174 lines
20 KiB
XML
174 lines
20 KiB
XML
<document id="C141CF92674767B863A2EE4CFFF0EF90" ID-DOI="10.11646/zootaxa.2851.1.1" ID-ISSN="11755334" ID-Zenodo-Dep="5286934" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629721501414" checkinUser="felipe" docAuthor="Blahnik, Roger J. & Holzenthal, Ralph W." docDate="2011" docId="03BE87970065FF8298B1FEE6FDF8C3B5" docLanguage="en" docName="zt02851p075.pdf" docOrigin="Zootaxa 2851 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.2851.1.1" docStyle="DocumentStyle:D239614CE4198176A422035174489AB1.4:Zootaxa.2009-2012.monograph" docStyleId="D239614CE4198176A422035174489AB1" docStyleName="Zootaxa.2009-2012.monograph" docStyleVersion="4" docTitle="Mortoniella bocaina Blahnik & Holzenthal 2011, new species" docType="treatment" docVersion="4" lastPageNumber="55" masterDocId="FF87FFEF0053FFB49826FF89FFBAC62F" masterDocTitle="Revision of the austral South American species of Mortoniella (Trichoptera: Glossosomatidae: Protoptilinae) 2851" masterLastPageNumber="75" masterPageNumber="1" pageNumber="55" updateTime="1699101759394" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:mods id="30E82CCB97457121A6874C189B418C3C" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="53DB671EE53581150AA4D02741BAC8D0">
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<mods:title id="EC0D236056D552C20319EFBED56D72D9">Revision of the austral South American species of Mortoniella (Trichoptera: Glossosomatidae: Protoptilinae) 2851</mods:title>
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<mods:name id="6BD217750E0EFA9075EC6D6C9D645227" type="personal">
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<mods:role id="986F446ABC70E36E76A8E8F1F073B731">
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<mods:roleTerm id="232E06951003C199707116F15B173DDD">Author</mods:roleTerm>
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<mods:namePart id="9DB761B3543E65C16706247C0EF14A98">Blahnik, Roger J.</mods:namePart>
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<mods:name id="3DBBED5B434B237FB9436CDEFF85D3EE" type="personal">
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<mods:roleTerm id="F3DA26B3296CAA589084BC6604F1B263">Author</mods:roleTerm>
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<mods:namePart id="85653875891BC75DEC8D7983075AA3A1">Holzenthal, Ralph W.</mods:namePart>
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<mods:typeOfResource id="3FF51D5E10583322BE5D826A2148972F">text</mods:typeOfResource>
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<mods:title id="85F3AF41CA3F6D6DF592AB900F0A5261">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="99E22E448BEE7ABDCDE9ADBCC8E790F4">
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<mods:date id="BF7695D435BF40E45506CE82587E0FF8">2011</mods:date>
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<mods:detail id="A9FF0A7870342F1775DF6B385C5E536A" type="pubDate">
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<mods:number id="886241EB8269C6D6103139E6176EA744">2011-04-29</mods:number>
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<mods:detail id="52952A0A05819D94347FE706CEDC5D85" type="volume">
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<mods:number id="CEB560249F6AB05FCBFBDD39CB7B1805">2851</mods:number>
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</mods:detail>
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<mods:detail id="7F328C777A90B82E2C8E84E133E2E3D2" type="issue">
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<mods:location id="2CF37BE18FF7CA041E957E5D9289F972">
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<mods:url id="5DD8AC32C3247AD63F18E5ACFFC485E0">https://biotaxa.org/Zootaxa/article/view/zootaxa.2851.1.1</mods:url>
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<mods:classification id="2082FE06C2539AF50B85D77CA025EA20">journal article</mods:classification>
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<mods:identifier id="48086CD0D8DC9621A0674653C4507678" type="DOI">10.11646/zootaxa.2851.1.1</mods:identifier>
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<mods:identifier id="903CF24FA15BE6854788250F6ACCD36D" type="ISSN">11755334</mods:identifier>
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<mods:identifier id="DEA6A1202DE5DF9490FE17B9FBAEA451" type="Zenodo-Dep">5286934</mods:identifier>
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<treatment id="03BE87970065FF8298B1FEE6FDF8C3B5" ID-DOI="http://doi.org/10.5281/zenodo.5293506" ID-Zenodo-Dep="5293506" LSID="urn:lsid:plazi:treatment:03BE87970065FF8298B1FEE6FDF8C3B5" httpUri="http://treatment.plazi.org/id/03BE87970065FF8298B1FEE6FDF8C3B5" lastPageNumber="55" pageId="54" pageNumber="55">
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<subSubSection id="C30D650A0065FF8298B1FEE6FD96C7A6" box="[151,556,367,393]" pageId="54" pageNumber="55" type="nomenclature">
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<paragraph id="8BA836810065FF8298B1FEE6FD96C7A6" blockId="54.[151,556,367,426]" box="[151,556,367,393]" pageId="54" pageNumber="55">
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<heading id="D0E081ED0065FF8298B1FEE6FD96C7A6" bold="true" box="[151,556,367,393]" fontSize="11" level="1" pageId="54" pageNumber="55" reason="1">
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<emphasis id="B963EA930065FF8298B1FEE6FD96C7A6" bold="true" box="[151,556,367,393]" pageId="54" pageNumber="55">
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<taxonomicName id="4C174D020065FF8298B1FEE6FE37C7A6" authority="Blahnik & Holzenthal, 2011" authorityName="Blahnik & Holzenthal" authorityYear="2011" box="[151,397,367,393]" class="Magnoliopsida" family="Apocynaceae" genus="Mortoniella" kingdom="Plantae" order="Gentianales" pageId="54" pageNumber="55" phylum="Tracheophyta" rank="species" species="bocaina" status="sp. nov.">
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<emphasis id="B963EA930065FF8298B1FEE6FE37C7A6" bold="true" box="[151,397,367,393]" italics="true" pageId="54" pageNumber="55">Mortoniella bocaina</emphasis>
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</taxonomicName>
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,
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<taxonomicNameLabel id="A25057E80065FF8299BAFEE6FD96C7A6" box="[412,556,367,393]" pageId="54" pageNumber="55" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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||
</paragraph>
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||
</subSubSection>
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<subSubSection id="C30D650A0065FF8298B1FE1BFCBAC2ED" pageId="54" pageNumber="55" type="description">
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<paragraph id="8BA836810065FF8298B1FE1BFEA1C785" blockId="54.[151,556,367,426]" box="[151,283,402,426]" pageId="54" pageNumber="55">
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<figureCitation id="132C2A040065FF8298B1FE1BFF49C785" box="[151,243,402,426]" captionStart="FIGURE 27" captionStartId="55.[151,250,1976,1999]" captionTargetBox="[224,1351,208,1950]" captionTargetId="figure-44@55.[209,1378,193,1956]" captionTargetPageId="55" captionText="FIGURE 27. Mortoniella bocaina, new species, male genitalia: A—lateral; B—segment IX and tergum X, dorsal; C—phallic ensemble, ventral; D—dorsal phallic spine, dorsal; E—ventral process of segment VI, lateral." figureDoi="http://doi.org/10.5281/zenodo.5287037" httpUri="https://zenodo.org/record/5287037/files/figure.png" pageId="54" pageNumber="55">Figs. 27</figureCitation>
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,
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<figureCitation id="132C2A040065FF8298D9FE1BFEA1C785" box="[255,283,402,426]" captionStart="FIGURES 30–32" captionStartId="59.[151,264,1954,1977]" captionTargetBox="[275,1319,189,1940]" captionTargetId="figure-62@59.[249,1335,179,1954]" captionTargetPageId="59" captionText="FIGURES 30–32. Mortoniella velasquezi species group, female genitalia: 30, Mortoniella tripuiensis, new species: A—tergum IX, dorsal; B—ventral; C—tergum VII, dorsal. 31, Mortoniella froehlichi, new species: tergum IX, dorsal. 32, Mortoniella bocaina, new species: tergum IX, dorsal." figureDoi="http://doi.org/10.5281/zenodo.5287047" httpUri="https://zenodo.org/record/5287047/files/figure.png" pageId="54" pageNumber="55">32</figureCitation>
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</paragraph>
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<paragraph id="8BA836810065FF8298B1FE53FB1EC471" blockId="54.[151,1436,474,1434]" pageId="54" pageNumber="55">
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Like the other 2 new species of the
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<taxonomicName id="4C174D020065FF829A02FE52FD04C7DC" box="[548,702,475,499]" class="Magnoliopsida" family="Apocynaceae" genus="Mortoniella" kingdom="Plantae" order="Gentianales" pageId="54" pageNumber="55" phylum="Tracheophyta" rank="species" species="velasquezi">
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<emphasis id="B963EA930065FF829A02FE52FD81C7DD" box="[548,571,475,498]" italics="true" pageId="54" pageNumber="55">M</emphasis>
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.
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<emphasis id="B963EA930065FF829A6FFE52FD04C7DC" box="[585,702,475,499]" italics="true" pageId="54" pageNumber="55">velasquezi</emphasis>
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</taxonomicName>
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group from
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<collectingCountry id="F30076110065FF829B6CFE53FC34C7DD" box="[842,910,474,498]" name="Brazil" pageId="54" pageNumber="55">Brazil</collectingCountry>
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described here, this species lacks scales on the hind wings. It is perhaps most likely to be confused with
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<taxonomicName id="4C174D020065FF829B3FFE76FC0EC439" authority="Blahnik & Holzenthal, 2011" authorityName="Blahnik & Holzenthal" authorityYear="2011" box="[793,948,511,534]" class="Magnoliopsida" family="Apocynaceae" genus="Mortoniella" kingdom="Plantae" order="Gentianales" pageId="54" pageNumber="55" phylum="Tracheophyta" rank="species" species="tripuiensis" status="sp. nov.">
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<emphasis id="B963EA930065FF829B3FFE76FC0EC439" box="[793,948,511,534]" italics="true" pageId="54" pageNumber="55">M. tripuiensis</emphasis>
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</taxonomicName>
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,
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<taxonomicNameLabel id="A25057E80065FF829BE4FE76FBBAC439" box="[962,1024,511,534]" pageId="54" pageNumber="55" rank="species">n. sp.</taxonomicNameLabel>
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, since both species lack the elongate and diagnostic endophallic spines found in
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<taxonomicName id="4C174D020065FF829A5FFDAAFCB0C414" authority="Blahnik & Holzenthal, 2011" authorityName="Blahnik & Holzenthal" authorityYear="2011" box="[633,778,547,571]" class="Magnoliopsida" family="Apocynaceae" genus="Mortoniella" kingdom="Plantae" order="Gentianales" pageId="54" pageNumber="55" phylum="Tracheophyta" rank="species" species="froehlichi" status="sp. nov.">
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<emphasis id="B963EA930065FF829A5FFDAAFCB0C414" box="[633,778,547,571]" italics="true" pageId="54" pageNumber="55">M. froehlichi</emphasis>
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</taxonomicName>
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,
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<taxonomicNameLabel id="A25057E80065FF829B3EFDAAFCE9C415" box="[792,851,547,570]" pageId="54" pageNumber="55" rank="species">n. sp.</taxonomicNameLabel>
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However,
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<taxonomicName id="4C174D020065FF829BEBFDAAFBF6C414" authorityName="Blahnik & Holzenthal" authorityYear="2011" box="[973,1100,547,571]" class="Magnoliopsida" family="Apocynaceae" genus="Mortoniella" kingdom="Plantae" order="Gentianales" pageId="54" pageNumber="55" phylum="Tracheophyta" rank="species" species="bocaina">
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<emphasis id="B963EA930065FF829BEBFDAAFBF6C414" box="[973,1100,547,571]" italics="true" pageId="54" pageNumber="55">M. bocaina</emphasis>
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</taxonomicName>
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does have a pair of short, apically rounded, endophallic spines, not found in
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<taxonomicName id="4C174D020065FF829A8AFDCEFCFDC471" authorityName="Blahnik & Holzenthal" authorityYear="2011" box="[684,839,583,606]" class="Magnoliopsida" family="Apocynaceae" genus="Mortoniella" kingdom="Plantae" order="Gentianales" pageId="54" pageNumber="55" phylum="Tracheophyta" rank="species" species="tripuiensis">
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<emphasis id="B963EA930065FF829A8AFDCEFCFDC471" box="[684,839,583,606]" italics="true" pageId="54" pageNumber="55">M. tripuiensis</emphasis>
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</taxonomicName>
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, which can help to diagnose it.
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</paragraph>
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<paragraph id="8BA836810065FF8298E1FDE3FAF2C4E5" blockId="54.[151,1436,474,1434]" pageId="54" pageNumber="55">
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Adult. Length of forewing: male 2.9–3.3, female 3.0–
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<quantity id="4CEF9B640065FF829B0EFDE3FC3BC4AD" box="[808,897,618,642]" metricMagnitude="-3" metricUnit="m" metricValue="3.2" pageId="54" pageNumber="55" unit="mm" value="3.2">3.2 mm</quantity>
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. Forewing with forks I, II, and III present, hind wing with fork II only. Spur formula 0:3:4. Overall color dark brown. Legs brown, tibial spurs somewhat darker in color, but not strongly contrasting with legs. Wing bar at anastamosis marked with white, contrasting setae.
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</paragraph>
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<paragraph id="8BA836810065FF8298E1FD5FFCBAC2ED" blockId="54.[151,1436,474,1434]" pageId="54" pageNumber="55">
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Male genitalia. Ventral process of segment VI posteriorly projecting, elongate, narrow, length more than 2 times width at base (
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<figureCitation id="132C2A040065FF8299A2FD73FE56C53D" box="[388,492,762,786]" captionStart="FIGURE 27" captionStartId="55.[151,250,1976,1999]" captionTargetBox="[224,1351,208,1950]" captionTargetId="figure-44@55.[209,1378,193,1956]" captionTargetPageId="55" captionText="FIGURE 27. Mortoniella bocaina, new species, male genitalia: A—lateral; B—segment IX and tergum X, dorsal; C—phallic ensemble, ventral; D—dorsal phallic spine, dorsal; E—ventral process of segment VI, lateral." figureDoi="http://doi.org/10.5281/zenodo.5287037" httpUri="https://zenodo.org/record/5287037/files/figure.png" pageId="54" pageNumber="55">Fig. 27E</figureCitation>
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). Segment IX moderately rounded anterolaterally, length greatest in ventral 1/2 (or near middle), posterolateral margin nearly linear; segment deeply excised dorsomesally and ventromesally, forming lateral lobes, lobes separated dorsomesally by about 1/2 width of segment. Tergum X moderate in length, simple in structure, with deep, broad, U-shaped mesal excision; apical lobes, as viewed laterally, distinctly projecting, rounded to subacute apically. Inferior appendages almost completely fused to phallic ensemble, basally with short, setose, anteriorly-directed lobes, apically apparently fused to phallicata and endophallic membrane, forming membranous or very lightly sclerotized lobes. Mesal pockets of fused inferior appendages very bulbously enlarged, with elongate, spine-like, posteriorly-directed apicoventral projections. Paramere appendages very narrow, elongate, about as long as dorsal phallic spine. Phallotheca very short; ventral rod-like appendages elongate, with widely flared apices. Dorsal phallic spine, as viewed dorsally, broad in middle, gradually narrowed and rounded apically; as viewed laterally, relatively narrow, sharply upturned at about apical 1/3rd. Phallicata membranous, continuous with endophallic membrane. Endophallic membrane simple in structure, with membranous apical and dorsal lobes, laterally with short, sclerotized, ventrally curved lobes.
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</paragraph>
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</subSubSection>
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<subSubSection id="C30D650A0065FF8298E1FB46FDF8C3B5" pageId="54" pageNumber="55" type="materials_examined">
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<paragraph id="8BA836810065FF8298E1FB46FEFDC301" blockId="54.[151,1436,474,1434]" pageId="54" pageNumber="55">
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<materialsCitation id="3B7F3CDC0065FF8298E1FB46FEFDC301" collectingDate="2002-03-03" collectionCode="MZUSP" collectorName="Cachoeira dos Posses" country="Brazil" elevation="1250" latitude="-22.77389" location="Parque Nacional da Serra da Bocaina" longLatPrecision="21" longitude="-44.604164" pageId="54" pageNumber="55" specimenCode="UMSP000069696" specimenCount="1" stateProvince="Sao Paulo" typeStatus="holotype">
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||
<emphasis id="B963EA930065FF8298E1FB46FD18C2C8" bold="true" box="[199,674,1230,1255]" pageId="54" pageNumber="55">
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||
<typeStatus id="54AC88230065FF8298E1FB46FE8EC2C8" box="[199,308,1231,1255]" pageId="54" pageNumber="55" type="holotype">Holotype</typeStatus>
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male:
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<collectingCountry id="F30076110065FF8299B3FB47FDBFC2C8" box="[405,517,1230,1255]" name="Brazil" pageId="54" pageNumber="55">BRAZIL</collectingCountry>
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||
:
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||
<collectingRegion id="49D3F8630065FF829A3FFB47FD27C2C8" box="[537,669,1230,1255]" country="Brazil" name="Sao Paulo" pageId="54" pageNumber="55">São Paulo</collectingRegion>
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||
:
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||
</emphasis>
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||
<location id="8EC8605A0065FF829A97FB46FB3AC2C9" LSID="urn:lsid:plazi:treatment:03BE87970065FF8298B1FEE6FDF8C3B5:8EC8605A0065FF829A97FB46FB3AC2C9" box="[689,1152,1230,1254]" country="Brazil" latitude="-22.77389" longLatPrecision="21" longitude="-44.604164" name="Parque Nacional da Serra da Bocaina" pageId="54" pageNumber="55" stateProvince="Sao Paulo">Parque Nacional da Serra da Bocaina</location>
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||
,
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<collectorName id="26E253570065FF829CB5FB47FA2DC2C9" box="[1171,1431,1230,1254]" pageId="54" pageNumber="55">Cachoeira dos Posses</collectorName>
|
||
,
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||
<geoCoordinate id="EE2350460065FF8298B1FB7BFEA3C325" box="[151,281,1266,1290]" degrees="22" direction="south" minutes="46" orientation="latitude" pageId="54" pageNumber="55" precision="15" seconds="26" value="-22.77389">22°46'26"S</geoCoordinate>
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||
,
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||
<geoCoordinate id="EE2350460065FF829903FB7BFE06C325" box="[293,444,1266,1290]" degrees="044" direction="west" minutes="36" orientation="longitude" pageId="54" pageNumber="55" precision="15" seconds="15" value="-44.604164">044°36'15"W</geoCoordinate>
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||
,
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||
<quantity id="4CEF9B640065FF8299EFFB7BFD9BC325" box="[457,545,1266,1290]" metricMagnitude="3" metricUnit="m" metricValue="1.25" pageId="54" pageNumber="55" unit="m" value="1250.0">
|
||
<elevation id="003AD1B20065FF8299EFFB7BFD9BC325" box="[457,545,1266,1290]" metricMagnitude="3" metricUnit="m" metricValue="1.25" pageId="54" pageNumber="55" unit="m" value="1250.0">1250 m</elevation>
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||
</quantity>
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||
,
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<date id="FFA910410065FF829A08FB7BFD21C325" box="[558,667,1266,1290]" pageId="54" pageNumber="55" value="2002-03-03">
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||
<collectingDate id="EFEDE9A90065FF829A08FB7BFD21C325" box="[558,667,1266,1290]" pageId="54" pageNumber="55" value="2002-03-03">3.iii.2002</collectingDate>
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||
</date>
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||
, Holzenthal, Blahnik, Paprocki, & Prather (
|
||
<specimenCode id="DBB19EFA0065FF829CAFFB7BFAEDC325" box="[1161,1367,1266,1290]" collectionCode="UMSP" country="USA" httpUri="http://grbio.org/cool/3avb-bd3e" name="University of Minnesota Insect Collection" pageId="54" pageNumber="55" type="University or college">UMSP000069696</specimenCode>
|
||
) (pinned) (
|
||
<collectionCode id="ED06AE440065FF8298FEFA9FFE86C301" box="[216,316,1302,1326]" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="54" pageNumber="55">MZUSP</collectionCode>
|
||
).
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="8BA836810065FF8298E1FAB2FA2CC37D" blockId="54.[151,1436,474,1434]" box="[199,1430,1338,1363]" pageId="54" pageNumber="55">
|
||
<materialsCitation id="3B7F3CDC0065FF8298E1FAB2FA28C37D" box="[199,1426,1338,1363]" collectionCode="UMSP" country="Brazil" pageId="54" pageNumber="55" specimenCount="4" specimenCount-female="3" specimenCount-male="1" stateProvince="Sao Paulo" typeStatus="paratype">
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||
<emphasis id="B963EA930065FF8298E1FAB2FDF1C37C" bold="true" box="[199,587,1338,1363]" pageId="54" pageNumber="55">
|
||
<typeStatus id="54AC88230065FF8298E1FAB2FEF9C37C" box="[199,323,1339,1363]" pageId="54" pageNumber="55" type="paratype">Paratypes</typeStatus>
|
||
:
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||
<collectingCountry id="F30076110065FF829969FAB3FE05C37C" box="[335,447,1338,1363]" name="Brazil" pageId="54" pageNumber="55">BRAZIL</collectingCountry>
|
||
:
|
||
<collectingRegion id="49D3F8630065FF8299EDFAB3FDFFC37C" box="[459,581,1338,1363]" country="Brazil" name="Sao Paulo" pageId="54" pageNumber="55">São Paulo</collectingRegion>
|
||
:
|
||
</emphasis>
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||
same locality and date as
|
||
<typeStatus id="54AC88230065FF829B57FAB3FC69C37D" box="[881,979,1338,1362]" pageId="54" pageNumber="55" type="holotype">holotype</typeStatus>
|
||
—
|
||
<specimenCount id="9D11FD080065FF829BDBFAB3FBF1C37D" box="[1021,1099,1338,1362]" pageId="54" pageNumber="55" type="male">1 male</specimenCount>
|
||
,
|
||
<specimenCount id="9D11FD080065FF829C70FAB3FB78C37D" box="[1110,1218,1338,1362]" pageId="54" pageNumber="55" type="female">3 females</specimenCount>
|
||
(pinned) (
|
||
<collectionCode id="ED06AE440065FF829D11FAB3FA30C37D" box="[1335,1418,1338,1362]" country="USA" httpUri="http://grbio.org/cool/3avb-bd3e" name="University of Minnesota Insect Collection" pageId="54" pageNumber="55" type="University or college">UMSP</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BA836810065FF8298E1FAD7FDF8C3B5" blockId="54.[151,1436,474,1434]" pageId="54" pageNumber="55">
|
||
Etymology. This species is named
|
||
<taxonomicName id="4C174D020065FF829A76FAD6FD6BC359" authorityName="Blahnik & Holzenthal" authorityYear="2011" box="[592,721,1374,1398]" class="Magnoliopsida" family="Apocynaceae" genus="Mortoniella" kingdom="Plantae" order="Gentianales" pageId="54" pageNumber="55" phylum="Tracheophyta" rank="species" species="bocaina">
|
||
<emphasis id="B963EA930065FF829A76FAD6FD6BC359" box="[592,721,1374,1398]" italics="true" pageId="54" pageNumber="55">M. bocaina</emphasis>
|
||
</taxonomicName>
|
||
, as a noun in apposition, for the name of the beautiful national park where this species was collected.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |