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<document id="CFA92A9FBC71E8760458D6091743D181" ID-CLB-Dataset="55730" ID-DOI="10.5852/ejt.2022.817.1767" ID-GBIF-Dataset="529dc516-47b9-424e-93ac-3787e407e100" ID-ISSN="2118-9773" ID-Zenodo-Dep="6517928" ID-ZooBank="FFB05CE7-3712-4FD0-91B4-FD84863CF9D7" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1651673830896" checkinUser="felipe" docAuthor="Yasuda, Kohei &amp; Yoshitomi, Hiroyuki" docDate="2022" docId="03BE8793BE4FFFC2FE17BEE483730193" docLanguage="en" docName="ejt-817_111-142.pdf" docOrigin="European Journal of Taxonomy 817 (1)" docSource="http://dx.doi.org/10.5852/ejt.2022.817.1767" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Georissus (Neogeorissus) japonicus Sato 1972" docType="treatment" docVersion="9" lastPageNumber="126" masterDocId="FF87FFEBBE42FFCDFFE8BF5B80470455" masterDocTitle="Revision of the genus Georissus (Coleoptera, Hydrophiloidea, Georissidae) of Japan" masterLastPageNumber="142" masterPageNumber="111" pageNumber="124" updateTime="1699329566334" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="75B26B66D6A68BC79E00A252625034C4">Revision of the genus Georissus (Coleoptera, Hydrophiloidea, Georissidae) of Japan</mods:title>
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<mods:namePart id="D7B668BA1A028175A00664E82BAE155E">Yasuda, Kohei</mods:namePart>
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<mods:affiliation id="13BB2009B9746A1B9D8980D733A89AE5">Entomological Laboratory, Faculty of Agriculture, Ehime University, Tarumi 3 - 5 - 7, Matsuyama, 790 - 8566 Japan.</mods:affiliation>
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<mods:namePart id="D4305577C3F165BA47FA2AF3CE5D25D1">Yoshitomi, Hiroyuki</mods:namePart>
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<mods:affiliation id="C6F6AE7AD9DBC48FE8CE0554E354C8CC">Entomological Laboratory, Faculty of Agriculture, Ehime University, Tarumi 3 - 5 - 7, Matsuyama, 790 - 8566 Japan.</mods:affiliation>
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<mods:title id="B9ADC4201134AC0F3D76E6F6D32C1E5C">European Journal of Taxonomy</mods:title>
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<mods:date id="B2ACCAF18F34F99C1CB3CC4ADFA69E95">2022</mods:date>
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<treatment id="03BE8793BE4FFFC2FE17BEE483730193" ID-DOI="http://doi.org/10.5281/zenodo.6518783" ID-GBIF-Taxon="195191634" ID-Zenodo-Dep="6518783" LSID="urn:lsid:plazi:treatment:03BE8793BE4FFFC2FE17BEE483730193" httpUri="http://treatment.plazi.org/id/03BE8793BE4FFFC2FE17BEE483730193" lastPageId="15" lastPageNumber="126" pageId="13" pageNumber="124">
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<taxonomicName id="4C174D06BE4FFFC0FE17BEE48473058C" ID-CoL="3FS2D" authority="Sato, 1972" authorityName="Sato" authorityYear="1972" box="[511,1076,446,473]" class="Insecta" family="Georissidae" genus="Georissus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="124" phylum="Arthropoda" rank="species" species="japonicus" subGenus="Neogeorissus">
<emphasis id="B963EA97BE4FFFC0FE17BEE483E8058C" bold="true" box="[511,943,446,473]" pageId="13" pageNumber="124">
<emphasis id="B963EA97BE4FFFC0FE17BEE48231058C" bold="true" box="[511,630,447,473]" italics="true" pageId="13" pageNumber="124">Georissus</emphasis>
(
<emphasis id="B963EA97BE4FFFC0FD6FBEE4836F058C" bold="true" box="[647,808,447,473]" italics="true" pageId="13" pageNumber="124">Neogeorissus</emphasis>
)
<emphasis id="B963EA97BE4FFFC0FCD2BEE483E8058C" bold="true" box="[826,943,447,473]" italics="true" pageId="13" pageNumber="124">japonicus</emphasis>
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<bibRefCitation id="EF864B74BE4FFFC0FC5FBEE48473058C" author="Sato M." box="[951,1076,447,473]" pageId="13" pageNumber="124" pagination="207 - 213" refId="ref14083" refString="Sato M. 1972. The Georissid beetles of Japan. Journal of Nagoya Women's College 18: 207 - 213." type="journal article" year="1972">Satô, 1972</bibRefCitation>
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<paragraph id="8BA83685BE4FFFC0FDCABEB8845605A9" blockId="13.[321,1266,446,544]" box="[546,1041,481,509]" pageId="13" pageNumber="124">[Japanese name: Yamato-marudoromushi]</paragraph>
</subSubSection>
<subSubSection id="C30D650EBE4FFFC0FEA9BD5E84B5064A" box="[321,1266,517,544]" pageId="13" pageNumber="124" type="description">
<paragraph id="8BA83685BE4FFFC0FEA9BD5E84B5064A" blockId="13.[321,1266,446,544]" box="[321,1266,517,544]" pageId="13" pageNumber="124">
<figureCitation id="132C2A00BE4FFFC0FEA9BD5E81E5064A" box="[321,418,517,543]" captionStart="Fig" captionStartId="7.[189,232,1886,1912]" captionTargetBox="[272,1331,265,1833]" captionTargetId="figure-11@7.[223,1365,265,1845]" captionTargetPageId="7" captionText="Fig. 2. Dorsal view of Georissus spp. A. G. (G.) canalifer Sharp, 1888. B. G. (G.) babai Satô, 1970. C. G. (Ni.) granulosus Satô, 1972. D. G. (Ne.) japonicus Satô, 1972. E. G. (Ne.) kurosawai Nakane, 1963. F. G. (Ne.) takahashii sp. nov., holotype, ♂ (EUMJ). G. G. (Ne.) satoi sp. nov., holotype (EUMJ). Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6517952" httpUri="https://zenodo.org/record/6517952/files/figure.png" pageId="13" pageNumber="124">Figs 2D</figureCitation>
,
<figureCitation id="132C2A00BE4FFFC0FE46BD5E826B0675" box="[430,556,517,544]" captionStart="Fig" captionStartId="9.[189,232,1886,1912]" captionTargetBox="[196,1392,306,1839]" captionTargetId="figure-11@9.[189,1398,268,1842]" captionTargetPageId="9" captionText="Fig. 3. Holotypes of Georissus spp. A–F. Dorsal habitus. G–L. Labels. A, K. G. (G.) babai Satô, 1970, ♂ (EUMJ). B, L. G. (Ni.) granulosus Satô, 1972, ♀ (EUMJ). C, H. G. (Ne.) kurosawai Nakane, 1963, ♂ (SEHU). D, J. G. (Ne.) japonicus Satô, 1972, ♂ (EUMJ). E, G. G. (Ne.) sakaii Satô, 1972, ♂ (EUMJ). F, I. G. (Ne.) katsuoi Nakane, 1995 (SEHU). Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6517956" httpUri="https://zenodo.org/record/6517956/files/figure.png" pageId="13" pageNumber="124">3D–F, J–L</figureCitation>
,
<figureCitation id="132C2A00BE4FFFC0FDD0BD5E8291064A" box="[568,726,517,544]" captionStart="Fig" captionStartId="14.[189,232,1876,1902]" captionTargetBox="[196,1386,277,1817]" captionTargetId="figure-18@14.[189,1398,264,1820]" captionTargetPageId="14" captionText="Fig. 4. Mouth parts and antennae of Georissus spp. A–C. Left maxillae. D–F. Labrum. G–I. Labial palpi. J–L. Left mandibles. M–O. Left antennae. A, D, G, J, M. G. (G.) canalifer Sharp, 1888. B, E, H, K, N. G. (Ni.) granulosus Satô, 1972. C, F, I, L, O. G. (Ne.) japonicus Satô, 1972. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6517962" httpUri="https://zenodo.org/record/6517962/files/figure.png" pageId="13" pageNumber="124">4C, F, I, L, O</figureCitation>
,
<figureCitation id="132C2A00BE4FFFC0FD0ABD5E834F0675" box="[738,776,517,544]" captionStart="Fig" captionStartId="16.[189,232,1822,1848]" captionTargetBox="[192,1395,266,1778]" captionTargetId="figure-17@16.[189,1398,266,1783]" captionTargetPageId="16" captionText="Fig. 5. Dorsal habitus of Georissus spp. A. G. (G.) canalifer Sharp, 1888. B. G. (G.) babai Satô, 1970. C. G. (Ni.) granulosus Satô, 1972. D. G. (Ne.) japonicus Satô, 1972. E–F. G. (Ne.) kurosawai Nakane, 1963 from Hokkaido (E) and Nagano (F). G–H. G. (Ne.) takahashii sp. nov. from Kochi (G) and Okinawa (H). I. G. (Ne.) satoi sp. nov." figureDoi="http://doi.org/10.5281/zenodo.6517964" httpUri="https://zenodo.org/record/6517964/files/figure.png" pageId="13" pageNumber="124">5D</figureCitation>
,
<figureCitation id="132C2A00BE4FFFC0FCFCBD5E837C064A" box="[788,827,517,543]" captionStart="Fig" captionStartId="18.[189,232,1882,1908]" captionTargetBox="[191,1394,563,1839]" captionTargetId="figure-90@18.[189,1398,562,1842]" captionTargetPageId="18" captionText="Fig. 6. Head of Georissus spp. A. G. (G.) canalifer Sharp, 1888. B. G. (G.) babai Satô, 1970. C. G. (Ni.) granulosus Satô, 1972. D. G. (Ne.) japonicus Satô, 1972. E–F. G. (Ne.) kurosawai Nakane, 1963 from Hokkaido (E) and Nagano (F). G–H. G. (Ne.) takahashii sp. nov. from Kochi (G) and Okinawa (H). I. G. (Ne.) satoi sp. nov." figureDoi="http://doi.org/10.5281/zenodo.6517968" httpUri="https://zenodo.org/record/6517968/files/figure.png" pageId="13" pageNumber="124">6D</figureCitation>
,
<figureCitation id="132C2A00BE4FFFC0FCAFBD5D832A0675" box="[839,877,518,544]" captionStart="Fig" captionStartId="19.[189,232,1879,1905]" captionTargetBox="[197,1389,792,1835]" captionTargetId="figure-118@19.[189,1398,783,1839]" captionTargetPageId="19" captionText="Fig. 7. Pronotum of Georissus spp. A. G. (G.) canalifer Sharp, 1888. B. G. (G.) babai Satô, 1970. C. G. (Ni.) granulosus Satô, 1972. D. G. (Ne.) japonicus Satô, 1972. E–F. G. (Ne.) kurosawai Nakane, 1963 from Hokkaido (E) and Nagano (F). G–H. G. (Ne.) takahashii sp. nov. from Kochi (G) and Okinawa (H). I. G. (Ne.) satoi sp. nov." figureDoi="http://doi.org/10.5281/zenodo.6517976" httpUri="https://zenodo.org/record/6517976/files/figure.png" pageId="13" pageNumber="124">7D</figureCitation>
,
<figureCitation id="132C2A00BE4FFFC0FC91BD5E83E7064A" box="[889,928,517,543]" captionStart="Fig" captionStartId="21.[191,234,1890,1916]" captionTargetBox="[193,1394,569,1846]" captionTargetId="figure-107@21.[189,1398,569,1852]" captionTargetPageId="21" captionText="Fig. 8. Ventrites of Georissus spp. A. G. (G.) canalifer Sharp, 1888. B. G. (G.) babai Satô, 1970. C. G. (Ni.) granulosus Satô, 1972. D. G. (Ne.) japonicus Satô, 1972. E–F. G. (Ne.) kurosawai Nakane, 1963 from Hokkaido (E) and Nagano (F). G–H. G. (Ne.) takahashii sp. nov. from Kochi (G) and Okinawa (F). I. G. (Ne.) satoi sp. nov. Abbreviation: vt = ventral teeth. Scale bars = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.6517982" httpUri="https://zenodo.org/record/6517982/files/figure.png" pageId="13" pageNumber="124">8D</figureCitation>
,
<figureCitation id="132C2A00BE4FFFC0FC44BD5E8395064A" box="[940,978,517,543]" captionStart="Fig" captionStartId="23.[189,232,1763,1789]" captionTargetBox="[239,1353,284,1696]" captionTargetId="figure-12@23.[189,1398,265,1704]" captionTargetPageId="23" captionText="Fig. 9. Male aedeagus of Georissus spp. A. G. (G.) canalifer Sharp, 1888. B. G. (G.) babai Satô, 1970. C. G. (Ni.) granulosus Satô, 1972. D. G. (Ne.) japonicus Satô, 1972. E. G. (Ne.) kurosawai Nakane, 1963. F–G. G. (Ne.) takahashii sp. nov. from Kochi (F) and Okinawa (G). H. G. (Ne.) satoi sp. nov. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6517986" httpUri="https://zenodo.org/record/6517986/files/figure.png" pageId="13" pageNumber="124">9D</figureCitation>
,
<figureCitation id="132C2A00BE4FFFC0FC37BD5E8455064A" box="[991,1042,517,543]" captionStart="Fig" captionStartId="24.[189,232,1780,1806]" captionTargetBox="[233,1337,288,1718]" captionTargetId="figure-18@24.[189,1398,264,1722]" captionTargetPageId="24" captionText="Fig. 10. Female ovipositor of Georissus spp. A. G. (G.) canalifer Sharp, 1888. B. G. (G.) babai Satô, 1970. C. G. (Ne.) japonicus Satô, 1972. D. G. (Ne.) kurosawai Nakane, 1963. E. G. (Ne.) takahashii sp. nov. F. G. (Ne.) satoi sp. nov. Abbreviations: gsty = gonostylus; ltg8 = latero-tergite 8. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6517988" httpUri="https://zenodo.org/record/6517988/files/figure.png" pageId="13" pageNumber="124">10C</figureCitation>
,
<figureCitation id="132C2A00BE4FFFC0FBF6BD5E84C8064A" box="[1054,1167,517,543]" captionStart="Fig" captionStartId="25.[189,232,1920,1946]" captionTargetBox="[213,1341,1093,1867]" captionTargetId="figure-278@25.[189,1398,1073,1882]" captionTargetPageId="25" captionText="Fig. 11. Female genital organs of subgenus Neogeorissus spp. A–D. Bursal sclerite. E–H. Spermatheca. A, E. G. (Ne.) japonicus Satô, 1972. B, F. G. (Ne.) kurosawai Nakane, 1963. C, G. G. (Ne.) takahashii sp. nov. D, H. G. (Ne.) satoi sp. nov. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6517990" httpUri="https://zenodo.org/record/6517990/files/figure.png" pageId="13" pageNumber="124">11A, 11E</figureCitation>
;
<tableCitation id="C695033EBE4FFFC0FB73BD5E84B5064A" box="[1179,1266,517,543]" captionStart="Table 2" captionStartId="20.[197,273,265,293]" captionTargetBox="[232,1343,364,817]" captionTargetPageId="20" captionText="Table 2. Measurements of Georissus (Neogeorissus) spp." httpUri="http://table.plazi.org/id/DF68660DBE56FFD9FF2DBE5283E20573" pageId="13" pageNumber="124" tableUuid="DF68660DBE56FFD9FF2DBE5283E20573">Table 2</tableCitation>
</paragraph>
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<paragraph id="8BA83685BE4FFFC0FF55BD1C836A0634" blockId="13.[189,926,583,680]" box="[189,813,583,609]" pageId="13" pageNumber="124">
<treatmentCitationGroup id="AB0711ABBE4FFFC0FF55BD1C836A0634" box="[189,813,583,609]" pageId="13" pageNumber="124">
<treatmentCitation id="0AB61094BE4FFFC0FF55BD1C836E0634" author="Sato M." box="[189,809,583,609]" page="210" pageId="13" pageNumber="124" year="1972">
<taxonomicName id="4C174D06BE4FFFC0FF55BD1C836E0634" ID-CoL="3FS2D" authority="Sato, 1972: 210" authorityName="Sato" authorityPageNumber="210" authorityYear="1972" box="[189,809,583,609]" class="Insecta" family="Georissidae" genus="Georissus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="124" phylum="Arthropoda" rank="species" species="japonicus" subGenus="Neogeorissus">
<emphasis id="B963EA97BE4FFFC0FF55BD1C81750634" box="[189,306,583,609]" italics="true" pageId="13" pageNumber="124">Georissus</emphasis>
(
<emphasis id="B963EA97BE4FFFC0FEABBD1C81A60635" box="[323,481,583,608]" italics="true" pageId="13" pageNumber="124">Neogeorissus</emphasis>
)
<emphasis id="B963EA97BE4FFFC0FE1BBD1C82220635" box="[499,613,583,608]" italics="true" pageId="13" pageNumber="124">japonicus</emphasis>
<bibRefCitation id="EF864B74BE4FFFC0FD84BD1C836E0634" author="Sato M." box="[620,809,583,609]" pageId="13" pageNumber="124" pagination="207 - 213" refId="ref14083" refString="Sato M. 1972. The Georissid beetles of Japan. Journal of Nagoya Women's College 18: 207 - 213." type="journal article" year="1972">Satô, 1972: 210</bibRefCitation>
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.
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</paragraph>
<paragraph id="8BA83685BE4FFFC0FF55BD31833E06D1" blockId="13.[189,926,583,680]" box="[189,889,618,644]" pageId="13" pageNumber="124">
<treatmentCitationGroup id="AB0711ABBE4FFFC0FF55BD31833E06D1" box="[189,889,618,644]" pageId="13" pageNumber="124">
<treatmentCitation id="0AB61094BE4FFFC0FF55BD3182BA06D1" author="Sato M." box="[189,765,618,644]" page="210" pageId="13" pageNumber="124" year="1972">
<taxonomicName id="4C174D06BE4FFFC0FF55BD3182BA06D1" ID-CoL="9G96B" authority="Sato, 1972: 210" authorityName="Sato" authorityPageNumber="210" authorityYear="1972" box="[189,765,618,644]" class="Insecta" family="Georissidae" genus="Georissus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="124" phylum="Arthropoda" rank="species" species="sakaii" subGenus="Neogeorissus">
<emphasis id="B963EA97BE4FFFC0FF55BD31817506D1" box="[189,306,618,644]" italics="true" pageId="13" pageNumber="124">Georissus</emphasis>
(
<emphasis id="B963EA97BE4FFFC0FEABBD31827E06D1" box="[323,569,618,644]" italics="true" pageId="13" pageNumber="124">Neogeorissus) sakaii</emphasis>
<bibRefCitation id="EF864B74BE4FFFC0FDA8BD3182BA06D1" author="Sato M." box="[576,765,618,644]" pageId="13" pageNumber="124" pagination="207 - 213" refId="ref14083" refString="Sato M. 1972. The Georissid beetles of Japan. Journal of Nagoya Women's College 18: 207 - 213." type="journal article" year="1972">Satô, 1972: 210</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
<emphasis id="B963EA97BE4FFFC0FCE0BD31833E06D1" bold="true" box="[776,889,618,644]" pageId="13" pageNumber="124">Syn. nov.</emphasis>
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BA83685BE4FFFC0FF55BDD583D906F2" blockId="13.[189,926,583,680]" box="[189,926,653,680]" pageId="13" pageNumber="124">
<treatmentCitationGroup id="AB0711ABBE4FFFC0FF55BDD583D906F2" box="[189,926,653,680]" pageId="13" pageNumber="124">
<treatmentCitation id="0AB61094BE4FFFC0FF55BDD5836506F2" author="Nakane T." box="[189,802,653,680]" page="42" pageId="13" pageNumber="124" year="1995">
<taxonomicName id="4C174D06BE4FFFC0FF55BDD5836506F2" ID-CoL="3FS45" authority="Nakane, 1995: 42" authorityName="Nakane" authorityPageNumber="42" authorityYear="1995" box="[189,802,653,680]" class="Insecta" family="Georissidae" genus="Georissus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="124" phylum="Arthropoda" rank="species" species="katsuoi" subGenus="Neogeorissus">
<emphasis id="B963EA97BE4FFFC0FF55BDD5817506FD" box="[189,306,654,680]" italics="true" pageId="13" pageNumber="124">Georissus</emphasis>
(
<emphasis id="B963EA97BE4FFFC0FEABBDD581A606F2" box="[323,481,654,679]" italics="true" pageId="13" pageNumber="124">Neogeorissus</emphasis>
)
<emphasis id="B963EA97BE4FFFC0FE1BBDD6820F06F2" box="[499,584,653,679]" italics="true" pageId="13" pageNumber="124">katsuoi</emphasis>
<bibRefCitation id="EF864B74BE4FFFC0FDA7BDD6836506F2" author="Nakane T." box="[591,802,653,680]" pageId="13" pageNumber="124" pagination="41 - 46" refId="ref13940" refString="Nakane T. 1995. Notes on some little-known beetles (Coleoptera) in Japan. 13. Kita-Kyushu no Konchu 42: 41 - 46." type="journal article" year="1995">Nakane, 1995: 42</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
<emphasis id="B963EA97BE4FFFC0FCC5BDD683D906F2" bold="true" box="[813,926,653,679]" pageId="13" pageNumber="124">Syn. nov.</emphasis>
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BA83685BE4FFFC0FF55BD9484D906BC" blockId="13.[189,1182,718,815]" box="[189,1182,718,745]" pageId="13" pageNumber="124">
<treatmentCitationGroup id="AB0711ABBE4FFFC0FF55BD9484D906BC" box="[189,1182,718,745]" pageId="13" pageNumber="124">
<taxonomicName id="4C174D06BE4FFFC0FF55BD94822206BD" ID-CoL="3FS2D" authorityName="Sato" authorityYear="1972" box="[189,613,719,745]" class="Insecta" family="Georissidae" genus="Georissus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="124" phylum="Arthropoda" rank="species" species="japonicus" subGenus="Neogeorissus">
<emphasis id="B963EA97BE4FFFC0FF55BD94817506BC" box="[189,306,719,745]" italics="true" pageId="13" pageNumber="124">Georissus</emphasis>
(
<emphasis id="B963EA97BE4FFFC0FEABBD9481A606BD" box="[323,481,719,744]" italics="true" pageId="13" pageNumber="124">Neogeorissus</emphasis>
)
<emphasis id="B963EA97BE4FFFC0FE1BBD94822206BD" box="[499,613,719,744]" italics="true" pageId="13" pageNumber="124">japonicus</emphasis>
</taxonomicName>
–
<treatmentCitation id="0AB61094BE4FFFC0FD6ABD94835E06BC" author="Sato M." box="[642,793,719,745]" page="3" pageId="13" pageNumber="124" year="1981">
<bibRefCitation id="EF864B74BE4FFFC0FD6ABD94835E06BC" author="Sato M." box="[642,793,719,745]" pageId="13" pageNumber="124" pagination="1 - 4" refId="ref14105" refString="Sato M. 1981. An outline of Japanese Georissidae. Coleopterist's News 53: 1 - 4. [In Japanese]" type="journal article" year="1981">Satô 1981: 3</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0AB61094BE4FFFC0FCCEBD9583E706BD" author="Sato M." box="[806,928,718,745]" page="206" pageId="13" pageNumber="124" year="1985">
<bibRefCitation id="EF864B74BE4FFFC0FCCEBD9583E706BD" author="Sato M." box="[806,928,718,745]" pageId="13" pageNumber="124" refId="ref14128" refString="Sato M. 1985. Georissidae. In: Ueno S., Kurosawa Y. &amp; Sato M. (eds) The Coleoptera of Japan in Color Vol. II. Hoikusha, Osaka. [In Japanese]" type="book" year="1985">1985: 206</bibRefCitation>
</treatmentCitation>
. —
<treatmentCitation id="0AB61094BE4FFFC0FC27BD9484DD06BC" author="Hansen M." box="[975,1178,719,745]" page="47" pageId="13" pageNumber="124" year="1999">
<bibRefCitation id="EF864B74BE4FFFC0FC27BD9484DD06BC" author="Hansen M." box="[975,1178,719,745]" pageId="13" pageNumber="124" refId="ref13193" refString="Hansen M. 1999. World Catalogue of Insects 2: Hydrophiloidea (s. str.) (Coleoptera). Apollo Books, Amsterdam." type="book" year="1999">Hansen 1999: 47</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BA83685BE4FFFC0FF55BDA984350759" blockId="13.[189,1182,718,815]" box="[189,1138,754,781]" pageId="13" pageNumber="124">
<treatmentCitationGroup id="AB0711ABBE4FFFC0FF55BDA984350759" box="[189,1138,754,781]" pageId="13" pageNumber="124">
<taxonomicName id="4C174D06BE4FFFC0FF55BDA9827E0759" ID-CoL="9G96B" authorityName="Sato" authorityYear="1972" box="[189,569,754,780]" class="Insecta" family="Georissidae" genus="Georissus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="124" phylum="Arthropoda" rank="species" species="sakaii" subGenus="Neogeorissus">
<emphasis id="B963EA97BE4FFFC0FF55BDA981750759" box="[189,306,754,780]" italics="true" pageId="13" pageNumber="124">Georissus</emphasis>
(
<emphasis id="B963EA97BE4FFFC0FEABBDA9827E0759" box="[323,569,754,780]" italics="true" pageId="13" pageNumber="124">Neogeorissus) sakaii</emphasis>
</taxonomicName>
–
<treatmentCitation id="0AB61094BE4FFFC0FDBEBDA982AA0759" author="Sato M." box="[598,749,754,780]" page="3" pageId="13" pageNumber="124" year="1981">
<bibRefCitation id="EF864B74BE4FFFC0FDBEBDA982AA0759" author="Sato M." box="[598,749,754,780]" pageId="13" pageNumber="124" pagination="1 - 4" refId="ref14105" refString="Sato M. 1981. An outline of Japanese Georissidae. Coleopterist's News 53: 1 - 4. [In Japanese]" type="journal article" year="1981">Satô 1981: 3</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0AB61094BE4FFFC0FD12BDA983330759" author="Sato M." box="[762,884,754,781]" page="206" pageId="13" pageNumber="124" year="1985">
<bibRefCitation id="EF864B74BE4FFFC0FD12BDA983330759" author="Sato M." box="[762,884,754,781]" pageId="13" pageNumber="124" refId="ref14128" refString="Sato M. 1985. Georissidae. In: Ueno S., Kurosawa Y. &amp; Sato M. (eds) The Coleoptera of Japan in Color Vol. II. Hoikusha, Osaka. [In Japanese]" type="book" year="1985">1985: 206</bibRefCitation>
</treatmentCitation>
. —
<treatmentCitation id="0AB61094BE4FFFC0FC4BBDA984290759" author="Hansen M." box="[931,1134,754,780]" page="49" pageId="13" pageNumber="124" year="1999">
<bibRefCitation id="EF864B74BE4FFFC0FC4BBDA984290759" author="Hansen M." box="[931,1134,754,780]" pageId="13" pageNumber="124" refId="ref13193" refString="Hansen M. 1999. World Catalogue of Insects 2: Hydrophiloidea (s. str.) (Coleoptera). Apollo Books, Amsterdam." type="book" year="1999">Hansen 1999: 49</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BA83685BE4FFFC0FF55BC4E8374077A" blockId="13.[189,1182,718,815]" box="[189,819,789,815]" pageId="13" pageNumber="124">
<treatmentCitationGroup id="AB0711ABBE4FFFC0FF55BC4E8374077A" box="[189,819,789,815]" pageId="13" pageNumber="124">
<taxonomicName id="4C174D06BE4FFFC0FF55BC4E820F077A" ID-CoL="3FS45" authorityName="Nakane" authorityYear="1995" box="[189,584,789,815]" class="Insecta" family="Georissidae" genus="Georissus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="124" phylum="Arthropoda" rank="species" species="katsuoi" subGenus="Neogeorissus">
<emphasis id="B963EA97BE4FFFC0FF55BC4E8175077A" box="[189,306,789,815]" italics="true" pageId="13" pageNumber="124">Georissus</emphasis>
(
<emphasis id="B963EA97BE4FFFC0FEABBC4D81A6077A" box="[323,481,790,815]" italics="true" pageId="13" pageNumber="124">Neogeorissus</emphasis>
)
<emphasis id="B963EA97BE4FFFC0FE1BBC4E820F077A" box="[499,584,789,815]" italics="true" pageId="13" pageNumber="124">katsuoi</emphasis>
</taxonomicName>
–
<treatmentCitation id="0AB61094BE4FFFC0FD8DBC4E8368077A" author="Hansen M." box="[613,815,789,815]" page="47" pageId="13" pageNumber="124" year="1999">
<bibRefCitation id="EF864B74BE4FFFC0FD8DBC4E8368077A" author="Hansen M." box="[613,815,789,815]" pageId="13" pageNumber="124" refId="ref13193" refString="Hansen M. 1999. World Catalogue of Insects 2: Hydrophiloidea (s. str.) (Coleoptera). Apollo Books, Amsterdam." type="book" year="1999">Hansen 1999: 47</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C30D650EBE4FFFC0FF55BC0D849307B3" pageId="13" pageNumber="124" type="diagnosis">
<paragraph id="8BA83685BE4FFFC0FF55BC0D81710725" blockId="13.[189,310,854,880]" box="[189,310,854,880]" pageId="13" pageNumber="124">
<heading id="D0E081E9BE4FFFC0FF55BC0D81710725" bold="true" box="[189,310,854,880]" fontSize="11" level="3" pageId="13" pageNumber="124" reason="3">
<emphasis id="B963EA97BE4FFFC0FF55BC0D81710725" bold="true" box="[189,310,854,880]" pageId="13" pageNumber="124">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BA83685BE4FFFC0FF55BCDE849307B3" blockId="13.[188,1398,901,998]" pageId="13" pageNumber="124">
This species is distinguished from other Japanese
<taxonomicName id="4C174D06BE4FFFC0FD16BCDD83DB07CA" authorityName="Sato" authorityYear="1972" box="[766,924,902,927]" class="Insecta" family="Georissidae" genus="Georissus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="136" phylum="Arthropoda" rank="subGenus" subGenus="Neogeorissus">
<emphasis id="B963EA97BE4FFFC0FD16BCDD83DB07CA" box="[766,924,902,927]" italics="true" pageId="13" pageNumber="124">Neogeorissus</emphasis>
</taxonomicName>
species by the combination of following characteristics; distinct granulation of pronotum and elytra; median lobe shorter than paramere (in contrast to median lobe longer than paramere in other Japanese species of
<taxonomicName id="4C174D06BE4FFFC0FBCCBC97848507B0" authorityName="Sato" authorityYear="1972" box="[1060,1218,972,997]" class="Insecta" family="Georissidae" genus="Georissus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="136" phylum="Arthropoda" rank="subGenus" subGenus="Neogeorissus">
<emphasis id="B963EA97BE4FFFC0FBCCBC97848507B0" box="[1060,1218,972,997]" italics="true" pageId="13" pageNumber="124">Neogeorissus</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="C30D650EBE4FFFC0FF55BB57851E0202" pageId="13" pageNumber="124" type="materials_examined">
<paragraph id="8BA83685BE4FFFC0FF55BB5781ED0073" blockId="13.[189,426,1036,1062]" box="[189,426,1036,1062]" pageId="13" pageNumber="124">
<heading id="D0E081E9BE4FFFC0FF55BB5781ED0073" bold="true" box="[189,426,1036,1062]" fontSize="11" level="3" pageId="13" pageNumber="124" reason="3">
<emphasis id="B963EA97BE4FFFC0FF55BB5781ED0073" bold="true" box="[189,426,1036,1062]" pageId="13" pageNumber="124">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="8BA83685BE4FFFC0FF02BB6082AD0000" blockId="13.[189,1326,1083,1145]" box="[234,746,1083,1110]" pageId="13" pageNumber="124">
<emphasis id="B963EA97BE4FFFC0FF02BB6082AD0000" bold="true" box="[234,746,1083,1110]" pageId="13" pageNumber="124">
<typeStatus id="54AC8827BE4FFFC0FF02BB60811B0000" box="[234,348,1083,1109]" pageId="13" pageNumber="124" type="holotype">Holotype</typeStatus>
of
<taxonomicName id="4C174D06BE4FFFC0FE6CBB6082AD0000" authority="Sato, 1972" authorityName="Sato" authorityYear="1972" box="[388,746,1083,1110]" class="Insecta" family="Georissidae" genus="Georissus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="124" phylum="Arthropoda" rank="species" species="japonicus" subGenus="Ne.">
<emphasis id="B963EA97BE4FFFC0FE6CBB6081E70000" bold="true" box="[388,416,1083,1109]" italics="true" pageId="13" pageNumber="124">G.</emphasis>
(
<emphasis id="B963EA97BE4FFFC0FE59BB67819C0003" bold="true" box="[433,475,1084,1110]" italics="true" pageId="13" pageNumber="124">Ne.</emphasis>
)
<emphasis id="B963EA97BE4FFFC0FE04BB6082260000" bold="true" box="[492,609,1083,1109]" italics="true" pageId="13" pageNumber="124">japonicus</emphasis>
<bibRefCitation id="EF864B74BE4FFFC0FD80BB6082AD0000" author="Sato M." box="[616,746,1083,1109]" pageId="13" pageNumber="124" pagination="207 - 213" refId="ref14083" refString="Sato M. 1972. The Georissid beetles of Japan. Journal of Nagoya Women's College 18: 207 - 213." type="journal article" year="1972">Satô, 1972</bibRefCitation>
</taxonomicName>
</emphasis>
</paragraph>
<paragraph id="8BA83685BE4FFFC0FF55BB048569002C" blockId="13.[189,1326,1083,1145]" box="[189,1326,1118,1145]" pageId="13" pageNumber="124">
<materialsCitation id="3B7F3CD8BE4FFFC0FF55BB04856D002C" ID-GBIF-Occurrence="3762863306" box="[189,1322,1118,1145]" collectingDate="1961-08-08" collectionCode="EUMJ" collectorName="M. Sato" country="Japan" location="Niigata Pref." pageId="13" pageNumber="124" specimenCount="1" specimenCount-male="1" stateProvince="Niigata">
<collectingCountry id="F3007615BE4FFFC0FF55BB048151002C" box="[189,278,1119,1145]" name="Japan" pageId="13" pageNumber="124">JAPAN</collectingCountry>
–
<emphasis id="B963EA97BE4FFFC0FEDBBB048202002D" bold="true" box="[307,581,1118,1145]" pageId="13" pageNumber="124">
Honshu:
<collectingRegion id="49D3F867BE4FFFC0FE4EBB058202002D" box="[422,581,1118,1144]" country="Japan" name="Niigata" pageId="13" pageNumber="124">Niigata Pref.</collectingRegion>
</emphasis>
•
<specimenCount id="9D11FD0CBE4FFFC0FDB6BB048230002C" box="[606,631,1119,1145]" pageId="13" pageNumber="124" type="male">♂</specimenCount>
; Murakami;
<date id="FFA91045BE4FFFC0FCE5BB0483E6002D" box="[781,929,1118,1145]" pageId="13" pageNumber="124" value="1961-08-08">
<collectingDate id="EFEDE9ADBE4FFFC0FCE5BB0483E6002D" box="[781,929,1118,1145]" pageId="13" pageNumber="124" value="1961-08-08">8 Aug. 1961</collectingDate>
</date>
;
<collectorName id="26E25353BE4FFFC0FC45BB04844D002C" box="[941,1034,1119,1145]" pageId="13" pageNumber="124">M. Satô</collectorName>
leg.;
<collectionCode id="ED06AE40BE4FFFC0FBA4BB0484DE002C" box="[1100,1177,1119,1145]" collectionName="Japan, Matsuyama, Ehime University" country="Japan" httpUri="http://grbio.org/cool/0wyw-sfpi" name="Ehime University" pageId="13" pageNumber="124">EUMJ</collectionCode>
(
<figureCitation id="132C2A00BE4FFFC0FB40BB058567002C" box="[1192,1312,1118,1145]" captionStart="Fig" captionStartId="9.[189,232,1886,1912]" captionTargetBox="[196,1392,306,1839]" captionTargetId="figure-11@9.[189,1398,268,1842]" captionTargetPageId="9" captionText="Fig. 3. Holotypes of Georissus spp. A–F. Dorsal habitus. G–L. Labels. A, K. G. (G.) babai Satô, 1970, ♂ (EUMJ). B, L. G. (Ni.) granulosus Satô, 1972, ♀ (EUMJ). C, H. G. (Ne.) kurosawai Nakane, 1963, ♂ (SEHU). D, J. G. (Ne.) japonicus Satô, 1972, ♂ (EUMJ). E, G. G. (Ne.) sakaii Satô, 1972, ♂ (EUMJ). F, I. G. (Ne.) katsuoi Nakane, 1995 (SEHU). Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6517956" httpUri="https://zenodo.org/record/6517956/files/figure.png" pageId="13" pageNumber="124">
Fig. 3
<collectionCode id="ED06AE40BE4FFFC0FB05BB048542002C" box="[1261,1285,1119,1145]" pageId="13" pageNumber="124">D</collectionCode>
,
<collectionCode id="ED06AE40BE4FFFC0FAFABB048567002C" box="[1298,1312,1119,1145]" pageId="13" pageNumber="124">J</collectionCode>
</figureCitation>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BA83685BE4FFFC0FF02BBFB82FB00EF" blockId="13.[189,1398,1183,1246]" box="[234,700,1183,1210]" pageId="13" pageNumber="124">
<emphasis id="B963EA97BE4FFFC0FF02BBFB82FB00EF" bold="true" box="[234,700,1183,1210]" pageId="13" pageNumber="124">
<typeStatus id="54AC8827BE4FFFC0FF02BBFB811B00EF" box="[234,348,1184,1210]" pageId="13" pageNumber="124" type="holotype">Holotype</typeStatus>
of
<taxonomicName id="4C174D06BE4FFFC0FE6CBBFB82FB00EF" authority="Sato, 1972" authorityName="Sato" authorityYear="1972" box="[388,700,1183,1210]" class="Insecta" family="Georissidae" genus="Georissus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="124" phylum="Arthropoda" rank="species" species="sakaii" subGenus="Ne.">
<emphasis id="B963EA97BE4FFFC0FE6CBBFB81E700EF" bold="true" box="[388,416,1184,1210]" italics="true" pageId="13" pageNumber="124">G.</emphasis>
(
<emphasis id="B963EA97BE4FFFC0FE59BBFB819C00EF" bold="true" box="[433,475,1184,1210]" italics="true" pageId="13" pageNumber="124">Ne.</emphasis>
)
<emphasis id="B963EA97BE4FFFC0FE04BBC4827300EF" bold="true" box="[492,564,1183,1210]" italics="true" pageId="13" pageNumber="124">sakaii</emphasis>
<bibRefCitation id="EF864B74BE4FFFC0FDD3BBC482FB00EF" author="Sato M." box="[571,700,1183,1210]" pageId="13" pageNumber="124" pagination="207 - 213" refId="ref14083" refString="Sato M. 1972. The Georissid beetles of Japan. Journal of Nagoya Women's College 18: 207 - 213." type="journal article" year="1972">Satô, 1972</bibRefCitation>
</taxonomicName>
</emphasis>
</paragraph>
<paragraph id="8BA83685BE4FFFC0FF55BB9885310088" blockId="13.[189,1398,1183,1246]" box="[189,1398,1219,1246]" pageId="13" pageNumber="124">
<materialsCitation id="3B7F3CD8BE4FFFC0FF55BB9885350088" ID-GBIF-Occurrence="3762863361" box="[189,1394,1219,1246]" collectingDate="1966-09-11" collectionCode="EUMJ" collectorName="M. Sakai" country="Japan" location="Tokushima Pref." pageId="13" pageNumber="124" specimenCount="1" specimenCount-male="1" stateProvince="Tokushima">
<collectingCountry id="F3007615BE4FFFC0FF55BB9881520088" box="[189,277,1219,1245]" name="Japan" pageId="13" pageNumber="124">JAPAN</collectingCountry>
–
<emphasis id="B963EA97BE4FFFC0FEC6BB9882370088" bold="true" box="[302,624,1219,1245]" pageId="13" pageNumber="124">
Shikoku:
<collectingRegion id="49D3F867BE4FFFC0FE4CBB9882370088" box="[420,624,1219,1245]" country="Japan" name="Tokushima" pageId="13" pageNumber="124">Tokushima Pref.</collectingRegion>
</emphasis>
•
<specimenCount id="9D11FD0CBE4FFFC0FD6DBB9F82D9008B" box="[645,670,1220,1246]" pageId="13" pageNumber="124" type="male">♂</specimenCount>
; Tsuda Beach;
<date id="FFA91045BE4FFFC0FCA1BB9883A50088" box="[841,994,1219,1245]" pageId="13" pageNumber="124" value="1966-09-11">
<collectingDate id="EFEDE9ADBE4FFFC0FCA1BB9883A50088" box="[841,994,1219,1245]" pageId="13" pageNumber="124" value="1966-09-11">11 Sep. 1966</collectingDate>
</date>
;
<collectorName id="26E25353BE4FFFC0FC04BB9F84130088" box="[1004,1108,1219,1246]" pageId="13" pageNumber="124">M. Sakai</collectorName>
leg.;
<collectionCode id="ED06AE40BE4FFFC0FB7ABB9F8499008B" box="[1170,1246,1220,1246]" collectionName="Japan, Matsuyama, Ehime University" country="Japan" httpUri="http://grbio.org/cool/0wyw-sfpi" name="Ehime University" pageId="13" pageNumber="124">EUMJ</collectionCode>
(
<figureCitation id="132C2A00BE4FFFC0FB04BB98852E0088" box="[1260,1385,1219,1245]" captionStart="Fig" captionStartId="9.[189,232,1886,1912]" captionTargetBox="[196,1392,306,1839]" captionTargetId="figure-11@9.[189,1398,268,1842]" captionTargetPageId="9" captionText="Fig. 3. Holotypes of Georissus spp. A–F. Dorsal habitus. G–L. Labels. A, K. G. (G.) babai Satô, 1970, ♂ (EUMJ). B, L. G. (Ni.) granulosus Satô, 1972, ♀ (EUMJ). C, H. G. (Ne.) kurosawai Nakane, 1963, ♂ (SEHU). D, J. G. (Ne.) japonicus Satô, 1972, ♂ (EUMJ). E, G. G. (Ne.) sakaii Satô, 1972, ♂ (EUMJ). F, I. G. (Ne.) katsuoi Nakane, 1995 (SEHU). Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6517956" httpUri="https://zenodo.org/record/6517956/files/figure.png" pageId="13" pageNumber="124">
Fig. 3
<collectionCode id="ED06AE40BE4FFFC0FAD9BB9885010088" box="[1329,1350,1219,1245]" pageId="13" pageNumber="124">E</collectionCode>
,
<collectionCode id="ED06AE40BE4FFFC0FAB8BB98852E0088" box="[1360,1385,1219,1245]" pageId="13" pageNumber="124">G</collectionCode>
</figureCitation>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BA83685BE4FFFC0FF02BA5E82B2014B" blockId="13.[189,1398,1284,1381]" box="[234,757,1284,1311]" pageId="13" pageNumber="124">
<emphasis id="B963EA97BE4FFFC0FF02BA5E82B2014B" bold="true" box="[234,757,1284,1311]" pageId="13" pageNumber="124">
<typeStatus id="54AC8827BE4FFFC0FF02BA5E811B014A" box="[234,348,1285,1311]" pageId="13" pageNumber="124" type="holotype">Holotype</typeStatus>
of
<taxonomicName id="4C174D06BE4FFFC0FE6CBA5F82B2014B" authority="Nakane, 1995" authorityName="Nakane" authorityYear="1995" box="[388,757,1284,1311]" class="Insecta" family="Georissidae" genus="Georissus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="124" phylum="Arthropoda" rank="species" species="katsuoi" subGenus="Ne.">
<emphasis id="B963EA97BE4FFFC0FE6CBA5F81E7014B" bold="true" box="[388,416,1284,1310]" italics="true" pageId="13" pageNumber="124">G.</emphasis>
(
<emphasis id="B963EA97BE4FFFC0FE59BA5E819C014A" bold="true" box="[433,475,1285,1311]" italics="true" pageId="13" pageNumber="124">Ne.</emphasis>
)
<emphasis id="B963EA97BE4FFFC0FE04BA5F8203014A" bold="true" box="[492,580,1284,1311]" italics="true" pageId="13" pageNumber="124">katsuoi</emphasis>
<bibRefCitation id="EF864B74BE4FFFC0FDA3BA5E82B2014B" author="Nakane T." box="[587,757,1284,1311]" pageId="13" pageNumber="124" pagination="41 - 46" refId="ref13940" refString="Nakane T. 1995. Notes on some little-known beetles (Coleoptera) in Japan. 13. Kita-Kyushu no Konchu 42: 41 - 46." type="journal article" year="1995">Nakane, 1995</bibRefCitation>
</taxonomicName>
</emphasis>
</paragraph>
<paragraph id="8BA83685BE4FFFC0FF55BA7381D10130" blockId="13.[189,1398,1284,1381]" pageId="13" pageNumber="124">
<materialsCitation id="3B7F3CD8BE4FFFC0FF55BA7381D50130" ID-GBIF-Occurrence="3762863328" collectingDate="1993-08-26" collectionCode="SEHU" collectorName="Saito" country="Japan" location="Torinoumi" municipality="Water-machi" pageId="13" pageNumber="124" specimenCount="1" stateProvince="Miyagi">
<collectingCountry id="F3007615BE4FFFC0FF55BA7381510117" box="[189,278,1320,1346]" name="Japan" pageId="13" pageNumber="124">JAPAN</collectingCountry>
–
<emphasis id="B963EA97BE4FFFC0FED9BA7382780114" bold="true" box="[305,575,1319,1346]" pageId="13" pageNumber="124">
Honshu:
<collectingRegion id="49D3F867BE4FFFC0FE4CBA7C82780114" box="[420,575,1319,1345]" country="Japan" name="Miyagi" pageId="13" pageNumber="124">Miyagi Pref.</collectingRegion>
</emphasis>
• sex unknown;
<collectingMunicipality id="6BCCACFFBE4FFFC0FCE9BA7383DD0117" box="[769,922,1320,1346]" pageId="13" pageNumber="124">Water-machi</collectingMunicipality>
,
<location id="8EC8605EBE4FFFC0FC4DBA7384630117" LSID="urn:lsid:plazi:treatment:03BE8793BE4FFFC2FE17BEE483730193:8EC8605EBE4FFFC0FC4DBA7384630117" box="[933,1060,1320,1346]" country="Japan" municipality="Water-machi" name="Torinoumi" pageId="13" pageNumber="124" stateProvince="Miyagi">Torinoumi</location>
;
<date id="FFA91045BE4FFFC0FBD8BA7384960117" box="[1072,1233,1320,1346]" pageId="13" pageNumber="124" value="1993-08-26">
<collectingDate id="EFEDE9ADBE4FFFC0FBD8BA7384960117" box="[1072,1233,1320,1346]" pageId="13" pageNumber="124" value="1993-08-26">26 Aug. 1993</collectingDate>
</date>
; K,
<collectorName id="26E25353BE4FFFC0FAE8BA73857B0117" box="[1280,1340,1320,1346]" pageId="13" pageNumber="124">Saito</collectorName>
leg.;
<collectionCode id="ED06AE40BE4FFFC0FF55BA10814D0130" box="[189,266,1355,1381]" pageId="13" pageNumber="124">SEHU</collectionCode>
(
<figureCitation id="132C2A00BE4FFFC0FEF2BA1081CF0130" box="[282,392,1355,1381]" captionStart="Fig" captionStartId="9.[189,232,1886,1912]" captionTargetBox="[196,1392,306,1839]" captionTargetId="figure-11@9.[189,1398,268,1842]" captionTargetPageId="9" captionText="Fig. 3. Holotypes of Georissus spp. A–F. Dorsal habitus. G–L. Labels. A, K. G. (G.) babai Satô, 1970, ♂ (EUMJ). B, L. G. (Ni.) granulosus Satô, 1972, ♀ (EUMJ). C, H. G. (Ne.) kurosawai Nakane, 1963, ♂ (SEHU). D, J. G. (Ne.) japonicus Satô, 1972, ♂ (EUMJ). E, G. G. (Ne.) sakaii Satô, 1972, ♂ (EUMJ). F, I. G. (Ne.) katsuoi Nakane, 1995 (SEHU). Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6517956" httpUri="https://zenodo.org/record/6517956/files/figure.png" pageId="13" pageNumber="124">
Fig. 3
<collectionCode id="ED06AE40BE4FFFC0FEB6BA1081370130" box="[350,368,1355,1381]" pageId="13" pageNumber="124">F</collectionCode>
,
<collectionCode id="ED06AE40BE4FFFC0FE94BA1081CF0130" box="[380,392,1355,1381]" pageId="13" pageNumber="124">I</collectionCode>
</figureCitation>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BA83685BE4FFFC0FF02BAD781E101F3" blockId="13.[189,1399,1420,1623]" box="[234,422,1420,1446]" pageId="13" pageNumber="124">
<emphasis id="B963EA97BE4FFFC0FF02BAD781E101F3" bold="true" box="[234,422,1420,1446]" pageId="13" pageNumber="124">Other material</emphasis>
</paragraph>
<paragraph id="8BA83685BE4FFFC0FF55BAEB851E0202" blockId="13.[189,1399,1420,1623]" pageId="13" pageNumber="124">
<materialsCitation id="3B7F3CD8BE4FFFC0FF55BAEB8208019C" ID-GBIF-Occurrence="3762863321" box="[189,591,1455,1482]" collectorName="Honshu" country="Japan" location="Niigata Pref." pageId="13" pageNumber="124" specimenCount="1" stateProvince="Niigata">
<collectingCountry id="F3007615BE4FFFC0FF55BAEB8151019F" box="[189,278,1456,1482]" name="Japan" pageId="13" pageNumber="124">JAPAN</collectingCountry>
–
<emphasis id="B963EA97BE4FFFC0FED0BAEB8208019C" bold="true" box="[312,591,1455,1482]" pageId="13" pageNumber="124">
<collectorName id="26E25353BE4FFFC0FED0BAEB81D9019F" box="[312,414,1456,1482]" pageId="13" pageNumber="124">Honshu</collectorName>
:
<collectingRegion id="49D3F867BE4FFFC0FE45BAF48208019C" box="[429,591,1455,1481]" country="Japan" name="Niigata" pageId="13" pageNumber="124">Niigata Pref.</collectingRegion>
</emphasis>
</materialsCitation>
•
<materialsCitation id="3B7F3CD8BE4FFFC0FD85BAEB813301B9" ID-GBIF-Occurrence="3762863377" collectingDate="1961-08-08" collectionCode="EUMJ" collectorName="Murakami &amp; M. Sato" country="Japan" location="Honshu" pageId="13" pageNumber="124" specimenCount="4" stateProvince="Shizuoka">
<specimenCount id="9D11FD0CBE4FFFC0FD85BAEB82EC019F" box="[621,683,1456,1482]" pageId="13" pageNumber="124" type="generic">4 ex.</specimenCount>
;
<collectorName id="26E25353BE4FFFC0FD53BAF4837E019C" box="[699,825,1455,1481]" pageId="13" pageNumber="124">Murakami</collectorName>
;
<date id="FFA91045BE4FFFC0FCAFBAEB8398019C" box="[839,991,1455,1482]" pageId="13" pageNumber="124" value="1961-08-08">
<collectingDate id="EFEDE9ADBE4FFFC0FCAFBAEB8398019C" box="[839,991,1455,1482]" pageId="13" pageNumber="124" value="1961-08-08">8 Aug. 1961</collectingDate>
</date>
;
<collectorName id="26E25353BE4FFFC0FC06BAEB840A019F" box="[1006,1101,1456,1482]" pageId="13" pageNumber="124">M. Satô</collectorName>
leg.;
<collectionCode id="ED06AE40BE4FFFC0FB7CBAEB84A3019F" box="[1172,1252,1456,1482]" collectionName="Japan, Matsuyama, Ehime University" country="Japan" httpUri="http://grbio.org/cool/0wyw-sfpi" name="Ehime University" pageId="13" pageNumber="124">EUMJ</collectionCode>
. –
<emphasis id="B963EA97BE4FFFC0FAE3BAEB813301B9" bold="true" pageId="13" pageNumber="124">
<location id="8EC8605EBE4FFFC0FAE3BAEB8536019F" LSID="urn:lsid:plazi:treatment:03BE8793BE4FFFC2FE17BEE483730193:8EC8605EBE4FFFC0FAE3BAEB8536019F" box="[1291,1393,1456,1482]" country="Japan" name="Honshu" pageId="13" pageNumber="124" stateProvince="Shizuoka">Honshu</location>
:
<collectingRegion id="49D3F867BE4FFFC0FF55BA89813301B9" box="[189,372,1490,1516]" country="Japan" name="Shizuoka" pageId="13" pageNumber="124">Shizuoka Pref.</collectingRegion>
</emphasis>
</materialsCitation>
•
<materialsCitation id="3B7F3CD8BE4FFFC0FE64BA8884C401B8" ID-GBIF-Occurrence="3762863323" box="[396,1155,1491,1518]" collectingDate="2001-05-25" collectionCode="EUMJ" collectorName="K. Hirai" country="Japan" location="Hamamatsu" pageId="13" pageNumber="124" specimenCount="1">
<specimenCount id="9D11FD0CBE4FFFC0FE64BA88818F01B8" box="[396,456,1491,1517]" pageId="13" pageNumber="124" type="generic">1 ex.</specimenCount>
;
<location id="8EC8605EBE4FFFC0FE3DBA88822301B8" LSID="urn:lsid:plazi:treatment:03BE8793BE4FFFC2FE17BEE483730193:8EC8605EBE4FFFC0FE3DBA88822301B8" box="[469,612,1491,1517]" country="Japan" name="Hamamatsu" pageId="13" pageNumber="124">Hamamatsu</location>
,
<location id="8EC8605EBE4FFFC0FD87BA88829901B8" LSID="urn:lsid:plazi:treatment:03BE8793BE4FFFC2FE17BEE483730193:8EC8605EBE4FFFC0FD87BA88829901B8" box="[623,734,1491,1517]" country="Japan" name="Ishiyama" pageId="13" pageNumber="124">Ishiyama</location>
;
<date id="FFA91045BE4FFFC0FD02BA8883CE01B8" box="[746,905,1491,1518]" pageId="13" pageNumber="124" value="2001-05-25">
<collectingDate id="EFEDE9ADBE4FFFC0FD02BA8883CE01B8" box="[746,905,1491,1518]" pageId="13" pageNumber="124" value="2001-05-25">25 May 2001</collectingDate>
</date>
;
<collectorName id="26E25353BE4FFFC0FC7DBA8883B301B8" box="[917,1012,1491,1517]" pageId="13" pageNumber="124">K. Hirai</collectorName>
leg.;
<collectionCode id="ED06AE40BE4FFFC0FBDEBA8884C401B8" box="[1078,1155,1491,1517]" collectionName="Japan, Matsuyama, Ehime University" country="Japan" httpUri="http://grbio.org/cool/0wyw-sfpi" name="Ehime University" pageId="13" pageNumber="124">EUMJ</collectionCode>
</materialsCitation>
•
<materialsCitation id="3B7F3CD8BE4FFFC0FB73BA88829B0244" ID-GBIF-Occurrence="3762863333" collectingDate="2001-10-04" collectionCode="EUMJ" collectorName="K. Hirai" country="Japan" location="Hamamatsu" pageId="13" pageNumber="124" specimenCount="1">
<specimenCount id="9D11FD0CBE4FFFC0FB73BA88849101B8" box="[1179,1238,1491,1517]" pageId="13" pageNumber="124" type="generic">1 ex.</specimenCount>
;
<location id="8EC8605EBE4FFFC0FB0BBA88853501B8" LSID="urn:lsid:plazi:treatment:03BE8793BE4FFFC2FE17BEE483730193:8EC8605EBE4FFFC0FB0BBA88853501B8" box="[1251,1394,1491,1517]" country="Japan" name="Hamamatsu" pageId="13" pageNumber="124">Hamamatsu</location>
,
<location id="8EC8605EBE4FFFC0FF55BAAD810D0245" LSID="urn:lsid:plazi:treatment:03BE8793BE4FFFC2FE17BEE483730193:8EC8605EBE4FFFC0FF55BAAD810D0245" box="[189,330,1526,1552]" country="Japan" name="Enshuhama" pageId="13" pageNumber="124">Enshuhama</location>
;
<date id="FFA91045BE4FFFC0FEBDBAAD81A50245" box="[341,482,1526,1552]" pageId="13" pageNumber="124" value="2001-10-04">
<collectingDate id="EFEDE9ADBE4FFFC0FEBDBAAD81A50245" box="[341,482,1526,1552]" pageId="13" pageNumber="124" value="2001-10-04">4 Oct. 2001</collectingDate>
</date>
;
<collectorName id="26E25353BE4FFFC0FE06BAAC820A0245" box="[494,589,1526,1553]" pageId="13" pageNumber="124">K. Hirai</collectorName>
leg.;
<collectionCode id="ED06AE40BE4FFFC0FD67BAAC829B0244" box="[655,732,1527,1553]" collectionName="Japan, Matsuyama, Ehime University" country="Japan" httpUri="http://grbio.org/cool/0wyw-sfpi" name="Ehime University" pageId="13" pageNumber="124">EUMJ</collectionCode>
</materialsCitation>
•
<materialsCitation id="3B7F3CD8BE4FFFC0FD1CBAAD81020261" ID-GBIF-Occurrence="3762863362" collectingDate="2001-10-13" collectionCode="EUMJ" collectorName="K. Hirai" country="Japan" location="Hamamatsu" pageId="13" pageNumber="124" specimenCount="3">
<specimenCount id="9D11FD0CBE4FFFC0FD1CBAAD83770244" box="[756,816,1526,1553]" pageId="13" pageNumber="124" type="generic">3 ex.</specimenCount>
;
<location id="8EC8605EBE4FFFC0FCD5BAAC838B0244" LSID="urn:lsid:plazi:treatment:03BE8793BE4FFFC2FE17BEE483730193:8EC8605EBE4FFFC0FCD5BAAC838B0244" box="[829,972,1527,1553]" country="Japan" name="Hamamatsu" pageId="13" pageNumber="124">Hamamatsu</location>
,
<location id="8EC8605EBE4FFFC0FC3FBAAD84230245" LSID="urn:lsid:plazi:treatment:03BE8793BE4FFFC2FE17BEE483730193:8EC8605EBE4FFFC0FC3FBAAD84230245" box="[983,1124,1526,1552]" country="Japan" name="Enshuhama" pageId="13" pageNumber="124">Enshuhama</location>
;
<date id="FFA91045BE4FFFC0FB87BAAD854C0245" box="[1135,1291,1526,1552]" pageId="13" pageNumber="124" value="2001-10-13">
<collectingDate id="EFEDE9ADBE4FFFC0FB87BAAD854C0245" box="[1135,1291,1526,1552]" pageId="13" pageNumber="124" value="2001-10-13">13 Oct. 2001</collectingDate>
</date>
;
<collectorName id="26E25353BE4FFFC0FAFEBAAC85310245" box="[1302,1398,1526,1553]" pageId="13" pageNumber="124">K. Hirai</collectorName>
leg.;
<collectionCode id="ED06AE40BE4FFFC0FF10B94181020261" box="[248,325,1562,1588]" collectionName="Japan, Matsuyama, Ehime University" country="Japan" httpUri="http://grbio.org/cool/0wyw-sfpi" name="Ehime University" pageId="13" pageNumber="124">EUMJ</collectionCode>
</materialsCitation>
•
<materialsCitation id="3B7F3CD8BE4FFFC0FEB6B94281BD0203" ID-GBIF-Occurrence="3762863379" collectingDate="2002-10-13" collectionCode="EUMJ" collectorName="Hamamatsu-shi &amp; Matsushima-cho &amp; Enshuhama &amp; K. Hirai" country="Japan" location="Kyushu" pageId="13" pageNumber="124" specimenCount="6" stateProvince="Fukuoka">
<specimenCount id="9D11FD0CBE4FFFC0FEB6B94281DE0261" box="[350,409,1561,1588]" pageId="13" pageNumber="124" type="generic">6 ex.</specimenCount>
;
<collectorName id="26E25353BE4FFFC0FE4EB94282260266" box="[422,609,1561,1587]" pageId="13" pageNumber="124">Hamamatsu-shi</collectorName>
,
<collectorName id="26E25353BE4FFFC0FD85B94283730266" box="[621,820,1561,1587]" pageId="13" pageNumber="124">Matsushima-cho</collectorName>
,
<collectorName id="26E25353BE4FFFC0FCD7B942838B0266" box="[831,972,1561,1587]" pageId="13" pageNumber="124">Enshuhama</collectorName>
;
<date id="FFA91045BE4FFFC0FC30B94284330266" box="[984,1140,1561,1587]" pageId="13" pageNumber="124" value="2002-10-13">
<collectingDate id="EFEDE9ADBE4FFFC0FC30B94284330266" box="[984,1140,1561,1587]" pageId="13" pageNumber="124" value="2002-10-13">13 Oct. 2002</collectingDate>
</date>
;
<collectorName id="26E25353BE4FFFC0FB68B94184A70266" box="[1152,1248,1561,1588]" pageId="13" pageNumber="124">K. Hirai</collectorName>
leg.;
<collectionCode id="ED06AE40BE4FFFC0FACAB94185350261" box="[1314,1394,1562,1588]" collectionName="Japan, Matsuyama, Ehime University" country="Japan" httpUri="http://grbio.org/cool/0wyw-sfpi" name="Ehime University" pageId="13" pageNumber="124">EUMJ</collectionCode>
. –
<emphasis id="B963EA97BE4FFFC0FF3BB96681BD0203" bold="true" box="[211,506,1596,1623]" pageId="13" pageNumber="124">
<location id="8EC8605EBE4FFFC0FF3BB966817E0202" LSID="urn:lsid:plazi:treatment:03BE8793BE4FFFC2FE17BEE483730193:8EC8605EBE4FFFC0FF3BB966817E0202" box="[211,313,1597,1623]" country="Japan" name="Kyushu" pageId="13" pageNumber="124" stateProvince="Fukuoka">Kyushu</location>
:
<collectingRegion id="49D3F867BE4FFFC0FEAEB96681BD0203" box="[326,506,1596,1623]" country="Japan" name="Fukuoka" pageId="13" pageNumber="124">Fukuoka Pref.</collectingRegion>
</emphasis>
</materialsCitation>
•
<materialsCitation id="3B7F3CD8BE4FFFC0FDFBB96685120202" ID-GBIF-Occurrence="3762863349" box="[531,1365,1596,1623]" collectingDate="2013-07-23" collectionCode="EUMJ" collectorName="M. Mori" country="Japan" location="Kouge-machi" pageId="13" pageNumber="124" specimenCount="8">
<specimenCount id="9D11FD0CBE4FFFC0FDFBB96682080202" box="[531,591,1597,1623]" pageId="13" pageNumber="124" type="generic">8 ex.</specimenCount>
;
<location id="8EC8605EBE4FFFC0FDB4B96782B80203" LSID="urn:lsid:plazi:treatment:03BE8793BE4FFFC2FE17BEE483730193:8EC8605EBE4FFFC0FDB4B96782B80203" box="[604,767,1596,1622]" country="Japan" name="Kouge-machi" pageId="13" pageNumber="124">Kouge-machi</location>
,
<location id="8EC8605EBE4FFFC0FCE2B96783F30203" LSID="urn:lsid:plazi:treatment:03BE8793BE4FFFC2FE17BEE483730193:8EC8605EBE4FFFC0FCE2B96783F30203" box="[778,948,1596,1622]" country="Japan" name="Shimotoubaru" pageId="13" pageNumber="124">Shimotoubaru</location>
;
<date id="FFA91045BE4FFFC0FC28B96684140202" box="[960,1107,1596,1623]" pageId="13" pageNumber="124" value="2013-07-23">
<collectingDate id="EFEDE9ADBE4FFFC0FC28B96684140202" box="[960,1107,1596,1623]" pageId="13" pageNumber="124" value="2013-07-23">23 Jul. 2013</collectingDate>
</date>
;
<collectorName id="26E25353BE4FFFC0FB88B96684840203" box="[1120,1219,1596,1623]" pageId="13" pageNumber="124">M. Mori</collectorName>
leg.;
<collectionCode id="ED06AE40BE4FFFC0FAEDB96685120202" box="[1285,1365,1597,1623]" collectionName="Japan, Matsuyama, Ehime University" country="Japan" httpUri="http://grbio.org/cool/0wyw-sfpi" name="Ehime University" pageId="13" pageNumber="124">EUMJ</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C30D650EBE4FFFC2FF55B925837E07D3" lastPageId="15" lastPageNumber="126" pageId="13" pageNumber="124" type="description">
<paragraph id="8BA83685BE4FFFC0FF55B925812C02CD" blockId="13.[189,363,1662,1688]" box="[189,363,1662,1688]" pageId="13" pageNumber="124">
<heading id="D0E081E9BE4FFFC0FF55B925812C02CD" bold="true" box="[189,363,1662,1688]" fontSize="11" level="3" pageId="13" pageNumber="124" reason="3">
<emphasis id="B963EA97BE4FFFC0FF55B925812C02CD" bold="true" box="[189,363,1662,1688]" pageId="13" pageNumber="124">Redescription</emphasis>
</heading>
</paragraph>
<paragraph id="8BA83685BE4FFFC0FF55B9F7852C0293" blockId="13.[189,1387,1708,1735]" box="[189,1387,1708,1735]" pageId="13" pageNumber="124">
COLORATION (
<figureCitation id="132C2A00BE4FFFC0FEB1B9F781F00293" box="[345,439,1708,1735]" captionStart="Fig" captionStartId="7.[189,232,1886,1912]" captionTargetBox="[272,1331,265,1833]" captionTargetId="figure-11@7.[223,1365,265,1845]" captionTargetPageId="7" captionText="Fig. 2. Dorsal view of Georissus spp. A. G. (G.) canalifer Sharp, 1888. B. G. (G.) babai Satô, 1970. C. G. (Ni.) granulosus Satô, 1972. D. G. (Ne.) japonicus Satô, 1972. E. G. (Ne.) kurosawai Nakane, 1963. F. G. (Ne.) takahashii sp. nov., holotype, ♂ (EUMJ). G. G. (Ne.) satoi sp. nov., holotype (EUMJ). Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6517952" httpUri="https://zenodo.org/record/6517952/files/figure.png" pageId="13" pageNumber="124">Fig. 2D</figureCitation>
). Black to dark brown, matt. Body surface often covered with grayish substance.
</paragraph>
<paragraph id="8BA83685BE4FFFC0FF55B9B5847C0324" blockId="13.[188,1399,1774,1906]" pageId="13" pageNumber="124">
HEAD (
<figureCitation id="132C2A00BE4FFFC0FEFBB9B58131035C" box="[275,374,1774,1801]" captionStart="Fig" captionStartId="18.[189,232,1882,1908]" captionTargetBox="[191,1394,563,1839]" captionTargetId="figure-90@18.[189,1398,562,1842]" captionTargetPageId="18" captionText="Fig. 6. Head of Georissus spp. A. G. (G.) canalifer Sharp, 1888. B. G. (G.) babai Satô, 1970. C. G. (Ni.) granulosus Satô, 1972. D. G. (Ne.) japonicus Satô, 1972. E–F. G. (Ne.) kurosawai Nakane, 1963 from Hokkaido (E) and Nagano (F). G–H. G. (Ne.) takahashii sp. nov. from Kochi (G) and Okinawa (H). I. G. (Ne.) satoi sp. nov." figureDoi="http://doi.org/10.5281/zenodo.6517968" httpUri="https://zenodo.org/record/6517968/files/figure.png" pageId="13" pageNumber="124">Fig. 6D</figureCitation>
). Clypeus uneven, provided with large granules sparsely;
<emphasis id="B963EA97BE4FFFC0FBABB9B58425035D" box="[1091,1122,1774,1800]" italics="true" pageId="13" pageNumber="124">slr</emphasis>
consisted of granules, reaching to posterior third of frons, posteriorly connected to
<emphasis id="B963EA97BE4FFFC0FC92B84A83EE037E" box="[890,937,1809,1835]" italics="true" pageId="13" pageNumber="124">smb</emphasis>
by an arc of sparsely arranged granules and formed a pair of half-rounded foveae;
<emphasis id="B963EA97BE4FFFC0FD20B86F82B0031B" box="[712,759,1844,1870]" italics="true" pageId="13" pageNumber="124">smb</emphasis>
consisted of tubercles and granules, anterior portion projecting to laterally;
<emphasis id="B963EA97BE4FFFC0FE21B80C81AC0324" box="[457,491,1879,1905]" italics="true" pageId="13" pageNumber="124">lsa</emphasis>
with a shallow groove along the posterior margin.
</paragraph>
<paragraph id="8BA83685BE4FFFC2FF55B8C28320053D" blockId="13.[189,1399,1944,2006]" lastBlockId="15.[188,1399,265,361]" lastPageId="15" lastPageNumber="126" pageId="13" pageNumber="124">
PRONOTUM (
<figureCitation id="132C2A00BE4FFFC0FEA2B8C381EC03E6" box="[330,427,1944,1971]" captionStart="Fig" captionStartId="19.[189,232,1879,1905]" captionTargetBox="[197,1389,792,1835]" captionTargetId="figure-118@19.[189,1398,783,1839]" captionTargetPageId="19" captionText="Fig. 7. Pronotum of Georissus spp. A. G. (G.) canalifer Sharp, 1888. B. G. (G.) babai Satô, 1970. C. G. (Ni.) granulosus Satô, 1972. D. G. (Ne.) japonicus Satô, 1972. E–F. G. (Ne.) kurosawai Nakane, 1963 from Hokkaido (E) and Nagano (F). G–H. G. (Ne.) takahashii sp. nov. from Kochi (G) and Okinawa (H). I. G. (Ne.) satoi sp. nov." figureDoi="http://doi.org/10.5281/zenodo.6517976" httpUri="https://zenodo.org/record/6517976/files/figure.png" pageId="13" pageNumber="124">Fig. 7D</figureCitation>
). Relatively large, widest at the middle. Anterior portion provided with granules densely;
<emphasis id="B963EA97BE4FFFC0FECEB8E0811A0380" box="[294,349,1979,2005]" italics="true" pageId="13" pageNumber="124">smlg</emphasis>
relatively wide, granulation dense, with longitudinal row of sparse pits. Posterior portion somewhat convex than anterior portion;
<emphasis id="B963EA97BE4DFFC2FD75BE5282FE0576" box="[669,697,265,291]" italics="true" pageId="15" pageNumber="126">cd</emphasis>
rhomboid, surrounded by granules; s
<emphasis id="B963EA97BE4DFFC2FB9ABE5284C80576" box="[1138,1167,265,291]" italics="true" pageId="15" pageNumber="126">dp</emphasis>
oblong oval, larger than
<emphasis id="B963EA97BE4DFFC2FF1DBE7781560513" box="[245,273,300,326]" italics="true" pageId="15" pageNumber="126">cd</emphasis>
, surrounded by granules;
<emphasis id="B963EA97BE4DFFC2FDD4BE7782190513" box="[572,606,300,326]" italics="true" pageId="15" pageNumber="126">slb</emphasis>
developed, with distinct granules;
<emphasis id="B963EA97BE4DFFC2FC1ABE7784480510" box="[1010,1039,300,325]" italics="true" pageId="15" pageNumber="126">pg</emphasis>
short and narrow, not reaching median portion of
<emphasis id="B963EA97BE4DFFC2FE7EBE1581BE053D" box="[406,505,334,360]" italics="true" pageId="15" pageNumber="126">sdp; sdn</emphasis>
developed, having a few teeth.
</paragraph>
<caption id="DF68660DBE4CFFC3FF55B80F853003E3" ID-DOI="http://doi.org/10.5281/zenodo.6517962" ID-Zenodo-Dep="6517962" httpUri="https://zenodo.org/record/6517962/files/figure.png" pageId="14" pageNumber="125" startId="14.[189,232,1876,1902]" targetBox="[196,1386,277,1817]" targetPageId="14">
<paragraph id="8BA83685BE4CFFC3FF55B80F853003E3" blockId="14.[188,1399,1876,1974]" pageId="14" pageNumber="125">
<emphasis id="B963EA97BE4CFFC3FF55B80F8149033B" bold="true" box="[189,270,1876,1902]" pageId="14" pageNumber="125">Fig. 4.</emphasis>
Mouth parts and antennae of
<taxonomicName id="4C174D06BE4CFFC3FD96B80E830B033B" box="[638,844,1876,1903]" class="Insecta" family="Georissidae" genus="Georissus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="125" phylum="Arthropoda" rank="species" species="undefined-a">
<emphasis id="B963EA97BE4CFFC3FD96B80E82B4033A" box="[638,755,1877,1903]" italics="true" pageId="14" pageNumber="125">Georissus</emphasis>
spp.
<emphasis id="B963EA97BE4CFFC3FCDFB80F830B033B" bold="true" box="[823,844,1876,1902]" pageId="14" pageNumber="125">A</emphasis>
</taxonomicName>
<emphasis id="B963EA97BE4CFFC3FCA5B80F8337033B" bold="true" box="[845,880,1876,1902]" pageId="14" pageNumber="125">–C</emphasis>
. Left maxillae.
<emphasis id="B963EA97BE4CFFC3FBDBB80E842E033A" bold="true" box="[1075,1129,1877,1903]" pageId="14" pageNumber="125">D–F</emphasis>
. Labrum.
<emphasis id="B963EA97BE4CFFC3FB01B80F855D033B" bold="true" box="[1257,1306,1876,1902]" pageId="14" pageNumber="125">G–I</emphasis>
. Labial palpi.
<emphasis id="B963EA97BE4CFFC3FEEEB823817103C7" bold="true" box="[262,310,1912,1938]" pageId="14" pageNumber="125">J–L</emphasis>
. Left mandibles.
<emphasis id="B963EA97BE4CFFC3FDEAB82C820403C4" bold="true" box="[514,579,1911,1937]" pageId="14" pageNumber="125">M–O</emphasis>
. Left antennae.
<emphasis id="B963EA97BE4CFFC3FD16B82C83E503C7" bold="true" box="[766,930,1911,1938]" pageId="14" pageNumber="125">A, D, G, J, M</emphasis>
.
<emphasis id="B963EA97BE4CFFC3FC58B823838203C7" box="[944,965,1912,1938]" italics="true" pageId="14" pageNumber="125">G</emphasis>
.
<taxonomicName id="4C174D06BE4CFFC3FC3BB823854C03C7" authority="Sharp, 1888" authorityName="Sharp" authorityYear="1888" box="[979,1291,1912,1938]" class="Insecta" family="Georissidae" genus="Georissus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="125" phylum="Arthropoda" rank="species" species="canalifer" subGenus="Georissus">
(
<emphasis id="B963EA97BE4CFFC3FC35B82383B503C7" box="[989,1010,1912,1938]" italics="true" pageId="14" pageNumber="125">G</emphasis>
.)
<emphasis id="B963EA97BE4CFFC3FBE2B823843303C7" box="[1034,1140,1912,1938]" italics="true" pageId="14" pageNumber="125">canalifer</emphasis>
<bibRefCitation id="EF864B74BE4CFFC3FB92B823854C03C7" author="Sharp D." box="[1146,1291,1912,1938]" pageId="14" pageNumber="125" pagination="242 - 245" refId="ref14169" refString="Sharp D. 1888. Descriptions of some new Coleoptera from Japan. The Annals and Magazine of Natural History Series 6 2: 242 - 245. https: // doi. org / 10.1080 / 00222938809460918" type="journal article" year="1888">Sharp, 1888</bibRefCitation>
</taxonomicName>
.
<emphasis id="B963EA97BE4CFFC3FAFDB82380B103E0" bold="true" pageId="14" pageNumber="125">B, E, H, K, N</emphasis>
.
<emphasis id="B963EA97BE4CFFC3FEEDB8C0815D03E0" box="[261,282,1947,1973]" italics="true" pageId="14" pageNumber="125">G</emphasis>
.
<taxonomicName id="4C174D06BE4CFFC3FEC0B8C0822903E0" authority="Sato, 1972" authorityName="Sato" authorityYear="1972" box="[296,622,1947,1974]" class="Insecta" family="Georissidae" genus="Georissus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="125" phylum="Arthropoda" rank="species" species="granulosus" subGenus="Ni .">
(
<emphasis id="B963EA97BE4CFFC3FEDAB8C7810903E0" box="[306,334,1948,1973]" italics="true" pageId="14" pageNumber="125">Ni</emphasis>
.)
<emphasis id="B963EA97BE4CFFC3FE8DB8C081A003E0" box="[357,487,1947,1973]" italics="true" pageId="14" pageNumber="125">granulosus</emphasis>
<bibRefCitation id="EF864B74BE4CFFC3FE06B8C0822903E0" author="Sato M." box="[494,622,1947,1973]" pageId="14" pageNumber="125" pagination="207 - 213" refId="ref14083" refString="Sato M. 1972. The Georissid beetles of Japan. Journal of Nagoya Women's College 18: 207 - 213." type="journal article" year="1972">Satô, 1972</bibRefCitation>
</taxonomicName>
.
<emphasis id="B963EA97BE4CFFC3FD90B8C0834D03E0" bold="true" box="[632,778,1947,1973]" pageId="14" pageNumber="125">C, F, I, L, O</emphasis>
.
<taxonomicName id="4C174D06BE4CFFC3FCF1B8C0843003E0" authority="Sato, 1972" authorityName="Sato" authorityYear="1972" box="[793,1143,1947,1974]" class="Insecta" family="Georissidae" genus="Georissus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="124" phylum="Arthropoda" rank="species" species="japonicus" subGenus="Ne.">
<emphasis id="B963EA97BE4CFFC3FCF1B8C0836903E0" box="[793,814,1947,1973]" italics="true" pageId="14" pageNumber="125">G</emphasis>
. (
<emphasis id="B963EA97BE4CFFC3FCAEB8C7832003E0" box="[838,871,1948,1973]" italics="true" pageId="14" pageNumber="125">Ne</emphasis>
.)
<emphasis id="B963EA97BE4CFFC3FC96B8C783B703E0" box="[894,1008,1948,1973]" italics="true" pageId="14" pageNumber="125">japonicus</emphasis>
<bibRefCitation id="EF864B74BE4CFFC3FC1FB8C0843003E0" author="Sato M." box="[1015,1143,1947,1973]" pageId="14" pageNumber="125" pagination="207 - 213" refId="ref14083" refString="Sato M. 1972. The Georissid beetles of Japan. Journal of Nagoya Women's College 18: 207 - 213." type="journal article" year="1972">Satô, 1972</bibRefCitation>
</taxonomicName>
. Scale bars = 0.1 mm.
</paragraph>
</caption>
<paragraph id="8BA83685BE4DFFC2FF55BED3818D05B2" blockId="15.[188,1399,392,487]" pageId="15" pageNumber="126">ELYTRA. Humeral bulges distinct. All intervals slightly elevated, provided with distinct lows of granules. Elytral punctures distinct. Lateral ridges on each elytron highly elevate, bearing distinct granules. Hind wings fully developed.</paragraph>
<paragraph id="8BA83685BE4DFFC2FF55BD5C84EE0611" blockId="15.[189,1399,518,580]" pageId="15" pageNumber="126">
ABDOMEN (
<figureCitation id="132C2A00BE4DFFC2FED7BD5D81D90675" box="[319,414,518,545]" captionStart="Fig" captionStartId="21.[191,234,1890,1916]" captionTargetBox="[193,1394,569,1846]" captionTargetId="figure-107@21.[189,1398,569,1852]" captionTargetPageId="21" captionText="Fig. 8. Ventrites of Georissus spp. A. G. (G.) canalifer Sharp, 1888. B. G. (G.) babai Satô, 1970. C. G. (Ni.) granulosus Satô, 1972. D. G. (Ne.) japonicus Satô, 1972. E–F. G. (Ne.) kurosawai Nakane, 1963 from Hokkaido (E) and Nagano (F). G–H. G. (Ne.) takahashii sp. nov. from Kochi (G) and Okinawa (F). I. G. (Ne.) satoi sp. nov. Abbreviation: vt = ventral teeth. Scale bars = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.6517982" httpUri="https://zenodo.org/record/6517982/files/figure.png" pageId="15" pageNumber="126">Fig. 8D</figureCitation>
). Lateral margin of ventrites uniformly narrowing posterior. Granules of ventrite 1 large and dense. Granulation of ventrites 2–4 indistinct, ventrite 5 distinct and sparse.
</paragraph>
<paragraph id="8BA83685BE4DFFC2FF55BD3881AF0752" blockId="15.[189,1399,610,775]" pageId="15" pageNumber="126">
MALE GENITALIA (
<figureCitation id="132C2A00BE4DFFC2FE67BD3981B60629" box="[399,497,610,637]" captionStart="Fig" captionStartId="23.[189,232,1763,1789]" captionTargetBox="[239,1353,284,1696]" captionTargetId="figure-12@23.[189,1398,265,1704]" captionTargetPageId="23" captionText="Fig. 9. Male aedeagus of Georissus spp. A. G. (G.) canalifer Sharp, 1888. B. G. (G.) babai Satô, 1970. C. G. (Ni.) granulosus Satô, 1972. D. G. (Ne.) japonicus Satô, 1972. E. G. (Ne.) kurosawai Nakane, 1963. F–G. G. (Ne.) takahashii sp. nov. from Kochi (F) and Okinawa (G). H. G. (Ne.) satoi sp. nov. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6517986" httpUri="https://zenodo.org/record/6517986/files/figure.png" pageId="15" pageNumber="126">Fig. 9D</figureCitation>
). Aedeagus
<quantity id="4CEF9B60BE4DFFC2FD60BD3982B20628" box="[648,757,610,637]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="15" pageNumber="126" unit="mm" value="0.3">0.30 mm</quantity>
long. Parameres shorter than phallobase, their basal portions combined as wide as anterior portion of phallobase; lateral margins bulbous at median portion; internal margins straight. Median lobe shorter than parameres, subparallel-sided, slightly angular at apex; basal struts short. Phallobase about 2.5 times as long as wide, sides paralleled: posterior portion with large basal foramen.
</paragraph>
<paragraph id="8BA83685BE4DFFC2FF55BC7C837E07D3" blockId="15.[189,1399,806,902]" pageId="15" pageNumber="126">
FEMALE GENITALIA. Latero-tergites 8 forming acute angle (
<figureCitation id="132C2A00BE4DFFC2FC69BC7D83B50715" box="[897,1010,806,832]" captionStart="Fig" captionStartId="24.[189,232,1780,1806]" captionTargetBox="[233,1337,288,1718]" captionTargetId="figure-18@24.[189,1398,264,1722]" captionTargetPageId="24" captionText="Fig. 10. Female ovipositor of Georissus spp. A. G. (G.) canalifer Sharp, 1888. B. G. (G.) babai Satô, 1970. C. G. (Ne.) japonicus Satô, 1972. D. G. (Ne.) kurosawai Nakane, 1963. E. G. (Ne.) takahashii sp. nov. F. G. (Ne.) satoi sp. nov. Abbreviations: gsty = gonostylus; ltg8 = latero-tergite 8. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6517988" httpUri="https://zenodo.org/record/6517988/files/figure.png" pageId="15" pageNumber="126">Fig. 10C</figureCitation>
); apex strongly incurved. Setae of gonostylus short. Spermatheca (
<figureCitation id="132C2A00BE4DFFC2FDB5BC1282820736" box="[605,709,841,867]" captionStart="Fig" captionStartId="25.[189,232,1920,1946]" captionTargetBox="[213,1341,1093,1867]" captionTargetId="figure-278@25.[189,1398,1073,1882]" captionTargetPageId="25" captionText="Fig. 11. Female genital organs of subgenus Neogeorissus spp. A–D. Bursal sclerite. E–H. Spermatheca. A, E. G. (Ne.) japonicus Satô, 1972. B, F. G. (Ne.) kurosawai Nakane, 1963. C, G. G. (Ne.) takahashii sp. nov. D, H. G. (Ne.) satoi sp. nov. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6517990" httpUri="https://zenodo.org/record/6517990/files/figure.png" pageId="15" pageNumber="126">Fig. 11E</figureCitation>
) curved oval, narrowing at apical portion. Bursal sclerite (
<figureCitation id="132C2A00BE4DFFC2FF20BC37817507D3" box="[200,306,876,902]" captionStart="Fig" captionStartId="25.[189,232,1920,1946]" captionTargetBox="[213,1341,1093,1867]" captionTargetId="figure-278@25.[189,1398,1073,1882]" captionTargetPageId="25" captionText="Fig. 11. Female genital organs of subgenus Neogeorissus spp. A–D. Bursal sclerite. E–H. Spermatheca. A, E. G. (Ne.) japonicus Satô, 1972. B, F. G. (Ne.) kurosawai Nakane, 1963. C, G. G. (Ne.) takahashii sp. nov. D, H. G. (Ne.) satoi sp. nov. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6517990" httpUri="https://zenodo.org/record/6517990/files/figure.png" pageId="15" pageNumber="126">Fig. 11A</figureCitation>
) large and asymmetrical, feebly sclerotized.
</paragraph>
</subSubSection>
<subSubSection id="C30D650EBE4DFFC2FF55BCF684C20160" pageId="15" pageNumber="126" type="discussion">
<paragraph id="8BA83685BE4DFFC2FF55BCF681680792" blockId="15.[189,303,941,967]" box="[189,303,941,967]" pageId="15" pageNumber="126">
<heading id="D0E081E9BE4DFFC2FF55BCF681680792" bold="true" box="[189,303,941,967]" fontSize="11" level="3" pageId="15" pageNumber="126" reason="3">
<emphasis id="B963EA97BE4DFFC2FF55BCF681680792" bold="true" box="[189,303,941,967]" pageId="15" pageNumber="126">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8BA83685BE4DFFC2FF55BC80835800F7" blockId="15.[189,1400,987,1187]" pageId="15" pageNumber="126">
<bibRefCitation id="EF864B74BE4DFFC2FF55BC80810E07A0" author="Sato M." box="[189,329,987,1013]" pageId="15" pageNumber="126" pagination="207 - 213" refId="ref14083" refString="Sato M. 1972. The Georissid beetles of Japan. Journal of Nagoya Women's College 18: 207 - 213." type="journal article" year="1972">Satô (1972)</bibRefCitation>
mentioned that
<taxonomicName id="4C174D06BE4DFFC2FDE7BC8083D707A0" authorityName="Sato" authorityYear="1972" box="[527,912,987,1013]" class="Insecta" family="Georissidae" genus="Georissus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="126" phylum="Arthropoda" rank="species" species="sakaii" subGenus="Neogeorissus">
<emphasis id="B963EA97BE4DFFC2FDE7BC8082C307A0" box="[527,644,987,1013]" italics="true" pageId="15" pageNumber="126">Georissus</emphasis>
(
<emphasis id="B963EA97BE4DFFC2FD70BC80837107A1" box="[664,822,987,1012]" italics="true" pageId="15" pageNumber="126">Neogeorissus</emphasis>
)
<emphasis id="B963EA97BE4DFFC2FCA2BC8083D707A0" box="[842,912,987,1013]" italics="true" pageId="15" pageNumber="126">sakaii</emphasis>
</taxonomicName>
was closely related to
<taxonomicName id="4C174D06BE4DFFC2FB41BC80850507A1" authorityName="Sato" authorityYear="1972" box="[1193,1346,987,1013]" class="Insecta" family="Georissidae" genus="Georissus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="126" phylum="Arthropoda" rank="species" species="japonicus">
<emphasis id="B963EA97BE4DFFC2FB41BC8084F907A0" box="[1193,1214,987,1013]" italics="true" pageId="15" pageNumber="126">G</emphasis>
.
<emphasis id="B963EA97BE4DFFC2FB38BC80850507A1" box="[1232,1346,987,1012]" italics="true" pageId="15" pageNumber="126">japonicus</emphasis>
</taxonomicName>
and distinguished by granules and foveae on body surface. Subsequently,
<bibRefCitation id="EF864B74BE4DFFC2FC31BCA684C0004D" author="Nakane T." box="[985,1159,1021,1048]" pageId="15" pageNumber="126" pagination="41 - 46" refId="ref13940" refString="Nakane T. 1995. Notes on some little-known beetles (Coleoptera) in Japan. 13. Kita-Kyushu no Konchu 42: 41 - 46." type="journal article" year="1995">Nakane (1995)</bibRefCitation>
described
<taxonomicName id="4C174D06BE4DFFC2FB17BCA58188006F" authorityName="Nakane" authorityYear="1995" class="Insecta" family="Georissidae" genus="Georissus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="126" phylum="Arthropoda" rank="species" species="katsuoi" subGenus="Neogeorissus">
<emphasis id="B963EA97BE4DFFC2FB17BCA58533004D" box="[1279,1396,1022,1048]" italics="true" pageId="15" pageNumber="126">Georissus</emphasis>
(
<emphasis id="B963EA97BE4DFFC2FF2FBB7A8122006F" box="[199,357,1057,1082]" italics="true" pageId="15" pageNumber="126">Neogeorissus</emphasis>
)
<emphasis id="B963EA97BE4DFFC2FE92BB7B8188006F" box="[378,463,1056,1082]" italics="true" pageId="15" pageNumber="126">katsuoi</emphasis>
</taxonomicName>
as a new species without comparison with the related species
<taxonomicName id="4C174D06BE4DFFC2FB3BBB7A8506006F" authorityName="Sato" authorityYear="1972" box="[1235,1345,1056,1083]" class="Insecta" family="Georissidae" genus="Georissus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="126" phylum="Arthropoda" rank="species" species="sakaii">
<emphasis id="B963EA97BE4DFFC2FB3BBB7A84AF006E" box="[1235,1256,1057,1083]" italics="true" pageId="15" pageNumber="126">G</emphasis>
.
<emphasis id="B963EA97BE4DFFC2FB13BB7B8506006F" box="[1275,1345,1056,1082]" italics="true" pageId="15" pageNumber="126">sakaii</emphasis>
</taxonomicName>
and
<taxonomicName id="4C174D06BE4DFFC2FF55BB1F81130008" authorityName="Sato" authorityYear="1972" box="[189,340,1092,1118]" class="Insecta" family="Georissidae" genus="Georissus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="126" phylum="Arthropoda" rank="species" species="japonicus">
<emphasis id="B963EA97BE4DFFC2FF55BB1F8095000B" box="[189,210,1092,1118]" italics="true" pageId="15" pageNumber="126">G</emphasis>
.
<emphasis id="B963EA97BE4DFFC2FF0ABB1F81130008" box="[226,340,1092,1117]" italics="true" pageId="15" pageNumber="126">japonicus</emphasis>
</taxonomicName>
. Both species were described based on only
<typeStatus id="54AC8827BE4DFFC2FC8FBB18839B0008" box="[871,988,1091,1117]" pageId="15" pageNumber="126" type="holotype">holotypes</typeStatus>
, and there have been no additional records until now. After a detailed comparison of
<typeStatus id="54AC8827BE4DFFC2FCEDBB3D832B00D5" box="[773,876,1126,1152]" pageId="15" pageNumber="126" type="holotype">holotype</typeStatus>
and additional specimens of the above three species, we conclude that they are the same species.
</paragraph>
<paragraph id="8BA83685BE4DFFC2FF55BB9281C600B6" blockId="15.[189,385,1225,1251]" box="[189,385,1225,1251]" pageId="15" pageNumber="126">
<heading id="D0E081E9BE4DFFC2FF55BB9281C600B6" bold="true" box="[189,385,1225,1251]" fontSize="11" level="3" pageId="15" pageNumber="126" reason="3">
<emphasis id="B963EA97BE4DFFC2FF55BB9281C600B6" bold="true" box="[189,385,1225,1251]" pageId="15" pageNumber="126">Biological notes</emphasis>
</heading>
</paragraph>
<paragraph id="8BA83685BE4DFFC2FF55BBAC84C20160" blockId="15.[189,1399,1271,1333]" pageId="15" pageNumber="126">
This species was collected from coastal sandy beaches (
<collectingRegion id="49D3F867BE4DFFC2FCB1BBAC838C0144" box="[857,971,1271,1297]" country="Japan" name="Shizuoka" pageId="15" pageNumber="126">Shizuoka</collectingRegion>
,
<collectingRegion id="49D3F867BE4DFFC2FC3FBBAC84190147" box="[983,1118,1271,1298]" country="Japan" name="Tokushima" pageId="15" pageNumber="126">Tokushima</collectingRegion>
), a river bank (
<bibRefCitation id="EF864B74BE4DFFC2FAFCBBAC81020161" author="Iwata T. &amp; Iwata Y." pageId="15" pageNumber="126" pagination="18 - 19" refId="ref13373" refString="Iwata T. &amp; Iwata Y. 2012. Additional record of Georissidae from Saitama Prefecture. Sayabane, New Series 6: 18 - 19. [In Japanese.]" type="book chapter" year="2012">Iwata &amp; Iwata 2012</bibRefCitation>
, in
<collectingRegion id="49D3F867BE4DFFC2FE87BA4181890161" box="[367,462,1306,1332]" country="Japan" name="Saitama" pageId="15" pageNumber="126">Saitama</collectingRegion>
by pit fall trap), or the shores of ponds (
<collectingRegion id="49D3F867BE4DFFC2FC4EBA4184460161" box="[934,1025,1306,1332]" country="Japan" name="Niigata" pageId="15" pageNumber="126">Niigata</collectingRegion>
,
<collectingRegion id="49D3F867BE4DFFC2FBE5BA41843E0160" box="[1037,1145,1306,1333]" country="Japan" name="Fukuoka" pageId="15" pageNumber="126">Fukuoka</collectingRegion>
).
</paragraph>
</subSubSection>
<subSubSection id="C30D650EBE4DFFC2FF55BA0083730193" pageId="15" pageNumber="126" type="distribution">
<paragraph id="8BA83685BE4DFFC2FF55BA0081110120" blockId="15.[189,342,1371,1397]" box="[189,342,1371,1397]" pageId="15" pageNumber="126">
<heading id="D0E081E9BE4DFFC2FF55BA0081110120" bold="true" box="[189,342,1371,1397]" fontSize="11" level="3" pageId="15" pageNumber="126" reason="3">
<emphasis id="B963EA97BE4DFFC2FF55BA0081110120" bold="true" box="[189,342,1371,1397]" pageId="15" pageNumber="126">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8BA83685BE4DFFC2FF55BAD283730193" blockId="15.[189,1399,1417,1479]" pageId="15" pageNumber="126">
<collectingCountry id="F3007615BE4DFFC2FF55BAD281E701F6" box="[189,416,1417,1443]" name="Russia" pageId="15" pageNumber="126">Russian Federation</collectingCountry>
(Lafer 1989);
<collectingCountry id="F3007615BE4DFFC2FDBBBAD282DD01F6" box="[595,666,1417,1443]" name="Japan" pageId="15" pageNumber="126">Japan</collectingCountry>
: Honshu (
<collectingRegion id="49D3F867BE4DFFC2FCFDBAD2832901F6" box="[789,878,1417,1443]" country="Japan" name="Miyagi" pageId="15" pageNumber="126">Miyagi</collectingRegion>
,
<collectingRegion id="49D3F867BE4DFFC2FC94BAD2839501F6" box="[892,978,1417,1443]" country="Japan" name="Ibaraki" pageId="15" pageNumber="126">Ibaraki</collectingRegion>
,
<collectingRegion id="49D3F867BE4DFFC2FC08BAD2847E01F6" box="[992,1081,1417,1443]" country="Japan" name="Niigata" pageId="15" pageNumber="126">Niigata</collectingRegion>
,
<collectingRegion id="49D3F867BE4DFFC2FBAFBAD284F301F6" box="[1095,1204,1417,1443]" country="Japan" name="Shizuoka" pageId="15" pageNumber="126">Shizuoka</collectingRegion>
Prefs), Shikoku (
<collectingRegion id="49D3F867BE4DFFC2FF2FBAF781C80192" box="[199,399,1452,1479]" country="Japan" name="Tokushima" pageId="15" pageNumber="126">Tokushima Pref.</collectingRegion>
), Kyushu (
<collectingRegion id="49D3F867BE4DFFC2FDE5BAF782FF0192" box="[525,696,1452,1479]" country="Japan" name="Fukuoka" pageId="15" pageNumber="126">Fukuoka Pref.</collectingRegion>
) (
<figureCitation id="132C2A00BE4DFFC2FD27BAF783610193" box="[719,806,1452,1478]" captionStart="Fig" captionStartId="27.[189,232,1952,1978]" captionTargetBox="[206,1384,303,1881]" captionTargetId="figure-11@27.[189,1398,290,1888]" captionTargetPageId="27" captionText="Fig. 13. Distribution map of Georissus spp. Russian localities for Georissus (G.) canalifer Sharp, 1888 and G. (Ne.) japonicus Satô, 1972 are omitted." figureDoi="http://doi.org/10.5281/zenodo.6518004" httpUri="https://zenodo.org/record/6518004/files/figure.png" pageId="15" pageNumber="126">Fig. 13</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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