226 lines
31 KiB
XML
226 lines
31 KiB
XML
<document id="CDA3DFC1DD705431035ABB5D9FE5947B" ID-CLB-Dataset="26407" ID-DOI="10.11646/zootaxa.4623.2.8" ID-GBIF-Dataset="db2b3bd4-2e27-4918-8fff-226b303c65b9" ID-ISSN="1175-5326" ID-Zenodo-Dep="3255499" ID-ZooBank="EEDC9982-ED9C-40BD-8584-CDADA572DDE" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1561449672174" checkinUser="plazi" docAuthor="Bocek, Matej & Adamkova, Kristyna" docDate="2019" docId="03AF335EC3676356FF06FCFCFBA4F938" docLanguage="en" docName="zootaxa.4623.2.8.pdf" docOrigin="Zootaxa 4623 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Schizotrichalus halmaherensis Bocek & Adamkova 2019, sp. nov." docType="treatment" docUuid="C82DA8EC-26B7-4623-BF8A-D1102ABC87F2" docUuidSource="ZooBank" docVersion="10" lastPageNumber="344" masterDocId="FF964B26C3646355FF91FF95FFD3FFEB" masterDocTitle="New species of Moluccan trichaline net-winged beetles, with remarks on the phylogenetic position and distribution of Schizotrichalus (Coleoptera: Lycidae: Metriorrhynchinae)" masterLastPageNumber="350" masterPageNumber="341" pageNumber="344" updateTime="1698736235721" updateUser="ExternalLinkService">
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<mods:titleInfo id="FA203E160E085A9AD0339AB577040C5D">
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<mods:title id="05049E107B1B1D7CC91E61314C868949">New species of Moluccan trichaline net-winged beetles, with remarks on the phylogenetic position and distribution of Schizotrichalus (Coleoptera: Lycidae: Metriorrhynchinae)</mods:title>
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<mods:roleTerm id="B895B2A005B2E03A067EC4A6DC45A46C">Author</mods:roleTerm>
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<mods:namePart id="336FB6E26CD3740EB62EC1C2E17D6F23">Bocek, Matej</mods:namePart>
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<mods:namePart id="708508C8B1D902E1CE7D73D6B74CF8A2">Adamkova, Kristyna</mods:namePart>
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<mods:typeOfResource id="72E7BB07F6FF466FC041D43796D04CB3">text</mods:typeOfResource>
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<mods:title id="839A23F40C3B7F5F012FBAAF050005AC">Zootaxa</mods:title>
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<mods:part id="3C6EC45603A045E3ED03A9BD3BF701A7">
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<mods:date id="4631B7DB86FA36765EB46B0A4FC4FB3E">2019</mods:date>
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<mods:number id="B1F5DD07D1FF5E4A0CBD5709AE322AD9">2019-06-25</mods:number>
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<mods:identifier id="10F5BDA43CA78073852E5FB8DCBAAAAB" type="DOI">10.11646/zootaxa.4623.2.8</mods:identifier>
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<treatment id="03AF335EC3676356FF06FCFCFBA4F938" ID-GBIF-Taxon="158380869" LSID="urn:lsid:zoobank.org:act:C82DA8EC-26B7-4623-BF8A-D1102ABC87F2" httpUri="http://treatment.plazi.org/id/03AF335EC3676356FF06FCFCFBA4F938" lastPageNumber="344" pageId="3" pageNumber="344">
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<subSubSection id="C31CD1C3C3676356FF06FCFCFDA2FC68" box="[151,625,873,900]" pageId="3" pageNumber="344" type="nomenclature">
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<paragraph id="8BB98248C3676356FF06FCFCFDA2FC68" blockId="3.[151,983,873,964]" box="[151,625,873,900]" pageId="3" pageNumber="344">
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<heading id="D0F13524C3676356FF06FCFCFDA2FC68" bold="true" box="[151,625,873,900]" fontSize="11" level="1" pageId="3" pageNumber="344" reason="1">
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<emphasis id="B9725E5AC3676356FF06FCFCFDA2FC68" bold="true" box="[151,625,873,900]" pageId="3" pageNumber="344">
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<taxonomicName id="4C06F9CBC3676356FF06FCFCFDDFFC6F" authority="Bocek & Adamkova, 2019" authorityName="Bocek & Adamkova" authorityYear="2019" box="[151,524,873,900]" class="Insecta" family="Lycidae" genus="Schizotrichalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="344" phylum="Arthropoda" rank="species" species="halmaherensis" status="sp. nov.">
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<emphasis id="B9725E5AC3676356FF06FCFCFDDFFC6F" bold="true" box="[151,524,873,900]" italics="true" pageId="3" pageNumber="344">Schizotrichalus halmaherensis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A241E321C3676356FD82FCFCFDA2FC68" box="[531,625,873,899]" pageId="3" pageNumber="344" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C31CD1C3C3676356FF06FC18FC04FC2F" pageId="3" pageNumber="344" type="description">
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<paragraph id="8BB98248C3676356FF06FC18FE7CFC4C" blockId="3.[151,983,873,964]" box="[151,431,909,935]" pageId="3" pageNumber="344">
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<figureCitation id="133D9ECDC3676356FF06FC18FE7CFC4C" box="[151,431,909,935]" captionStart="FIGURES 2–16" captionStartId="5.[151,264,1674,1699]" captionTargetBox="[153,1429,647,1644]" captionTargetId="figure@5.[151,1436,669,1654]" captionTargetPageId="5" captionText="FIGURES 2–16. General appearance and diagnostic characters used for the species delimitation. (2–5) General appearance. (2) Schizotrichalus halmaherensis sp. nov., male; (3) ditto, female; (4) Eniclases kusyi sp. nov., male; (5) E. moluccanus, male. (6–9) Pronota (6) S. halmaherensis sp. nov., male; (7) ditto, female; (8) E. moluccanus, male; (9) E. kusyi sp. nov., male. (10–12) Male genitalia (10) S. halmaherensis sp. nov.; (11) E. kusyi sp. nov.; (12) E. moluccanus; (13–16) Basal antennomeres (13) S. halmaherensis sp. nov., male; (14) ditto, female; (15) E. kusyi sp. nov., male; (16) E. moluccanus, male. Scale bars (2–5) 2 mm; (6–16) 0.5 mm." httpUri="https://zenodo.org/record/3255503/files/figure.png" pageId="3" pageNumber="344">Figs 2–3, 6–7, 10, 13–14</figureCitation>
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</paragraph>
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<paragraph id="8BB98248C3676356FF06FC3FFC04FC2F" blockId="3.[151,983,873,964]" box="[151,983,938,964]" pageId="3" pageNumber="344">
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<uri id="FF978E4AC3676356FF06FC3FFC04FC2F" box="[151,983,938,964]" pageId="3" pageNumber="344">
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urn:lsid:zoobank.org:act:
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<uuid id="FFA0B89DC3676356FE38FC3FFC04FC2F" box="[425,983,938,964]" pageId="3" pageNumber="344">C82DA8EC-26B7-4623-BF8A-D1102ABC87F2</uuid>
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</uri>
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</paragraph>
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</subSubSection>
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<subSubSection id="C31CD1C3C3676356FF06FB99FAFEFB84" pageId="3" pageNumber="344" type="materials_examined">
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<paragraph id="8BB98248C3676356FF06FB99FAFEFB84" blockId="3.[151,1437,1036,1747]" pageId="3" pageNumber="344">
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<emphasis id="B9725E5AC3676356FF06FB99FE57FBCC" bold="true" box="[151,388,1036,1063]" pageId="3" pageNumber="344">Material examined.</emphasis>
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<materialsCitation id="3B6E8815C3676356FE1FFB98FDDFFBA0" ID-GBIF-Occurrence="2269395182" collectionCode="LMBC" collectorName="Halmahera, SE & Gn. Jailolo" country="Indonesia" elevation="460" latitude="1.0756667" location="Indonesia" longLatPrecision="13" longitude="127.4475" pageId="3" pageNumber="344" specimenCount="1" typeStatus="holotype">
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<typeStatus id="54BD3CEAC3676356FE1FFB98FE29FBCC" box="[398,506,1037,1063]" pageId="3" pageNumber="344" type="holotype">Holotype</typeStatus>
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. Male (A02461;
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<figureCitation id="133D9ECDC3676356FD50FB98FCD4FBCC" box="[705,775,1037,1063]" captionStart="FIGURES 2–16" captionStartId="5.[151,264,1674,1699]" captionTargetBox="[153,1429,647,1644]" captionTargetId="figure@5.[151,1436,669,1654]" captionTargetPageId="5" captionText="FIGURES 2–16. General appearance and diagnostic characters used for the species delimitation. (2–5) General appearance. (2) Schizotrichalus halmaherensis sp. nov., male; (3) ditto, female; (4) Eniclases kusyi sp. nov., male; (5) E. moluccanus, male. (6–9) Pronota (6) S. halmaherensis sp. nov., male; (7) ditto, female; (8) E. moluccanus, male; (9) E. kusyi sp. nov., male. (10–12) Male genitalia (10) S. halmaherensis sp. nov.; (11) E. kusyi sp. nov.; (12) E. moluccanus; (13–16) Basal antennomeres (13) S. halmaherensis sp. nov., male; (14) ditto, female; (15) E. kusyi sp. nov., male; (16) E. moluccanus, male. Scale bars (2–5) 2 mm; (6–16) 0.5 mm." httpUri="https://zenodo.org/record/3255503/files/figure.png" pageId="3" pageNumber="344">Fig. 2</figureCitation>
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),
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<collectingCountry id="F311C2D8C3676356FCB1FB98FC43FBCC" box="[800,912,1037,1063]" name="Indonesia" pageId="3" pageNumber="344">Indonesia</collectingCountry>
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,
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<collectorName id="26F3E79EC3676356FC0FFB98FB98FBCC" box="[926,1099,1037,1063]" pageId="3" pageNumber="344">Halmahera, SE</collectorName>
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slope of
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<collectorName id="26F3E79EC3676356FB2DFB98FAEFFBCC" box="[1212,1340,1037,1063]" pageId="3" pageNumber="344">Gn. Jailolo</collectorName>
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,
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<elevation id="002B657BC3676356FADAFB98FA44FBCC" box="[1355,1431,1037,1063]" metricMagnitude="2" metricUnit="m" metricValue="4.6" pageId="3" pageNumber="344" unit="m" value="460.0">
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<quantity id="4CFE2FADC3676356FADAFB98FA44FBCC" box="[1355,1431,1037,1063]" metricMagnitude="2" metricUnit="m" metricValue="4.6" pageId="3" pageNumber="344" unit="m" value="460.0">460 m</quantity>
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</elevation>
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,
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<geoCoordinate id="EE32E48FC3676356FF06FBA5FED1FBA0" box="[151,258,1072,1099]" degrees="1" direction="north" minutes="4.54" orientation="latitude" pageId="3" pageNumber="344" precision="9" value="1.0756667">1°4.54’N</geoCoordinate>
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,
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<geoCoordinate id="EE32E48FC3676356FE9EFBA5FE4DFBA0" box="[271,414,1072,1099]" degrees="127" direction="east" minutes="26.85" orientation="longitude" pageId="3" pageNumber="344" precision="9" value="127.4475">127°26.85’E</geoCoordinate>
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||
(
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<collectionCode id="ED171A8DC3676356FE3FFBA4FDD7FBA0" box="[430,516,1073,1099]" pageId="3" pageNumber="344">LMBC</collectionCode>
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)
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||
</materialsCitation>
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.
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<materialsCitation id="3B6E8815C3676356FD89FBA4FD92FB84" ID-GBIF-Occurrence="2269395184" country="Indonesia" county="Tobaru" elevation="190" latitude="0.32633334" location="Halmahera" longLatPrecision="13" longitude="127.828" municipality="Weda" pageId="3" pageNumber="344" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
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<typeStatus id="54BD3CEAC3676356FD89FBA4FD5AFBA0" box="[536,649,1073,1099]" pageId="3" pageNumber="344" type="paratype">Paratypes</typeStatus>
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.
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<specimenCount id="9D0049C1C3676356FD05FBA4FD24FBA0" box="[660,759,1073,1099]" pageId="3" pageNumber="344" type="female">1 female</specimenCount>
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(
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<accessionNumber id="94551FABC3676356FC99FBA4FCB4FBA0" box="[776,871,1073,1099]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/A02428" pageId="3" pageNumber="344" type="EnaNcbi">A02428</accessionNumber>
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,
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<figureCitation id="133D9ECDC3676356FCE2FBA4FC64FBA0" box="[883,951,1073,1099]" captionStart="FIGURES 2–16" captionStartId="5.[151,264,1674,1699]" captionTargetBox="[153,1429,647,1644]" captionTargetId="figure@5.[151,1436,669,1654]" captionTargetPageId="5" captionText="FIGURES 2–16. General appearance and diagnostic characters used for the species delimitation. (2–5) General appearance. (2) Schizotrichalus halmaherensis sp. nov., male; (3) ditto, female; (4) Eniclases kusyi sp. nov., male; (5) E. moluccanus, male. (6–9) Pronota (6) S. halmaherensis sp. nov., male; (7) ditto, female; (8) E. moluccanus, male; (9) E. kusyi sp. nov., male. (10–12) Male genitalia (10) S. halmaherensis sp. nov.; (11) E. kusyi sp. nov.; (12) E. moluccanus; (13–16) Basal antennomeres (13) S. halmaherensis sp. nov., male; (14) ditto, female; (15) E. kusyi sp. nov., male; (16) E. moluccanus, male. Scale bars (2–5) 2 mm; (6–16) 0.5 mm." httpUri="https://zenodo.org/record/3255503/files/figure.png" pageId="3" pageNumber="344">Fig. 3</figureCitation>
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),
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<collectingCountry id="F311C2D8C3676356FC5EFBA4FBECFBA0" box="[975,1087,1073,1099]" name="Indonesia" pageId="3" pageNumber="344">Indonesia</collectingCountry>
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,
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<location id="8ED9D493C3676356FBDDFBA4FB19FBA0" LSID="urn:lsid:plazi:treatment:03AF335EC3676356FF06FCFCFBA4F938:8ED9D493C3676356FBDDFBA4FB19FBA0" box="[1100,1226,1073,1099]" country="Indonesia" county="Tobaru" latitude="0.32633334" longLatPrecision="13" longitude="127.828" municipality="Weda" name="Halmahera" pageId="3" pageNumber="344">Halmahera</location>
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, road
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<collectingMunicipality id="6BDD1832C3676356FA81FBA4FA9CFBA0" box="[1296,1359,1073,1099]" pageId="3" pageNumber="344">Weda</collectingMunicipality>
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to
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<collectingCounty id="62D8FAC4C3676356FAE7FBA4FF1FFB84" pageId="3" pageNumber="344">Tobaru</collectingCounty>
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,
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<elevation id="002B657BC3676356FF46FBC0FEF3FB84" box="[215,288,1109,1135]" metricMagnitude="2" metricUnit="m" metricValue="1.9" pageId="3" pageNumber="344" unit="m" value="190.0">
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<quantity id="4CFE2FADC3676356FF46FBC0FEF3FB84" box="[215,288,1109,1135]" metricMagnitude="2" metricUnit="m" metricValue="1.9" pageId="3" pageNumber="344" unit="m" value="190.0">190 m</quantity>
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</elevation>
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,
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<geoCoordinate id="EE32E48FC3676356FEBDFBC1FE77FB84" box="[300,420,1108,1135]" degrees="0" direction="north" minutes="19.58" orientation="latitude" pageId="3" pageNumber="344" precision="9" value="0.32633334">0°19.58’N</geoCoordinate>
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,
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<geoCoordinate id="EE32E48FC3676356FE3EFBC0FD92FB84" box="[431,577,1109,1135]" degrees="127" direction="east" minutes="49.68" orientation="longitude" pageId="3" pageNumber="344" precision="9" value="127.828">127°49.68’E</geoCoordinate>
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||
</materialsCitation>
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||
,
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<materialsCitation id="3B6E8815C3676356FDDDFBC0FAFAFB84" ID-GBIF-Occurrence="2269395183" box="[588,1321,1108,1135]" collectionCode="LMBC" collectorName="Halmahera, SE & Gn. Jailolo" country="Indonesia" elevation="460" latitude="1.0756667" location="Indonesia" longLatPrecision="13" longitude="127.4475" pageId="3" pageNumber="344" specimenCount="2" specimenCount-male="2" typeStatus="paratype">
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||
<specimenCount id="9D0049C1C3676356FDDDFBC0FD71FB84" box="[588,674,1109,1135]" pageId="3" pageNumber="344" type="male">2 males</specimenCount>
|
||
(A02465–66), the same locality as the holotype (
|
||
<collectionCode id="ED171A8DC3676356FB5DFBC0FAF1FB84" box="[1228,1314,1109,1135]" pageId="3" pageNumber="344">LMBC</collectionCode>
|
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)
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||
</materialsCitation>
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||
.
|
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</paragraph>
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||
</subSubSection>
|
||
<subSubSection id="C31CD1C3C3676356FF56FBEDFB41FAAC" pageId="3" pageNumber="344" type="diagnosis">
|
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<paragraph id="8BB98248C3676356FF56FBEDFB41FAAC" blockId="3.[151,1437,1036,1747]" pageId="3" pageNumber="344">
|
||
<emphasis id="B9725E5AC3676356FF56FBEDFE91FB78" bold="true" box="[199,322,1144,1171]" pageId="3" pageNumber="344">Diagnosis.</emphasis>
|
||
<taxonomicName id="4C06F9CBC3676356FED6FBECFD4FFB78" authorityName="Bocek & Adamkova" authorityYear="2019" box="[327,668,1145,1171]" class="Insecta" family="Lycidae" genus="Schizotrichalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="344" phylum="Arthropoda" rank="species" species="halmaherensis">
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<emphasis id="B9725E5AC3676356FED6FBECFD4FFB78" box="[327,668,1145,1171]" italics="true" pageId="3" pageNumber="344">Schizotrichalus halmaherensis</emphasis>
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</taxonomicName>
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is the only Halmaheran species of the genus and superficially resembles
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<taxonomicName id="4C06F9CBC3676356FF5BFB08FE11FB5C" authorityName="Kleine" authorityYear="1930" box="[202,450,1181,1207]" class="Insecta" family="Lycidae" genus="Eniclases" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="344" phylum="Arthropoda" rank="species" species="moluccanus">
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<emphasis id="B9725E5AC3676356FF5BFB08FE11FB5C" box="[202,450,1181,1207]" italics="true" pageId="3" pageNumber="344">Eniclases moluccanus</emphasis>
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</taxonomicName>
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, possibly a member of the same local mimetic complex.
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<taxonomicName id="4C06F9CBC3676356FBA7FB08FB20FB5C" authorityName="Bocek & Adamkova" authorityYear="2019" box="[1078,1267,1181,1207]" class="Insecta" family="Lycidae" genus="Schizotrichalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="344" phylum="Arthropoda" rank="species" species="halmaherensis">
|
||
<emphasis id="B9725E5AC3676356FBA7FB08FB20FB5C" box="[1078,1267,1181,1207]" italics="true" pageId="3" pageNumber="344">S. halmaherensis</emphasis>
|
||
</taxonomicName>
|
||
differs from
|
||
<taxonomicName id="4C06F9CBC3676356FA15FB0BFECEFB30" authorityName="Kleine" authorityYear="1930" class="Insecta" family="Lycidae" genus="Eniclases" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="344" phylum="Arthropoda" rank="species" species="moluccanus">
|
||
<emphasis id="B9725E5AC3676356FA15FB0BFECEFB30" italics="true" pageId="3" pageNumber="344">E. moluccanus</emphasis>
|
||
</taxonomicName>
|
||
in characters used for the generic definition: the presence of five pronotal areoles (
|
||
<figureCitation id="133D9ECDC3676356FB25FB54FACAFB30" box="[1204,1305,1216,1243]" captionStart="FIGURES 2–16" captionStartId="5.[151,264,1674,1699]" captionTargetBox="[153,1429,647,1644]" captionTargetId="figure@5.[151,1436,669,1654]" captionTargetPageId="5" captionText="FIGURES 2–16. General appearance and diagnostic characters used for the species delimitation. (2–5) General appearance. (2) Schizotrichalus halmaherensis sp. nov., male; (3) ditto, female; (4) Eniclases kusyi sp. nov., male; (5) E. moluccanus, male. (6–9) Pronota (6) S. halmaherensis sp. nov., male; (7) ditto, female; (8) E. moluccanus, male; (9) E. kusyi sp. nov., male. (10–12) Male genitalia (10) S. halmaherensis sp. nov.; (11) E. kusyi sp. nov.; (12) E. moluccanus; (13–16) Basal antennomeres (13) S. halmaherensis sp. nov., male; (14) ditto, female; (15) E. kusyi sp. nov., male; (16) E. moluccanus, male. Scale bars (2–5) 2 mm; (6–16) 0.5 mm." httpUri="https://zenodo.org/record/3255503/files/figure.png" pageId="3" pageNumber="344">Figs 5–8</figureCitation>
|
||
), characteristic prolonged, slender phallus without extensive membranous sac (
|
||
<figureCitation id="133D9ECDC3676356FC03FB70FBC2FB14" box="[914,1041,1253,1279]" captionStart="FIGURES 2–16" captionStartId="5.[151,264,1674,1699]" captionTargetBox="[153,1429,647,1644]" captionTargetId="figure@5.[151,1436,669,1654]" captionTargetPageId="5" captionText="FIGURES 2–16. General appearance and diagnostic characters used for the species delimitation. (2–5) General appearance. (2) Schizotrichalus halmaherensis sp. nov., male; (3) ditto, female; (4) Eniclases kusyi sp. nov., male; (5) E. moluccanus, male. (6–9) Pronota (6) S. halmaherensis sp. nov., male; (7) ditto, female; (8) E. moluccanus, male; (9) E. kusyi sp. nov., male. (10–12) Male genitalia (10) S. halmaherensis sp. nov.; (11) E. kusyi sp. nov.; (12) E. moluccanus; (13–16) Basal antennomeres (13) S. halmaherensis sp. nov., male; (14) ditto, female; (15) E. kusyi sp. nov., male; (16) E. moluccanus, male. Scale bars (2–5) 2 mm; (6–16) 0.5 mm." httpUri="https://zenodo.org/record/3255503/files/figure.png" pageId="3" pageNumber="344">Figs 10, 12</figureCitation>
|
||
) and the males of
|
||
<taxonomicName id="4C06F9CBC3676356FB4FFB70FA4FFB14" authorityName="Bocek & Adamkova" authorityYear="2019" box="[1246,1436,1253,1279]" class="Insecta" family="Lycidae" genus="Schizotrichalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="344" phylum="Arthropoda" rank="species" species="halmaherensis">
|
||
<emphasis id="B9725E5AC3676356FB4FFB70FA4FFB14" box="[1246,1436,1253,1279]" italics="true" pageId="3" pageNumber="344">S. halmaherensis</emphasis>
|
||
</taxonomicName>
|
||
have flabellate antennae (
|
||
<figureCitation id="133D9ECDC3676356FE24FA9CFD49FACF" box="[437,666,1289,1316]" captionStart="FIGURES 2–16" captionStartId="5.[151,264,1674,1699]" captionTargetBox="[153,1429,647,1644]" captionTargetId="figure@5.[151,1436,669,1654]" captionTargetPageId="5" captionText="FIGURES 2–16. General appearance and diagnostic characters used for the species delimitation. (2–5) General appearance. (2) Schizotrichalus halmaherensis sp. nov., male; (3) ditto, female; (4) Eniclases kusyi sp. nov., male; (5) E. moluccanus, male. (6–9) Pronota (6) S. halmaherensis sp. nov., male; (7) ditto, female; (8) E. moluccanus, male; (9) E. kusyi sp. nov., male. (10–12) Male genitalia (10) S. halmaherensis sp. nov.; (11) E. kusyi sp. nov.; (12) E. moluccanus; (13–16) Basal antennomeres (13) S. halmaherensis sp. nov., male; (14) ditto, female; (15) E. kusyi sp. nov., male; (16) E. moluccanus, male. Scale bars (2–5) 2 mm; (6–16) 0.5 mm." httpUri="https://zenodo.org/record/3255503/files/figure.png" pageId="3" pageNumber="344">Figs 2–3, 13–14, 16</figureCitation>
|
||
). Papuan congeneric,
|
||
<emphasis id="B9725E5AC3676356FC03FA9CFBF8FAC8" box="[914,1067,1289,1315]" italics="true" pageId="3" pageNumber="344">
|
||
<taxonomicName id="4C06F9CBC3676356FC03FA9CFBF4FAC8" authorityName="Waterhouse" authorityYear="1879" box="[914,1063,1289,1315]" class="Insecta" family="Lycidae" genus="Trichalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="344" phylum="Arthropoda" rank="species" species="nigrescens">S. nigrescens</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
differs from
|
||
<taxonomicName id="4C06F9CBC3676356FB50FA9CFAACFAC8" authorityName="Bocek & Adamkova" authorityYear="2019" box="[1217,1407,1289,1315]" class="Insecta" family="Lycidae" genus="Schizotrichalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="344" phylum="Arthropoda" rank="species" species="halmaherensis">
|
||
<emphasis id="B9725E5AC3676356FB50FA9CFAACFAC8" box="[1217,1407,1289,1315]" italics="true" pageId="3" pageNumber="344">S. halmaherensis</emphasis>
|
||
</taxonomicName>
|
||
in serrate male antennae and uniform dark brown coloration (
|
||
<bibRefCitation id="EF97FFB9C3676356FCB9FAB8FC23FAAC" author="Waterhouse, C. O." box="[808,1008,1325,1351]" pageId="3" pageNumber="344" refId="ref7129" refString="Waterhouse, C. O. (1879) Illustration of the typical specimens of Coleoptera in the collection of the British Museum. Part I. - Lycidae. British Museum, London, 93 pp." type="book" year="1879">Waterhouse 1879</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EF97FFB9C3676356FC6AFAB8FB55FAAC" author="Bocak, L." box="[1019,1158,1325,1351]" pageId="3" pageNumber="344" pagination="315 - 351" refId="ref5210" refString="Bocak, L. (2002) Revision and phylogenetic analysis of Metriorrhynchinae. European Journal of Entomology, 99, 315 - 351. https: // doi. org / 10.14411 / eje. 2002.043" type="journal article" year="2002">Bocak 2002</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31CD1C3C3676356FF56FAC5FB36F9A8" pageId="3" pageNumber="344" type="description">
|
||
<paragraph id="8BB98248C3676356FF56FAC5FB36F9A8" blockId="3.[151,1437,1036,1747]" pageId="3" pageNumber="344">
|
||
<emphasis id="B9725E5AC3676356FF56FAC5FE8AFA80" bold="true" box="[199,345,1360,1387]" pageId="3" pageNumber="344">Description.</emphasis>
|
||
Male. Body length
|
||
<quantity id="4CFE2FADC3676356FDA5FAC5FD19FA80" box="[564,714,1360,1387]" metricMagnitude="-2" metricUnit="m" metricValue="1.035" metricValueMax="1.1199999999999999" metricValueMin="0.95" pageId="3" pageNumber="344" unit="mm" value="10.35" valueMax="11.2" valueMin="9.5">9.5–11.2 mm</quantity>
|
||
. Head, antennae and legs completely black, pronotum orange, almost half of elytra orange with sharp transition between brightly colored humeral half and black colored apex. Head small with hemispherically prominent eyes, their diameter 1.03 times interocular distance, antennomeres 3–10 flabellate with long lamellae (
|
||
<figureCitation id="133D9ECDC3676356FE57FA28FDCAFA3C" box="[454,537,1469,1495]" captionStart="FIGURES 2–16" captionStartId="5.[151,264,1674,1699]" captionTargetBox="[153,1429,647,1644]" captionTargetId="figure@5.[151,1436,669,1654]" captionTargetPageId="5" captionText="FIGURES 2–16. General appearance and diagnostic characters used for the species delimitation. (2–5) General appearance. (2) Schizotrichalus halmaherensis sp. nov., male; (3) ditto, female; (4) Eniclases kusyi sp. nov., male; (5) E. moluccanus, male. (6–9) Pronota (6) S. halmaherensis sp. nov., male; (7) ditto, female; (8) E. moluccanus, male; (9) E. kusyi sp. nov., male. (10–12) Male genitalia (10) S. halmaherensis sp. nov.; (11) E. kusyi sp. nov.; (12) E. moluccanus; (13–16) Basal antennomeres (13) S. halmaherensis sp. nov., male; (14) ditto, female; (15) E. kusyi sp. nov., male; (16) E. moluccanus, male. Scale bars (2–5) 2 mm; (6–16) 0.5 mm." httpUri="https://zenodo.org/record/3255503/files/figure.png" pageId="3" pageNumber="344">Fig. 13</figureCitation>
|
||
). Pronotum trapezoidal, wider than long, with five areoles and two characteristic bulges on each of its lateral margins (
|
||
<figureCitation id="133D9ECDC3676356FDA9FA74FD4DFA17" box="[568,670,1505,1532]" captionStart="FIGURES 2–16" captionStartId="5.[151,264,1674,1699]" captionTargetBox="[153,1429,647,1644]" captionTargetId="figure@5.[151,1436,669,1654]" captionTargetPageId="5" captionText="FIGURES 2–16. General appearance and diagnostic characters used for the species delimitation. (2–5) General appearance. (2) Schizotrichalus halmaherensis sp. nov., male; (3) ditto, female; (4) Eniclases kusyi sp. nov., male; (5) E. moluccanus, male. (6–9) Pronota (6) S. halmaherensis sp. nov., male; (7) ditto, female; (8) E. moluccanus, male; (9) E. kusyi sp. nov., male. (10–12) Male genitalia (10) S. halmaherensis sp. nov.; (11) E. kusyi sp. nov.; (12) E. moluccanus; (13–16) Basal antennomeres (13) S. halmaherensis sp. nov., male; (14) ditto, female; (15) E. kusyi sp. nov., male; (16) E. moluccanus, male. Scale bars (2–5) 2 mm; (6–16) 0.5 mm." httpUri="https://zenodo.org/record/3255503/files/figure.png" pageId="3" pageNumber="344">Figs 6–7</figureCitation>
|
||
). Elytra with nine costae at humeri, primary costa 1 short, secondary costae well-developed, elytral cells subquadrate, regular. Phallus long, very slender, mostly membranous, with cuplike apex, phallobase without visible membranous sac. Female. Antennae acutely serrate (
|
||
<figureCitation id="133D9ECDC3676356FB15F9BCFB0BF9A8" box="[1156,1240,1577,1603]" captionStart="FIGURES 2–16" captionStartId="5.[151,264,1674,1699]" captionTargetBox="[153,1429,647,1644]" captionTargetId="figure@5.[151,1436,669,1654]" captionTargetPageId="5" captionText="FIGURES 2–16. General appearance and diagnostic characters used for the species delimitation. (2–5) General appearance. (2) Schizotrichalus halmaherensis sp. nov., male; (3) ditto, female; (4) Eniclases kusyi sp. nov., male; (5) E. moluccanus, male. (6–9) Pronota (6) S. halmaherensis sp. nov., male; (7) ditto, female; (8) E. moluccanus, male; (9) E. kusyi sp. nov., male. (10–12) Male genitalia (10) S. halmaherensis sp. nov.; (11) E. kusyi sp. nov.; (12) E. moluccanus; (13–16) Basal antennomeres (13) S. halmaherensis sp. nov., male; (14) ditto, female; (15) E. kusyi sp. nov., male; (16) E. moluccanus, male. Scale bars (2–5) 2 mm; (6–16) 0.5 mm." httpUri="https://zenodo.org/record/3255503/files/figure.png" pageId="3" pageNumber="344">Fig. 14</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31CD1C3C3676356FF56F9D9FD04F98C" box="[199,727,1612,1639]" pageId="3" pageNumber="344" type="distribution">
|
||
<paragraph id="8BB98248C3676356FF56F9D9FD04F98C" blockId="3.[151,1437,1036,1747]" box="[199,727,1612,1639]" pageId="3" pageNumber="344">
|
||
<emphasis id="B9725E5AC3676356FF56F9D9FDD1F98C" bold="true" box="[199,514,1612,1639]" pageId="3" pageNumber="344">Distribution and diversity.</emphasis>
|
||
Halmahera Island.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31CD1C3C3676356FF56F9E4FDD6F944" pageId="3" pageNumber="344" type="description">
|
||
<paragraph id="8BB98248C3676356FF56F9E4FDD6F944" blockId="3.[151,1437,1036,1747]" pageId="3" pageNumber="344">
|
||
<emphasis id="B9725E5AC3676356FF56F9E4FEADF960" bold="true" box="[199,382,1649,1675]" pageId="3" pageNumber="344">Measurements.</emphasis>
|
||
BL
|
||
<quantity id="4CFE2FADC3676356FE20F9E5FD9BF960" box="[433,584,1648,1675]" metricMagnitude="-2" metricUnit="m" metricValue="1.035" metricValueMax="1.1199999999999999" metricValueMin="0.95" pageId="3" pageNumber="344" unit="mm" value="10.35" valueMax="11.2" valueMin="9.5">9.5–11.2 mm</quantity>
|
||
, EL
|
||
<quantity id="4CFE2FADC3676356FDEEF9E4FCDAF960" box="[639,777,1649,1675]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" metricValueMax="8.2" metricValueMin="7.8" pageId="3" pageNumber="344" unit="mm" value="8.0" valueMax="8.2" valueMin="7.8">7.8–8.2 mm</quantity>
|
||
, WH
|
||
<quantity id="4CFE2FADC3676356FCDCF9E5FC04F960" box="[845,983,1648,1675]" metricMagnitude="-3" metricUnit="m" metricValue="3.4" metricValueMax="3.5" metricValueMin="3.3" pageId="3" pageNumber="344" unit="mm" value="3.4" valueMax="3.5" valueMin="3.3">3.3–3.5 mm</quantity>
|
||
, PL
|
||
<quantity id="4CFE2FADC3676356FB9DF9E4FB61F960" box="[1036,1202,1649,1675]" metricMagnitude="-3" metricUnit="m" metricValue="1.815" metricValueMax="1.84" metricValueMin="1.79" pageId="3" pageNumber="344" unit="mm" value="1.815" valueMax="1.84" valueMin="1.79">1.79–1.84 mm</quantity>
|
||
, PW
|
||
<quantity id="4CFE2FADC3676356FB60F9E5FA4BF960" box="[1265,1432,1648,1675]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" metricValueMax="2.53" metricValueMin="2.47" pageId="3" pageNumber="344" unit="mm" value="2.5" valueMax="2.53" valueMin="2.47">2.47–2.53 mm</quantity>
|
||
, Edist
|
||
<quantity id="4CFE2FADC3676356FF49F900FEEDF944" box="[216,318,1685,1711]" metricMagnitude="-4" metricUnit="m" metricValue="7.4" pageId="3" pageNumber="344" unit="mm" value="0.74">0.74 mm</quantity>
|
||
, Ediam
|
||
<quantity id="4CFE2FADC3676356FE08F900FDD3F944" box="[409,512,1685,1711]" metricMagnitude="-4" metricUnit="m" metricValue="7.6" pageId="3" pageNumber="344" unit="mm" value="0.76">0.76 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31CD1C3C3676356FF56F92CFBA4F938" box="[199,1143,1721,1747]" pageId="3" pageNumber="344" type="etymology">
|
||
<paragraph id="8BB98248C3676356FF56F92CFBA4F938" blockId="3.[151,1437,1036,1747]" box="[199,1143,1721,1747]" pageId="3" pageNumber="344">
|
||
<emphasis id="B9725E5AC3676356FF56F92CFE9DF938" bold="true" box="[199,334,1721,1747]" pageId="3" pageNumber="344">Etymology.</emphasis>
|
||
The specific name refers to Halmahera, the
|
||
<typeStatus id="54BD3CEAC3676356FCAFF92CFCBDF938" box="[830,878,1721,1747]" pageId="3" pageNumber="344">type</typeStatus>
|
||
locality of this species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |