518 lines
75 KiB
XML
518 lines
75 KiB
XML
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<mods:title id="4FB6F5F450A48ECEA8E5340A158C8ADB">Revision of the genus Georissus (Coleoptera, Hydrophiloidea, Georissidae) of Japan</mods:title>
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<mods:namePart id="F87783443F9DCA4FC7570148984B4E39">Yasuda, Kohei</mods:namePart>
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<mods:affiliation id="DF5E16CE3C23FA87EB7AD9338DD62C5B">Entomological Laboratory, Faculty of Agriculture, Ehime University, Tarumi 3 - 5 - 7, Matsuyama, 790 - 8566 Japan.</mods:affiliation>
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<mods:nameIdentifier id="5F4058D151A0DF23422534571024CFF9" type="email">koheizari303@gmail.com</mods:nameIdentifier>
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<mods:namePart id="F6C8B8D5732412A17359AE56A6265B62">Yoshitomi, Hiroyuki</mods:namePart>
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<mods:affiliation id="A79A0904DBD5A158B1A9DED02C5A6872">Entomological Laboratory, Faculty of Agriculture, Ehime University, Tarumi 3 - 5 - 7, Matsuyama, 790 - 8566 Japan.</mods:affiliation>
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<heading id="D0E081E9BE40FFCFFDB3BD2E839E06DA" box="[603,985,628,655]" centered="true" fontSize="11" level="2" pageId="2" pageNumber="113" reason="2">
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Genus
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<taxonomicName id="4C174D06BE40FFCFFD45BD2F839E06DA" ID-CoL="8VRPM" authority="Latreille, 1809" authorityName="Latreille" authorityYear="1809" box="[685,985,628,655]" class="Insecta" family="Georissidae" genus="Georissus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="113" phylum="Arthropoda" rank="genus">
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<emphasis id="B963EA97BE40FFCFFD45BD2F836306DB" bold="true" box="[685,804,628,654]" italics="true" pageId="2" pageNumber="113">Georissus</emphasis>
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<bibRefCitation id="EF864B74BE40FFCFFCC4BD2F839E06DA" author="Latreille P. A." box="[812,985,628,655]" pageId="2" pageNumber="113" refId="ref13513" refString="Latreille P. A. 1809. Genera Crustaceorum et Insectorum, secundum ordinem naturalem in Familias disposita, iconibus exemplisque plurimus explicate, Vol. 4. A. Koenig, Paris and Strassburg [Parisiis et Argentorati]. https: // doi. org / 10.5962 / bhl. title. 65741" type="book" year="1809">Latreille, 1809</bibRefCitation>
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</taxonomicName>
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<treatmentCitation id="0AB61094BE40FFCFFF55BDED826E0685" author="Latreille P. A." box="[189,553,694,720]" page="377" pageId="2" pageNumber="113" year="1809">
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<taxonomicName id="4C174D06BE40FFCFFF55BDED826E0685" ID-CoL="8VRPM" authority="Latreille, 1809: 377" authorityName="Latreille" authorityPageNumber="377" authorityYear="1809" box="[189,553,694,720]" class="Insecta" family="Georissidae" genus="Georissus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="113" phylum="Arthropoda" rank="genus">
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<emphasis id="B963EA97BE40FFCFFF55BDED81750685" box="[189,306,694,720]" italics="true" pageId="2" pageNumber="113">Georissus</emphasis>
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<bibRefCitation id="EF864B74BE40FFCFFED3BDED826E0685" author="Latreille P. A." box="[315,553,694,720]" pageId="2" pageNumber="113" refId="ref13513" refString="Latreille P. A. 1809. Genera Crustaceorum et Insectorum, secundum ordinem naturalem in Familias disposita, iconibus exemplisque plurimus explicate, Vol. 4. A. Koenig, Paris and Strassburg [Parisiis et Argentorati]. https: // doi. org / 10.5962 / bhl. title. 65741" type="book" year="1809">Latreille, 1809: 377</bibRefCitation>
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</taxonomicName>
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.
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<typeStatus id="54AC8827BE40FFCFFDDCBDEC82290684" box="[564,622,695,721]" pageId="2" pageNumber="113">Type</typeStatus>
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species:
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<taxonomicName id="4C174D06BE40FFCFFD36BDED842D0685" authority="Fabricius, 1798" authorityName="Fabricius" authorityYear="1798" box="[734,1130,694,720]" class="Insecta" family="Tenebrionidae" genus="Pimelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="113" phylum="Arthropoda" rank="species" species="pygmaea">
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<emphasis id="B963EA97BE40FFCFFD36BDED83EC0685" box="[734,939,694,720]" italics="true" pageId="2" pageNumber="113">Pimelia pygmaea</emphasis>
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Fabricius, 1798
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</taxonomicName>
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(=
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<taxonomicName id="4C174D06BE40FFCFFB7EBDED813106A6" authority="Rossi, 1794" authorityName="Rossi" authorityYear="1794" class="Insecta" family="Byrrhidae" genus="Byrrhus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="113" phylum="Arthropoda" rank="species" species="crenulatus">
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<emphasis id="B963EA97BE40FFCFFB7EBDED85300685" box="[1174,1399,694,720]" italics="true" pageId="2" pageNumber="113">Byrrhus crenulatus</emphasis>
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Rossi, 1794
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</taxonomicName>
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) [monotypy].
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</paragraph>
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<paragraph id="8BA83685BE40FFCFFF55BDA7850A0743" blockId="2.[189,1399,694,791]" box="[189,1357,764,791]" pageId="2" pageNumber="113">
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<treatmentCitation id="0AB61094BE40FFCFFF55BDA782010743" author="Illiger J. C. W." box="[189,582,764,790]" page="297" pageId="2" pageNumber="113" year="1807">
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<taxonomicName id="4C174D06BE40FFCFFF55BDA782010743" ID-CoL="8XK9H" authority="Illiger, 1807 a: 297" authorityName="Illiger" authorityPageNumber="297" authorityYear="1807" box="[189,582,764,790]" class="Insecta" family="Georissidae" genus="Cathammistes" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="113" phylum="Arthropoda" rank="genus">
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<emphasis id="B963EA97BE40FFCFFF55BDA781240743" box="[189,355,764,790]" italics="true" pageId="2" pageNumber="113">Cathammistes</emphasis>
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<bibRefCitation id="EF864B74BE40FFCFFE83BDA782010743" author="Illiger J. C. W." box="[363,582,764,790]" pageId="2" pageNumber="113" pagination="296 - 317" refId="ref13313" refString="Illiger J. C. W. 1807 a. Nachlese zu den Bemerkungen, Berichtigungen und Zusatzen zu Fabricii Systema Eleutheratorum. Magazin fur Insektenkunde 6: 296 - 317." type="journal article" year="1807">Illiger, 1807a: 297</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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;
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<treatmentCitation id="0AB61094BE40FFCFFDBABDA782900743" author="Illiger J. C. W." box="[594,727,764,790]" page="322" pageId="2" pageNumber="113" year="1807">
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<bibRefCitation id="EF864B74BE40FFCFFDBABDA782900743" author="Illiger J. C. W." box="[594,727,764,790]" pageId="2" pageNumber="113" pagination="318 - 350" refId="ref13345" refString="Illiger J. C. W. 1807 b. Vorschlag zur Aufnahme im Fabricischen Systeme fehlender Kafergattungen. Magazin fur Insektenkunde 6: 318 - 350." type="journal article" year="1807">1807b: 322</bibRefCitation>
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</treatmentCitation>
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[suppressed, cf.
|
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<treatmentCitation id="0AB61094BE40FFCFFC75BDA784120742" author="ICZN" box="[925,1109,764,791]" page="58" pageId="2" pageNumber="113" year="1998">
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<bibRefCitation id="EF864B74BE40FFCFFC75BDA784120742" author="ICZN" box="[925,1109,764,791]" pageId="2" pageNumber="113" pagination="58 - 60" refId="ref13253" refString="ICZN 1998. Opinion 1891. Crenitis Bedel, 1881, Georissus Latreille, 1809 and Oosternum Sharp, 1882 (Insecta, Coleoptera): conserved. Bulletin of Zoological Nomenclature 55: 58 - 60. Available from https: // www. biodiversitylibrary. org / page / 12441426 [accessed 22 Mar. 2022]." type="journal article" year="1998">ICZN 1998: 58</bibRefCitation>
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</treatmentCitation>
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]. Gender: masculine.
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="8BA83685BE40FFCFFF55BC65832D070C" blockId="2.[189,874,829,857]" box="[189,874,829,857]" pageId="2" pageNumber="113">
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<treatmentCitationGroup id="AB0711ABBE40FFCFFF55BC65832D070C" box="[189,874,829,857]" pageId="2" pageNumber="113">
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<taxonomicName id="4C174D06BE40FFCFFF55BC658170070D" ID-CoL="8VXTS" box="[189,311,830,856]" class="Insecta" family="Georissidae" genus="Georyssus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="113" phylum="Arthropoda" rank="genus">
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<emphasis id="B963EA97BE40FFCFFF55BC658170070D" box="[189,311,830,856]" italics="true" pageId="2" pageNumber="113">Georyssus</emphasis>
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</taxonomicName>
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[unjustified emendation] –
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<treatmentCitation id="0AB61094BE40FFCFFD93BC658321070C" author="Stephens J. F." box="[635,870,830,857]" page="105" pageId="2" pageNumber="113" year="1828">
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<bibRefCitation id="EF864B74BE40FFCFFD93BC658321070C" author="Stephens J. F." box="[635,870,830,857]" pageId="2" pageNumber="113" refId="ref14422" refString="Stephens J. F. 1828 - 1829. Illustrations of British Entomology … Mandibulata, Vol. 2. Baldwin & Cradock, London. https: // doi. org / 10.5962 / bhl. title. 8133" type="book" year="1828">Stephens 1828: 105</bibRefCitation>
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</treatmentCitation>
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.
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection id="C30D650EBE40FFC9FF55BC2482A3059B" lastPageId="4" lastPageNumber="115" pageId="2" pageNumber="113" type="description">
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<paragraph id="8BA83685BE40FFCFFF55BC24825607CC" blockId="2.[189,529,895,921]" box="[189,529,895,921]" pageId="2" pageNumber="113">
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<heading id="D0E081E9BE40FFCFFF55BC24825607CC" bold="true" box="[189,529,895,921]" fontSize="11" level="3" pageId="2" pageNumber="113" reason="3">
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<emphasis id="B963EA97BE40FFCFFF55BC24825607CC" bold="true" box="[189,529,895,921]" pageId="2" pageNumber="113">Supplementary description</emphasis>
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</heading>
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</paragraph>
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||
<paragraph id="8BA83685BE40FFCFFF55BCF583D0079D" blockId="2.[189,919,942,968]" box="[189,919,942,968]" pageId="2" pageNumber="113">BODY. Black to reddish brown, oval, strongly convex dorsally.</paragraph>
|
||
<paragraph id="8BA83685BE40FFCFFF55BCB484F8013C" blockId="2.[187,1402,1007,1386]" pageId="2" pageNumber="113">
|
||
CLYPEUS. Uneven, with granules or carinae on its surface; articulated carinae arising at antero-lateral portion of clypeus and reaching anterior portion of frons (
|
||
<emphasis id="B963EA97BE40FFCFFD1EBB4983530079" box="[758,788,1042,1068]" italics="true" pageId="2" pageNumber="113">slr</emphasis>
|
||
in
|
||
<figureCitation id="132C2A00BE40FFCFFCD0BB4983D50078" box="[824,914,1042,1069]" captionStart="Fig" captionStartId="3.[189,232,1888,1914]" captionTargetBox="[189,1394,269,1844]" captionTargetId="figure-11@3.[189,1398,269,1844]" captionTargetPageId="3" captionText="Fig. 1. Georissus (Ne.) kurosawai Nakane, 1963, showing terminology. A. Head. B. Pronotum. Abbreviations: cd = central depression; lsa = lateral smooth area; pg = posterior grooves; sdn = subbasal dentation; sdp = subbasal depression; slb = sublateral bulge; slr = sublateral ridge; smb = submedian bulge; smlg = shallow median longitudinal groove." figureDoi="http://doi.org/10.5281/zenodo.6517950" httpUri="https://zenodo.org/record/6517950/files/figure.png" pageId="2" pageNumber="113">Fig. 1A</figureCitation>
|
||
); a pair of distinct bulges (
|
||
<emphasis id="B963EA97BE40FFCFFB2BBB4984B50079" box="[1219,1266,1042,1068]" italics="true" pageId="2" pageNumber="113">smb</emphasis>
|
||
in
|
||
<figureCitation id="132C2A00BE40FFCFFAFDBB49852A0078" box="[1301,1389,1042,1069]" captionStart="Fig" captionStartId="3.[189,232,1888,1914]" captionTargetBox="[189,1394,269,1844]" captionTargetId="figure-11@3.[189,1398,269,1844]" captionTargetPageId="3" captionText="Fig. 1. Georissus (Ne.) kurosawai Nakane, 1963, showing terminology. A. Head. B. Pronotum. Abbreviations: cd = central depression; lsa = lateral smooth area; pg = posterior grooves; sdn = subbasal dentation; sdp = subbasal depression; slb = sublateral bulge; slr = sublateral ridge; smb = submedian bulge; smlg = shallow median longitudinal groove." figureDoi="http://doi.org/10.5281/zenodo.6517950" httpUri="https://zenodo.org/record/6517950/files/figure.png" pageId="2" pageNumber="113">Fig. 1A</figureCitation>
|
||
) on antero-median portion of frons (in
|
||
<taxonomicName id="4C174D06BE40FFCFFD81BB6D837F001A" box="[617,824,1078,1103]" class="Insecta" family="Georissidae" genus="Georyssus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="121" phylum="Arthropoda" rank="subGenus" subGenus="Nipponogeorissus">
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||
<emphasis id="B963EA97BE40FFCFFD81BB6D837F001A" box="[617,824,1078,1103]" italics="true" pageId="2" pageNumber="113">Nipponogeorissus</emphasis>
|
||
</taxonomicName>
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||
and
|
||
<taxonomicName id="4C174D06BE40FFCFFC86BB6D844E001A" box="[878,1033,1078,1103]" class="Insecta" family="Georissidae" genus="Georyssus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="136" phylum="Arthropoda" rank="subGenus" subGenus="Neogeorissus">
|
||
<emphasis id="B963EA97BE40FFCFFC86BB6D844E001A" box="[878,1033,1078,1103]" italics="true" pageId="2" pageNumber="113">Neogeorissus</emphasis>
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||
</taxonomicName>
|
||
); sides of frons smooth, lacking granules (
|
||
<emphasis id="B963EA97BE40FFCFFED9BB0281140026" box="[305,339,1113,1139]" italics="true" pageId="2" pageNumber="113">lsa</emphasis>
|
||
in
|
||
<figureCitation id="132C2A00BE40FFCFFE95BB02819D0026" box="[381,474,1113,1139]" captionStart="Fig" captionStartId="3.[189,232,1888,1914]" captionTargetBox="[189,1394,269,1844]" captionTargetId="figure-11@3.[189,1398,269,1844]" captionTargetPageId="3" captionText="Fig. 1. Georissus (Ne.) kurosawai Nakane, 1963, showing terminology. A. Head. B. Pronotum. Abbreviations: cd = central depression; lsa = lateral smooth area; pg = posterior grooves; sdn = subbasal dentation; sdp = subbasal depression; slb = sublateral bulge; slr = sublateral ridge; smb = submedian bulge; smlg = shallow median longitudinal groove." figureDoi="http://doi.org/10.5281/zenodo.6517950" httpUri="https://zenodo.org/record/6517950/files/figure.png" pageId="2" pageNumber="113">Fig. 1A</figureCitation>
|
||
). Eyes developed, not protuberant. Antennae (
|
||
<figureCitation id="132C2A00BE40FFCFFC14BB0284C70026" box="[1020,1152,1113,1139]" captionStart="Fig" captionStartId="14.[189,232,1876,1902]" captionTargetBox="[196,1386,277,1817]" captionTargetId="figure-18@14.[189,1398,264,1820]" captionTargetPageId="14" captionText="Fig. 4. Mouth parts and antennae of Georissus spp. A–C. Left maxillae. D–F. Labrum. G–I. Labial palpi. J–L. Left mandibles. M–O. Left antennae. A, D, G, J, M. G. (G.) canalifer Sharp, 1888. B, E, H, K, N. G. (Ni.) granulosus Satô, 1972. C, F, I, L, O. G. (Ne.) japonicus Satô, 1972. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6517962" httpUri="https://zenodo.org/record/6517962/files/figure.png" pageId="2" pageNumber="113">Fig. 4M–O</figureCitation>
|
||
) short, 9-segmented, with 3-segmented pubescent club (7-segmented with one-segmented club in Australian species:
|
||
<bibRefCitation id="EF864B74BE40FFCFFAF3BB2780BC00EC" author="Fikacek M." pageId="2" pageNumber="113" pagination="251 - 257" refId="ref12979" refString="Fikacek M. 2019. 17. Georissidae Castelnau, 1840. In: Slipinski A. & Lawrence J. F. (eds) Australian Beetles Vol. 2 Archostemata, Myxophaga, Adephaga, Polyphaga (part): 251 - 257. CSIRO Publishing." type="book chapter" year="2019">Fikáček 2019</bibRefCitation>
|
||
). Mandibles (
|
||
<figureCitation id="132C2A00BE40FFCFFE74BBC4824800EC" box="[412,527,1183,1209]" captionStart="Fig" captionStartId="14.[189,232,1876,1902]" captionTargetBox="[196,1386,277,1817]" captionTargetId="figure-18@14.[189,1398,264,1820]" captionTargetPageId="14" captionText="Fig. 4. Mouth parts and antennae of Georissus spp. A–C. Left maxillae. D–F. Labrum. G–I. Labial palpi. J–L. Left mandibles. M–O. Left antennae. A, D, G, J, M. G. (G.) canalifer Sharp, 1888. B, E, H, K, N. G. (Ni.) granulosus Satô, 1972. C, F, I, L, O. G. (Ne.) japonicus Satô, 1972. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6517962" httpUri="https://zenodo.org/record/6517962/files/figure.png" pageId="2" pageNumber="113">Fig. 4J–L</figureCitation>
|
||
) large and well sclerotized, partly concealed beneath clypeus and labrum; antero-lateral margin forming obtuse angle; apex pointed; internal margin with microtrichia except apex. Maxillary palpi (
|
||
<figureCitation id="132C2A00BE40FFCFFE95BBBD81BE0155" box="[381,505,1254,1280]" captionStart="Fig" captionStartId="14.[189,232,1876,1902]" captionTargetBox="[196,1386,277,1817]" captionTargetId="figure-18@14.[189,1398,264,1820]" captionTargetPageId="14" captionText="Fig. 4. Mouth parts and antennae of Georissus spp. A–C. Left maxillae. D–F. Labrum. G–I. Labial palpi. J–L. Left mandibles. M–O. Left antennae. A, D, G, J, M. G. (G.) canalifer Sharp, 1888. B, E, H, K, N. G. (Ni.) granulosus Satô, 1972. C, F, I, L, O. G. (Ne.) japonicus Satô, 1972. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6517962" httpUri="https://zenodo.org/record/6517962/files/figure.png" pageId="2" pageNumber="113">Fig. 4A–C</figureCitation>
|
||
) 4-segmented; palpomere 4 longest and nearly same length as palpomeres 1–3 combined. Labrum (
|
||
<figureCitation id="132C2A00BE40FFCFFE4EBA5282670176" box="[422,544,1289,1315]" captionStart="Fig" captionStartId="14.[189,232,1876,1902]" captionTargetBox="[196,1386,277,1817]" captionTargetId="figure-18@14.[189,1398,264,1820]" captionTargetPageId="14" captionText="Fig. 4. Mouth parts and antennae of Georissus spp. A–C. Left maxillae. D–F. Labrum. G–I. Labial palpi. J–L. Left mandibles. M–O. Left antennae. A, D, G, J, M. G. (G.) canalifer Sharp, 1888. B, E, H, K, N. G. (Ni.) granulosus Satô, 1972. C, F, I, L, O. G. (Ne.) japonicus Satô, 1972. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6517962" httpUri="https://zenodo.org/record/6517962/files/figure.png" pageId="2" pageNumber="113">Fig. 4D–F</figureCitation>
|
||
) semicircular, widest at base; lateral margins fringed with spines. Mentum subtriangular, strongly narrowed anteriorly, with long setae on disc. Labial palpi (
|
||
<figureCitation id="132C2A00BE40FFCFFB93BA7784A80113" box="[1147,1263,1324,1350]" captionStart="Fig" captionStartId="14.[189,232,1876,1902]" captionTargetBox="[196,1386,277,1817]" captionTargetId="figure-18@14.[189,1398,264,1820]" captionTargetPageId="14" captionText="Fig. 4. Mouth parts and antennae of Georissus spp. A–C. Left maxillae. D–F. Labrum. G–I. Labial palpi. J–L. Left mandibles. M–O. Left antennae. A, D, G, J, M. G. (G.) canalifer Sharp, 1888. B, E, H, K, N. G. (Ni.) granulosus Satô, 1972. C, F, I, L, O. G. (Ne.) japonicus Satô, 1972. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6517962" httpUri="https://zenodo.org/record/6517962/files/figure.png" pageId="2" pageNumber="113">Fig. 4G–I</figureCitation>
|
||
) as long as mentum; apical palpomere oval to oblong oval, densely bearing spines in interior margin.
|
||
</paragraph>
|
||
<paragraph id="8BA83685BE40FFCFFF55BACA81C7022B" blockId="2.[188,1399,1425,1662]" pageId="2" pageNumber="113">
|
||
PRONOTUM. Half oval to trapezoidal, divided into anterior and posterior portions; anterior portion irregularly arranged with granules or short grooves and pits, with shallow median longitudinal groove (
|
||
<emphasis id="B963EA97BE40FFCFFF2FBA8C80B901A4" box="[199,254,1495,1521]" italics="true" pageId="2" pageNumber="113">smlg</emphasis>
|
||
in
|
||
<figureCitation id="132C2A00BE40FFCFFEC3BA8C81CE01A4" box="[299,393,1495,1521]" captionStart="Fig" captionStartId="3.[189,232,1888,1914]" captionTargetBox="[189,1394,269,1844]" captionTargetId="figure-11@3.[189,1398,269,1844]" captionTargetPageId="3" captionText="Fig. 1. Georissus (Ne.) kurosawai Nakane, 1963, showing terminology. A. Head. B. Pronotum. Abbreviations: cd = central depression; lsa = lateral smooth area; pg = posterior grooves; sdn = subbasal dentation; sdp = subbasal depression; slb = sublateral bulge; slr = sublateral ridge; smb = submedian bulge; smlg = shallow median longitudinal groove." figureDoi="http://doi.org/10.5281/zenodo.6517950" httpUri="https://zenodo.org/record/6517950/files/figure.png" pageId="2" pageNumber="113">Fig. 1B</figureCitation>
|
||
); posterior portion more convex than anterior portion, with large fovea on anteromedian portion (
|
||
<emphasis id="B963EA97BE40FFCFFE68BAA181DB0241" box="[384,412,1530,1556]" italics="true" pageId="2" pageNumber="113">cd</emphasis>
|
||
in
|
||
<figureCitation id="132C2A00BE40FFCFFE28BAA1825E0241" box="[448,537,1530,1556]" captionStart="Fig" captionStartId="3.[189,232,1888,1914]" captionTargetBox="[189,1394,269,1844]" captionTargetId="figure-11@3.[189,1398,269,1844]" captionTargetPageId="3" captionText="Fig. 1. Georissus (Ne.) kurosawai Nakane, 1963, showing terminology. A. Head. B. Pronotum. Abbreviations: cd = central depression; lsa = lateral smooth area; pg = posterior grooves; sdn = subbasal dentation; sdp = subbasal depression; slb = sublateral bulge; slr = sublateral ridge; smb = submedian bulge; smlg = shallow median longitudinal groove." figureDoi="http://doi.org/10.5281/zenodo.6517950" httpUri="https://zenodo.org/record/6517950/files/figure.png" pageId="2" pageNumber="113">Fig. 1B</figureCitation>
|
||
); posterior sides of
|
||
<emphasis id="B963EA97BE40FFCFFCE8BAA1835B0241" box="[768,796,1530,1556]" italics="true" pageId="2" pageNumber="113">cd</emphasis>
|
||
with a pair of distinct foveae (
|
||
<emphasis id="B963EA97BE40FFCFFB6BBAA184EB0241" box="[1155,1196,1530,1556]" italics="true" pageId="2" pageNumber="113">sdp</emphasis>
|
||
in
|
||
<figureCitation id="132C2A00BE40FFCFFB38BAA1856F0241" box="[1232,1320,1530,1556]" captionStart="Fig" captionStartId="3.[189,232,1888,1914]" captionTargetBox="[189,1394,269,1844]" captionTargetId="figure-11@3.[189,1398,269,1844]" captionTargetPageId="3" captionText="Fig. 1. Georissus (Ne.) kurosawai Nakane, 1963, showing terminology. A. Head. B. Pronotum. Abbreviations: cd = central depression; lsa = lateral smooth area; pg = posterior grooves; sdn = subbasal dentation; sdp = subbasal depression; slb = sublateral bulge; slr = sublateral ridge; smb = submedian bulge; smlg = shallow median longitudinal groove." figureDoi="http://doi.org/10.5281/zenodo.6517950" httpUri="https://zenodo.org/record/6517950/files/figure.png" pageId="2" pageNumber="113">Fig. 1B</figureCitation>
|
||
) (only in
|
||
<taxonomicName id="4C174D06BE40FFCFFF32B945813F0262" box="[218,376,1566,1591]" class="Insecta" family="Georissidae" genus="Georyssus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="136" phylum="Arthropoda" rank="subGenus" subGenus="Neogeorissus">
|
||
<emphasis id="B963EA97BE40FFCFFF32B945813F0262" box="[218,376,1566,1591]" italics="true" pageId="2" pageNumber="113">Neogeorissus</emphasis>
|
||
</taxonomicName>
|
||
); sublateral potion of posterior portion with distinct bulge (
|
||
<emphasis id="B963EA97BE40FFCFFBC3B946840A0262" box="[1067,1101,1565,1591]" italics="true" pageId="2" pageNumber="113">slb</emphasis>
|
||
in
|
||
<figureCitation id="132C2A00BE40FFCFFB98B946848E026D" box="[1136,1225,1565,1592]" captionStart="Fig" captionStartId="3.[189,232,1888,1914]" captionTargetBox="[189,1394,269,1844]" captionTargetId="figure-11@3.[189,1398,269,1844]" captionTargetPageId="3" captionText="Fig. 1. Georissus (Ne.) kurosawai Nakane, 1963, showing terminology. A. Head. B. Pronotum. Abbreviations: cd = central depression; lsa = lateral smooth area; pg = posterior grooves; sdn = subbasal dentation; sdp = subbasal depression; slb = sublateral bulge; slr = sublateral ridge; smb = submedian bulge; smlg = shallow median longitudinal groove." figureDoi="http://doi.org/10.5281/zenodo.6517950" httpUri="https://zenodo.org/record/6517950/files/figure.png" pageId="2" pageNumber="113">Fig. 1B</figureCitation>
|
||
); behind of
|
||
<emphasis id="B963EA97BE40FFCFFABBB94685320262" box="[1363,1397,1565,1591]" italics="true" pageId="2" pageNumber="113">slb</emphasis>
|
||
with deep groove (
|
||
<emphasis id="B963EA97BE40FFCFFE7FB91A81F3020F" box="[407,436,1601,1626]" italics="true" pageId="2" pageNumber="113">pg</emphasis>
|
||
in
|
||
<figureCitation id="132C2A00BE40FFCFFE30B91A8276020E" box="[472,561,1601,1627]" captionStart="Fig" captionStartId="3.[189,232,1888,1914]" captionTargetBox="[189,1394,269,1844]" captionTargetId="figure-11@3.[189,1398,269,1844]" captionTargetPageId="3" captionText="Fig. 1. Georissus (Ne.) kurosawai Nakane, 1963, showing terminology. A. Head. B. Pronotum. Abbreviations: cd = central depression; lsa = lateral smooth area; pg = posterior grooves; sdn = subbasal dentation; sdp = subbasal depression; slb = sublateral bulge; slr = sublateral ridge; smb = submedian bulge; smlg = shallow median longitudinal groove." figureDoi="http://doi.org/10.5281/zenodo.6517950" httpUri="https://zenodo.org/record/6517950/files/figure.png" pageId="2" pageNumber="113">Fig. 1B</figureCitation>
|
||
); sides somewhat protruding laterally, having dentation (
|
||
<emphasis id="B963EA97BE40FFCFFB21B91A84B5020E" box="[1225,1266,1601,1627]" italics="true" pageId="2" pageNumber="113">sdn</emphasis>
|
||
in
|
||
<figureCitation id="132C2A00BE40FFCFFAFDB91A852A020E" box="[1301,1389,1601,1627]" captionStart="Fig" captionStartId="3.[189,232,1888,1914]" captionTargetBox="[189,1394,269,1844]" captionTargetId="figure-11@3.[189,1398,269,1844]" captionTargetPageId="3" captionText="Fig. 1. Georissus (Ne.) kurosawai Nakane, 1963, showing terminology. A. Head. B. Pronotum. Abbreviations: cd = central depression; lsa = lateral smooth area; pg = posterior grooves; sdn = subbasal dentation; sdp = subbasal depression; slb = sublateral bulge; slr = sublateral ridge; smb = submedian bulge; smlg = shallow median longitudinal groove." figureDoi="http://doi.org/10.5281/zenodo.6517950" httpUri="https://zenodo.org/record/6517950/files/figure.png" pageId="2" pageNumber="113">Fig. 1B</figureCitation>
|
||
) in
|
||
<taxonomicName id="4C174D06BE40FFCFFF33B93E813E022B" box="[219,377,1637,1662]" class="Insecta" family="Georissidae" genus="Georyssus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="136" phylum="Arthropoda" rank="subGenus" subGenus="Neogeorissus">
|
||
<emphasis id="B963EA97BE40FFCFFF33B93E813E022B" box="[219,377,1637,1662]" italics="true" pageId="2" pageNumber="113">Neogeorissus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BA83685BE40FFCFFF55B9FD81960353" blockId="2.[189,1400,1701,1798]" pageId="2" pageNumber="113">
|
||
LEGS. Protrochantins concealed. Procoxae large, plate-like, concealing prosternum. Femora somewhat flattened. Tibiae slender, especially protibiae rather flattened. Tarsal formula 5-5-5 (5-4-
|
||
<quantity id="4CEF9B60BE40FFCFFB21B99384BF02B7" box="[1225,1272,1736,1763]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="2" pageNumber="113" unit="in" value="4.0">4 in</quantity>
|
||
Australian species:
|
||
<bibRefCitation id="EF864B74BE40FFCFFEC9B9B781820353" author="Fikacek M." box="[289,453,1772,1798]" pageId="2" pageNumber="113" pagination="251 - 257" refId="ref12979" refString="Fikacek M. 2019. 17. Georissidae Castelnau, 1840. In: Slipinski A. & Lawrence J. F. (eds) Australian Beetles Vol. 2 Archostemata, Myxophaga, Adephaga, Polyphaga (part): 251 - 257. CSIRO Publishing." type="book chapter" year="2019">Fikáček 2019</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BA83685BE40FFCFFF55B87582EF033F" blockId="2.[189,1398,1837,1899]" pageId="2" pageNumber="113">ELYTRA. Provided with ten punctate rows. All or alternate intervals slightly to highly raised. Hind wings well developed or absent in some species.</paragraph>
|
||
<paragraph id="8BA83685BE40FFCFFF55B8C984E2039A" blockId="2.[189,1398,1938,2000]" pageId="2" pageNumber="113">
|
||
VENTRITES. Ventrites 1 and 2 connate; ventrite 1 large, with distinct granules; posterior margin of ventrite 1 and anterior margin of ventrite 2 with a pair of ventral teeth (
|
||
<emphasis id="B963EA97BE40FFCFFBE8B8ED8452039A" box="[1024,1045,1974,1999]" italics="true" pageId="2" pageNumber="113">vt</emphasis>
|
||
in
|
||
<figureCitation id="132C2A00BE40FFCFFBD3B8EE84DF039A" box="[1083,1176,1973,1999]" captionStart="Fig" captionStartId="21.[191,234,1890,1916]" captionTargetBox="[193,1394,569,1846]" captionTargetId="figure-107@21.[189,1398,569,1852]" captionTargetPageId="21" captionText="Fig. 8. Ventrites of Georissus spp. A. G. (G.) canalifer Sharp, 1888. B. G. (G.) babai Satô, 1970. C. G. (Ni.) granulosus Satô, 1972. D. G. (Ne.) japonicus Satô, 1972. E–F. G. (Ne.) kurosawai Nakane, 1963 from Hokkaido (E) and Nagano (F). G–H. G. (Ne.) takahashii sp. nov. from Kochi (G) and Okinawa (F). I. G. (Ne.) satoi sp. nov. Abbreviation: vt = ventral teeth. Scale bars = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.6517982" httpUri="https://zenodo.org/record/6517982/files/figure.png" pageId="2" pageNumber="113">Fig. 8A</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="DF68660DBE41FFCEFF55B83B835403B1" ID-DOI="http://doi.org/10.5281/zenodo.6517950" ID-Zenodo-Dep="6517950" httpUri="https://zenodo.org/record/6517950/files/figure.png" pageId="3" pageNumber="114" startId="3.[189,232,1888,1914]" targetBox="[189,1394,269,1844]" targetPageId="3">
|
||
<paragraph id="8BA83685BE41FFCEFF55B83B835403B1" blockId="3.[188,1399,1888,2020]" pageId="3" pageNumber="114">
|
||
<emphasis id="B963EA97BE41FFCEFF55B83B8154032F" bold="true" box="[189,275,1888,1914]" pageId="3" pageNumber="114">Fig. 1.</emphasis>
|
||
<taxonomicName id="4C174D06BE41FFCEFECAB83A8360032F" authority="Nakane, 1963" authorityName="Nakane" authorityYear="1963" box="[290,807,1888,1915]" class="Insecta" family="Georissidae" genus="Georissus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="114" phylum="Arthropoda" rank="species" species="kurosawai" subGenus="Ne.">
|
||
<emphasis id="B963EA97BE41FFCEFECAB83A822D032F" box="[290,618,1888,1915]" italics="true" pageId="3" pageNumber="114">Georissus (Ne.) kurosawai</emphasis>
|
||
<bibRefCitation id="EF864B74BE41FFCEFD92B83B8360032F" author="Nakane T." box="[634,807,1888,1914]" pageId="3" pageNumber="114" pagination="2" refId="ref13863" refString="Nakane T. 1963. Hydrophilidae. In: Nakane T., Ohbayashi K., Nomura S. & Kurosawa Y. (eds) Iconographia Insetorum Japonicorum Colore naturali edita Vol. 2 (Coleoptera). Hokuryukan, Tokyo. [In Japanese.]" type="book chapter" year="1963">Nakane, 1963</bibRefCitation>
|
||
</taxonomicName>
|
||
, showing terminology.
|
||
<emphasis id="B963EA97BE41FFCEFBB9B83B8421032F" bold="true" box="[1105,1126,1888,1914]" pageId="3" pageNumber="114">A</emphasis>
|
||
. Head.
|
||
<emphasis id="B963EA97BE41FFCEFB39B83B84A2032F" bold="true" box="[1233,1253,1888,1914]" pageId="3" pageNumber="114">B</emphasis>
|
||
. Pronotum. Abbreviations:
|
||
<emphasis id="B963EA97BE41FFCEFE9AB8D881C903C8" box="[370,398,1923,1949]" italics="true" pageId="3" pageNumber="114">cd</emphasis>
|
||
= central depression;
|
||
<emphasis id="B963EA97BE41FFCEFD63B8D882EA03C8" box="[651,685,1923,1949]" italics="true" pageId="3" pageNumber="114">lsa</emphasis>
|
||
= lateral smooth area;
|
||
<emphasis id="B963EA97BE41FFCEFC5EB8DF839403C8" box="[950,979,1924,1949]" italics="true" pageId="3" pageNumber="114">pg</emphasis>
|
||
= posterior grooves;
|
||
<emphasis id="B963EA97BE41FFCEFB22B8D884B403C8" box="[1226,1267,1923,1949]" italics="true" pageId="3" pageNumber="114">sdn</emphasis>
|
||
= subbasal dentation;
|
||
<emphasis id="B963EA97BE41FFCEFED4B8FC81220394" box="[316,357,1959,1985]" italics="true" pageId="3" pageNumber="114">sdp</emphasis>
|
||
= subbasal depression;
|
||
<emphasis id="B963EA97BE41FFCEFD6FB8FC82EE0394" box="[647,681,1959,1985]" italics="true" pageId="3" pageNumber="114">slb</emphasis>
|
||
= sublateral bulge;
|
||
<emphasis id="B963EA97BE41FFCEFC73B8FC83FD0394" box="[923,954,1959,1985]" italics="true" pageId="3" pageNumber="114">slr</emphasis>
|
||
= sublateral ridge;
|
||
<emphasis id="B963EA97BE41FFCEFB4FB8FC84910394" box="[1191,1238,1959,1985]" italics="true" pageId="3" pageNumber="114">smb</emphasis>
|
||
= submedian bulge;
|
||
<emphasis id="B963EA97BE41FFCEFEE6B891810203B1" box="[270,325,1994,2020]" italics="true" pageId="3" pageNumber="114">smlg</emphasis>
|
||
= shallow median longitudinal groove.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BA83685BE46FFC9FF55BE5283B10576" blockId="4.[189,1014,265,291]" box="[189,1014,265,291]" pageId="4" pageNumber="115">
|
||
MALE GENITALIA (
|
||
<figureCitation id="132C2A00BE46FFC9FE6FBE5281890576" box="[391,462,265,291]" captionStart="Fig" captionStartId="23.[189,232,1763,1789]" captionTargetBox="[239,1353,284,1696]" captionTargetId="figure-12@23.[189,1398,265,1704]" captionTargetPageId="23" captionText="Fig. 9. Male aedeagus of Georissus spp. A. G. (G.) canalifer Sharp, 1888. B. G. (G.) babai Satô, 1970. C. G. (Ni.) granulosus Satô, 1972. D. G. (Ne.) japonicus Satô, 1972. E. G. (Ne.) kurosawai Nakane, 1963. F–G. G. (Ne.) takahashii sp. nov. from Kochi (F) and Okinawa (G). H. G. (Ne.) satoi sp. nov. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6517986" httpUri="https://zenodo.org/record/6517986/files/figure.png" pageId="4" pageNumber="115">Fig. 9</figureCitation>
|
||
). Aedeagus trilobate
|
||
<typeStatus id="54AC8827BE46FFC9FD2FBE5282BB0576" box="[711,764,265,291]" pageId="4" pageNumber="115">type</typeStatus>
|
||
, almost symmetrical.
|
||
</paragraph>
|
||
<paragraph id="8BA83685BE46FFC9FF55BE1082A3059B" blockId="4.[188,1399,330,463]" pageId="4" pageNumber="115">
|
||
FEMALE GENITALIA. Gonocoxite (
|
||
<figureCitation id="132C2A00BE46FFC9FDD4BE1182D30530" box="[572,660,330,357]" captionStart="Fig" captionStartId="24.[189,232,1780,1806]" captionTargetBox="[233,1337,288,1718]" captionTargetId="figure-18@24.[189,1398,264,1722]" captionTargetPageId="24" captionText="Fig. 10. Female ovipositor of Georissus spp. A. G. (G.) canalifer Sharp, 1888. B. G. (G.) babai Satô, 1970. C. G. (Ne.) japonicus Satô, 1972. D. G. (Ne.) kurosawai Nakane, 1963. E. G. (Ne.) takahashii sp. nov. F. G. (Ne.) satoi sp. nov. Abbreviations: gsty = gonostylus; ltg8 = latero-tergite 8. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6517988" httpUri="https://zenodo.org/record/6517988/files/figure.png" pageId="4" pageNumber="115">Fig. 10</figureCitation>
|
||
) long, almost cross each other posteriorly. Gonostylus short, provided with two long setae at apex. Median sclerite unilobed, densely bearing long setae. Spermatheca (
|
||
<figureCitation id="132C2A00BE46FFC9FF2CBECA810805FE" box="[196,335,401,427]" captionStart="Fig" captionStartId="25.[189,232,1920,1946]" captionTargetBox="[213,1341,1093,1867]" captionTargetId="figure-278@25.[189,1398,1073,1882]" captionTargetPageId="25" captionText="Fig. 11. Female genital organs of subgenus Neogeorissus spp. A–D. Bursal sclerite. E–H. Spermatheca. A, E. G. (Ne.) japonicus Satô, 1972. B, F. G. (Ne.) kurosawai Nakane, 1963. C, G. G. (Ne.) takahashii sp. nov. D, H. G. (Ne.) satoi sp. nov. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6517990" httpUri="https://zenodo.org/record/6517990/files/figure.png" pageId="4" pageNumber="115">Fig. 11E–H</figureCitation>
|
||
) slightly sclerotized, elongated oval. Bursal sclerite (
|
||
<figureCitation id="132C2A00BE46FFC9FC29BECA840805FE" box="[961,1103,401,427]" captionStart="Fig" captionStartId="25.[189,232,1920,1946]" captionTargetBox="[213,1341,1093,1867]" captionTargetId="figure-278@25.[189,1398,1073,1882]" captionTargetPageId="25" captionText="Fig. 11. Female genital organs of subgenus Neogeorissus spp. A–D. Bursal sclerite. E–H. Spermatheca. A, E. G. (Ne.) japonicus Satô, 1972. B, F. G. (Ne.) kurosawai Nakane, 1963. C, G. G. (Ne.) takahashii sp. nov. D, H. G. (Ne.) satoi sp. nov. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6517990" httpUri="https://zenodo.org/record/6517990/files/figure.png" pageId="4" pageNumber="115">Fig. 11A–D</figureCitation>
|
||
) somewhat developed in subgenus
|
||
<taxonomicName id="4C174D06BE46FFC9FED9BEEE8188059B" authorityName="Sato" authorityYear="1972" box="[305,463,437,462]" class="Insecta" family="Georissidae" genus="Georissus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="115" phylum="Arthropoda" rank="subGenus" subGenus="Neogeorissus">
|
||
<emphasis id="B963EA97BE46FFC9FED9BEEE8188059B" box="[305,463,437,462]" italics="true" pageId="4" pageNumber="115">Neogeorissus</emphasis>
|
||
</taxonomicName>
|
||
, slightly asymmetrical.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30D650EBE46FFC9FF55BEAD83170645" box="[189,848,501,528]" pageId="4" pageNumber="115" type="multiple">
|
||
<paragraph id="8BA83685BE46FFC9FF55BEAD83170645" blockId="4.[189,848,501,528]" box="[189,848,501,528]" pageId="4" pageNumber="115">
|
||
<heading id="D0E081E9BE46FFC9FF55BEAD83170645" bold="true" box="[189,848,501,528]" fontSize="11" level="3" pageId="4" pageNumber="115" reason="3">
|
||
<emphasis id="B963EA97BE46FFC9FF55BEAD83170645" bold="true" box="[189,848,501,528]" pageId="4" pageNumber="115">
|
||
Key to the species of the genus
|
||
<taxonomicName id="4C174D06BE46FFC9FDA8BEAE82F0065A" authorityName="Latreille" authorityYear="1809" box="[576,695,501,527]" class="Insecta" family="Georissidae" genus="Georissus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="115" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B963EA97BE46FFC9FDA8BEAE82F0065A" bold="true" box="[576,695,501,527]" italics="true" pageId="4" pageNumber="115">Georissus</emphasis>
|
||
</taxonomicName>
|
||
from
|
||
<collectingCountry id="F3007615BE46FFC9FCEBBEAD83170645" box="[771,848,502,528]" name="Japan" pageId="4" pageNumber="115">Japan</collectingCountry>
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30D650EBE46FFC9FF55BD7F85300213" pageId="4" pageNumber="115" type="key">
|
||
<key id="FF8661B4BE46FFC9FF55BD7F85300213" pageId="4" pageNumber="115">
|
||
<keyStep id="30E32E20BE46FFC9FF55BD7F8530069E" pageId="4" pageNumber="115">
|
||
<paragraph id="8BA83685BE46FFC9FF55BD7F853006D1" blockId="4.[189,1401,548,1606]" pageId="4" pageNumber="115">
|
||
<keyLead id="30E695B0BE46FFC9FF55BD7F853006D1" pageId="4" pageNumber="115">
|
||
1. Posterior portion of pronotum with a shallow median longitudinal groove, lacking granules or
|
||
<emphasis id="B963EA97BE46FFC9FAB5BD7F853E066B" box="[1373,1401,548,574]" italics="true" pageId="4" pageNumber="115">cd</emphasis>
|
||
and
|
||
<emphasis id="B963EA97BE46FFC9FEF4BD1C81020634" box="[284,325,583,609]" italics="true" pageId="4" pageNumber="115">sdp</emphasis>
|
||
. Elytra bearing seriate large punctures. Elytral interstices smooth. Aedeagus broad, median lobe short (
|
||
<figureCitation id="132C2A00BE46FFC9FE87BD3181B606D1" box="[367,497,618,644]" captionStart="Fig" captionStartId="23.[189,232,1763,1789]" captionTargetBox="[239,1353,284,1696]" captionTargetId="figure-12@23.[189,1398,265,1704]" captionTargetPageId="23" captionText="Fig. 9. Male aedeagus of Georissus spp. A. G. (G.) canalifer Sharp, 1888. B. G. (G.) babai Satô, 1970. C. G. (Ni.) granulosus Satô, 1972. D. G. (Ne.) japonicus Satô, 1972. E. G. (Ne.) kurosawai Nakane, 1963. F–G. G. (Ne.) takahashii sp. nov. from Kochi (F) and Okinawa (G). H. G. (Ne.) satoi sp. nov. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6517986" httpUri="https://zenodo.org/record/6517986/files/figure.png" pageId="4" pageNumber="115">Fig. 9A–B</figureCitation>
|
||
). Subgenus
|
||
<taxonomicName id="4C174D06BE46FFC9FD96BD3083E006D1" ID-CoL="8VRPM" authority="Latreille, 1809" authorityName="Latreille" authorityYear="1809" box="[638,935,618,645]" class="Insecta" family="Georissidae" genus="Georissus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="115" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B963EA97BE46FFC9FD96BD3082B406D0" box="[638,755,619,645]" italics="true" pageId="4" pageNumber="115">Georissus</emphasis>
|
||
<bibRefCitation id="EF864B74BE46FFC9FD12BD3183E006D1" author="Latreille P. A." box="[762,935,618,644]" pageId="4" pageNumber="115" refId="ref13513" refString="Latreille P. A. 1809. Genera Crustaceorum et Insectorum, secundum ordinem naturalem in Familias disposita, iconibus exemplisque plurimus explicate, Vol. 4. A. Koenig, Paris and Strassburg [Parisiis et Argentorati]. https: // doi. org / 10.5962 / bhl. title. 65741" type="book" year="1809">Latreille, 1809</bibRefCitation>
|
||
</taxonomicName>
|
||
............................................................ 2
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BA83685BE46FFC9FF29BDD58530069E" blockId="4.[189,1401,548,1606]" pageId="4" pageNumber="115">
|
||
<keyLead id="30E695B0BE46FFC9FF29BDD58530069E" pageId="4" pageNumber="115">
|
||
– Posterior portion of pronotum provided with granules or foveae. Elytra bearing distinct granules. Elytral interstices raised. Aedeagus narrow, median lobe short or long (
|
||
<figureCitation id="132C2A00BE46FFC9FBC6BDEA84EB069E" box="[1070,1196,689,715]" captionStart="Fig" captionStartId="23.[189,232,1763,1789]" captionTargetBox="[239,1353,284,1696]" captionTargetId="figure-12@23.[189,1398,265,1704]" captionTargetPageId="23" captionText="Fig. 9. Male aedeagus of Georissus spp. A. G. (G.) canalifer Sharp, 1888. B. G. (G.) babai Satô, 1970. C. G. (Ni.) granulosus Satô, 1972. D. G. (Ne.) japonicus Satô, 1972. E. G. (Ne.) kurosawai Nakane, 1963. F–G. G. (Ne.) takahashii sp. nov. from Kochi (F) and Okinawa (G). H. G. (Ne.) satoi sp. nov. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6517986" httpUri="https://zenodo.org/record/6517986/files/figure.png" pageId="4" pageNumber="115">Fig. 9C–H</figureCitation>
|
||
) ...................... 3
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30E32E20BE46FFC9FF55BDA9853007E9" pageId="4" pageNumber="115">
|
||
<paragraph id="8BA83685BE46FFC9FF55BDA985300706" blockId="4.[189,1401,548,1606]" pageId="4" pageNumber="115">
|
||
<keyLead id="30E695B0BE46FFC9FF55BDA985300706" pageId="4" pageNumber="115">
|
||
2. Granules of clypeus distinct, large and dense (
|
||
<figureCitation id="132C2A00BE46FFC9FCF5BDA9833A0758" box="[797,893,754,781]" captionStart="Fig" captionStartId="18.[189,232,1882,1908]" captionTargetBox="[191,1394,563,1839]" captionTargetId="figure-90@18.[189,1398,562,1842]" captionTargetPageId="18" captionText="Fig. 6. Head of Georissus spp. A. G. (G.) canalifer Sharp, 1888. B. G. (G.) babai Satô, 1970. C. G. (Ni.) granulosus Satô, 1972. D. G. (Ne.) japonicus Satô, 1972. E–F. G. (Ne.) kurosawai Nakane, 1963 from Hokkaido (E) and Nagano (F). G–H. G. (Ne.) takahashii sp. nov. from Kochi (G) and Okinawa (H). I. G. (Ne.) satoi sp. nov." figureDoi="http://doi.org/10.5281/zenodo.6517968" httpUri="https://zenodo.org/record/6517968/files/figure.png" pageId="4" pageNumber="115">Fig. 6B</figureCitation>
|
||
). Posterior portion of pronotum lacking a longitudinal groove (
|
||
<figureCitation id="132C2A00BE46FFC9FE09BC4E82790765" box="[481,574,789,816]" captionStart="Fig" captionStartId="19.[189,232,1879,1905]" captionTargetBox="[197,1389,792,1835]" captionTargetId="figure-118@19.[189,1398,783,1839]" captionTargetPageId="19" captionText="Fig. 7. Pronotum of Georissus spp. A. G. (G.) canalifer Sharp, 1888. B. G. (G.) babai Satô, 1970. C. G. (Ni.) granulosus Satô, 1972. D. G. (Ne.) japonicus Satô, 1972. E–F. G. (Ne.) kurosawai Nakane, 1963 from Hokkaido (E) and Nagano (F). G–H. G. (Ne.) takahashii sp. nov. from Kochi (G) and Okinawa (H). I. G. (Ne.) satoi sp. nov." figureDoi="http://doi.org/10.5281/zenodo.6517976" httpUri="https://zenodo.org/record/6517976/files/figure.png" pageId="4" pageNumber="115">Fig. 7B</figureCitation>
|
||
). Internal margins of parameres covered with long setae (
|
||
<figureCitation id="132C2A00BE46FFC9FB08BC4E857D0765" box="[1248,1338,789,816]" captionStart="Fig" captionStartId="23.[189,232,1763,1789]" captionTargetBox="[239,1353,284,1696]" captionTargetId="figure-12@23.[189,1398,265,1704]" captionTargetPageId="23" captionText="Fig. 9. Male aedeagus of Georissus spp. A. G. (G.) canalifer Sharp, 1888. B. G. (G.) babai Satô, 1970. C. G. (Ni.) granulosus Satô, 1972. D. G. (Ne.) japonicus Satô, 1972. E. G. (Ne.) kurosawai Nakane, 1963. F–G. G. (Ne.) takahashii sp. nov. from Kochi (F) and Okinawa (G). H. G. (Ne.) satoi sp. nov. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6517986" httpUri="https://zenodo.org/record/6517986/files/figure.png" pageId="4" pageNumber="115">Fig. 9B</figureCitation>
|
||
) ..... ......................................................................................................................
|
||
<emphasis id="B963EA97BE46FFC9FBBCBC6384370706" bold="true" box="[1108,1136,824,851]" pageId="4" pageNumber="115">
|
||
<emphasis id="B963EA97BE46FFC9FBBCBC63842E0707" bold="true" box="[1108,1129,824,850]" italics="true" pageId="4" pageNumber="115">G</emphasis>
|
||
.
|
||
</emphasis>
|
||
<taxonomicName id="4C174D06BE46FFC9FB9FBC6385300706" ID-CoL="3FRYW" authority="Sato, 1970" authorityName="Sato" authorityYear="1970" box="[1143,1399,824,851]" class="Insecta" family="Georissidae" genus="Georissus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="115" phylum="Arthropoda" rank="species" species="babai" subGenus="Georissus">
|
||
<emphasis id="B963EA97BE46FFC9FB9FBC6384B50706" bold="true" box="[1143,1266,824,851]" pageId="4" pageNumber="115">
|
||
(
|
||
<emphasis id="B963EA97BE46FFC9FB69BC6384D10707" bold="true" box="[1153,1174,824,850]" italics="true" pageId="4" pageNumber="115">G</emphasis>
|
||
.)
|
||
<emphasis id="B963EA97BE46FFC9FB47BC6384B50706" bold="true" box="[1199,1266,824,851]" italics="true" pageId="4" pageNumber="115">babai</emphasis>
|
||
</emphasis>
|
||
<bibRefCitation id="EF864B74BE46FFC9FB11BC6285300706" author="Sato M." box="[1273,1399,825,851]" pageId="4" pageNumber="115" pagination="199 - 200" refId="ref14044" refString="Sato M. 1970. Une nouvelle espece de genre Georyssus [sic!] Latreille au Japon (Coleoptera: Georyssidae [sic!]). Journal of Nagoya Women's College 16: 199 - 200." type="journal article" year="1970">Satô, 1970</bibRefCitation>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BA83685BE46FFC9FF29BC07853007E9" blockId="4.[189,1401,548,1606]" pageId="4" pageNumber="115">
|
||
<keyLead id="30E695B0BE46FFC9FF29BC07853007E9" pageId="4" pageNumber="115">
|
||
– Granules of clypeus indistinct, small and sparse (
|
||
<figureCitation id="132C2A00BE46FFC9FCD5BC0783D80722" box="[829,927,860,887]" captionStart="Fig" captionStartId="18.[189,232,1882,1908]" captionTargetBox="[191,1394,563,1839]" captionTargetId="figure-90@18.[189,1398,562,1842]" captionTargetPageId="18" captionText="Fig. 6. Head of Georissus spp. A. G. (G.) canalifer Sharp, 1888. B. G. (G.) babai Satô, 1970. C. G. (Ni.) granulosus Satô, 1972. D. G. (Ne.) japonicus Satô, 1972. E–F. G. (Ne.) kurosawai Nakane, 1963 from Hokkaido (E) and Nagano (F). G–H. G. (Ne.) takahashii sp. nov. from Kochi (G) and Okinawa (H). I. G. (Ne.) satoi sp. nov." figureDoi="http://doi.org/10.5281/zenodo.6517968" httpUri="https://zenodo.org/record/6517968/files/figure.png" pageId="4" pageNumber="115">Fig. 6A</figureCitation>
|
||
). Posterior portion of pronotum having a shallow longitudinal groove (
|
||
<figureCitation id="132C2A00BE46FFC9FD8CBC24828107CC" box="[612,710,895,921]" captionStart="Fig" captionStartId="19.[189,232,1879,1905]" captionTargetBox="[197,1389,792,1835]" captionTargetId="figure-118@19.[189,1398,783,1839]" captionTargetPageId="19" captionText="Fig. 7. Pronotum of Georissus spp. A. G. (G.) canalifer Sharp, 1888. B. G. (G.) babai Satô, 1970. C. G. (Ni.) granulosus Satô, 1972. D. G. (Ne.) japonicus Satô, 1972. E–F. G. (Ne.) kurosawai Nakane, 1963 from Hokkaido (E) and Nagano (F). G–H. G. (Ne.) takahashii sp. nov. from Kochi (G) and Okinawa (H). I. G. (Ne.) satoi sp. nov." figureDoi="http://doi.org/10.5281/zenodo.6517976" httpUri="https://zenodo.org/record/6517976/files/figure.png" pageId="4" pageNumber="115">Fig. 7A</figureCitation>
|
||
). Internal margins of parameres covered with short setae (
|
||
<figureCitation id="132C2A00BE46FFC9FF1BBCF9810807E9" box="[243,335,930,956]" captionStart="Fig" captionStartId="23.[189,232,1763,1789]" captionTargetBox="[239,1353,284,1696]" captionTargetId="figure-12@23.[189,1398,265,1704]" captionTargetPageId="23" captionText="Fig. 9. Male aedeagus of Georissus spp. A. G. (G.) canalifer Sharp, 1888. B. G. (G.) babai Satô, 1970. C. G. (Ni.) granulosus Satô, 1972. D. G. (Ne.) japonicus Satô, 1972. E. G. (Ne.) kurosawai Nakane, 1963. F–G. G. (Ne.) takahashii sp. nov. from Kochi (F) and Okinawa (G). H. G. (Ne.) satoi sp. nov. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6517986" httpUri="https://zenodo.org/record/6517986/files/figure.png" pageId="4" pageNumber="115">Fig. 9A</figureCitation>
|
||
) ..............................................................................................
|
||
<emphasis id="B963EA97BE46FFC9FBF1BCF9847207E9" bold="true" box="[1049,1077,930,956]" pageId="4" pageNumber="115">
|
||
<emphasis id="B963EA97BE46FFC9FBF1BCF9846907E9" bold="true" box="[1049,1070,930,956]" italics="true" pageId="4" pageNumber="115">G</emphasis>
|
||
.
|
||
</emphasis>
|
||
<taxonomicName id="4C174D06BE46FFC9FBD5BCF9853007E9" ID-CoL="3FRYY" authority="Sharp, 1888" authorityName="Sharp" authorityYear="1888" box="[1085,1399,930,957]" class="Insecta" family="Georissidae" genus="Georissus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="115" phylum="Arthropoda" rank="species" species="canalifer" subGenus="Georissus">
|
||
<emphasis id="B963EA97BE46FFC9FBD5BCF984A607E8" bold="true" box="[1085,1249,930,957]" pageId="4" pageNumber="115">
|
||
(
|
||
<emphasis id="B963EA97BE46FFC9FBAFBCF9841B07E9" bold="true" box="[1095,1116,930,956]" italics="true" pageId="4" pageNumber="115">G</emphasis>
|
||
.)
|
||
<emphasis id="B963EA97BE46FFC9FB9CBCF984A607E8" bold="true" box="[1140,1249,930,957]" italics="true" pageId="4" pageNumber="115">canalifer</emphasis>
|
||
</emphasis>
|
||
<bibRefCitation id="EF864B74BE46FFC9FB01BCF9853007E9" author="Sharp D." box="[1257,1399,930,956]" pageId="4" pageNumber="115" pagination="242 - 245" refId="ref14169" refString="Sharp D. 1888. Descriptions of some new Coleoptera from Japan. The Annals and Magazine of Natural History Series 6 2: 242 - 245. https: // doi. org / 10.1080 / 00222938809460918" type="journal article" year="1888">Sharp, 1888</bibRefCitation>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30E32E20BE46FFC9FF55BCBF853000FB" pageId="4" pageNumber="115">
|
||
<paragraph id="8BA83685BE46FFC9FF55BCBF85300011" blockId="4.[189,1401,548,1606]" pageId="4" pageNumber="115">
|
||
<keyLead id="30E695B0BE46FFC9FF55BCBF85300011" pageId="4" pageNumber="115">
|
||
3. Posterior portion of pronotum densely bearing granules, lacking
|
||
<emphasis id="B963EA97BE46FFC9FBEEBCBF846507AB" box="[1030,1058,996,1022]" italics="true" pageId="4" pageNumber="115">cd</emphasis>
|
||
and
|
||
<emphasis id="B963EA97BE46FFC9FB8CBCBF84CA07AB" box="[1124,1165,996,1022]" italics="true" pageId="4" pageNumber="115">sdp</emphasis>
|
||
. Pronotum lacking
|
||
<emphasis id="B963EA97BE46FFC9FF02BB5C81540074" box="[234,275,1031,1057]" italics="true" pageId="4" pageNumber="115">sdn</emphasis>
|
||
(
|
||
<figureCitation id="132C2A00BE46FFC9FECEBB5C81C10074" box="[294,390,1031,1057]" captionStart="Fig" captionStartId="19.[189,232,1879,1905]" captionTargetBox="[197,1389,792,1835]" captionTargetId="figure-118@19.[189,1398,783,1839]" captionTargetPageId="19" captionText="Fig. 7. Pronotum of Georissus spp. A. G. (G.) canalifer Sharp, 1888. B. G. (G.) babai Satô, 1970. C. G. (Ni.) granulosus Satô, 1972. D. G. (Ne.) japonicus Satô, 1972. E–F. G. (Ne.) kurosawai Nakane, 1963 from Hokkaido (E) and Nagano (F). G–H. G. (Ne.) takahashii sp. nov. from Kochi (G) and Okinawa (H). I. G. (Ne.) satoi sp. nov." figureDoi="http://doi.org/10.5281/zenodo.6517976" httpUri="https://zenodo.org/record/6517976/files/figure.png" pageId="4" pageNumber="115">Fig. 7C</figureCitation>
|
||
). Aedeagus extremely narrow, phallobase posteriorly widening (
|
||
<figureCitation id="132C2A00BE46FFC9FB65BB5C84AA0074" box="[1165,1261,1031,1057]" captionStart="Fig" captionStartId="23.[189,232,1763,1789]" captionTargetBox="[239,1353,284,1696]" captionTargetId="figure-12@23.[189,1398,265,1704]" captionTargetPageId="23" captionText="Fig. 9. Male aedeagus of Georissus spp. A. G. (G.) canalifer Sharp, 1888. B. G. (G.) babai Satô, 1970. C. G. (Ni.) granulosus Satô, 1972. D. G. (Ne.) japonicus Satô, 1972. E. G. (Ne.) kurosawai Nakane, 1963. F–G. G. (Ne.) takahashii sp. nov. from Kochi (F) and Okinawa (G). H. G. (Ne.) satoi sp. nov. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6517986" httpUri="https://zenodo.org/record/6517986/files/figure.png" pageId="4" pageNumber="115">Fig. 9C</figureCitation>
|
||
). Subgenus
|
||
<taxonomicName id="4C174D06BE46FFC9FF02BB7082040011" ID-CoL="8XRHK" authority="Sato, 1972" authorityName="Sato" authorityYear="1972" box="[234,579,1066,1092]" class="Insecta" family="Georissidae" genus="Georissus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="115" phylum="Arthropoda" rank="subGenus" subGenus="Nipponogeorissus">
|
||
<emphasis id="B963EA97BE46FFC9FF02BB7081F90011" box="[234,446,1067,1092]" italics="true" pageId="4" pageNumber="115">Nipponogeorissus</emphasis>
|
||
<bibRefCitation id="EF864B74BE46FFC9FE2EBB7182040011" author="Sato M." box="[454,579,1066,1092]" pageId="4" pageNumber="115" pagination="207 - 213" refId="ref14083" refString="Sato M. 1972. The Georissid beetles of Japan. Journal of Nagoya Women's College 18: 207 - 213." type="journal article" year="1972">Satô, 1972</bibRefCitation>
|
||
</taxonomicName>
|
||
<emphasis id="B963EA97BE46FFC9FDB8BB70845B0011" box="[592,1052,1066,1092]" italics="true" pageId="4" pageNumber="115">
|
||
...........................................................
|
||
<emphasis id="B963EA97BE46FFC9FBEFBB71845B0011" bold="true" box="[1031,1052,1066,1092]" pageId="4" pageNumber="115">G</emphasis>
|
||
</emphasis>
|
||
.
|
||
<taxonomicName id="4C174D06BE46FFC9FBC3BB7185300011" ID-CoL="3FS3F" authority="Sato, 1972" authorityName="Sato" authorityYear="1972" box="[1067,1399,1065,1092]" class="Insecta" family="Georissidae" genus="Georissus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="115" phylum="Arthropoda" rank="species" species="granulosus" subGenus="Ni .">
|
||
(
|
||
<emphasis id="B963EA97BE46FFC9FBDDBB7184150011" bold="true" box="[1077,1106,1066,1092]" italics="true" pageId="4" pageNumber="115">Ni</emphasis>
|
||
.)
|
||
<emphasis id="B963EA97BE46FFC9FB82BB7284B60011" bold="true" box="[1130,1265,1065,1092]" italics="true" pageId="4" pageNumber="115">granulosus</emphasis>
|
||
<bibRefCitation id="EF864B74BE46FFC9FB11BB7185300011" author="Sato M." box="[1273,1399,1066,1092]" pageId="4" pageNumber="115" pagination="207 - 213" refId="ref14083" refString="Sato M. 1972. The Georissid beetles of Japan. Journal of Nagoya Women's College 18: 207 - 213." type="journal article" year="1972">Satô, 1972</bibRefCitation>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BA83685BE46FFC9FF29BB15853000FB" blockId="4.[189,1401,548,1606]" pageId="4" pageNumber="115">
|
||
<keyLead id="30E695B0BE46FFC9FF29BB15853000FB" pageId="4" pageNumber="115">
|
||
– Posterior portion of pronotum provided with granules, present
|
||
<emphasis id="B963EA97BE46FFC9FC09BB1683BA0032" box="[993,1021,1101,1127]" italics="true" pageId="4" pageNumber="115">cd</emphasis>
|
||
and
|
||
<emphasis id="B963EA97BE46FFC9FBD3BB1684230032" box="[1083,1124,1101,1127]" italics="true" pageId="4" pageNumber="115">sdp</emphasis>
|
||
. Pronotum having
|
||
<emphasis id="B963EA97BE46FFC9FAA6BB1685300032" box="[1358,1399,1101,1127]" italics="true" pageId="4" pageNumber="115">sdn</emphasis>
|
||
(
|
||
<figureCitation id="132C2A00BE46FFC9FF1BBB2B811400DE" box="[243,339,1136,1163]" captionStart="Fig" captionStartId="3.[189,232,1888,1914]" captionTargetBox="[189,1394,269,1844]" captionTargetId="figure-11@3.[189,1398,269,1844]" captionTargetPageId="3" captionText="Fig. 1. Georissus (Ne.) kurosawai Nakane, 1963, showing terminology. A. Head. B. Pronotum. Abbreviations: cd = central depression; lsa = lateral smooth area; pg = posterior grooves; sdn = subbasal dentation; sdp = subbasal depression; slb = sublateral bulge; slr = sublateral ridge; smb = submedian bulge; smlg = shallow median longitudinal groove." figureDoi="http://doi.org/10.5281/zenodo.6517950" httpUri="https://zenodo.org/record/6517950/files/figure.png" pageId="4" pageNumber="115">Fig. 1B</figureCitation>
|
||
). Aedeagus narrow, phallobase subparallel-sided (
|
||
<figureCitation id="132C2A00BE46FFC9FC51BB2B840600DE" box="[953,1089,1136,1163]" captionStart="Fig" captionStartId="23.[189,232,1763,1789]" captionTargetBox="[239,1353,284,1696]" captionTargetId="figure-12@23.[189,1398,265,1704]" captionTargetPageId="23" captionText="Fig. 9. Male aedeagus of Georissus spp. A. G. (G.) canalifer Sharp, 1888. B. G. (G.) babai Satô, 1970. C. G. (Ni.) granulosus Satô, 1972. D. G. (Ne.) japonicus Satô, 1972. E. G. (Ne.) kurosawai Nakane, 1963. F–G. G. (Ne.) takahashii sp. nov. from Kochi (F) and Okinawa (G). H. G. (Ne.) satoi sp. nov. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6517986" httpUri="https://zenodo.org/record/6517986/files/figure.png" pageId="4" pageNumber="115">Fig. 9D–H</figureCitation>
|
||
). Subgenus
|
||
<taxonomicName id="4C174D06BE46FFC9FB30BB2A812F00FB" ID-CoL="8XRHJ" authority="Sato, 1972" authorityName="Sato" authorityYear="1972" class="Insecta" family="Georissidae" genus="Georissus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="115" phylum="Arthropoda" rank="subGenus" subGenus="Neogeorissus">
|
||
<emphasis id="B963EA97BE46FFC9FB30BB2A853100DF" box="[1240,1398,1137,1162]" italics="true" pageId="4" pageNumber="115">Neogeorissus</emphasis>
|
||
<bibRefCitation id="EF864B74BE46FFC9FF02BBCF812F00FB" author="Sato M." box="[234,360,1172,1198]" pageId="4" pageNumber="115" pagination="207 - 213" refId="ref14083" refString="Sato M. 1972. The Georissid beetles of Japan. Journal of Nagoya Women's College 18: 207 - 213." type="journal article" year="1972">Satô, 1972</bibRefCitation>
|
||
</taxonomicName>
|
||
......................................................................................................................................... 4
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30E32E20BE46FFC9FF55BB8E85300146" pageId="4" pageNumber="115">
|
||
<paragraph id="8BA83685BE46FFC9FF55BB8E853000A5" blockId="4.[189,1401,548,1606]" box="[189,1399,1237,1264]" pageId="4" pageNumber="115">
|
||
<keyLead id="30E695B0BE46FFC9FF55BB8E853000A5" box="[189,1399,1237,1264]" pageId="4" pageNumber="115">
|
||
4. Elytral interstices 2, 4, 6 highly raised (
|
||
<figureCitation id="132C2A00BE46FFC9FD5EBB8E837100A5" box="[694,822,1237,1264]" captionStart="Fig" captionStartId="16.[189,232,1822,1848]" captionTargetBox="[192,1395,266,1778]" captionTargetId="figure-17@16.[189,1398,266,1783]" captionTargetPageId="16" captionText="Fig. 5. Dorsal habitus of Georissus spp. A. G. (G.) canalifer Sharp, 1888. B. G. (G.) babai Satô, 1970. C. G. (Ni.) granulosus Satô, 1972. D. G. (Ne.) japonicus Satô, 1972. E–F. G. (Ne.) kurosawai Nakane, 1963 from Hokkaido (E) and Nagano (F). G–H. G. (Ne.) takahashii sp. nov. from Kochi (G) and Okinawa (H). I. G. (Ne.) satoi sp. nov." figureDoi="http://doi.org/10.5281/zenodo.6517964" httpUri="https://zenodo.org/record/6517964/files/figure.png" pageId="4" pageNumber="115">Fig. 5G–H</figureCitation>
|
||
) ...............................
|
||
<emphasis id="B963EA97BE46FFC9FBC7BB8E855100A5" bold="true" box="[1071,1302,1237,1264]" pageId="4" pageNumber="115">
|
||
<emphasis id="B963EA97BE46FFC9FBC7BB8E840300BA" bold="true" box="[1071,1092,1237,1263]" italics="true" pageId="4" pageNumber="115">G</emphasis>
|
||
.
|
||
<taxonomicName id="4C174D06BE46FFC9FBBBBB8E855100A5" ID-CoL="9GD46" authority="Yasuda & Yoshitomi, 2022" authorityName="Yasuda & Yoshitomi" authorityYear="2022" box="[1107,1302,1237,1264]" class="Insecta" family="Georissidae" genus="Georissus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="115" phylum="Arthropoda" rank="species" species="takahashii" status="sp. nov." subGenus="Ne">
|
||
(
|
||
<emphasis id="B963EA97BE46FFC9FBB4BB8E843900BA" bold="true" box="[1116,1150,1237,1263]" italics="true" pageId="4" pageNumber="115">Ne</emphasis>
|
||
.)
|
||
<emphasis id="B963EA97BE46FFC9FB7FBB8E855100A5" bold="true" box="[1175,1302,1237,1264]" italics="true" pageId="4" pageNumber="115">takahashii</emphasis>
|
||
</taxonomicName>
|
||
</emphasis>
|
||
<taxonomicNameLabel id="A25057ECBE46FFC9FAF5BB8D853000A5" box="[1309,1399,1238,1264]" pageId="4" pageNumber="115" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BA83685BE46FFC9FF29BBA285300146" blockId="4.[189,1401,548,1606]" box="[193,1399,1272,1299]" pageId="4" pageNumber="115">
|
||
<keyLead id="30E695B0BE46FFC9FF29BBA285300146" box="[193,1399,1272,1299]" pageId="4" pageNumber="115">
|
||
– All elytral interstices equally raised (
|
||
<figureCitation id="132C2A00BE46FFC9FD73BBA3836C0146" box="[667,811,1272,1299]" captionStart="Fig" captionStartId="16.[189,232,1822,1848]" captionTargetBox="[192,1395,266,1778]" captionTargetId="figure-17@16.[189,1398,266,1783]" captionTargetPageId="16" captionText="Fig. 5. Dorsal habitus of Georissus spp. A. G. (G.) canalifer Sharp, 1888. B. G. (G.) babai Satô, 1970. C. G. (Ni.) granulosus Satô, 1972. D. G. (Ne.) japonicus Satô, 1972. E–F. G. (Ne.) kurosawai Nakane, 1963 from Hokkaido (E) and Nagano (F). G–H. G. (Ne.) takahashii sp. nov. from Kochi (G) and Okinawa (H). I. G. (Ne.) satoi sp. nov." figureDoi="http://doi.org/10.5281/zenodo.6517964" httpUri="https://zenodo.org/record/6517964/files/figure.png" pageId="4" pageNumber="115">Fig. 5D–F, I</figureCitation>
|
||
) .......................................................................... 5
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30E32E20BE46FFC9FF55BA62853001CF" pageId="4" pageNumber="115">
|
||
<paragraph id="8BA83685BE46FFC9FF55BA6285300122" blockId="4.[189,1401,548,1606]" pageId="4" pageNumber="115">
|
||
<keyLead id="30E695B0BE46FFC9FF55BA6285300122" pageId="4" pageNumber="115">
|
||
5. Coloration of body black. Elytral granules indistinct (
|
||
<figureCitation id="132C2A00BE46FFC9FCB6BA6183910101" box="[862,982,1337,1364]" captionStart="Fig" captionStartId="16.[189,232,1822,1848]" captionTargetBox="[192,1395,266,1778]" captionTargetId="figure-17@16.[189,1398,266,1783]" captionTargetPageId="16" captionText="Fig. 5. Dorsal habitus of Georissus spp. A. G. (G.) canalifer Sharp, 1888. B. G. (G.) babai Satô, 1970. C. G. (Ni.) granulosus Satô, 1972. D. G. (Ne.) japonicus Satô, 1972. E–F. G. (Ne.) kurosawai Nakane, 1963 from Hokkaido (E) and Nagano (F). G–H. G. (Ne.) takahashii sp. nov. from Kochi (G) and Okinawa (H). I. G. (Ne.) satoi sp. nov." figureDoi="http://doi.org/10.5281/zenodo.6517964" httpUri="https://zenodo.org/record/6517964/files/figure.png" pageId="4" pageNumber="115">Fig. 5E–F</figureCitation>
|
||
) ..................................................... .......................................................................................................
|
||
<emphasis id="B963EA97BE46FFC9FC0EBA0684440122" bold="true" box="[998,1027,1373,1399]" pageId="4" pageNumber="115">
|
||
<emphasis id="B963EA97BE46FFC9FC0EBA0683BC0122" bold="true" box="[998,1019,1373,1399]" italics="true" pageId="4" pageNumber="115">G</emphasis>
|
||
.
|
||
</emphasis>
|
||
<taxonomicName id="4C174D06BE46FFC9FBE2BA0785300122" ID-CoL="3FS2G" authority="Nakane, 1963" authorityName="Nakane" authorityYear="1963" box="[1034,1399,1372,1399]" class="Insecta" family="Georissidae" genus="Georissus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="115" phylum="Arthropoda" rank="species" species="kurosawai" subGenus="Ne">
|
||
<emphasis id="B963EA97BE46FFC9FBE2BA07848A0122" bold="true" box="[1034,1229,1372,1399]" pageId="4" pageNumber="115">
|
||
(
|
||
<emphasis id="B963EA97BE46FFC9FBFCBA0684710122" bold="true" box="[1044,1078,1373,1399]" italics="true" pageId="4" pageNumber="115">Ne</emphasis>
|
||
.)
|
||
<emphasis id="B963EA97BE46FFC9FBA7BA07848A0122" bold="true" box="[1103,1229,1372,1399]" italics="true" pageId="4" pageNumber="115">kurosawai</emphasis>
|
||
</emphasis>
|
||
<bibRefCitation id="EF864B74BE46FFC9FB3CBA0685300122" author="Nakane T." box="[1236,1399,1373,1399]" pageId="4" pageNumber="115" pagination="2" refId="ref13863" refString="Nakane T. 1963. Hydrophilidae. In: Nakane T., Ohbayashi K., Nomura S. & Kurosawa Y. (eds) Iconographia Insetorum Japonicorum Colore naturali edita Vol. 2 (Coleoptera). Hokuryukan, Tokyo. [In Japanese.]" type="book chapter" year="1963">Nakane, 1963</bibRefCitation>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BA83685BE46FFC9FF29BADA853001CF" blockId="4.[189,1401,548,1606]" box="[193,1399,1408,1435]" pageId="4" pageNumber="115">
|
||
<keyLead id="30E695B0BE46FFC9FF29BADA853001CF" box="[193,1399,1408,1435]" pageId="4" pageNumber="115">
|
||
– Coloration of body dark reddish brown. Elytral granules distinct (
|
||
<figureCitation id="132C2A00BE46FFC9FC04BADB841801CE" box="[1004,1119,1408,1435]" captionStart="Fig" captionStartId="16.[189,232,1822,1848]" captionTargetBox="[192,1395,266,1778]" captionTargetId="figure-17@16.[189,1398,266,1783]" captionTargetPageId="16" captionText="Fig. 5. Dorsal habitus of Georissus spp. A. G. (G.) canalifer Sharp, 1888. B. G. (G.) babai Satô, 1970. C. G. (Ni.) granulosus Satô, 1972. D. G. (Ne.) japonicus Satô, 1972. E–F. G. (Ne.) kurosawai Nakane, 1963 from Hokkaido (E) and Nagano (F). G–H. G. (Ne.) takahashii sp. nov. from Kochi (G) and Okinawa (H). I. G. (Ne.) satoi sp. nov." figureDoi="http://doi.org/10.5281/zenodo.6517964" httpUri="https://zenodo.org/record/6517964/files/figure.png" pageId="4" pageNumber="115">Fig. 5D, I</figureCitation>
|
||
) ................................ 6
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30E32E20BE46FFC9FF55BA9A85300213" pageId="4" pageNumber="115">
|
||
<paragraph id="8BA83685BE46FFC9FF55BA9A853001AA" blockId="4.[189,1401,548,1606]" pageId="4" pageNumber="115">
|
||
<keyLead id="30E695B0BE46FFC9FF55BA9A853001AA" pageId="4" pageNumber="115">
|
||
6. Coloration of body black to dark brown. Granulation of pronotum distinct (
|
||
<figureCitation id="132C2A00BE46FFC9FB84BA9A848B0189" box="[1132,1228,1473,1500]" captionStart="Fig" captionStartId="19.[189,232,1879,1905]" captionTargetBox="[197,1389,792,1835]" captionTargetId="figure-118@19.[189,1398,783,1839]" captionTargetPageId="19" captionText="Fig. 7. Pronotum of Georissus spp. A. G. (G.) canalifer Sharp, 1888. B. G. (G.) babai Satô, 1970. C. G. (Ni.) granulosus Satô, 1972. D. G. (Ne.) japonicus Satô, 1972. E–F. G. (Ne.) kurosawai Nakane, 1963 from Hokkaido (E) and Nagano (F). G–H. G. (Ne.) takahashii sp. nov. from Kochi (G) and Okinawa (H). I. G. (Ne.) satoi sp. nov." figureDoi="http://doi.org/10.5281/zenodo.6517976" httpUri="https://zenodo.org/record/6517976/files/figure.png" pageId="4" pageNumber="115">Fig. 7D</figureCitation>
|
||
). Median lobe shorter than parameres (
|
||
<figureCitation id="132C2A00BE46FFC9FDEFBABE822401AA" box="[519,611,1509,1535]" captionStart="Fig" captionStartId="23.[189,232,1763,1789]" captionTargetBox="[239,1353,284,1696]" captionTargetId="figure-12@23.[189,1398,265,1704]" captionTargetPageId="23" captionText="Fig. 9. Male aedeagus of Georissus spp. A. G. (G.) canalifer Sharp, 1888. B. G. (G.) babai Satô, 1970. C. G. (Ni.) granulosus Satô, 1972. D. G. (Ne.) japonicus Satô, 1972. E. G. (Ne.) kurosawai Nakane, 1963. F–G. G. (Ne.) takahashii sp. nov. from Kochi (F) and Okinawa (G). H. G. (Ne.) satoi sp. nov. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6517986" httpUri="https://zenodo.org/record/6517986/files/figure.png" pageId="4" pageNumber="115">Fig. 9D</figureCitation>
|
||
) ........................................................
|
||
<taxonomicName id="4C174D06BE46FFC9FBFCBABE853001AA" ID-CoL="3FS2D" authority="Sato, 1972" authorityName="Sato" authorityYear="1972" box="[1044,1399,1508,1535]" class="Insecta" family="Georissidae" genus="Georissus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="124" phylum="Arthropoda" rank="species" species="japonicus" subGenus="Ne.">
|
||
<emphasis id="B963EA97BE46FFC9FBFCBABE84B601AA" bold="true" box="[1044,1265,1508,1535]" pageId="4" pageNumber="115">
|
||
<emphasis id="B963EA97BE46FFC9FBFCBABE846E01AA" bold="true" box="[1044,1065,1509,1535]" italics="true" pageId="4" pageNumber="115">G</emphasis>
|
||
. (
|
||
<emphasis id="B963EA97BE46FFC9FBAABABE842301AA" bold="true" box="[1090,1124,1509,1535]" italics="true" pageId="4" pageNumber="115">Ne</emphasis>
|
||
.)
|
||
<emphasis id="B963EA97BE46FFC9FB94BABE84B601AA" bold="true" box="[1148,1265,1509,1535]" italics="true" pageId="4" pageNumber="115">japonicus</emphasis>
|
||
</emphasis>
|
||
<bibRefCitation id="EF864B74BE46FFC9FB11BABE853001AA" author="Sato M." box="[1273,1399,1509,1535]" pageId="4" pageNumber="115" pagination="207 - 213" refId="ref14083" refString="Sato M. 1972. The Georissid beetles of Japan. Journal of Nagoya Women's College 18: 207 - 213." type="journal article" year="1972">Satô, 1972</bibRefCitation>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BA83685BE46FFC9FF29B95385300213" blockId="4.[189,1401,548,1606]" pageId="4" pageNumber="115">
|
||
<keyLead id="30E695B0BE46FFC9FF29B95385300213" pageId="4" pageNumber="115">
|
||
– Coloration of dark reddish brown. Granulation of pronotum indistinct (
|
||
<figureCitation id="132C2A00BE46FFC9FBC2B953843B0277" box="[1066,1148,1544,1570]" captionStart="Fig" captionStartId="19.[189,232,1879,1905]" captionTargetBox="[197,1389,792,1835]" captionTargetId="figure-118@19.[189,1398,783,1839]" captionTargetPageId="19" captionText="Fig. 7. Pronotum of Georissus spp. A. G. (G.) canalifer Sharp, 1888. B. G. (G.) babai Satô, 1970. C. G. (Ni.) granulosus Satô, 1972. D. G. (Ne.) japonicus Satô, 1972. E–F. G. (Ne.) kurosawai Nakane, 1963 from Hokkaido (E) and Nagano (F). G–H. G. (Ne.) takahashii sp. nov. from Kochi (G) and Okinawa (H). I. G. (Ne.) satoi sp. nov." figureDoi="http://doi.org/10.5281/zenodo.6517976" httpUri="https://zenodo.org/record/6517976/files/figure.png" pageId="4" pageNumber="115">Fig. 7I</figureCitation>
|
||
). Median lobe longer than parameres (
|
||
<figureCitation id="132C2A00BE46FFC9FE45B970824E0210" box="[429,521,1579,1605]" captionStart="Fig" captionStartId="23.[189,232,1763,1789]" captionTargetBox="[239,1353,284,1696]" captionTargetId="figure-12@23.[189,1398,265,1704]" captionTargetPageId="23" captionText="Fig. 9. Male aedeagus of Georissus spp. A. G. (G.) canalifer Sharp, 1888. B. G. (G.) babai Satô, 1970. C. G. (Ni.) granulosus Satô, 1972. D. G. (Ne.) japonicus Satô, 1972. E. G. (Ne.) kurosawai Nakane, 1963. F–G. G. (Ne.) takahashii sp. nov. from Kochi (F) and Okinawa (G). H. G. (Ne.) satoi sp. nov. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6517986" httpUri="https://zenodo.org/record/6517986/files/figure.png" pageId="4" pageNumber="115">Fig. 9H</figureCitation>
|
||
) .................................................................................
|
||
<emphasis id="B963EA97BE46FFC9FB9CB97085520210" bold="true" box="[1140,1301,1579,1605]" pageId="4" pageNumber="115">
|
||
<emphasis id="B963EA97BE46FFC9FB9CB97084CE0210" bold="true" box="[1140,1161,1579,1605]" italics="true" pageId="4" pageNumber="115">G</emphasis>
|
||
.
|
||
<taxonomicName id="4C174D06BE46FFC9FB7FB97085520210" ID-CoL="9GD45" authority="Yasuda & Yoshitomi, 2022" authorityName="Yasuda & Yoshitomi" authorityYear="2022" box="[1175,1301,1579,1605]" class="Insecta" family="Georissidae" genus="Georissus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="115" phylum="Arthropoda" rank="species" species="satoi" status="sp. nov." subGenus="Ne">
|
||
(
|
||
<emphasis id="B963EA97BE46FFC9FB49B97084840210" bold="true" box="[1185,1219,1579,1605]" italics="true" pageId="4" pageNumber="115">Ne</emphasis>
|
||
.)
|
||
<emphasis id="B963EA97BE46FFC9FB34B97085520210" bold="true" box="[1244,1301,1579,1605]" italics="true" pageId="4" pageNumber="115">satoi</emphasis>
|
||
</taxonomicName>
|
||
</emphasis>
|
||
<taxonomicNameLabel id="A25057ECBE46FFC9FAF4B97785300213" box="[1308,1399,1580,1606]" pageId="4" pageNumber="115" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
</key>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |