478 lines
85 KiB
XML
478 lines
85 KiB
XML
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<mods:title id="5A620C30D9D3581F01780B7669CE04BA">Preliminary re-examination of genus-level taxonomy of the pollen beetle subfamily Meligethinae (Coleoptera: Nitidulidae)</mods:title>
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<mods:namePart id="4117813B44AE1D762A2F3AEA46DAD58C">Audisio, Paolo</mods:namePart>
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<mods:affiliation id="EE68ECA3736A42F385FD18314CCE5B5E">Dipartimento di Biologia Animale e dell’Uomo, Sapienza Rome University, via A. Borelli, 50, I- 00161 Rome, Italy; e-mail: paolo. audisio @ uniroma 1. it</mods:affiliation>
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<mods:namePart id="F62FFA359906457E61059AA6DB39E408">Cline, Andrew Richard</mods:namePart>
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<mods:affiliation id="6B7ABE798CEE9C5F30192EC727F76464">Dipartimento di Biologia Animale e dell’Uomo, Sapienza Rome University, viale Dell’Università, 32, I- 00185 Rome, Italy; e-mail: alessio. debiase @ uniroma 1. it</mods:affiliation>
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<mods:namePart id="7E72E4AD047EB2679E7ED470F1BAC87A">Mancini, Emiliano</mods:namePart>
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<mods:affiliation id="D7B1232AE3E5D961EBDE363A942CC5F7">Dipartimento di Scienze di Sanità Pubblica, Sapienza Rome University, Piazzale Aldo Moro 5, I- 00185, Rome, Italy; e-mail: emiliano. mancini @ uniroma 1. it</mods:affiliation>
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<mods:namePart id="6A2D6900979B27C35921772189BFE4E3">Trizzino, Marco</mods:namePart>
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<mods:affiliation id="41468C48355D32E481BFD3A099F51064">Dipartimento di Biologia Animale e dell’Uomo, Sapienza Rome University, via A. Borelli, 50, I- 00161 Rome, Italy; e-mail: paolo. audisio @ uniroma 1. it</mods:affiliation>
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<mods:namePart id="29270EAD8F0F4EC8731C5F729FDEE13C">Costantini, Lorenzo</mods:namePart>
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<mods:affiliation id="F49536B61294A820DD33026EEA17FCFD">Museo Nazionale d’Arte Orientale, Servizio di Bioarcheologia e Microscopia, via Merulana 248, I- 00185 Rome, Italy; e-mail: l. costantin @ mclink. it</mods:affiliation>
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<mods:namePart id="25027EFF6763CB5A7C156DB5992DD45F">Strika, Sirio</mods:namePart>
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<mods:affiliation id="674F16D6AB8D6197BBFF874CD6F8566F">Museo Nazionale d’Arte Orientale, Servizio di Bioarcheologia e Microscopia, via Merulana 248, I- 00185 Rome, Italy; e-mail: l. costantin @ mclink. it</mods:affiliation>
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<mods:namePart id="0984B13A4290A3E3629609B4D02979D8">Lamanna, Francesco</mods:namePart>
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<mods:affiliation id="B68ED1C0B11B95B7047F3248E5E6830D">Dipartimento di Biologia Animale e dell’Uomo, Sapienza Rome University, via A. Borelli, 50, I- 00161 Rome, Italy; e-mail: paolo. audisio @ uniroma 1. it</mods:affiliation>
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<mods:namePart id="3A3F2266F698B63746430AE106ADD738">Cerretti, Pierfilippo</mods:namePart>
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<mods:affiliation id="9643C094C44997BE24F6C5690B599610">Dipartimento di Biologia Animale e dell’Uomo, Sapienza Rome University, via A. Borelli, 50, I- 00161 Rome, Italy; e-mail: paolo. audisio @ uniroma 1. it</mods:affiliation>
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<mods:title id="300682342F4AFF8C6FD4C85939C93C55">Acta Entomologica Musei Nationalis Pragae</mods:title>
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<mods:date id="32885C5B226A7F2F087F50429BA44D50">2009</mods:date>
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<mods:number id="D83DBD2E582224CB621C6B0912131CA0">2009-12-15</mods:number>
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<mods:number id="72354FFA580DB6CE744E4FC9729877AC">49</mods:number>
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4.
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<taxonomicName id="4C174D59F67AFF94BACBFD0BFD5EFCAC" authority="Kirejtshuk, 1989" authorityName="Kirejtshuk" authorityYear="1989" box="[448,747,767,791]" class="Insecta" family="Nitidulidae" genus="Lariopsis" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="357" phylum="Arthropoda" rank="genus">
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<emphasis id="B963EAC8F67AFF94BACBFD0BFD93FCAC" bold="true" box="[448,550,767,791]" italics="true" pageId="16" pageNumber="357">Lariopsis</emphasis>
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<bibRefCitation id="EF864B2BF67AFF94B926FD0BFD5EFCAC" author="KIREJTSHUK A. G." box="[557,747,767,791]" pageId="16" pageNumber="357" pagination="64 - 89" refId="ref89287" refString="KIREJTSHUK A. G. 1989: Novye taksony zhukov-blestyanok (Coleoptera, Nitidulidae) vostochnogo polushariya (Chast' III). [New taxa of the Nitidulidae (Coleoptera) of the East hemisphere (Part III)]. Trudy Zoologicheskogo Instituta, Akademiya Nauk SSSR 208: 64 - 89 (in Russian)." type="journal article" year="1989">Kirejtshuk, 1989</bibRefCitation>
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stat. nov.
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(
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<figureCitation id="132C2A5FF67AFF94B944FCE9FD1AFC8B" box="[591,687,797,817]" captionStart="Fig" captionStartId="18.[151,182,1492,1512]" captionTargetBox="[168,1109,431,1469]" captionTargetId="figure-10@18.[167,1109,431,1470]" captionTargetPageId="18" captionText="Fig. 4. Lariopsis Kirejtshuk, 1989: a – L.vultuosus (Kirejtshuk & Audisio, 1995); b–k – L. variabilis (Reitter, 1872). a – male habitus (length 3.3 mm); b, c – dorso-lateral view of head; d – ventral view of head and anterior portion of prosternum; e – prosternal process and mesoventrite; f – middle leg with outer margin of mesotibia; g – microsetae on middle posterior margin of pronotum; h – exposed portion of last visible abdominal ventrite; k – caudal marginal lines of metacoxal cavities. Scale bars: Figs. b, c, d, e, h, k = 100 μm; Fig. f = 30 μm; Fig. g = 10 μm." figureDoi="http://doi.org/10.5281/zenodo.5319344" httpUri="https://zenodo.org/record/5319344/files/figure.png" pageId="16" pageNumber="357">Figs. 4 a–k</figureCitation>
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)
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</paragraph>
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<paragraph id="8BA836DAF67AFF94BB9CFCB5FC30FCE8" blockId="16.[151,901,797,851]" box="[151,901,832,851]" pageId="16" pageNumber="357">
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<treatmentCitationGroup id="AB0711F4F67AFF94BB9CFCB5FC30FCE8" box="[151,901,832,851]" pageId="16" pageNumber="357">
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<treatmentCitation id="0AB610CBF67AFF94BB9CFCB5FE28FCE8" author="KIREJTSHUK A. G." box="[151,413,832,851]" page="86" pageId="16" pageNumber="357" year="1989">
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<taxonomicName id="4C174D59F67AFF94BB9CFCB5FE28FCE8" authority="Kirejtshuk, 1989: 86" authorityName="Kirejtshuk" authorityPageNumber="86" authorityYear="1989" box="[151,413,832,851]" class="Insecta" family="Nitidulidae" genus="Lariopsis" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="357" phylum="Arthropoda" rank="genus">
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<emphasis id="B963EAC8F67AFF94BB9CFCB5FF5DFCE8" box="[151,232,833,851]" italics="true" pageId="16" pageNumber="357">Lariopsis</emphasis>
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<bibRefCitation id="EF864B2BF67AFF94BBE6FCB4FE28FCE8" author="KIREJTSHUK A. G." box="[237,413,832,851]" pageId="16" pageNumber="357" pagination="64 - 89" refId="ref89287" refString="KIREJTSHUK A. G. 1989: Novye taksony zhukov-blestyanok (Coleoptera, Nitidulidae) vostochnogo polushariya (Chast' III). [New taxa of the Nitidulidae (Coleoptera) of the East hemisphere (Part III)]. Trudy Zoologicheskogo Instituta, Akademiya Nauk SSSR 208: 64 - 89 (in Russian)." type="journal article" year="1989">Kirejtshuk, 1989: 86</bibRefCitation>
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</taxonomicName>
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(described as a subgenus of
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<taxonomicName id="4C174D59F67AFF94B999FCB5FCCEFCE8" ID-CoL="5NQ7" authority="Stephens, 1830" authorityName="Stephens" authorityYear="1830" box="[658,891,832,851]" class="Insecta" family="Nitidulidae" genus="Meligethes" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="357" phylum="Arthropoda" rank="genus">
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<emphasis id="B963EAC8F67AFF94B999FCB5FD45FCE8" box="[658,752,833,851]" italics="true" pageId="16" pageNumber="357">Meligethes</emphasis>
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<bibRefCitation id="EF864B2BF67AFF94B9FEFCB4FCCEFCE8" author="STEPHENS J. F." box="[757,891,832,851]" pageId="16" pageNumber="357" pagination="16 - 19" refId="ref92184" refString="STEPHENS J. F. 1830: Illustrations of British Entomology; or, a Synopsis of Indigenous Insects: containing their generic and specific distinctions; with an account of their metamorphoses, times of appearance, localities, food, and economy, as far as practicable. Mandibulata, vol. III. Baldwin and Craddock, London, 379 pp, pls. 16 - 19." type="book chapter" year="1830">Stephens, 1830</bibRefCitation>
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</taxonomicName>
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).
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</paragraph>
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<subSubSection id="C30D6551F67AFF96BB9CFC9FFC2BFE2F" lastPageId="18" lastPageNumber="359" pageId="16" pageNumber="357" type="type_taxon">
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<paragraph id="8BA836DAF67AFF94BB9CFC9FFD99FC18" blockId="16.[151,1127,875,931]" pageId="16" pageNumber="357">
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<emphasis id="B963EAC8F67AFF94BB9CFC9FFE9FFC38" bold="true" box="[151,298,875,899]" pageId="16" pageNumber="357">
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<typeStatus id="54AC8878F67AFF94BB9CFC9FFF7AFC38" box="[151,207,875,899]" pageId="16" pageNumber="357">Type</typeStatus>
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species.
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</emphasis>
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<taxonomicName id="4C174D59F67AFF94BA24FC98FD61FC38" authority="Reitter, 1872: 248" authorityName="Reitter" authorityPageNumber="248" authorityYear="1872" box="[303,724,875,899]" class="Insecta" family="Nitidulidae" genus="Meligethes" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="357" phylum="Arthropoda" rank="species" species="variabilis">
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<emphasis id="B963EAC8F67AFF94BA24FC98FDA5FC38" box="[303,528,876,899]" italics="true" pageId="16" pageNumber="357">Meligethes variabilis</emphasis>
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Reitter, 1872: 248
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</taxonomicName>
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||
(by original designation) [=
|
||
<taxonomicName id="4C174D59F67AFF94BF09FC98FE14FC18" authority="(Reitter, 1872)" authorityName="Audisio & Cline & Biase & Antonini & Mancini & Trizzino & Costantini & Strika & Lamanna & Cerretti" authorityYear="2009" baseAuthorityName="Reitter" baseAuthorityYear="1872" class="Insecta" family="Nitidulidae" genus="Lariopsis" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="357" phylum="Arthropoda" rank="species" species="variabilis" status="comb. nov.">
|
||
<emphasis id="B963EAC8F67AFF94BF09FC98FF48FC18" italics="true" pageId="16" pageNumber="357">Lariopsis variabilis</emphasis>
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||
(Reitter, 1872)
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||
</taxonomicName>
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<taxonomicNameLabel id="A25057B3F67AFF94BAACFC7FFD97FC18" box="[423,546,907,931]" pageId="16" pageNumber="357" rank="species">comb. nov.</taxonomicNameLabel>
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||
].
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</paragraph>
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<paragraph id="8BA836DAF67AFF94BB9CFC4EFCBBFC49" blockId="16.[151,1127,954,1586]" pageId="16" pageNumber="357">
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<emphasis id="B963EAC8F67AFF94BB9CFC4EFD89FC69" bold="true" box="[151,572,954,978]" pageId="16" pageNumber="357">Generic redescription and diagnosis.</emphasis>
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Inclusive species vary greatly in size (
|
||
<quantity id="4CEF9B3FF67AFF94B8E8FC4EFBD3FC69" box="[995,1126,954,978]" metricMagnitude="-3" metricUnit="m" metricValue="2.9" metricValueMax="3.5" metricValueMin="2.3" pageId="16" pageNumber="357" unit="mm" value="2.9" valueMax="3.5" valueMin="2.3">2.3–3.5 mm</quantity>
|
||
length), and share the following combination of characters.
|
||
</paragraph>
|
||
<paragraph id="8BA836DAF67AFF94BBBEFC0FFC80FB49" blockId="16.[151,1127,954,1586]" pageId="16" pageNumber="357">
|
||
<emphasis id="B963EAC8F67AFF94BBBEFC0FFE50FBA9" box="[181,485,1019,1042]" italics="true" pageId="16" pageNumber="357">Body color and pubescence</emphasis>
|
||
: pubescence golden to silvery-whitish, highly variable, 1) short, fine, and recumbent or 2) long, suberect, and partially obscuring the variably colored dorsal body surface (yellowish, reddish, brown, blackish, or blackish with orange spots on elytra:
|
||
<figureCitation id="132C2A5FF67AFF94BBEAFBAEFE99FBC9" box="[225,300,1114,1138]" captionStart="Fig" captionStartId="18.[151,182,1492,1512]" captionTargetBox="[168,1109,431,1469]" captionTargetId="figure-10@18.[167,1109,431,1470]" captionTargetPageId="18" captionText="Fig. 4. Lariopsis Kirejtshuk, 1989: a – L.vultuosus (Kirejtshuk & Audisio, 1995); b–k – L. variabilis (Reitter, 1872). a – male habitus (length 3.3 mm); b, c – dorso-lateral view of head; d – ventral view of head and anterior portion of prosternum; e – prosternal process and mesoventrite; f – middle leg with outer margin of mesotibia; g – microsetae on middle posterior margin of pronotum; h – exposed portion of last visible abdominal ventrite; k – caudal marginal lines of metacoxal cavities. Scale bars: Figs. b, c, d, e, h, k = 100 μm; Fig. f = 30 μm; Fig. g = 10 μm." figureDoi="http://doi.org/10.5281/zenodo.5319344" httpUri="https://zenodo.org/record/5319344/files/figure.png" pageId="16" pageNumber="357">Fig. 4a</figureCitation>
|
||
); pronotal and elytral sides narrowly flattened, typically the same color as disc. Lateral margin of pronotum and elytra with a series of faintly distinct, small and short setae, each seta usually 0.3–0.6× as long as those on elytral disc; posterior margin of pronotum comprising moderately long, usually distally trifid or tetrafid microsetae, microsetae uniformly distributed along middle region anterior to scutellum (
|
||
<figureCitation id="132C2A5FF67AFF94B9D2FB2EFC9CFB49" box="[729,809,1242,1266]" captionStart="Fig" captionStartId="18.[151,182,1492,1512]" captionTargetBox="[168,1109,431,1469]" captionTargetId="figure-10@18.[167,1109,431,1470]" captionTargetPageId="18" captionText="Fig. 4. Lariopsis Kirejtshuk, 1989: a – L.vultuosus (Kirejtshuk & Audisio, 1995); b–k – L. variabilis (Reitter, 1872). a – male habitus (length 3.3 mm); b, c – dorso-lateral view of head; d – ventral view of head and anterior portion of prosternum; e – prosternal process and mesoventrite; f – middle leg with outer margin of mesotibia; g – microsetae on middle posterior margin of pronotum; h – exposed portion of last visible abdominal ventrite; k – caudal marginal lines of metacoxal cavities. Scale bars: Figs. b, c, d, e, h, k = 100 μm; Fig. f = 30 μm; Fig. g = 10 μm." figureDoi="http://doi.org/10.5281/zenodo.5319344" httpUri="https://zenodo.org/record/5319344/files/figure.png" pageId="16" pageNumber="357">
|
||
Fig.
|
||
<quantity id="4CEF9B3FF67AFF94B800FB2EFC9CFB49" box="[779,809,1242,1266]" metricMagnitude="-3" metricUnit="kg" metricValue="4.0" pageId="16" pageNumber="357" unit="g" value="4.0">4g</quantity>
|
||
</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BA836DAF67AFF95BBBEFB0FFF4CFE28" blockId="16.[151,1127,954,1586]" lastBlockId="17.[113,1089,219,1619]" lastPageId="17" lastPageNumber="358" pageId="16" pageNumber="357">
|
||
<emphasis id="B963EAC8F67AFF94BBBEFB0FFEE0FAA9" box="[181,341,1275,1298]" italics="true" pageId="16" pageNumber="357">Dorsal habitus</emphasis>
|
||
: body more or less convex, variably shaped, usually moderately short and wide, oval, or narrower and more parallel-sided (
|
||
<figureCitation id="132C2A5FF67AFF94B995FAEEFD59FA89" box="[670,748,1306,1330]" captionStart="Fig" captionStartId="18.[151,182,1492,1512]" captionTargetBox="[168,1109,431,1469]" captionTargetId="figure-10@18.[167,1109,431,1470]" captionTargetPageId="18" captionText="Fig. 4. Lariopsis Kirejtshuk, 1989: a – L.vultuosus (Kirejtshuk & Audisio, 1995); b–k – L. variabilis (Reitter, 1872). a – male habitus (length 3.3 mm); b, c – dorso-lateral view of head; d – ventral view of head and anterior portion of prosternum; e – prosternal process and mesoventrite; f – middle leg with outer margin of mesotibia; g – microsetae on middle posterior margin of pronotum; h – exposed portion of last visible abdominal ventrite; k – caudal marginal lines of metacoxal cavities. Scale bars: Figs. b, c, d, e, h, k = 100 μm; Fig. f = 30 μm; Fig. g = 10 μm." figureDoi="http://doi.org/10.5281/zenodo.5319344" httpUri="https://zenodo.org/record/5319344/files/figure.png" pageId="16" pageNumber="357">Fig. 4a</figureCitation>
|
||
); dorsal punctures on discal portion of pronotum larger than eye facets, moderately to deeply impressed and densely distributed; anterior margin of clypeus moderately to strongly arcuately emarginate, simple, i.e. without small distinct bulge medially, and distinctly bordered (
|
||
<figureCitation id="132C2A5FF67AFF94B9D4FA8EFC98FA29" box="[735,813,1402,1426]" captionStart="Fig" captionStartId="18.[151,182,1492,1512]" captionTargetBox="[168,1109,431,1469]" captionTargetId="figure-10@18.[167,1109,431,1470]" captionTargetPageId="18" captionText="Fig. 4. Lariopsis Kirejtshuk, 1989: a – L.vultuosus (Kirejtshuk & Audisio, 1995); b–k – L. variabilis (Reitter, 1872). a – male habitus (length 3.3 mm); b, c – dorso-lateral view of head; d – ventral view of head and anterior portion of prosternum; e – prosternal process and mesoventrite; f – middle leg with outer margin of mesotibia; g – microsetae on middle posterior margin of pronotum; h – exposed portion of last visible abdominal ventrite; k – caudal marginal lines of metacoxal cavities. Scale bars: Figs. b, c, d, e, h, k = 100 μm; Fig. f = 30 μm; Fig. g = 10 μm." figureDoi="http://doi.org/10.5281/zenodo.5319344" httpUri="https://zenodo.org/record/5319344/files/figure.png" pageId="16" pageNumber="357">Fig. 4c</figureCitation>
|
||
), with circum-ocular furrows (occipital sulci) on dorsal side of head shallow but nearly completely developed anteriorly and posteriorly (
|
||
<figureCitation id="132C2A5FF67AFF94BA4EFA4EFE00FA69" box="[325,437,1466,1490]" captionStart="Fig" captionStartId="18.[151,182,1492,1512]" captionTargetBox="[168,1109,431,1469]" captionTargetId="figure-10@18.[167,1109,431,1470]" captionTargetPageId="18" captionText="Fig. 4. Lariopsis Kirejtshuk, 1989: a – L.vultuosus (Kirejtshuk & Audisio, 1995); b–k – L. variabilis (Reitter, 1872). a – male habitus (length 3.3 mm); b, c – dorso-lateral view of head; d – ventral view of head and anterior portion of prosternum; e – prosternal process and mesoventrite; f – middle leg with outer margin of mesotibia; g – microsetae on middle posterior margin of pronotum; h – exposed portion of last visible abdominal ventrite; k – caudal marginal lines of metacoxal cavities. Scale bars: Figs. b, c, d, e, h, k = 100 μm; Fig. f = 30 μm; Fig. g = 10 μm." figureDoi="http://doi.org/10.5281/zenodo.5319344" httpUri="https://zenodo.org/record/5319344/files/figure.png" pageId="16" pageNumber="357">Figs. 4b, c</figureCitation>
|
||
); eyes large and usually moderately projecting laterally (
|
||
<figureCitation id="132C2A5FF67AFF94BF00FA4EFF74FA49" captionStart="Fig" captionStartId="18.[151,182,1492,1512]" captionTargetBox="[168,1109,431,1469]" captionTargetId="figure-10@18.[167,1109,431,1470]" captionTargetPageId="18" captionText="Fig. 4. Lariopsis Kirejtshuk, 1989: a – L.vultuosus (Kirejtshuk & Audisio, 1995); b–k – L. variabilis (Reitter, 1872). a – male habitus (length 3.3 mm); b, c – dorso-lateral view of head; d – ventral view of head and anterior portion of prosternum; e – prosternal process and mesoventrite; f – middle leg with outer margin of mesotibia; g – microsetae on middle posterior margin of pronotum; h – exposed portion of last visible abdominal ventrite; k – caudal marginal lines of metacoxal cavities. Scale bars: Figs. b, c, d, e, h, k = 100 μm; Fig. f = 30 μm; Fig. g = 10 μm." figureDoi="http://doi.org/10.5281/zenodo.5319344" httpUri="https://zenodo.org/record/5319344/files/figure.png" pageId="16" pageNumber="357">Figs. 4a, b, c</figureCitation>
|
||
); pronotum with faintly distinct posterior angles, rounded to obtuse and never directed posteriorly (
|
||
<figureCitation id="132C2A5FF67AFF94BA12FA0EFED2F9A9" box="[281,359,1530,1554]" captionStart="Fig" captionStartId="18.[151,182,1492,1512]" captionTargetBox="[168,1109,431,1469]" captionTargetId="figure-10@18.[167,1109,431,1470]" captionTargetPageId="18" captionText="Fig. 4. Lariopsis Kirejtshuk, 1989: a – L.vultuosus (Kirejtshuk & Audisio, 1995); b–k – L. variabilis (Reitter, 1872). a – male habitus (length 3.3 mm); b, c – dorso-lateral view of head; d – ventral view of head and anterior portion of prosternum; e – prosternal process and mesoventrite; f – middle leg with outer margin of mesotibia; g – microsetae on middle posterior margin of pronotum; h – exposed portion of last visible abdominal ventrite; k – caudal marginal lines of metacoxal cavities. Scale bars: Figs. b, c, d, e, h, k = 100 μm; Fig. f = 30 μm; Fig. g = 10 μm." figureDoi="http://doi.org/10.5281/zenodo.5319344" httpUri="https://zenodo.org/record/5319344/files/figure.png" pageId="16" pageNumber="357">Fig. 4a</figureCitation>
|
||
); scutellum regularly punctured on most of exposed portion; elytra with simple punctation, never transversely strigose; elytral humeral angle faintly distinct, not protruding laterally (
|
||
<figureCitation id="132C2A5FF67BFF95BA44FF2FFE28FF4F" box="[335,413,219,244]" captionStart="Fig" captionStartId="18.[151,182,1492,1512]" captionTargetBox="[168,1109,431,1469]" captionTargetId="figure-10@18.[167,1109,431,1470]" captionTargetPageId="18" captionText="Fig. 4. Lariopsis Kirejtshuk, 1989: a – L.vultuosus (Kirejtshuk & Audisio, 1995); b–k – L. variabilis (Reitter, 1872). a – male habitus (length 3.3 mm); b, c – dorso-lateral view of head; d – ventral view of head and anterior portion of prosternum; e – prosternal process and mesoventrite; f – middle leg with outer margin of mesotibia; g – microsetae on middle posterior margin of pronotum; h – exposed portion of last visible abdominal ventrite; k – caudal marginal lines of metacoxal cavities. Scale bars: Figs. b, c, d, e, h, k = 100 μm; Fig. f = 30 μm; Fig. g = 10 μm." figureDoi="http://doi.org/10.5281/zenodo.5319344" httpUri="https://zenodo.org/record/5319344/files/figure.png" pageId="17" pageNumber="358">Fig. 4a</figureCitation>
|
||
); elytral humeral striae absent; elytral pre-sutural striae visible, originating at scutellar vertex, terminating at elytral apex, and delimiting on each elytron a faintly distinct, flat, not raised sutural border, border widest at posterior third, slightly narrower than proximal width of 3
|
||
<superScript id="7C629B92F67BFF95BAA3FECEFE00FEF2" attach="left" box="[424,437,314,329]" fontSize="6" pageId="17" pageNumber="358">rd</superScript>
|
||
antennomere; elytral apices truncately rounded in both sexes (
|
||
<figureCitation id="132C2A5FF67BFF95BB72FEAFFF7CFECF" box="[121,201,347,372]" captionStart="Fig" captionStartId="18.[151,182,1492,1512]" captionTargetBox="[168,1109,431,1469]" captionTargetId="figure-10@18.[167,1109,431,1470]" captionTargetPageId="18" captionText="Fig. 4. Lariopsis Kirejtshuk, 1989: a – L.vultuosus (Kirejtshuk & Audisio, 1995); b–k – L. variabilis (Reitter, 1872). a – male habitus (length 3.3 mm); b, c – dorso-lateral view of head; d – ventral view of head and anterior portion of prosternum; e – prosternal process and mesoventrite; f – middle leg with outer margin of mesotibia; g – microsetae on middle posterior margin of pronotum; h – exposed portion of last visible abdominal ventrite; k – caudal marginal lines of metacoxal cavities. Scale bars: Figs. b, c, d, e, h, k = 100 μm; Fig. f = 30 μm; Fig. g = 10 μm." figureDoi="http://doi.org/10.5281/zenodo.5319344" httpUri="https://zenodo.org/record/5319344/files/figure.png" pageId="17" pageNumber="358">Fig. 4a</figureCitation>
|
||
); pygidium partially exposed, moderately convex, apically rounded in both sexes (
|
||
<figureCitation id="132C2A5FF67BFF95BB72FE8FFF59FE28" box="[121,236,379,404]" captionStart="Fig" captionStartId="18.[151,182,1492,1512]" captionTargetBox="[168,1109,431,1469]" captionTargetId="figure-10@18.[167,1109,431,1470]" captionTargetPageId="18" captionText="Fig. 4. Lariopsis Kirejtshuk, 1989: a – L.vultuosus (Kirejtshuk & Audisio, 1995); b–k – L. variabilis (Reitter, 1872). a – male habitus (length 3.3 mm); b, c – dorso-lateral view of head; d – ventral view of head and anterior portion of prosternum; e – prosternal process and mesoventrite; f – middle leg with outer margin of mesotibia; g – microsetae on middle posterior margin of pronotum; h – exposed portion of last visible abdominal ventrite; k – caudal marginal lines of metacoxal cavities. Scale bars: Figs. b, c, d, e, h, k = 100 μm; Fig. f = 30 μm; Fig. g = 10 μm." figureDoi="http://doi.org/10.5281/zenodo.5319344" httpUri="https://zenodo.org/record/5319344/files/figure.png" pageId="17" pageNumber="358">Figs. 4a, h</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BA836DAF67BFF95BB9BFE69FE9BFCC8" blockId="17.[113,1089,219,1619]" pageId="17" pageNumber="358">
|
||
<emphasis id="B963EAC8F67BFF95BB9BFE69FE80FE0F" box="[144,309,413,436]" italics="true" pageId="17" pageNumber="358">Ventral habitus</emphasis>
|
||
: antennal furrows markedly delimited, nearly parallel-sided, or slightly divergent posteriorly; mentum subpentagonal (
|
||
<figureCitation id="132C2A5FF67BFF95B976FE4FFD64FE68" box="[637,721,443,467]" captionStart="Fig" captionStartId="18.[151,182,1492,1512]" captionTargetBox="[168,1109,431,1469]" captionTargetId="figure-10@18.[167,1109,431,1470]" captionTargetPageId="18" captionText="Fig. 4. Lariopsis Kirejtshuk, 1989: a – L.vultuosus (Kirejtshuk & Audisio, 1995); b–k – L. variabilis (Reitter, 1872). a – male habitus (length 3.3 mm); b, c – dorso-lateral view of head; d – ventral view of head and anterior portion of prosternum; e – prosternal process and mesoventrite; f – middle leg with outer margin of mesotibia; g – microsetae on middle posterior margin of pronotum; h – exposed portion of last visible abdominal ventrite; k – caudal marginal lines of metacoxal cavities. Scale bars: Figs. b, c, d, e, h, k = 100 μm; Fig. f = 30 μm; Fig. g = 10 μm." figureDoi="http://doi.org/10.5281/zenodo.5319344" httpUri="https://zenodo.org/record/5319344/files/figure.png" pageId="17" pageNumber="358">Fig. 4d</figureCitation>
|
||
); prosternal antennal furrows on anterior margin of prosternum almost completely obliterated (
|
||
<figureCitation id="132C2A5FF67BFF95B81EFE2FFCD2FE48" box="[789,871,475,499]" captionStart="Fig" captionStartId="18.[151,182,1492,1512]" captionTargetBox="[168,1109,431,1469]" captionTargetId="figure-10@18.[167,1109,431,1470]" captionTargetPageId="18" captionText="Fig. 4. Lariopsis Kirejtshuk, 1989: a – L.vultuosus (Kirejtshuk & Audisio, 1995); b–k – L. variabilis (Reitter, 1872). a – male habitus (length 3.3 mm); b, c – dorso-lateral view of head; d – ventral view of head and anterior portion of prosternum; e – prosternal process and mesoventrite; f – middle leg with outer margin of mesotibia; g – microsetae on middle posterior margin of pronotum; h – exposed portion of last visible abdominal ventrite; k – caudal marginal lines of metacoxal cavities. Scale bars: Figs. b, c, d, e, h, k = 100 μm; Fig. f = 30 μm; Fig. g = 10 μm." figureDoi="http://doi.org/10.5281/zenodo.5319344" httpUri="https://zenodo.org/record/5319344/files/figure.png" pageId="17" pageNumber="358">Fig. 4d</figureCitation>
|
||
); prosternal process relatively narrow, subapical portion strongly dilated, 2.3–2.5× as wide as maximum width of 1
|
||
<superScript id="7C629B92F67BFF95BB90FDEEFF10FD92" attach="left" box="[155,165,538,553]" fontSize="6" pageId="17" pageNumber="358">st</superScript>
|
||
antennomere, apex arcuately convex (
|
||
<figureCitation id="132C2A5FF67BFF95B936FDEFFD3FFD8F" box="[573,650,539,564]" captionStart="Fig" captionStartId="18.[151,182,1492,1512]" captionTargetBox="[168,1109,431,1469]" captionTargetId="figure-10@18.[167,1109,431,1470]" captionTargetPageId="18" captionText="Fig. 4. Lariopsis Kirejtshuk, 1989: a – L.vultuosus (Kirejtshuk & Audisio, 1995); b–k – L. variabilis (Reitter, 1872). a – male habitus (length 3.3 mm); b, c – dorso-lateral view of head; d – ventral view of head and anterior portion of prosternum; e – prosternal process and mesoventrite; f – middle leg with outer margin of mesotibia; g – microsetae on middle posterior margin of pronotum; h – exposed portion of last visible abdominal ventrite; k – caudal marginal lines of metacoxal cavities. Scale bars: Figs. b, c, d, e, h, k = 100 μm; Fig. f = 30 μm; Fig. g = 10 μm." figureDoi="http://doi.org/10.5281/zenodo.5319344" httpUri="https://zenodo.org/record/5319344/files/figure.png" pageId="17" pageNumber="358">Fig. 4e</figureCitation>
|
||
); lateral borders of prosternal process not delimiting furrows, terminating at base of prosternal process (
|
||
<figureCitation id="132C2A5FF67BFF95B806FDCFFCE9FDEF" box="[781,860,571,596]" captionStart="Fig" captionStartId="18.[151,182,1492,1512]" captionTargetBox="[168,1109,431,1469]" captionTargetId="figure-10@18.[167,1109,431,1470]" captionTargetPageId="18" captionText="Fig. 4. Lariopsis Kirejtshuk, 1989: a – L.vultuosus (Kirejtshuk & Audisio, 1995); b–k – L. variabilis (Reitter, 1872). a – male habitus (length 3.3 mm); b, c – dorso-lateral view of head; d – ventral view of head and anterior portion of prosternum; e – prosternal process and mesoventrite; f – middle leg with outer margin of mesotibia; g – microsetae on middle posterior margin of pronotum; h – exposed portion of last visible abdominal ventrite; k – caudal marginal lines of metacoxal cavities. Scale bars: Figs. b, c, d, e, h, k = 100 μm; Fig. f = 30 μm; Fig. g = 10 μm." figureDoi="http://doi.org/10.5281/zenodo.5319344" httpUri="https://zenodo.org/record/5319344/files/figure.png" pageId="17" pageNumber="358">Fig. 4e</figureCitation>
|
||
); posterior margin of mesoventrite simple, never incised medially, posteriorly slightly convex (
|
||
<figureCitation id="132C2A5FF67BFF95B88DFDAFFC60FDCF" box="[902,981,603,628]" captionStart="Fig" captionStartId="18.[151,182,1492,1512]" captionTargetBox="[168,1109,431,1469]" captionTargetId="figure-10@18.[167,1109,431,1470]" captionTargetPageId="18" captionText="Fig. 4. Lariopsis Kirejtshuk, 1989: a – L.vultuosus (Kirejtshuk & Audisio, 1995); b–k – L. variabilis (Reitter, 1872). a – male habitus (length 3.3 mm); b, c – dorso-lateral view of head; d – ventral view of head and anterior portion of prosternum; e – prosternal process and mesoventrite; f – middle leg with outer margin of mesotibia; g – microsetae on middle posterior margin of pronotum; h – exposed portion of last visible abdominal ventrite; k – caudal marginal lines of metacoxal cavities. Scale bars: Figs. b, c, d, e, h, k = 100 μm; Fig. f = 30 μm; Fig. g = 10 μm." figureDoi="http://doi.org/10.5281/zenodo.5319344" httpUri="https://zenodo.org/record/5319344/files/figure.png" pageId="17" pageNumber="358">Fig. 4e</figureCitation>
|
||
); variably developed male impressions on metaventrite and hypopygial tubercles (
|
||
<figureCitation id="132C2A5FF67BFF95B874FD8FFC64FD28" box="[895,977,635,659]" captionStart="Fig" captionStartId="18.[151,182,1492,1512]" captionTargetBox="[168,1109,431,1469]" captionTargetId="figure-10@18.[167,1109,431,1470]" captionTargetPageId="18" captionText="Fig. 4. Lariopsis Kirejtshuk, 1989: a – L.vultuosus (Kirejtshuk & Audisio, 1995); b–k – L. variabilis (Reitter, 1872). a – male habitus (length 3.3 mm); b, c – dorso-lateral view of head; d – ventral view of head and anterior portion of prosternum; e – prosternal process and mesoventrite; f – middle leg with outer margin of mesotibia; g – microsetae on middle posterior margin of pronotum; h – exposed portion of last visible abdominal ventrite; k – caudal marginal lines of metacoxal cavities. Scale bars: Figs. b, c, d, e, h, k = 100 μm; Fig. f = 30 μm; Fig. g = 10 μm." figureDoi="http://doi.org/10.5281/zenodo.5319344" httpUri="https://zenodo.org/record/5319344/files/figure.png" pageId="17" pageNumber="358">Fig. 4h</figureCitation>
|
||
); first two visible abdominal ventrites simple in both sexes, without tufts of setae; caudal marginal lines of metacoxal cavities simple, parallel and contiguous to posterior margin of metacoxal cavities, without arched impression of outer ‘axillary’ line (
|
||
<figureCitation id="132C2A5FF67BFF95B9E2FD2FFC8FFD48" box="[745,826,731,755]" captionStart="Fig" captionStartId="18.[151,182,1492,1512]" captionTargetBox="[168,1109,431,1469]" captionTargetId="figure-10@18.[167,1109,431,1470]" captionTargetPageId="18" captionText="Fig. 4. Lariopsis Kirejtshuk, 1989: a – L.vultuosus (Kirejtshuk & Audisio, 1995); b–k – L. variabilis (Reitter, 1872). a – male habitus (length 3.3 mm); b, c – dorso-lateral view of head; d – ventral view of head and anterior portion of prosternum; e – prosternal process and mesoventrite; f – middle leg with outer margin of mesotibia; g – microsetae on middle posterior margin of pronotum; h – exposed portion of last visible abdominal ventrite; k – caudal marginal lines of metacoxal cavities. Scale bars: Figs. b, c, d, e, h, k = 100 μm; Fig. f = 30 μm; Fig. g = 10 μm." figureDoi="http://doi.org/10.5281/zenodo.5319344" httpUri="https://zenodo.org/record/5319344/files/figure.png" pageId="17" pageNumber="358">Fig. 4k</figureCitation>
|
||
); ‘axillary’ space on first abdominal ventrite moderately developed, ‘axillary’ angle widely obtuse (
|
||
<figureCitation id="132C2A5FF67BFF95B88EFD0FFC60FCA8" box="[901,981,763,787]" captionStart="Fig" captionStartId="18.[151,182,1492,1512]" captionTargetBox="[168,1109,431,1469]" captionTargetId="figure-10@18.[167,1109,431,1470]" captionTargetPageId="18" captionText="Fig. 4. Lariopsis Kirejtshuk, 1989: a – L.vultuosus (Kirejtshuk & Audisio, 1995); b–k – L. variabilis (Reitter, 1872). a – male habitus (length 3.3 mm); b, c – dorso-lateral view of head; d – ventral view of head and anterior portion of prosternum; e – prosternal process and mesoventrite; f – middle leg with outer margin of mesotibia; g – microsetae on middle posterior margin of pronotum; h – exposed portion of last visible abdominal ventrite; k – caudal marginal lines of metacoxal cavities. Scale bars: Figs. b, c, d, e, h, k = 100 μm; Fig. f = 30 μm; Fig. g = 10 μm." figureDoi="http://doi.org/10.5281/zenodo.5319344" httpUri="https://zenodo.org/record/5319344/files/figure.png" pageId="17" pageNumber="358">Fig. 4k</figureCitation>
|
||
); moderately large, short and shallowly impressed arched impressions on basal portion of last visible abdominal ventrite, frequently partially obscured by distal portion of penultimate abdominal ventrite (
|
||
<figureCitation id="132C2A5FF67BFF95BBDAFCAFFE94FCC8" box="[209,289,859,883]" captionStart="Fig" captionStartId="18.[151,182,1492,1512]" captionTargetBox="[168,1109,431,1469]" captionTargetId="figure-10@18.[167,1109,431,1470]" captionTargetPageId="18" captionText="Fig. 4. Lariopsis Kirejtshuk, 1989: a – L.vultuosus (Kirejtshuk & Audisio, 1995); b–k – L. variabilis (Reitter, 1872). a – male habitus (length 3.3 mm); b, c – dorso-lateral view of head; d – ventral view of head and anterior portion of prosternum; e – prosternal process and mesoventrite; f – middle leg with outer margin of mesotibia; g – microsetae on middle posterior margin of pronotum; h – exposed portion of last visible abdominal ventrite; k – caudal marginal lines of metacoxal cavities. Scale bars: Figs. b, c, d, e, h, k = 100 μm; Fig. f = 30 μm; Fig. g = 10 μm." figureDoi="http://doi.org/10.5281/zenodo.5319344" httpUri="https://zenodo.org/record/5319344/files/figure.png" pageId="17" pageNumber="358">Fig. 4h</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BA836DAF67BFF95BB9BFC89FDFCFA08" blockId="17.[113,1089,219,1619]" pageId="17" pageNumber="358">
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<emphasis id="B963EAC8F67BFF95BB9BFC89FEA7FC2F" box="[144,274,893,916]" italics="true" pageId="17" pageNumber="358">Appendages</emphasis>
|
||
: male 1
|
||
<superScript id="7C629B92F67BFF95BA6FFC8EFEDBFC32" attach="left" box="[356,366,890,905]" fontSize="6" pageId="17" pageNumber="358">st</superScript>
|
||
antennomere 0.8–1.0× as long as width of protibiae excluding distal teeth (
|
||
<figureCitation id="132C2A5FF67BFF95BBB8FC6FFE96FC0F" box="[179,291,923,948]" captionStart="Fig" captionStartId="18.[151,182,1492,1512]" captionTargetBox="[168,1109,431,1469]" captionTargetId="figure-10@18.[167,1109,431,1470]" captionTargetPageId="18" captionText="Fig. 4. Lariopsis Kirejtshuk, 1989: a – L.vultuosus (Kirejtshuk & Audisio, 1995); b–k – L. variabilis (Reitter, 1872). a – male habitus (length 3.3 mm); b, c – dorso-lateral view of head; d – ventral view of head and anterior portion of prosternum; e – prosternal process and mesoventrite; f – middle leg with outer margin of mesotibia; g – microsetae on middle posterior margin of pronotum; h – exposed portion of last visible abdominal ventrite; k – caudal marginal lines of metacoxal cavities. Scale bars: Figs. b, c, d, e, h, k = 100 μm; Fig. f = 30 μm; Fig. g = 10 μm." figureDoi="http://doi.org/10.5281/zenodo.5319344" httpUri="https://zenodo.org/record/5319344/files/figure.png" pageId="17" pageNumber="358">Figs. 4a, c</figureCitation>
|
||
); 3
|
||
<superScript id="7C629B92F67BFF95BA4EFC6EFEE7FC12" attach="left" box="[325,338,922,937]" fontSize="6" pageId="17" pageNumber="358">rd</superScript>
|
||
antennomere in both sexes usually 2.3–2.4× as long as wide, 1.2–1.3× longer and distinctly thinner than 2
|
||
<superScript id="7C629B92F67BFF95BAE1FC4EFE4FFC72" attach="left" box="[490,506,954,969]" fontSize="6" pageId="17" pageNumber="358">nd</superScript>
|
||
antennomere (
|
||
<figureCitation id="132C2A5FF67BFF95B991FC4FFCB9FC6F" box="[666,780,955,980]" captionStart="Fig" captionStartId="18.[151,182,1492,1512]" captionTargetBox="[168,1109,431,1469]" captionTargetId="figure-10@18.[167,1109,431,1470]" captionTargetPageId="18" captionText="Fig. 4. Lariopsis Kirejtshuk, 1989: a – L.vultuosus (Kirejtshuk & Audisio, 1995); b–k – L. variabilis (Reitter, 1872). a – male habitus (length 3.3 mm); b, c – dorso-lateral view of head; d – ventral view of head and anterior portion of prosternum; e – prosternal process and mesoventrite; f – middle leg with outer margin of mesotibia; g – microsetae on middle posterior margin of pronotum; h – exposed portion of last visible abdominal ventrite; k – caudal marginal lines of metacoxal cavities. Scale bars: Figs. b, c, d, e, h, k = 100 μm; Fig. f = 30 μm; Fig. g = 10 μm." figureDoi="http://doi.org/10.5281/zenodo.5319344" httpUri="https://zenodo.org/record/5319344/files/figure.png" pageId="17" pageNumber="358">Figs. 4a, c</figureCitation>
|
||
); 4
|
||
<superScript id="7C629B92F67BFF95B824FC4EFC8EFC72" attach="left" box="[815,827,954,969]" fontSize="6" pageId="17" pageNumber="358">th</superScript>
|
||
and 5
|
||
<superScript id="7C629B92F67BFF95B876FC4EFC3CFC72" attach="left" box="[893,905,954,969]" fontSize="6" pageId="17" pageNumber="358">th</superScript>
|
||
antennomeres in both sexes subequal, short, slightly longer than wide; antennal club compact, small, simple, comprising last 3 antennomeres in both sexes (8
|
||
<superScript id="7C629B92F67BFF95B966FC0EFDCCFBB2" attach="left" box="[621,633,1018,1033]" fontSize="6" pageId="17" pageNumber="358">th</superScript>
|
||
antennomere moderately widened, 0.6× as wide as 9
|
||
<superScript id="7C629B92F67BFF95BBD9FBEEFF6BFB92" attach="right" box="[210,222,1050,1065]" fontSize="6" pageId="17" pageNumber="358">th</superScript>
|
||
antennomere) (
|
||
<figureCitation id="132C2A5FF67BFF95BA75FBEFFE79FB88" box="[382,460,1051,1075]" captionStart="Fig" captionStartId="18.[151,182,1492,1512]" captionTargetBox="[168,1109,431,1469]" captionTargetId="figure-10@18.[167,1109,431,1470]" captionTargetPageId="18" captionText="Fig. 4. Lariopsis Kirejtshuk, 1989: a – L.vultuosus (Kirejtshuk & Audisio, 1995); b–k – L. variabilis (Reitter, 1872). a – male habitus (length 3.3 mm); b, c – dorso-lateral view of head; d – ventral view of head and anterior portion of prosternum; e – prosternal process and mesoventrite; f – middle leg with outer margin of mesotibia; g – microsetae on middle posterior margin of pronotum; h – exposed portion of last visible abdominal ventrite; k – caudal marginal lines of metacoxal cavities. Scale bars: Figs. b, c, d, e, h, k = 100 μm; Fig. f = 30 μm; Fig. g = 10 μm." figureDoi="http://doi.org/10.5281/zenodo.5319344" httpUri="https://zenodo.org/record/5319344/files/figure.png" pageId="17" pageNumber="358">Fig. 4b</figureCitation>
|
||
), much narrower than width of protibiae, sexual dimorphism absent; labial palpi moderately long (
|
||
<figureCitation id="132C2A5FF67BFF95B90FFBCFFDE0FBE8" box="[516,597,1083,1107]" captionStart="Fig" captionStartId="18.[151,182,1492,1512]" captionTargetBox="[168,1109,431,1469]" captionTargetId="figure-10@18.[167,1109,431,1470]" captionTargetPageId="18" captionText="Fig. 4. Lariopsis Kirejtshuk, 1989: a – L.vultuosus (Kirejtshuk & Audisio, 1995); b–k – L. variabilis (Reitter, 1872). a – male habitus (length 3.3 mm); b, c – dorso-lateral view of head; d – ventral view of head and anterior portion of prosternum; e – prosternal process and mesoventrite; f – middle leg with outer margin of mesotibia; g – microsetae on middle posterior margin of pronotum; h – exposed portion of last visible abdominal ventrite; k – caudal marginal lines of metacoxal cavities. Scale bars: Figs. b, c, d, e, h, k = 100 μm; Fig. f = 30 μm; Fig. g = 10 μm." figureDoi="http://doi.org/10.5281/zenodo.5319344" httpUri="https://zenodo.org/record/5319344/files/figure.png" pageId="17" pageNumber="358">Fig. 4d</figureCitation>
|
||
), terminal segment 1.9–2.0× as long as wide; maxillary palpi moderately long and slender (
|
||
<figureCitation id="132C2A5FF67BFF95B95FFBAFFD16FBC8" box="[596,675,1115,1139]" captionStart="Fig" captionStartId="18.[151,182,1492,1512]" captionTargetBox="[168,1109,431,1469]" captionTargetId="figure-10@18.[167,1109,431,1470]" captionTargetPageId="18" captionText="Fig. 4. Lariopsis Kirejtshuk, 1989: a – L.vultuosus (Kirejtshuk & Audisio, 1995); b–k – L. variabilis (Reitter, 1872). a – male habitus (length 3.3 mm); b, c – dorso-lateral view of head; d – ventral view of head and anterior portion of prosternum; e – prosternal process and mesoventrite; f – middle leg with outer margin of mesotibia; g – microsetae on middle posterior margin of pronotum; h – exposed portion of last visible abdominal ventrite; k – caudal marginal lines of metacoxal cavities. Scale bars: Figs. b, c, d, e, h, k = 100 μm; Fig. f = 30 μm; Fig. g = 10 μm." figureDoi="http://doi.org/10.5281/zenodo.5319344" httpUri="https://zenodo.org/record/5319344/files/figure.png" pageId="17" pageNumber="358">Fig. 4d</figureCitation>
|
||
), terminal segment 2.5–2.6× as long as wide; mandible usually mid-sized (
|
||
<figureCitation id="132C2A5FF67BFF95BAEEFB8FFD81FB28" box="[485,564,1147,1171]" captionStart="Fig" captionStartId="18.[151,182,1492,1512]" captionTargetBox="[168,1109,431,1469]" captionTargetId="figure-10@18.[167,1109,431,1470]" captionTargetPageId="18" captionText="Fig. 4. Lariopsis Kirejtshuk, 1989: a – L.vultuosus (Kirejtshuk & Audisio, 1995); b–k – L. variabilis (Reitter, 1872). a – male habitus (length 3.3 mm); b, c – dorso-lateral view of head; d – ventral view of head and anterior portion of prosternum; e – prosternal process and mesoventrite; f – middle leg with outer margin of mesotibia; g – microsetae on middle posterior margin of pronotum; h – exposed portion of last visible abdominal ventrite; k – caudal marginal lines of metacoxal cavities. Scale bars: Figs. b, c, d, e, h, k = 100 μm; Fig. f = 30 μm; Fig. g = 10 μm." figureDoi="http://doi.org/10.5281/zenodo.5319344" httpUri="https://zenodo.org/record/5319344/files/figure.png" pageId="17" pageNumber="358">Fig. 4d</figureCitation>
|
||
), apex moderately acuminate, sexual dimorphism absent; tarsal claws strongly toothed at base (as in
|
||
<figureCitation id="132C2A5FF67BFF95B982FB6FFD59FB0F" box="[649,748,1179,1204]" captionStart="Fig" captionStartId="62.[151,182,1415,1435]" captionTargetBox="[152,1126,217,1360]" captionTargetId="figure-10@62.[151,1127,217,1360]" captionTargetPageId="62" captionText="Fig. 17. Aristogethes Audisio & Cline, gen. nov.: a – A. translatus (Grouvelle, 1913); b – A. pecten (Audisio, Kirk-Spriggs & Kirejtshuk, 1998); c, e–k – A. pubescens (Reitter, 1872); d, m – A. marshalli (Grouvelle, 1914). a, b – male habitus (a – length 2.6 mm; b – length 2.4 mm); c, d – dorsal view of head; e – ventral view of head and anterior portion of prosternum; f – prosternal process and mesoventrite; g – anterior portion of scutellum and microsetae on middle of posterior margin of pronotum; h – exposed portion of last visible abdominal ventrite; k – caudal marginal line of metacoxal cavity; m – last tarsomere of middle leg. Scale bars: Figs. d, f, h, k, m = 100 μm; Fig. g = 20 μm." figureDoi="http://doi.org/10.5281/zenodo.5363178" httpUri="https://zenodo.org/record/5363178/files/figure.png" pageId="17" pageNumber="358">
|
||
Fig.
|
||
<quantity id="4CEF9B3FF67BFF95B9B2FB68FD59FB0F" box="[697,748,1180,1204]" metricMagnitude="1" metricUnit="m" metricValue="1.7" pageId="17" pageNumber="358" unit="m" value="17.0">17m</quantity>
|
||
</figureCitation>
|
||
); tarsi of normal size and shape, 0.6–0.7× as long as corresponding tibiae (
|
||
<figureCitation id="132C2A5FF67BFF95B93CFB4FFD33FB6F" box="[567,646,1211,1236]" captionStart="Fig" captionStartId="18.[151,182,1492,1512]" captionTargetBox="[168,1109,431,1469]" captionTargetId="figure-10@18.[167,1109,431,1470]" captionTargetPageId="18" captionText="Fig. 4. Lariopsis Kirejtshuk, 1989: a – L.vultuosus (Kirejtshuk & Audisio, 1995); b–k – L. variabilis (Reitter, 1872). a – male habitus (length 3.3 mm); b, c – dorso-lateral view of head; d – ventral view of head and anterior portion of prosternum; e – prosternal process and mesoventrite; f – middle leg with outer margin of mesotibia; g – microsetae on middle posterior margin of pronotum; h – exposed portion of last visible abdominal ventrite; k – caudal marginal lines of metacoxal cavities. Scale bars: Figs. b, c, d, e, h, k = 100 μm; Fig. f = 30 μm; Fig. g = 10 μm." figureDoi="http://doi.org/10.5281/zenodo.5319344" httpUri="https://zenodo.org/record/5319344/files/figure.png" pageId="17" pageNumber="358">Fig. 4a</figureCitation>
|
||
); protibiae with a series of large, wide at base, uneven, moderately sharp teeth on lateral margin (
|
||
<figureCitation id="132C2A5FF67BFF95B9CEFB2FFC83FB48" box="[709,822,1243,1268]" captionStart="Fig" captionStartId="18.[151,182,1492,1512]" captionTargetBox="[168,1109,431,1469]" captionTargetId="figure-10@18.[167,1109,431,1470]" captionTargetPageId="18" captionText="Fig. 4. Lariopsis Kirejtshuk, 1989: a – L.vultuosus (Kirejtshuk & Audisio, 1995); b–k – L. variabilis (Reitter, 1872). a – male habitus (length 3.3 mm); b, c – dorso-lateral view of head; d – ventral view of head and anterior portion of prosternum; e – prosternal process and mesoventrite; f – middle leg with outer margin of mesotibia; g – microsetae on middle posterior margin of pronotum; h – exposed portion of last visible abdominal ventrite; k – caudal marginal lines of metacoxal cavities. Scale bars: Figs. b, c, d, e, h, k = 100 μm; Fig. f = 30 μm; Fig. g = 10 μm." figureDoi="http://doi.org/10.5281/zenodo.5319344" httpUri="https://zenodo.org/record/5319344/files/figure.png" pageId="17" pageNumber="358">Figs. 4a, e</figureCitation>
|
||
; Fig.
|
||
<quantity id="4CEF9B3FF67BFF95B879FB28FC1DFB48" box="[882,936,1243,1268]" metricMagnitude="0" metricUnit="m" metricValue="1.8796" pageId="17" pageNumber="358" unit="in" value="74.0">74 in</quantity>
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<bibRefCitation id="EF864B2BF67BFF95B8A4FB28FF4AFAA8" author="KIREJTSHUK A. G. & AUDISIO P." pageId="17" pageNumber="358" pagination="191 - 254" refId="ref90005" refString="KIREJTSHUK A. G. & AUDISIO P. 1995: Preliminary revision of South African Meligethes subgenus Lariopsis (Coleoptera: Nitidulidae, Meligethinae). Fragmenta Entomologica 27: 191 - 254." type="journal article" year="1995">KIREJTSHUK & AUDISIO 1995</bibRefCitation>
|
||
); meso- and metatibiae on lateral margin bearing a nearly double and uneven row of large and robust brown spurs (
|
||
<figureCitation id="132C2A5FF67BFF95BACEFAEFFDADFA88" box="[453,536,1307,1331]" captionStart="Fig" captionStartId="18.[151,182,1492,1512]" captionTargetBox="[168,1109,431,1469]" captionTargetId="figure-10@18.[167,1109,431,1470]" captionTargetPageId="18" captionText="Fig. 4. Lariopsis Kirejtshuk, 1989: a – L.vultuosus (Kirejtshuk & Audisio, 1995); b–k – L. variabilis (Reitter, 1872). a – male habitus (length 3.3 mm); b, c – dorso-lateral view of head; d – ventral view of head and anterior portion of prosternum; e – prosternal process and mesoventrite; f – middle leg with outer margin of mesotibia; g – microsetae on middle posterior margin of pronotum; h – exposed portion of last visible abdominal ventrite; k – caudal marginal lines of metacoxal cavities. Scale bars: Figs. b, c, d, e, h, k = 100 μm; Fig. f = 30 μm; Fig. g = 10 μm." figureDoi="http://doi.org/10.5281/zenodo.5319344" httpUri="https://zenodo.org/record/5319344/files/figure.png" pageId="17" pageNumber="358">Figs. 4f</figureCitation>
|
||
), U-shaped sinuosity absent at distal third; meso- and metatibiae moderately slender, flat (
|
||
<figureCitation id="132C2A5FF67BFF95BAF9FACFFDF5FAEF" box="[498,576,1339,1364]" captionStart="Fig" captionStartId="18.[151,182,1492,1512]" captionTargetBox="[168,1109,431,1469]" captionTargetId="figure-10@18.[167,1109,431,1470]" captionTargetPageId="18" captionText="Fig. 4. Lariopsis Kirejtshuk, 1989: a – L.vultuosus (Kirejtshuk & Audisio, 1995); b–k – L. variabilis (Reitter, 1872). a – male habitus (length 3.3 mm); b, c – dorso-lateral view of head; d – ventral view of head and anterior portion of prosternum; e – prosternal process and mesoventrite; f – middle leg with outer margin of mesotibia; g – microsetae on middle posterior margin of pronotum; h – exposed portion of last visible abdominal ventrite; k – caudal marginal lines of metacoxal cavities. Scale bars: Figs. b, c, d, e, h, k = 100 μm; Fig. f = 30 μm; Fig. g = 10 μm." figureDoi="http://doi.org/10.5281/zenodo.5319344" httpUri="https://zenodo.org/record/5319344/files/figure.png" pageId="17" pageNumber="358">Fig. 4a</figureCitation>
|
||
), never distinctly subtrapezoidal or axe-shaped; sexual dimorphism variably expressed in metatibiae, i.e. more distinctly sinuate in males, and tarsal plates of prolegs moderately wider in males; posterior margin of metafemora simple in both sexes, without tubercles or projections.
|
||
</paragraph>
|
||
<paragraph id="8BA836DAF67BFF95BB9BFA49FCBFF9E8" blockId="17.[113,1089,219,1619]" pageId="17" pageNumber="358">
|
||
<emphasis id="B963EAC8F67BFF95BB9BFA49FE9FFA6F" box="[144,298,1469,1492]" italics="true" pageId="17" pageNumber="358">Male genitalia</emphasis>
|
||
: processes along inner side of parameres absent (
|
||
<figureCitation id="132C2A5FF67BFF95B825FA4FFC1EFA6F" box="[814,939,1467,1492]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStart-3="Fig" captionStartId-0="62.[151,182,1415,1435]" captionStartId-1="66.[151,182,1111,1131]" captionStartId-2="70.[151,182,1491,1511]" captionStartId-3="72.[151,182,1463,1483]" captionTargetBox-0="[152,1126,217,1360]" captionTargetBox-1="[169,1067,225,1090]" captionTargetBox-2="[185,1092,408,1471]" captionTargetBox-3="[196,1082,403,1444]" captionTargetId-0="figure-10@62.[151,1127,217,1360]" captionTargetId-1="figure-175@66.[151,1111,216,1094]" captionTargetId-2="figure-10@70.[185,1093,408,1471]" captionTargetId-3="figure-10@72.[195,1083,403,1444]" captionTargetPageId-0="62" captionTargetPageId-1="66" captionTargetPageId-2="70" captionTargetPageId-3="72" captionText-0="Fig. 17. Aristogethes Audisio & Cline, gen. nov.: a – A. translatus (Grouvelle, 1913); b – A. pecten (Audisio, Kirk-Spriggs & Kirejtshuk, 1998); c, e–k – A. pubescens (Reitter, 1872); d, m – A. marshalli (Grouvelle, 1914). a, b – male habitus (a – length 2.6 mm; b – length 2.4 mm); c, d – dorsal view of head; e – ventral view of head and anterior portion of prosternum; f – prosternal process and mesoventrite; g – anterior portion of scutellum and microsetae on middle of posterior margin of pronotum; h – exposed portion of last visible abdominal ventrite; k – caudal marginal line of metacoxal cavity; m – last tarsomere of middle leg. Scale bars: Figs. d, f, h, k, m = 100 μm; Fig. g = 20 μm." captionText-1="Fig. 18. JelinekigethesAudisio & Cline, gen. nov.: a–e – J. danielssoni (Audisio, 1995).a – male habitus; b–c – male genitalia; d – ovipositor; e – male protibia. Figs. a–e – refer to AUDISIO (1995) for scale." captionText-2="Fig. 19. Astylogethes Kirejtshuk, 1992: a–h – A. subrugosus (Gyllenhal, 1808). a – male habitus (length 2.4 mm); b – dorsal view of head; c – ventral view of head and anterior portion of prosternum; d – prosternal process and mesoventrite; e – outer margin of protibia; f – exposed portion of last visible abdominal ventrite; g – scutellum and microsetae on middle of posterior margin of pronotum; h – caudal marginal line of metacoxal cavity. Scale bars: Figs. c, d, f = 100 μm; Fig. e = 20 μm." captionText-3="Fig. 20. Stachygethes Audisio & Cline, gen. nov.: a, c, f, k – S. ruficornis (Marsham, 1802); b – S. variolosus (Easton, 1964); d, e, g, h – S. assimilis (Sturm, 1845). a, b – male habitus (a – length 2.5 mm; b – length 2.5 mm); c – dorsal view of head; d – ventral view of head and anterior portion of prosternum; e – scutellum and microsetae on middle of posterior margin of pronotum; f – caudal marginal line of metacoxal cavity; g – prosternal process and mesoventrite; h – exposed portion of last visible abdominal ventrite; k – outer margin of mesotibia. Scale bars: Figs. d, f, g, h = 100 μm; Fig. e = 30 μm; Fig. k = 20 μm." figureDoi-0="http://doi.org/10.5281/zenodo.5363178" figureDoi-1="http://doi.org/10.5281/zenodo.5363180" figureDoi-2="http://doi.org/10.5281/zenodo.5363182" figureDoi-3="http://doi.org/10.5281/zenodo.5363184" httpUri-0="https://zenodo.org/record/5363178/files/figure.png" httpUri-1="https://zenodo.org/record/5363180/files/figure.png" httpUri-2="https://zenodo.org/record/5363182/files/figure.png" httpUri-3="https://zenodo.org/record/5363184/files/figure.png" pageId="17" pageNumber="358">
|
||
Figs.
|
||
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in
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<bibRefCitation id="EF864B2BF67BFF95B8C1FA48FEAAFA48" author="KIREJTSHUK A. G. & AUDISIO P." pageId="17" pageNumber="358" pagination="191 - 254" refId="ref90005" refString="KIREJTSHUK A. G. & AUDISIO P. 1995: Preliminary revision of South African Meligethes subgenus Lariopsis (Coleoptera: Nitidulidae, Meligethinae). Fragmenta Entomologica 27: 191 - 254." type="journal article" year="1995">KIREJTSHUK & AUDISIO 1995</bibRefCitation>
|
||
), with nearly transversely truncate distal margin, and without deep median longitudinal desclerotization from proximal portion of tegmen extending to medial distal Vshaped excision; median lobe of aedeagus without lateral emargination, subtruncate to bluntly acuminate distally, with arcuate shallow emargination distally.
|
||
</paragraph>
|
||
<paragraph id="8BA836DAF678FF96BBBEFF29FC2BFE2F" blockId="18.[151,1127,220,404]" pageId="18" pageNumber="359">
|
||
<emphasis id="B963EAC8F678FF96BBBEFF29FEDDFF4F" box="[181,360,221,244]" italics="true" pageId="18" pageNumber="359">Female genitalia</emphasis>
|
||
(
|
||
<emphasis id="B963EAC8F678FF96BA7EFF29FE55FF4F" box="[373,480,221,244]" italics="true" pageId="18" pageNumber="359">ovipositor</emphasis>
|
||
): moderately large; styli long and distinct, simple, cylindrical, more distinctly pigmented distally, inserted close to apex of contiguous gonostyloids; each gonostyloid moderately sclerotized and pigmented distally, with a simple, never indentate outer portion of basicoxite (Fig.
|
||
<quantity id="4CEF9B3FF678FF96BABFFEC8FE5FFEEF" box="[436,490,316,340]" metricMagnitude="0" metricUnit="m" metricValue="1.1938" pageId="18" pageNumber="359" unit="in" value="47.0">47 in</quantity>
|
||
<bibRefCitation id="EF864B2BF678FF96BAFBFEC8FCACFEE8" author="KIREJTSHUK A. G. & AUDISIO P." box="[496,793,315,340]" pageId="18" pageNumber="359" pagination="191 - 254" refId="ref90005" refString="KIREJTSHUK A. G. & AUDISIO P. 1995: Preliminary revision of South African Meligethes subgenus Lariopsis (Coleoptera: Nitidulidae, Meligethinae). Fragmenta Entomologica 27: 191 - 254." type="journal article" year="1995">KIREJTSHUK & AUDISIO 1995</bibRefCitation>
|
||
), and a single, narrow, scarcely pigmented and sclerotized arcuate area along outer subdistal portion of gonostyloids. ‘Central point’ of ovipositor centrally located, without proximad directed spicule.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF686652F678FF96BB9CFA20FBB4F9E8" ID-DOI="http://doi.org/10.5281/zenodo.5319344" ID-Zenodo-Dep="5319344" httpUri="https://zenodo.org/record/5319344/files/figure.png" pageId="18" pageNumber="359" startId="18.[151,182,1492,1512]" targetBox="[168,1109,431,1469]" targetPageId="18">
|
||
<paragraph id="8BA836DAF678FF96BB9CFA20FBB4F9E8" blockId="18.[151,1129,1492,1619]" pageId="18" pageNumber="359">
|
||
<emphasis id="B963EAC8F678FF96BB9CFA20FF7BFA53" bold="true" box="[151,206,1492,1512]" pageId="18" pageNumber="359">Fig. 4.</emphasis>
|
||
<taxonomicName id="4C174D59F678FF96BBD9FA22FE0DFA53" authority="Kirejtshuk, 1989" authorityName="Kirejtshuk" authorityYear="1989" box="[210,440,1493,1512]" class="Insecta" family="Nitidulidae" genus="Lariopsis" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="359" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B963EAC8F678FF96BBD9FA22FE96FA53" box="[210,291,1494,1512]" italics="true" pageId="18" pageNumber="359">Lariopsis</emphasis>
|
||
<bibRefCitation id="EF864B2BF678FF96BA2CFA21FE0DFA53" author="KIREJTSHUK A. G." box="[295,440,1493,1512]" pageId="18" pageNumber="359" pagination="64 - 89" refId="ref89287" refString="KIREJTSHUK A. G. 1989: Novye taksony zhukov-blestyanok (Coleoptera, Nitidulidae) vostochnogo polushariya (Chast' III). [New taxa of the Nitidulidae (Coleoptera) of the East hemisphere (Part III)]. Trudy Zoologicheskogo Instituta, Akademiya Nauk SSSR 208: 64 - 89 (in Russian)." type="journal article" year="1989">Kirejtshuk, 1989</bibRefCitation>
|
||
</taxonomicName>
|
||
:
|
||
<emphasis id="B963EAC8F678FF96BAB4FA21FE7FFA53" bold="true" box="[447,458,1493,1512]" pageId="18" pageNumber="359">a</emphasis>
|
||
–
|
||
<taxonomicName id="4C174D59F678FF96BAD7FA22FC8AFA53" authority="(Kirejtshuk & Audisio, 1995)" authorityName="Audisio & Cline & Biase & Antonini & Mancini & Trizzino & Costantini & Strika & Lamanna & Cerretti" authorityYear="2009" baseAuthorityName="Kirejtshuk & Audisio" baseAuthorityYear="1995" box="[476,831,1493,1512]" class="Insecta" family="Nitidulidae" genus="Lariopsis" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="359" phylum="Arthropoda" rank="species" species="vultuosus">
|
||
<emphasis id="B963EAC8F678FF96BAD7FA22FDF7FA53" box="[476,578,1494,1512]" italics="true" pageId="18" pageNumber="359">L.vultuosus</emphasis>
|
||
(
|
||
<bibRefCitation id="EF864B2BF678FF96B947FA21FC8FFA53" author="KIREJTSHUK A. G. & AUDISIO P." box="[588,826,1493,1512]" pageId="18" pageNumber="359" pagination="191 - 254" refId="ref90005" refString="KIREJTSHUK A. G. & AUDISIO P. 1995: Preliminary revision of South African Meligethes subgenus Lariopsis (Coleoptera: Nitidulidae, Meligethinae). Fragmenta Entomologica 27: 191 - 254." type="journal article" year="1995">Kirejtshuk & Audisio, 1995</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
;
|
||
<emphasis id="B963EAC8F678FF96B84CFA21FCDFFA53" bold="true" box="[839,874,1493,1512]" pageId="18" pageNumber="359">b–k</emphasis>
|
||
–
|
||
<taxonomicName id="4C174D59F678FF96B877FA22FBD0FA53" authority="(Reitter, 1872)" authorityName="Audisio & Cline & Biase & Antonini & Mancini & Trizzino & Costantini & Strika & Lamanna & Cerretti" authorityYear="2009" baseAuthorityName="Reitter" baseAuthorityYear="1872" box="[892,1125,1493,1512]" class="Insecta" family="Nitidulidae" genus="Lariopsis" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="359" phylum="Arthropoda" rank="species" species="variabilis">
|
||
<emphasis id="B963EAC8F678FF96B877FA22FC51FA53" box="[892,996,1494,1512]" italics="true" pageId="18" pageNumber="359">L. variabilis</emphasis>
|
||
(Reitter, 1872)
|
||
</taxonomicName>
|
||
.
|
||
<emphasis id="B963EAC8F678FF96BB9CFA04FF17F9B8" bold="true" box="[151,162,1520,1539]" pageId="18" pageNumber="359">a</emphasis>
|
||
– male habitus (length 3.3 mm);
|
||
<emphasis id="B963EAC8F678FF96BAB0FA04FE72F9B8" bold="true" box="[443,455,1520,1539]" pageId="18" pageNumber="359">b</emphasis>
|
||
,
|
||
<emphasis id="B963EAC8F678FF96BADAFA04FE6FF9B8" bold="true" box="[465,474,1520,1539]" pageId="18" pageNumber="359">c</emphasis>
|
||
– dorso-lateral view of head;
|
||
<emphasis id="B963EAC8F678FF96B9DCFA04FD56F9B8" bold="true" box="[727,739,1520,1539]" pageId="18" pageNumber="359">d</emphasis>
|
||
– ventral view of head and anterior portion of prosternum;
|
||
<emphasis id="B963EAC8F678FF96BA08F9FFFEB9F9A5" bold="true" box="[259,268,1547,1566]" pageId="18" pageNumber="359">e</emphasis>
|
||
– prosternal process and mesoventrite;
|
||
<emphasis id="B963EAC8F678FF96B955F9FEFDD0F9A5" bold="true" box="[606,613,1546,1566]" pageId="18" pageNumber="359">f</emphasis>
|
||
– middle leg with outer margin of mesotibia;
|
||
<emphasis id="B963EAC8F678FF96B8E7F9FFFC42F9A5" bold="true" box="[1004,1015,1547,1566]" pageId="18" pageNumber="359">g</emphasis>
|
||
– microsetae on middle posterior margin of pronotum;
|
||
<emphasis id="B963EAC8F678FF96BAFEF9D1FDB4F983" bold="true" box="[501,513,1573,1592]" pageId="18" pageNumber="359">h</emphasis>
|
||
– exposed portion of last visible abdominal ventrite;
|
||
<emphasis id="B963EAC8F678FF96B8CAF9D1FC78F983" bold="true" box="[961,973,1573,1592]" pageId="18" pageNumber="359">k</emphasis>
|
||
– caudal marginal lines of metacoxal cavities. Scale bars: Figs.
|
||
<emphasis id="B963EAC8F678FF96B91DF9B4FD97F9E8" bold="true" box="[534,546,1600,1619]" pageId="18" pageNumber="359">b</emphasis>
|
||
,
|
||
<emphasis id="B963EAC8F678FF96B926F9B4FD83F9E8" bold="true" box="[557,566,1600,1619]" pageId="18" pageNumber="359">c</emphasis>
|
||
,
|
||
<emphasis id="B963EAC8F678FF96B94AF9B4FDF8F9E8" bold="true" box="[577,589,1600,1619]" pageId="18" pageNumber="359">d</emphasis>
|
||
,
|
||
<emphasis id="B963EAC8F678FF96B953F9B4FDD4F9E8" bold="true" box="[600,609,1600,1619]" pageId="18" pageNumber="359">e</emphasis>
|
||
,
|
||
<emphasis id="B963EAC8F678FF96B967F9B4FDCDF9E8" bold="true" box="[620,632,1600,1619]" pageId="18" pageNumber="359">h</emphasis>
|
||
,
|
||
<emphasis id="B963EAC8F678FF96B989F9B4FD3BF9E8" bold="true" box="[642,654,1600,1619]" pageId="18" pageNumber="359">k</emphasis>
|
||
= 100 μm; Fig.
|
||
<emphasis id="B963EAC8F678FF96B812F9CBFC95F9E8" bold="true" box="[793,800,1599,1619]" pageId="18" pageNumber="359">f</emphasis>
|
||
= 30 μm; Fig.
|
||
<emphasis id="B963EAC8F678FF96B8AAF9B4FC19F9E8" bold="true" box="[929,940,1600,1619]" pageId="18" pageNumber="359">g</emphasis>
|
||
= 10 μm.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C30D6551F679FF97BB7AFF28FF50FECF" pageId="19" pageNumber="360" type="etymology">
|
||
<paragraph id="8BA836DAF679FF97BB7AFF28FF50FECF" blockId="19.[113,1089,219,724]" pageId="19" pageNumber="360">
|
||
<emphasis id="B963EAC8F679FF97BB7AFF28FF47FF4F" bold="true" box="[113,242,220,244]" pageId="19" pageNumber="360">Etymology.</emphasis>
|
||
The generic name was derived from
|
||
<taxonomicName id="4C174D59F679FF97B980FF29FCD2FF48" authority="Scopoli, 1763" authorityName="Scopoli" authorityYear="1763" box="[651,871,219,244]" class="Insecta" family="Nitidulidae" genus="Laria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="360" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B963EAC8F679FF97B980FF29FD73FF4F" box="[651,710,221,244]" italics="true" pageId="19" pageNumber="360">Laria</emphasis>
|
||
<bibRefCitation id="EF864B2BF679FF97B9C4FF28FCD2FF48" author="SCOPOLI J." box="[719,871,219,244]" pageId="19" pageNumber="360" refId="ref91932" refString="SCOPOLI J. 1763: Entomologia Carniolica exhibens Insecta Carniolae indigena et distributa in ordines, genera, species, varietates. Methodo Linnaeana. Joannis Thomae Trattner, Vindobonae [Wien], xxxii + 420 pp." type="book" year="1763">Scopoli, 1763</bibRefCitation>
|
||
</taxonomicName>
|
||
, the ancient generic name of members of the present-day
|
||
<taxonomicName id="4C174D59F679FF97BAF1FF08FD32FEAF" authorityName="C. G. Thomson" authorityYear="1859" box="[506,647,252,276]" class="Insecta" family="Nitidulidae" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="360" phylum="Arthropoda" rank="subFamily" subFamily="Meligethinae">Meligethinae</taxonomicName>
|
||
genus
|
||
<taxonomicName id="4C174D59F679FF97B9DAFF09FC1EFEAF" authority="Stephens, 1829" authorityName="Stephens" authorityYear="1829" box="[721,939,252,276]" class="Insecta" family="Nitidulidae" genus="Pria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="360" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B963EAC8F679FF97B9DAFF09FCB5FEAF" box="[721,768,253,276]" italics="true" pageId="19" pageNumber="360">Pria</emphasis>
|
||
Stephens, 1829
|
||
</taxonomicName>
|
||
, to emphasize the general aspect of a few, small-sized, and yellowish-colored species of
|
||
<taxonomicName id="4C174D59F679FF97B88FFEE9FBB9FE8F" authority="Audisio & Cline & Biase & Antonini & Mancini & Trizzino & Costantini & Strika & Lamanna & Cerretti, 2009" authorityName="Audisio & Cline & Biase & Antonini & Mancini & Trizzino & Costantini & Strika & Lamanna & Cerretti" authorityYear="2009" box="[900,1036,285,308]" class="Insecta" family="Nitidulidae" genus="Neolariopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="360" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B963EAC8F679FF97B88FFEE9FBB9FE8F" box="[900,1036,285,308]" italics="true" pageId="19" pageNumber="360">Neolariopsis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25057B3F679FF97BF18FEE8FF2AFEE8" pageId="19" pageNumber="360" rank="genus">gen. nov.</taxonomicNameLabel>
|
||
(previously included in
|
||
<taxonomicName id="4C174D59F679FF97BAABFEC9FDB0FEEF" authorityName="Kirejtshuk" authorityYear="1989" box="[416,517,317,340]" class="Insecta" family="Nitidulidae" genus="Lariopsis" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="360" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B963EAC8F679FF97BAABFEC9FDB0FEEF" box="[416,517,317,340]" italics="true" pageId="19" pageNumber="360">Lariopsis</emphasis>
|
||
</taxonomicName>
|
||
) that superficially resemble members of
|
||
<taxonomicName id="4C174D59F679FF97B8BDFEC9FC50FEEF" authorityName="Stephens" authorityYear="1829" box="[950,997,317,340]" class="Insecta" family="Nitidulidae" genus="Pria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="360" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B963EAC8F679FF97B8BDFEC9FC50FEEF" box="[950,997,317,340]" italics="true" pageId="19" pageNumber="360">Pria</emphasis>
|
||
</taxonomicName>
|
||
. Gender masculine.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30D6551F679FF97BB7AFE8FFC11FCAC" pageId="19" pageNumber="360" type="biology_ecology">
|
||
<paragraph id="8BA836DAF679FF97BB7AFE8FFD5BFE6F" blockId="19.[113,1089,219,724]" pageId="19" pageNumber="360">
|
||
<emphasis id="B963EAC8F679FF97BB7AFE8FFF78FE28" bold="true" box="[113,205,379,403]" pageId="19" pageNumber="360">Biology.</emphasis>
|
||
The two inclusive species are strictly associated for larval development with inflorescences (capitula) of
|
||
<taxonomicName id="4C174D59F679FF97BA67FE68FE51FE0F" box="[364,484,412,436]" class="Magnoliopsida" family="Asteraceae" kingdom="Plantae" order="Asterales" pageId="19" pageNumber="360" phylum="Tracheophyta" rank="family">Asteraceae</taxonomicName>
|
||
, e.g. those of
|
||
<taxonomicName id="4C174D59F679FF97B975FE69FD64FE0F" box="[638,721,413,436]" class="Magnoliopsida" family="Asteraceae" genus="Arctotis" kingdom="Plantae" order="Asterales" pageId="19" pageNumber="360" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="B963EAC8F679FF97B975FE69FD64FE0F" box="[638,721,413,436]" italics="true" pageId="19" pageNumber="360">Arctotis</emphasis>
|
||
</taxonomicName>
|
||
L.,
|
||
<taxonomicName id="4C174D59F679FF97B80BFE69FC7EFE0F" authority="Wendl." authorityName="Wendl." box="[768,971,412,436]" class="Magnoliopsida" family="Asteraceae" genus="Arctotheca" kingdom="Plantae" order="Asterales" pageId="19" pageNumber="360" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="B963EAC8F679FF97B80BFE69FCC1FE0F" box="[768,884,413,436]" italics="true" pageId="19" pageNumber="360">Arctotheca</emphasis>
|
||
Wendl.
|
||
</taxonomicName>
|
||
, and allied genera (
|
||
<bibRefCitation id="EF864B2BF679FF97BBCCFE48FE44FE68" author="KIREJTSHUK A. G. & AUDISIO P." box="[199,497,443,468]" pageId="19" pageNumber="360" pagination="191 - 254" refId="ref90005" refString="KIREJTSHUK A. G. & AUDISIO P. 1995: Preliminary revision of South African Meligethes subgenus Lariopsis (Coleoptera: Nitidulidae, Meligethinae). Fragmenta Entomologica 27: 191 - 254." type="journal article" year="1995">KIREJTSHUK & AUDISIO 1995</bibRefCitation>
|
||
, and unpublished data).
|
||
</paragraph>
|
||
<paragraph id="8BA836DAF679FF97BB7AFE2FFEA3FD2F" blockId="19.[113,1089,219,724]" pageId="19" pageNumber="360">
|
||
<emphasis id="B963EAC8F679FF97BB7AFE2FFED1FE48" bold="true" box="[113,356,475,499]" pageId="19" pageNumber="360">Phylogenetic position.</emphasis>
|
||
<taxonomicName id="4C174D59F679FF97BA62FE29FE79FE4F" authorityName="Kirejtshuk" authorityYear="1989" box="[361,460,477,500]" class="Insecta" family="Nitidulidae" genus="Lariopsis" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="360" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B963EAC8F679FF97BA62FE29FE79FE4F" box="[361,460,477,500]" italics="true" pageId="19" pageNumber="360">Lariopsis</emphasis>
|
||
</taxonomicName>
|
||
is more closely related to
|
||
<taxonomicName id="4C174D59F679FF97B9DEFE29FCEEFE4F" authority="Audisio & Cline & Biase & Antonini & Mancini & Trizzino & Costantini & Strika & Lamanna & Cerretti, 2009" authorityName="Audisio & Cline & Biase & Antonini & Mancini & Trizzino & Costantini & Strika & Lamanna & Cerretti" authorityYear="2009" box="[725,859,477,500]" class="Insecta" family="Nitidulidae" genus="Neolariopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="360" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B963EAC8F679FF97B9DEFE29FCEEFE4F" box="[725,859,477,500]" italics="true" pageId="19" pageNumber="360">Neolariopsis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25057B3F679FF97B854FE28FC0AFE48" box="[863,959,476,500]" pageId="19" pageNumber="360" rank="genus">gen. nov.</taxonomicNameLabel>
|
||
,
|
||
<taxonomicName id="4C174D59F679FF97B8C3FE29FF0FFDAF" authority="Audisio & Cline & Biase & Antonini & Mancini & Trizzino & Costantini & Strika & Lamanna & Cerretti, 2009" authorityName="Audisio & Cline & Biase & Antonini & Mancini & Trizzino & Costantini & Strika & Lamanna & Cerretti" authorityYear="2009" class="Insecta" family="Nitidulidae" genus="Odontholariopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="360" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B963EAC8F679FF97B8C3FE29FF0FFDAF" italics="true" pageId="19" pageNumber="360">Odontholariopsis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25057B3F679FF97BBB4FE08FE97FDA8" box="[191,290,508,532]" pageId="19" pageNumber="360" rank="genus">gen. nov.</taxonomicNameLabel>
|
||
, and
|
||
<taxonomicName id="4C174D59F679FF97BA5CFE09FE55FDAF" authority="Audisio & Cline & Biase & Antonini & Mancini & Trizzino & Costantini & Strika & Lamanna & Cerretti, 2009" authorityName="Audisio & Cline & Biase & Antonini & Mancini & Trizzino & Costantini & Strika & Lamanna & Cerretti" authorityYear="2009" box="[343,480,509,532]" class="Insecta" family="Nitidulidae" genus="Asterogethes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="360" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B963EAC8F679FF97BA5CFE09FE55FDAF" box="[343,480,509,532]" italics="true" pageId="19" pageNumber="360">Asterogethes</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25057B3F679FF97BAEDFE08FDFCFDA8" box="[486,585,508,532]" pageId="19" pageNumber="360" rank="genus">gen. nov.</taxonomicNameLabel>
|
||
; collectively forming a newly defined
|
||
<taxonomicName id="4C174D59F679FF97B8D7FE09FBF4FDAF" authorityName="Kirejtshuk" authorityYear="1989" box="[988,1089,509,532]" class="Insecta" family="Nitidulidae" genus="Lariopsis" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="360" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B963EAC8F679FF97B8D7FE09FBF4FDAF" box="[988,1089,509,532]" italics="true" pageId="19" pageNumber="360">Lariopsis</emphasis>
|
||
</taxonomicName>
|
||
generic complex, which is supported by both adult morphology and molecular data (
|
||
<bibRefCitation id="EF864B2BF679FF97B8EEFDE8FF50FDEF" author="TRIZZINO M. & AUDISIO P. & ANTONINI G. & DE BIASE A. & MANCINI E." pageId="19" pageNumber="360" pagination="215 - 226" refId="ref92518" refString="TRIZZINO M., AUDISIO P., ANTONINI G., DE BIASE A. & MANCINI E. 2009: Comparative analysis of sequences and secondary structures of the rRNA internal transcribed spacer 2 (ITS 2) in pollen-beetles of the subfamily Meligethinae (Coleoptera, Nitidulidae): potential use of slippage-derived sequences in molecular systematics. Molecular Phylogenetics and Evolution 51: 215 - 226." type="journal article" year="2009">TRIZZINO et al. 2009</bibRefCitation>
|
||
). Phylogenetic relationships of
|
||
<taxonomicName id="4C174D59F679FF97B938FDC9FD2DFDEF" authorityName="Kirejtshuk" authorityYear="1989" box="[563,664,573,596]" class="Insecta" family="Nitidulidae" genus="Lariopsis" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="360" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B963EAC8F679FF97B938FDC9FD2DFDEF" box="[563,664,573,596]" italics="true" pageId="19" pageNumber="360">Lariopsis</emphasis>
|
||
</taxonomicName>
|
||
with
|
||
<emphasis id="B963EAC8F679FF97B9D8FDC9FBA1FDEF" box="[723,1044,573,596]" italics="true" pageId="19" pageNumber="360">
|
||
<taxonomicName id="4C174D59F679FF97B9D8FDC9FCC6FDEF" authorityName="Reitter" authorityYear="1871" box="[723,883,573,596]" class="Insecta" family="Nitidulidae" genus="Acanthogethes" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="360" phylum="Arthropoda" rank="genus">Acanthogethes</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C174D59F679FF97B876FDC9FBBAFDEF" authorityName="Scholz" authorityYear="1932" box="[893,1039,573,596]" class="Insecta" family="Nitidulidae" genus="Clypeogethes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="360" phylum="Arthropoda" rank="genus">Clypeogethes</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
and other ancestral lineages of
|
||
<taxonomicName id="4C174D59F679FF97BA87FDA8FDACFDCF" authorityName="C. G. Thomson" authorityYear="1859" box="[396,537,604,628]" class="Insecta" family="Nitidulidae" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="360" phylum="Arthropoda" rank="subFamily" subFamily="Meligethinae">Meligethinae</taxonomicName>
|
||
remain unclear, and are only weakly supported with molecular data.
|
||
</paragraph>
|
||
<paragraph id="8BA836DAF679FF97BB7AFD68FDC9FD68" blockId="19.[113,1089,219,724]" pageId="19" pageNumber="360">
|
||
<emphasis id="B963EAC8F679FF97BB7AFD68FD91FD08" bold="true" box="[113,548,667,692]" pageId="19" pageNumber="360">Taxonomy and geographic distribution.</emphasis>
|
||
<taxonomicName id="4C174D59F679FF97B922FD69FD39FD0F" authorityName="Kirejtshuk" authorityYear="1989" box="[553,652,669,692]" class="Insecta" family="Nitidulidae" genus="Lariopsis" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="360" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B963EAC8F679FF97B922FD69FD39FD0F" box="[553,652,669,692]" italics="true" pageId="19" pageNumber="360">Lariopsis</emphasis>
|
||
</taxonomicName>
|
||
includes two described species distributed in Southern Africa (
|
||
<bibRefCitation id="EF864B2BF679FF97BA4EFD48FDDAFD68" author="KIREJTSHUK A. G. & AUDISIO P." box="[325,623,699,724]" pageId="19" pageNumber="360" pagination="191 - 254" refId="ref90005" refString="KIREJTSHUK A. G. & AUDISIO P. 1995: Preliminary revision of South African Meligethes subgenus Lariopsis (Coleoptera: Nitidulidae, Meligethinae). Fragmenta Entomologica 27: 191 - 254." type="journal article" year="1995">KIREJTSHUK & AUDISIO 1995</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BA836DAF679FF97BB7AFD1EFCDBFD47" blockId="19.[113,932,745,791]" box="[113,878,745,764]" pageId="19" pageNumber="360">
|
||
<taxonomicName id="4C174D59F679FF97BB7AFD1EFE2EFD47" authority="(Reitter, 1872)" authorityName="Audisio & Cline & Biase & Antonini & Mancini & Trizzino & Costantini & Strika & Lamanna & Cerretti" authorityYear="2009" baseAuthorityName="Reitter" baseAuthorityYear="1872" box="[113,411,745,764]" class="Insecta" family="Nitidulidae" genus="Lariopsis" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="360" phylum="Arthropoda" rank="species" species="variabilis" status="comb. nov.">
|
||
<emphasis id="B963EAC8F679FF97BB7AFD1EFEACFD47" box="[113,281,746,764]" italics="true" pageId="19" pageNumber="360">Lariopsis variabilis</emphasis>
|
||
(Reitter, 1872)
|
||
</taxonomicName>
|
||
<emphasis id="B963EAC8F679FF97BAABFD1DFDB7FD47" bold="true" box="[416,514,745,764]" pageId="19" pageNumber="360">
|
||
<taxonomicNameLabel id="A25057B3F679FF97BAABFD1DFDB7FD47" box="[416,514,745,764]" pageId="19" pageNumber="360" rank="species">comb. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
<collectingCountry id="F300764AF679FF97B9BAFD1DFC94FD47" box="[689,801,745,764]" name="South Africa" pageId="19" pageNumber="360">South Africa</collectingCountry>
|
||
: W
|
||
<collectingRegion id="49D3F838F679FF97B849FD1DFCDBFD47" box="[834,878,745,764]" country="South Africa" name="Western Cape" pageId="19" pageNumber="360">Cape</collectingRegion>
|
||
</paragraph>
|
||
<paragraph id="8BA836DAF679FF97BB7AFCF1FC11FCAC" blockId="19.[113,932,745,791]" box="[113,932,772,791]" pageId="19" pageNumber="360">
|
||
<taxonomicName id="4C174D59F679FF97BB7AFCF1FDADFCAC" authority="(Kirejtshuk & Audisio, 1995)" authorityName="Audisio & Cline & Biase & Antonini & Mancini & Trizzino & Costantini & Strika & Lamanna & Cerretti" authorityYear="2009" baseAuthorityName="Kirejtshuk & Audisio" baseAuthorityYear="1995" box="[113,536,772,791]" class="Insecta" family="Nitidulidae" genus="Lariopsis" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="360" phylum="Arthropoda" rank="species" species="vultuosus" status="comb. nov.">
|
||
<emphasis id="B963EAC8F679FF97BB7AFCF1FEADFCAC" box="[113,280,773,791]" italics="true" pageId="19" pageNumber="360">Lariopsis vultuosus</emphasis>
|
||
(
|
||
<bibRefCitation id="EF864B2BF679FF97BA28FCF0FDA6FCAC" author="KIREJTSHUK A. G. & AUDISIO P." box="[291,531,772,791]" pageId="19" pageNumber="360" pagination="191 - 254" refId="ref90005" refString="KIREJTSHUK A. G. & AUDISIO P. 1995: Preliminary revision of South African Meligethes subgenus Lariopsis (Coleoptera: Nitidulidae, Meligethinae). Fragmenta Entomologica 27: 191 - 254." type="journal article" year="1995">Kirejtshuk & Audisio, 1995</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
<emphasis id="B963EAC8F679FF97B915FCF0FD35FCAC" bold="true" box="[542,640,772,791]" pageId="19" pageNumber="360">
|
||
<taxonomicNameLabel id="A25057B3F679FF97B915FCF0FD35FCAC" box="[542,640,772,791]" pageId="19" pageNumber="360" rank="species">comb. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
<collectingCountry id="F300764AF679FF97B9BAFCF0FC94FCAC" box="[689,801,772,791]" name="South Africa" pageId="19" pageNumber="360">South Africa</collectingCountry>
|
||
: W and E
|
||
<collectingRegion id="49D3F838F679FF97B873FCF0FC11FCAC" box="[888,932,772,791]" country="South Africa" name="Western Cape" pageId="19" pageNumber="360">Cape</collectingRegion>
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |