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<document id="C392204EFA097332F8FED78A4269080A" ID-CLB-Dataset="54546" ID-DOI="10.5852/ejt.2022.821.1799" ID-GBIF-Dataset="62057142-b60a-43c5-b7cd-947fc7615635" ID-ISSN="2118-9773" ID-Zenodo-Dep="6595468" ID-ZooBank="C85A43AA-6753-4851-8CEE-88AAE6DF069B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1653931612537" checkinUser="felipe" docAuthor="Ranasinghe, Sasanka, Eberle, Jonas, Athukorala, Namal, Benjamin, Suresh P. &amp; Ahrens, Dirk" docDate="2022" docId="03A99E29FF973067FDA5E9684D3A1332" docLanguage="en" docName="ejt-821_57-101.pdf" docOrigin="European Journal of Taxonomy 821 (1)" docSource="http://zoobank.org/c85a43aa-6753-4851-8cee-88aae6df069b" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Maladera deenstana Ranasinghe &amp; Eberle &amp; Athukorala &amp; Benjamin &amp; Ahrens 2022, sp. nov." docType="treatment" docUuid="054FA7C8-DD22-49C4-8C43-B41E610ADDF6" docUuidSource="ZooBank" docVersion="7" lastPageNumber="88" masterDocId="FF90E651FF8B3078FFC8EC494C7C155C" masterDocTitle="New species of Sericini from Sri Lanka (Coleoptera, Scarabaeidae). Part II" masterLastPageNumber="101" masterPageNumber="57" pageNumber="85" updateTime="1699338309283" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="6B1D887D78DC14804B830F3F019E7495">New species of Sericini from Sri Lanka (Coleoptera, Scarabaeidae). Part II</mods:title>
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<mods:namePart id="541255F31DB136FC8292C1E8C357A318">Ranasinghe, Sasanka</mods:namePart>
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<mods:affiliation id="4B23727C1277BC46C64514A6F08ADC7F">Zoologisches Forschungsmuseum A. Koenig, Adenauerallee 127, 53113 Bonn, Germany.</mods:affiliation>
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<mods:namePart id="C851D35C2F3EE6A697E10F11C550A537">Eberle, Jonas</mods:namePart>
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<mods:affiliation id="4AB8CCF067AC05723D2B388258F3ED04">Zoologisches Forschungsmuseum A. Koenig, Adenauerallee 127, 53113 Bonn, Germany.</mods:affiliation>
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<mods:namePart id="FDD0FC2EF01F0572A7969A361F8DBB53">Athukorala, Namal</mods:namePart>
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<mods:affiliation id="60E012E640A7057421C7C29546A2D0B4">National Institute of Fundamental Studies, Hantana Road, 20000 Kandy, Sri Lanka.</mods:affiliation>
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<mods:namePart id="4C91CAE6B4F7A69A24A1D8916FBA6EA3">Benjamin, Suresh P.</mods:namePart>
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<mods:affiliation id="2A0CBC1E5BC8B563CDAE93DE219E489C">Zoologisches Forschungsmuseum A. Koenig, Adenauerallee 127, 53113 Bonn, Germany. &amp; National Institute of Fundamental Studies, Hantana Road, 20000 Kandy, Sri Lanka.</mods:affiliation>
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<mods:namePart id="7A2547EB2CBAE03F2B3BB93629FCF7E1">Ahrens, Dirk</mods:namePart>
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<mods:affiliation id="FE0C796D165C5700F25032A4D072F6A1">Zoologisches Forschungsmuseum A. Koenig, Adenauerallee 127, 53113 Bonn, Germany.</mods:affiliation>
<mods:nameIdentifier id="B564E95064A4A700C33350E7B57E8F3A" type="email">ahrens.dirk_col@gmx.de,d.ahrens@leibniz-zfmk.de</mods:nameIdentifier>
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<mods:title id="46E3EE8C858C777A1A4F79EBE8A20546">European Journal of Taxonomy</mods:title>
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<mods:date id="E1E261E82AA3F6BB7341E1E5627C294C">2022</mods:date>
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<treatment id="03A99E29FF973067FDA5E9684D3A1332" ID-DOI="http://doi.org/10.5281/zenodo.7473387" ID-GBIF-Taxon="207927100" ID-Zenodo-Dep="7473387" LSID="urn:lsid:zoobank.org:act:054FA7C8-DD22-49C4-8C43-B41E610ADDF6" httpUri="http://treatment.plazi.org/id/03A99E29FF973067FDA5E9684D3A1332" lastPageId="31" lastPageNumber="88" pageId="28" pageNumber="85">
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<paragraph id="8BBF2F3FFF973064FDA5E9684FBA1060" blockId="28.[361,1226,1313,1411]" box="[621,966,1313,1340]" pageId="28" pageNumber="85">
<heading id="D0F79853FF973064FDA5E9684FBA1060" box="[621,966,1313,1340]" centered="true" fontSize="11" level="2" pageId="28" pageNumber="85" reason="2">
<taxonomicName id="4C0054BCFF973064FDA5E9684F181060" authority="Ranasinghe &amp; Eberle &amp; Athukorala &amp; Benjamin &amp; Ahrens, 2022" authorityName="Ranasinghe &amp; Eberle &amp; Athukorala &amp; Benjamin &amp; Ahrens" authorityYear="2022" box="[621,868,1313,1340]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="85" phylum="Arthropoda" rank="species" species="deenstana" status="sp. nov.">
<emphasis id="B974F32DFF973064FDA5E9684F181060" bold="true" box="[621,868,1313,1340]" italics="true" pageId="28" pageNumber="85">Maladera deenstana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2474E56FF973064FCA4E96B4FBA1060" box="[876,966,1314,1340]" pageId="28" pageNumber="85" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
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<subSubSection id="C31A7CB4FF973064FEA1E90C4FFD10DE" pageId="28" pageNumber="85" type="description">
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</uri>
</paragraph>
<paragraph id="8BBF2F3FFF973064FD7BE9214FFD10DE" blockId="28.[361,1226,1313,1411]" box="[691,897,1384,1411]" pageId="28" pageNumber="85">
<figureCitation id="133B33BAFF973064FD7BE9214F4810DE" box="[691,820,1384,1410]" captionStart="Fig" captionStartId="27.[189,232,1921,1947]" captionTargetBox="[204,1388,789,1860]" captionTargetId="figure-195@27.[189,1398,770,1883]" captionTargetPageId="27" captionText="Fig. 6. AD. Maladera dambullana sp. nov., holotype, ♂ (ZFMK). EH. M. deenstana sp. nov., holotype, ♂ (ZFMK). A, E. Aedeagus, left side lateral view. C, G. Aedeagus, right side lateral view. B, F. Parameres, dorsal view. D, H. Habitus (not to scale). Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6595481" httpUri="https://zenodo.org/record/6595481/files/figure.png" pageId="28" pageNumber="85">Figs 6EH</figureCitation>
,
<figureCitation id="133B33BAFF973064FC88E9204F2010DF" box="[832,860,1385,1411]" captionStart="Fig" captionStartId="29.[189,232,1917,1943]" captionTargetBox="[189,1398,1076,1874]" captionTargetId="figure-244@29.[189,1398,1076,1874]" captionTargetPageId="29" captionText="Fig. 7. Distribution of ten new species. A. Selaserica fabriziae sp. nov. B. Sel. sororinitida sp. nov. C. Neoserica pophami sp. nov. D. Maladera haniel sp. nov. E. M. kishi sp. nov. F. M. windy sp. nov. G. M. karunaratnae sp. nov. H. M. hiyarensis sp. nov. I. M. dambullana sp. nov. J. M. deenstana sp. nov." figureDoi="http://doi.org/10.5281/zenodo.6595483" httpUri="https://zenodo.org/record/6595483/files/figure.png" pageId="28" pageNumber="85">7J</figureCitation>
,
<figureCitation id="133B33BAFF973064FCA0E9214FFD10DE" box="[872,897,1384,1410]" captionStart="Fig" captionStartId="30.[189,232,1885,1911]" captionTargetBox="[235,1353,265,1847]" captionTargetId="figure-16@30.[235,1353,265,1847]" captionTargetPageId="30" captionText="Fig. 8. Photographs of the habitats of the new species.A. Selaserica fabriziae sp. nov. B. Sel. sororinitida sp. nov. C. Neoserica pophami sp. nov. D. Maladera haniel sp. nov. and M. kishi sp. nov. E. M. windy sp. nov. F. M. karunaratnae sp. nov. G. M. hiyarensis sp. nov.H. M. dambullana sp. nov. I. M. deenstana sp. nov." figureDoi="http://doi.org/10.5281/zenodo.6595485" httpUri="https://zenodo.org/record/6595485/files/figure.png" pageId="28" pageNumber="85">8I</figureCitation>
</paragraph>
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<paragraph id="8BBF2F3FFF973064FF75E9E04D4A109F" blockId="28.[189,310,1449,1475]" box="[189,310,1449,1475]" pageId="28" pageNumber="85">
<heading id="D0F79853FF973064FF75E9E04D4A109F" bold="true" box="[189,310,1449,1475]" fontSize="11" level="3" pageId="28" pageNumber="85" reason="3">
<emphasis id="B974F32DFF973064FF75E9E04D4A109F" bold="true" box="[189,310,1449,1475]" pageId="28" pageNumber="85">Diagnosis</emphasis>
</heading>
</paragraph>
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<taxonomicName id="4C0054BCFF973064FF75E9914DCF10AE" authority="Ranasinghe &amp; Eberle &amp; Athukorala &amp; Benjamin &amp; Ahrens, 2022" authorityName="Ranasinghe &amp; Eberle &amp; Athukorala &amp; Benjamin &amp; Ahrens" authorityYear="2022" box="[189,435,1496,1522]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="85" phylum="Arthropoda" rank="species" species="deenstana" status="sp. nov.">
<emphasis id="B974F32DFF973064FF75E9914DCF10AE" box="[189,435,1496,1522]" italics="true" pageId="28" pageNumber="85">Maladera deenstana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2474E56FF973064FE76E9904E6010AF" box="[446,540,1497,1523]" pageId="28" pageNumber="85" rank="species">sp. nov.</taxonomicNameLabel>
is in external shape and shape of aedeagus similar to
<taxonomicName id="4C0054BCFF973064FB74E9904D07134A" authority="(Brenske, 1898)" baseAuthorityName="Brenske" baseAuthorityYear="1898" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="85" phylum="Arthropoda" rank="species" species="weligamana">
<emphasis id="B974F32DFF973064FB74E990490A10AE" box="[1212,1398,1496,1522]" italics="true" pageId="28" pageNumber="85">M. weligamana</emphasis>
(Brenske, 1898)
</taxonomicName>
. The new species differs from
<taxonomicName id="4C0054BCFF973064FD1BE9B54FFB1349" baseAuthorityName="Brenske" baseAuthorityYear="1898" box="[723,903,1531,1557]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="85" phylum="Arthropoda" rank="species" species="weligamana">
<emphasis id="B974F32DFF973064FD1BE9B54FFB1349" box="[723,903,1531,1557]" italics="true" pageId="28" pageNumber="85">M. weligamana</emphasis>
</taxonomicName>
by the mesally more compressed aedeagus and having the distal part less narrowed towards apex (all lateral view); the apical part is less narrowed a quarter before apex than in
<taxonomicName id="4C0054BCFF973064FDDDEA0A4EB71300" baseAuthorityName="Brenske" baseAuthorityYear="1898" box="[533,715,1602,1628]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="85" phylum="Arthropoda" rank="species" species="weligamana">
<emphasis id="B974F32DFF973064FDDDEA0A4EB71300" box="[533,715,1602,1628]" italics="true" pageId="28" pageNumber="85">M. weligamana</emphasis>
</taxonomicName>
(dorsal view).
</paragraph>
</subSubSection>
<subSubSection id="C31A7CB4FF973064FF75EACA494B1391" box="[189,1335,1667,1741]" pageId="28" pageNumber="85" type="etymology">
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<heading id="D0F79853FF973064FF75EACA4D3813C1" bold="true" box="[189,324,1667,1693]" fontSize="11" level="3" pageId="28" pageNumber="85" reason="3">
<emphasis id="B974F32DFF973064FF75EACA4D3813C1" bold="true" box="[189,324,1667,1693]" pageId="28" pageNumber="85">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8BBF2F3FFF973064FF75EAFB494B1391" blockId="28.[189,1335,1714,1741]" box="[189,1335,1714,1741]" pageId="28" pageNumber="85">
The new species is named after its
<typeStatus id="54BB919DFF973064FD9FEAFB4EF61390" box="[599,650,1714,1740]" pageId="28" pageNumber="85">type</typeStatus>
locality Deenston (adjective in the nominative singular).
</paragraph>
</subSubSection>
<subSubSection id="C31A7CB4FF973064FF75EABA48E612DF" pageId="28" pageNumber="85" type="materials_examined">
<paragraph id="8BBF2F3FFF973064FF75EABA4D101251" blockId="28.[189,364,1779,1805]" box="[189,364,1779,1805]" pageId="28" pageNumber="85">
<heading id="D0F79853FF973064FF75EABA4D101251" bold="true" box="[189,364,1779,1805]" fontSize="11" level="3" pageId="28" pageNumber="85" reason="3">
<emphasis id="B974F32DFF973064FF75EABA4D101251" bold="true" box="[189,364,1779,1805]" pageId="28" pageNumber="85">Type material</emphasis>
</heading>
</paragraph>
<paragraph id="8BBF2F3FFF973064FF22EB6B48E612DF" blockId="28.[189,1399,1826,1923]" pageId="28" pageNumber="85">
<materialsCitation id="3B682562FF973064FF22EB6B48E612DF" ID-GBIF-Occurrence="4003031386" collectingDate="2019-02-20" collectingMethod="Black light" collectionCode="ZFMK" collectorName="Eberle &amp; Ranasinghe" country="Sri Lanka" county="Knuckles South" elevation="1108" latitude="7.330824" location="Sri Lanka" longLatPrecision="1" longitude="80.86203" municipality="Deenston" pageId="28" pageNumber="85" specimenCode="X-SR0187" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<emphasis id="B974F32DFF973064FF22EB6B4D201260" bold="true" box="[234,348,1826,1852]" pageId="28" pageNumber="85">
<typeStatus id="54BB919DFF973064FF22EB6B4D201260" box="[234,348,1826,1852]" pageId="28" pageNumber="85" type="holotype">Holotype</typeStatus>
</emphasis>
<collectingCountry id="F3176FAFFF973064FF75EB0C4D271203" box="[189,347,1861,1887]" name="Sri Lanka" pageId="28" pageNumber="85">SRI LANKA</collectingCountry>
<specimenCount id="9D06E4B6FF973064FEB4EB0F4DE9123C" box="[380,405,1862,1888]" pageId="28" pageNumber="85" type="male"></specimenCount>
; “
<specimenCode id="DBA68744FF973064FE67EB0C4E4F1203" box="[431,563,1861,1887]" pageId="28" pageNumber="85">X-SR0187</specimenCode>
,
<location id="8EDF79E4FF973064FD8AEB0C4EC01203" LSID="urn:lsid:plazi:treatment:03A99E29FF973067FDA5E9684D3A1332:8EDF79E4FF973064FD8AEB0C4EC01203" box="[578,700,1861,1887]" country="Sri Lanka" county="Knuckles South" latitude="7.330824" longLatPrecision="1" longitude="80.86203" municipality="Deenston" name="Sri Lanka" pageId="28" pageNumber="85">Sri Lanka</location>
,
<location id="8EDF79E4FF973064FD04EB0C4FFC1203" LSID="urn:lsid:plazi:treatment:03A99E29FF973067FDA5E9684D3A1332:8EDF79E4FF973064FD04EB0C4FFC1203" box="[716,896,1861,1887]" country="Sri Lanka" county="Knuckles South" latitude="7.330824" longLatPrecision="1" longitude="80.86203" municipality="Deenston" name="Kandy District" pageId="28" pageNumber="85">Kandy District</location>
,
<collectingMunicipality id="6BDBB545FF973064FC58EB0F487E123C" box="[912,1026,1862,1888]" pageId="28" pageNumber="85">Deenston</collectingMunicipality>
,
<collectingCounty id="62DE57B3FF973064FBD9EB0C48AF1203" box="[1041,1235,1861,1887]" pageId="28" pageNumber="85">Knuckles South</collectingCounty>
;
<geoCoordinate id="EE3449F8FF973064FB2BEB0C490E1203" box="[1251,1394,1861,1887]" degrees="7.330824" direction="north" orientation="latitude" pageId="28" pageNumber="85" precision="1" value="7.330824">7.330824ºN</geoCoordinate>
,
<geoCoordinate id="EE3449F8FF973064FF75EB214D2412DE" box="[189,344,1896,1922]" degrees="80.862032" direction="east" orientation="longitude" pageId="28" pageNumber="85" precision="1" value="80.86203">80.862032ºE</geoCoordinate>
;
<quantity id="4CF882DAFF973064FEACEB214DC512DE" box="[356,441,1896,1922]" metricMagnitude="3" metricUnit="m" metricValue="1.108" pageId="28" pageNumber="85" unit="m" value="1108.0">
<elevation id="002DC80CFF973064FEACEB214DC512DE" box="[356,441,1896,1922]" metricMagnitude="3" metricUnit="m" metricValue="1.108" pageId="28" pageNumber="85" unit="m" value="1108.0">1108m</elevation>
</quantity>
;
<date id="FFBE09FFFF973064FE0CEB214E3A12DE" box="[452,582,1896,1922]" pageId="28" pageNumber="85" value="2019-02-20">
<collectingDate id="EFFAF017FF973064FE0CEB214E3A12DE" box="[452,582,1896,1922]" pageId="28" pageNumber="85" value="2019-02-20">20-II-2019</collectingDate>
</date>
;
<collectorName id="26F54AE9FF973064FD9BEB214EDC12DE" box="[595,672,1896,1922]" pageId="28" pageNumber="85">Eberle</collectorName>
and
<collectorName id="26F54AE9FF973064FD10EB214F1D12DE" box="[728,865,1896,1922]" pageId="28" pageNumber="85">Ranasinghe</collectorName>
leg.;
<collectingMethod id="52415728FF973064FC6BEB21485712DE" box="[931,1067,1896,1922]" pageId="28" pageNumber="85">Black light</collectingMethod>
”;
<collectionCode id="ED11B7FAFF973064FB89EB2048EA12DF" box="[1089,1174,1897,1923]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="28" pageNumber="85" type="Museum">ZFMK</collectionCode>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C31A7CB4FF963067FF75ED414D4510A1" lastPageId="31" lastPageNumber="88" pageId="29" pageNumber="86" type="description">
<paragraph id="8BBF2F3FFF963065FF75ED414D32147E" blockId="29.[189,334,264,290]" box="[189,334,264,290]" pageId="29" pageNumber="86">
<heading id="D0F79853FF963065FF75ED414D32147E" bold="true" box="[189,334,264,290]" fontSize="11" level="3" pageId="29" pageNumber="86" reason="3">
<emphasis id="B974F32DFF963065FF75ED414D32147E" bold="true" box="[189,334,264,290]" pageId="29" pageNumber="86">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8BBF2F3FFF963065FF75ED71485B140E" blockId="29.[189,1063,311,338]" box="[189,1063,311,338]" pageId="29" pageNumber="86">
MEASUREMENTS. Length:
<quantity id="4CF882DAFF963065FE28ED7E4E41140E" box="[480,573,311,338]" metricMagnitude="-3" metricUnit="m" metricValue="8.2" pageId="29" pageNumber="86" unit="mm" value="8.2">8.2 mm</quantity>
, length of elytra:
<quantity id="4CF882DAFF963065FCC3ED7E4F14140E" box="[779,872,311,338]" metricMagnitude="-3" metricUnit="m" metricValue="6.1" pageId="29" pageNumber="86" unit="mm" value="6.1">6.1 mm</quantity>
, width:
<quantity id="4CF882DAFF963065FC0DED7E485E140E" box="[965,1058,311,338]" metricMagnitude="-3" metricUnit="m" metricValue="4.9" pageId="29" pageNumber="86" unit="mm" value="4.9">4.9 mm</quantity>
.
</paragraph>
<paragraph id="8BBF2F3FFF963065FF75ED304F1414EA" blockId="29.[187,1399,376,438]" pageId="29" pageNumber="86">
HABITUS (
<figureCitation id="133B33BAFF963065FEE5ED314DFB14CF" box="[301,391,376,403]" captionStart="Fig" captionStartId="27.[189,232,1921,1947]" captionTargetBox="[204,1388,789,1860]" captionTargetId="figure-195@27.[189,1398,770,1883]" captionTargetPageId="27" captionText="Fig. 6. AD. Maladera dambullana sp. nov., holotype, ♂ (ZFMK). EH. M. deenstana sp. nov., holotype, ♂ (ZFMK). A, E. Aedeagus, left side lateral view. C, G. Aedeagus, right side lateral view. B, F. Parameres, dorsal view. D, H. Habitus (not to scale). Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6595481" httpUri="https://zenodo.org/record/6595481/files/figure.png" pageId="29" pageNumber="86">Fig. 6H</figureCitation>
) AND COLOURATION. Body oval, dark brown, antenna yellow, dorsal surface dull, densely minutely setose, elytra with moderately dense, short setae.
</paragraph>
<paragraph id="8BBF2F3FFF963065FF75ED974D50160B" blockId="29.[189,1401,477,855]" pageId="29" pageNumber="86">HEAD. Labroclypeus subtrapezoidal, distinctly wider than long, widest at base, lateral margins weakly convex and strongly convergent to widely rounded anterior angles, lateral border and ocular canthus producing an indistinct blunt angle, margins moderately reflexed, anteriorly distinctly emarginate medially; surface shiny, convexly elevated medially, finely and densely punctate, distance between punctures equal their diameter, with a few erect setae; frontoclypeal suture fine and weakly angled medially; smooth area in front of eye approximately twice as wide as long; ocular canthus short and narrow, finely and densely punctate, with a single short terminal seta. Frons dull, with fine, irregularly sparse punctures, with a few longer, adpressed setae on disc and beside eyes. Eyes moderately large, ratio of diameter/interocular width: 0.66. Antenna yellow, with ten antennomeres; club with three antennomeres, little shorter than remaining antennomeres combined. Mentum elevated and anteriorly flattened.</paragraph>
<paragraph id="8BBF2F3FFF963067FF75EF3648591435" blockId="29.[189,1399,894,991]" lastBlockId="31.[188,1399,265,362]" lastPageId="31" lastPageNumber="88" pageId="29" pageNumber="86">PRONOTUM. Moderately wide, widest at base, lateral margins weakly convex and evenly narrowed anteriorly, anterior angles moderately produced and moderately sharp, anterior marginal line fine and complete, anterior margin weakly produced medially; surface finely and densely punctate, with minute setae in punctures, otherwise glabrous; anterior and lateral borders sparsely setose, basal margin without marginal line; hypomeron ventrobasally carinate and slightly produced ventrally. Scutellum short and wide, triangular, with fine and dense punctures, along midline impunctate.</paragraph>
<caption id="DF7F7FB7FF963065FF75EB34490B1282" ID-DOI="http://doi.org/10.5281/zenodo.6595483" ID-Zenodo-Dep="6595483" httpUri="https://zenodo.org/record/6595483/files/figure.png" pageId="29" pageNumber="86" startId="29.[189,232,1917,1943]" targetBox="[189,1398,1076,1874]" targetPageId="29">
<paragraph id="8BBF2F3FFF963065FF75EB34490B1282" blockId="29.[189,1399,1917,2014]" pageId="29" pageNumber="86">
<emphasis id="B974F32DFF963065FF75EB344D7212C4" bold="true" box="[189,270,1917,1944]" pageId="29" pageNumber="86">Fig. 7.</emphasis>
Distribution of ten new species.
<emphasis id="B974F32DFF963065FD6BEB344EC412CB" bold="true" box="[675,696,1917,1943]" pageId="29" pageNumber="86">A</emphasis>
.
<taxonomicName id="4C0054BCFF963065FD02EB344FCB12CB" authority="Ranasinghe &amp; Eberle &amp; Athukorala &amp; Benjamin &amp; Ahrens, 2022" authorityName="Ranasinghe &amp; Eberle &amp; Athukorala &amp; Benjamin &amp; Ahrens" authorityYear="2022" box="[714,951,1917,1943]" class="Insecta" family="Melolonthidae" genus="Selaserica" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="86" phylum="Arthropoda" rank="species" species="fabriziae" status="sp. nov.">
<emphasis id="B974F32DFF963065FD02EB344FCB12CB" box="[714,951,1917,1943]" italics="true" pageId="29" pageNumber="86">Selaserica fabriziae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2474E56FF963065FC0AEB37486312C4" box="[962,1055,1918,1944]" pageId="29" pageNumber="86" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="B974F32DFF963065FBE1EB37484112C4" bold="true" box="[1065,1085,1918,1944]" pageId="29" pageNumber="86">B</emphasis>
.
<taxonomicName id="4C0054BCFF963065FB86EB34497312CB" authority="Ranasinghe &amp; Eberle &amp; Athukorala &amp; Benjamin &amp; Ahrens, 2022" authorityName="Ranasinghe &amp; Eberle &amp; Athukorala &amp; Benjamin &amp; Ahrens" authorityYear="2022" box="[1102,1295,1917,1943]" class="Insecta" family="Melolonthidae" genus="Selaserica" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="86" phylum="Arthropoda" rank="species" species="sororinitida" status="sp. nov.">
<emphasis id="B974F32DFF963065FB86EB34497312CB" box="[1102,1295,1917,1943]" italics="true" pageId="29" pageNumber="86">Sel. sororinitida</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2474E56FF963065FAD1EB37490B12C4" box="[1305,1399,1918,1944]" pageId="29" pageNumber="86" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="B974F32DFF963065FF75EBE94CAE12E6" bold="true" box="[189,210,1952,1978]" pageId="29" pageNumber="86">C</emphasis>
.
<taxonomicName id="4C0054BCFF963065FF2AEBE84DB512E7" authority="Ranasinghe &amp; Eberle &amp; Athukorala &amp; Benjamin &amp; Ahrens, 2022" authorityName="Ranasinghe &amp; Eberle &amp; Athukorala &amp; Benjamin &amp; Ahrens" authorityYear="2022" box="[226,457,1953,1979]" class="Insecta" family="Melolonthidae" genus="Neoserica" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="86" phylum="Arthropoda" rank="species" species="pophami" status="sp. nov.">
<emphasis id="B974F32DFF963065FF2AEBE84DB512E7" box="[226,457,1953,1979]" italics="true" pageId="29" pageNumber="86">Neoserica pophami</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2474E56FF963065FE1AEBE84E5212E7" box="[466,558,1953,1979]" pageId="29" pageNumber="86" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="B974F32DFF963065FDF0EBE84E3012E7" bold="true" box="[568,588,1953,1979]" pageId="29" pageNumber="86">D</emphasis>
.
<taxonomicName id="4C0054BCFF963065FD94EBE84F5E12E7" authority="Ranasinghe &amp; Eberle &amp; Athukorala &amp; Benjamin &amp; Ahrens, 2022" authorityName="Ranasinghe &amp; Eberle &amp; Athukorala &amp; Benjamin &amp; Ahrens" authorityYear="2022" box="[604,802,1953,1979]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="86" phylum="Arthropoda" rank="species" species="haniel" status="sp. nov.">
<emphasis id="B974F32DFF963065FD94EBE84F5E12E7" box="[604,802,1953,1979]" italics="true" pageId="29" pageNumber="86">Maladera haniel</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2474E56FF963065FCE3EBE84FFB12E7" box="[811,903,1953,1979]" pageId="29" pageNumber="86" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="B974F32DFF963065FC59EBE84FDF12E7" bold="true" box="[913,931,1953,1979]" pageId="29" pageNumber="86">E</emphasis>
.
<taxonomicName id="4C0054BCFF963065FC7BEBE8486F12E7" authority="Ranasinghe &amp; Eberle &amp; Athukorala &amp; Benjamin &amp; Ahrens, 2022" authorityName="Ranasinghe &amp; Eberle &amp; Athukorala &amp; Benjamin &amp; Ahrens" authorityYear="2022" box="[947,1043,1953,1979]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="86" phylum="Arthropoda" rank="species" species="kishi" status="sp. nov.">
<emphasis id="B974F32DFF963065FC7BEBE8486F12E7" box="[947,1043,1953,1979]" italics="true" pageId="29" pageNumber="86">M. kishi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2474E56FF963065FBD4EBE8480412E7" box="[1052,1144,1953,1979]" pageId="29" pageNumber="86" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="B974F32DFF963065FB4AEBE848EE12E7" bold="true" box="[1154,1170,1953,1979]" pageId="29" pageNumber="86">F</emphasis>
.
<taxonomicName id="4C0054BCFF963065FB6BEBE8496E12E7" authority="Ranasinghe &amp; Eberle &amp; Athukorala &amp; Benjamin &amp; Ahrens, 2022" authorityName="Ranasinghe &amp; Eberle &amp; Athukorala &amp; Benjamin &amp; Ahrens" authorityYear="2022" box="[1187,1298,1953,1979]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="86" phylum="Arthropoda" rank="species" species="windy" status="sp. nov.">
<emphasis id="B974F32DFF963065FB6BEBE8496E12E7" box="[1187,1298,1953,1979]" italics="true" pageId="29" pageNumber="86">M. windy</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2474E56FF963065FAD3EBE8490B12E7" box="[1307,1399,1953,1979]" pageId="29" pageNumber="86" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="B974F32DFF963065FF77EB8A4CA81281" bold="true" box="[191,212,1987,2013]" pageId="29" pageNumber="86">G</emphasis>
.
<taxonomicName id="4C0054BCFF963065FF29EB8C4DDB1282" authority="Ranasinghe &amp; Eberle &amp; Athukorala &amp; Benjamin &amp; Ahrens, 2022" authorityName="Ranasinghe &amp; Eberle &amp; Athukorala &amp; Benjamin &amp; Ahrens" authorityYear="2022" box="[225,423,1988,2014]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="86" phylum="Arthropoda" rank="species" species="karunaratnae" status="sp. nov.">
<emphasis id="B974F32DFF963065FF29EB8C4DDB1282" box="[225,423,1988,2014]" italics="true" pageId="29" pageNumber="86">M. karunaratnae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2474E56FF963065FE64EB8D4E791282" box="[428,517,1988,2014]" pageId="29" pageNumber="86" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="B974F32DFF963065FDC1EB8D4E5C1282" bold="true" box="[521,544,1988,2014]" pageId="29" pageNumber="86">H</emphasis>
.
<taxonomicName id="4C0054BCFF963065FDE7EB8C4EB71282" authority="Ranasinghe &amp; Eberle &amp; Athukorala &amp; Benjamin &amp; Ahrens, 2022" authorityName="Ranasinghe &amp; Eberle &amp; Athukorala &amp; Benjamin &amp; Ahrens" authorityYear="2022" box="[559,715,1988,2014]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="86" phylum="Arthropoda" rank="species" species="hiyarensis" status="sp. nov.">
<emphasis id="B974F32DFF963065FDE7EB8C4EB71282" box="[559,715,1988,2014]" italics="true" pageId="29" pageNumber="86">M. hiyarensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2474E56FF963065FD1AEB8D4F571282" box="[722,811,1988,2014]" pageId="29" pageNumber="86" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="B974F32DFF963065FCE7EB8D4F461282" bold="true" box="[815,826,1988,2014]" pageId="29" pageNumber="86">I</emphasis>
.
<taxonomicName id="4C0054BCFF963065FC81EB8C4F871282" authority="Ranasinghe &amp; Eberle &amp; Athukorala &amp; Benjamin &amp; Ahrens, 2022" authorityName="Ranasinghe &amp; Eberle &amp; Athukorala &amp; Benjamin &amp; Ahrens" authorityYear="2022" box="[841,1019,1988,2014]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="86" phylum="Arthropoda" rank="species" species="dambullana" status="sp. nov.">
<emphasis id="B974F32DFF963065FC81EB8C4F871282" box="[841,1019,1988,2014]" italics="true" pageId="29" pageNumber="86">M. dambullana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2474E56FF963065FBC9EB8D48261282" box="[1025,1114,1988,2014]" pageId="29" pageNumber="86" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="B974F32DFF963065FB96EB8D48081282" bold="true" box="[1118,1140,1988,2014]" pageId="29" pageNumber="86">J.</emphasis>
<taxonomicName id="4C0054BCFF963065FBB3EB8C49641282" authority="Ranasinghe &amp; Eberle &amp; Athukorala &amp; Benjamin &amp; Ahrens, 2022" authorityName="Ranasinghe &amp; Eberle &amp; Athukorala &amp; Benjamin &amp; Ahrens" authorityYear="2022" box="[1147,1304,1988,2014]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="86" phylum="Arthropoda" rank="species" species="deenstana" status="sp. nov.">
<emphasis id="B974F32DFF963065FBB3EB8C49641282" box="[1147,1304,1988,2014]" italics="true" pageId="29" pageNumber="86">M. deenstana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2474E56FF963065FAD6EB8D490B1282" box="[1310,1399,1988,2014]" pageId="29" pageNumber="86" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</caption>
<caption id="DF7F7FB7FF953066FF75EB144D6412BE" ID-DOI="http://doi.org/10.5281/zenodo.6595485" ID-Zenodo-Dep="6595485" httpUri="https://zenodo.org/record/6595485/files/figure.png" pageId="30" pageNumber="87" startId="30.[189,232,1885,1911]" targetBox="[235,1353,265,1847]" targetPageId="30">
<paragraph id="8BBF2F3FFF953066FF75EB144D6412BE" blockId="30.[189,1398,1885,2018]" pageId="30" pageNumber="87">
<emphasis id="B974F32DFF953066FF75EB144D75122B" bold="true" box="[189,265,1885,1911]" pageId="30" pageNumber="87">Fig. 8.</emphasis>
Photographs of the habitats of the new species.
<emphasis id="B974F32DFF953066FCE4EB144F3D122B" bold="true" box="[812,833,1885,1911]" pageId="30" pageNumber="87">A</emphasis>
.
<taxonomicName id="4C0054BCFF953066FC87EB17484B1224" authority="Ranasinghe &amp; Eberle &amp; Athukorala &amp; Benjamin &amp; Ahrens, 2022" authorityName="Ranasinghe &amp; Eberle &amp; Athukorala &amp; Benjamin &amp; Ahrens" authorityYear="2022" box="[847,1079,1886,1912]" class="Insecta" family="Melolonthidae" genus="Selaserica" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="87" phylum="Arthropoda" rank="species" species="fabriziae" status="sp. nov.">
<emphasis id="B974F32DFF953066FC87EB17484B1224" box="[847,1079,1886,1912]" italics="true" pageId="30" pageNumber="87">Selaserica fabriziae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2474E56FF953066FBF5EB1748E91224" box="[1085,1173,1886,1912]" pageId="30" pageNumber="87" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="B974F32DFF953066FB51EB1748D11224" bold="true" box="[1177,1197,1886,1912]" pageId="30" pageNumber="87">B</emphasis>
.
<taxonomicName id="4C0054BCFF953066FB73EB17490A1224" authority="Ranasinghe &amp; Eberle &amp; Athukorala &amp; Benjamin &amp; Ahrens, 2022" authorityName="Ranasinghe &amp; Eberle &amp; Athukorala &amp; Benjamin &amp; Ahrens" authorityYear="2022" box="[1211,1398,1886,1912]" class="Insecta" family="Melolonthidae" genus="Selaserica" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="87" phylum="Arthropoda" rank="species" species="sororinitida" status="sp. nov.">
<emphasis id="B974F32DFF953066FB73EB17490A1224" box="[1211,1398,1886,1912]" italics="true" pageId="30" pageNumber="87">Sel. sororinitida</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2474E56FF953066FF75EBC84D6412C7" box="[189,280,1921,1947]" pageId="30" pageNumber="87" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="B974F32DFF953066FEE9EBC84D4A12C7" bold="true" box="[289,310,1921,1947]" pageId="30" pageNumber="87">C</emphasis>
.
<taxonomicName id="4C0054BCFF953066FE8CEBCB4E5612C7" authority="Ranasinghe &amp; Eberle &amp; Athukorala &amp; Benjamin &amp; Ahrens, 2022" authorityName="Ranasinghe &amp; Eberle &amp; Athukorala &amp; Benjamin &amp; Ahrens" authorityYear="2022" box="[324,554,1921,1947]" class="Insecta" family="Melolonthidae" genus="Neoserica" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="87" phylum="Arthropoda" rank="species" species="pophami" status="sp. nov.">
<emphasis id="B974F32DFF953066FE8CEBCB4E5612C7" box="[324,554,1921,1947]" italics="true" pageId="30" pageNumber="87">Neoserica pophami</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2474E56FF953066FDFAEBC84EF112C7" box="[562,653,1921,1947]" pageId="30" pageNumber="87" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="B974F32DFF953066FD5EEBC84ED712C7" bold="true" box="[662,683,1921,1947]" pageId="30" pageNumber="87">D</emphasis>
.
<taxonomicName id="4C0054BCFF953066FD71EBC84F0212C7" authority="Ranasinghe &amp; Eberle &amp; Athukorala &amp; Benjamin &amp; Ahrens, 2022" authorityName="Ranasinghe &amp; Eberle &amp; Athukorala &amp; Benjamin &amp; Ahrens" authorityYear="2022" box="[697,894,1921,1947]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="87" phylum="Arthropoda" rank="species" species="haniel" status="sp. nov.">
<emphasis id="B974F32DFF953066FD71EBC84F0212C7" box="[697,894,1921,1947]" italics="true" pageId="30" pageNumber="87">Maladera haniel</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2474E56FF953066FC4EEBC84F9E12C7" box="[902,994,1921,1947]" pageId="30" pageNumber="87" rank="species">sp. nov.</taxonomicNameLabel>
and
<taxonomicName id="4C0054BCFF953066FBD4EBCB480712C7" authority="Ranasinghe &amp; Eberle &amp; Athukorala &amp; Benjamin &amp; Ahrens, 2022" authorityName="Ranasinghe &amp; Eberle &amp; Athukorala &amp; Benjamin &amp; Ahrens" authorityYear="2022" box="[1052,1147,1921,1947]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="87" phylum="Arthropoda" rank="species" species="kishi" status="sp. nov.">
<emphasis id="B974F32DFF953066FBD4EBCB480712C7" box="[1052,1147,1921,1947]" italics="true" pageId="30" pageNumber="87">M. kishi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2474E56FF953066FB4BEBC848A212C7" box="[1155,1246,1921,1947]" pageId="30" pageNumber="87" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="B974F32DFF953066FB2EEBC8488512C7" bold="true" box="[1254,1273,1921,1947]" pageId="30" pageNumber="87">E</emphasis>
.
<taxonomicName id="4C0054BCFF953066FAC1EBCB490A12C7" authority="Ranasinghe &amp; Eberle &amp; Athukorala &amp; Benjamin &amp; Ahrens, 2022" authorityName="Ranasinghe &amp; Eberle &amp; Athukorala &amp; Benjamin &amp; Ahrens" authorityYear="2022" box="[1289,1398,1921,1947]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="87" phylum="Arthropoda" rank="species" species="windy" status="sp. nov.">
<emphasis id="B974F32DFF953066FAC1EBCB490A12C7" box="[1289,1398,1921,1947]" italics="true" pageId="30" pageNumber="87">M. windy</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2474E56FF953066FF75EBEC4D6A12E3" box="[189,278,1957,1983]" pageId="30" pageNumber="87" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="B974F32DFF953066FED2EBED4D5012E2" bold="true" box="[282,300,1956,1982]" pageId="30" pageNumber="87">F</emphasis>
.
<taxonomicName id="4C0054BCFF953066FEF2EBEC4E7C12E2" authority="Ranasinghe &amp; Eberle &amp; Athukorala &amp; Benjamin &amp; Ahrens, 2022" authorityName="Ranasinghe &amp; Eberle &amp; Athukorala &amp; Benjamin &amp; Ahrens" authorityYear="2022" box="[314,512,1956,1982]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="87" phylum="Arthropoda" rank="species" species="karunaratnae" status="sp. nov.">
<emphasis id="B974F32DFF953066FEF2EBEC4E7C12E2" box="[314,512,1956,1982]" italics="true" pageId="30" pageNumber="87">M. karunaratnae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2474E56FF953066FDCDEBEC4E2212E3" box="[517,606,1957,1983]" pageId="30" pageNumber="87" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="B974F32DFF953066FDAAEBED4E0512E2" bold="true" box="[610,633,1956,1982]" pageId="30" pageNumber="87">G</emphasis>
.
<taxonomicName id="4C0054BCFF953066FD4FEBEC4F5812E2" authority="Ranasinghe &amp; Eberle &amp; Athukorala &amp; Benjamin &amp; Ahrens, 2022" authorityName="Ranasinghe &amp; Eberle &amp; Athukorala &amp; Benjamin &amp; Ahrens" authorityYear="2022" box="[647,804,1956,1982]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="87" phylum="Arthropoda" rank="species" species="hiyarensis" status="sp. nov.">
<emphasis id="B974F32DFF953066FD4FEBEC4F5812E2" box="[647,804,1956,1982]" italics="true" pageId="30" pageNumber="87">M. hiyarensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2474E56FF953066FCE2EBEC4FFF12E3" box="[810,899,1957,1983]" pageId="30" pageNumber="87" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="B974F32DFF953066FC4EEBED4FE112E2" bold="true" box="[902,925,1956,1982]" pageId="30" pageNumber="87">H</emphasis>
.
<taxonomicName id="4C0054BCFF953066FC64EBEC482112E2" authority="Ranasinghe &amp; Eberle &amp; Athukorala &amp; Benjamin &amp; Ahrens, 2022" authorityName="Ranasinghe &amp; Eberle &amp; Athukorala &amp; Benjamin &amp; Ahrens" authorityYear="2022" box="[940,1117,1956,1982]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="87" phylum="Arthropoda" rank="species" species="dambullana" status="sp. nov.">
<emphasis id="B974F32DFF953066FC64EBEC482112E2" box="[940,1117,1956,1982]" italics="true" pageId="30" pageNumber="87">M. dambullana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2474E56FF953066FBABEBEC48C012E3" box="[1123,1212,1957,1983]" pageId="30" pageNumber="87" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="B974F32DFF953066FB08EBED48B712E2" bold="true" box="[1216,1227,1956,1982]" pageId="30" pageNumber="87">I</emphasis>
.
<taxonomicName id="4C0054BCFF953066FB12EBEC490A12E2" authority="Ranasinghe &amp; Eberle &amp; Athukorala &amp; Benjamin &amp; Ahrens, 2022" authorityName="Ranasinghe &amp; Eberle &amp; Athukorala &amp; Benjamin &amp; Ahrens" authorityYear="2022" box="[1242,1398,1956,1982]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="87" phylum="Arthropoda" rank="species" species="deenstana" status="sp. nov.">
<emphasis id="B974F32DFF953066FB12EBEC490A12E2" box="[1242,1398,1956,1982]" italics="true" pageId="30" pageNumber="87">M. deenstana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2474E56FF953066FF75EB814D6412BE" box="[189,280,1992,2018]" pageId="30" pageNumber="87" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph id="8BBF2F3FFF943067FF75EDD84E581764" blockId="31.[189,1399,401,568]" pageId="31" pageNumber="88">ELYTRA. Short oval, widest at posterior third, striae indistinctly impressed, finely and densely punctate, intervals flat, with fine, dense punctures, with minute setae in punctures and a few moderately dense, short setae on lateral intervals and posterior half; epipleural edge fine, ending at the weakly convex external apical angle of elytra, epipleura densely setose; apical border membranous, apex covered with a rim of short microtrichomes.</paragraph>
<paragraph id="8BBF2F3FFF943067FF75EE294D351610" blockId="31.[188,1400,607,844]" pageId="31" pageNumber="88">
VENTRAL SURFACE. Moderately shiny, thorax and metacoxa with large and dense punctures, densely minutely setose, metacoxa with numerous long setae laterally; each abdominal sternite, in addition to evenly distributed fine and dense punctures bearing each a fine seta, with a distinct transverse row of coarse punctures each bearing a long and more robust seta, penultimate sternite apically with a narrow, shiny smooth chitinous border. Mesosternum between mesocoxae as wide as mesofemur, with a semicircular ridge bearing robust setae. Ratio of length of metepisternum/metacoxa: 1/2.0. Pygidium lacking in
<typeStatus id="54BB919DFF943067FF12EF7B4D391610" box="[218,325,818,844]" pageId="31" pageNumber="88" type="holotype">holotype</typeStatus>
.
</paragraph>
<paragraph id="8BBF2F3FFF943067FF75EF3D495F100B" blockId="31.[188,1400,884,1368]" pageId="31" pageNumber="88">
LEGS. Short and wide; femur with two longitudinal rows of setae, finely and densely punctate, densely setose; metafemur shiny, anterior edge acute, lacking an adjacent serrated line, ventral surface densely punctate and setose; posterior ventral margin straight, strongly widened in apical half and very indistinctly serrate apically; posterior dorsal margin not serrated, densely setose. Metatibia short, widest at middle, posteriorly slightly narrowed, ratio width/length: 1/2.9, dorsally sharply carinate, with two groups of spines, basal one shortly behind middle, apical one at four fifths of metatibial length, basally beside dorsal margin with two single punctures with serrated margins, each bearing single spines; lateral face longitudinal convex, with sparse, fine punctures and with minute setae in punctures; ventral margin finely serrate, with five strong spines equidistant from each other; medial face smooth, apex interiorly near tarsal articulation shallowly truncate. Meso- and metatarsomeres impunctate and glabrous dorsally, ventrally with sparse, short setae; metatarsomeres ventrally glabrous, with a strongly serrated ridge, beside which is a strong longitudinal carina; first metatarsomere as long as two following tarsomeres combined and as long as dorsal tibial spur. Protibia short, bidentate. All claws symmetrical, feebly curved and long, with normally developed basal tooth, distal protarsomeres lacking in
<typeStatus id="54BB919DFF943067FB7CE9744963100B" box="[1204,1311,1341,1367]" pageId="31" pageNumber="88" type="holotype">holotype</typeStatus>
.
</paragraph>
<paragraph id="8BBF2F3FFF943067FF75E9364DB810C5" blockId="31.[189,452,1407,1433]" box="[189,452,1407,1433]" pageId="31" pageNumber="88">
AEDEAGUS.
<figureCitation id="133B33BAFF943067FE8AE9364DBC10C5" box="[322,448,1407,1433]" captionStart="Fig" captionStartId="27.[189,232,1921,1947]" captionTargetBox="[204,1388,789,1860]" captionTargetId="figure-195@27.[189,1398,770,1883]" captionTargetPageId="27" captionText="Fig. 6. AD. Maladera dambullana sp. nov., holotype, ♂ (ZFMK). EH. M. deenstana sp. nov., holotype, ♂ (ZFMK). A, E. Aedeagus, left side lateral view. C, G. Aedeagus, right side lateral view. B, F. Parameres, dorsal view. D, H. Habitus (not to scale). Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6595481" httpUri="https://zenodo.org/record/6595481/files/figure.png" pageId="31" pageNumber="88">Fig. 6EG</figureCitation>
.
</paragraph>
<paragraph id="8BBF2F3FFF943067FF22E9894D391086" blockId="31.[189,325,1472,1533]" box="[234,325,1472,1498]" pageId="31" pageNumber="88">
<emphasis id="B974F32DFF943067FF22E9894D391086" bold="true" box="[234,325,1472,1498]" pageId="31" pageNumber="88">Female</emphasis>
</paragraph>
<paragraph id="8BBF2F3FFF943067FF75E9AA4D4510A1" blockId="31.[189,325,1472,1533]" box="[189,313,1507,1533]" pageId="31" pageNumber="88">Unknown.</paragraph>
</subSubSection>
<subSubSection id="C31A7CB4FF943067FF75EA6D4D3A1332" box="[189,342,1572,1646]" pageId="31" pageNumber="88" type="distribution">
<paragraph id="8BBF2F3FFF943067FF75EA6D4D2A1362" blockId="31.[189,342,1572,1598]" box="[189,342,1572,1598]" pageId="31" pageNumber="88">
<heading id="D0F79853FF943067FF75EA6D4D2A1362" bold="true" box="[189,342,1572,1598]" fontSize="11" level="3" pageId="31" pageNumber="88" reason="3">
<emphasis id="B974F32DFF943067FF75EA6D4D2A1362" bold="true" box="[189,342,1572,1598]" pageId="31" pageNumber="88">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8BBF2F3FFF943067FF75EA1A4D3A1332" blockId="31.[189,326,1619,1646]" box="[189,326,1619,1646]" pageId="31" pageNumber="88">
See
<figureCitation id="133B33BAFF943067FF27EA1A4D3D1332" box="[239,321,1619,1646]" captionStart="Fig" captionStartId="29.[189,232,1917,1943]" captionTargetBox="[189,1398,1076,1874]" captionTargetId="figure-244@29.[189,1398,1076,1874]" captionTargetPageId="29" captionText="Fig. 7. Distribution of ten new species. A. Selaserica fabriziae sp. nov. B. Sel. sororinitida sp. nov. C. Neoserica pophami sp. nov. D. Maladera haniel sp. nov. E. M. kishi sp. nov. F. M. windy sp. nov. G. M. karunaratnae sp. nov. H. M. hiyarensis sp. nov. I. M. dambullana sp. nov. J. M. deenstana sp. nov." figureDoi="http://doi.org/10.5281/zenodo.6595483" httpUri="https://zenodo.org/record/6595483/files/figure.png" pageId="31" pageNumber="88">Fig. 7J</figureCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>