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<document id="DEB8B590C14DF255EE8D7AAE2C35236C" ID="10.11646/zootaxa.4277.1.2SLASH11353" ID-CLB-Dataset="32839" ID-DOI="10.11646/zootaxa.4277.1.2" ID-GBIF-Dataset="bc734cb4-fc74-4e84-82a7-715d00735b7a" ID-ISSN="1175-5326" ID-Zenodo-Dep="808980" ID-ZooBank="60FF6986-6FD7-473C-932E-9DF7AB1580FC" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1497510936264" checkinUser="plazi" docAuthor="Tuzovskij, Petr V." docDate="2017" docId="03BE87932104D654FF7CFC1152FCFC36" docLanguage="en" docName="zootaxa.4277.1.2SLASH11353.pdf" docOrigin="Zootaxa 4277 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Hygrobates nigromaculatus Lebert 1879" docType="treatment" docVersion="7" lastPageNumber="24" masterDocId="FF87FFEB2105D653FFEBFFF55205FFB6" masterDocTitle="On the systematics of the water mites Hygrobates nigromaculatus Lebert, 1879 and H. setosus Besseling, 1942 (Acari, Hydrachnidia, Hygrobatidae)" masterLastPageNumber="31" masterPageNumber="17" pageNumber="18" updateTime="1698440525943" updateUser="ExternalLinkService">
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<mods:title id="B7EBD80C5BF248A6AAD3E1E7C7E69317">On the systematics of the water mites Hygrobates nigromaculatus Lebert, 1879 and H. setosus Besseling, 1942 (Acari, Hydrachnidia, Hygrobatidae)</mods:title>
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<mods:name id="8A7D94A5B9BAB954FDA424FDB33DB6A7" type="personal">
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<mods:namePart id="28A4E55E4C843A45AAD9F8199EECE466">Tuzovskij, Petr V.</mods:namePart>
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<mods:date id="E3F0545E131A3EEC6267E74C61723BBC">2017</mods:date>
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<treatment id="03BE87932104D654FF7CFC1152FCFC36" ID-DOI="http://doi.org/10.5281/zenodo.6022536" ID-GBIF-Taxon="131878627" ID-Zenodo-Dep="6022536" LSID="urn:lsid:plazi:treatment:03BE87932104D654FF7CFC1152FCFC36" httpUri="http://treatment.plazi.org/id/03BE87932104D654FF7CFC1152FCFC36" lastPageId="7" lastPageNumber="24" pageId="1" pageNumber="18">
<subSubSection id="C30D650E2104D652FF7CFC11531DFB96" pageId="1" pageNumber="18" type="nomenclature">
<paragraph id="8BA836852104D652FF7CFC115089FC49" blockId="1.[151,652,996,1056]" box="[151,652,996,1023]" pageId="1" pageNumber="18">
<heading id="D0E081E92104D652FF7CFC115089FC49" bold="true" box="[151,652,996,1023]" fontSize="11" level="1" pageId="1" pageNumber="18" reason="1">
<taxonomicName id="4C174D062104D652FF7CFC115089FC49" authority="Lebert, 1879" authorityName="Lebert" authorityYear="1879" box="[151,652,996,1023]" class="Arachnida" family="Hygrobatidae" genus="Hygrobates" kingdom="Animalia" order="Trombidiformes" pageId="1" pageNumber="18" phylum="Arthropoda" rank="species" species="nigromaculatus">
<emphasis id="B963EA972104D652FF7CFC115089FC49" bold="true" box="[151,652,996,1023]" pageId="1" pageNumber="18">
<emphasis id="B963EA972104D652FF7CFC1153E2FC48" bold="true" box="[151,487,996,1022]" italics="true" pageId="1" pageNumber="18">Hygrobates nigromaculatus</emphasis>
Lebert, 1879
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BA836852104D652FF7CFBF2531DFB96" blockId="1.[151,652,996,1056]" box="[151,280,1031,1056]" pageId="1" pageNumber="18">
(
<figureCitation id="132C2A002104D652FF74FBF25315FB96" box="[159,272,1031,1056]" captionStart-0="FIGURES 1 2" captionStart-1="FIGURES 3 10" captionStart-2="FIGURES 11 16" captionStart-3="FIGURES 17 23" captionStart-4="FIGURES 24 26" captionStartId-0="2.[151,264,1058,1081]" captionStartId-1="3.[151,264,1952,1975]" captionStartId-2="4.[151,264,1198,1221]" captionStartId-3="5.[151,264,1424,1447]" captionStartId-4="6.[151,264,1952,1975]" captionTargetBox-0="[176,1396,201,1034]" captionTargetBox-1="[201,1384,199,1917]" captionTargetBox-2="[162,1422,197,1168]" captionTargetBox-3="[155,1428,223,1387]" captionTargetBox-4="[183,1416,202,1916]" captionTargetId-0="figure@2.[170,1408,193,1041]" captionTargetId-1="figure@3.[185,1402,193,1932]" captionTargetId-2="figure@4.[150,1436,193,1177]" captionTargetId-3="figure@5.[155,1440,215,1397]" captionTargetId-4="figure@6.[161,1425,193,1932]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionTargetPageId-3="5" captionTargetPageId-4="6" captionText-0="FIGURES 1 2. Hygobates nigromaculatus Lebert, 1879, larva: 1, dorsal view; 2, ventral view. Scale bar: 100 µm." captionText-1="FIGURES 3 10. Hygobates nigromaculatus Lebert, 1879, larva: 3, chelicera, dorsal view; 4, chela, lateral view; 5, pedipalp, lateral view; 6, Leg I; 7, Leg II; 8, Leg III; 9, claws of leg I; 10, claws of leg III. Scale bars: 3, 6 8 = 50 µm, 4 5, 9 10 = 20 µm." captionText-2="FIGURES 11 16. Hygobates nigromaculatus Lebert, 1879, deutonymph: 11, seta Fch; 12, idiosoma, ventral view; 13, pedipalp, lateral view; 14, I-Leg- 4 6; 15, IV-Leg- 4 6, 16, claw. Scale bars: 11, 13, 14 15 = 50 µm, 12 = 100 µm, 16 = 20 µm." captionText-3="FIGURES 17 23. Hygobates nigromaculatus Lebert, 1879, female: 17, seta Fch; 18, coxal plates I IV, ventral view; 19, genital field; 20 21, genital plate; 22, pedipalp, lateral view; 23, I-Leg- 4 6. Scale bars: 17, 22 = 50 µm, 18 = 200 µm, 19 21, 23 = 100 µm." captionText-4="FIGURES 24 26. Hygobates nigromaculatus Lebert, 1879, male: 24 25, genital plate; 26, ejaculatory complex. Scale bars: 24, 25 = 100 µm, 26 = 50 µm." httpUri-0="https://zenodo.org/record/808982/files/figure.png" httpUri-1="https://zenodo.org/record/808984/files/figure.png" httpUri-2="https://zenodo.org/record/808986/files/figure.png" httpUri-3="https://zenodo.org/record/808988/files/figure.png" httpUri-4="https://zenodo.org/record/808990/files/figure.png" pageId="1" pageNumber="18">Figs 126</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C30D650E2104D652FF7CFBBA5315FAAB" pageId="1" pageNumber="18" type="materials_examined">
<paragraph id="8BA836852104D652FF7CFBBA5315FAAB" blockId="1.[151,1437,1103,2028]" pageId="1" pageNumber="18">
<materialsCitation id="3B7F3CD82104D652FF7CFBBA560EFB3A" ID-GBIF-Occurrence="1563886348" collectingDate="2000" collectingDateMax="2002" collectingDateMin="2000" country="Russia" county="Nekouz District" location="Pogorelka" municipality="Rybinsk" pageId="1" pageNumber="18" specimenCount="53" specimenCount-female="13" specimenCount-larva="26" specimenCount-male="14" stateProvince="Yaroslavl Province">
<emphasis id="B963EA972104D652FF7CFBBA537EFBDE" bold="true" box="[151,379,1103,1128]" pageId="1" pageNumber="18">Material examined</emphasis>
.
<specimenCount id="9D11FD0C2104D652FE67FBA5500EFBDE" box="[396,523,1103,1128]" pageId="1" pageNumber="18" type="female">13 females</specimenCount>
,
<specimenCount id="9D11FD0C2104D652FDF2FBA55087FBDE" box="[537,642,1103,1128]" pageId="1" pageNumber="18" type="male">14 males</specimenCount>
, 17 deutonymphs and
<specimenCount id="9D11FD0C2104D652FC6CFBBA51F3FBDE" box="[903,1014,1103,1128]" pageId="1" pageNumber="18" type="larva">26 larvae</specimenCount>
:
<collectingCountry id="F30076152104D652FBE9FBBA5655FBDE" box="[1026,1104,1103,1128]" name="Russia" pageId="1" pageNumber="18">Russia</collectingCountry>
,
<collectingRegion id="49D3F8672104D652FBB5FBBA573DFBDE" box="[1118,1336,1103,1128]" country="Russia" name="Yaroslavl'" pageId="1" pageNumber="18">Yaroslavl Province</collectingRegion>
,
<collectingCounty id="62C94E092104D652FAAEFBBA52EBFB3B" pageId="1" pageNumber="18">Nekouz District</collectingCounty>
,
<collectingMunicipality id="6BCCACFF2104D652FF16FB81535CFB3B" box="[253,345,1140,1165]" pageId="1" pageNumber="18">Rybinsk</collectingMunicipality>
reservoir, near village
<location id="8EC8605E2104D652FD82FB8150DBFB3B" LSID="urn:lsid:plazi:treatment:03BE87932104D654FF7CFC1152FCFC36:8EC8605E2104D652FD82FB8150DBFB3B" box="[617,734,1140,1165]" country="Russia" county="Nekouz District" municipality="Rybinsk" name="Pogorelka" pageId="1" pageNumber="18" stateProvince="Yaroslavl Province">Pogorelka</location>
,
<location id="8EC8605E2104D652FD06FB81517AFB3A" LSID="urn:lsid:plazi:treatment:03BE87932104D654FF7CFC1152FCFC36:8EC8605E2104D652FD06FB81517AFB3A" box="[749,895,1140,1164]" country="Russia" county="Nekouz District" municipality="Rybinsk" name="June - August" pageId="1" pageNumber="18" stateProvince="Yaroslavl Province">JuneAugust</location>
<collectingDate id="EFEDE9AD2104D652FC62FB81560EFB3A" box="[905,1035,1140,1164]" pageId="1" pageNumber="18" value="2000" valueMax="2002">
<date id="FFA910452104D652FC62FB81560EFB3A" box="[905,1035,1140,1164]" pageId="1" pageNumber="18" value="2000" valueMax="2002">20002002</date>
</collectingDate>
</materialsCitation>
;
<materialsCitation id="3B7F3CD82104D652FBF1FB815092FB06" ID-GBIF-Occurrence="1563886350" collectingDate="1975-08-03" country="Russia" location="Pleshcheevo Lake" pageId="1" pageNumber="18" specimenCount="1" specimenCount-male="1" stateProvince="Yaroslavl Province">
<specimenCount id="9D11FD0C2104D652FBF1FB815669FB3B" box="[1050,1132,1140,1165]" pageId="1" pageNumber="18" type="male">1 male</specimenCount>
, 1 deutonymph:
<collectingRegion id="49D3F8672104D652FAD9FB8152F8FB06" country="Russia" name="Yaroslavl'" pageId="1" pageNumber="18">Yaroslavl Province</collectingRegion>
,
<location id="8EC8605E2104D652FEE0FB6253D9FB06" LSID="urn:lsid:plazi:treatment:03BE87932104D654FF7CFC1152FCFC36:8EC8605E2104D652FEE0FB6253D9FB06" box="[267,476,1175,1200]" country="Russia" name="Pleshcheevo Lake" pageId="1" pageNumber="18" stateProvince="Yaroslavl Province">Pleshcheevo Lake</location>
,
<collectingDate id="EFEDE9AD2104D652FE02FB6D5092FB06" box="[489,663,1175,1200]" pageId="1" pageNumber="18" value="1975-08-03">
<date id="FFA910452104D652FE02FB6D5092FB06" box="[489,663,1175,1200]" pageId="1" pageNumber="18" value="1975-08-03">3 August 1975</date>
</collectingDate>
</materialsCitation>
;
<materialsCitation id="3B7F3CD82104D652FD4EFB6D5315FAAB" ID-GBIF-Occurrence="1563886351" collectingDate="1992-06" country="Russia" county="Stavropol District" location="Park" municipality="National" pageId="1" pageNumber="18" specimenCount="10" specimenCount-female="4" specimenCount-male="6" stateProvince="Samara Province">
<specimenCount id="9D11FD0C2104D652FD4EFB6D5112FB06" box="[677,791,1175,1200]" pageId="1" pageNumber="18" type="female">4 females</specimenCount>
,
<specimenCount id="9D11FD0C2104D652FCCEFB625187FB06" box="[805,898,1175,1200]" pageId="1" pageNumber="18" type="male">6 males</specimenCount>
:
<collectingRegion id="49D3F8672104D652FC64FB6D5651FB06" box="[911,1108,1175,1200]" country="Russia" name="Samara" pageId="1" pageNumber="18">Samara Province</collectingRegion>
,
<collectingCounty id="62C94E092104D652FB8AFB62572BFB06" box="[1121,1326,1175,1200]" pageId="1" pageNumber="18">Stavropol District</collectingCounty>
,
<collectingMunicipality id="6BCCACFF2104D652FAD7FB625799FB06" box="[1340,1436,1175,1200]" pageId="1" pageNumber="18">National</collectingMunicipality>
natural
<location id="8EC8605E2104D652FF07FB49531AFB63" LSID="urn:lsid:plazi:treatment:03BE87932104D654FF7CFC1152FCFC36:8EC8605E2104D652FF07FB49531AFB63" box="[236,287,1212,1237]" country="Russia" county="Stavropol District" municipality="National" name="Park" pageId="1" pageNumber="18" stateProvince="Samara Province">Park</location>
<location id="8EC8605E2104D652FEDAFB495005FB63" LSID="urn:lsid:plazi:treatment:03BE87932104D654FF7CFC1152FCFC36:8EC8605E2104D652FEDAFB495005FB63" box="[305,512,1212,1237]" country="Russia" county="Stavropol District" municipality="National" name="Samarskaya Luka" pageId="1" pageNumber="18" stateProvince="Samara Province">Samarskaya Luka</location>
”,
<location id="8EC8605E2104D652FDFCFB495095FB63" LSID="urn:lsid:plazi:treatment:03BE87932104D654FF7CFC1152FCFC36:8EC8605E2104D652FDFCFB495095FB63" box="[535,656,1212,1237]" country="Russia" county="Stavropol District" municipality="National" name="Kuibyshev" pageId="1" pageNumber="18" stateProvince="Samara Province">Kuibyshev</location>
reservoir near village
<location id="8EC8605E2104D652FC65FB4951FDFB63" LSID="urn:lsid:plazi:treatment:03BE87932104D654FF7CFC1152FCFC36:8EC8605E2104D652FC65FB4951FDFB63" box="[910,1016,1212,1237]" country="Russia" county="Stavropol District" municipality="National" name="Koltsovo" pageId="1" pageNumber="18" stateProvince="Samara Province">Koltsovo</location>
,
<collectingDate id="EFEDE9AD2104D652FBE8FB48567CFB62" box="[1027,1145,1212,1236]" pageId="1" pageNumber="18" value="1992-06">
<date id="FFA910452104D652FBE8FB48567CFB62" box="[1027,1145,1212,1236]" pageId="1" pageNumber="18" value="1992-06">June 1992</date>
</collectingDate>
. Larvae were reared from six females, the duration of the embryonic period was 1015 days; 15 deutonymphs were reared from larvae in laboratory.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C30D650E2104D652FF2CFAD2530CFA1B" pageId="1" pageNumber="18" type="diagnosis">
<paragraph id="8BA836852104D652FF2CFAD2530CFA1B" blockId="1.[151,1437,1103,2028]" pageId="1" pageNumber="18">
<emphasis id="B963EA972104D652FF2CFAD2539CFAF6" bold="true" box="[199,409,1319,1344]" pageId="1" pageNumber="18">Diagnosis. Larva.</emphasis>
Dorsal plate broad (L/W ratio 1.081.25) with convex lateral margins, coxal groups fused to each other medially but suture line between them present;
<emphasis id="B963EA972104D652FCB3FAB85188FAD2" box="[856,909,1357,1380]" italics="true" pageId="1" pageNumber="18">tmas</emphasis>
on coxae III absent; setae
<emphasis id="B963EA972104D652FB2EFAB856DBFAD2" box="[1221,1246,1357,1380]" italics="true" pageId="1" pageNumber="18">Pi</emphasis>
and
<emphasis id="B963EA972104D652FAF2FAB85732FAD2" box="[1305,1335,1357,1380]" italics="true" pageId="1" pageNumber="18">Pe</emphasis>
equal in length; number of thickened distal setae from trochanter to tarsus on legs segments: I: 1-2-2-1-0, II: 0-2-3-4-0, III: 0-2-4-5-0.
</paragraph>
</subSubSection>
<subSubSection id="C30D650E2104D654FF2CFA4251AAFED2" lastPageId="7" lastPageNumber="24" pageId="1" pageNumber="18" type="description">
<paragraph id="8BA836852104D652FF2CFA425352F9D6" blockId="1.[151,1437,1103,2028]" pageId="1" pageNumber="18">
<emphasis id="B963EA972104D652FF2CFA425354FA66" bold="true" box="[199,337,1463,1488]" pageId="1" pageNumber="18">Description</emphasis>
. Dorsal plate broad (L/W ratio 1.081.25), covering large part of dorsum, acute anteriorly, with convex lateral margins (
<figureCitation id="132C2A002104D652FE4EFA2953E3FA42" box="[421,486,1500,1525]" captionStart="FIGURES 1 2" captionStartId="2.[151,264,1058,1081]" captionTargetBox="[176,1396,201,1034]" captionTargetId="figure@2.[170,1408,193,1041]" captionTargetPageId="2" captionText="FIGURES 1 2. Hygobates nigromaculatus Lebert, 1879, larva: 1, dorsal view; 2, ventral view. Scale bar: 100 µm." httpUri="https://zenodo.org/record/808982/files/figure.png" pageId="1" pageNumber="18">Fig. 1</figureCitation>
); simple setae long and thick, but setae
<emphasis id="B963EA972104D652FC4DFA2951D7FA42" box="[934,978,1500,1524]" italics="true" pageId="1" pageNumber="18">Fch</emphasis>
slightly longer than
<emphasis id="B963EA972104D652FB51FA2856D0FA42" box="[1210,1237,1501,1524]" italics="true" pageId="1" pageNumber="18">Vi,</emphasis>
trichobothria
<emphasis id="B963EA972104D652FA9DFA2852B6F9A1" italics="true" pageId="1" pageNumber="18">Fp, Oi</emphasis>
thin, short and equal in length. Eight pairs of setae (
<emphasis id="B963EA972104D652FCECF9F5565AF9A1" box="[775,1119,1536,1560]" italics="true" pageId="1" pageNumber="18">Oe, Hi, He, Sci, Sce, Li, Le, Si</emphasis>
) situated in soft membrane;
<emphasis id="B963EA972104D652FF7CF9D152E1F98A" box="[151,228,1572,1596]" italics="true" pageId="1" pageNumber="18">Oe, Hi</emphasis>
and
<emphasis id="B963EA972104D652FEF5F9D15340F98D" box="[286,325,1572,1595]" italics="true" pageId="1" pageNumber="18">Sce</emphasis>
subequal in length and longer than other setae situated in the membrane. Surface of dorsal plate with reticulation.
</paragraph>
<paragraph id="8BA836852104D652FF2CF9995655F836" blockId="1.[151,1437,1103,2028]" pageId="1" pageNumber="18">
Both coxal groups fused to each other medially but suture line between them present (
<figureCitation id="132C2A002104D652FB7DF99956D9F932" box="[1174,1244,1644,1669]" captionStart="FIGURES 1 2" captionStartId="2.[151,264,1058,1081]" captionTargetBox="[176,1396,201,1034]" captionTargetId="figure@2.[170,1408,193,1041]" captionTargetPageId="2" captionText="FIGURES 1 2. Hygobates nigromaculatus Lebert, 1879, larva: 1, dorsal view; 2, ventral view. Scale bar: 100 µm." httpUri="https://zenodo.org/record/808982/files/figure.png" pageId="1" pageNumber="18">Fig. 2</figureCitation>
). Coxal setae C1 and C2 relatively short, thin and subequal in length; C3 located on coxae III anteriolaterally, slightly shorter than C4 but longer than anterior coxal setae. Transverse muscle attachment scar on coxae III absent. Setae
<emphasis id="B963EA972104D652FAC0F9405741F97A" box="[1323,1348,1717,1740]" italics="true" pageId="1" pageNumber="18">Pi</emphasis>
and
<emphasis id="B963EA972104D652FA95F940579EF97A" box="[1406,1435,1717,1740]" italics="true" pageId="1" pageNumber="18">Pe</emphasis>
short, thin and equal in length. Urstigmae relatively large and oval shaped. Setae
<emphasis id="B963EA972104D652FBAAF92D565EF959" box="[1089,1115,1752,1775]" italics="true" pageId="1" pageNumber="18">Ci</emphasis>
(
<figureCitation id="132C2A002104D652FB86F92256B4F946" box="[1133,1201,1751,1776]" captionStart="FIGURES 3 10" captionStartId="3.[151,264,1952,1975]" captionTargetBox="[201,1384,199,1917]" captionTargetId="figure@3.[185,1402,193,1932]" captionTargetPageId="3" captionText="FIGURES 3 10. Hygobates nigromaculatus Lebert, 1879, larva: 3, chelicera, dorsal view; 4, chela, lateral view; 5, pedipalp, lateral view; 6, Leg I; 7, Leg II; 8, Leg III; 9, claws of leg I; 10, claws of leg III. Scale bars: 3, 6 8 = 50 µm, 4 5, 9 10 = 20 µm." httpUri="https://zenodo.org/record/808984/files/figure.png" pageId="1" pageNumber="18">Fig. 3</figureCitation>
) very long and well thickened, inserted on moderately long projections. Excretory pore plate wider than long (L/W ratio 0.450.50), slightly convex anteriorly, “W” shaped posteriorly and with muscle attachment scars anteriorly; excretory pore located near the center of plate between bases of setae
<emphasis id="B963EA972104D652FCCAF8B1514DF8ED" box="[801,840,1860,1883]" italics="true" pageId="1" pageNumber="18">Ae;</emphasis>
bases of setae
<emphasis id="B963EA972104D652FC14F8B1561DF8ED" box="[1023,1048,1860,1883]" italics="true" pageId="1" pageNumber="18">Ai</emphasis>
located close to anterior margin; distance between bases of setae
<emphasis id="B963EA972104D652FE14F89D504DF8C9" box="[511,584,1896,1919]" italics="true" pageId="1" pageNumber="18">AeAe</emphasis>
two times larger than distance between
<emphasis id="B963EA972104D652FBE2F89D564CF8C9" box="[1033,1097,1896,1919]" italics="true" pageId="1" pageNumber="18">AiAi</emphasis>
.
</paragraph>
<paragraph id="8BA836852104D652FF2CF87952B0F85A" blockId="1.[151,1437,1103,2028]" pageId="1" pageNumber="18">
Basal segments of chelicerae fused to each other medially (
<figureCitation id="132C2A002104D652FC60F87951D1F812" box="[907,980,1932,1957]" captionStart="FIGURES 3 10" captionStartId="3.[151,264,1952,1975]" captionTargetBox="[201,1384,199,1917]" captionTargetId="figure@3.[185,1402,193,1932]" captionTargetPageId="3" captionText="FIGURES 3 10. Hygobates nigromaculatus Lebert, 1879, larva: 3, chelicera, dorsal view; 4, chela, lateral view; 5, pedipalp, lateral view; 6, Leg I; 7, Leg II; 8, Leg III; 9, claws of leg I; 10, claws of leg III. Scale bars: 3, 6 8 = 50 µm, 4 5, 9 10 = 20 µm." httpUri="https://zenodo.org/record/808984/files/figure.png" pageId="1" pageNumber="18">Fig. 3</figureCitation>
), with slightly convex lateral margins, anterior edges more or less convex. Chela relatively small, crescentshaped, with two small subapical teeth (
<figureCitation id="132C2A002104D652FA84F84552ADF85A" captionStart="FIGURES 3 10" captionStartId="3.[151,264,1952,1975]" captionTargetBox="[201,1384,199,1917]" captionTargetId="figure@3.[185,1402,193,1932]" captionTargetPageId="3" captionText="FIGURES 3 10. Hygobates nigromaculatus Lebert, 1879, larva: 3, chelicera, dorsal view; 4, chela, lateral view; 5, pedipalp, lateral view; 6, Leg I; 7, Leg II; 8, Leg III; 9, claws of leg I; 10, claws of leg III. Scale bars: 3, 6 8 = 50 µm, 4 5, 9 10 = 20 µm." httpUri="https://zenodo.org/record/808984/files/figure.png" pageId="1" pageNumber="18">Fig. 4</figureCitation>
).
</paragraph>
<caption id="DF68660D2107D651FF7CFBD75727FB8E" box="[151,1314,1058,1081]" httpUri="https://zenodo.org/record/808982/files/figure.png" pageId="2" pageNumber="19" targetBox="[176,1396,201,1034]" targetPageId="2">
<paragraph id="8BA836852107D651FF7CFBD75727FB8E" blockId="2.[151,1314,1058,1081]" box="[151,1314,1058,1081]" pageId="2" pageNumber="19">
<emphasis id="B963EA972107D651FF7CFBD7533FFB81" bold="true" box="[151,314,1058,1081]" pageId="2" pageNumber="19">FIGURES 12.</emphasis>
<taxonomicName id="4C174D062107D651FEABFBD750DEFB8E" authority="Lebert, 1879" authorityName="Lebert" authorityYear="1879" box="[320,731,1058,1080]" class="Arachnida" family="Hygrobatidae" genus="Hygobates" kingdom="Animalia" order="Trombidiformes" pageId="2" pageNumber="19" phylum="Arthropoda" rank="species" species="nigromaculatus">
<emphasis id="B963EA972107D651FEABFBD75054FB8E" box="[320,593,1058,1080]" italics="true" pageId="2" pageNumber="19">Hygobates nigromaculatus</emphasis>
Lebert, 1879
</taxonomicName>
, larva: 1, dorsal view; 2, ventral view. Scale bar: 100 µm.
</paragraph>
</caption>
<paragraph id="8BA836852107D651FF2CFBAA5031FB52" blockId="2.[151,1437,1119,2033]" pageId="2" pageNumber="19">
Pedipalps stout (
<figureCitation id="132C2A002107D651FE6EFBAA53C9FBCE" box="[389,460,1119,1144]" captionStart="FIGURES 3 10" captionStartId="3.[151,264,1952,1975]" captionTargetBox="[201,1384,199,1917]" captionTargetId="figure@3.[185,1402,193,1932]" captionTargetPageId="3" captionText="FIGURES 3 10. Hygobates nigromaculatus Lebert, 1879, larva: 3, chelicera, dorsal view; 4, chela, lateral view; 5, pedipalp, lateral view; 6, Leg I; 7, Leg II; 8, Leg III; 9, claws of leg I; 10, claws of leg III. Scale bars: 3, 6 8 = 50 µm, 4 5, 9 10 = 20 µm." httpUri="https://zenodo.org/record/808984/files/figure.png" pageId="2" pageNumber="19">Fig. 5</figureCitation>
): P1 very short, without setae; P2 relatively large, with single dorsal seta near the middle of segment; P3 with a very long, thick lateroproximal seta and a short, fine dorsoproximal seta; P4 with two unequal setae and a massive dorsodistal claw; P5 small, with one rather long solenidion and seven unequal simple setae, one of them very short.
</paragraph>
<paragraph id="8BA836852107D651FF2CFB05510BFA20" blockId="2.[151,1437,1119,2033]" pageId="2" pageNumber="19">
Legs fivesegmented. Shape and arrangement of setae, excluding eupathidia, as shown in
<figureCitation id="132C2A002107D651FB5FFB1A571DFABE" box="[1204,1304,1263,1288]" captionStart="FIGURES 3 10" captionStartId="3.[151,264,1952,1975]" captionTargetBox="[201,1384,199,1917]" captionTargetId="figure@3.[185,1402,193,1932]" captionTargetPageId="3" captionText="FIGURES 3 10. Hygobates nigromaculatus Lebert, 1879, larva: 3, chelicera, dorsal view; 4, chela, lateral view; 5, pedipalp, lateral view; 6, Leg I; 7, Leg II; 8, Leg III; 9, claws of leg I; 10, claws of leg III. Scale bars: 3, 6 8 = 50 µm, 4 5, 9 10 = 20 µm." httpUri="https://zenodo.org/record/808984/files/figure.png" pageId="2" pageNumber="19">Figs 68</figureCitation>
. Number of thickened setae from trochanter to tarsus: IIII-Leg: 1-2-2-1-0, II: 0-2-3-4-0, III: 0-2-4-5-0. Solenidion on II-Leg-5 located near the middle of segment (
<figureCitation id="132C2A002107D651FDDBFAC35071FAF9" box="[560,628,1334,1359]" captionStart="FIGURES 3 10" captionStartId="3.[151,264,1952,1975]" captionTargetBox="[201,1384,199,1917]" captionTargetId="figure@3.[185,1402,193,1932]" captionTargetPageId="3" captionText="FIGURES 3 10. Hygobates nigromaculatus Lebert, 1879, larva: 3, chelicera, dorsal view; 4, chela, lateral view; 5, pedipalp, lateral view; 6, Leg I; 7, Leg II; 8, Leg III; 9, claws of leg I; 10, claws of leg III. Scale bars: 3, 6 8 = 50 µm, 4 5, 9 10 = 20 µm." httpUri="https://zenodo.org/record/808984/files/figure.png" pageId="2" pageNumber="19">Fig. 7</figureCitation>
), and solenidion or solenidia on other segments of legs situated distally. Claws of legs III (
<figureCitation id="132C2A002107D651FEB4FAAF53B5FAC4" box="[351,432,1370,1395]" captionStart="FIGURES 3 10" captionStartId="3.[151,264,1952,1975]" captionTargetBox="[201,1384,199,1917]" captionTargetId="figure@3.[185,1402,193,1932]" captionTargetPageId="3" captionText="FIGURES 3 10. Hygobates nigromaculatus Lebert, 1879, larva: 3, chelicera, dorsal view; 4, chela, lateral view; 5, pedipalp, lateral view; 6, Leg I; 7, Leg II; 8, Leg III; 9, claws of leg I; 10, claws of leg III. Scale bars: 3, 6 8 = 50 µm, 4 5, 9 10 = 20 µm." httpUri="https://zenodo.org/record/808984/files/figure.png" pageId="2" pageNumber="19">Fig. 10</figureCitation>
) larger than claws of legs I and II (
<figureCitation id="132C2A002107D651FCDEFAAF517CFAC4" box="[821,889,1370,1395]" captionStart="FIGURES 3 10" captionStartId="3.[151,264,1952,1975]" captionTargetBox="[201,1384,199,1917]" captionTargetId="figure@3.[185,1402,193,1932]" captionTargetPageId="3" captionText="FIGURES 3 10. Hygobates nigromaculatus Lebert, 1879, larva: 3, chelicera, dorsal view; 4, chela, lateral view; 5, pedipalp, lateral view; 6, Leg I; 7, Leg II; 8, Leg III; 9, claws of leg I; 10, claws of leg III. Scale bars: 3, 6 8 = 50 µm, 4 5, 9 10 = 20 µm." httpUri="https://zenodo.org/record/808984/files/figure.png" pageId="2" pageNumber="19">Fig. 9</figureCitation>
). Central claw and lateral claws in all legs nearly subequal in size and with two hair-like projections each.
</paragraph>
<paragraph id="8BA836852107D651FF2CFA575674F938" blockId="2.[151,1437,1119,2033]" pageId="2" pageNumber="19">
Measurements (n=10). Dorsal plate L 195240, W 180220; setae
<emphasis id="B963EA972107D651FC52FA54563AFA0E" box="[953,1087,1441,1465]" italics="true" pageId="2" pageNumber="19">Fch, Hi, Sci</emphasis>
L 5161,setae
<emphasis id="B963EA972107D651FB06FA575709FA0F" box="[1261,1292,1442,1465]" italics="true" pageId="2" pageNumber="19">Fp</emphasis>
and
<emphasis id="B963EA972107D651FAA8FA54575AFA0E" box="[1347,1375,1441,1464]" italics="true" pageId="2" pageNumber="19">Oi</emphasis>
L 9 12, setae
<emphasis id="B963EA972107D651FF14FA305311FA6A" box="[255,276,1477,1500]" italics="true" pageId="2" pageNumber="19">Vi</emphasis>
L 4855, setae
<emphasis id="B963EA972107D651FE2DFA3053E2FA6A" box="[454,487,1477,1500]" italics="true" pageId="2" pageNumber="19">Oe</emphasis>
L 5768; setae
<emphasis id="B963EA972107D651FD73FA3050BCFA6A" box="[664,697,1477,1500]" italics="true" pageId="2" pageNumber="19">He</emphasis>
and
<emphasis id="B963EA972107D651FD05FA305111FA6A" box="[750,788,1477,1500]" italics="true" pageId="2" pageNumber="19">Sce</emphasis>
L 4148; setae
<emphasis id="B963EA972107D651FC2DFA30562EFA6A" box="[966,1067,1477,1500]" italics="true" pageId="2" pageNumber="19">Li, Le, Si</emphasis>
and
<emphasis id="B963EA972107D651FB8BFA30567FFA6A" box="[1120,1146,1477,1500]" italics="true" pageId="2" pageNumber="19">Se</emphasis>
L 3245, setae
<emphasis id="B963EA972107D651FAC0FA305740FA6A" box="[1323,1349,1477,1500]" italics="true" pageId="2" pageNumber="19">Ci</emphasis>
L 145 160, setae
<emphasis id="B963EA972107D651FEE4FA1C532DF9B6" box="[271,296,1513,1536]" italics="true" pageId="2" pageNumber="19">Pi</emphasis>
and
<emphasis id="B963EA972107D651FE8BFA1C537BF9B6" box="[352,382,1513,1536]" italics="true" pageId="2" pageNumber="19">Pe</emphasis>
L 1923, setae C1 and C2 L 3542, setae C3 L 4155, setae C4 L 5762; medial margins of coxae IIII L 1 53175; gnathosomal bay L 4448; urstigma L 1922; excretory pore plate L 4245, W 8996; capitulum L 7080; basal segments of chelicerae L 6774, W 3840, chela L 1921; pedipalpal segments (P15) L: 35, 3032, 1719, 67, 56; legs segments L: ILeg-15: 2532, 3236, 3236, 3548, 4155; IILeg-15: 2838, 3539, 3548, 4452, 5161; IIILeg-15: 3842, 3842, 4148, 5761, 5771.
</paragraph>
<paragraph id="8BA836852107D651FF2CF96C5619F8D5" blockId="2.[151,1437,1119,2033]" pageId="2" pageNumber="19">
<emphasis id="B963EA972107D651FF2CF96C536CF904" bold="true" box="[199,361,1689,1714]" pageId="2" pageNumber="19">Deutonymph.</emphasis>
Idiosoma oval in shape and somewhat flattened dorsoventrally. Integument soft and finely striated. Trichobothria
<emphasis id="B963EA972107D651FE70F94B53BFF963" box="[411,442,1726,1749]" italics="true" pageId="2" pageNumber="19">Fp</emphasis>
,
<emphasis id="B963EA972107D651FE21F94853E3F962" box="[458,486,1725,1748]" italics="true" pageId="2" pageNumber="19">Oi</emphasis>
and setae
<emphasis id="B963EA972107D651FD8AF94B507FF963" box="[609,634,1726,1749]" italics="true" pageId="2" pageNumber="19">Pi</emphasis>
not associated with glandularia, other idiosomal setae associated with glandularia. Setae
<emphasis id="B963EA972107D651FE9EF91553A4F94E" box="[373,417,1760,1784]" italics="true" pageId="2" pageNumber="19">Fch</emphasis>
(
<figureCitation id="132C2A002107D651FE5CF9155008F94F" box="[439,525,1760,1785]" captionStart="FIGURES 11 16" captionStartId="4.[151,264,1198,1221]" captionTargetBox="[162,1422,197,1168]" captionTargetId="figure@4.[150,1436,193,1177]" captionTargetPageId="4" captionText="FIGURES 11 16. Hygobates nigromaculatus Lebert, 1879, deutonymph: 11, seta Fch; 12, idiosoma, ventral view; 13, pedipalp, lateral view; 14, I-Leg- 4 6; 15, IV-Leg- 4 6, 16, claw. Scale bars: 11, 13, 14 15 = 50 µm, 12 = 100 µm, 16 = 20 µm." httpUri="https://zenodo.org/record/808986/files/figure.png" pageId="2" pageNumber="19">Fig. 11</figureCitation>
) much thicker than other idiosomal setae. Anterior coxal plates with short apodemes, posteromedial margin convexity rounded (
<figureCitation id="132C2A002107D651FD18F8F65143F8AA" box="[755,838,1795,1820]" captionStart="FIGURES 11 16" captionStartId="4.[151,264,1198,1221]" captionTargetBox="[162,1422,197,1168]" captionTargetId="figure@4.[150,1436,193,1177]" captionTargetPageId="4" captionText="FIGURES 11 16. Hygobates nigromaculatus Lebert, 1879, deutonymph: 11, seta Fch; 12, idiosoma, ventral view; 13, pedipalp, lateral view; 14, I-Leg- 4 6; 15, IV-Leg- 4 6, 16, claw. Scale bars: 11, 13, 14 15 = 50 µm, 12 = 100 µm, 16 = 20 µm." httpUri="https://zenodo.org/record/808986/files/figure.png" pageId="2" pageNumber="19">Fig. 12</figureCitation>
). Coxal plate IV subtriangular in shape with rounded medial margin. Genital field with two plates, bearing two unequal acetabula (ac-2 slightly larger than ac-1) and three short, thin setae each; genital sclerite small and located between the plates.
</paragraph>
<paragraph id="8BA836852107D651FF2CF89B569EF847" blockId="2.[151,1437,1119,2033]" pageId="2" pageNumber="19">
Pedipalp moderately long (
<figureCitation id="132C2A002107D651FE17F89B5057F831" box="[508,594,1902,1927]" captionStart="FIGURES 11 16" captionStartId="4.[151,264,1198,1221]" captionTargetBox="[162,1422,197,1168]" captionTargetId="figure@4.[150,1436,193,1177]" captionTargetPageId="4" captionText="FIGURES 11 16. Hygobates nigromaculatus Lebert, 1879, deutonymph: 11, seta Fch; 12, idiosoma, ventral view; 13, pedipalp, lateral view; 14, I-Leg- 4 6; 15, IV-Leg- 4 6, 16, claw. Scale bars: 11, 13, 14 15 = 50 µm, 12 = 100 µm, 16 = 20 µm." httpUri="https://zenodo.org/record/808986/files/figure.png" pageId="2" pageNumber="19">Fig. 13</figureCitation>
): P-1 without setae, P-2 with one dorsoproximal seta and two dorsodistal setae, ventral margin straight or slightly concave distally forming a right angle, a few denticles covering distal 3/5 of ventral margin of P-2; P-3 with two dorsodistal thick subequal setae, ventral margin convex with a few denticles covering distal half of segment; P-4 ventral setae on the same level near middle of segment.
</paragraph>
<caption id="DF68660D2106D650FF7CF8555793F860" httpUri="https://zenodo.org/record/808984/files/figure.png" pageId="3" pageNumber="20" targetBox="[201,1384,199,1917]" targetPageId="3">
<paragraph id="8BA836852106D650FF7CF8555793F860" blockId="3.[151,1436,1952,2006]" pageId="3" pageNumber="20">
<emphasis id="B963EA972106D650FF7CF855534DF800" bold="true" box="[151,328,1952,1975]" pageId="3" pageNumber="20">FIGURES 310.</emphasis>
<taxonomicName id="4C174D062106D650FEA4F85450E9F801" authority="Lebert, 1879" authorityName="Lebert" authorityYear="1879" box="[335,748,1953,1975]" class="Arachnida" family="Hygrobatidae" genus="Hygobates" kingdom="Animalia" order="Trombidiformes" pageId="3" pageNumber="20" phylum="Arthropoda" rank="species" species="nigromaculatus">
<emphasis id="B963EA972106D650FEA4F8545064F801" box="[335,609,1953,1975]" italics="true" pageId="3" pageNumber="20">Hygobates nigromaculatus</emphasis>
Lebert, 1879
</taxonomicName>
, larva: 3, chelicera, dorsal view; 4, chela, lateral view; 5, pedipalp, lateral view; 6, Leg I; 7, Leg II; 8, Leg III; 9, claws of leg I; 10, claws of leg III. Scale bars: 3, 68=50 µm, 45, 910 =20 µm.
</paragraph>
</caption>
<caption id="DF68660D2101D657FF7CFB5B5791FB55" httpUri="https://zenodo.org/record/808986/files/figure.png" pageId="4" pageNumber="21" targetBox="[162,1422,197,1168]" targetPageId="4">
<paragraph id="8BA836852101D657FF7CFB5B5791FB55" blockId="4.[151,1436,1198,1251]" pageId="4" pageNumber="21">
<emphasis id="B963EA972101D657FF7CFB5B535DFB72" bold="true" box="[151,344,1198,1221]" pageId="4" pageNumber="21">FIGURES 1116.</emphasis>
<taxonomicName id="4C174D062101D657FE8FFB5B5115FB72" authority="Lebert, 1879" authorityName="Lebert" authorityYear="1879" box="[356,784,1198,1220]" class="Arachnida" family="Hygrobatidae" genus="Hygobates" kingdom="Animalia" order="Trombidiformes" pageId="4" pageNumber="21" phylum="Arthropoda" rank="species" species="nigromaculatus">
<emphasis id="B963EA972101D657FE8FFB5B507FFB72" box="[356,634,1198,1220]" italics="true" pageId="4" pageNumber="21">Hygobates nigromaculatus</emphasis>
Lebert, 1879
</taxonomicName>
, deutonymph: 11, seta
<emphasis id="B963EA972101D657FBFBFB5B5632FB72" box="[1040,1079,1198,1220]" italics="true" pageId="4" pageNumber="21">Fch</emphasis>
; 12, idiosoma, ventral view; 13, pedipalp, lateral view; 14, I-Leg-46; 15, IV-Leg-46, 16, claw. Scale bars: 11, 13, 1415 = 50 µm, 12 = 100 µm, 16 = 20 µm.
</paragraph>
</caption>
<paragraph id="8BA836852101D657FF2CFAE451D1FA23" blockId="4.[151,1437,1296,2005]" pageId="4" pageNumber="21">
Legs sixsegmented without swimming setae. I-Leg-4 with two subequal relatively long distoventral setae, I- Leg-5 with two short unequal distoventral setae (
<figureCitation id="132C2A002101D657FD21FAC1511AFAFA" box="[714,799,1332,1357]" captionStart="FIGURES 11 16" captionStartId="4.[151,264,1198,1221]" captionTargetBox="[162,1422,197,1168]" captionTargetId="figure@4.[150,1436,193,1177]" captionTargetPageId="4" captionText="FIGURES 11 16. Hygobates nigromaculatus Lebert, 1879, deutonymph: 11, seta Fch; 12, idiosoma, ventral view; 13, pedipalp, lateral view; 14, I-Leg- 4 6; 15, IV-Leg- 4 6, 16, claw. Scale bars: 11, 13, 14 15 = 50 µm, 12 = 100 µm, 16 = 20 µm." httpUri="https://zenodo.org/record/808986/files/figure.png" pageId="4" pageNumber="21">Fig. 14</figureCitation>
); IV-Leg-4 with one long and three relatively short and thick distal setae, IV-Leg-5 with one short and two long, thick setae (
<figureCitation id="132C2A002101D657FC49FAAD51F3FAC7" box="[930,1014,1368,1393]" captionStart="FIGURES 11 16" captionStartId="4.[151,264,1198,1221]" captionTargetBox="[162,1422,197,1168]" captionTargetId="figure@4.[150,1436,193,1177]" captionTargetPageId="4" captionText="FIGURES 11 16. Hygobates nigromaculatus Lebert, 1879, deutonymph: 11, seta Fch; 12, idiosoma, ventral view; 13, pedipalp, lateral view; 14, I-Leg- 4 6; 15, IV-Leg- 4 6, 16, claw. Scale bars: 11, 13, 14 15 = 50 µm, 12 = 100 µm, 16 = 20 µm." httpUri="https://zenodo.org/record/808986/files/figure.png" pageId="4" pageNumber="21">Fig. 15</figureCitation>
). Claws of all legs with long external and short internal clawlets, lamella with concave ventral margin (
<figureCitation id="132C2A002101D657FC98FA8951C2FA23" box="[883,967,1404,1429]" captionStart="FIGURES 11 16" captionStartId="4.[151,264,1198,1221]" captionTargetBox="[162,1422,197,1168]" captionTargetId="figure@4.[150,1436,193,1177]" captionTargetPageId="4" captionText="FIGURES 11 16. Hygobates nigromaculatus Lebert, 1879, deutonymph: 11, seta Fch; 12, idiosoma, ventral view; 13, pedipalp, lateral view; 14, I-Leg- 4 6; 15, IV-Leg- 4 6, 16, claw. Scale bars: 11, 13, 14 15 = 50 µm, 12 = 100 µm, 16 = 20 µm." httpUri="https://zenodo.org/record/808986/files/figure.png" pageId="4" pageNumber="21">Fig. 16</figureCitation>
).
</paragraph>
<paragraph id="8BA836852101D657FF2CFA545798F9FF" blockId="4.[151,1437,1296,2005]" pageId="4" pageNumber="21">
Measurements (n=10). Idiosoma L 240510; seta
<emphasis id="B963EA972101D657FCFDFA555147FA0E" box="[790,834,1440,1464]" italics="true" pageId="4" pageNumber="21">Fch</emphasis>
L3538; coxal plates I + capitulum mL 125155; genital plates L 4254, W 3036, genital acetabula (ac-12) L/W 1824/2430, 2430/3033; cheliceral segments L: base 100105, chela 4250; pedipalp segments (P15) L: 1719, 4854, 4245, 6572, 2127; legs segments L: I-Leg-16: 3042, 4248, 4255, 6072, 6578, 7280; II-Leg-16: 3042, 4250, 5460, 7580, 85105, 9097; III-Leg-16: 3045, 4250, 6066, 8496, 95100; IV-Leg-16: 6066, 5460, 7290, 115140, 125145, 115135.
</paragraph>
<paragraph id="8BA836852101D657FF2CF9A15624F96E" blockId="4.[151,1437,1296,2005]" pageId="4" pageNumber="21">
<emphasis id="B963EA972101D657FF2CF9A15320F9DB" bold="true" box="[199,293,1620,1645]" pageId="4" pageNumber="21">Female.</emphasis>
Dorsum similar to deutonymph. Setae
<emphasis id="B963EA972101D657FD19F9A1511BF9DA" box="[754,798,1620,1644]" italics="true" pageId="4" pageNumber="21">Fch</emphasis>
(
<figureCitation id="132C2A002101D657FCDBF9A15180F9DA" box="[816,901,1620,1645]" captionStart="FIGURES 17 23" captionStartId="5.[151,264,1424,1447]" captionTargetBox="[155,1428,223,1387]" captionTargetId="figure@5.[155,1440,215,1397]" captionTargetPageId="5" captionText="FIGURES 17 23. Hygobates nigromaculatus Lebert, 1879, female: 17, seta Fch; 18, coxal plates I IV, ventral view; 19, genital field; 20 21, genital plate; 22, pedipalp, lateral view; 23, I-Leg- 4 6. Scale bars: 17, 22 = 50 µm, 18 = 200 µm, 19 21, 23 = 100 µm." httpUri="https://zenodo.org/record/808988/files/figure.png" pageId="4" pageNumber="21">Fig. 17</figureCitation>
) thicker than others idiosomal setae. Anterior coxal plates with short apodemes, posteromedial margin usually broadly rounded (
<figureCitation id="132C2A002101D657FB81F98D56CDF926" box="[1130,1224,1656,1681]" captionStart="FIGURES 17 23" captionStartId="5.[151,264,1424,1447]" captionTargetBox="[155,1428,223,1387]" captionTargetId="figure@5.[155,1440,215,1397]" captionTargetPageId="5" captionText="FIGURES 17 23. Hygobates nigromaculatus Lebert, 1879, female: 17, seta Fch; 18, coxal plates I IV, ventral view; 19, genital field; 20 21, genital plate; 22, pedipalp, lateral view; 23, I-Leg- 4 6. Scale bars: 17, 22 = 50 µm, 18 = 200 µm, 19 21, 23 = 100 µm." httpUri="https://zenodo.org/record/808988/files/figure.png" pageId="4" pageNumber="21">Figs 18</figureCitation>
). Coxal plates IV subtriangular. Genital plates as long as or slightly shorter than gonopore, with 1522 setae each; acetabula in a triangular position, medial margin slightly indented near the centre (
<figureCitation id="132C2A002101D657FC79F9355610F96E" box="[914,1045,1728,1753]" captionStart="FIGURES 17 23" captionStartId="5.[151,264,1424,1447]" captionTargetBox="[155,1428,223,1387]" captionTargetId="figure@5.[155,1440,215,1397]" captionTargetPageId="5" captionText="FIGURES 17 23. Hygobates nigromaculatus Lebert, 1879, female: 17, seta Fch; 18, coxal plates I IV, ventral view; 19, genital field; 20 21, genital plate; 22, pedipalp, lateral view; 23, I-Leg- 4 6. Scale bars: 17, 22 = 50 µm, 18 = 200 µm, 19 21, 23 = 100 µm." httpUri="https://zenodo.org/record/808988/files/figure.png" pageId="4" pageNumber="21">Figs 1921</figureCitation>
).
</paragraph>
<paragraph id="8BA836852101D657FF2CF9115361F83A" blockId="4.[151,1437,1296,2005]" pageId="4" pageNumber="21">
P-1 with a single dorsodistal seta; P-2 usually with six (occasionally five or seven) thick dorsal setae, ventral margin straight or slightly concave distally forming a right angle, denticles covering distal 2/3 or 3/5 of ventral margin of segment; P-3 with two dorsoproximal, two dorsodistal thick subequal setae and single thin distal seta, ventral margin straight, denticles covering distal 2/3 of segment; P-4 ventral setae on the same level near middle of segment (
<figureCitation id="132C2A002101D657FEEFF881535DF83A" box="[260,344,1908,1933]" captionStart="FIGURES 17 23" captionStartId="5.[151,264,1424,1447]" captionTargetBox="[155,1428,223,1387]" captionTargetId="figure@5.[155,1440,215,1397]" captionTargetPageId="5" captionText="FIGURES 17 23. Hygobates nigromaculatus Lebert, 1879, female: 17, seta Fch; 18, coxal plates I IV, ventral view; 19, genital field; 20 21, genital plate; 22, pedipalp, lateral view; 23, I-Leg- 4 6. Scale bars: 17, 22 = 50 µm, 18 = 200 µm, 19 21, 23 = 100 µm." httpUri="https://zenodo.org/record/808988/files/figure.png" pageId="4" pageNumber="21">Fig. 22</figureCitation>
).
</paragraph>
<paragraph id="8BA836852101D657FF2CF86C53CFF862" blockId="4.[151,1437,1296,2005]" pageId="4" pageNumber="21">
Legs slender, I-Leg-4 with two unequal, rather long, distoventral setae, I-Leg-5 with two short unequal distoventral setae (
<figureCitation id="132C2A002101D657FE82F84953B8F862" box="[361,445,1980,2005]" captionStart="FIGURES 17 23" captionStartId="5.[151,264,1424,1447]" captionTargetBox="[155,1428,223,1387]" captionTargetId="figure@5.[155,1440,215,1397]" captionTargetPageId="5" captionText="FIGURES 17 23. Hygobates nigromaculatus Lebert, 1879, female: 17, seta Fch; 18, coxal plates I IV, ventral view; 19, genital field; 20 21, genital plate; 22, pedipalp, lateral view; 23, I-Leg- 4 6. Scale bars: 17, 22 = 50 µm, 18 = 200 µm, 19 21, 23 = 100 µm." httpUri="https://zenodo.org/record/808988/files/figure.png" pageId="4" pageNumber="21">Fig. 23</figureCitation>
).
</paragraph>
<caption id="DF68660D2100D656FF7CFA655325FA52" httpUri="https://zenodo.org/record/808988/files/figure.png" pageId="5" pageNumber="22" targetBox="[155,1428,223,1387]" targetPageId="5">
<paragraph id="8BA836852100D656FF7CFA655325FA52" blockId="5.[151,1435,1424,1508]" pageId="5" pageNumber="22">
<emphasis id="B963EA972100D656FF7CFA655352FA13" bold="true" box="[151,343,1424,1447]" pageId="5" pageNumber="22">FIGURES 1723.</emphasis>
<taxonomicName id="4C174D062100D656FE8AFA655103FA10" authority="Lebert, 1879" authorityName="Lebert" authorityYear="1879" box="[353,774,1424,1446]" class="Arachnida" family="Hygrobatidae" genus="Hygobates" kingdom="Animalia" order="Trombidiformes" pageId="5" pageNumber="22" phylum="Arthropoda" rank="species" species="nigromaculatus">
<emphasis id="B963EA972100D656FE8AFA655070FA10" box="[353,629,1424,1446]" italics="true" pageId="5" pageNumber="22">Hygobates nigromaculatus</emphasis>
Lebert, 1879
</taxonomicName>
, female: 17, seta
<emphasis id="B963EA972100D656FC2EFA6551E9FA10" box="[965,1004,1424,1446]" italics="true" pageId="5" pageNumber="22">Fch</emphasis>
; 18, coxal plates IIV, ventral view; 19, genital field; 2021, genital plate; 22, pedipalp, lateral view; 23, I-Leg-46. Scale bars: 17, 22 = 50 µm, 18 = 200 µm, 1921, 23 = 100 µm.
</paragraph>
</caption>
<paragraph id="8BA836852100D656FF2CF9F951E2F96E" blockId="5.[151,1436,1547,1968]" pageId="5" pageNumber="22">
Measurements (n=10). Idiosoma L 10001450; seta
<emphasis id="B963EA972100D656FCCDF9FE5157F995" box="[806,850,1547,1571]" italics="true" pageId="5" pageNumber="22">Fch</emphasis>
L3545; coxal plates I + capitulum mL 300330; genital plates L 150175, W 6590, genital acetabula (ac-13) L/W 5065/2535, 5575/3035, 6075/4255; cheliceral segments L: base 225265, chela 110125; pedipalp segments (P15) L: 3042, 108115, 8590, 145 165, 4048; legs segments L: I-Leg-16: 6275, 90115, 110140, 160210, 1 85215, 165205; II-Leg-16: 6590, 100115, 135155, 210240, 225250, 195240; III-Leg-16: 7090, 100150, 160190, 260290, 280315, 245 275; IV-Leg-16: 130165, 135165, 220250, 305350, 310350, 275330.
</paragraph>
<paragraph id="8BA836852100D656FF2CF91653DBF806" blockId="5.[151,1436,1547,1968]" pageId="5" pageNumber="22">
<emphasis id="B963EA972100D656FF2CF916530EF94A" bold="true" box="[199,267,1763,1788]" pageId="5" pageNumber="22">Male.</emphasis>
Genital plate wider than long, anterior margin with a small, knob-shaped projection, posterior margin indented, with a short rounded or bifurcated median projection, with 1728 pairs of setae; acetabula in a triangular position, in most cases ac-3 larger than ac-2 and ac-1 (
<figureCitation id="132C2A002100D656FCE2F8DE5159F8F2" box="[777,860,1835,1860]" captionStart="FIGURES 24 26" captionStartId="6.[151,264,1952,1975]" captionTargetBox="[183,1416,202,1916]" captionTargetId="figure@6.[161,1425,193,1932]" captionTargetPageId="6" captionText="FIGURES 24 26. Hygobates nigromaculatus Lebert, 1879, male: 24 25, genital plate; 26, ejaculatory complex. Scale bars: 24, 25 = 100 µm, 26 = 50 µm." httpUri="https://zenodo.org/record/808990/files/figure.png" pageId="5" pageNumber="22">Fig. 24</figureCitation>
) but occasionally all acetabula subequal (
<figureCitation id="132C2A002100D656FAD1F8DE578AF8F2" box="[1338,1423,1835,1860]" captionStart="FIGURES 24 26" captionStartId="6.[151,264,1952,1975]" captionTargetBox="[183,1416,202,1916]" captionTargetId="figure@6.[161,1425,193,1932]" captionTargetPageId="6" captionText="FIGURES 24 26. Hygobates nigromaculatus Lebert, 1879, male: 24 25, genital plate; 26, ejaculatory complex. Scale bars: 24, 25 = 100 µm, 26 = 50 µm." httpUri="https://zenodo.org/record/808990/files/figure.png" pageId="5" pageNumber="22">Fig. 25</figureCitation>
). Ejaculatory complex (
<figureCitation id="132C2A002100D656FE7AF8BA53E1F8DE" box="[401,484,1871,1896]" captionStart="FIGURES 24 26" captionStartId="6.[151,264,1952,1975]" captionTargetBox="[183,1416,202,1916]" captionTargetId="figure@6.[161,1425,193,1932]" captionTargetPageId="6" captionText="FIGURES 24 26. Hygobates nigromaculatus Lebert, 1879, male: 24 25, genital plate; 26, ejaculatory complex. Scale bars: 24, 25 = 100 µm, 26 = 50 µm." httpUri="https://zenodo.org/record/808990/files/figure.png" pageId="5" pageNumber="22">Fig. 26</figureCitation>
) with long anterior arms and comparatively short posterior ones, proximal chamber large subrectangular, with straight anterior margin, pair of rudimentary proximal horns extending to anterior margin of proximal chamber.
</paragraph>
<caption id="DF68660D2103D655FF7CF85553C2F860" httpUri="https://zenodo.org/record/808990/files/figure.png" pageId="6" pageNumber="23" targetBox="[183,1416,202,1916]" targetPageId="6">
<paragraph id="8BA836852103D655FF7CF85553C2F860" blockId="6.[151,1436,1952,2006]" pageId="6" pageNumber="23">
<emphasis id="B963EA972103D655FF7CF8555353F800" bold="true" box="[151,342,1952,1975]" pageId="6" pageNumber="23">FIGURES 2426.</emphasis>
<taxonomicName id="4C174D062103D655FEB6F85450FBF801" authority="Lebert, 1879" authorityName="Lebert" authorityYear="1879" box="[349,766,1953,1975]" class="Arachnida" family="Hygrobatidae" genus="Hygobates" kingdom="Animalia" order="Trombidiformes" pageId="6" pageNumber="23" phylum="Arthropoda" rank="species" species="nigromaculatus">
<emphasis id="B963EA972103D655FEB6F854506AF801" box="[349,623,1953,1975]" italics="true" pageId="6" pageNumber="23">Hygobates nigromaculatus</emphasis>
Lebert, 1879
</taxonomicName>
, male: 2425, genital plate; 26, ejaculatory complex. Scale bars: 24, 25 = 100 µm, 26 = 50 µm.
</paragraph>
</caption>
<paragraph id="8BA836852102D654FF2CFF6D51AAFED2" blockId="7.[151,1437,151,896]" pageId="7" pageNumber="24">
Measurements (n=10). Idiosoma L 8001150; seta
<emphasis id="B963EA972102D654FCF5FF6D514FFF06" box="[798,842,152,176]" italics="true" pageId="7" pageNumber="24">Fch</emphasis>
L3042; coxal plates I + capitulum mL 270320; genital plate L 150165, W 190225, genital acetabula (ac-13) L/W 5060/3642, 5080/3538, 5585/4250; cheliceral segments L: base 180250, chela 90105; pedipalp segments (P15) L: 3035, 95115, 6585, 125140, 4048; legs segments L: I-Leg-16: 5065, 80100, 100125, 145155, 160175, 145175; II-Leg-16: 5065, 85 100, 110140, 165215, 190225, 180215; III-Leg-16: 6590, 100115, 130170, 205250, 240290, 200250; IV-Leg-16: 125150, 120150, 175215, 235315, 265315, 230270.
</paragraph>
</subSubSection>
<subSubSection id="C30D650E2102D654FF2CFE9A52FCFC36" pageId="7" pageNumber="24" type="discussion">
<paragraph id="8BA836852102D654FF2CFE9A517EFDAE" blockId="7.[151,1437,151,896]" pageId="7" pageNumber="24">
<emphasis id="B963EA972102D654FF2CFE9A533EFE3E" bold="true" box="[199,315,367,392]" pageId="7" pageNumber="24">Remarks.</emphasis>
The morphology of the larva described by
<bibRefCitation id="EF864B742102D654FCAAFE9A5600FE3E" author="Wainstein" box="[833,1029,367,392]" pageId="7" pageNumber="24" refString="Wainstein, B. A. (1980) Opredelitel' lichinok vodyanykh kleshchey [Key to water mite larvae]. Nauka, Leningrad, 238 pp. [in Russian]" type="book" year="1980">Wainstein (1980)</bibRefCitation>
as
<taxonomicName id="4C174D062102D654FBD8FE845758FE3E" authority="Thor, 1898" authorityName="Thor" authorityYear="1898" box="[1075,1373,367,392]" class="Arachnida" family="Hygrobatidae" genus="Hygobates" kingdom="Animalia" order="Trombidiformes" pageId="7" pageNumber="24" phylum="Arthropoda" rank="species" species="longiporus">
<emphasis id="B963EA972102D654FBD8FE8456D6FE3E" box="[1075,1235,368,392]" italics="true" pageId="7" pageNumber="24">H. longiporus</emphasis>
Thor, 1898
</taxonomicName>
is in perfect agreement with the description of the larva given above. The combination of the dorsal plate broad with convex lateral margin, absent
<emphasis id="B963EA972102D654FE02FE4C501BFE66" box="[489,542,441,464]" italics="true" pageId="7" pageNumber="24">tmas</emphasis>
on the coxal plates III, subequal setae
<emphasis id="B963EA972102D654FC3EFE4D51EBFE79" box="[981,1006,440,463]" italics="true" pageId="7" pageNumber="24">Pi</emphasis>
and
<emphasis id="B963EA972102D654FBCDFE4C5646FE66" box="[1062,1091,441,464]" italics="true" pageId="7" pageNumber="24">Pe</emphasis>
and I-Leg-2/3 with two heavy setae each is diagnostic for the larva
<emphasis id="B963EA972102D654FDB4FE28513BFE42" box="[607,830,476,500]" italics="true" pageId="7" pageNumber="24">
<taxonomicName id="4C174D062102D654FDB4FE28513CFE42" authorityName="Lebert" authorityYear="1879" box="[607,825,476,500]" class="Arachnida" family="Hygrobatidae" genus="Hygobates" kingdom="Animalia" order="Trombidiformes" pageId="7" pageNumber="24" phylum="Arthropoda" rank="species" species="nigromaculatus">H. nigromaculatus</taxonomicName>
.
</emphasis>
Therefore, the larva described under the name
<taxonomicName id="4C174D062102D654FA6AFE2853DDFDAE" authority="Wainstein 1980" authorityName="Wainstein" authorityYear="1980" class="Arachnida" family="Hygrobatidae" genus="Hygobates" kingdom="Animalia" order="Trombidiformes" pageId="7" pageNumber="24" phylum="Arthropoda" rank="species" species="longiporus">
<emphasis id="B963EA972102D654FA6AFE285315FDAE" italics="true" pageId="7" pageNumber="24">H. longiporus</emphasis>
(
<bibRefCitation id="EF864B742102D654FEF5FE0A53D5FDAE" author="Wainstein" box="[286,464,511,536]" pageId="7" pageNumber="24" refString="Wainstein, B. A. (1980) Opredelitel' lichinok vodyanykh kleshchey [Key to water mite larvae]. Nauka, Leningrad, 238 pp. [in Russian]" type="book" year="1980">Wainstein 1980</bibRefCitation>
)
</taxonomicName>
probably refer to
<emphasis id="B963EA972102D654FD49FDF4517EFDAE" box="[674,891,512,536]" italics="true" pageId="7" pageNumber="24">
<taxonomicName id="4C174D062102D654FD49FDF45172FDAE" authorityName="Lebert" authorityYear="1879" box="[674,887,512,536]" class="Arachnida" family="Hygrobatidae" genus="Hygobates" kingdom="Animalia" order="Trombidiformes" pageId="7" pageNumber="24" phylum="Arthropoda" rank="species" species="nigromaculatus">H. nigromaculatus</taxonomicName>
.
</emphasis>
</paragraph>
<paragraph id="8BA836852102D654FF2CFDD15011FD33" blockId="7.[151,1437,151,896]" pageId="7" pageNumber="24">
The dorsal plate in larvae
<taxonomicName id="4C174D062102D654FE05FDD050C5FD8A" authorityName="Lebert" authorityYear="1879" box="[494,704,548,572]" class="Arachnida" family="Hygrobatidae" genus="Hygobates" kingdom="Animalia" order="Trombidiformes" pageId="7" pageNumber="24" phylum="Arthropoda" rank="species" species="nigromaculatus">
<emphasis id="B963EA972102D654FE05FDD050C5FD8A" box="[494,704,548,572]" italics="true" pageId="7" pageNumber="24">H. nigromaculatus</emphasis>
</taxonomicName>
from the Netherlands is larger, L 274321 µm and 292300 µm, respectively (
<bibRefCitation id="EF864B742102D654FEC5FDBD508EFDD6" author="Van" box="[302,651,583,608]" pageId="7" pageNumber="24" refString="Van Hezewijk, M. J. &amp; Davids, C. (1985) The larvae of three water mite species of the genus Hygrobates and their development (Acari, Hydrachnellae). Bulletin zoologisch Museum, Universiteit van Amsterdam, 10 (14), 97 - 105." type="journal article" year="1985">Van Hezewijk &amp; Davids 1985</bibRefCitation>
,
<bibRefCitation id="EF864B742102D654FD73FDB2519AFDD6" author="Martin" box="[664,927,583,608]" pageId="7" pageNumber="24" refString="Martin, P. &amp; Davids, C. (2002) Life history strategies of Hygrobates nigromaculatus, a widespread palaearctic water mite (Acari, Hydrachnidia, Hygrobatidae). In: Bernini, F., Nannelli, G., Nuccaci, G. &amp; de Lillo, E. (Eds.), Acarid Phylogeny and Evolution. Adaptations in mites and ticks. Kluwer Academic Publishers, Dordrecht, pp. 101 - 110. https: // doi. org / 10.1007 / 978 - 94 - 017 - 0611 - 7 _ 11" type="book chapter" year="2002">Martin &amp; Davids 2002</bibRefCitation>
) than Yaroslavl larvae, others morphometric data without essential differences.
</paragraph>
<paragraph id="8BA836852102D654FF7CFD7A564FFCA3" blockId="7.[151,1437,151,896]" pageId="7" pageNumber="24">
The description of the deutonymph of
<taxonomicName id="4C174D062102D654FD83FD64513AFD1E" authorityName="Lebert" authorityYear="1879" box="[616,831,656,680]" class="Arachnida" family="Hygrobatidae" genus="Hygobates" kingdom="Animalia" order="Trombidiformes" pageId="7" pageNumber="24" phylum="Arthropoda" rank="species" species="nigromaculatus">
<emphasis id="B963EA972102D654FD83FD64513AFD1E" box="[616,831,656,680]" italics="true" pageId="7" pageNumber="24">H. nigromaculatus</emphasis>
</taxonomicName>
has been made based on two specimens from the European part of
<collectingCountry id="F30076152102D654FEB2FD4153A1FD7B" box="[345,420,692,717]" name="Russia" pageId="7" pageNumber="24">Russia</collectingCountry>
and two specimens from the Asian part of
<collectingCountry id="F30076152102D654FC68FD4151CBFD7B" box="[899,974,692,717]" name="Russia" pageId="7" pageNumber="24">Russia</collectingCountry>
(
<collectingRegion id="49D3F8672102D654FC37FD41565AFD7B" box="[988,1119,692,717]" country="Russia" name="Kamchatka" pageId="7" pageNumber="24">Kamchatka</collectingRegion>
) (
<bibRefCitation id="EF864B742102D654FB9EFD41572DFD7A" author="Tuzovskij" box="[1141,1320,692,717]" pageId="7" pageNumber="24" refString="Tuzovskij, P. V. (1990) Opredelitel' deutonymphs vodyanykh kleshchey [Key to water mite deutonymphs]. Nauka, Moscow, 238 pp. [in Russian]" type="book" year="1990">Tuzovskij 1990</bibRefCitation>
). The new description is based only on European specimens. The Asian specimens differ from European specimens by the presence of swimming seta on IIIV-Leg-5 and, probably, belong to another species.
</paragraph>
<paragraph id="8BA836852102D654FF2CFCEA52FCFC36" blockId="7.[151,1437,151,896]" pageId="7" pageNumber="24">
The morphometric data of adults
<emphasis id="B963EA972102D654FDD1FCD45114FC8E" box="[570,785,800,824]" italics="true" pageId="7" pageNumber="24">
<taxonomicName id="4C174D062102D654FDD1FCD4510BFC8E" authorityName="Lebert" authorityYear="1879" box="[570,782,800,824]" class="Arachnida" family="Hygrobatidae" genus="Hygobates" kingdom="Animalia" order="Trombidiformes" pageId="7" pageNumber="24" phylum="Arthropoda" rank="species" species="nigromaculatus">H. nigromaculatus</taxonomicName>
,
</emphasis>
except of genital plate of male after Gerecke
<emphasis id="B963EA972102D654FAE7FCD45741FC8E" box="[1292,1348,800,824]" italics="true" pageId="7" pageNumber="24">et al.</emphasis>
(1960), are without essential differences from the known descriptions (
<bibRefCitation id="EF864B742102D654FC98FCB15602FCEA" author="Martin" box="[883,1031,836,861]" pageId="7" pageNumber="24" refString="Martin, P., Dabert, M. &amp; Dabert, J. (2010) Molecular evidence for species separation in the water mite Hygrobates nigromaculatus Lebert, 1879 (Acari, Hydrachnidia): evolutionary consequences of the loss of larval parasitism. Aquatic Sciences, 32, 347 - 360." type="journal article" year="2010">Martin 2010</bibRefCitation>
,
<bibRefCitation id="EF864B742102D654FBFDFCB15704FCEB" author="Gerecke" box="[1046,1281,836,861]" pageId="7" pageNumber="24" refString="Gerecke, R., Gledhill, T., Pesic, V. &amp; Smit, H. (2016) 8. Acari: Hydrachnidia III. In: Gerecke, R. (Ed.), Suβwasserfauna von Mitteleuropa, 7 / 2 - 3. Springer, Verlag, Berlin, Heidelberg, pp. 1 - 429." type="journal article" year="2016">
Gerecke
<emphasis id="B963EA972102D654FB96FCB056BCFCEA" box="[1149,1209,836,860]" italics="true" pageId="7" pageNumber="24">et al.</emphasis>
2016
</bibRefCitation>
). See general remarks.
</paragraph>
</subSubSection>
</treatment>
</document>