageId="4" captionText="FIGURE 2. Digital images of live animals showing color in life. A. Tritonia nigritigris sp. nov., holotype (LACM 3553), in situ, right side invaginations indicated with arrows. B. Tritonia sp. (SIO-BIC M12395), lab photo. C. Dendronotus claguei sp. nov., holotype (LACM 3555), in situ. D. Ziminella vrijenhoeki sp. nov., holotype (SIO-BIC M12137), lab photo. E. Ziminella vrijenhoeki sp. nov., paratype (SIO-BIC M12135), lab photo. F. Cuthona methana sp. nov., holotype (SIO-BIC M12413), lab photo. G. Cuthona methana sp. nov., paratype (SIO-BIC M12412), lab photo. H. Aeolidia libitinaria sp. nov., holotype (SIO- BIC M12138), lab photo." httpUri="https://zenodo.org/record/2611759/files/figure.png" pageId="9" pageNumber="410">Fig. 2B</figureCitation>
|
||
). Lateral margins with 8 branched appendages (cerata), tripinnate, decreasing in size towards the posterior end (
|
||
<figureCitation id="132C2A5FFFAFDE079F699EEC79FA5EE8" box="[1136,1228,402,427]" captionStart="FIGURE 2" captionStartId="4.[151,250,1838,1860]" captionTargetBox="[151,1436,193,1817]" captionTargetId="figure@4.[151,1436,193,1817]" captionTargetPageId="4" captionText="FIGURE 2. Digital images of live animals showing color in life. A. Tritonia nigritigris sp. nov., holotype (LACM 3553), in situ, right side invaginations indicated with arrows. B. Tritonia sp. (SIO-BIC M12395), lab photo. C. Dendronotus claguei sp. nov., holotype (LACM 3555), in situ. D. Ziminella vrijenhoeki sp. nov., holotype (SIO-BIC M12137), lab photo. E. Ziminella vrijenhoeki sp. nov., paratype (SIO-BIC M12135), lab photo. F. Cuthona methana sp. nov., holotype (SIO-BIC M12413), lab photo. G. Cuthona methana sp. nov., paratype (SIO-BIC M12412), lab photo. H. Aeolidia libitinaria sp. nov., holotype (SIO- BIC M12138), lab photo." httpUri="https://zenodo.org/record/2611759/files/figure.png" pageId="9" pageNumber="410">Fig. 2A</figureCitation>
|
||
). Dorsum smooth, lacking tubercles or projections. Velum wide, triangular, with 14 conical, simple projections. Rhinophores elongate, narrow, each with a smooth, long stalk. Rhinophoral clubs with 5 tripinnate papillae, forming a circle closed posteriorly by a much more elongate papilla, covered with lamellae on its posterior end (
|
||
<figureCitation id="132C2A5FFFAFDE079FA79E8078225D54" box="[1214,1300,510,535]" captionStart="FIGURE 3" captionStartId="5.[151,250,1211,1233]" captionTargetBox="[151,1437,193,1189]" captionTargetId="figure@5.[151,1437,193,1189]" captionTargetPageId="5" captionText="FIGURE 3. Photographs of the external morphology of preserved specimens of Tritonioidea and Dendronotoidea. A. Holotype of Tritonia nigritigris sp. nov. (LACM 3553), lateral view. B. Paratype of Dendronotus claguei sp. nov. (LACM 3555), lateral view. C–D. Tritonia sp. (SIO-BIC M12395), C. dorsal view of the head, D. lateral view. Abbreviations: a, anus; pe, penis. Arrow indicates damage on the right side of the body." httpUri="https://zenodo.org/record/2611761/files/figure.png" pageId="9" pageNumber="410">Fig. 3C</figureCitation>
|
||
), located on anterior end of notum. Rhinophoral sheaths low, with smooth edges. Foot smooth, elongate posteriorly, visible dorsally at posterior end and in notum invaginations. Anal opening on right side of body (
|
||
<figureCitation id="132C2A5FFFAFDE079FBB9D3879C85D1C" box="[1186,1278,582,607]" captionStart="FIGURE 3" captionStartId="5.[151,250,1211,1233]" captionTargetBox="[151,1437,193,1189]" captionTargetId="figure@5.[151,1437,193,1189]" captionTargetPageId="5" captionText="FIGURE 3. Photographs of the external morphology of preserved specimens of Tritonioidea and Dendronotoidea. A. Holotype of Tritonia nigritigris sp. nov. (LACM 3553), lateral view. B. Paratype of Dendronotus claguei sp. nov. (LACM 3555), lateral view. C–D. Tritonia sp. (SIO-BIC M12395), C. dorsal view of the head, D. lateral view. Abbreviations: a, anus; pe, penis. Arrow indicates damage on the right side of the body." httpUri="https://zenodo.org/record/2611761/files/figure.png" pageId="9" pageNumber="410">Fig. 3D</figureCitation>
|
||
). Background color translucent white, with a concentration of orange pigment on the central dorsum. Body contractions concentrate the pigment, forming an irregular network of red lines. Notum edge translucent white with the lateral appendages (cerata) opaque white to cream. Velum and velar appendages translucent white. Rhinophores with translucent white stalks and beige sheath clubs. Foot uniform translucent white.
|
||
</paragraph>
|
||
<paragraph id="8BA836DAFFAFDE079BDE9D847FF05CA8" blockId="9.[151,1437,258,1651]" pageId="9" pageNumber="410">
|
||
Digestive system with large, muscular buccal bulb (
|
||
<figureCitation id="132C2A5FFFAFDE0798339D847E455C50" box="[810,883,762,787]" captionStart="FIGURE 7" captionStartId="10.[151,250,1948,1970]" captionTargetBox="[437,1113,226,1851]" captionTargetId="figure@10.[408,1162,193,1897]" captionTargetPageId="10" captionText="FIGURE 7. Drawing of the internal anatomy of a specimen of Tritonia sp. (SIO-BIC M12395). Abbreviations: bb, buccal bulb; cns, central nervous system; dg, digestive gland; es, esophagus; i, intestine; sg, salivary gland; st, stomach." httpUri="https://zenodo.org/record/2611769/files/figure.png" pageId="9" pageNumber="410">Fig. 7</figureCitation>
|
||
). Esophagus narrow, connecting anteriorly into buccal bulb, where two large, flat salivary glands connect. Intestine emerging dorsally from the stomach, forming a loop towards the right side of body where it opens into the anus (
|
||
<figureCitation id="132C2A5FFFAFDE0798939C3C7ED05C18" box="[906,998,834,859]" captionStart="FIGURE 3" captionStartId="5.[151,250,1211,1233]" captionTargetBox="[151,1437,193,1189]" captionTargetId="figure@5.[151,1437,193,1189]" captionTargetPageId="5" captionText="FIGURE 3. Photographs of the external morphology of preserved specimens of Tritonioidea and Dendronotoidea. A. Holotype of Tritonia nigritigris sp. nov. (LACM 3553), lateral view. B. Paratype of Dendronotus claguei sp. nov. (LACM 3555), lateral view. C–D. Tritonia sp. (SIO-BIC M12395), C. dorsal view of the head, D. lateral view. Abbreviations: a, anus; pe, penis. Arrow indicates damage on the right side of the body." httpUri="https://zenodo.org/record/2611761/files/figure.png" pageId="9" pageNumber="410">Fig. 3D</figureCitation>
|
||
). Radular formula 44 × 49.1.
|
||
<quantity id="4CEF9B3FFFAFDE079E2D9C3D78475C18" box="[1332,1393,834,859]" metricMagnitude="0" metricUnit="m" metricValue="1.2446" pageId="9" pageNumber="410" unit="in" value="49.0">49 in</quantity>
|
||
the single specimen examined. Rachidian teeth broad, with a short, wide central cusp, shorter than tooth, and 1–2 denticles or folds on each side of central cusp (
|
||
<figureCitation id="132C2A5FFFAFDE0799A79CF47E2D5CE0" box="[702,795,906,931]" captionStart="FIGURE 8" captionStartId="11.[151,250,1136,1158]" captionTargetBox="[151,1436,193,1114]" captionTargetId="figure@11.[151,1436,193,1114]" captionTargetPageId="11" captionText="FIGURE 8. SEMs of the radula and jaws of the single specimen of Tritonia sp. (SIO-BIC M12395). A. Rachidian and inner lateral radular teeth. B. Lateral radular teeth. C. Outer radular teeth. D. Jaw. E. Jaw masticatory border." httpUri="https://zenodo.org/record/2611771/files/figure.png" pageId="9" pageNumber="410">Fig. 8A</figureCitation>
|
||
). Lateral teeth hook-shaped, with sharp, elongate cusps lacking denticles (
|
||
<figureCitation id="132C2A5FFFAFDE079A7B9CD07C8C5C84" box="[354,442,942,967]" captionStart="FIGURE 8" captionStartId="11.[151,250,1136,1158]" captionTargetBox="[151,1436,193,1114]" captionTargetId="figure@11.[151,1436,193,1114]" captionTargetPageId="11" captionText="FIGURE 8. SEMs of the radula and jaws of the single specimen of Tritonia sp. (SIO-BIC M12395). A. Rachidian and inner lateral radular teeth. B. Lateral radular teeth. C. Outer radular teeth. D. Jaw. E. Jaw masticatory border." httpUri="https://zenodo.org/record/2611771/files/figure.png" pageId="9" pageNumber="410">Fig. 8B</figureCitation>
|
||
). Outer lateral teeth very elongate with reduced bases (
|
||
<figureCitation id="132C2A5FFFAFDE079F069CD079415C84" box="[1055,1143,942,967]" captionStart="FIGURE 8" captionStartId="11.[151,250,1136,1158]" captionTargetBox="[151,1436,193,1114]" captionTargetId="figure@11.[151,1436,193,1114]" captionTargetPageId="11" captionText="FIGURE 8. SEMs of the radula and jaws of the single specimen of Tritonia sp. (SIO-BIC M12395). A. Rachidian and inner lateral radular teeth. B. Lateral radular teeth. C. Outer radular teeth. D. Jaw. E. Jaw masticatory border." httpUri="https://zenodo.org/record/2611771/files/figure.png" pageId="9" pageNumber="410">Fig. 8C</figureCitation>
|
||
). Jaws elongate (
|
||
<figureCitation id="132C2A5FFFAFDE079E2F9CD078B85C84" box="[1334,1422,942,967]" captionStart="FIGURE 8" captionStartId="11.[151,250,1136,1158]" captionTargetBox="[151,1436,193,1114]" captionTargetId="figure@11.[151,1436,193,1114]" captionTargetPageId="11" captionText="FIGURE 8. SEMs of the radula and jaws of the single specimen of Tritonia sp. (SIO-BIC M12395). A. Rachidian and inner lateral radular teeth. B. Lateral radular teeth. C. Outer radular teeth. D. Jaw. E. Jaw masticatory border." httpUri="https://zenodo.org/record/2611771/files/figure.png" pageId="9" pageNumber="410">Fig. 8D</figureCitation>
|
||
), with a differentiated masticatory border (
|
||
<figureCitation id="132C2A5FFFAFDE07997B9CAC7F8C5CA8" box="[610,698,978,1003]" captionStart="FIGURE 8" captionStartId="11.[151,250,1136,1158]" captionTargetBox="[151,1436,193,1114]" captionTargetId="figure@11.[151,1436,193,1114]" captionTargetPageId="11" captionText="FIGURE 8. SEMs of the radula and jaws of the single specimen of Tritonia sp. (SIO-BIC M12395). A. Rachidian and inner lateral radular teeth. B. Lateral radular teeth. C. Outer radular teeth. D. Jaw. E. Jaw masticatory border." httpUri="https://zenodo.org/record/2611771/files/figure.png" pageId="9" pageNumber="410">Fig. 8E</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BA836DAFFAFDE079BDE9C887EA65B70" blockId="9.[151,1437,258,1651]" pageId="9" pageNumber="410">
|
||
Reproductive system absent, specimen was probably castrated as suggested by damage on the right side of the body visible in the live (
|
||
<figureCitation id="132C2A5FFFAFDE079ABE9B647CCB5B70" box="[423,509,1050,1075]" captionStart="FIGURE 2" captionStartId="4.[151,250,1838,1860]" captionTargetBox="[151,1436,193,1817]" captionTargetId="figure@4.[151,1436,193,1817]" captionTargetPageId="4" captionText="FIGURE 2. Digital images of live animals showing color in life. A. Tritonia nigritigris sp. nov., holotype (LACM 3553), in situ, right side invaginations indicated with arrows. B. Tritonia sp. (SIO-BIC M12395), lab photo. C. Dendronotus claguei sp. nov., holotype (LACM 3555), in situ. D. Ziminella vrijenhoeki sp. nov., holotype (SIO-BIC M12137), lab photo. E. Ziminella vrijenhoeki sp. nov., paratype (SIO-BIC M12135), lab photo. F. Cuthona methana sp. nov., holotype (SIO-BIC M12413), lab photo. G. Cuthona methana sp. nov., paratype (SIO-BIC M12412), lab photo. H. Aeolidia libitinaria sp. nov., holotype (SIO- BIC M12138), lab photo." httpUri="https://zenodo.org/record/2611759/files/figure.png" pageId="9" pageNumber="410">Fig. 2B</figureCitation>
|
||
) and preserved (
|
||
<figureCitation id="132C2A5FFFAFDE0799A19B647E395B70" box="[696,783,1050,1075]" captionStart="FIGURE 3" captionStartId="5.[151,250,1211,1233]" captionTargetBox="[151,1437,193,1189]" captionTargetId="figure@5.[151,1437,193,1189]" captionTargetPageId="5" captionText="FIGURE 3. Photographs of the external morphology of preserved specimens of Tritonioidea and Dendronotoidea. A. Holotype of Tritonia nigritigris sp. nov. (LACM 3553), lateral view. B. Paratype of Dendronotus claguei sp. nov. (LACM 3555), lateral view. C–D. Tritonia sp. (SIO-BIC M12395), C. dorsal view of the head, D. lateral view. Abbreviations: a, anus; pe, penis. Arrow indicates damage on the right side of the body." httpUri="https://zenodo.org/record/2611761/files/figure.png" pageId="9" pageNumber="410">Fig. 3D</figureCitation>
|
||
) specimen.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30D6551FFAFDE079BDE9B4078AA5A48" pageId="9" pageNumber="410" type="biology_ecology">
|
||
<paragraph id="8BA836DAFFAFDE079BDE9B4078AA5B80" blockId="9.[151,1437,258,1651]" pageId="9" pageNumber="410">
|
||
<emphasis id="B963EAC8FFAFDE079BDE9B407C115B14" bold="true" box="[199,295,1086,1111]" pageId="9" pageNumber="410">Biology.</emphasis>
|
||
This species was collected at
|
||
<quantity id="4CEF9B3FFFAFDE0799619B407FF75B15" box="[632,705,1086,1111]" metricMagnitude="2" metricUnit="m" metricValue="5.87" pageId="9" pageNumber="410" unit="m" value="587.0">587 m</quantity>
|
||
, from Hydrate Ridge, an area of methane seeps about 44 nautical miles off
|
||
<collectingRegion id="49D3F838FFAFDE079A1F9B1D7C5D5B39" box="[262,363,1123,1146]" country="United Kingdom" name="Newport" pageId="9" pageNumber="410">Newport</collectingRegion>
|
||
,
|
||
<collectingRegion id="49D3F838FFAFDE079A619B1D7CFA5B38" box="[376,460,1123,1147]" country="United States of America" name="Oregon" pageId="9" pageNumber="410">Oregon</collectingRegion>
|
||
(
|
||
<figureCitation id="132C2A5FFFAFDE079AC49B1C7F175B38" box="[477,545,1122,1147]" captionStart="FIGURE 1" captionStartId="2.[151,250,1630,1652]" captionTargetBox="[151,1436,195,1606]" captionTargetId="figure@2.[151,1436,195,1606]" captionTargetPageId="2" captionText="FIGURE 1. Distribution map of the collection localities in this paper including depth ranges." httpUri="https://zenodo.org/record/2611757/files/figure.png" pageId="9" pageNumber="410">Fig. 1</figureCitation>
|
||
). It came from an inactive, experimental rock (Levin Lab E12, rock F), upon which an unidentified soft coral (SIO-BIC Co2330) was growing. Other organisms collected from the soft coral were an unidentified amphipod (SIO-BIC C11265)
|
||
<materialsCitation id="3B7F3C87FFAFDE0799CC9BD478AE5B80" ID-GBIF-Occurrence="2029167908" box="[725,1432,1194,1219]" collectionCode="SIO-BIC" pageId="9" pageNumber="410" specimenCode="A2679" specimenCount="1">
|
||
and the polychaete worm
|
||
<taxonomicName id="4C174D59FFAFDE0798EC9BD479F25B80" box="[1013,1220,1194,1219]" class="Polychaeta" family="Syllidae" genus="Eusyllis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phyllodocida" pageId="9" pageNumber="410" phylum="Annelida" rank="species" species="nuchalata">
|
||
<emphasis id="B963EAC8FFAFDE0798EC9BD479F25B80" box="[1013,1220,1194,1219]" italics="true" pageId="9" pageNumber="410">Eusyllis nuchalata</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<collectionCode id="ED06AE1FFFAFDE079FCB9BD5780F5B80" box="[1234,1337,1195,1219]" pageId="9" pageNumber="410">SIO-BIC</collectionCode>
|
||
<specimenCode id="DBB19EA1FFAFDE079E269BD478A65B80" box="[1343,1424,1194,1219]" collectionCode="SIO-BIC" pageId="9" pageNumber="410">A2679</specimenCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BA836DAFFAFDE079BDE9BB078AA5A48" blockId="9.[151,1437,258,1651]" pageId="9" pageNumber="410">
|
||
<emphasis id="B963EAC8FFAFDE079BDE9BB07CFA5BA4" bold="true" box="[199,460,1230,1255]" pageId="9" pageNumber="410">Phylogenetic position.</emphasis>
|
||
<taxonomicName id="4C174D59FFAFDE079ACA9BB17F1B5BA5" authorityName="Cuvier" authorityYear="1797" box="[467,557,1231,1254]" class="Gastropoda" family="Tritoniidae" genus="Tritonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="9" pageNumber="410" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B963EAC8FFAFDE079ACA9BB17F1B5BA5" box="[467,557,1231,1254]" italics="true" pageId="9" pageNumber="410">Tritonia</emphasis>
|
||
</taxonomicName>
|
||
sp. is placed in a clade including Eastern Pacific species of
|
||
<taxonomicName id="4C174D59FFAFDE079FD49BB1781E5BA5" authorityName="Cuvier" authorityYear="1797" box="[1229,1320,1231,1254]" class="Gastropoda" family="Tritoniidae" genus="Tritonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="9" pageNumber="410" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B963EAC8FFAFDE079FD49BB1781E5BA5" box="[1229,1320,1231,1254]" italics="true" pageId="9" pageNumber="410">Tritonia</emphasis>
|
||
</taxonomicName>
|
||
such as
|
||
<taxonomicName id="4C174D59FFAFDE079E9E9BB17CA65A48" authority="(Stearns, 1873)" baseAuthorityName="Stearns" baseAuthorityYear="1873" class="Gastropoda" family="Tritoniidae" genus="Tritonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="9" pageNumber="410" phylum="Mollusca" rank="species" species="festiva">
|
||
<emphasis id="B963EAC8FFAFDE079E9E9BB17DD75A48" italics="true" pageId="9" pageNumber="410">T. festiva</emphasis>
|
||
(Stearns, 1873)
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C174D59FFAFDE079ADF9B8A7E7F5A48" authority="(Pallas, 1788)" baseAuthorityName="Pallas" baseAuthorityYear="1788" box="[454,841,1266,1291]" class="Gastropoda" family="Tritoniidae" genus="Tritonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="9" pageNumber="410" phylum="Mollusca" rank="species" species="tetraquetra">
|
||
<emphasis id="B963EAC8FFAFDE079ADF9B8A7F935A48" box="[454,677,1268,1291]" italics="true" pageId="9" pageNumber="410">Tritonia tetraquetra</emphasis>
|
||
(
|
||
<bibRefCitation id="EF864B2BFFAFDE0799AD9B8C7E775A48" author="Pallas, P. S." box="[692,833,1266,1291]" pageId="9" pageNumber="410" pagination="229 - 249" refId="ref15780" refString="Pallas, P. S. (1788) Marina varia nova et rariora. Nova Acta Academiae Scientiarum Imperialis Petropolitanea, 2, 229 - 249." type="journal article" year="1788">Pallas, 1788</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, as well as two other unidentified specimens (
|
||
<figureCitation id="132C2A5FFFAFDE079E539B8C78B85A48" box="[1354,1422,1266,1291]" captionStart="FIGURE 6" captionStartId="8.[151,250,1949,1971]" captionTargetBox="[391,1173,195,1901]" captionTargetId="figure@8.[388,1173,195,1901]" captionTargetPageId="8" captionText="FIGURE 6. Bayesian consensus phylogenetic analyses of Tritonioidea and Dendronotoidea, with branches collapsed for clarity. The phylogenetic position of the species described herein (in bold) are represented in grey ovals. Posterior probability and maximum likelihood bootstrap values indicated for each branch." httpUri="https://zenodo.org/record/2611767/files/figure.png" pageId="9" pageNumber="410">Fig. 6</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30D6551FFAFDE079BDE9A687C175930" pageId="9" pageNumber="410" type="discussion">
|
||
<paragraph id="8BA836DAFFAFDE079BDE9A687E945A34" blockId="9.[151,1437,258,1651]" pageId="9" pageNumber="410">
|
||
<emphasis id="B963EAC8FFAFDE079BDE9A687C0D5A6C" bold="true" box="[199,315,1302,1327]" pageId="9" pageNumber="410">Remarks.</emphasis>
|
||
<taxonomicName id="4C174D59FFAFDE079A5A9A697CAB5A6D" authorityName="Cuvier" authorityYear="1797" box="[323,413,1303,1326]" class="Gastropoda" family="Tritoniidae" genus="Tritonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="9" pageNumber="410" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B963EAC8FFAFDE079A5A9A697CAB5A6D" box="[323,413,1303,1326]" italics="true" pageId="9" pageNumber="410">Tritonia</emphasis>
|
||
</taxonomicName>
|
||
sp. appears to be different from other species of
|
||
<taxonomicName id="4C174D59FFAFDE0798C89A69791D5A6D" authorityName="Cuvier" authorityYear="1797" box="[977,1067,1303,1326]" class="Gastropoda" family="Tritoniidae" genus="Tritonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="9" pageNumber="410" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B963EAC8FFAFDE0798C89A69791D5A6D" box="[977,1067,1303,1326]" italics="true" pageId="9" pageNumber="410">Tritonia</emphasis>
|
||
</taxonomicName>
|
||
described to date. However, the single specimen examined lacked a reproductive system. Damage on the right side of the body may suggest the animal was either harmed during collection or castrated by a parasite.
|
||
</paragraph>
|
||
<paragraph id="8BA836DAFFAFDE079BDE9AFC7C175930" blockId="9.[151,1437,258,1651]" pageId="9" pageNumber="410">
|
||
The external morphology of
|
||
<taxonomicName id="4C174D59FFAFDE0799159AFA7F505AD8" authorityName="Cuvier" authorityYear="1797" box="[524,614,1412,1435]" class="Gastropoda" family="Tritoniidae" genus="Tritonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="9" pageNumber="410" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B963EAC8FFAFDE0799159AFA7F505AD8" box="[524,614,1412,1435]" italics="true" pageId="9" pageNumber="410">Tritonia</emphasis>
|
||
</taxonomicName>
|
||
sp. resembles that of
|
||
<taxonomicName id="4C174D59FFAFDE0798469AFA7DE45AFC" authority="Bertsch & Mozqueira Osuna, 1986" authorityName="Bertsch & Mozqueira Osuna" authorityYear="1986" class="Gastropoda" family="Tritoniidae" genus="Tritonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="9" pageNumber="410" phylum="Mollusca" rank="species" species="myrakeenae">
|
||
<emphasis id="B963EAC8FFAFDE0798469AFA797E5AD8" box="[863,1096,1410,1435]" italics="true" pageId="9" pageNumber="410">Tritonia myrakeenae</emphasis>
|
||
<bibRefCitation id="EF864B2BFFAFDE079F489AFC7DE45AFC" author="Bertsch, H. & Mozqueira Osuna, A." pageId="9" pageNumber="410" pagination="46 - 49" refId="ref13431" refString="Bertsch, H. & Mozqueira Osuna, A. (1986) A new species of Tritonia (Nudibranchia) from southern California and Baja California. The Nautilus, 100, 46 - 49. https: // doi. org / 10.5962 / bhl. part. 26492" type="journal article" year="1986">Bertsch & Mozqueira Osuna, 1986</bibRefCitation>
|
||
</taxonomicName>
|
||
, also found in the Eastern Pacific, but the latter lacks red pigment on the dorsum and often has two white pigment patches posterior to the head (
|
||
<bibRefCitation id="EF864B2BFFAFDE0799549AB47EEB5AA0" author="Bertsch, H. & Mozqueira Osuna, A." box="[589,989,1482,1507]" pageId="9" pageNumber="410" pagination="46 - 49" refId="ref13431" refString="Bertsch, H. & Mozqueira Osuna, A. (1986) A new species of Tritonia (Nudibranchia) from southern California and Baja California. The Nautilus, 100, 46 - 49. https: // doi. org / 10.5962 / bhl. part. 26492" type="journal article" year="1986">Bertsch & Mozqueira Osuna, 1986</bibRefCitation>
|
||
), absent in
|
||
<taxonomicName id="4C174D59FFAFDE079F479AB2798E5AA0" authorityName="Cuvier" authorityYear="1797" box="[1118,1208,1484,1507]" class="Gastropoda" family="Tritoniidae" genus="Tritonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="9" pageNumber="410" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B963EAC8FFAFDE079F479AB2798E5AA0" box="[1118,1208,1484,1507]" italics="true" pageId="9" pageNumber="410">Tritonia</emphasis>
|
||
</taxonomicName>
|
||
sp. The radula of
|
||
<taxonomicName id="4C174D59FFAFDE079E9E9AB27C285944" authorityName="Bertsch & Mozqueira Osuna" authorityYear="1986" class="Gastropoda" family="Tritoniidae" genus="Tritonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="9" pageNumber="410" phylum="Mollusca" rank="species" species="myrakeenae">
|
||
<emphasis id="B963EAC8FFAFDE079E9E9AB27C285944" italics="true" pageId="9" pageNumber="410">T. myrakeenae</emphasis>
|
||
</taxonomicName>
|
||
is also different from that of
|
||
<taxonomicName id="4C174D59FFAFDE07996E9A917FE45945" authorityName="Cuvier" authorityYear="1797" box="[631,722,1519,1542]" class="Gastropoda" family="Tritoniidae" genus="Tritonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="9" pageNumber="410" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B963EAC8FFAFDE07996E9A917FE45945" box="[631,722,1519,1542]" italics="true" pageId="9" pageNumber="410">Tritonia</emphasis>
|
||
</taxonomicName>
|
||
sp. The former has a rachidian tooth with an elongate central cusp and two strong lateral denticles (
|
||
<bibRefCitation id="EF864B2BFFAFDE079922996C7EF05968" author="Bertsch, H. & Mozqueira Osuna, A." box="[571,966,1554,1579]" pageId="9" pageNumber="410" pagination="46 - 49" refId="ref13431" refString="Bertsch, H. & Mozqueira Osuna, A. (1986) A new species of Tritonia (Nudibranchia) from southern California and Baja California. The Nautilus, 100, 46 - 49. https: // doi. org / 10.5962 / bhl. part. 26492" type="journal article" year="1986">Bertsch & Mozqueira Osuna, 1986</bibRefCitation>
|
||
), whereas
|
||
<taxonomicName id="4C174D59FFAFDE079F22996A79A35968" authorityName="Cuvier" authorityYear="1797" box="[1083,1173,1556,1579]" class="Gastropoda" family="Tritoniidae" genus="Tritonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="9" pageNumber="410" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B963EAC8FFAFDE079F22996A79A35968" box="[1083,1173,1556,1579]" italics="true" pageId="9" pageNumber="410">Tritonia</emphasis>
|
||
</taxonomicName>
|
||
sp. has a much broader rachidian with a shorter central cusp. Unfortunately, no molecular data for
|
||
<taxonomicName id="4C174D59FFAFDE079F37994979EE590C" authorityName="Bertsch & Mozqueira Osuna" authorityYear="1986" box="[1070,1240,1590,1615]" class="Gastropoda" family="Tritoniidae" genus="Tritonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="9" pageNumber="410" phylum="Mollusca" rank="species" species="myrakeenae">
|
||
<emphasis id="B963EAC8FFAFDE079F37994979EE590C" box="[1070,1240,1590,1615]" italics="true" pageId="9" pageNumber="410">T. myrakeenae</emphasis>
|
||
</taxonomicName>
|
||
is available for comparison.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |